subfamily tribe genus species species author date country valid available current valid name original combination fossil taxonomic history Formicinae Camponotini Camponotus irritans melanogaster TRUE TRUE FALSE

melanogaster. Camponotus (Tanaemyrmex) irritans var. melanogaster Stitz, 1938: 114 (q.) MICRONESIA (Caroline Is).

Myrmicinae Formicoxenini Romblonella yapensis TRUE TRUE FALSE

yapensis. Romblonella yapensis Smith, M.R. 1953a: 78 (w.) MICRONESIA (Caroline Is).

Formicinae Camponotini Camponotus TRUE TRUE FALSE

Extant: 1,058 species, 495 subspecies

Fossil: 31 species

Camponotus Mayr, 1861: 35. Type-species: Formica ligniperda, by subsequent designation of Bingham, 1903: 347.

Taxonomic history

[Type-species not Formica herculeana, unjustified subsequent designation by Forel, 1914a: 259, repeated in Arnold, 1922: 612.]

Camponotus in Formicinae: Mayr, 1861: 35 [Formicidae]; Mayr, 1862: 652 (in key) [Formicidae]; Mayr, 1865: 6 [Formicidae]; Mayr, 1868c: 26 [Formicidae].

Camponotus in Camponotinae: Forel, 1878: 367 [Camponotidae]; Emery & Forel, 1879a: 447 [Camponotidae]; Forel, 1892j: 223 [Camponotidae]; Dalla Torre, 1893: 171.

Camponotus in Camponotinae, Camponotini: Forel, 1886f: 141; Forel, 1893a: 165; Forel, 1895b: 101; Emery, 1895j: 772; Forel, 1899c: 130; Ashmead, 1905b: 384; Wheeler, W.M. 1910g: 144; Forel, 1912i: 89; Wheeler, W.M. 1915h: 138; Forel, 1917: 250; Arnold, 1922: 610.

Camponotus in Formicinae, Camponotini: Bondroit, 1918: 65; Wheeler, W.M. 1922a: 703; Emery, 1925b: 59; all subsequent authors except the following.

Camponotus in Formicinae, Formica genus group: Agosti, 1991: 295.

Camponotus as subgenus of Formica: Smith, F. 1871a: 306.

Camponotus as genus: all authors except the entry above.

Junior synonym of CAMPONOTUS

†Paleosminthurus Pierce & Gibron, 1962: 146. Type-species: †Paleosminthurus juliae, by monotypy.

Taxonomic history

†Paleosminthurus in Collembola, family †Paleosminthuridae: Pierce & Gibron, 1962: 146.

†Paleosminthurus incertae sedis in Formicidae: Najt, 1987: 152.

†Paleosminthurus incertae sedis in Formicinae: Hölldobler & Wilson, 1990: 18.

†Paleosminthurus in †Paleosmithurinae: Bolton, 1994: 187 [provisional subfamily status]; Bolton, 1995b: 14, 311; Bolton, 2003: 76, 263.

†Paleosminthurus as junior synonym of Camponotus: Snelling (pers. comm., 2004). [Synonymy remained unpublished at his death in 2008.]

†Shanwangella Zhang, 1989: 307. Type-species: †Shanwangella palaeoptera, by original designation.

Taxonomic history

†Shanwangella as junior synonym of Camponotus: Hong & Wu, 2000: 20.

Subgenera of CAMPONOTUS include the nominal plus the following.

All the subgenera were given as provisional junior synonyms of Camponotus by Brown, 1973b: 179 - 185. Brown's list was repeated in Hölldobler & Wilson, 1990: 18 but with all subgenera listed as junior synonyms; subgeneric status was reinstated in Bolton, 1994: 50. These consecutive entries are not repeated individually below.

Subgenus CAMPONOTUS (COLOBOPSIS)

Colobopsis Mayr, 1861: 38. Type-species: Formica truncata, by subsequent designation of Bingham, 1903: 342.

Taxonomic history

Colobopsis in Formicinae: Mayr, 1861: 38 [Formicidae]; Mayr, 1862: 652 (in key) [Formicidae]; Mayr, 1865: 7 [Formicidae].

Colobopsis in Camponotinae: Forel, 1878: 368 [Camponotidae]; Emery & Forel, 1897: 449 [Camponotidae].

Colobopsis in Camponotinae, Camponotini: Forel, 1886f: 193; Emery, 1895j: 772; Ashmead, 1905b: 384.

Colobopsis in Formicinae, Camponotini: Bondroit, 1918: 65; Emery, 1925b: 144; all subsequent authors.

Colobopsis as genus: Mayr, 1861: 38; Mayr, 1862: 652; Mayr, 1865: 7; Mayr, 1870b: 940; Forel, 1878: 368; Forel, 1879a: 125; Emery & Forel, 1879a: 449; André, 1882b: 159; Bingham, 1903: 342; Forel, 1886f: 193; Ashmead, 1905b: 384; Bondroit, 1918: 66; Jaffe, 1993: 14.

Colobopsis as subgenus of Camponotus: Emery, 1889b: 517; Dalla Torre, 1893: 219; Forel, 1893a: 165; Forel, 1893b: 435; Emery, 1895j: 772; Wheeler, W.M. 1904b: 139; Wheeler, W.M. 1910g: 144; Forel, 1912i: 90; Forel, 1914a: 263; Forel, 1917: 251; Emery, 1920b: 247; Arnold, 1922: 613; Wheeler, W.M. 1922a: 708; Emery, 1925b: 144; Donisthorpe, 1943f: 634; Creighton, 1950a: 390; Chapman & Capco, 1951: 222; Kempf, 1972a: 42; Smith, D.R., 1979: 1433; Bolton, 1994: 50.

Junior synonyms of CAMPONOTUS (COLOBOPSIS)

Condylomyrma Santschi, 1928c: 72 [as subgenus of Camponotus]. Type-species: Camponotus (Condylomyrma) bryani, by monotypy.

Taxonomic history

Condylomyrma as subgenus of Camponotus: Santschi, 1928c: 72.

Condylomyrma as junior synonym of Colobopsis: Wheeler, W.M. 1934e: 422 (in text).

[Campylomyrma Wheeler, W.M. 1934e: 421 (in text), incorrect subsequent spelling.]

Dolophra Wu, J. & Wang, 1994: 35. Type-species: Dolophra politae, by original designation.

Taxonomic history

Dolophra as genus: Wu, J. & Wang, 1994: 35; Wu, J. & Wang, 1995: 158.

Dolophra as junior synonym of Camponotus: Bolton, 1995b: 27.

Dolophra as junior synonym of Camponotus (Colobopsis): Bolton, 2003: 113, 268.

Subgenus CAMPONOTUS (MAYRIA)

Mayria Forel, 1878: 369. Type-species: Mayria madagascarensis (junior secondary homonym in Camponotus, replaced by Camponotus repens), by monotypy.

Taxonomic history

Mayria in Camponotinae: Forel, 1878: 369 [Camponotidae]; Dalla Torre, 1893: 219.

Mayria in Camponotinae, Camponotini: Forel, 1893a: 165; Emery, 1895j: 772; Ashmead, 1905b: 384; Wheeler, W.M. 1910g: 144.

Mayria as subgenus of Camponotus: Forel, 1894e: 227; Forel, 1914a: 262; Forel, 1917: 251; Wheeler, W.M. 1922a: 706; Emery, 1925b: 121; all subsequent authors.

Junior synonym of CAMPONOTUS (MAYRIA)

Myrmosaga Forel, 1912i: 92 [as subgenus of Camponotus]. Type-species: Camponotus kelleri, by subsequent designation of Wheeler, W.M. 1913a: 81.

Taxonomic history

[Type-species not Camponotus quadrimaculatus, unjustified subsequent designation by Forel, 1914a: 260; repeated in Arnold, 1922: 613.]

Myrmosaga as subgenus of Camponotus Forel, 1912i: 92; Forel, 1914a: 260; Forel, 1917: 250; Emery, 1920b: 241; Arnold, 1922: 613; Wheeler, W.M. 1922a: 706; all subsequent authors.

Myrmosaga as junior synonym of Mayria: Emery, 1925b: 121.

Subgenus CAMPONOTUS (RHINOMYRMEX)

Rhinomyrmex Forel, 1886f: 192. Type-species: Rhinomyrmex klaesii, by monotypy.

Taxonomic history

Rhinomyrmex in Camponotinae: Forel, 1886f: 192 [Camponotii]; Dalla Torre, 1893: 219.

Rhinomyrmex in Camponotinae, Camponotini: Forel, 1893a: 165; Emery, 1895j: 772; Ashmead, 1905b: 384; Wheeler, W.M. 1910g: 144.

Rhinomyrmex as genus: Forel, 1886f: 192; Forel, 1893a: 165; Dalla Torre, 1893: 219; Emery, 1895j: 772; Wheeler, W.M. 1910g: 144.

Rhinomyrmex as subgenus of Camponotus: Emery, 1896d: 374; Forel, 1914a: 264; Forel, 1917: 251; Emery, 1920b: 247; Wheeler, W.M. 1922a: 708; Emery, 1925b: 142; all subsequent authors.

Subgenus CAMPONOTUS (DENDROMYRMEX)

Dendromyrmex Emery, 1895j: 772. Type-species: Formica chartifex, by subsequent designation of Wheeler, W.M. 1911f: 161.

Taxonomic history

Dendromyrmex in Camponotinae, Camponotini: Emery, 1895j: 772; Ashmead, 1905b: 384; Forel, 1912i: 89; Wheeler, W.M. 1910g: 144; Forel, 1917: 250.

Dendromyrmex in Formicinae, Camponotini: Wheeler, W.M. 1922a: 700; Emery, 1925b: 172; all subsequent authors except the following.

Dendromyrmex in Formicinae, Formica genus group: Agosti, 1991: 295.

Dendromyrmex as subgenus of Camponotus: Fernández, 2002: 51.

Subgenus CAMPONOTUS (DINOMYRMEX)

Dinomyrmex Ashmead, 1905b: 384. Type-species: Formica gigas, by original designation.

Taxonomic history

Dinomyrmex in Camponotinae, Camponotini: Ashmead, 1905b: 384.

Dinomyrmex as genus: Ashmead, 1905b: 384.

Dinomyrmex as subgenus of Camponotus: Forel, 1913b: 350; Forel, 1914a: 259; Forel, 1917: 250; Emery, 1920b: 236; Arnold, 1922: 612; Wheeler, W.M. 1922a: 704; Emery, 1925b: 69; all subsequent authors.

Junior synonym of CAMPONOTUS (DINOMYRMEX)

Myrmogigas Forel, 1912i: 91 [as subgenus of Camponotus]. Type-species: Formica gigas, by subsequent designation of Wheeler, W.M. 1913a: 81.

Taxonomic history

Myrmogigas as subgenus of Camponotus: Forel, 1912i: 91.

Myrmogigas as junior synonym of Dinomyrmex: Forel, 1913b: 350 (in text); Forel, 1914a: 259; Wheeler, W.M. 1922a: 704. [Dinomyrmex and Myrmogigas share the same type-species, synonymy is therefore absolute.]

Subgenus CAMPONOTUS (TANAEMYRMEX)

Tanaemyrmex Ashmead, 1905b: 384. Type-species: Formica longipes (junior primary homonym in Formica, replaced by Camponotus etiolipes), by original designation.

Taxonomic history

Tanaemyrmex in Camponotinae, Camponotini: Ashmead, 1905b: 384.

Tanaemyrmex as genus: Ashmead, 1905b: 384.

Tanaemyrmex as subgenus of Camponotus: Emery, 1925b: 75; all subsequent authors.

Junior synonym of CAMPONOTUS (TANAEMYRMEX)

Myrmoturba Forel, 1912i: 91 [as subgenus of Camponotus]. Type-species: Formica maculata, by subsequent designation of Wheeler, W.M. 1913a: 82.

Taxonomic history

Myrmoturba as subgenus of Camponotus: Forel, 1912i: 91; Forel, 1914a: 259; Forel, 1917: 250; Emery, 1920b: 235; Arnold, 1922: 612; Wheeler, W.M. 1922a: 704.

Myrmoturba as junior synonym of Tanaemyrmex: Emery, 1925b: 75; all subsequent authors.

Subgenus CAMPONOTUS (ORTHONOTOMYRMEX)

Orthonotomyrmex Ashmead, 1906: 31.

Taxonomic history

[Replacement name for Orthonotus Ashmead, 1905b: 384; junior homonym of Orthonotus Westwood, in Stephens, 1829: 344 (Hemiptera). Type-species not Formica lateralis, unjustified subsequent designation by Forel, 1914a: 264; repeated in Arnold, 1922: 613.]

Orthonotomyrmex as subgenus of Camponotus: Forel, 1913i: 350 (in text); Forel, 1914a: 264; Forel, 1917: 251; Emery, 1920b: 244; Arnold, 1922: 613; Wheeler, W.M. 1922a: 707; Emery, 1925b: 124; all subsequent authors.

Homonym replaced by ORTHONOTOMYRMEX

Orthonotus Ashmead, 1905b: 384. Type-species: Formica sericea, by original designation.

Taxonomic history

[Junior homonym of Orthonotus Westwood, in Stephens, 1829: 344 (Hemiptera).]

Orthonotus in Camponotinae, Camponotini: Ashmead, 1905b: 384.

Subgenus CAMPONOTUS (MYRMAMBLYS)

Myrmamblys Forel, 1912i: 90 [as subgenus of Camponotus]. Type-species: Camponotus reticulatus, by subsequent designation of Wheeler, W.M. 1913a: 80.

Taxonomic history

[Type-species not Camponotus fastigatus, unjustified subsequent designation by Forel, 1914a: 263; repeated in Arnold, 1922: 613.]

Myrmamblys as subgenus of Camponotus: Forel, 1912i: 90; Forel, 1914a: 263; Forel, 1917: 251; Emery, 1920b: 248; Arnold, 1922: 613; Wheeler, W.M. 1922a: 708; Emery, 1925b: 136; Santschi, 1926c: 601; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOTHRIX)

Myrmothrix Forel, 1912i: 91 [as subgenus of Camponotus]. Type-species: Formica abdominalis (junior primary homonym; Formica atriceps is first available replacement name), by subsequent designation of Wheeler, W.M. 1913a: 81.

Taxonomic history

[Type-species not Formica rufipes, unjustified subsequent designation by Forel, 1914a: 260.]

Myrmothrix as subgenus of Camponotus: Forel, 1912i: 91; Forel, 1914a: 260; Forel, 1917: 250; Emery, 1920b: 236; Emery, 1925b: 107; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOSERICUS)

Myrmosericus Forel, 1912i: 91 [as subgenus of Camponotus]. Type-species: Formica rufoglauca, by subsequent designation of Wheeler, W.M. 1913a: 81.

Taxonomic history

Myrmosericus as subgenus of Camponotus: Forel, 1912i: 91; Forel, 1914a: 259; Forel, 1917: 250; Emery, 1920b: 236; Arnold, 1922: 613; Wheeler, W.M. 1922a: 704; Emery, 1925b: 104; all subsequent authors.

[See note under Myrmosaulus.]

Subgenus CAMPONOTUS (MYRMOPHYMA)

Myrmophyma Forel, 1912i: 91 [as subgenus of Camponotus]. Type-species: Camponotus capito, by subsequent designation of Wheeler, W.M. 1913a: 81.

Taxonomic history

[Type-species not Camponotus quadrisectus, unjustified subsequent designation by Forel, 1914a: 261; repeated in Arnold, 1922: 612.]

Myrmophyma as subgenus of Camponotus: Forel, 1912i: 91; Forel, 1914a: 261; Forel, 1917: 250; Emery, 1920b: 239; Arnold, 1922: 612; Wheeler, W.M. 1922a: 706; Emery, 1925b: 109; all subsequent authors.

Junior synonym of CAMPONOTUS (MYRMOPHYMA)

Myrmocamelus Forel, 1914a: 261 [as subgenus of Camponotus]. Type-species: Formica ephippium, by original designation.

Taxonomic history

[Type-species not Camponotus gambeyi, unjustified subsequent designation by Forel, 1922: 101. Myrmocamelus also described as new by Forel, 1915b: 102.]

Myrmocamelus as subgenus of Camponotus: Forel, 1914a: 261; Forel, 1915b: 102; Forel, 1917: 250; Forel, 1922: 101.

Myrmocamelus as junior synonym of Myrmophyma: Emery, 1920b: 257; Wheeler, W.M. 1921a: 18; Emery, 1925b: 109.

Subgenus CAMPONOTUS (MYRMOTREMA)

Myrmotrema Forel, 1912i: 91 [as subgenus of Camponotus]. Type-species: Camponotus foraminosus, by subsequent designation of Wheeler, W.M. 1913a: 81.

Taxonomic history

Myrmotrema as subgenus of Camponotus: Forel, 1912i: 91; Forel, 1914a: 262; Forel, 1917: 250; Emery, 1920b: 245; Arnold, 1922: 613; Wheeler, W.M. 1922a: 707; Emery, 1925b: 130; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOBRACHYS)

Myrmobrachys Forel, 1912i: 91 [as subgenus of Camponotus]. Type-species: Formica senex, by subsequent designation of Wheeler, W.M. 1913a: 81.

Taxonomic history

Myrmobrachys as subgenus of Camponotus: Forel, 1912i: 91; Forel, 1914a: 262; Forel, 1917: 251; Emery, 1920b: 251; Wheeler, W.M. 1922a: 709; Emery, 1925b: 161; all subsequent authors.

Subgenus CAMPONOTUS (MYRMENTOMA)

Myrmentoma Forel, 1912i: 92 [as subgenus of Camponotus]. Type-species: Formica lateralis, by subsequent designation of Wheeler, W.M. 1913a: 80.

Taxonomic history

Myrmentoma as junior synonym of Orthonotomyrmex: Forel, 1913b: 350 (in text); Forel, 1914a: 264; Wheeler, W.M. 1922a: 970.

Myrmentoma as subgenus of Camponotus: Forel, 1912i: 92; Emery, 1920b: 243; Wheeler, W.M. 1922a: 707; Emery, 1925a: 62; Emery, 1925b: 116; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOSPHINCTA)

Myrmosphincta Forel, 1912i: 92 [as subgenus of Camponotus]. Type-species: Formica sexguttata, by subsequent designation of Wheeler, W.M. 1913a: 81.

Taxonomic history

[Type-species not Formica cinerascens, unjustified subsequent designation by Forel, 1914a: 265; repeated in Arnold, 1922: 614.]

Myrmosphincta as subgenus of Camponotus: Forel, 1912i: 92; Forel, 1914a: 265; Forel, 1917: 251; Emery, 1920b: 238; Arnold, 1922: 614; Wheeler, W.M. 1922a: 708; Emery, 1925b: 151; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOTARSUS)

Myrmotarsus Forel, 1912i: 92 [as subgenus of Camponotus]. Type-species: Formica mistura, by subsequent designation of Wheeler, W.M. 1913a: 81.

Taxonomic history

[Type-species not Formica irritabilis, unjustified subsequent designation by Forel, 1914a: 260.]

Myrmotarsus as subgenus of Camponotus: Forel, 1912i: 92; Forel, 1914a: 260; Forel, 1917: 250; Emery, 1920b: 238; Wheeler, W.M. 1922a: 705; Emery, 1925b: 134; all subsequent authors.

Subgenus CAMPONOTUS (MYRMEPOMIS)

Myrmepomis Forel, 1912i: 92 [as subgenus of Camponotus]. Type-species: Formica sericeiventris, by subsequent designation of Wheeler, W.M. 1913a: 81.

Taxonomic history

[Type-species not Formica fulvopilosa, unjustified subsequent designation by Forel, 1914a: 265; repeated in Arnold, 1922: 614.]

Myrmepomis as subgenus of Camponotus: Forel, 1912i: 92; Forel, 1914a: 265; Forel, 1917: 251; Emery, 1920b: 245; Arnold, 1922: 614; Wheeler, W.M. 1922a: 705; Emery, 1925b: 170; all subsequent authors.

Junior synonym of CAMPONOTUS (MYRMEPOMIS)

Myrmolophus Emery, 1920b: 237 [as subgenus of Camponotus]. Type-species: Formica sericeiventris, by original designation.

Taxonomic history

Myrmolophus as subgenus of Camponotus: Emery, 1920b: 237.

Myrmolophus as junior synonym of Myrmepomis: Wheeler, W.M. 1921a: 17; Wheeler, W.M. 1922a: 705; Emery, 1925b: 170. [Myrmepomis and Myrmolophus share the same type-species, synonymy is therefore absolute.]

Subgenus CAMPONOTUS (MYRMOGONIA)

Myrmogonia Forel, 1912i: 92 [as subgenus of Camponotus]. Type-species: Camponotus laminatus, by subsequent designation of Wheeler, W.M. 1913a: 81.

Taxonomic history

Myrmogonia as subgenus of Camponotus: Forel, 1912i: 92; Forel, 1914a: 261; Forel, 1917: 250; Emery, 1920b: 240; Wheeler, W.M. 1922a: 706; Emery, 1925b: 143; all subsequent authors.

Subgenus CAMPONOTUS (MYRMEURYNOTA)

Myrmeurynota Forel, 1912i: 92 [as subgenus of Camponotus]. Type-species: Camponotus eurynotus, by subsequent designation of Wheeler, W.M. 1913a: 81.

Taxonomic history

[Type-species not Camponotus gilviventris, unjustified subsequent designation by Forel, 1914a: 266.]

Myrmeurynota as subgenus of Camponotus: Forel, 1912i: 92; Forel, 1914a: 266; Forel, 1917: 251; Emery, 1920b: 251; Wheeler, W.M. 1922a: 710; Emery, 1925b: 167; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOPSAMMA)

Myrmopsamma Forel, 1914a: 261 [as subgenus of Camponotus]. Type-species: Camponotus mystaceus, by original designation.

Taxonomic history

Myrmopsamma as subgenus of Camponotus: Forel, 1914a: 261; Forel, 1917: 250; Emery, 1920b: 246; Arnold, 1922: 612; Wheeler, W.M. 1922a: 708; Emery, 1925b: 70; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOMALIS)

Myrmomalis Forel, 1914a: 263 [as subgenus of Camponotus]. Type-species: Camponotus depressus, by original designation.

Taxonomic history

Myrmomalis as subgenus of Camponotus: Forel, 1914a: 263; Forel, 1917: 251; Emery, 1920b: 251; Wheeler, W.M. 1922a: 710; Emery, 1925b: 169; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOPLATYS)

Myrmoplatys Forel, 1916: 460 [as subgenus of Camponotus]. Type-species: Camponotus korthalsiae, by original designation.

Taxonomic history

Myrmoplatys as genus: Wheeler, W.M. 1936e: 219 (in text); Donisthorpe, 1943f: 670.

Myrmoplatys as subgenus of Camponotus: Forel, 1916: 460; Forel, 1917: 251; Emery, 1920b: 238; Wheeler, W.M. 1922a: 705; Emery, 1925b: 135; Bolton, 1994: 50.

Subgenus CAMPONOTUS (MYRMAPHAENUS)

Myrmaphaenus Emery, 1920b: 237 [as subgenus of Camponotus]. Type-species: Camponotus leydigi, by original designation.

Taxonomic history

Myrmaphaenus as subgenus of Camponotus: Emery, 1920b: 237; Wheeler, W.M. 1922a: 705; Emery, 1925b: 152; all subsequent authors.

Junior synonyms of CAMPONOTUS (MYRMAPHAENUS)

Paracolobopsis Emery, 1920b: 249 [as subgenus of Camponotus]. Type-species: Camponotus salvini, by original designation.

Taxonomic history

Paracolobopsis as subgenus of Camponotus: Emery, 1920b: 249; Wheeler, W.M. 1922a: 709.

Paracolobopsis as junior synonym of Myrmaphaenus: Emery, 1925b: 152; Kempf, 1972a: 44.

Neomyrmamblys Wheeler, W.M. 1921a: 19 [as subgenus of Camponotus]. Type-species: Camponotus fastigatus, by subsequent designation of Santschi, 1921f: 311.

Taxonomic history

Neomyrmamblys as subgenus of Camponotus: Wheeler, W.M. 1921a: 19; Wheeler, W.M. 1922a: 708.

Neomyrmamblys as junior synonym of Myrmaphaenus: Emery, 1925b: 152; Kempf, 1972a: 44.

Subgenus CAMPONOTUS (MYRMONESITES)

Myrmonesites Emery, 1920b: 242 [as subgenus of Camponotus]. Type-species: Camponotus putatus, by original designation.

Taxonomic history

Myrmonesites as subgenus of Camponotus: Emery, 1920b: 242; Wheeler, W.M. 1922a: 706; Emery, 1925b: 123.

[Myrmensites Donisthorpe, 1943f: 666, incorrect subsequent spelling.]

Subgenus CAMPONOTUS (MYRMOPYTIA)

Myrmopytia Emery, 1920b: 243 [as subgenus of Camponotus]. Type-species: Camponotus imitator, by original designation.

Taxonomic history

Myrmopytia as subgenus of Camponotus: Emery, 1920b: 243; Wheeler, W.M. 1922a: 707; Emery, 1925b: 114; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOTEMNUS)

Myrmotemnus Emery, 1920b: 246 [as subgenus of Camponotus]. Type-species: Camponotus moeschi, by original designation.

Taxonomic history

Myrmotemnus as junior synonym of Myrmamblys: Wheeler, W.M. 1921a: 19; Wheeler, W.M. 1922a: 708; Emery, 1925b: 137; Donisthorpe, 1943f: 671 (anachronism).

Myrmotemnus as subgenus of Camponotus: Emery, 1920b: 246; Santschi, 1926c: 601.

