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Fossil: 1 valid subspecies
Radoboj, Croatia; in Early Miocene (Burdigalian, 16-20.4 Ma) micritic limestone.
Lectotype UMJG-P 77.499 and other specimens (UMJG-P 77.543, 77.644b, 77.565) in Universalmuseum Joanneum, Geology and Paleontology, Graz, Austria; other material in Geologische Bundesanstalt and in Naturhistorisches Museum, Vienna, Austria.
*One specimen of subspecies parschlugianus in Paleontology Department of the Museum National d'Histoire Naturelle, Paris, France.
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