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Species: Nesomyrmex angulatus   (Mayr, 1862) 

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Nesomyrmex angulatus lybica

Taxonomic History (provided by Barry Bolton, 2020)

Extant: 1 valid subspecies

Leptothorax angulatus Mayr, 1862 PDF: 739 (w.) EGYPT. Palearctic. AntCat AntWiki HOL

Taxonomic history

Santschi, 1914b: 107 (q.).
Combination in Nesomyrmex (Goniothorax): Emery, 1896g PDF: 58 (footnote); in Nesomyrmex: Bolton, 2003 PDF: 272.
Current subspecies: nominal plus N. a. lybica.


  Geographic regions (According to curated Geolocale/Taxon lists):
    Africa: Comoros, Egypt, Ethiopia, Kenya, Madagascar, Mayotte, Mozambique, Senegal, South Africa, Tanzania, Tunisia, Zimbabwe
    Asia: Saudi Arabia
  Biogeographic regions (According to curated Bioregion/Taxon lists):
    Afrotropical, Malagasy, Palearctic

Distribution Notes:

Reference for Kenya if not type: Bolton 1982


Caste of types: (w)

Taxonomic Treatment (provided by Plazi)

Treatment Citation: Mbanyana, N. & Robertson, H. G., 2008, Review of the ant genus Nesomyrmex (Hymenoptera: Formicidae: Myrmicinae) in southern Africa., African Natural History 4, pp. 35-55

Nesomyrmex angulatusHNS (Mayr, 1862) Leptothorax angulatus MayrHNS, 1862: 739. For type information and synonyms see Bolton (1982). There is some debate on the web ( nesomyrmex_ angulatusHNS /nesomyrmex_ angulatusHNS.htm - accessed 9 September 2008) over the validity of some of the synonyms but there is no attempt to resolve this here.

Fig. 1a -c

Description of worker

HL 0.672-0.811, HW 0.565-0.692, HW1 0.610-0.736, CI 78-87, SL 0.487-0.613, SI 84-92, PW 0.423-0.541, ML 0.785-1.087, EL 0.169-0.197, EI 28-30 (5 of 81 measured).

Description matches that of Bolton (1982) except for few additional characters. Sculpture on the dorsum of head is not only reticulate-punctulate but also with longitudinal striations, which are well defined near the inner margin of the eyes.


Nesomyrmex angulatusHNS is easily distinguished from other species in the angulatusHNS species-group by the mesosoma, which forms a single, uninterrupted flat surface without any trace of metanotal groove; in the other species, the promesonotum is separated from propodeum by metanotal groove or impression.


Nests in cavities of dead wood on trees in savanna habitats.

Material examined South Africa: KwaZulu-Natal: St Lucia, 05 August 1945, J.C. Faure, SAM-HYM-C004716; Mkuzi [= Mkuze], 16 May 1946, J.C. Faure, SAM-HYM-C004717; Mkuze Game Reserve, Mantuma Rest Camp, 27°36'S 32°13'E, collected from the nest, 26 February 1988, H.G. Robertson, SAM-HYM-C001007; 4.5 km SW of Golela, 27°20'03"S 31°50'09"E, 20 November 1996, Acacia niloticaHNS, F. Grobbelaar & I.M. Millar, SAM-HYM-C010657; Sordwana, 27°33'S 32°40'E, 20 May 1946, A.R.I., SAM-HYM-C011707; Pongola, [27°23'S 31°37'E], 13 July 1967, Dr Karney, SAM-HYM-C015108; Limpopo: Lapellala [= Lapalala], [23°53'S 20°28'E], ex foragers, M. Villet, SAM-HYM-C 000541; Letaba estate, 23° 51'S 30°18'E, Collected ex nest, 09 February 1988, H.G. Robertson, SAM-HYM-C000937; Nylsvley, 24°40'S 28°43'E, ex foragers, 16 April 1987, H.G. Robertson,SAM-HYM-C000481.Zimbabwe: Khami River, 30 November 1913, G. Arnold, SAM-HYM-C004711; Sawmills , 31 October 1919, SAM-HYM-C004712; Victoria Falls, December 1938, Nat. Museum S. Rhodesia [= National Museum of Southern Rhodesia], SAM-HYM-C004715; Melsetter, 03 October 1950, Nat. Museum S. Rhodesia [= National Museum of Southern Rhodesia], SAM-HYM-C 004718; Zambesi valley,7 kmSEofAngwa Bridge, 16°08'S 30°15'E, 18 June -09 September 1988, J. Weyrich, SAM-HYM-C005668; Kenya: Segera Ranch, Laikipia Plateau , 0°15'N 36°50'E, 25 February 1994, collected in Acacia woodland on black cotton soils, J. Pruetz, SAM-HYM-C 017849; Tanzania: Mkomazi Game Reserve, Pangaro, 3°54'37"S 37°47'05"E, 09 December 1995, Foragers on dead wood of Acacia tree. Open woodland, H.G. Robertson, SAM-HYM-C008655.

Specimen Habitat Summary

Found most commonly in these habitats: 6 times found in miombo woodland, 6 times found in dry forest, 4 times found in coastal thicket, 6 times found in coastal scrub, 7 times found in mangrove, coastal scrub, 3 times found in forest on inselberg, 3 times found in coastal dry forest scrub, 1 times found in beach dune veg., closed times found in woodland, 2 times found in Open Miombo Woodland, ...

Found most commonly in these microhabitats: 12 times ex dead twig above ground, 24 times on low vegetation, 2 times ex dead bamboo, 1 times UV Black light, 1 times nest in dead stem, 2 times ground forager(s), 1 times Ex dead limb stub of Acacia etbaica, 1 times ex dead branch above ground, 1 times ex dead twig, 1 times arboreal foragers, 1 times Arboreal, ...

Collected most commonly using these methods: 3 times beating low vegetation, 4 times Hand collected, 2 times search, 3 times Dead wood, 3 times Pitfall trap, 2 times beating vegetation (2 hours), 1 times beating Acacia, 1 times By Hand, 1 times Hand.

Elevations: collected from 1 - 1892 meters, 197 meters average

Collect Date Range: collected between 1911-10-01 and 2019-09-16

Type specimens: Holotype of Leptothorax latinodis: casent0914925; Lectotype of Leptothorax angulatus: casent0914922; syntype of Leptothorax angulatus concolor: casent0912893; syntype of Leptothorax angulatus ilgii: casent0908991

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