Subgenus CAMPONOTUS (PSEUDOCOLOBOPSIS)

Pseudocolobopsis Emery, 1920b: 249 [as subgenus of Camponotus]. Type-species: Camponotus macrocephalus, by original designation.

Taxonomic history

Pseudocolobopsis as subgenus of Camponotus: Emery, 1920b: 249; Wheeler, W.M. 1922a: 709; Emery, 1925b: 157; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOSTENUS)

Myrmostenus Emery, 1920b: 250 [as subgenus of Camponotus]. Type-species: Camponotus mirabilis, by original designation.

Taxonomic history

Myrmostenus as subgenus of Camponotus: Emery, 1920b: 250; Wheeler, W.M. 1922a: 709; Emery, 1925b: 161; all subsequent authors.

Subgenus CAMPONOTUS (HYPERCOLOBOPSIS)

Hypercolobopsis Emery, 1920b: 250 [as subgenus of Camponotus]. Type-species: Colobopsis paradoxa, by original designation.

Taxonomic history

Hypercolobopsis as subgenus of Camponotus: Emery, 1920b: 250; Wheeler, W.M. 1922a: 709; Emery, 1925b: 160; all subsequent authors.

Junior synonym of CAMPONOTUS (HYPERCOLOBOPSIS)

Neocolobopsis Borgmeier, 1928b: 65 [as subgenus of Camponotus]. Type-species: Camponotus (Neocolobopsis) scrobifer (junior synonym of Camponotus coriolanus), by original designation.

Taxonomic history

Neocolobopsis as subgenus of Camponotus Borgmeier, 1928b: 65.

Neocolobopsis as junior synonym of Hypercolobopsis: Kempf, 1968b: 411; Kempf, 1972a: 43.

Subgenus CAMPONOTUS (MYRMOPIROMIS)

Myrmopiromis Wheeler, W.M. 1921a: 17 [as subgenus of Camponotus]. Type-species: Formica fulvopilosa, by subsequent designation of Wheeler, W.M. 1922a: 707.

Taxonomic history

Myrmopiromis as subgenus of Camponotus: Wheeler, W.M. 1921a: 17; Wheeler, W.M. 1922a: 707; Emery, 1925b: 1127.

Subgenus CAMPONOTUS (MYRMOSAULUS)

Myrmosaulus Wheeler, W.M. 1921a: 18 [as subgenus of Camponotus]. Type-species: Formica cinerascens, by subsequent designation of Wheeler, W.M. 1922a: 705.

Taxonomic history

[Type-species not Formica singularis, unjustified subsequent designation by Emery, 1925b: 113, repeated in Donisthorpe, 1943f: 670. Note that Donisthorpe, 1932c: 445, states that C. cinerascens belongs in subgenus Myrmosericus. If so then Myrmosaulus would fall as a junior synonym of Myrmosericus.]

Myrmosaulus as subgenus of Camponotus: Wheeler, W.M. 1921a: 18; Wheeler, W.M. 1922a: 705; Emery, 1925b: 112; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOCLADOECUS)

Myrmocladoecus Wheeler, W.M. 1921a: 19 [as subgenus of Camponotus]. Type-species: Camponotus latangulus, by subsequent designation of Wheeler, W.M. 1922a: 709.

Taxonomic history

Myrmocladoecus as subgenus of Camponotus: Wheeler, W.M. 1921a: 19; Wheeler, W.M. 1922a: 709; Emery, 1925b: 166; all subsequent authors.

Subgenus CAMPONOTUS (MANNIELLA)

Manniella Wheeler, W.M. 1921a: 19 [as subgenus of Camponotus]. Type-species: Camponotus sphaericus, by original designation.

Taxonomic history

Manniella subgenus of Camponotus: Wheeler, W.M. 1921a: 19; Wheeler, W.M. 1922a: 710; Emery, 1925b: 159; all subsequent authors.

Subgenus CAMPONOTUS (MYRMISOLEPIS)

Myrmisolepis Santschi, 1921f: 310 [as subgenus of Camponotus]. Type-species: Camponotus epinotalis, by original designation.

Taxonomic history

Myrmisolepis as subgenus of Camponotus: Santschi, 1921f: 310; Emery, 1925b: 133; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOPELTA)

Myrmopelta Santschi, 1921f: 310 [as subgenus of Camponotus]. Type-species: Camponotus arminius, by original designation.

Taxonomic history

Myrmopelta as junior synonym of Myrmamblys: Emery, 1925b: 137.

Myrmopelta as subgenus of Camponotus: Santschi, 1921f: 310; Wheeler, W.M. 1922a: 704 (footnote); Santschi, 1926a: 16; Santschi, 1926c: 603; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOPLATYPUS)

Myrmoplatypus Santschi, 1921f: 311 [as subgenus of Camponotus]. Type-species: Camponotus platytarsus, by original designation.

Taxonomic history

Myrmoplatypus as subgenus of Camponotus: Santschi, 1921f: 311; Emery, 1925b: 169; all subsequent authors.

Subgenus CAMPONOTUS (MYRMEPINOTUS)

Myrmepinotus Santschi, 1921f: 312 [as subgenus of Camponotus]. Type-species: Camponotus echinoploides, by original designation.

Taxonomic history

Myrmepinotus as subgenus of Camponotus: Santschi, 1921f: 312; Emery, 1925b: 126; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOXYGENYS)

Myrmoxygenys Emery, 1925b: 70 [as subgenus of Camponotus]. Type-species: Camponotus caesar, by monotypy.

Taxonomic history

Myrmoxygenys as subgenus of Camponotus: Emery, 1925b: 70; all subsequent authors.

Subgenus CAMPONOTUS (KARAVAIEVIA)

Karavaievia Emery, 1925b: 115 [as subgenus of Camponotus]. Type-species: Camponotus exsectus, by original designation.

Taxonomic history

Karavaievia as subgenus of Camponotus: Emery, 1925b: 115; all subsequent authors.

Subgenus CAMPONOTUS (MYRMODIRACHIS)

Myrmodirachis Emery, 1925b: 168 [as subgenus of Camponotus]. Type-species: Camponotus heathi, by original designation.

Taxonomic history

Myrmodirachis as subgenus of Camponotus: Emery, 1925b: 168; all subsequent authors.

Subgenus CAMPONOTUS (MYRMESPERA)

Myrmespera Santschi, 1926b: 247 [as subgenus of Camponotus]. Type-species: Camponotus (Myrmespera) debellator, by subsequent designation of Santschi, 1926c: 604.

Taxonomic history

Myrmespera as subgenus of Camponotus: Santschi, 1926b: 247; Santschi, 1926c: 603; all subsequent authors.

Subgenus CAMPONOTUS (PARAMYRMAMBLYS)

Paramyrmamblys Santschi, 1926c: 604 [as subgenus of Camponotus].Type-species: Camponotus (Myrmamblys) ostiarius, by original designation.

Taxonomic history

Paramyrmamblys as subgenus of Camponotus: Santschi, 1926c: 604; all subsequent authors.

Subgenus CAMPONOTUS (MYRMACRHAPHE)

Myrmacrhaphe Santschi, 1926c: 607 [as subgenus of Camponotus]. Type-species: Camponotus conradti, by original designation.

Taxonomic history

Myrmacrhaphe as subgenus of Camponotus: Santschi, 1926c: 607; all subsequent authors.

Subgenus CAMPONOTUS (THLIPSEPINOTUS)

Thlipsepinotus Santschi, 1928e: 483 [as subgenus of Camponotus]. Type-species: Camponotus claripes, by original designation.

Taxonomic history

Thlipsepinotus as subgenus of Camponotus: Santschi, 1928e: 483; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOPALPELLA)

Myrmopalpella Stärcke, 1934: 30 [as subgenus of Camponotus]. Type-species: Camponotus megalonyx, by monotypy.

Taxonomic history

Myrmopalpella as genus: Wheeler, W.M. 1936e: 219 (in text); Donisthorpe, 1943f: 669.

Myrmopalpella as subgenus of Camponotus: Bolton, 1995b: 37.

Genus Camponotus references

Mayr, 1861: 35 (Europe species key); Roger, 1863b: 1, 9 (Camponotus, Colobopsis catalogues); Mayr, 1863: 397, 403 (Camponotus, Colobopsis catalogues); Mayr, 1865: 6, 7 (Camponotus, Colobopsis diagnosis); Mayr, 1867a: 67 (Colobopsis diagnois); Mayr, 1868c: 27 (†Baltic Amber species key); Mayr, 1870a: 373 (Colombia + Panama (= New Grenada) species key); Mayr, 1870b: 941 (Colobopsis species key); André, 1874: 175 (Europe species key); Forel, 1874: 38 (Switzerland species key); Mayr, 1876: 58 (Australia species key); Mayr, 1878: 869 (all Dendromyrmex species key (as part of Camponotus)); Forel, 1878: 367, 368 (Camponotus, Colobopsis diagnoses); André, 1882a: 138 (Europe & Algeria species key); André, 1882b: 159 (Europe & Algeria Colobopsis species key); Cresson, 1887: 255 (U.S.A. Camponotus, Colobopsis catalogue); Provancher, 1887: 228 (Canada species key); Nasonov, 1889: 57 (Russia species key); Forel, 1891b: 71, 216 (Madagascar species key); Forel, 1892j: 223 (India & Sri Lanka species key); Dalla Torre, 1893: 219 (Rhinomyrmex, Mayria, Camponotus catalogues); Emery, 1893i: 667 (North America species key); Forel, 1893b: 435 (India & Sri Lanka C. (Colobopsis) species key); Emery, 1895j: 772 (synoptic classification); Emery, 1896d: 370, 378 (species groups, catalogue); Emery, 1898a: 225 (additions to 1896a catalogue); Emery, 1903 (South America species key); Bingham, 1903: 343, 348 (India, Sri Lanka & Burma Colobopsis, Camponotus species keys); Ruzsky, 1905b: 185 (Russian Empire species key); Emery, 1908a: 183 (Palaearctic C. herculeanus & C. maculatus groups species key); Bondroit, 1910: 486 (Belgium species key); Wheeler, W.M. 1910d: 297 (North America species key); Forel, 1914a: 259, 266 (subgenera diagnoses, species listed by subgenus); Stitz, 1914: 94 (Central Europe species key); Santschi, 1915c: 266 (C. (Myrmotrema) species key); Emery, 1916b: 227 (Italy species key); Wheeler, W.M. 1916m: 600 (U.S.A., Connecticut species key); Mann, 1916: 489 (Dendromyrmex, all species key); Bondroit, 1918 (France & Belgium species key); Emery, 1920b: 235 (subgenera, diagoses); Emery, 1920c: 8 (Old World C. maculatus group, checklist & key); Wheeler, W.M. 1921a: 16 (subgenera, notes); Santschi, 1921f: 310 (subgenera, notes); Mann, 1921: 489, 499 (Fiji Is C. (Myrmogonia) & C. (Colobopsis) species keys); Wheeler, W.M. 1922a: 704 (subgenera synopsis); Wheeler, W.M. 1922a: 232, 948, 1039 (diagnosis, Afrotropical, Malagasy catalogues); Arnold, 1922: 610, 612, 619, 662, 673 (diagnosis, South Africa subgenera synopsis, South Africa C. (Tanaemyrmex), C. (Myrmophyma), C. (Myrmopsamma) species keys); Arnold, 1924: 683, 700, 709, 724, 735 (South Africa C. (Myrmosericus), C. (Myrmamblys), C. (Orthonotomyrmex), C. (Myrmotrema), C. (Myrmepomis) species keys); Emery, 1925a: 62 (Palaearctic C. lateralis group, key); Emery, 1925b: 59 (diagnosis, subgenera key, catalogue); Emery, 1925b: 69 (C. (Dinomyrmex) diagnosis, catalogue); Emery, 1925b: 70 (C. (Myrmoxygenys) & C. (Myrmopsamma) diagnoses, catalogues); Emery, 1925b: 71 (C. (Camponotus) diagnosis, catalogue); Emery, 1925b: 75 (C. (Tanaemyrmex) diagnosis, catalogue); Emery, 1925b: 104 (C. (Myrmosericus) diagnosis, catalogue); Emery, 1925b: 107 (C. (Myrmothrix) diagnosis, catalogue); Emery, 1925b: 109 (C. (Myrmophyma) diagnosis, catalogue); Emery, 1925b: 112 (C. (Myrmosaulus) diagnosis, catalogue); Emery, 1925b: 114 (C. (Myrmopytia) diagnosis, catalogue); Emery, 1925b: 115 (C. (Karavaievia) diagnosis, catalogue); Emery, 1925b: 116 (C. (Myrmentoma) diagnosis, catalogue); Emery, 1925b: 121 (C. (Mayria) diagnosis, catalogue); Emery, 1925b: 123 (C. (Myrmonesites) diagnosis, catalogue); Emery, 1925b: 124 (C. (Orthonotomyrmex) diagnosis, catalogue); Emery, 1925b: 126 (C. (Myrmepinotus) diagnosis, catalogue); Emery, 1925b: 127 (C. (Myrmopiromis) diagnosis, catalogue); Emery, 1925b: 130 (C. (Myrmotrema) diagnosis, catalogue); Emery, 1925b: 133 (C. (Myrmisolepis) diagnosis, catalogue); Emery, 1925b: 134 (C. (Myrmotarsus) diagnosis, catalogue); Emery, 1925b: 135 (C. (Myrmoplatys) diagnosis, catalogue); Emery, 1925b: 136 (C. (Myrmamblys) diagnosis, catalogue); Emery, 1925b: 142 (C. (Rhinomyrmex) diagnosis, catalogue); Emery, 1925b: 143 (C. (Myrmogonia) diagnosis, catalogue); Emery, 1925b: 144 (C. (Colobopsis) diagnosis, catalogue); Emery, 1925b: 151 (C. (Myrmosphincta) diagnosis, catalogue); Emery, 1925b: 152 (C. (Myrmaphaenus) diagnosis, catalogue); Emery, 1925b: 157 (C. (Pseudocolobopsis) diagnosis, catalogue); Emery, 1925b: 159 (C. (Manniella) diagnosis, catalogue); Emery, 1925b: 160 (C. (Hypercolobopsis) diagnosis, catalogue); Emery, 1925b: 161 (C. (Myrmostenus) & C. (Myrmobrachys) diagnoses, catalogues); Emery, 1925b: 166 (C. (Myrmocladoecus) diagnosis, catalogue); Emery, 1925b: 167 (C. (Myrmeurynota) diagnosis, catalogue); Emery, 1925b: 168 (C. (Myrmodirachis) diagnosis, catalogue); Emery, 1925b: 169 (C. (Myrmomalis) & C. (Myrmoplatypus) diagnoses, catalogues); Emery, 1925b: 170 (C. (Myrmepomis) diagnosis, catalogue); Emery, 1925b: 172 (Dendromyrmex diagnosis, catalogue); Santschi, 1926a: 16, 21 (C. (Myrmopelta) diagnosis, species key); Karavaiev, 1927c: 275 (Ukraine species key); Arnol'di, 1933b: 601 (Russia species key); Menozzi, 1933b: 81 (Israel species key); Menozzi, 1935c: 330 (Chile species key); Karavaiev, 1936: 175 (Ukraine species key); Finzi, 1936: 190 (Egypt species key); Menozzi, 1939a: 316 (Himalaya & Tibet species key); Stitz, 1939: 236 (Germany species key); Kratochvíl, 1941: 109 (Central Europe species key); Novák & Sadil, 1941: 109 (Central Europe species key); Cole, 1942: 387 (U.S.A., Utah species key); Donisthorpe, 1932g: 248 (C. (Orthonotomyrmex) notes); Holgersen, 1943b: 174 (Norway species key); Holgersen, 1944: 199 (Norway species key); Buren, 1944a: 293 (U.S.A., Iowa species key); Creighton, 1950a: 362 (North America species keys); Chapman & Capco, 1951: 220 (Asia checklist); Yasumatsu & Brown, 1951: 29 (C. herculeanus complex); Kusnezov, 1952d: 192 (Argentina species key); Yasumatsu & Brown, 1957: 45 (C. herculeanus complex); Pisarski, 1961a: 175 (Poland species key); Gregg, 1963: 655 (U.S.A., Colorado species key); Wheeler, G.C. & Wheeler, J. 1963: 163 (U.S.A., North Dakota species key); Arnol'di, 1967: 1817 (former U.S.S.R. C. (Camponotus) species key); Bernard, 1967: 330 (diagnosis, Western Europe species key); Kempf, 1972a: 42, 95 (Neotropical catalogue); Hashmi, 1973: 1 (C. (Myrmothrix) all species revision, key); Alayo, 1974: 27 (Cuba species key); Snelling & Hunt, 1976: 117 (Chile species key); Tarbinsky, 1976: 144 (Kyrghyzstan species key); Boven, 1977: 130 (Belgium species key); Kutter, 1977c: 191 (Switzerland species key); Arnol'di & Dlussky, 1978: 551 (former European U.S.S.R. species key); Collingwood, 1978: 90 (Iberian Peninsula species key); Collingwood, 1979: 87 (Fennoscandia & Denmark species key); Smith, D.R. 1979: 1424 (North America catalogue); Allred, 1982: 445 (U.S.A., Utah species key); Gösswald, 1985: 264 (Germany species key); Collingwood, 1985: 275 (Saudi Arabia species key); Taylor & Brown, D.R. 1985: 109 (Australia catalogue); Dumpert, 1986: 571 (C. (Karavaievia) species key); Wheeler, G.C. & Wheeler, J. 1986g: 60 (U.S.A., Nevada species key); Agosti & Collingwood, 1987b: 282 (Balkans species key); Taylor, 1987a: 10, 16 (Australia, New Caledonia, New Zealand checklists); Snelling, R.R. 1988: 57 (Nearctic C. (Myrmentoma) species key); Wang, C., Xiao & Wu, 1989a: 222 (China species key); Dlussky, Soyunov & Zabelin, 1990: 125 (Turkmenistan species key); Kupyanskaya, 1990: 164 (Far Eastern Russia species key); Brandão, 1991: 332 (Neotropical catalogue); Morisita, Kubota, Onoyama, et al. 1991: 38 (Japan species key); Terayama, 1991: 165 (Japan C. (Paramyrmamblys) species); Atanasov & Dlussky, 1992: 207 (Bulgaria species key); Wang, C. & Wu, 1994: 28 (China species key); Radchenko, 1994b: 116 (South Siberia species key); Arakelian, 1994: 82 (Armenia species key); Bolton, 1994: 50 (synoptic classification); Bolton, 1995a: 1048 (census); Bolton, 1995b: 83, 169 (catalogue); Douwes, 1995: 92 (Sweden species key); Dumpert, Maschwitz, Weissflog et al. 1995: 104 (C. (Karavaievia) species key); Kupyanskaya, 1995: 352 (Far Eastern Russia species key); Wu, J. & Wang, 1995: 168 (China species key); Collingwood & Agosti, 1996: 370 (Saudi Arabia species key); Seifert, 1996b: 170 (Central Europe species key); Radchenko, 1996b: 1195 (Asiatic Palaearctic species key); Cagniant, 1996b: 87 (Morocco species key); McArthur & Adams, 1996: 18 (Australia C. nigriceps group, key); Mackay, 1997: 194 (C. (Myrmostenus) species key); Mackay & Mackay, 1997b: 319 and Snelling, R.R. 2000: 607 (C. montivagus complex, keys); Robertson & Zachariades, 1997: 2 (C. fulvopilosus group, key); Snelling & Torres, 1998: 9 (Puerto Rico & Virgin Is species key); Collingwood & Prince, 1998: 24 (Portugal species key); Terayama, 1999b: 27 (Japan species key); Shattuck, 1999: 91 (Australia synopsis); Zhou, 2001b: 200 (China, Guangxi species key); McArthur & Shattuck, 2001: 27 (Australia C. macrocephalus group, key); Czechowski, Radchenko & Czechowska, 2002: 153 (Poland species key); Fernández, 2002: 58 (C. (Dendromyrmex) diagnosis, all species revision, key); Mackay & Mackay, 2002: 248 (U.S.A., New Mexico species key); Shattuck & McArthur, 2002: 65 (Australia C. wiederkehri group key); Bolton, 2003: 112 (synopsis); Shattuck, 2005: 3 (C. aureopilus group key); Radchenko, 2005b: 188 (North Korea species key); Hansen & Klotz, 2005: 75 (North American “Carpenter Ants” key); Snelling, R.R. 2006: 96 (U.S.A. C. festinatus complex, key); Karmaly & Narendran, 2006: 29 (India species, key); Dumpert, Maschwitz & Weissflog, 2006: 77 (C. (Karavaievia) species, keys); Seifert, 2007: 152 (North and Central Europe species key); McArthur, 2007a: 290 (Australia species key); Shattuck & Janda, 2009: 252 (C. aureopilus group key); Mackay & Delsinne, 2009: 496 (New World C. maculatus complex, key); Terayama, 2009: 216 (Taiwan species key); Heterick, 2009: 60 (south-western Australia species key); Ionescu-Hirsch, 2009: 59 (Israel species key); McArthur, 2010: 18 (south Australia species key).

Cerapachyinae Cerapachyini Cerapachys TRUE TRUE FALSE

146 species

Cerapachys Smith, F. 1857a: 74. Type-species: Cerapachys antennatus, by subsequent designation of Bingham, 1903: 28.

Taxonomic history

Cerapachys in Myrmicinae: Smith, F. 1857a: 74 [Myrmicidae]; Smith, F. 1861b: 47 [Myrmicidae]; Smith, F. 1871a: 329 [Myrmicidae].

Cerapachys in Ponerinae: Dalla Torre, 1893: 17.

Cerapachys in Dorylinae, Cerapachyini: Emery, 1895j: 765; Emery, 1901a: 34.

Cerapachys in Ponerinae, Cerapachyini: Forel, 1893a: 162 (also misspelled Orapachys, same page); Forel, 1900d: 331; Ashmead, 1905b: 382; Wheeler, W.M. 1910g: 137; Emery, 1911d: 8; Arnold, 1915: 11; Forel, 1917: 239; Brown, 1975: 18; Snelling, R.R. 1981: 389; Dlussky & Fedoseeva, 1988: 79; Hölldobler & Wilson, 1990: 10.

Cerapachys in Cerapachyinae, Cerapachyini: Wheeler, W.M. 1902d: 185; Wheeler, W.M. 1922a: 639; Donisthorpe, 1943f: 632; Chapman & Capco, 1951: 17; Kempf, 1972a: 76; Wheeler, G.C. & Wheeler, J. 1985: 256; Bolton, 1990a: 67; Bolton, 1990c: 1357; Jaffe, 1993: 9; Bolton, 1994: 19; Bolton, 2003: 34.

Junior synonyms of CERAPACHYS

Syscia Roger, 1861a: 19. Type-species: Syscia typhla, by monotypy.

Taxonomic history

Syscia in Ponerinae: Mayr, 1862: 714 (in key) [Poneridae]; Mayr, 1865: 15 [Poneridae]; Dalla Torre, 1893: 17.

Syscia in Dorylinae, Cerapachyini: Emery, 1895j: 765.

Syscia in Ponerinae, Cerapachyini: Forel, 1893a: 162; Forel, 1900d: 329; Ashmead, 1905b: 382.

Syscia as genus: Dalla Torre, 1893: 17; Forel, 1900d: 329; Bingham, 1903: 31.

Syscia as subgenus of Cerapachys: Wheeler, W.M. 1902d: 185; Emery, 1902c: 24; Wheeler, W.M. 1910g: 137; Emery, 1911d: 10; Forel, 1917: 239; Wheeler, W.M. 1922a: 639; Donisthorpe, 1943g: 730.

Syscia as junior synonym of Cerapachys: Kempf, 1972a: 76; Brown, 1975: 18; all subsequent authors.

Ooceraea Roger, 1862a: 248. Type-species: Ooceraea fragosa, by monotypy.

Taxonomic history

Ooceraea in Myrmicinae: Mayr, 1865: 24 [Myrmicidae].

Ooceraea in Myrmicidae, Pheidolidae: Emery, 1877a: 81.

Ooceraea in Poneridae: Smith, F. 1871a: 324.

Ooceraea in Ponerinae: Dalla Torre, 1893: 17.

Ooceraea in Ponerinae, Cerapachyini: Forel, 1893a: 162.

Ooceraea in Dorylinae, Cerapachyini: Emery, 1895j: 765; Emery, 1901a: 34.

Ooceraea in Ecitoninae, Ecitonini: Ashmead, 1905b: 381; Ashmead, 1906: 25.

Ooceraea as genus: Roger, 1862a: 248; Smith, F. 1871a: 324; Dalla Torre, 1893: 17; Forel, 1900d: 329; Bingham, 1903: 31.

Ooceraea as subgenus of Cerapachys: Emery, 1902c: 24; Wheeler, W.M. 1902d: 185; Wheeler, W.M. 1910g: 137; Emery, 1911d: 10; Forel, 1917: 239; Wheeler, W.M. 1922a: 639; Donisthorpe, 1943f: 678.

Ooceraea as junior synonym of Cerapachys: Brown, 1973b: 183 [provisional]; Brown, 1975: 19; all subsequent authors.

Lioponera Mayr, 1879: 666. Type-species: Lioponera longitarsus, by monotypy.

Taxonomic history

Lioponera in Ponerinae: Dalla Torre, 1893: 17.

Lioponera in Pachycondylinae, Lioponerini: Ashmead, 1905b: 382.

Lioponera in Ponerinae, Cerapachyini: Forel, 1893a: 162; Forel, 1900d: 329; Wheeler, W.M. 1910g: 137; Emery, 1911d: 11; Forel, 1917: 239.

Lioponera in Dorylinae, Cerapachyini: Emery, 1895j: 765.

Lioponera in Cerapachyinae, Cerapachyini: Wheeler, W.M. 1902d: 185; Wheeler, W.M. 1922a: 639; Donisthorpe, 1943f: 657.

Lioponera as subgenus of Cerapachys: Forel, 1892l: 244.

Lioponera as genus: Mayr, 1879: 666; Dalla Torre, 1893: 17; Forel, 1900d: 329; Bingham, 1903: 26; Emery, 1911d: 11; Wheeler, W.M. 1922a: 639; Donisthorpe, 1943f: 657.

Lioponera as junior synonym of Cerapachys: Brown, 1975: 19; all subsequent authors.

Parasyscia Emery, in André, 1882c: 235. Type-species: Parasyscia piochardi, by monotypy.

Taxonomic history

Parasyscia in Ponerinae: Dalla Torre, 1893: 17.

Parasyscia in Dorylinae, Cerapachyini: Emery, 1895j: 765.

Parasyscia in Ponerinae, Cerapachyini: Forel, 1893a: 162 [Cerapachysii]; Ashmead, 1905b: 382.

Parasyscia as genus: Dalla Torre, 1893: 17.

Parasyscia as subgenus of Cerapachys: Forel, 1892l: 243; Wheeler, W.M. 1902d: 185; Wheeler, W.M. 1910g: 137; Emery, 1911d: 9; Forel, 1917: 239; Wheeler, W.M. 1922a: 639; Donisthorpe, 1943f: 681; Creighton, 1950a: 57.

Parasyscia as junior synonym of Cerapachys: Kempf, 1972a: 76; Brown, 1975: 18; all subsequent authors.

Phyracaces Emery, 1902c: 23. Type-species: Cerapachys mayri, by original designation.

Taxonomic history

Phyracaces in Ponerinae, Cerapachyini: Ashmead, 1905b: 382; Wheeler, W.M. 1910g: 137; Emery, 1911d: 10; Arnold, 1915: 17; Forel, 1917: 239.

Phyracaces in Cerapachyinae, Cerapachyini: Wheeler, W.M. 1902d: 185; Wheeler, W.M. 1922a: 639; Donisthorpe, 1943f: 683.

Phyracaces as junior synonym of Cerapachys: Brown, 1973b: 183 [provisional]; Brown, 1975: 19; all subsequent authors.

Cysias Emery, 1902c: 24 [as subgenus of Cerapachys]. Type-species: Ooceraea papuana, by original designation.

Taxonomic history

Cysias in Ponerinae, Cerapachyini: Ashmead, 1905b: 382.

Cysias as genus: Ashmead, 1905b: 382.

Cysias as subgenus of Cerapachys: Emery, 1902c: 24; Wheeler, W.M. 1910g: 137; Donisthorpe, 1943f: 637.

Cysias as junior synonym of Syscia: Emery, 1911d: 10; Brown, 1975: 19; all subsequent authors.

Ceratopachys Schulz, W.A. 1906: 155, unjustified emendation of Cerapachys.

Taxonomic history

Ceratopachys as junior synonym of Cerapachys: Wheeler, W.M. 1922a: 755.

Chrysapace Crawley, 1924: 380. Type-species: Chrysapace jacobsoni (junior secondary homonym in Cerapachys, replaced by Cerapachys crawleyi), by monotypy.

Taxonomic history

Chrysapace in Ponerinae: Crawley, 1924: 380.

Chrysapace in Cerapachyinae, Cerapachyini: Donisthorpe, 1943f: 633.

Chrysapace as junior synonym of Cerapachys: Brown, 1975: 19; all subsequent authors.

Neophyracaces Clark, 1941: 76. Type-species: Phyracaces clarus, by original designation.

Taxonomic history

Neophyracaces in Cerapachyinae: Clark, 1941: 76.

Neophyracaces as junior synonym of Cerapachys: Brown, 1973b: 183 [provisional]; Brown, 1975: 19; all subsequent authors.

Yunodorylus Xu, 2000b: 297. Type-species: Yunodorylus sexspinus, by original designation.

Taxonomic history

Yunodorylus in Dorylinae: Xu, 2000b: 297.

Yunodorylus in Cerapachyinae, Cerapachyini: Bolton, 2003: 141, 268.

Yunodorylus as junior synonym of Cerapachys: Bolton, 2003: 141, 268; Borowiec, 2009: 57.

Genus Cerapachys references

Roger, 1863b: 21, 25 (catalogue); Mayr, 1863: 403, 439, 455 (catalogue); Mayr, 1865: 15, 24 (Syscia, Ooceraea diagnoses); André, 1882c: 236 (Europe & Algeria species); Dalla Torre, 1893: 17 (Lioponera, Cerapachys, Parasyscia, Ooceraea, Syscia catalogues); Forel, 1900d: 331 (India & Sri Lanka species key); Bingham, 1903: 27, 28 (India, Sri Lanka & Burma Lioponera, Cerapachys species keys); Emery, 1911d: 8 (diagnosis, subgenera key, catalogue); Emery, 1911d: 9 (C. (Parasyscia) diagnosis, catalogue); Emery, 1911d: 10 (C. (Ooceraea) & C. (Syscia) diagnoses, catalogues); Emery, 1911d: 10 (Phyracaces diagnosis, catalogue); Emery, 1911d: 11 (Lioponera diagnosis, catalogue); Arnold, 1915: 11 (diagnosis, South Africa species key); Wheeler, W.M. 1918a: 239 (Australia Phyracaces species key); Mann, 1921: 408 (Fiji species key); Wheeler, W.M. 1922a: 52, 53, 639 (Cerapachys & Phyracaces diagnoses, subgenera key); Wheeler, W.M. 1924d: 224 (Chrysapace, comments); Arnold, 1926: 193 (South Africa species, revised key); Donisthorpe, 1939a: 253 (Lioponera diagnosis); Chapman & Capco, 1951: 17, 21, 22 (Asia Cerapachys, Lioponera, Phyracaces checklists); Wilson, 1959b: 44, 52, 55, 56 (Melanesia, Fiji Is, New Guinea & New Caledonia species keys); Kempf, 1972a: 76 (Neotropical catalogue); Brown, 1975: 18, 26 (diagnosis, keys); Smith, D.R. 1979: 1333 (North America catalogue); Taylor & Brown, D.R. 1985: 23 (Australia catalogue); Taylor, 1987a: 16 (Australia, New Caledonia checklist); Terayama, Kubota, Sakai & Kawazoe, 1988: 35 (Taiwan species key); Morisita, Kubota, Onoyama, et al. 1989: 30 (Japan species key); Bolton, 1995a: 1048 (census); Bolton, 1995b: 142 (catalogue); Wu, J. & Wang, 1995: 47 (China species key); Collingwood & Agosti, 1996: 312 (Saudi Arabia species key); Terayama, 1996: 18 (Japan species key); Shattuck, 1999: 60 (Australia synopsis); Mackay & Mackay, 2002: 36 (U.S.A., New Mexico species key); Borowiec, 2009: 46 (C. sexspinus group key); Terayama, 2009: 122 (Taiwan species key); Heterick, 2009: 126 (south-western Australia species key).

Myrmicinae Dacetini Strumigenys TRUE TRUE FALSE

Extant: 487 species

Fossil:

Strumigenys Smith, F. 1860c: 72. Type-species: Strumigenys mandibularis, by monotypy.

Taxonomic history

Strumigenys in Myrmicidae: Smith, F. 1871a: 334; Cresson, 1887: 262.

Strumigenys in Myrmicidae, Cryptoceridae: Emery, 1877a: 81.

Strumigenys in Myrmicinae: Mayr, 1865: 26 [Myrmicidae]; Emery & Forel, 1879a: 464 [Myrmicidae]; Dalla Torre, 1893: 145.

Strumigenys in Cryptoceridae, Dacetini: Ashmead, 1905b: 384.

Strumigenys in Myrmicinae, Dacetini: Forel, 1892d: 344; Forel, 1893a: 164; Forel, 1895b: 133; Emery, 1895j: 770; Forel, 1899c: 42; Wheeler, W.M. 1910g: 141; Emery, 1914a: 42; Forel, 1917: 246; Arnold, 1917: 372; Wheeler, W.M. 1922a: 668; Emery, 1924d: 319; Brown, 1948e: 108; all subsequent authors.

Junior synonyms of STRUMIGENYS

Labidogenys Roger, 1862a: 249. Type-species: Labidogenys lyroessa, by monotypy.

Taxonomic history

Labidogenys in Myrmicinae, Dacetini: Donisthorpe, 1943f: 654; Brown, 1948e: 102.

Labidogenys as genus: Roger, 1862a: 249; Brown, 1948e: 102.

Labidogenys as junior synonym of Strumigenys: Roger, 1863b: 40; Emery & Forel, 1879a: 464; Dalla Torre, 1893: 145; Bingham, 1903: 147; Emery, 1924d: 320; Brown & Wilson, 1959b: 281; Brown, 1960b: 38; Bolton, 1983: 358; Bolton, 1999: 1673; Bolton, 2000: 492.

Proscopomyrmex Patrizi, 1946: 294. Type-species: Proscopomyrmex londianensis, by monotypy.

Taxonomic history

Proscopomyrmex in Myrmicinae, Dacetini: Patrizi, 1946: 294 [Dacetonini].

Proscopomyrmex as junior synonym of Strumigenys: Brown, 1949d: 15; Bolton, 1999: 1673; Bolton, 2000: 492.

Eneria Donisthorpe, 1948d: 598. Type-species: Eneria excisa (junior synonym of Strumigenys loriae), by original designation.

Taxonomic history

Eneria as genus: Chapman & Capco, 1951: 104 (anachronism).

Eneria as junior synonym of Strumigenys: Brown, 1949d: 15; Bolton, 1999: 1673; Bolton, 2000: 492.

Quadristruma Brown, 1949b: 47. Type-species: Epitritus emmae, by original designation.

Taxonomic history

Quadristruma in Myrmicinae, Dacetini: Brown, 1949b: 47; all subsequent authors.

Quadristruma as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 12; Bolton, 1999: 1673; Bolton, 2000: 492.

Genus Strumigenys references

Roger, 1863b: 40 (catalogue); Mayr, 1863: 424, 455 (catalogue); Mayr, 1865: 26 (diagnosis); Forel, 1886a: 216 (all species key); Mayr, 1887: 568 (all species key); Dalla Torre, 1893: 145 (catalogue); Emery, 1895c: 325 (North America species key); Emery, 1897c: 574 (New Guinea species key); Forel, 1903a: 707 (India & Sri Lanka species key); Bingham, 1903: 147 (India, Sri Lanka & Burma species key); Wheeler, W.M. 1908a: 146 (New World species key); Santschi, 1913b: 257 (Afrotropical species key); Arnold, 1917: 373 (diagnosis, South Africa species key); Mann, 1921: 467 (Fiji Is species key); Wheeler, W.M. 1922a: 917, 1034 (Afrotropical, Malagasy catalogues); Emery, 1924d: 319 (diagnosis, catalogue); Brown, 1948e: 108 (diagnosis, review of genus); Brown, 1949d: 21 (Japan, China & Taiwan species key); Creighton, 1950a: 301 (North America species key); Chapman & Capco, 1951: 104, 107 (Asia Eneria, Strumigenys (Strumigenys) checklists); Brown, 1954k: 10 (Afrotropical species revision, key); Brown, 1958h: 38 (New Zealand species); Brown, 1962c: 238 (Neotropical species synopsis, key); Kempf, 1972a: 226, 243 (Neotropical Quadristruma, Strumigenys catalogues); Alayo, 1974: 19 (Cuba species key); Kempf, 1976b: 43 (additions to Brown, 1962c key); Smith, D.R. 1979: 1403, 1409 (North America Strumigenys, Quadristruma catalogues); Bolton, 1983: 358, 400 (Afrotropical Strumigenys, Quadristruma species revisions, keys); Taylor & Brown, D.R. 1985: 85, 87 (Australia Quadristruma, Strumigenys catalogues); Taylor, 1987a: 66, 76 (Australia, New Caledonia & New Zealand checklists); Terayama & Kubota, 1989: 780 (Taiwan species key); Brandão, 1991: 380 (Neotropical catalogue); Arakelian & Dlussky, 1991: 150 (former U.S.S.R. species key); Morisita, Kubota, et al. 1992: 63 (Japan species key); Dlussky, 1993: 53 (Fiji Is, Tonga and Samoa species key); Bolton, 1994: 105 (synoptic classification); Bolton, 1995a: 1053 (census); Bolton, 1995b: 377, 395 (catalogue); Gronenberg, 1996: 2021 (mandible morphology); Lin & Wu, 1996: 139 (Taiwan species key); Lattke & Goitía, 1997: 370 (Venezuela species key); Shattuck, 1999: 160, 167 (Australia synopsis); Bolton, 1999: 1657, 1673 (morphology, diagnosis, review of genus, phylogeny); Bolton, 2000: 496, 583, 716, 942 (Neotropical-Nearctic, Afrotropical, Malesian-Oriental-East Palaearctic, Austral species revisions, keys); Fisher, 2000: 614 (Malagasy species revision, key); Zhou, 2001b: 78 (China, Guangxi species key); Lin & Wu, 2001: 166 (Taiwan species key); Zhou & Xu, 2003: 737 (China species key); Yoshimura & Onoyama, 2007: 676 (Japan species key); Terayama, 2009: 142 (Taiwan species key); Heterick, 2009: 176 (south-western Australia species key).

Myrmicinae Stenammini Lordomyrma TRUE TRUE FALSE

27 species

Lordomyrma Emery, 1897c: 591. Type-species: Lordomyrma furcifera, by subsequent designation of Wheeler, W.M. 1911f: 166.

Taxonomic history

[Type-species not Podomyrma caledonica, incorrect subsequent designation by Wheeler, W.M. 1919f: 98.]

Lordomyrma in Myrmicinae, Myrmicini: Wheeler, W.M. 1910g: 139.

Lordomyrma in Myrmicinae, Myrmecinini: Emery, 1912b: 105; Emery, 1914a: 41; Forel, 1917: 244; Wheeler, W.M. 1922a: 663; Emery, 1924d: 238 [subtribe Podomyrmini]; all subsequent authors to those below; Kugler, C. 1994: 25.

Lordomyrma in Myrmicinae, Pheidolini: Dlussky & Fedoseeva, 1988: 80; Hölldobler & Wilson, 1990: 16.

Lordomyrma in Myrmicinae, Stenammini: Bolton, 1994: 106.

Junior synonyms of LORDOMYRMA

Promeranoplus Emery, 1914f: 412. Type-species: Promeranoplus rouxi, by monotypy.

Taxonomic history

Promeranoplus in Myrmicinae, Meranoplini: Emery, 1914a: 41; Forel, 1917: 244; Wheeler, W.M. 1922a: 663; Emery, 1924d: 225; Wheeler, W.M. 1934a: 176; Wheeler, W.M. 1935a: 6 (in key); all subsequent authors to the following.

Promeranoplus in Myrmicinae, Calyptomyrmecini: Dlussky & Fedoseeva, 1988: 80.

Promeranoplus as junior synonym of Lordomyrma: Hölldobler & Wilson, 1990: 14; Bolton, 1994: 106; Taylor, 2009: 17.

Prodicroaspis Emery, 1914f: 414. Type-species: Prodicroaspis sarasini, by monotypy.

Taxonomic history

Prodicroaspis in Myrmicinae, Meranoplini: Emery, 1914a: 41; Forel, 1917: 244; Wheeler, W.M. 1922a: 664; Emery, 1924d: 223; Wheeler, W.M. 1934a: 176; Wheeler, W.M. 1935a: 6 (in key); all subsequent authors to the following.

Prodicroaspis in Myrmicinae, Calyptomyrmecini: Dlussky & Fedoseeva, 1988: 80.

Prodicroaspis as junior synonym of Lordomyrma: Hölldobler & Wilson, 1990: 14; Bolton, 1994: 106; Taylor, 2009: 16.

Genus Lordomyrma references

Wheeler, W.M. 1919f: 98 (diagnosis, all species key); Emery, 1924d: 223 (Prodicroaspis diagnosis, catalogue); Emery, 1924d: 223 (Promeranoplus diagnosis, catalogue); Emery, 1924d: 238 (diagnosis, catalogue); Donisthorpe, 1941a: 37 (all species key); Chapman & Capco, 1951: 116 (Asia checklist); Taylor & Brown, D.R. 1985: 65 (Australia catalogue); Taylor, 1987a: 37, 65, 66 (Australia, New Caledonia checklists); Bolton, 1995a: 1050 (census); Bolton, 1995b: 248 (catalogue); Kugler, C. 1997: 193 (sting structure); Shattuck, 1999: 137 (Australia synopsis); Sarnat, 2006: 13 (Fiji Is species, key); Taylor, 2009: 16 (notes on synonyms).

Myrmicinae Pheidolini Pheidole TRUE TRUE FALSE

Extant: 970 species, 142 subspecies

Fossil: 4 species

Pheidole Westwood, 1839: 219. Type-species: Atta providens, by monotypy.

Taxonomic history

Pheidole in Poneridae, Attidae: Smith, F. 1858b: 172.

Pheidole in Attidae: Smith, F. 1860a: 74; Smith, F. 1862a: 49.

Pheidole in Myrmicidae: Smith, F. 1871a: 331; Cresson, 1887: 262.

Pheidole in Myrmicidae, Pheidolidae: Emery, 1877a: 81.

Pheidole in Myrmicinae: Mayr, 1861: 69 [Myrmicidae]; Mayr, 1865: 22 [Myrmicidae]; Emery & Forel, 1879a: 463 [Myrmicidae]; Dalla Torre, 1893: 88.

Pheidole in Myrmicinae, Myrmicini: Forel, 1895b: 127; Emery, 1895j: 769; Forel, 1899c: 64; Ashmead, 1905b: 383; Wheeler, W.M. 1910g: 139; Kempf, 1972a: 183 (anachronism).

Pheidole in Myrmicinae, Pheidolini: Forel, 1893a: 165; Emery, 1914a: 40; Forel, 1917: 241; Arnold, 1920a: 414; Emery, 1921f: 77; Wheeler, W.M. 1922a: 661; all subsequent authors except Kempf, 1972a: 183, above.

Junior synonyms of PHEIDOLE

Oecophthora Heer, 1852: 15. Type-species: Oecophthora pusilla (junior synonym of Formica megacephala), by monotypy.

Taxonomic history

Oecophthora in Myrmicinae: Mayr, 1855: 453 [Myrmicidae].

Oecophthora as junior synonym of Pheidole: Smith, F. 1858a: 282; Smith, F. 1858b: 172; Roger, 1863b: 30.

Ischnomyrmex Mayr, 1862: 738. Type-species: Myrmica longipes, by monotypy.

Taxonomic history

Ischnomyrmex in Myrmicinae: Mayr, 1865: 18 [Myrmicidae]; Dalla Torre, 1893: 98.

Ischnomyrmex in Myrmicinae, Myrmicini: Emery, 1895j: 769; Ashmead, 1905b: 383; Wheeler, W.M. 1910g: 140.

Ischnomyrmex in Myrmicinae, Pheidolini: Emery, 1914a: 40; Forel, 1917: 241; Emery, 1922e: 113; Wheeler, W.M. 1922a: 661; all subsequent authors.

Ischnomyrmex as junior synonym of Aphaenogaster: Emery & Forel, 1879a: 461; Bingham, 1903: 270.

Ischnomyrmex as subgenus of Aphaenogaster: Forel, 1891b: 166; Dalla Torre, 1893: 98; Forel, 1899c: 59; Emery, 1908c: 313; Wheeler, W.M. 1910g: 140.

Ischnomyrmex as subgenus of Stenamma: Emery, 1895c: 298; Emery, 1895j: 769; Forel, 1902h: 439; Forel, 1903a: 693.

Ischnomyrmex as subgenus of Pheidole: Forel, 1913k: 49.

Ischnomyrmex as genus: Mayr, 1862: 738; Emery, 1914a: 40; Viehmeyer, 1914d: 601; Forel, 1917: 241; Wheeler, W.M. 1922a: 661; Emery, 1922e: 113. Stitz, 1911a: 367; Kusnezov, 1952b: 10 (in key).

Ischnomyrmex as junior synonym of Pheidole: Brown, 1973b: 181 [provisional]; Smith, D.R. 1979: 1365; Snelling, R.R. 1981: 395.

Leptomyrma Motschoulsky, 1863: 17. Type-species: Leptomyrma gracilipes, by monotypy.

Taxonomic history

Leptomyrma in Myrmicidae: Smith, F. 1871a: 329.

Leptomyrma as junior synonym of Pheidole: Emery, 1892b: 166; all subsequent authors.

Pheidolacanthinus Smith, F. 1865: 75. Type-species: Pheidolacanthinus armatus (junior synonym of Myrmica quadrispinosa), by monotypy.

Taxonomic history

Pheidolacanthinus in Myrmicidae: Smith, F. 1871a: 333.

Pheidolacanthinus in Myrmicinae: Dalla Torre, 1893: 98.

Pheidolacanthinus as subgenus of Pheidole: Emery, 1915i: 190; Forel, 1917: 241; Emery, 1921f: 81; Wheeler, W.M. 1922a: 673; all subsequent authors to the following.

Pheidolacanthinus as junior synonym of Pheidole: Brown, 1973b: 183 [provisional]; Smith, D.R. 1979: 1365; Snelling, R.R. 1981: 395.

Ceratopheidole Pergande, 1896: 889 [as subgenus of Pheidole]. Type-species: Pheidole (Ceratopheidole) granulata, by monotypy.

Taxonomic history

Ceratopheidole in Myrmicinae, Pheidolini: Emery, 1914a: 40; Forel, 1917: 241; Emery, 1922e: 112; Wheeler, W.M. 1922a: 661; all subsequent authors.

Ceratopheidole as genus: Emery, 1914a: 40; Forel, 1917: 241; Emery, 1922e: 112; Wheeler, W.M. 1922a: 661; Chapman & Capco, 1951: 135; Wu, J. & Wang, 1995: 104 (anachronism).

Ceratopheidole as subgenus of Pheidole: Pergande, 1896: 889; Kempf, 1972a: 183.

Ceratopheidole as junior synonym of Pheidole: Brown, 1973b: 179 [provisional]; Smith, D.R. 1979: 1365; Snelling, R.R. 1981: 395; Wilson, 2003: 6.

Epipheidole Wheeler, W.M. 1903f: 664. Type-species: Epipheidole inquilina, by monotypy.

Taxonomic history

[Epipheidole also described as new by Wheeler, W.M. 1904a: 14.]

Epipheidole in Myrmicinae, Stenammini: Ashmead, 1905b: 383.

Epipheidole in Myrmicinae, Myrmicini: Wheeler, W.M. 1910g: 140.

Epipheidole in Myrmicinae, Pheidolini: Emery, 1914a: 40; Forel, 1917: 241; Emery, 1922e: 114; Wheeler, W.M. 1922a: 661; Creighton, 1950a: 192.

Epipheidole as junior synonym of Pheidole: Cole, 1965: 174; Smith, D.R. 1979: 1366.

Phidole Bingham, 1903: 220, unjustified emendation of Pheidole.

Taxonomic history

Phidole as junior synonym of Pheidole: Wheeler, 1922a: 806.

Sympheidole Wheeler, W.M. 1904a: 7. Type-species: Sympheidole elecebra, by monotypy.

Taxonomic history

Sympheidole in Myrmicinae, Myrmicini: Wheeler, W.M. 1910g: 140.

Sympheidole in Myrmicinae, Stenammini: Ashmead, 1905b: 383.

Sympheidole in Myrmicinae, Pheidolini: Emery, 1914a: 40; Forel, 1917: 241; Emery, 1922e: 115; Wheeler, W.M. 1922a: 661; all subsequent authors to the following.

Sympheidole as junior synonym of Pheidole: Brown, 1973b: 185 [provisional]; Smith, D.R. 1979: 1365; Snelling, R.R. 1981: 395; Wilson, 2003: 6.

Phidola Schulz, W.A. 1906: 155, unjustified emendation of Pheidole.

Taxonomic history

Phidola as junior synonym of Pheidole: Smith, D.R. 1979: 1365.

Allopheidole Forel, 1912f: 237 [as subgenus of Pheidole]. Type-species: Pheidole kingi (junior synonym of Pheidole tepicana), by subsequent designation of Wheeler, W.M. 1913a: 79.

Taxonomic history

Allopheidole as subgenus of Pheidole: Forel, 1912f: 237; Forel, 1917: 241.

Allopheidole as junior synonym of Pheidole: Emery, 1915i: 190; Emery, 1921f: 84; Smith, D.R. 1979: 1366.

Decapheidole Forel, 1912f: 237 [as subgenus of Pheidole]. Type-species: Pheidole perpusilla, by subsequent designation of Wheeler, W.M. 1913a: 80.

Taxonomic history

Decapheidole in Myrmicinae, Pheidolini: Wheeler, W.M. 1922a: 661.

Decapheidole as genus: Wheeler, W.M. 1922a: 661; Borgmeier, 1927c: 78.

Decapheidole as subgenus of Pheidole: Forel, 1912f: 237, Forel, 1917: 241; Emery, 1922e: 112; Kempf, 1972a: 183.

Decapheidole as junior synonym of Pheidole: Brown, 1973b: 180 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395; Wilson, 2003: 6.

Isopheidole Forel, 1912m: 765 [as subgenus of Pheidole]. Type-species: Myrmica longipes, by monotypy.

Taxonomic history

Isopheidole as junior synonym of Ischnomyrmex: Forel, 1913b: 350 (in text); Forel, 1913k: 49; Emery, 1922e: 113; Wheeler, W.M. 1922a: 680 (in key). [Ischnomyrmex and Isopheidole share the same type-species, synonymy is therefore absolute.]

Elasmopheidole Forel, 1913k: 43 [as subgenus of Pheidole]. Type-species: Pheidole aberrans, by subsequent designation of Emery, 1915i: 190.

Taxonomic history

Elasmopheidole as subgenus of Pheidole: Forel, 1913k: 43; Emery, 1915i: 190; Forel, 1917: 241; Emery, 1922e: 110; Wheeler, W.M. 1922a: 673; all subsequent authors to the following.

Elasmopheidole as junior synonym of Pheidole: Brown, 1973b: 180 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395.

Cardiopheidole Wheeler, W.M. 1914b: 48 [as subgenus of Pheidole]. Type-species: Pheidole vaslitii, by original designation.

Taxonomic history

Cardiopheidole as subgenus of Pheidole: Wheeler, W.M. 1914b: 48; Forel, 1917: 241.

Cardiopheidole as junior synonym of Pheidole: Emery, 1915i: 190; Emery, 1921f: 84; Smith, D.R. 1979: 1366.

Anergatides Wasmann, 1915b: 281. Type-species: Anergatides kohli (junior secondary homonym in Pheidole, replaced by Pheidole neokohli), by monotypy.

Taxonomic history

Anergatides in Myrmicinae, Pheidolini: Forel, 1917: 243 [subtribe Anergatini]; Emery, 1922e: 116 [subtribe Anergatidini]; Donisthorpe, 1943f: 623.

Anergatides in Myrmicinae, Solenopsidini: Wheeler, W.M. 1922a: 663; Wheeler, G.C. & Wheeler, J. 1985: 257 (anachronism).

Anergatides as junior synonym of Pheidole: Brown, 1973b: 178 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395; Wilson, 1984: 328; Bolton, 1994: 106.

Parapheidole Emery, 1915d: 68. Type-species: Aphaenogaster oculata, by monotypy.

Taxonomic history

Parapheidole in Myrmicinae, Pheidolini: Forel, 1917: 241; Emery, 1922e: 115; Wheeler, W.M. 1922a: 661; all subsequent authors.

Parapheidole as junior synonym of Pheidole: Brown, 1973b: 183 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395.

Macropheidole Emery, 1915i: 190 [as subgenus of Pheidole]. Type-species: Pheidole fimbriata, by monotypy.

Taxonomic history

Macropheidole as subgenus of Pheidole: Emery, 1915i: 190; Forel, 1917: 241; Emery, 1921f: 81; Wheeler, W.M. 1922a: 673; all subsequent authors to the following.

Macropheidole as junior synonym of Pheidole: Smith, M.R. 1951a: 800; Brown, 1973b: 181 [provisional]; Smith, D.R. 1979: 1366.

Scrobopheidole Emery, 1915i: 190 [as subgenus of Pheidole]. Type-species: Pheidole scrobifera, by monotypy.

Taxonomic history

Scrobopheidole as subgenus of Pheidole: Forel, 1917: 241; Emery, 1922e: 112; Wheeler, W.M. 1922a: 673; all subsequent authors to the following.

Scrobopheidole as junior synonym of Pheidole: Brown, 1973b: 184 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395; Wilson, 2003: 6.

Stegopheidole Emery, 1915i: 190 [as subgenus of Pheidole]. Type-species: Pheidole (Elasmopheidole) upeneci, by monotypy.

Taxonomic history

Stegopheidole as subgenus of Pheidole: Emery, 1921f: 83; Wheeler, W.M. 1922a: 673; all subsequent authors to the following.

Stegopheidole as junior synonym of Pheidole: Brown, 1973b: 185 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395.

Trachypheidole Emery, 1915i: 190 [as subgenus of Pheidole]. Type-species: Pheidole bicornis, by original designation.

Taxonomic history

Trachypheidole as subgenus of Pheidole: Forel, 1917: 241; Emery, 1922e: 111; Wheeler, W.M. 1922a: 673; all subsequent authors to the following.

Trachypheidole as junior synonym of Pheidole: Brown, 1973b: 185 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395; Wilson, 2003: 6.

Electropheidole Mann, 1921: 438 [as subgenus of Pheidole]. Type-species: Pheidole (Electropheidole) roosevelti, by subsequent designation of Donisthorpe, 1943f: 642.

Taxonomic history

Electropheidole as subgenus of Pheidole: Mann, 1921: 438; Wheeler, W.M. 1922a: 672; all subsequent authors to the following.

Electropheidole as junior synonym of Pheidole: Brown, 1973b: 180 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395.

Bruchomyrma Santschi, 1922d: 248. Type-species: Bruchomyrma acutidens, by monotypy.

Taxonomic history

Bruchomyrma in Myrmicinae, Pheidolini: Donisthorpe, 1943f: 629.

Bruchomyrma in Myrmicinae, Myrmicini: Kempf, 1972a: 42.

Bruchomyrma as junior synonym of Pheidole: Brown, 1973b: 179 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395; Wilson, 1984: 327; Bolton, 1994: 106; Wilson, 2003: 6.

Bruchomyrma as genus: Dlussky & Fedoseeva, 1988: 80 (anachronism).

Cephalomorium Forel, 1922: 91 [as subgenus of Tetramorium]. Type-species: Tetramorium (Cephalomorium) bahai, by monotypy.

Taxonomic history

Cephalomorium as junior synonym of Hendecapheidole: Santschi, 1925d: 228.

Hendecapheidole Wheeler, W.M. 1922d: 3 [as subgenus of Pheidole]. Type-species: Pheidole tachigaliae, by original designation.

Taxonomic history

Hendecapheidole as junior synonym of Pheidole: Brown, 1973b: 181 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395; Wilson, 2003: 6.

Gallardomyrma Bruch, 1932: 271. Type-species: Gallardomyrma argentina, by original designation.

Taxonomic history

Gallardomyrma in Myrmicinae, Pheidolini: Donisthorpe, 1943f: 646.

Gallardomyrma in Myrmicinae, Myrmicini: Kempf, 1972a: 110.

Gallardomyrma as junior synonym of Pheidole: Brown, 1973b: 180 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395; Wilson, 1984: 327; Wilson, 2003: 6.

Gallardomyrma as genus: Kempf, 1972a: 110; Dlussky & Fedoseeva, 1988: 80 (anachronism).

Conothoracoides Strand, 1935: 176.

Taxonomic history

[Replacement name for Conothorax Karavaiev; junior homonym of Conothorax Jekel, 1854: 9 bis (foldout pages) (Coleoptera).]

Conothoracoides as junior synonym of Pheidole: Brown, 1973b: 179 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395.

Homonym replaced by Conothoracoides

Conothorax Karavaiev, 1935a: 75. Type-species: Conothorax bilobum, by monotypy.

Taxonomic history

[Conothorax Karavaiev junior homonym of Conothorax Jekel, 1854: 9 bis (foldout pages) (Coleoptera).]

Conothorax in Myrmicinae, Pheidolini: Karavaiev, 1935a: 75.

Eriopheidole Kusnezov, 1952a: 10. Type-species: Eriopheidole symbiotica (unresolved junior secondary homonym in Pheidole), by monotypy.

Taxonomic history

Eriopheidole in Myrmicinae, Pheidolini: Kusnezov, 1952a: 10.

Eriopheidole in Myrmicinae, Myrmicini: Kempf, 1972a: 107.

Eriopheidole junior synonym of Pheidole: Brown, 1973b: 180 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395; Bolton, 1994: 106.

Eriopheidole as genus: Kempf, 1972a: 107; Dlussky & Fedoseeva, 1988: 80 (anachronism).

Xenoaphaenogaster Baroni Urbani, 1964b: 50. Type-species: Xenoaphaenogaster inquilina (junior synonym of Myrmica pallidula), by original designation.

Taxonomic history

Xenoaphaenogaster as junior synonym of Monomorium: Brown, 1973b: 185 [provisional].

Xenoaphaenogaster as junior synonym of Pheidole: Bolton, 1987: 291.

Genus Pheidole references

Smith, F. 1858b: 172 (diagnosis); Mayr, 1861: 70 (Europe species key); Roger, 1863b: 27, 30 (catalogue); Mayr, 1863: 424, 440 (Ischnomyrmex, Pheidole catalogues); Mayr, 1865: 18, 22 (Ischnomyrmex, Pheidole diagnoses); Mayr, 1867a: 92, 96 (Ischnomyrmex, Pheidole diagnoses); Mayr, 1870b: 977, 979 (Australia & New World species keys); Mayr, 1876: 102 (Australia species key); André, 1883b: 382 (Europe & Algeria species key); Mayr, 1879: 674 (Asia species key); Mayr, 1887: 582 (New World species key); Cresson, 1887: 262 (U.S.A., catalogue); Nasonov, 1889: 75 (Russia species key); Dalla Torre, 1893: 88, 98 (Pheidole, Pheidolacanthinus catalogues); Emery, 1895c: 288 (North America species key); Mayr, 1896: 241 (Africa species, partial key); Emery, 1896g: 80 (P. flavens group, key); Forel, 1902c: 165 (India species key); Forel, 1902f: 523 (India & Sri Lanka species key); Bingham, 1903: 223 (India, Sri Lanka & Burma species key); Wheeler, W.M. 1908a: 133 (Puerto Rico species key); Arnold, 1920a: 414 (diagnosis, South Africa species key); Emery, 1921f: 78 (diagnosis, subgenera key, catalogue); Emery, 1921f: 81 (P. (Macropheidole) & P. (Pheidolacanthinus) diagnoses, catalogues); Emery, 1921f: 83 (P. (Stegopheidole) diagnosis, catalogue); Mann, 1921: 442 (Fiji Is species key); Emery, 1922e: 110 (P. (Elasmopheidole) diagnosis, catalogue); Emery, 1922e: 111 (P. (Trachypheidole) diagnosis, catalogue); Emery, 1922e: 112 (P. (Scrobopheidole), P. (Decapheidole) & P. (Ceratopheidole) diagnoses, catalogues); Emery, 1922e: 113 (Ischnomyrmex diagnosis, catalogue); Emery, 1922e: 114 (Epipheidole diagnosis, catalogue); Emery, 1922e: 115 (Sympheidole, Parapheidole diagnoses, catalogues); Emery, 1922e: 116 (Anergatides diagnosis, catalogue); Wheeler, W.M. 1922a: 126, 672 (diagnosis, subgenera key); Wheeler, W.M. 1922a: 806, 879 (Afrotropical Pheidole, Anergatides catalogues); Wheeler, W.M. 1922a: 1017, 1021 (Malagasy Pheidole, Parapheidole catalogues); Gallardo, 1932c: 179 (Argentina Elasmopheidole species key); Finzi, 1936: 165 (Egypt species key); Menozzi, 1939a: 298 (Himalaya species key); Cole, 1942: 362 (U.S.A., Utah species key); Buren, 1944a: 285 (U.S.A., Iowa species key); Creighton, 1950a: 163 (North America species key); Creighton, 1950a: 192, 194 (North America Epipheidole, Sympheidole species); Chapman & Capco, 1951: 135, 137 (Asia Ceratopheidole, Conothorax, Ischnomyrmex, Pheidole checklists); Kusnezov, 1952b: 61 (Argentina species key); Smith, M.R. 1955c: 302 (P. (Hendecapheidole) species key); Gregg, 1959: 9 (U.S.A. species key); Gregg, 1963: 407 (U.S.A., Colorado species key); Wheeler, G.C. & Wheeler, J. 1963: 126 (U.S.A., North Dakota species key); Bernard, 1967: 149 (diagnosis, Western Europe species key); Gregg, 1969: 95 (addition to Gregg, 1959 U.S.A. key); Kempf, 1972a: 183 (Neotropical catalogue); Alayo, 1974: 11 (Cuba species key); Smith, D.R. 1979: 1365 (North America catalogue); Allred, 1982: 441 (U.S.A., Utah species key); Ogata, 1982: 191 (Japan species key); Collingwood, 1985: 253 (Saudi Arabia species key); Naves, 1985: 54 (U.S.A., Florida species key); Taylor & Brown, D.R. 1985: 74 (Australia catalogue); Wheeler, G.C. & Wheeler, J. 1986g: 41 (U.S.A., Nevada species key); Taylor, 1987a: 53 (Australia, New Caledonia & New Zealand checklists); Agosti & Collingwood, 1987b: 271 (Balkans species key); Hölldobler & Wilson, 1990: 15 (synoptic classification); Kupyanskaya, 1990: 123 (Far Eastern Russia species key); Brandão, 1991: 368 (Neotropical catalogue); Morisita, Kubota, et al. 1992: 22 (Japan species key); Bolton, 1994: 106 (synoptic classification); Kupyanskaya, 1995: 345 (Far Eastern Russia species key); Bolton, 1995a: 1051 (census); Bolton, 1995b: 316 (catalogue); Wu, J. & Wang, 1995: 99 (China species key); Collingwood & Agosti, 1996: 322 (Saudi Arabia species key); Park & Kim, 2000: 109 (Korea species key); Shattuck, 1999: 152 (Australia synopsis); Eguchi, 1999: 103 (Borneo P. longipes group, key); Ward, 2000: 96 (U.S.A. P. hyatti complex, key); Zhou, 2001b: 121 (China, Guangxi species key); Eguchi, 2001b: 10 (Borneo species key); ; Mackay & Mackay, 2002: 149 (U.S.A., New Mexico species key)Wilson, 2003: 27 (New World, all species revision, keys); Sarnat, 2008: 8 (Fiji Is P. roosevelti group, key); Eguchi, 2008: 8 (North Vietnam species key); Terayama, 2009: 171 (Taiwan species key); Heterick, 2009: 169 (south-western Australia species key); Sarnat & Moreau, 2011: 114 (P. roosevelti group phylogeny and radiation).

Genera incertae sedis in Pheidolini

Myrmicinae Adelomyrmecini Adelomyrmex hirsutus TRUE TRUE FALSE

hirsutus. Adelomyrmex (Arctomyrmex) hirsutus Mann, 1921: 458, fig. 21 (w.). FIJI IS. See also: Fernández, 2003b: 18.

Formicinae Camponotini Camponotus bryani TRUE TRUE FALSE

bryani. Camponotus (Condylomyrma) bryani Santschi, 1928c: 72, fig. 2 (w.) FIJI IS. Combination in C. (Colobopsis): Wheeler, W.M. 1934e: 423.

Formicinae Camponotini Camponotus cristatus TRUE TRUE FALSE

2 subspecies

cristatus. Camponotus cristatus Mayr, 1866a: 489, fig. 3 (w.) FIJI IS. Mann, 1921: 479 (s.q.m.). Combination in C. (Myrmogonia): Forel, 1912i: 92. Current subspecies: nominal plus nagasau, sadinus.

Formicinae Camponotini Camponotus dentatus TRUE TRUE FALSE

1 subspecies

dentatus. Colobopsis dentata Mayr, 1866a: 492, fig. 5 (w.) FIJI IS. Mann, 1921: 491 (s.q.m.). Combination in Camponotus: Dalla Torre, 1893: 228; in C. (Myrmamblys): Forel, 1914a: 271; in C. (Myrmotemnus): Emery, 1920b: 258; in C. (Colobopsis): Mann, 1921: 491. Current subspecies: nominal plus fijianus.

Formicinae Camponotini Camponotus laminatus TRUE TRUE FALSE

1 subspecies

laminatus. Camponotus laminatus Mayr, 1866a: 489, fig. 4 (w.) FIJI IS. Mann, 1921: 477 (s.q.m.). Combination in C. (Myrmogonia): Forel, 1912i: 92. Current subspecies: nominal plus levuanus.

Formicinae Camponotini Camponotus lauensis TRUE TRUE FALSE

lauensis. Camponotus (Myrmogonia) lauensis Mann, 1921: 488, fig. 35 (w.) FIJI IS.

Formicinae Camponotini Camponotus maafui TRUE TRUE FALSE

maafui. Camponotus (Myrmogonia) maafui Mann, 1921: 482, fig. 31 (s.w.) FIJI IS.

Formicinae Camponotini Camponotus manni TRUE TRUE FALSE

1 subspecies

manni. Camponotus (Colobopsis) manni Wheeler, W.M. 1934e: 418, fig. 4 (w.) FIJI IS. Current subspecies: nominal plus umbratilis.

Formicinae Camponotini Camponotus maudella TRUE TRUE FALSE

1 subspecies

maudella. Camponotus (Colobopsis) maudella Mann, 1921: 496, fig. 38 (s.w.q.m.) FIJI IS. Current subspecies: nominal plus seemanni.

Formicinae Camponotini Camponotus oceanicus TRUE TRUE FALSE

oceanicus. Colobopsis oceanica Mayr, 1870b: 943 (q.) FIJI IS. Combination in Camponotus: Dalla Torre, 1893: 245; in C. (Myrmamblys): Forel, 1914a: 272; in C.(Myrmotemnus): Emery, 1920b: 258; in C. (Colobopsis): Emery, 1925b: 146.

Formicinae Camponotini Camponotus rotumanus TRUE TRUE FALSE

rotumanus. Camponotus (Myrmamblys) rotumanus Wilson & Taylor, 1967: 98, fig. 82 (w.) FIJI IS.

Formicinae Camponotini Camponotus schmeltzi TRUE TRUE FALSE

3 subspecies

schmeltzi. Camponotus schmeltzi Mayr, 1866a: 490 (w.) FIJI IS. Mann, 1921: 484 (s.). Combination in C. (Myrmogonia): Forel, 1912i: 92. Current subspecies: nominal plus kadi, loloma, trotteri.

Formicinae Camponotini Camponotus vitiensis TRUE TRUE FALSE

vitiensis. Camponotus (Colobopsis) vitiensis Mann, 1921: 490, fig. 36 (w.) FIJI IS. Donisthorpe, 1946b: 69 (s.q.m.).

Formicinae Camponotini Camponotus manni umbratilis TRUE TRUE FALSE

umbratilis. Camponotus (Colobopsis) manni subsp. umbratilis Wheeler, W.M. 1934e: 420, fig. 4 (w.m.) FIJI IS.

Formicinae Camponotini Camponotus schmeltzi trotteri TRUE TRUE FALSE

trotteri. Camponotus (Myrmogonia) schmeltzii subsp. trotteri Mann, 1921: 486, fig. 34 (w.) FIJI IS.

Formicinae Camponotini Camponotus maudella seemanni TRUE TRUE FALSE

seemanni. Camponotus (Colobopsis) maudella var. seemanni Mann, 1921: 498 (s.w.q.) FIJI IS.

Formicinae Camponotini Camponotus cristatus sadinus TRUE TRUE FALSE

sadinus. Camponotus (Myrmogonia) cristatus subsp. sadina Mann, 1921: 482 (s.w.) FIJI IS.

Formicinae Camponotini Camponotus schmeltzi kadi TRUE TRUE FALSE

kadi. Camponotus (Myrmogonia) schmeltzii var. kadi Mann, 1921: 485 (s.w.m.) FIJI IS.

Formicinae Camponotini Camponotus laminatus levuanus TRUE TRUE FALSE

levuanus. Camponotus (Myrmogonia) laminatus var. levuanus Mann, 1921: 479 (w.) FIJI IS.

Formicinae Camponotini Camponotus schmeltzi loloma TRUE TRUE FALSE

loloma. Camponotus (Myrmogonia) schmeltzii var. loloma Mann, 1921: 486, fig. 33 (w.) FIJI IS.

Formicinae Camponotini Camponotus cristatus nagasau TRUE TRUE FALSE

nagasau. Camponotus (Myrmogonia) cristatus var. nagasau Mann, 1921: 482 (s.w.) FIJI IS.

Myrmicinae Formicoxenini Cardiocondyla nuda TRUE TRUE FALSE

1 subspecies

nuda. Leptothorax nudus Mayr, 1866a: 508 (w.) FIJI IS. Emery, 1897c: 588 (q. ergatoid m.); Wheeler, G.C. & Wheeler, J. 1973a: 27 (l.); Imai, et al. 1984: 6 (k.). Combination in Cardiocondyla: Forel, 1881: 6. Current subspecies: nominal plus sculptinodis. See also: Seifert, 2003a: 245.

Cerapachyinae Cerapachyini Cerapachys cryptus TRUE TRUE FALSE

cryptus. Cerapachys (Syscia) cryptus Mann, 1921: 408, fig. 1 (w.q.) FIJI IS. Wheeler, G.C. 1950: 105 (l.).

Cerapachyinae Cerapachyini Cerapachys fuscior TRUE TRUE FALSE

fuscior. Cerapachys (Syscia) cryptus subsp. fuscior Mann, 1921: 410 (w.) FIJI IS. Raised to species: Wilson, 1959b: 55.

Cerapachyinae Cerapachyini Cerapachys lindrothi TRUE TRUE FALSE

lindrothi. Cerapachys (Cerapachys) lindrothi Wilson, 1959b: 52 (w.q.) FIJI IS.

Cerapachyinae Cerapachyini Cerapachys majusculus TRUE TRUE FALSE

majusculus. Cerapachys (Cerapachys) majusculus Mann, 1921: 408 (w.) FIJI IS.

Cerapachyinae Cerapachyini Cerapachys sculpturatus TRUE TRUE FALSE

sculpturatus. Cerapachys (Cerapachys) vitiensis subsp. sculpturatus Mann, 1921: 407 (w.) FIJI IS. Raised to species: Wilson, 1959b: 53.

Cerapachyinae Cerapachyini Cerapachys vitiensis TRUE TRUE FALSE

vitiensis. Cerapachys (Cerapachys) vitiensis Mann, 1921: 406 (w.) FIJI IS.

Cerapachyinae Cerapachyini Cerapachys zimmermani TRUE TRUE FALSE

zimmermani. Cerapachys (Cerapachys) zimmermani Wilson, 1959b: 54 (w.) FIJI IS.

Myrmicinae Basicerotini Eurhopalothrix insidiatrix TRUE TRUE FALSE

insidiatrix. Eurhopalothix insidiatrix Taylor, 1980b: 238, figs. 11-14 (w.q.) FIJI IS.

Ectatomminae Ectatommini Gnamptogenys aterrima TRUE TRUE FALSE

aterrima. Wheeleripone aterrima Mann, 1921: 411, fig. 2 (w.q.m.) FIJI IS. Combination in Gnamptogenys: Brown, 1958g: 227. See also: Lattke, 2004: 60.

Ponerinae Ponerini Hypoponera monticola TRUE TRUE FALSE

monticola. Ponera monticola Mann, 1921: 418, fig. 5 (w.) FIJI IS. Santschi, 1928c: 67 (q.). Combination in Hypoponera: Taylor, 1967a: 12.

Ponerinae Ponerini Hypoponera turaga TRUE TRUE FALSE

turaga. Ponera turaga Mann, 1921: 416 (w.q.) FIJI IS. Combination in Hypoponera: Dlussky, 1994: 53 (in list); Bolton, 1995b: 216.

Ponerinae Ponerini Hypoponera vitiensis TRUE TRUE FALSE

vitiensis. Ponera vitiensis Mann, 1921: 414, fig. 4 (w.q.) FIJI IS. Combination in Hypoponera: Bolton, 1995b: 216. Given as junior synonym of confinis by Dlussky, 1994: 53 (in list only); unconfirmed.

Ponerinae Ponerini Leptogenys foveopunctata TRUE TRUE FALSE

foveopunctata. Leptogenys (Lobopelta) foveopunctata Mann, 1921: 421 (w.) FIJI IS.

Ponerinae Ponerini Leptogenys fugax TRUE TRUE FALSE

fugax. Leptogenys (Lobopelta) fugax Mann, 1921: 422, fig. 7 (w.) FIJI IS.

Ponerinae Ponerini Leptogenys humiliata TRUE TRUE FALSE

humiliata. Leptogenys (Lobopelta) humiliata Mann, 1921: 421 (w.) FIJI IS.

Ponerinae Ponerini Leptogenys letilae TRUE TRUE FALSE

letilae. Leptogenys (Lobopelta) letilae Mann, 1921: 419, fig. 6 (w.q.m.) FIJI IS.

Ponerinae Ponerini Leptogenys navua TRUE TRUE FALSE

navua. Leptogenys (Lobopelta) navua Mann, 1921: 423, figs. 8, 9 (w.m.) FIJI IS.

Ponerinae Ponerini Leptogenys vitiensis TRUE TRUE FALSE

vitiensis. Leptogenys (Lobopelta) vitiensis Mann, 1921: 424, fig. 10 (w.) FIJI IS.

Myrmicinae Stenammini Lordomyrma curvata TRUE TRUE FALSE

curvata. Lordomyrma curvata Sarnat, 2006: 15, figs. 2, 3 (w.) FIJI IS.

Myrmicinae Stenammini Lordomyrma desupra TRUE TRUE FALSE

desupra. Lordomyrma desupra Sarnat, 2006: 17, figs. 4, 5 (w.) FIJI IS.

Myrmicinae Stenammini Lordomyrma levifrons TRUE TRUE FALSE

levifrons. Rogeria (Irogera) tortuosa subsp. levifrons Mann, 1921: 453 (w.) FIJI IS. Combination in Lordomyrma: Kugler, C. 1994: 26. Raised to species: Sarnat, 2006: 20.

Myrmicinae Stenammini Lordomyrma polita TRUE TRUE FALSE

polita. Rogeria (Irogera) tortuosa subsp. polita Mann, 1921: 453 (w.) FIJI IS. Combination in Lordomyrma: Kugler, C. 1994: 26. Raised to species: Sarnat, 2006: 21.

Myrmicinae Stenammini Lordomyrma rugosa TRUE TRUE FALSE

rugosa. Rogeria (Irogera) rugosa Mann, 1921: 455, fig. 20 (w.) FIJI IS. Combination in Lordomyrma: Kugler, C. 1994: 26. See also: Sarnat, 2006: 23.

Myrmicinae Stenammini Lordomyrma stoneri TRUE TRUE FALSE

stoneri. Rogeria (Irogera) tortuosa subsp. stoneri Mann, 1925b: 5 (w.) FIJI IS. Combination in Lordomyrma: Kugler, C. 1994: 26. Raised to species: Sarnat, 2006: 25.

Myrmicinae Stenammini Lordomyrma striatella TRUE TRUE FALSE

striatella. Rogeria (Irogera) striatella Mann, 1921: 454, fig. 19 (w.m.) FIJI IS. Combination in Lordomyrma: Kugler, C. 1994: 26. See also: Sarnat, 2006: 27.

Myrmicinae Stenammini Lordomyrma sukuna TRUE TRUE FALSE

sukuna. Lordomyrma sukuna Sarnat, 2006: 29, figs. 16, 17 (w.) FIJI IS.

Myrmicinae Stenammini Lordomyrma tortuosa TRUE TRUE FALSE

tortuosa. Rogeria (Irogera) tortuosa Mann, 1921: 452, fig. 18 (w.) FIJI IS. Combination in Lordomyrma: Kugler, C. 1994: 26. See also: Sarnat, 2006: 33.

Myrmicinae Stenammini Lordomyrma vanua TRUE TRUE FALSE

vanua. Lordomyrma vanua Lucky & Sarnat, 2008: 42, figs. 1, 4, 5 (w.) FIJI IS.

Myrmicinae Stenammini Lordomyrma vuda TRUE TRUE FALSE

vuda. Lordomyrma vuda Sarnat, 2006: 34, figs. 20, 21 (w.) FIJI IS.

Myrmicinae Solenopsidini Monomorium vitiense TRUE TRUE FALSE

vitiense. Monomorium (Monomorium) vitiensis Mann, 1921: 444 (w.) FIJI IS. Given as junior synonym of talpa by Dlussky, 1994: 54 (in list only); unconfirmed.

Myrmicinae Myrmecinini Myrmecina cacabau TRUE TRUE FALSE

cacabau. Archaeomyrmex cacabau Mann, 1921: 449, fig. 17 (w.) FIJI IS. Combination in Myrmecina: Brown, 1971a: 1. See also: Taylor, 1980d: 122.

Formicinae Plagiolepidini Nylanderia vitiensis TRUE TRUE FALSE

vitiensis. Prenolepis (Nylanderia) vitiensis Mann, 1921: 474, fig. 28 (w.m.) FIJI IS. Combination in Paratrechina (Nylanderia): Emery, 1925b: 221; in Nylanderia: LaPolla, Brady & Shattuck, 2010a: 127. Given as junior synonym of vaga by Dlussky, 1994: 55 (in list only); unconfirmed.

Dolichoderinae Leptomyrmecini Ochetellus sororis TRUE TRUE FALSE

sororis. Iridomyrmex sororis Mann, 1921: 469, fig. 26 (w.q.m.) FIJI IS. Combination in Ochetellus: Shattuck, 1992a: 17.

Ponerinae Ponerini Odontomachus angulatus TRUE TRUE FALSE

angulatus. Odontomachus angulatus Mayr, 1866a: 500, pl., fig. 10 (w.) FIJI IS. Mann, 1921: 427 (q.). Senior synonym of politus: Brown, 1976a: 102. See also: Brown, 1976a: 124.

Ponerinae Ponerini Odontomachus simillimus TRUE TRUE FALSE

simillimus. Odontomachus simillimus Smith, F. 1858b: 80, pl. 5, figs. 8, 9 (q.) FIJI IS. Mayr, 1867a: 79 (w.); Karavaiev, 1925c: 294 (m.); Wheeler, G.C. & Wheeler, J. 1980: 530 (l.); Imai, Brown, et al. 1984: 67 (k.); Tjan, Imai, et al. 1986: 57 (k.). Junior synonym of haematodus: Roger, 1861a: 24; Mayr, 1865: 64; Emery, 1911d: 114. Revived from synonymy: Wilson, 1959a: 499; Brown, 1976a: 106. Senior synonym of fuscipennis: Wilson, 1959a: 499; of breviceps, pallidicornis: Brown, 1976a: 106. See also: Brown, 1976a: 165; Fisher & Smith, 2008: 15; Sorger & Zettel, 2011: 158.

Formicinae Plagiolepidini Paraparatrechina oceanica TRUE TRUE FALSE

oceanica. Prenolepis (Nylanderia) oceanica Mann, 1921: 476 (w.) FIJI IS. Combination in Paratrechina (Nylanderia): Emery, 1925b: 221; in Paraparatrechina: LaPolla, Brady & Shattuck, 2010a: 128.

Myrmicinae Pheidolini Pheidole bula TRUE TRUE FALSE

bula. Pheidole bula Sarnat, 2008: 12, figs. 32-34, 53-55, 74-76 (s.w.q.) FIJI IS.

Myrmicinae Pheidolini Pheidole caldwelli TRUE TRUE FALSE

caldwelli. Pheidole caldwelli Mann, 1921: 434, fig. 14 (s.w.q.) FIJI IS.

Myrmicinae Pheidolini Pheidole colaensis TRUE TRUE FALSE

colaensis. Pheidole (Electropheidole) colaensis Mann, 1921: 441 (s.w.) FIJI IS. Sarnat, 2008: 17 (q.).

Myrmicinae Pheidolini Pheidole furcata TRUE TRUE FALSE

furcata. Pheidole furcata Sarnat, 2008: 19, figs. 38-40, 59-61, 80-82 (s.w.q.) FIJI IS.

Myrmicinae Pheidolini Pheidole knowlesi TRUE TRUE FALSE

1 subspecies

knowlesi. Pheidole knowlesi Mann, 1921: 436, fig. 13 (s.w.q.) FIJI IS. Current subspecies: nominal plus extensa.

Myrmicinae Pheidolini Pheidole oceanica TRUE TRUE FALSE

oceanica. Pheidole oceanica Mayr, 1866a: 510 (s.w.q.m.) FIJI IS. Senior synonym of boraborensis, pattesoni, upoluana: Wilson & Taylor, 1967: 48.

Myrmicinae Pheidolini Pheidole onifera TRUE TRUE FALSE

onifera. Pheidole onifera Mann, 1921: 427, fig. 12 (s.w.) FIJI IS.

Myrmicinae Pheidolini Pheidole pegasus TRUE TRUE FALSE

pegasus. Pheidole pegasus Sarnat, 2008: 21, figs. 41-43, 62-64, 83-85 (s.w.q.) FIJI IS.

Myrmicinae Pheidolini Pheidole roosevelti TRUE TRUE FALSE

roosevelti. Pheidole (Electropheidole) roosevelti Mann, 1921: 438, fig. 15 (s.w.q.) FIJI IS. See also: Sarnat, 2008: 22.

Myrmicinae Pheidolini Pheidole simplispinosa TRUE TRUE FALSE

simplispinosa. Pheidole simplispinosa Sarnat, 2008: 26, figs. 47-49, 68-70, 89-91 (s.w.q.) FIJI IS.

Myrmicinae Pheidolini Pheidole uncagena TRUE TRUE FALSE

uncagena. Pheidole uncagena Sarnat, 2008: 28, figs. 50-52, 71-73 (s.w.) FIJI IS.

Myrmicinae Pheidolini Pheidole vatu TRUE TRUE FALSE

vatu. Pheidole vatu Mann, 1921: 431, fig. 13 (s.w.) FIJI IS.

Myrmicinae Pheidolini Pheidole wilsoni TRUE TRUE FALSE

wilsoni. Pheidole wilsoni Mann, 1921: 433 (s.w.) FIJI IS.

Myrmicinae Pheidolini Pheidole knowlesi extensa TRUE TRUE FALSE

extensa. Pheidole knowlesi subsp. extensa Mann, 1921: 438 (s.w.) FIJI IS.

Dolichoderinae Leptomyrmecini Philidris nagasau TRUE TRUE FALSE

2 subspecies

nagasau. Iridomyrmex nagasau Mann, 1921: 470, fig. 27 (w.) FIJI IS. Combination in Philidris: Shattuck, 1992a: 18. Current subspecies: nominal plus agnata, alticola.

Dolichoderinae Leptomyrmecini Philidris nagasau alticola TRUE TRUE FALSE

alticola. Iridomyrmex nagasau subsp. alticola Mann, 1921: 472 (w.) FIJI IS. Combination in Philidris: Shattuck, 1992a: 18.

Dolichoderinae Leptomyrmecini Philidris nagasau agnata TRUE TRUE FALSE

agnata. Iridomyrmex nagasau subsp. agnatus Mann, 1921: 472 (w.) FIJI IS. Combination in Philidris: Shattuck, 1992a: 18.

Myrmicinae Formicoxenini Poecilomyrma senirewae TRUE TRUE FALSE

1 subspecies

senirewae. Poecilomyrma senirewae Mann, 1921: 446, fig. 16 (w.) FIJI IS. Current subspecies: nominal plus myrmecodiae.

Myrmicinae Formicoxenini Poecilomyrma senirewae myrmecodiae TRUE TRUE FALSE

myrmecodiae. Poecilomyrma senirewae subsp. myrmecodiae Mann, 1921: 448 (w.) FIJI IS.

Formicinae Camponotini Polyrhachis rotumana TRUE TRUE FALSE

rotumana. Polyrhachis (Chariomyrma) rotumana Wilson & Taylor, 1967: 99, fig. 83 (w.) FIJI IS.

Ponerinae Ponerini Ponera colaensis TRUE TRUE FALSE

colaensis. Ponera colaensis Mann, 1921: 417 (w.) FIJI IS. Santschi, 1928c: 68 (q.). See also: Taylor, 1967a: 58.

Ponerinae Ponerini Ponera manni TRUE TRUE FALSE

manni. Ponera manni Taylor, 1967a: 86, figs. 76, 77 (w.) FIJI IS.

Myrmicinae Myrmecinini Pristomyrmex mandibularis TRUE TRUE FALSE

mandibularis. Pristomyrmex mandibularis Mann, 1921: 444 (w.) FIJI IS. See also: Wang, M. 2003: 505.

Proceratiinae Proceratiini Proceratium oceanicum TRUE TRUE FALSE

oceanicum. Proceratium oceanicum De Andrade, in Baroni Urbani & De Andrade, 2003b: 310, fig. 127 (w.) FIJI IS.

Proceratiinae Proceratiini Proceratium relictum TRUE TRUE FALSE

relictum. Proceratium relictum Mann, 1921: 413, fig. 3 (w.q.) FIJI IS. See also: Baroni Urbani & De Andrade, 2003b: 306.

Myrmicinae Dacetini Pyramica trauma TRUE TRUE FALSE

trauma. Pyramica trauma Bolton, 2000: 408 (w.) FIJI IS.

Myrmicinae Formicoxenini Romblonella vitiensis TRUE TRUE FALSE

vitiensis. Romblonella vitiensis Smith, M.R. 1953a: 79 (w.) FIJI IS.

Myrmicinae Formicoxenini Romblonella scrobifera liogaster TRUE TRUE FALSE

liogaster. Tetramorium scrobiferum var. liogaster Santschi, 1928c: 69 (w.) FIJI IS. Combination in Romblonella: Bolton, 1995b: 382.

Myrmicinae Dacetini Strumigenys basiliska TRUE TRUE FALSE

basiliska. Strumigenys basiliska Bolton, 2000: 750 (w.) FIJI IS.

Myrmicinae Dacetini Strumigenys chernovi TRUE TRUE FALSE

chernovi. Strumigenys chernovi Dlussky, 1993: 57, figs. 2, 3 (w.q.) FIJI IS. See also: Bolton, 2000: 805.

Myrmicinae Dacetini Strumigenys daithma TRUE TRUE FALSE

daithma. Strumigenys daithma Bolton, 2000: 756 (w.) FIJI IS.

Myrmicinae Dacetini Strumigenys ekasura TRUE TRUE FALSE

ekasura. Strumigenys ekasura Bolton, 2000: 807 (w.) FIJI IS.

Myrmicinae Dacetini Strumigenys frivola TRUE TRUE FALSE

frivola. Strumigenys frivola Bolton, 2000: 817 (w.) FIJI IS.

Myrmicinae Dacetini Strumigenys jepsoni TRUE TRUE FALSE

jepsoni. Strumigenys jepsoni Mann, 1921: 462, fig. 22 (w.) FIJI IS. See also: Bolton, 2000: 809.

Myrmicinae Dacetini Strumigenys nidifex TRUE TRUE FALSE

nidifex. Strumigenys nidifex Mann, 1921: 464, fig. 23 (w.q.m.) FIJI IS. Wheeler, G.C. & Wheeler, J. 1955a: 140 (l.). See also: Brown, 1971c: 81; Bolton, 2000: 905.

Myrmicinae Dacetini Strumigenys panaulax TRUE TRUE FALSE

panaulax. Strumigenys panaulax Bolton, 2000: 811 (w.) FIJI IS.

Myrmicinae Dacetini Strumigenys praefecta TRUE TRUE FALSE

praefecta. Strumigenys praefecta Bolton, 2000: 826 (w.q.) FIJI IS.

Myrmicinae Dacetini Strumigenys scelesta TRUE TRUE FALSE

scelesta. Strumigenys scelestus Mann, 1921: 463, fig. 22 (w.) FIJI IS. See also: Bolton, 2000: 812.

Myrmicinae Dacetini Strumigenys sulcata TRUE TRUE FALSE

sulcata. Strumigenys sulcata Bolton, 2000: 828 (w.q.) FIJI IS.

Dolichoderinae Tapinomini Technomyrmex vitiensis TRUE TRUE FALSE

vitiensis. Technomyrmex albipes var. vitiensis Mann, 1921: 473 (w.) FIJI IS. Santschi, 1928a: 52 (m.). Junior synonym of albipes: Wilson & Taylor, 1967: 82. Revived from synonymy, raised to species and senior synonym of rufescens: Bolton, 2007a: 104.

Formicinae Camponotini Camponotus reticulatus fullawayi TRUE TRUE FALSE

fullawayi. Camponotus reticulatus subsp. fullawayi Wheeler, W.M. 1912a: 47 (s.w.) GUAM.

Myrmicinae Pheidolini Pheidole recondita TRUE TRUE FALSE

recondita. Pheidole recondita Clouse, 2007a: 10, pl. 4 (s.w.) GUAM I. [Also described as new by Clouse, 2007b: 197, 242.]

Myrmicinae Pheidolini Pheidole recondita TRUE TRUE FALSE

recondita. Pheidole recondita Clouse, 2007a: 10, pl. 4 (s.w.) GUAM I. [Also described as new by Clouse, 2007b: 197, 242.]

Amblyoponinae Amblyoponini Amblyopone zwaluwenburgi TRUE TRUE FALSE

zwaluwenburgi. Stigmatomma (Fulakora) zwaluwenburgi Williams, 1946: 639 (w.) HAWAII. Combination in Amblyopone: Brown, 1960a: 169.

Myrmicinae Formicoxenini Cardiocondyla minutior TRUE TRUE FALSE

minutior. Cardiocondyla nuda var. minutior Forel, 1899a: 120 (w.) HAWAII. Wheeler, W.M. 1922f: 317 (q.); Heinze, 1999: 251 (polymorphic m.). Subspecies of nuda: Smith, M.R. 1944a: 38. Creighton, 1950a: 198. Junior synonym of nuda: Wilson & Taylor, 1967: 55. Revived from synonymy: Heinze, 1999: 251. Senior synonym of tsukuyomi: Seifert, 2003a: 283.

Ponerinae Ponerini Hypoponera zwaluwenburgi TRUE TRUE FALSE

zwaluwenburgi. Pseudocryptopone zwaluwenburgi Wheeler, W.M. 1933g: 14, fig. 5 (w.) HAWAII. Combination in Ponera: Wilson, 1957b: 375; in Hypoponera: Taylor, 1967a: 12.

Myrmicinae Solenopsidini Monomorium liliuokalanii TRUE TRUE FALSE

liliuokalanii. Monomorium minutum var. liliuokalanii Forel, 1899a: 119 (w.) HAWAII. Subspecies of minutum: Forel, 1909d: 225. Junior synonym of minutum: Wilson & Taylor, 1967: 67. Revived from synonymy, raised to species and senior synonym of samoanum: Bolton, 1987: 288.

Formicinae Plagiolepidini Nylanderia sharpii TRUE TRUE FALSE

sharpii. Prenolepis sharpii Forel, 1899a: 121, fig. 1 (w.q.m.) HAWAII. Combination in Paratrechina (Nylanderia): Emery, 1925b: 221; in Nylanderia: LaPolla, Brady & Shattuck, 2010a: 127.

Ponerinae Ponerini Ponera swezeyi TRUE TRUE FALSE

swezeyi. Pseudocryptopone swezeyi Wheeler, W.M. 1933g: 16, fig. 6 (w.q.) HAWAII. Combination in Ponera: Wilson, 1957b: 370. See also: Taylor, 1967a: 85.

Myrmicinae Formicoxenini Romblonella townesi TRUE TRUE FALSE

townesi. Romblonella townesi Smith, M.R. 1953a: 80 (w.) MARIANAS IS.

Myrmicinae Dacetini Pyramica mumfordi TRUE TRUE FALSE

mumfordi. Strumigenys (Cephaloxys) inezi subsp. mumfordi Wheeler, W.M. 1932c: 160 (w.) FRENCH POLYNESIA (Marquesas Is). Combination in Smithistruma: Brown, 1948e: 105; in Pyramica: Bolton, 1999: 1673. Raised to species: Brown, 1953g: 124. See also: Brown, 1964a: 195; Bolton, 2000: 401.

Dolichoderinae TRUE TRUE FALSE

Extant: 28 genera, 692 species, 130 subspecies

Fossil: 16 genera, 103 species

Dolichoderidae Forel, 1878: 364. Type-genus: Dolichoderus.

Taxonomic history

Dolichoderinae as family: Emery, 1894g: 378 [Dolichoderidae]; Ashmead, 1905b: 384 [Dolichoderidae]; Novák & Sadil, 1941: 94 [Dolichoderidae]; Bernard, 1951: 1071 [Dolichoderidae]; Bernard, 1953b: 255 [Dolichoderidae].

Dolichoderinae as tribe of Formicidae: André, 1882a: 127 [Dolichoderidae].

Dolichoderinae as subfamily of Formicidae: Forel, 1878: 364 [Dolichoderidae]; Emery & Forel, 1879a: 454 [Dolichoderidae]; Nasonov, 1889: 26 [Dolichoderidae]; Forel, 1892j: 220 [Dolichoderidae]; Forel, 1893a: 165; Dalla Torre, 1893: 156; Forel, 1895b: 107 [Dolichoderidae]; Emery, 1895j: 771 [subfamily spelled Dolichoderini]; Emery, 1896e: 186; Forel, 1899c: 98; Bingham, 1903: 288; Wheeler, W.M. 1910g: 142; Emery, 1913a: 2; Arnold, 1915: 143; Wheeler, W.M. 1915h: 71; Wheeler, 1915g: 811 [Dolichoderides]; Arnold, 1915: 143; Donisthorpe, 1915d: 178; Forel, 1917: 247; Escherich, 1917: 2 [Dolichoderini]; Bondroit, 1918: 86 [Dolichoderitae]; Wheeler, W.M. 1920: 53; Wheeler, W.M. 1922a: 199; Borgmeier, 1923: 80; Karavaiev, 1936: 163; Smith, M.R. 1951a: 833; Clark, 1951: 16; Brown, 1954e: 29; Kempf, 1972a: 266; Wheeler, G.C. & Wheeler, J. 1972a: 41; Brown, 1973b: 169; all subsequent authors.

Dolichoderinae as formicomorph subfamily of Formicidae: Bolton, 2003: 18, 80.

Dolichoderinae as formicoid subfamily of Formicidae: Brady, Schultz, et al. 2006: 18173; Moreau, Bell, et al. 2006: 102.

Dolichoderinae as formicoid dolichoderomorph subfamily of Formicidae: Ward, 2007a: 556.

Tribes (extant) of Dolichoderinae: Bothriomyrmecini, Dolichoderini, Leptomyrmecini, Tapinomini.

Tribes (extinct) incertae sedis in Dolichoderinae: †Miomyrmecini, †Zherichiniini.

Genera (extinct) incertae sedis in Dolichoderinae: †Alloiomma, †Asymphylomyrmex, †Elaeomyrmex, †Elaphrodites, †Eldermyrmex, †Emplastus, †Eotapinoma, †Eurymyrmex, †Kotshkorkia, †Leptomyrmula, †Miomyrmex, †Petraeomyrmex, †Proiridomyrmex, †Protazteca, †Zherichinius.

Subfamily Dolichoderinae and tribes references, world

Forel, 1878: 364, 380 (diagnosis, genera); Dalla Torre, 1893: 156 (catalogue); Emery, 1895j: 771 (synoptic classification); Emery, 1896e: 186 (genera key); Handlirsch, 1907: 869 (†fossil taxa catalogue); Wheeler, W.M. 1910g: 142 (diagnosis); Emery, 1913a: 2, 6 (diagnosis, tribe key, catalogue); Emery, 1913a: 17 (Tapinomini diagnosis, genera key, catalogue); Arnold, 1915: 144, (diagnosis); Gallardo, 1916a: 3 (diagnosis); Forel, 1917: 247 (synoptic classification); Forel, 1921c: 136 (diagnosis); Wheeler, W.M. 1922a: 199, 687, 688 (diagnosis, tribe key, Tapinomini genera key); Brown & Nutting, 1950: 127 (venation, phylogeny); Brown, 1954e: 29 (phylogeny); Pavan, 1955: 135 (gastral organs); Eisner, 1957: 453 (proventriculus morphology); Bernard, 1967: 246 (diagnosis); Gotwald, 1969: 118 (mouthparts morphology); Wheeler, G.C. & Wheeler, J. 1972a: 41 (diagnosis); Brown, 1973b: 169 (genera & distribution); Wheeler, G.C. & Wheeler, J. 1976b: 61 (larvae, review & synthesis); Snelling, R.R. 1981: 401 (synoptic classification); Dazzini Valcurone & Fanfani, 1985: 1 (gastral glands); Wheeler, G.C. & Wheeler, J. 1985: 258 (synoptic classification); Billen, 1986: 173 (Dufour's gland); Billen, 1987: 278 (abdominal glands); Baroni Urbani & Wilson, 1987: 1 (†fossil Leptomyrmecini); Dlussky & Fedoseeva, 1988: 77 (synoptic classification); Hölldobler & Wilson, 1990: 9 (synoptic classification, genera keys); Baroni Urbani, Bolton & Ward, 1992: 316 (phylogeny); Shattuck, 1992b: 199 (higher classification, phylogeny); Shattuck, 1992c: 20 (revision of subfamily, genera key); Shattuck, 1994: 3 (catalogue); Bolton, 1994: 22 (diagnosis, synoptic classification, genera keys); Shattuck, 1995: 217 (phylogeny, genera); Bolton, 1995a: 1038 (census); Bolton, 1995b: 10 (catalogue); Wenseleers, Schoeters, et al., 1998: 121 (cloacal gland); Brandão, Baroni Urbani, et al. 1999: 411 (phylogeny, genera); Chiotis, Jermiin & Crozier, 2000: 108 (phylogeny, genera); Dlussky & Rasnitsyn, 2002: 416 (diagnosis for impression fossils); Bolton, 2003: 18, 80 (diagnosis, synopsis); Dubovikov, 2005a: 91 (tribes of Dolichoderinae); Brady, Schultz, et al. 2006: 18173 (phylogeny); Moreau, Bell, et al. 2006: 102 (phylogeny); Ward, Brady, et al. 2010: 342 (phylogeny, tribe-rank classification).

Regional and national faunas with keys

André, 1882a: 127 (Europe & Algeria); Nasonov, 1889: 50 (Russia); Forel, 1891b: 9 (Madagascar genera); Forel, 1895e: 460 (India & Sri Lanka); Bingham, 1903: 288 (India, Sri Lanka & Burma); Ruzsky, 1905b: 102 (Russian Empire); Bondroit, 1910: 488 (Belgium); Wheeler, W.M. 1910g: 560 (North America genera); Stitz, 1914: 77 (Central Europe); Gallardo, 1915: 34 (Argentina genera); Forel, 1915d: 40 (Switzerland); Arnold, 1915: 145 (South Africa); Donisthorpe, 1915d: 178 (Britain); Gallardo, 1916a: 12 (Argentina); Emery, 1916b: 208 (Italy); Wheeler, W.M. 1916m: 589 (U.S.A., Connecticut); Bondroit, 1918: 86 (France & Belgium); Gallardo, 1919b: 253 (Argentina genera); Soudek, 1922: 58 (Czechoslovakia); Stärcke, 1926: 117 (Netherlands); Karavaiev, 1927c: 271 (Ukraine); Donisthorpe, 1927b: 199 (Britain); Menozzi & Russo, 1930: 172 (Dominican Republic); Arnol'di, 1933b: 600 (Russia); Menozzi, 1933b: 90 (Israel genera); Karavaiev, 1936: 164 (Ukraine); Smith, M.R. 1937: 861 (Puerto Rico); Stitz, 1939: 209 (Germany); Kratochvíl, 1941: 94 (Central Europe); Novák & Sadil, 1941: 94 (Central Europe); Cole, 1942: 370 (U.S.A., Utah); Smith, M.R. 1943f: 309 (U.S.A. males); Buren, 1944a: 290 (U.S.A., Iowa); Smith, M.R. 1947f: 592 (U.S.A. genera); Creighton, 1950a: 330 (North America); Kusnezov, 1956: 27 (Argentina); Brown, 1958h: 29 (New Zealand); Kusnezov, 1959: 40 (Neotropical genera); Gregg, 1963: 341 (U.S.A., Colorado); Wheeler, G.C. & Wheeler, J. 1963: 149 (U.S.A., North Dakota); Bernard, 1967: 246 (Western Europe); Wilson & Taylor, 1967: 17 (Polynesia); Bolton, 1973a: 329 (West Africa genera); Bolton & Collingwood, 1975: 3 (Britain); Boven, 1970b: 24 (Netherlands); Snelling, R.R. & Hunt, 1976: 90 (Chile); Tarbinsky, 1976: 118 (Kyrghyzstan); Boven, 1977: 123 (Belgium); Kutter, 1977c: 170 (Switzerland); Arnol'di & Dlussky, 1978: 546 (former European U.S.S.R.); Collingwood, 1978: 80 (Iberian Peninsula); Collingwood, 1979: 32 (Fennoscandia & Denmark); Greenslade, 1979: 28 (South Australia genera); Schembri & Collingwood, 1981: 420 (Malta); Allred, 1982: 443 (U.S.A., Utah); Baroni Urbani, 1984: 80 (Neotropical genera); Gösswald, 1985: 314 (Germany); Collingwood, 1985: 242 (Saudi Arabia); Wheeler, G.C. & Wheeler, J. 1986g: 55 (U.S.A., Nevada); Agosti & Collingwood, 1987b: 278 (Balkans); Dlussky, Soyunov & Zabelin, 1990: 168 (Turkmenistan); Kupyanskaya, 1990: 152 (Far Eastern Russia); Morisita, Kubota, Onoyama, et al., 1991: 2 (Japan); Atanasov & Dlussky, 1992: 49 (Bulgaria); Lattke, in Jaffe, 1993: 147 (Neotropical genera); Arakelian, 1994: 70 (Armenia); Wu, J. & Wang, 1995: 115 (China genera); Kupyanskaya, 1995: 331 (Far Eastern Russia); Collingwood & Agosti, 1996: 359 (Saudi Arabia); Seifert, 1996b: 161 (Central Europe); Skinner & Allen, 1996: 40 (Britain); Collingwood & Prince, 1998: 20 (Portugal); Kim & Kim, 1999: 18 (Korea); Shattuck, 1999: 26, 64 (Australia genera, synopsis); Andersen, 2000: 59 (northern Australia genera); Zhou, 2001b: 149 (China, Guangxi); Czechowski, Radchenko & Czechowska, 2002: 134 (Poland); Aktaç & Radchenko, 2002: 54 (Turkey genera); Yoshimura & Onoyama, 2002: 425 (Japan genera, males); Mackay & Mackay, 2002: 221 (U.S.A., New Mexico); Palacio & Fernández, in Fernández, 2003d: 239 (Neotropical genera); Radchenko, 2005b: 186 (North Korea); Coovert, 2005: 102 (U.S.A., Ohio); Clouse, 2007b: 190 (Micronesia); Seifert, 2007: 147 (North and Central Europe); Fisher, 2009: 51 (Malagasy genera); Terayama, 2009: 196 (Taiwan); Heterick, 2009: 28 (south-western Australia genera); Boer, 2010: 14 (Benelux); Yoshimura & Fisher, 2011: 7 (Malagasy males key).

Formicinae TRUE TRUE FALSE

Extant: 49 genera, 2,831 species, 839 subspecies

Fossil: 29 genera, 193 species, 1 subspecies

Formicariae Latreille, 1809: 124. Type-genus: Formica.

Taxonomic history

Formicinae as group name: Latreille, 1809: 124 [Formicariae]; Lepeletier de Saint-Fargeau, 1835: 197 [Formicites]; Nylander, 1846a: 877 [Formicae].

Formicinae as family: Smith, F. 1858b: 1 [Formicidae]; Smith, F. 1861b: 36 [Formicidae]; Smith, 1871a: 302 [Formicidae]; André, 1882a: 125 [Formicidae]; Saunders, 1896: 18 [Formicidae]; Novák & Sadil, 1941: 97 [Formicidae]; Bernard, 1951: 1073 [Formicidae]; Bernard, 1953b: 256 [Formicidae].

Formicinae as tribe of Formicidae: André, 1874: 167 [Formicidae].

Formicinae as subfamily of Formicidae: Mayr, 1855: 286, 299 [Formicidae]; Mayr, 1861: 21 [Formicidae]; Mayr, 1862: 651 [Formicidae]; Mayr, 1865: 6 [Formicidae]; Mayr, 1868c: 24 [Formicidae]; Forel, 1870: 307 [Formicidae]; Forel, 1874: 21 [Formicidae]; Emery, 1877a: 70 [Formicidae]; André, 1881b: 54 [Formicidae]; Ashmead, 1905b: 384; Bondroit, 1918: 17 [Formicitae]; Wheeler, W.M. 1920: 53; Wheeler, W.M. 1922a: 210; Emery, 1925b: 2; Karavaiev, 1936: 172; Clark, 1951: 16; Brown, 1954e: 29; Kempf, 1972a: 266; Brown, 1973b: 169; all subsequent authors.

Formicinae as formicomorph subfamily of Formicidae: Bolton, 2003: 20, 93.

Formicinae as formicoid subfamily of Formicidae: Brady, Schultz, et al. 2006: 18173; Moreau, Bell, et al. 2006: 102.

Formicinae as formicoid formicomorph subfamily of Formicidae: Ward, 2007a: 556.

Tribes of Formicinae: Camponotini, Formicini, Gesomyrmecini, Gigantiopini, Lasiini, Melophorini, Myrmecorhynchini, Myrmoteratini, Notostigmatini, Oecophyllini, Plagiolepidini.

Genera (extinct) incertae sedis in Formicinae: †Camponotites, †Drymomyrmex, †Imhoffia, †Kyromyrma, †Leucotaphus, †Protrechina, †Zhangidris.

Hong (2002) genera (extinct) incertae sedis in Formicinae: †Curtipalpulus, †Eoleptocerites, †Eurytarsites, †Fushuniformica, †Huaxiaformica, †Leptogasteritus, †Liaoformica, †Longiformica, †Magnogasterites, †Orbicapitia, †Ovalicapito, †Ovaligastrula, †Sinoformica, †Sinotenuicapito, †Wilsonia (unresolved junior homonym).

Subfamily Formicinae references, world

Mayr, 1862: 651 (genera key); Mayr, 1865: 6 (diagnosis); Handlirsch, 1907: 859 (†fossil taxa catalogue); Dalla Torre, 1893: 171 (catalogue); Emery, 1895j: 772 (synoptic classification); Emery, 1896e: 187 (genera key); Wheeler, W.M. 1910g: 143 (diagnosis); Forel, 1912i: 88 (tribes key); Forel, 1917: 248 (synoptic classification); Arnold, 1920a: 551 (diagnosis); Forel, 1921c: 139 (diagnosis); Wheeler, W.M. 1922a: 210, 691 (diagnosis, tribes key); Emery, 1925b: 2 (diagnosis, tribe key, catalogue); Brown & Nutting, 1950: 127 (venation, phylogeny); Eisner, 1957: 465 (proventriculus morphology); Hung & Brown, 1966: 198 (gastric apex, structure); Bernard, 1967: 267 (diagnosis); Gotwald, 1969: 120 (mouthparts morphology); Wheeler, G.C. & Wheeler, J. 1972a: 41 (diagnosis); Brown, 1973b: 169 (genera, distribution); Wheeler, G.C. & Wheeler, J. 1976b: 62 (larvae, review & synthesis); Snelling, R.R. 1981: 402 (synoptic classification); Wheeler, G.C. & Wheeler, J. 1985: 258 (synoptic classification); Billen, 1986: 173 (Dufour's gland); Dlussky & Fedoseeva, 1988: 77 (synoptic classification); Hölldobler & Wilson, 1990: 9 (synoptic classification, genera keys); Agosti, 1991: 295 (genus group diagnoses); Shattuck, 1992b: 201 (phylogeny); Baroni Urbani, Bolton & Ward, 1992: 317 (phylogeny); Bolton, 1994: 42 (diagnosis, synoptic classification, genera keys); Bolton, 1995a: 1039 (census); Bolton, 1995b: 11 (catalogue); Wenseleers, Schoeters, et al. 1998: 121 (cloacal gland); Dlussky & Rasnitsyn, 2002: 417 (diagnosis for impression fossils); Bolton, 2003: 20, 93 (diagnosis, synopsis); Brady, Schultz, et al. 2006: 18173 (phylogeny); Moreau, Bell, et al. 2006: 102 (phylogeny).

Regional and national faunas with keys

Mayr, 1855: 299 (Austria); Mayr, 1861: 25 (Europe); Mayr, 1868c: 25 (†Baltic Amber); André, 1874: 167 (Europe); Forel, 1874: 22 (Switzerland); Saunders, E. 1880: 203 (Britain); André, 1882a: 126 (Europe & Algeria); Provancher, 1887: 225 (Canada); Cresson, 1887: 94 (U.S.A. genera); Nasonov, 1889: 50 (Russia); Forel, 1891b: 8 (Madagascar genera); Lameere, 1892: 62 (Belgium); Forel, 1892j: 220 (India & Sri Lanka); Bingham, 1903: 308 (India, Sri Lanka & Burma); Ruzsky, 1905b: 100 (Russian Empire); Wasmann, 1906: 7 (Luxemburg); Bondroit, 1910: 481 (Belgium); Wheeler, W.M. 1910g: 560 (North America genera); Stitz, 1914: 80 (Central Europe); Gallardo, 1915: 35 (Argentina genera); Forel, 1915d: 45 (Switzerland); Donisthorpe, 1915d: 184 (Britain); Emery, 1916b: 216 (Italy); Wheeler, W.M. 1916m: 590 (U.S.A., Connecticut); Bondroit, 1918: 17 (France & Belgium); Arnold, 1920a: 552 (South Africa); Kutter, 1920b: 134 (Switzerland); Soudek, 1922: 61 (Czechoslovakia); Lomnicki, 1925a: 160 (Poland); Stärcke, 1926: 118, 146 (Netherlands); Karavaiev, 1927c: 273 (Ukraine); Donisthorpe, 1927b: 205 (Britain); Menozzi & Russo, 1930: 172 (Dominican Republic); Arnol'di, 1933b: 601 (Russia); Menozzi, 1933b: 90 (Israel genera); Karavaiev, 1936: 173 (Ukraine); Smith, M.R. 1937: 865 (Puerto Rico); Stitz, 1939: 230 (Germany); Kratochvíl, 1941: 97 (Central Europe); Novák & Sadil, 1941: 97 (Central Europe); Cole, 1942: 373 (U.S.A., Utah); Smith, M.R. 1943f: 309 (U.S.A., males); Buren, 1944a: 292 (U.S.A., Iowa); Holgersen, 1943b: 173 (Norway); Holgersen, 1944: 199 (Norway); Smith, M.R. 1947f: 599 (U.S.A. genera); Boven, 1947: 181 (Belgium); Creighton, 1950a: 355 (North America); Kusnezov, 1956: 31 (Argentina); Brown, 1958h: 42 (New Zealand); Boven, 1959: 11 (Netherlands); Gregg, 1963: 447 (U.S.A., Colorado); Wheeler, G.C. & Wheeler, J. 1963: 160 (U.S.A., North Dakota); Collingwood, 1964: 104 (Britain); Bernard, 1967: 268 (Western Europe); Wilson & Taylor, 1967: 17 (Polynesia); Boven, 1970b: 26 (Netherlands); Kempf, 1972a: 266 (Neotropical, synoptic classification); Bolton, 1973a: 329 (West Africa genera); Bolton & Collingwood, 1975: 3 (Britain); Snelling & Hunt, 1976: 104 (Chile); Tarbinsky, 1976: 126 (Kyrghyzstan); Boven, 1977: 126 (Belgium); Kutter, 1977c: 183 (Switzerland); Arnol'di & Dlussky, 1978: 548 (former European U.S.S.R.); Collingwood, 1978: 88 (Iberian Peninsula); Collingwood, 1979: 85 (Fennoscandia & Denmark); Greenslade, 1979: 32 (South Australia genera); Schembri & Collingwood, 1981: 436 (Malta); Prins, 1983: 8 (Southern Africa genera); Allred, 1982: 444 (U.S.A., Utah); Verhaeghe, Deligne, et al., 1984: 106 (Belgium genera); Baroni Urbani, 1984: 81 (Neotropical genera); Gösswald, 1985: 263 (Germany); Collingwood, 1985: 273 (Saudi Arabia); Wheeler, G.C. & Wheeler, J. 1986g: 58 (U.S.A., Nevada); Nilsson & Douwes, 1987: 68 (Norway); Agosti & Collingwood, 1987b: 279 (Balkans); Dlussky, Soyunov & Zabelin, 1990: 124 (Turkmenistan); Kupyanskaya, 1990: 162 (Far Eastern Russia); Morisita, Kubota, Onoyama, et al., 1991: 10 (Japan); Atanasov & Dlussky, 1992: 49 (Bulgaria); Shattuck, 1992b: 199 (higher classification, phylogeny); Lattke, in Jaffe, 1993: 150 (Neotropical genera); Arakelian, 1994: 76 (Armenia); Wu, J. & Wang, 1995: 125 (China genera); Kupyanskaya, 1995: 332 (Far Eastern Russia); Collingwood & Agosti, 1996: 361 (Saudi Arabia); Seifert, 1996b: 166 (Central Europe); Skinner & Allen, 1996: 41 (Britain); Collingwood & Prince, 1998: 21 (Portugal); Shattuck, 1999: 25, 86 (Australia genera, synopsis); Andersen, 2000: 68 (northern Australia genera); Zhou, 2001b: 165 (China, Guangxi); Czechowski, Radchenko & Czechowska, 2002: 147 (Poland); Aktaç & Radchenko, 2002: 54 (Turkey genera); Yoshimura & Onoyama, 2002: 425 (Japan genera, males); Mackay & Mackay, 2002: 236 (U.S.A., New Mexico); Palacio & Fernández, in Fernández, 2003d: 242 (Neotropical genera); Radchenko, 2005b: 187 (North Korea); Coovert, 2005: 113 (U.S.A., Ohio); Clouse, 2007b: 190 (Micronesia); Seifert, 2007: 150 (North and Central Europe); Terayama, 2009: 202 (Taiwan); Heterick, 2009: 30 (south-western Australia genera); Boer, 2010: 17 (Benelux).

Myrmicinae TRUE TRUE FALSE

Extant: 141 genera, 6,087 species, 864 subspecies

Fossil: 31 genera, 130 species

Myrmicites Lepeletier de Saint-Fargeau, 1835: 169. Type-genus: Myrmica.

Taxonomic history

Myrmicinae as group name: Lepeletier de Saint-Fargeau, 1835: 169 [Myrmicites]; Nylander, 1846a: 877 [Myrmicae].

Myrmicinae as family: Smith, F. 1851: 4 [Myrmicidae]; Smith, F. 1861b: 45 [Myrmicidae]; Smith, F. 1871a: 324 [Myrmicidae]; André, 1882a: 125 [Myrmicidae]; Cresson, 1887: 93 [Myrmicidae]; Emery, 1894g: 383 [Myrmicidae]; Saunders, 1896: 18 [Myrmicidae]; Ashmead, 1905b: 383 [Myrmicidae]; Novák & Sadil, 1941: 71 [Myrmicidae]; Bernard, 1951: 1058 [Myrmicidae]; Bernard, 1953b: 222 [Myrmicidae].

Myrmicinae as subfamily of Poneridae: Smith, F. 1858b: 114 [Myrmicidae].

Myrmicinae as tribe of Formicidae: André, 1874: 167 [Myrmicidae].

Myrmicinae as subfamily of Myrmicidae: Ashmead, 1905b: 383.

Myrmicinae as subfamily of Formicidae: Mayr, 1855: 290, 299 [Myrmicidae]; Smith, F. 1857a: 70 [Myrmicidae]; Mayr, 1861: 21 [Myrmicidae]; Smith, F. 1862b: 33 [Myrmicidae]; Mayr, 1862: 738 [Myrmicidae]; Mayr, 1865: 17 [Myrmicidae]; Mayr, 1868c: 24 [Myrmicidae]; Forel, 1870: 307 [Myrmicidae]; Forel, 1874: 22 [Myrmicidae]; Emery, 1877a: 70 [Myrmicidae]; Forel, 1878: 367 [Myrmicidae]; Emery & Forel, 1879a: 456 [Myrmicidae]; André, 1881b: 64 [Myrmicidae]; Nasonov, 1889: 28 [Myrmicidae]; Forel, 1891b: 11 [Myrmicidae]; Forel, 1892j: 220 [Myrmicidae]; Forel, 1893a: 163; Dalla Torre, 1893: 53; Emery, 1895j: 768 [subfamily spelled Myrmicini]; Emery, 1896e: 179; Forel, 1899c: 30; Forel, 1902f: 520; Bingham, 1903: 105; Wheeler, W.M. 1910g: 138; Emery, 1914a: 29; Wheeler, 1915g: 806 [Myrmicides]; Wheeler, W.M. 1915h: 40; Donisthorpe, 1915d: 74; Arnold, 1916: 166; Escherich, 1917: 2 [Myrmicini]; Forel, 1917: 240 [subfamily spelled Myrmicini]; Bondroit, 1918: 14 [Myrmicitae]; Wheeler, W.M. 1920: 53; Wheeler, W.M. 1922a: 124; Emery, 1921f: 3; Karavaiev, 1934: 59; Clark, 1951: 16; Brown, 1954e: 28; Wheeler, G.C. & Wheeler, J. 1972a: 40; Brown, 1973b: 166; all subsequent authors.

Myrmicinae as myrmicomorph subfamily of Formicidae: Bolton, 2003: 52, 182.

Myrmicinae as formicoid subfamily of Formicidae: Moreau, Bell, et al. 2006: 102; Brady, Schultz, et al. 2006: 18173.

Myrmicinae as formicoid myrmicomorph subfamily of Formicidae: Ward, 2007a: 556.

Tribes of Myrmicinae: Adelomyrmecini, Ankylomyrmini, Attini, Basicerotini, Blepharidattini, Cataulacini, Cephalotini, Crematogastrini, Dacetini, Formicoxenini, Lenomyrmecini, Liomyrmecini, Melissotarsini, Meranoplini, Metaponini, Myrmecinini, Myrmicariini, Myrmicini, Paratopulini, Phalacromyrmecini, Pheidolini, Solenopsidini, Stegomyrmecini, Stenammini, Tetramoriini.

Genera (extant) incertae sedis in Myrmicinae: Diaphoromyrma, Tyrannomyrmex.

Genera (extinct) incertae sedis in Myrmicinae: †Afromyrma, †Attopsis, †Cephalomyrmex, †Electromyrmex, †Eocenidris, †Eomyrmex, †Fallomyrma, †Miosolenopsis, †Myrmecites, †Solenopsites.

Hong (2002) genera (extinct) incertae sedis in Myrmicinae: †Brachytarsites, †Clavipetiola, †Fushunomyrmex, †Orbigastrula, †Quadrulicapito, †Quineangulicapito, †Sinomyrmex, †Sphaerogasterites, †Wumyrmex.

Collective group name in Myrmicinae: †Myrmicites.

Subfamily Myrmicinae references, world

Mayr, 1865: 17 (diagnosis); Mayr, 1867a: 91 (diagnosis); Forel, 1878: 367 (diagnosis); Handlirsch, 1907: 872 (†fossil taxa catalogue); Dalla Torre, 1893: 53 (catalogue); Emery, 1895j: 768 (diagnosis); Emery, 1896e: 179 (genera key); Wheeler, W.M. 1910g: 138 (diagnosis); Emery, 1912b: 101 (phylogeny); Emery, 1914a: 34 (phylogeny, tribe key); Arnold, 1916: 164 (diagnosis); Forel, 1917: 240 (synoptic classification); Forel, 1921c: 139 (diagnosis); Emery, 1921f: 3 (diagnosis, tribes & genera key, catalogue); Wheeler, W.M. 1922a: 124, 655 (diagnosis, tribes key); Brown & Nutting, 1950: 126 (venation, phylogeny); Brown, 1954e: 28 (phylogeny); Eisner, 1957: 477 (proventriculus morphology); Bernard, 1967: 93 (diagnosis); Gotwald, 1969: 99 (mouthparts morphology); Wheeler, G.C. & Wheeler, J. 1972a: 40 (diagnosis); Brown, 1973b: 166 (genera, distribution); Wheeler, G.C. & Wheeler, J. 1976b: 52 (larvae, review & synthesis); Kugler, C. 1978a: 413 (sting structure); Kugler, C. 1978b: 267 (pygidial glands); Kugler, C. 1979c: 117 (sting, evolution); Snelling, R.R. 1981: 393 (synoptic classification); Caetano, F.H. 1984: 257 (digestive tract, morphology); Wheeler, G.C. & Wheeler, J. 1985: 257 (synoptic classification); Billen, 1986: 167 (Dufour's gland); Dlussky & Fedoseeva, 1988: 79 (synoptic classification); Hölldobler & Wilson, 1990: 9 onward (synoptic classification, genera keys); Baroni Urbani, Bolton & Ward, 1992: 317 (phylogeny); Bolton, 1994: 75 (diagnosis, synoptic classification, genera keys); Bolton, 1995a: 1040 (census); Bolton, 1995b: 13 (catalogue); Hashimoto, 1996: 354 (phylogenetic position); Baroni Urbani, 2000: 480 (phylogeny); Dlussky & Rasnitsyn, 2002: 422 (diagnosis for wingless fossils); Bolton, 2003: 52, 182 (diagnosis, synopsis); Moreau, Bell, et al. 2006: 102 (phylogeny); Brady, Schultz, et al. 2006: 18173 (phylogeny); Ward, 2007a: 556 (classification).

Regional and national faunas with keys

Mayr, 1855: 391 (Austria); Mayr, 1861: 29 (Europe); Mayr, 1868c: 79 (†Baltic Amber); André, 1874: 171 (Europe); Forel, 1874: 29 (Switzerland); Saunders, E. 1880: 213 (Britain); André, 1882c: 256 (Europe & Algeria); Cresson, 1887: 98 (U.S.A. genera); Provancher, 1887: 243 (Canada); Nasonov, 1889: 54 (Russia); Forel, 1891b: 11 (Madagascar genera); Lameere, 1892: 66 (Belgium); Forel, 1902f: 520 (India & Sri Lanka genera); Bingham, 1903: 105 (India, Sri Lanka & Burma); Ruzsky, 1905b: 103 (Russian Empire); Wasmann, 1906: 13 (Luxemburg); Bondroit, 1910: 490 (Belgium); Wheeler, W.M. 1910g: 558 (North America genera); Stitz, 1914: 55 (Central Europe); Gallardo, 1915: 32 (Argentina genera); Forel, 1915d: 8 (Switzerland); Donisthorpe, 1915d: 74 (Britain); Arnold, 1916: 166, 170 (South Africa tribes, genera); Emery, 1916b: 112 (Italy); Wheeler, W.M. 1916m: 581 (U.S.A., Connecticut); Bondroit, 1918: 90 (France & Belgium); Kutter, 1920b: 144 (Switzerland); Soudek, 1922: 20 (Czechoslovakia); Stärcke, 1926: 84 (Netherlands); Karavaiev, 1927c: 256 (Ukraine); Donisthorpe, 1927b: 77 (Britain); Menozzi & Russo, 1930: 170 (Dominican Republic); Gallardo, 1932b: 91 (Argentina, tribes); Arnol'di, 1933b: 596 (Russia); Menozzi, 1933b: 88 (Israel genera); Karavaiev, 1934: 60 (Ukraine); Smith, M.R. 1937: 829 (Puerto Rico); Stitz, 1939: 63 (Germany); Kratochvíl, 1941: 71 (Central Europe); Novák & Sadil, 1941: 71 (Central Europe); Cole, 1942: 360 (U.S.A., Utah); Smith, M.R. 1943f: 291 (U.S.A., males); Holgersen, 1943b: 166 (Norway); Holgersen, 1944: 198 (Norway); Buren, 1944a: 281 (U.S.A., Iowa); Smith, M.R. 1947f: 543 (U.S.A. genera); Boven, 1947: 170 (Belgium); Creighton, 1950a: 83 (North America); Kusnezov, 1956: 15 (Argentina); Brown, 1958h: 25 (New Zealand); Boven, 1959: 7 (Netherlands); Gregg, 1963: 288 (U.S.A., Colorado); Wheeler, G.C. & Wheeler, J. 1963: 92 (U.S.A., North Dakota); Collingwood, 1964: 94 (Britain); Bernard, 1967: 95 (Western Europe); Wilson & Taylor, 1967: 13 (Polynesia); Boven, 1970b: 9 (Netherlands); Kempf, 1972a: 263 (Neotropical, synoptic classification); Bolton, 1973a: 325 (West Africa genera); Bolton & Collingwood, 1975: 3 (Britain); Snelling, R.R. & Hunt, 1976: 70 (Chile); Tarbinsky, 1976: 19 (Kyrghyzstan); Boven, 1977: 69 (Belgium); Kutter, 1977c: 31 (Switzerland); Arnol'di & Dlussky, 1978: 524 (former European U.S.S.R.); Collingwood, 1978: 75 (Iberian Peninsula); Collingwood, 1979: 36 (Fennoscandia & Denmark); Greenslade, 1979: 20 (South Australia genera); Schembri & Collingwood, 1981: 423 (Malta); Allred, 1982: 438 (U.S.A., Utah); Baroni Urbani, 1984: 76 (Neotropical genera); Verhaeghe, Deligne, et al. 1984: 112 (Belgium genera); Gösswald, 1985: 289 (Germany); Collingwood, 1985: 245 (Saudi Arabia); Wheeler, G.C. & Wheeler, J. 1986g: 20 (U.S.A., Nevada); Nilsson & Douwes, 1987: 57 (Norway); Agosti & Collingwood, 1987b: 265 (Balkans); Dlussky, Soyunov & Zabelin, 1990: 181 (Turkmenistan); Kupyanskaya, 1990: 89 (Far Eastern Russia); Ogata, 1991b: 61 (Japan genera); Morisita, Kubota, et al. 1992: 1 (Japan); Atanasov & Dlussky, 1992: 51 (Bulgaria); Lattke, in Jaffe, 1993: 153 (Neotropical genera); Arakelian, 1994: 15 (Armenia); Wu, J. & Wang, 1995: 57 (China genera); Kupyanskaya, 1995: 327 (Far Eastern Russia); Collingwood & Agosti, 1996: 308 (Saudi Arabia); Seifert, 1996b: 108 (Central Europe); Skinner & Allen, 1996: 43 (Britain); Collingwood & Prince, 1998: 10 (Portugal); Shattuck, 1999: 39, 122 (Australia genera, synopsis); Andersen, 2000: 37 (northern Australia genera); Zhou, 2001b: 69 (China, Guangxi); Czechowski, Radchenko & Czechowska, 2002: 135 (Poland); Aktaç & Radchenko, 2002: 55 (Turkey genera); Yoshimura & Onoyama, 2002: 424 (Japan genera, males); Mackay & Mackay, 2002: 58 (U.S.A., New Mexico); Palacio & Fernández, in Fernández, 2003d: 244 (Neotropical genera and synopsis); Coovert, 2005: 32 (U.S.A., Ohio); Radchenko, 2005b: 184 (North Korea); Clouse, 2007b: 190 (Micronesia); Seifert, 2007: 110 (North and Central Europe); Terayama, 2009: 131 (Taiwan); Heterick, 2009: 36 (south-western Australia genera); Boer, 2010: 46 (Benelux); Eguchi, Bui & Yamane, 2011: 8 (Vietnam genera).

Ponerinae TRUE TRUE FALSE

Extant: 27 genera, 1,033 species, 155 subspecies

Fossil: 9 genera, 68 species, 2 subspecies

Ponerites Lepeletier de Saint-Fargeau, 1835: 185. Type-genus: Ponera.

Taxonomic history

Ponerinae as group name: Lepeletier de Saint-Fargeau, 1835: 185 [Ponérites].

Ponerinae as family: Smith, F. 1851: 6 [Poneridae]; Smith, F. 1858b: 76 [Poneridae]; Smith, F. 1861b: 44 [Poneridae]; Smith, F. 1871a: 320 [Poneridae]; André, 1882a: 125 [Poneridae]; Cresson, 1887: 93 [Poneridae]; Emery, 1894g: 379 [Poneridae]; Saunders, 1896: 18 [Poneridae]; Ashmead, 1905b: 382 [Poneridae]; Novák & Sadil, 1941: 70 [Poneridae]; Bernard, 1951: 1042 [Poneridae]; Bernard, 1953b: 185 [Poneridae].

Ponerinae as tribe of Formicidae: André, 1874: 167 [Poneridae].

Ponerinae as subfamily of Poneridae: Ashmead, 1905b: 382.

Ponerinae as subfamily of Formicidae: Mayr, 1855: 289, 299 [Poneridae]; Mayr, 1861: 21 [Poneridae]; Mayr, 1862: 712 [Poneridae]; Smith, F. 1857a: 64 [Poneridae]; Smith, F. 1862b: 31 [Poneridae]; Mayr, 1865: 11 [Poneridae]; Mayr, 1868c: 24 [Poneridae]; Forel, 1870: 307 [Poneridae]; Forel, 1874: 21 [Poneridae]; Emery, 1877a: 70 [Poneridae]; Forel, 1878: 366 [Poneridae]; Emery & Forel, 1879a: 455 [Poneridae]; André, 1881b: 64 [Poneridae]; Forel, 1892j: 220 [Poneridae]; Dalla Torre, 1893: 13; Forel, 1893a: 162; Forel, 1895b: 111 [Poneridae]; Nasonov, 1889: 27 [Poneridae]; Emery, 1895j: 766 [subfamily spelled Ponerini]; Emery, 1896e: 176; Forel, 1899c: 2; Emery, 1901a: 36; Bingham, 1903: 23; Wheeler, W.M. 1910g: 134; Emery, 1911d: 2; Wheeler, 1915g: 805 [Ponerides]; Donisthorpe, 1915d: 65; Wheeler, W.M. 1915h: 25; Arnold, 1915: 9; Escherich, 1917: 2 [Ponerini]; Forel, 1917: 235; Bondroit, 1918: 78 [Poneritae]; Wheeler, W.M. 1920: 53; Wheeler, W.M. 1922a: 56, 632, 640; Borgmeier, 1923: 37; Karavaiev, 1934: 49; Clark, 1951: 15; Brown, 1954e: 24; Wheeler, G.C. & Wheeler, J. 1972a: 39; Brown, 1973b: 165; all subsequent authors.

Ponerinae as poneromorph subfamily of Formicidae: Bolton, 2003: 42, 156.

Ponerinae as poneroid subfamily of Formicidae: Ouellette, Fisher, et al. 2006: 365; Brady, Schultz, et al. 2006: 18173; Moreau, Bell et al. 2006: 102; Ward, 2007a: 555.

Tribes of Ponerinae: Platythyreini, Ponerini, Thaumatomyrmecini.

Genera (extinct) incertae sedis in Ponerinae: †Afropone, †Eogorgites, †Eoponerites, †Furcisutura, †Longicapitia.

Subfamily Ponerinae references, world

Smith, F. 1858b: 76 (diagnosis); Mayr, 1862: 712 (genera key); Mayr, 1865: 11 (Odontomachidae, Poneridae diagnoses), Mayr, 1867a: 79, 81 (Odontomachidae, Poneridae diagnoses); Forel, 1878: 366 (diagnosis); Dalla Torre, 1893: 13 (world catalogue); Emery, 1895j: 766 (diagnosis); Emery, 1896e: 176 (genera key); Handlirsch, 1907: 879 (†fossil taxa catalogue); Wheeler, W.M. 1910g: 134 (diagnosis); Emery, 1911d: 3 (diagnosis, tribes key); Forel, 1917: 235 (synoptic classification); Forel, 1921c: 133 (diagnosis); Wheeler, W.M. 1922a: 56, 636 (diagnosis, tribes key); Brown & Nutting, 1950: 124 (venation, phylogeny); Brown, 1954e: 24 (phylogeny); Eisner, 1957: 475 (proventriculus morphology); Bernard, 1967: 79 (diagnosis); Gotwald, 1969: 25 (mouthparts morphology); Wheeler, G.C. & Wheeler, J. 1972a: 39 (diagnosis); Brown, 1973b: 165 (genera, distribution); Wheeler, G.C. & Wheeler, J. 1976b: 48 (larvae, review & synthesis); Snelling, R.R. 1981: 387 (synoptic classification); Wheeler, G.C. & Wheeler, J. 1985: 256 (synoptic classification); Billen, 1986: 168 (Dufour's gland); Fanfani & Dazzini Valcurone, 1986: 115 (exocrine glands synopsis); Dlussky & Fedoseeva, 1988: 78 (synoptic classification); Hölldobler & Wilson, 1990: 9 onward (synoptic classification, genera keys); Baroni Urbani, Bolton & Ward, 1992: 317 (phylogeny); Bolton, 1994: 153 (diagnosis, synoptic classification, genera keys); Bolton, 1995a: 1042 (census); Bolton, 1995b: 14 (catalogue); Grimaldi, Agosti & Carpenter, 1997: 20 (†Cretaceous genera); Dlussky & Rasnitsyn, 2002: 422 (diagnosis for wingless fossils); Bolton, 2003: 42, 156 (diagnosis, synopsis); Ouellette, Fisher et al. 2006: 359 (phylogeny); Brady, Schultz, et al. 2006: 18173 (phylogeny); Moreau, Bell et al. 2006: 102 (phylogeny); Ward, 2007a: 555 (classification).

.

Regional and national faunas with keys

Mayr, 1861: 28 (Europe); Mayr, 1868c: 69 (†Baltic Amber); André, 1874: 170 (Europe); Forel, 1874: 29 (Switzerland); André, 1882b: 227 (Europe & Algeria); Cresson, 1887: 97 (U.S.A., genera); Provancher, 1887: 238 (Canada); Nasonov, 1889: 54 (Russia); Forel, 1891b: 9 (Madagascar genera); Forel, 1900c: 52 (India & Sri Lanka); Bingham, 1903: 23 (India, Sri Lanka & Burma); Ruzsky, 1905b: 108 (Russian Empire); Wheeler, W.M. 1910g: 557 (North America genera); Bondroit, 1910: 489 (Belgium); Stitz, 1914: 54 (Central Europe); Gallardo, 1915: 31 (Argentina genera); Forel, 1915d: 6 (Switzerland); Arnold, 1915: 10 (South Africa); Donisthorpe, 1915d: 65 (Britain); Emery, 1916b: 98 (Italy); Wheeler, W.M. 1916m: 580 (U.S.A., Connecticut); Bondroit, 1918: 79 (France & Belgium); Gallardo, 1918b: 4 (Argentina); Soudek, 1922: 18 (Czechoslovakia); Stärcke, 1926: 82 (Netherlands); Donisthorpe, 1927b: 68 (Britain); Menozzi & Russo, 1930: 169 (Dominican Republic); Arnol'di, 1933b: 596 (Russia); Karavaiev, 1934: 50 (Ukraine); Smith, M.R. 1937: 823 (Puerto Rico); Stitz, 1939: 58 (Germany); Smith, M.R. 1943f: 278 (U.S.A. males); Buren, 1944a: 279 (U.S.A., Iowa); Smith, M.R. 1947f: 530 (U.S.A. genera); Creighton, 1950a: 31 (North America); Kusnezov, 1956: 11 (Argentina); Brown, 1958h: 11 (New Zealand); Gregg, 1963: 280 (U.S.A., Colorado); Bernard, 1967: 80 (Western Europe); Wilson & Taylor, 1967: 10 (Polynesia); Kempf, 1972a: 262 (Neotropical synoptic classification); Bolton, 1973a: 323 (West Africa genera); Bolton & Collingwood, 1975: 3 (Britain); Boven, 1977: 66 (Belgium); Kutter, 1977c: 21 (Switzerland); Arnol'di & Dlussky, 1978: 522 (former European U.S.S.R.); Collingwood, 1978: 74 (Iberian Peninsula); Collingwood, 1979: 29 (Fennoscandia & Denmark); Greenslade, 1979: 14 (South Australia genera); Francoeur, 1979: 30 (Canada, Québec); Schembri & Collingwood, 1981: 417 (Malta); Allred, 1982: 438 (U.S.A., Utah); Baroni Urbani, 1984: 75 (Neotropical genera); Collingwood, 1985: 236 (Saudi Arabia); Wheeler, G.C. & Wheeler, J. 1986g: 17 (U.S.A., Nevada); Agosti & Collingwood, 1987b: 264 (Balkans); Ogata, 1987: 101 (Japan genera); Morisita, Kubota, et al. 1989: 8 (Japan); Dlussky, Soyunov & Zabelin, 1990: 173 (Turkmenistan); Kupyanskaya, 1990: 85 (Far Eastern Russia); Atanasov & Dlussky, 1992: 51 (Bulgaria); Lattke, in Jaffe, 1993: 166 (Neotropical genera); Arakelian, 1994: 10 (Armenia); Wu, J. & Wang, 1995: 31 (China genera); Collingwood & Agosti, 1996: 308 (Saudi Arabia); Seifert, 1996b: 107 (Central Europe); Collingwood & Prince, 1998: 10 (Portugal); Kim, Kim & Kim, 1998: 145 (Korea); Shattuck, 1999: 52, 179 (Australia genera synopsis); Andersen, 2000: 19 (northern Australia genera); Zhou, 2001b: 22 (China, Guangxi); Czechowski, Radchenko & Czechowska, 2002: 133 (Poland); Aktaç & Radchenko, 2002: 53 (Turkey genera); Yoshimura & Onoyama, 2002: 436 (Japan genera, males); Mackay & Mackay, 2002: 27 (U.S.A., New Mexico); Csösz, 2003: 147 (Carpathian Basin); Palacio & Fernández, in Fernández, 2003d: 256 (Neotropical genera); Radchenko, 2005b: 184 (North Korea); Coovert, 2005: 23 (U.S.A., Ohio); Yoshimura & Fisher, 2007: 27 (Malagasy genera, males); Clouse, 2007b: 190 (Micronesia); Seifert, 2007: 108 (North and Central Europe); Bolton & Fisher, 2008c: 32 (Afrotropical genera); Fernández & Arias-Penna, 2008: 32 (Neotropical genera); Dlussky, 2009: 1071 (Eocene amber); Terayama, 2009: 101 (Taiwan genera); Heterick, 2009: 34 (south-western Australia genera); Boer, 2010: 68 (Benelux).

Formicinae Camponotini Camponotus eperiamorum TRUE TRUE FALSE

eperiamorum. Camponotus eperiamorum Clouse, 2007a: 3, pl. 1A-C (w.q.m.) MICRONESIA (Pohnpei I.). [Also described as new by Clouse, 2007b: 202, 213.]

Formicinae Camponotini Camponotus erythrocephalus TRUE TRUE FALSE

erythrocephalus. Camponotus erythrocephalus Clouse, 2007a: 6, pl. 1D-F (w.) MICRONESIA (Yap I.). [Also described as new by Clouse, 2007b: 201, 214.]

Formicinae Camponotini Camponotus marianensis TRUE TRUE FALSE

marianensis. Camponotus marianensis Clouse, 2007a: 7, pl. 2D-F, 3A (w.) MICRONESIA (Yap I.). [Also described as new by Clouse, 2007b: 201, 214.]

Formicinae Camponotini Camponotus irritans melanogaster TRUE TRUE FALSE

melanogaster. Camponotus (Tanaemyrmex) irritans var. melanogaster Stitz, 1938: 114 (q.) MICRONESIA (Caroline Is).

Myrmicinae Myrmecinini Pristomyrmex largus TRUE TRUE FALSE

largus. Pristomyrmex largus Wang, M. 2003: 497, figs. 213-214 (w.q.) MICRONESIA (Ponape I.).

Myrmicinae Formicoxenini Romblonella yapensis TRUE TRUE FALSE

yapensis. Romblonella yapensis Smith, M.R. 1953a: 78 (w.) MICRONESIA (Caroline Is).

Myrmicinae Stenammini Vollenhovia kaselela TRUE TRUE FALSE

kaselela. Vollenhovia kaselela Clouse, 2007a: 13, pl. 5 (w.) MICRONESIA (Pohnpei I.). [Also described as new by Clouse, 2007b: 192, 234.]

Myrmicinae Stenammini Vollenhovia mwereka TRUE TRUE FALSE

mwereka. Vollenhovia mwereka Clouse, 2007a: 15, pl. 6A-C (w.) MICRONESIA (Pohnpei I.). [Also described as new by Clouse, 2007b: 192, 234.]

Myrmicinae Stenammini Vollenhovia pwidikidika TRUE TRUE FALSE

pwidikidika. Vollenhovia pwidikidika Clouse, 2007a: 17, pl. 6D-F (w.) MICRONESIA (Pohnpei I.). [Also described as new by Clouse, 2007b: 192, 225.]

Ponerinae Ponerini Ponera bableti TRUE TRUE FALSE

bableti. Ponera bableti Perrault, 1993: 334 (w.) FRENCH POLYNESIA (Mururoa I.).

Formicinae Camponotini Camponotus flavicomans TRUE TRUE FALSE

flavicomans. Camponotus flavicomans Clouse, 2007a: 6, pl. 2A-C (w.) PALAU I. [Also described as new by Clouse, 2007b: 201, 214.]

Formicinae Camponotini Camponotus peleliuensis TRUE TRUE FALSE

peleliuensis. Camponotus peleliuensis Clouse, 2007a: 9, pl. 3B-D (w.) PALAU (Peleliu I.). [Also described as new by Clouse, 2007b: 201, 214.]

Formicinae Camponotini Camponotus irritans kubaryi TRUE TRUE FALSE

kubaryi. Camponotus kubaryi Mayr, 1876: 66 (s.w.) PALAU IS. Combination in C. (Myrmoturba): Forel, 1914a: 267; in C. (Tanaemyrmex): Emery, 1925b: 94. Subspecies of maculatus: Emery, 1896d: 370. Currently subspecies of irritans: Emery, 1920c: 7; Karavaiev, 1929c: 236.

Leptanillinae Leptanillini Leptanilla palauensis TRUE TRUE FALSE

palauensis. Probolomyrmex palauensis Smith, M.R. 1953c: 128, figs. 1, 2 (m.) PALAU IS. Combination in Leptanilla: Taylor, 1965d: 363. See also: Petersen, 1968: 596.

Myrmicinae Myrmecinini Pristomyrmex minusculus TRUE TRUE FALSE

minusculus. Pristomyrmex minusculus Wang, M. 2003: 507, figs. 229-232 (w.q.) PALAU IS (Peleliu I.).

Myrmicinae Formicoxenini Romblonella palauensis TRUE TRUE FALSE

palauensis. Romblonella palauensis Smith, M.R. 1953f: 164 (w.m.) PALAU IS.

Myrmicinae Dacetini Strumigenys stemonixys TRUE TRUE FALSE

stemonixys. Strumigenys stemonixys Brown, 1971c: 81, figs. 2, 4 (w.) PALAU IS. See also: Bolton, 2000: 906.

Formicinae Camponotini Camponotus flavicomans TRUE TRUE FALSE

flavicomans. Camponotus flavicomans Clouse, 2007a: 6, pl. 2A-C (w.) PALAU I. [Also described as new by Clouse, 2007b: 201, 214.]

Formicinae Camponotini Camponotus irritans kubaryi TRUE TRUE FALSE

kubaryi. Camponotus kubaryi Mayr, 1876: 66 (s.w.) PALAU IS. Combination in C. (Myrmoturba): Forel, 1914a: 267; in C. (Tanaemyrmex): Emery, 1925b: 94. Subspecies of maculatus: Emery, 1896d: 370. Currently subspecies of irritans: Emery, 1920c: 7; Karavaiev, 1929c: 236.

Leptanillinae Leptanillini Leptanilla palauensis TRUE TRUE FALSE

palauensis. Probolomyrmex palauensis Smith, M.R. 1953c: 128, figs. 1, 2 (m.) PALAU IS. Combination in Leptanilla: Taylor, 1965d: 363. See also: Petersen, 1968: 596.

Myrmicinae Myrmecinini Pristomyrmex minusculus TRUE TRUE FALSE

minusculus. Pristomyrmex minusculus Wang, M. 2003: 507, figs. 229-232 (w.q.) PALAU IS (Peleliu I.).

Myrmicinae Formicoxenini Romblonella palauensis TRUE TRUE FALSE

palauensis. Romblonella palauensis Smith, M.R. 1953f: 164 (w.m.) PALAU IS.

Myrmicinae Dacetini Strumigenys stemonixys TRUE TRUE FALSE

stemonixys. Strumigenys stemonixys Brown, 1971c: 81, figs. 2, 4 (w.) PALAU IS. See also: Bolton, 2000: 906.

Formicinae Camponotini Camponotus irritans kubaryi TRUE TRUE FALSE

kubaryi. Camponotus kubaryi Mayr, 1876: 66 (s.w.) PALAU IS. Combination in C. (Myrmoturba): Forel, 1914a: 267; in C. (Tanaemyrmex): Emery, 1925b: 94. Subspecies of maculatus: Emery, 1896d: 370. Currently subspecies of irritans: Emery, 1920c: 7; Karavaiev, 1929c: 236.

Leptanillinae Leptanillini Leptanilla palauensis TRUE TRUE FALSE

palauensis. Probolomyrmex palauensis Smith, M.R. 1953c: 128, figs. 1, 2 (m.) PALAU IS. Combination in Leptanilla: Taylor, 1965d: 363. See also: Petersen, 1968: 596.

Myrmicinae Myrmecinini Pristomyrmex minusculus TRUE TRUE FALSE

minusculus. Pristomyrmex minusculus Wang, M. 2003: 507, figs. 229-232 (w.q.) PALAU IS (Peleliu I.).

Myrmicinae Formicoxenini Romblonella palauensis TRUE TRUE FALSE

palauensis. Romblonella palauensis Smith, M.R. 1953f: 164 (w.m.) PALAU IS.

Myrmicinae Dacetini Strumigenys stemonixys TRUE TRUE FALSE

stemonixys. Strumigenys stemonixys Brown, 1971c: 81, figs. 2, 4 (w.) PALAU IS. See also: Bolton, 2000: 906.

Myrmicinae Stenammini Vollenhovia cristata TRUE TRUE FALSE

cristata. Aratromyrmex cristatus Stitz, 1938: 108 (q.) PHILIPPINES (Palawan I.). Combination in Vollenhovia: Baroni Urbani, 1980a: 95.

Myrmicinae Stenammini Vollenhovia luctuosa major TRUE TRUE FALSE

major. Aratromyrmex luctuosus var. major Stitz, 1938: 108 (q.) PHILIPPINES (Palawan I.). Combination in Vollenhovia: Baroni Urbani, 1980a: 95.

Myrmicinae Dacetini Strumigenys TRUE TRUE FALSE

Extant: 487 species

Fossil:

Strumigenys Smith, F. 1860c: 72. Type-species: Strumigenys mandibularis, by monotypy.

Taxonomic history

Strumigenys in Myrmicidae: Smith, F. 1871a: 334; Cresson, 1887: 262.

Strumigenys in Myrmicidae, Cryptoceridae: Emery, 1877a: 81.

Strumigenys in Myrmicinae: Mayr, 1865: 26 [Myrmicidae]; Emery & Forel, 1879a: 464 [Myrmicidae]; Dalla Torre, 1893: 145.

Strumigenys in Cryptoceridae, Dacetini: Ashmead, 1905b: 384.

Strumigenys in Myrmicinae, Dacetini: Forel, 1892d: 344; Forel, 1893a: 164; Forel, 1895b: 133; Emery, 1895j: 770; Forel, 1899c: 42; Wheeler, W.M. 1910g: 141; Emery, 1914a: 42; Forel, 1917: 246; Arnold, 1917: 372; Wheeler, W.M. 1922a: 668; Emery, 1924d: 319; Brown, 1948e: 108; all subsequent authors.

Junior synonyms of STRUMIGENYS

Labidogenys Roger, 1862a: 249. Type-species: Labidogenys lyroessa, by monotypy.

Taxonomic history

Labidogenys in Myrmicinae, Dacetini: Donisthorpe, 1943f: 654; Brown, 1948e: 102.

Labidogenys as genus: Roger, 1862a: 249; Brown, 1948e: 102.

Labidogenys as junior synonym of Strumigenys: Roger, 1863b: 40; Emery & Forel, 1879a: 464; Dalla Torre, 1893: 145; Bingham, 1903: 147; Emery, 1924d: 320; Brown & Wilson, 1959b: 281; Brown, 1960b: 38; Bolton, 1983: 358; Bolton, 1999: 1673; Bolton, 2000: 492.

Proscopomyrmex Patrizi, 1946: 294. Type-species: Proscopomyrmex londianensis, by monotypy.

Taxonomic history

Proscopomyrmex in Myrmicinae, Dacetini: Patrizi, 1946: 294 [Dacetonini].

Proscopomyrmex as junior synonym of Strumigenys: Brown, 1949d: 15; Bolton, 1999: 1673; Bolton, 2000: 492.

Eneria Donisthorpe, 1948d: 598. Type-species: Eneria excisa (junior synonym of Strumigenys loriae), by original designation.

Taxonomic history

Eneria as genus: Chapman & Capco, 1951: 104 (anachronism).

Eneria as junior synonym of Strumigenys: Brown, 1949d: 15; Bolton, 1999: 1673; Bolton, 2000: 492.

Quadristruma Brown, 1949b: 47. Type-species: Epitritus emmae, by original designation.

Taxonomic history

Quadristruma in Myrmicinae, Dacetini: Brown, 1949b: 47; all subsequent authors.

Quadristruma as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 12; Bolton, 1999: 1673; Bolton, 2000: 492.

Genus Strumigenys references

Roger, 1863b: 40 (catalogue); Mayr, 1863: 424, 455 (catalogue); Mayr, 1865: 26 (diagnosis); Forel, 1886a: 216 (all species key); Mayr, 1887: 568 (all species key); Dalla Torre, 1893: 145 (catalogue); Emery, 1895c: 325 (North America species key); Emery, 1897c: 574 (New Guinea species key); Forel, 1903a: 707 (India & Sri Lanka species key); Bingham, 1903: 147 (India, Sri Lanka & Burma species key); Wheeler, W.M. 1908a: 146 (New World species key); Santschi, 1913b: 257 (Afrotropical species key); Arnold, 1917: 373 (diagnosis, South Africa species key); Mann, 1921: 467 (Fiji Is species key); Wheeler, W.M. 1922a: 917, 1034 (Afrotropical, Malagasy catalogues); Emery, 1924d: 319 (diagnosis, catalogue); Brown, 1948e: 108 (diagnosis, review of genus); Brown, 1949d: 21 (Japan, China & Taiwan species key); Creighton, 1950a: 301 (North America species key); Chapman & Capco, 1951: 104, 107 (Asia Eneria, Strumigenys (Strumigenys) checklists); Brown, 1954k: 10 (Afrotropical species revision, key); Brown, 1958h: 38 (New Zealand species); Brown, 1962c: 238 (Neotropical species synopsis, key); Kempf, 1972a: 226, 243 (Neotropical Quadristruma, Strumigenys catalogues); Alayo, 1974: 19 (Cuba species key); Kempf, 1976b: 43 (additions to Brown, 1962c key); Smith, D.R. 1979: 1403, 1409 (North America Strumigenys, Quadristruma catalogues); Bolton, 1983: 358, 400 (Afrotropical Strumigenys, Quadristruma species revisions, keys); Taylor & Brown, D.R. 1985: 85, 87 (Australia Quadristruma, Strumigenys catalogues); Taylor, 1987a: 66, 76 (Australia, New Caledonia & New Zealand checklists); Terayama & Kubota, 1989: 780 (Taiwan species key); Brandão, 1991: 380 (Neotropical catalogue); Arakelian & Dlussky, 1991: 150 (former U.S.S.R. species key); Morisita, Kubota, et al. 1992: 63 (Japan species key); Dlussky, 1993: 53 (Fiji Is, Tonga and Samoa species key); Bolton, 1994: 105 (synoptic classification); Bolton, 1995a: 1053 (census); Bolton, 1995b: 377, 395 (catalogue); Gronenberg, 1996: 2021 (mandible morphology); Lin & Wu, 1996: 139 (Taiwan species key); Lattke & Goitía, 1997: 370 (Venezuela species key); Shattuck, 1999: 160, 167 (Australia synopsis); Bolton, 1999: 1657, 1673 (morphology, diagnosis, review of genus, phylogeny); Bolton, 2000: 496, 583, 716, 942 (Neotropical-Nearctic, Afrotropical, Malesian-Oriental-East Palaearctic, Austral species revisions, keys); Fisher, 2000: 614 (Malagasy species revision, key); Zhou, 2001b: 78 (China, Guangxi species key); Lin & Wu, 2001: 166 (Taiwan species key); Zhou & Xu, 2003: 737 (China species key); Yoshimura & Onoyama, 2007: 676 (Japan species key); Terayama, 2009: 142 (Taiwan species key); Heterick, 2009: 176 (south-western Australia species key).

Myrmicinae Adelomyrmecini Adelomyrmex samoanus TRUE TRUE FALSE

samoanus. Adelomyrmex (Arctomyrmex) samoanus Wilson & Taylor, 1967: 77, fig. 62 (w.) SAMOA. See also: Fernández, 2003b: 27.

Ponerinae Ponerini Anochetus graeffei TRUE TRUE FALSE

graeffei. Anochetus graeffei Mayr, 1870b: 961 (w.) SAMOA. Mayr, 1876: 86 (q.); Wheeler, G.C. & Wheeler, J. 1971b: 1212 (l.); Imai, Brown, et al. 1984: 5 (k.). Senior synonym of amati, minutus, oceanicus, punctiventris: Wilson, 1959a: 507; of rudis, ruginotus, taylori: Brown, 1978c: 557 (see also p. 586).

Formicinae Camponotini Camponotus chloroticus TRUE TRUE FALSE

chloroticus. Camponotus maculatus subsp. chloroticus Emery, 1897d: 574 (w.) NEW GUINEA. Emery, 1914f: 424 (m.). Combination in C. (Myrmoturba): Emery, 1914f: 424; in C. (Tanaemyrmex): Emery, 1925b: 94. Subspecies of irritans: Emery, 1920c: 7; Karavaiev, 1933a: 316. Raised to species, senior synonym of chlorogaster, sanctaecrucis and material of the unavailable name samoaensis referred here: Wilson & Taylor, 1967: 93.

Formicinae Camponotini Camponotus flavolimbatus TRUE TRUE FALSE

flavolimbatus. Camponotus (Colobopsis) flavolimbatus Viehmeyer, 1922: 218 (q.) SAMOA. Senior synonym of buxtoni, nautarum: Wilson & Taylor, 1967: 97.

Formicinae Camponotini Camponotus navigator TRUE TRUE FALSE

navigator. Camponotus (Myrmoturba) navigator Wilson & Taylor, 1967: 94, fig. 77 (s.w.q.) SAMOA.

Myrmicinae Basicerotini Eurhopalothrix procera TRUE TRUE FALSE

procera. Rhopalothrix procera Emery, 1897c: 572, pl. 14, fig. 18 (w.q.) NEW GUINEA. Combination in Eurhopalothrix: Brown & Kempf, 1960: 225. Senior synonym of angulinodis, ballionii, borneensis, kokodensis, malua, manni, samoana, subdentatus and material of the unavailable name melanotica referred here: Brown & Kempf, 1960: 225.

Myrmicinae Solenopsidini Monomorium liliuokalanii TRUE TRUE FALSE

liliuokalanii. Monomorium minutum var. liliuokalanii Forel, 1899a: 119 (w.) HAWAII. Subspecies of minutum: Forel, 1909d: 225. Junior synonym of minutum: Wilson & Taylor, 1967: 67. Revived from synonymy, raised to species and senior synonym of samoanum: Bolton, 1987: 288.

Ponerinae Ponerini Pachycondyla insulana TRUE TRUE FALSE

insulana. Ponera insulana Mayr, 1876: 87 (w.) SAMOA. Combination in Pachycondyla (Ectomomyrmex): Emery, 1901a: 46; in Ectomomyrmex: Wilson, 1958d: 364; in Pachycondyla: Brown, in Bolton, 1995b: 306.

Myrmicinae Pheidolini Pheidole aana TRUE TRUE FALSE

aana. Pheidole aana Wilson & Taylor, 1967: 42, fig. 31 (s.w.) SAMOA.

Myrmicinae Pheidolini Pheidole atua TRUE TRUE FALSE

atua. Pheidole atua Wilson & Taylor, 1967: 43, fig. 32 (s.w.) SAMOA.

Ponerinae Ponerini Ponera loi TRUE TRUE FALSE

loi. Ponera loi Taylor, 1967a: 60, figs. 52-55 (w.q.) SAMOA.

Ponerinae Ponerini Ponera woodwardi TRUE TRUE FALSE

woodwardi. Ponera woodwardi Taylor, 1967a: 62, figs. 7, 55-58 (w.q.m.l.) SAMOA.

Myrmicinae Dacetini Pyramica epipola TRUE TRUE FALSE

epipola. Pyramica epipola Bolton, 2000: 399 (w.) SAMOA.

Myrmicinae Stenammini Rogeria exsulans TRUE TRUE FALSE

exsulans. Rogeria exsulans Wilson & Taylor, 1967: 74, fig. 60 (w.) SAMOA. See also: Kugler, C. 1994: 64.

Myrmicinae Dacetini Strumigenys godeffroyi TRUE TRUE FALSE

godeffroyi. Strumigenys godeffroyi Mayr, 1866a: 516 (w.) SAMOA. Mayr, 1887: 569 (q.); Imai, Brown, et al. 1984: 68 (k.). Senior synonym of butteli, indica: Brown, 1949d: 17; of geococci: Bolton, 2000: 791. See also: Bingham, 1903: 149.

Myrmicinae Dacetini Strumigenys mailei TRUE TRUE FALSE

mailei. Strumigenys mailei Wilson & Taylor, 1967: 38, fig. 28 (w.) SAMOA. Dlussky, 1993: 59 (q.). See also: Bolton, 2000: 823.

Dolichoderinae Tapinomini Tapinoma wheeleri TRUE TRUE FALSE

wheeleri. Zatapinoma wheeleri Mann, 1935: 36, fig. 2 (s.) SAMOA. Combination in Tapinoma: Shattuck, 1992c: 152.

Myrmicinae Stenammini Vollenhovia pacifica TRUE TRUE FALSE

pacifica. Vollenhovia pacifica Wilson & Taylor, 1967: 56, fig. 41 (w.q.m.) SAMOA.

Myrmicinae Stenammini Vollenhovia samoensis TRUE TRUE FALSE

samoensis. Vollenhovia samoensis Mayr, 1876: 99 (w.m.) SAMOA. Senior synonym of agilis: Wilson & Taylor, 1967: 57.

Formicinae Camponotini Polyrhachis TRUE TRUE FALSE

Extant: 614 species, 86 subspecies

Fossil: 1 species

Polyrhachis Smith, F. 1857a: 58. Type-species: Formica bihamata, by original designation.

Taxonomic history

[Polyrhachis Shuckard, in Swainson & Shuckard, 1840: 172, nomen nudum.]

Polyrhachis in Formicidae: Smith, F. 1858b: 58.

Polyrhachis in Formicinae: Mayr, 1862: 652 [Formicidae]; Mayr, 1865: 6 [Formicidae].

Polyrhachis in Camponotinae: Forel, 1878: 367 [Camponotidae]; Dalla Torre, 1893: 257.

Polyrhachis in Camponotinae, Polyrhachidini: Ashmead, 1905b: 384.

Polyrhachis in Camponotinae, Camponotini: Forel, 1886f: 194; Forel, 1893a: 165; Emery, 1895j: 772; Wheeler, W.M. 1910g: 144; Forel, 1912i: 89; Forel, 1917: 251; Arnold, 1924: 741.

Polyrhachis in Formicinae, Camponotini: Wheeler, W.M. 1922a: 701; Emery, 1925b: 175; all subsequent authors except the following.

Polyrhachis in Formicinae, Formica genus group: Agosti, 1991: 295.

Polyrhachis as subgenus of Myrma: Wheeler, W.M. 1911c: 860; Wheeler, W.M. 1911f: 170. [Myrma as subgenus of Polyrhachis: Forel, 1915b: 106; Forel, 1917: 251; Wheeler, W.M. 1922a: 702, 993; Emery, 1925b: 198; all subsequent authors (see note under Myrma).]

[Polyrachis Arnold, 1924: 741, incorrect subsequent spelling.]

Subgenera of POLYRHACHIS include the nominal plus the following.

All the earlier subgenera were given as provisional junior synonyms of Polyrhachis by Brown, 1973b: 178 - 184. Brown's list was repeated in Hölldobler & Wilson, 1990: 19 but with all subgenera listed as junior synonyms; subgeneric status was reinstated in Bolton, 1994: 50. These consecutive entries are not repeated individually below.

Subgenus POLYRHACHIS (MYRMA)

Myrma Billberg, 1820: 104. Type-species: Formica militaris, by subsequent designation of Wheeler, W.M. 1911c: 859.

Taxonomic history

[Although Myrma antedates Polyrhachis, the former has been treated as a subgenus of the latter since Forel, 1917: 251 (see Hung, 1967b: 396 for history). Dorow, Kohout & Taylor, 1997: 236 proposed the precedence of Polyrhachis over Myrma and that precedence was established by the ICZN (Opinion 1919) 1999: 92.]

Polyrhachis as subgenus of Myrma: Wheeler, W.M. 1911c: 860; Wheeler, W.M. 1911f: 170.

Myrma as subgenus of Polyrhachis: Forel, 1915b: 106; Forel, 1917: 251; Wheeler, W.M. 1922a: 702, 993; Emery, 1925b: 198; all subsequent authors, see note above.

Junior synonyms of POLYRHACHIS (MYRMA)

Hoplomyrmus Gerstäcker, 1859: 262. Type-species: Hoplomyrmus schistaceus, by monotypy.

Taxonomic history

[Hoplomyrmus also described as new by Gerstäcker, 1862: 508.]

Hoplomyrmus as junior synonym of Polyrhachis: Roger, 1861b: 174; Roger, 1863b: 6; Mayr, 1863: 446; Dalla Torre, 1893: 257.

Hoplomyrmus as junior synonym of Myrma: Wheeler, W.M. 1911c: 860; Wheeler, W.M. 1922a: 993; Emery, 1925b: 198; Bolton, 1973b: 288.

Pseudocyrtomyrma Emery, 1921e: 18 [as subgenus of Polyrhachis]. Type-species: Polyrhachis revoili, by original designation.

Taxonomic history

Pseudocyrtomyrma as subgenus of Polyrhachis: Emery, 1921e: 18; Emery, 1925b: 206.

Pseudocyrtomyrma as junior synonym of Myrma: Bolton, 1973b: 288.

Anoplomyrma Chapman, 1963: 258 [as subgenus of Polyrhachis]. Type-species: Polyrhachis (Anoplomyrma) parabiotica, by monotypy.

Taxonomic history

Anoplomyrma as junior synonym of Myrma: Dorow, 1995: 30.

Subgenus POLYRHACHIS (HEMIOPTICA)

Hemioptica Roger, 1862a: 238. Type-species: Hemioptica scissa, by monotypy.

Taxonomic history

Hemioptica in Formicinae: Mayr, 1862: 769 [Formicidae]; Mayr, 1865: 6 [Formicidae].

Hemioptica in Camponotinae: Dalla Torre, 1893: 271.

Hemioptica in Camponotinae, Camponotini: Emery, 1895j: 772; Wheeler, W.M. 1910g: 144; all subsequent authors except the following.

Hemioptica in Camponotinae, Polyrhachidini: Ashmead, 1905b: 384.

Hemioptica as junior synonym of Polyrhachis: Forel, 1878: 368.

Hemioptica as subgenus of Myrma: Wheeler, W.M. 1911c: 860; Wheeler, W.M. 1911f: 164.

Hemioptica as genus: Roger, 1862a: 238; Mayr, 1862: 769; Mayr, 1865: 6; Dalla Torre, 1893: 271; Emery, 1895j: 772; Ashmead, 1905b: 384; Wheeler, W.M. 1910g: 144; Emery, 1925b: 209; Donisthorpe, 1943f: 649; Chapman & Capco, 1951: 255.

Hemioptica as subgenus of Polyrhachis: Mayr, 1872: 138; Forel, 1915b: 107; Forel, 1917: 251; Wheeler, W.M. 1922a: 701; Dorow & Kohout, 1995: 95; Bolton, 1995b: 31; Dorow, 1995: 29.

Subgenus POLYRHACHIS (HAGIOMYRMA)

Hagiomyrma Wheeler, W.M. 1911c: 860 [as subgenus of Polyrhachis]. Type-species: Formica ammon, by original designation.

Taxonomic history

Hagiomyrma as subgenus of Polyrhachis: Wheeler, W.M. 1911c: 860; Forel, 1917: 251; Wheeler, W.M. 1922a: 702; Emery, 1925b: 184; all subsequent authors.

Subgenus POLYRHACHIS (HIRTOMYRMA)

Hirtomyrma Kohout, 2010: 199 [as subgenus of Polyrhachis]. Type-species: Polyrhachis hirta, by original designation.

Taxonomic history

Hirtomyrma as subgenus of Polyrhachis: Kohout, 2010: 199.

Subgenus POLYRHACHIS (CAMPOMYRMA)

Campomyrma Wheeler, W.M. 1911g: 860 [as subgenus of Polyrhachis]. Type-species: Polyrhachis clypeata (junior synonym of Formica exercita), by original designation.

Taxonomic history

Campomyrma as subgenus of Polyrhachis: Forel, 1917: 251; Wheeler, W.M. 1922a: 702; Emery, 1925b: 178; all subsequent authors.

Subgenus POLYRHACHIS (MYRMOTHRINAX)

Myrmothrinax Forel, 1915b: 107 [as subgenus of Polyrhachis]. Type-species: Polyrhachis thrinax, by original designation.

Taxonomic history

Myrmothrinax as subgenus of Polyrhachis: Forel, 1915b: 107; Forel, 1917: 251; Wheeler, W.M. 1922a: 704; Emery, 1925b: 178; all subsequent authors.

Junior synonym of POLYRHACHIS (MYRMOTHRINAX)

Evelyna Donisthorpe, 1937c: 273 [as subgenus of Polyrhachis]. Type-species: Polyrhachis (Evelyna) cheesmanae, by original designation.

Taxonomic history

Evelyna as junior synonym of Myrmothrinax: Hung, 1967b: 402.

Subgenus POLYRHACHIS (CYRTOMYRMA)

Cyrtomyrma Forel, 1915b: 107 [as subgenus of Polyrhachis]. Type-species: Formica rastellata, by original designation.

Taxonomic history

Cyrtomyrma as subgenus of Polyrhachis: Forel, 1915b: 107; Forel, 1917: 251; Wheeler, W.M. 1922a: 701; Emery, 1925b: 207; all subsequent authors.

Subgenus POLYRHACHIS (MYRMHOPLA)

Myrmhopla Forel, 1915b: 107 [as subgenus of Polyrhachis]. Type-species: Formica armata, by original designation.

Taxonomic history

Myrmhopla as subgenus of Polyrhachis: Forel, 1915b: 107; Forel, 1917: 251; Wheeler, W.M. 1922a: 701; Emery, 1925b: 190; all subsequent authors.

Junior synonyms of POLYRHACHIS (MYRMHOPLA)

Cephalomyrma Karavaiev, 1935a: 115 [as subgenus of Polyrhachis]. Type-species: Polyrhachis (Cephalomyrma) stylifera, by monotypy.

Taxonomic history

Cephalomyrma as junior synonym of Myrmhopla: Hung, 1967b: 402.

Florencea Donisthorpe, 1937a: 624 [as subgenus of Polyrhachis]. Type-species: Polyrhachis (Florencea) kirkae (junior synonym of Polyrhachis nigriceps), by original designation.

Taxonomic history

Florencea as junior synonym of Myrmhopla: Hung, 1967b: 402.

Subgenus POLYRHACHIS (CHARIOMYRMA)

Chariomyrma Forel, 1915b: 107 [as subgenus of Polyrhachis]. Type-species: Polyrhachis guerini, by original designation.

Taxonomic history

Chariomyrma as subgenus of Polyrhachis: Forel, 1915b: 107; Forel, 1917: 251; Wheeler, W.M. 1922a: 702; Emery, 1925b: 185; all subsequent authors.

Subgenus POLYRHACHIS (HEDOMYRMA)

Hedomyrma Forel, 1915b: 107 [as subgenus of Polyrhachis]. Type-species: Polyrhachis ornata, by original designation.

Taxonomic history

Hedomyrma as subgenus of Polyrhachis: Forel, 1915b: 107; Forel, 1917: 251; Wheeler, W.M. 1922a: 702; Emery, 1925b: 189; all subsequent authors.

Junior synonyms of POLYRHACHIS (HEDOMYRMA)

Dolichorhachis Mann, 1919: 386 [as subgenus of Polyrhachis]. Type-species: Polyrhachis (Dolichorhachis) malaensis, by monotypy.

Taxonomic history

Dolichorhachis as subgenus of Polyrhachis: Mann, 1919: 386; Wheeler, W.M. 1922a: 702; Emery, 1925b: 188.

Dolichorhachis as junior synonym of Hedomyrma: Dorow, 1995: 26.

Morleyidris Donisthorpe, 1944c: 64 [as subgenus of Polyrhachis]. Type-species: Polyrhachis (Morleyidris) trina, by original designation.

Taxonomic history

Morleyidris as junior synonym of Hedomyrma: Hung, 1967b: 402.

Subgenus POLYRHACHIS (MYRMATOPA)

Myrmatopa Forel, 1915b: 107 [as subgenus of Polyrhachis]. Type-species: Polyrhachis schang, by original designation.

Taxonomic history

Myrmatopa as subgenus of Polyrhachis: Forel, 1917: 251; Wheeler, W.M. 1922a: 702; Emery, 1925b: 180; all subsequent authors.

Junior synonym of POLYRHACHIS (MYRMATOPA)

Irenea Donisthorpe, 1938e: 502 [as subgenus of Dolichoderus]. Type-species: Dolichoderus (Irenea) omyrmex, by original designation.

Taxonomic history

Irenea in Dolichoderinae, Dolichoderini: Donisthorpe, 1938e: 502; Donisthorpe, 1943f: 653; Chapman & Capco, 1951: 186.

Irenea as subgenus of Dolichoderus: Donisthorpe, 1938e: 502.

Irenea as genus: Donisthorpe, 1941h: 59; Chapman & Capco, 1951: 186.

Irenea as junior synonym of Polyrhachis (Myrmatopa): Bolton, 1994: 50; Dorow, 1995: 42.

Subgenus POLYRHACHIS (AULACOMYRMA)

Aulacomyrma Emery, 1921e: 17 [as subgenus of Polyrhachis]. Type-species: Polyrhachis porcata, by original designation.

Taxonomic history

Aulacomyrma as subgenus of Polyrhachis: Emery, 1921e: 17; Emery, 1925b: 197; all subsequent authors.

Junior synonym of POLYRHACHIS (AULACOMYRMA)

Johnia Karavaiev, 1927e: 43 [as subgenus of Polyrhachis]. Type-species: Polyrhachis (Johnia) schizospina, by monotypy.

Taxonomic history

Johnia as junior synonym of Aulacomyrma: Hung, 1967b: 402.

Genus Polyrhachis references

Smith, F. 1858b: 58 (diagnosis); Roger, 1863b: 6 (catalogue); Mayr, 1863: 422, 443 (Hemioptica, Polyrhachis catalogues); Mayr, 1865: 6 (Polyrhachis, Hemioptica diagnoses); Mayr, 1867a: 40, 41 (diagnosis, species groups (as "turmae")); Mayr, 1876: 68 (Australia species key); Forel, 1878: 368 (diagnosis); Mayr, 1879: 648 (species groups); André, 1887: 286 (Afrotropical species key); Dalla Torre, 1893: 257, 271 (Polyrhachis, Hemioptica catalogues); Forel, 1893c: 17 (India & Sri Lanka species key); Emery, 1896d: 378 (catalogue); Emery, 1897d: 579, 582 (P. relucens & P. guerini groups, keys); Emery, 1898a: 227 (additions to 1896a catalogue); Bingham, 1903: 383 (India, Sri Lanka & Burma species key); Santschi, 1910h: 284 (P. rastellata group, key); Viehmeyer, 1914b: 50, 54 (P. rastellata & P. hostilis groups, keys); Wheeler, W.M. 1922a: 256, 701, 992 (diagnosis, subgenera key, Afrotropical catalogue); Arnold, 1924: 741 (diagnosis, South Africa species key); Emery, 1925b: 175 (diagnosis, subgenera key, catalogue); Emery, 1925b: 178 (P. (Campomyrma) diagnosis, catalogue); Emery, 1925b: 180 (P. (Myrmatopa) diagnosis, catalogue); Emery, 1925b: 181 (P. (Polyrhachis) diagnosis, catalogue); Emery, 1925b: 182 (P. (Myrmothrinax) diagnosis, catalogue); Emery, 1925b: 184 (P. (Hagiomyrma) diagnosis, catalogue); Emery, 1925b: 185 (P. (Chariomyrma) diagnosis, catalogue); Emery, 1925b: 188 (P. (Dolichorhachis) diagnosis, catalogue); Emery, 1925b: 189 (P. (Hedomyrma) diagnosis, catalogue); Emery, 1925b: 190 (P. (Myrmhopla) diagnosis, catalogue); Emery, 1925b: 197 (P. (Aulacomyrma) diagnosis, catalogue); Emery, 1925b: 198 (P. (Myrma) diagnosis, catalogue); Emery, 1925b: 206 (P. (Pseudocyrtomyrma) diagnosis, catalogue); Emery, 1925b: 207 (P. (Cyrtomyrma) diagnosis, catalogue); Emery, 1925b: 209 (Hemioptica diagnosis, catalogue); Donisthorpe, 1938c: 246 (P. (Cyrtomyrma) review); Santschi, 1939c: 13 (P. (Pseudocyrtomyrma) species key); Chapman & Capco, 1951: 186, 255, 256 (Asia Irenea, Hemioptica, Polyrhachis checklists); Hung, 1962: 24 (Taiwan species key); Hung, 1967b: 395 (review of subgenera); Hung, 1970: 5 (P. (Polyrhachis) species revision, key); Bolton, 1973b: 289 (Afrotropical species revision, key); Bolton, 1975c: 1 (P. sexspinosa group, key); Taylor & Brown, D.R. 1985: 131 (Australia catalogue); Taylor, 1987a: 59 (Australia, New Caledonia checklist); Kohout, 1987: 169 (Philippines P. sexspinosa group, key); Kohout, 1989: 515 (Australia P. relucens group, key); Kohout, 1990: 500 (P. viehmeyeri group, key); Kohout & Taylor, 1990: 509 (Australia checklist); Morisita, Kubota, Onoyama, et al. 1991: 45 (Japan species key); Wang C. & Wu, 1991: 596 (China species key); Dorow & Kohout, 1995: 96 (P. (Hemioptica) species key); Bolton, 1994: 50 (synoptic classification); Bolton, 1995a: 1052 (census); Bolton, 1995b: 342 (catalogue); Dorow, 1995: 9 (subgenera revision, subgenera key, catalogue); Dorow, 1995: 10-11 (P. (Myrmhopla) species groups, key; P. arachne-, P. cryptoceroides- P. flavoflagellata-, P. furcata- and P. hector species groups, keys); Dorow, 1995: 12 (P. (Aulacomyrma) synopsis); Dorow, 1995: 13 (P. (Campomoyrma) synopsis); Dorow, 1995: 16 (P. (Chariomyrma) synopsis); Dorow, 1995: 21 (P. (Cyrtomyrma) synopsis); Dorow, 1995: 24 (P. (Hagiomyrma) synopsis); Dorow, 1995: 26 (P. (Hedomyrma) synopsis); Dorow, 1995: 28 (P. (Hemioptica) synopsis); Dorow, 1995: 30 (P. (Myrma) synopsis); Dorow, 1995: 42 (P. (Myrmatopa) synopsis); Dorow, 1995: 45 (P. (Myrmhopla) synopsis); Dorow, 1995: 61 (P. (Myrmothrinax) synopsis); Dorow, 1995: 63 (P. (Polyrhachis) synopsis); Wu, J. & Wang, 1995: 159 (China species key); Shattuck, 1999: 107 (Australia synopsis); Zhou, 2001b: 190 (China, Guangxi species key); Xu, 2002b: 523 (China P. (Cyrtomyrma) species key; Bolton, 2003: 122 (synopsis); Karmaly, 2004: 542 (India species key); Kohout, 2006a: 146 (P. cryptoceroides group, key); Kohout, 2006b: 87 (P. Cyrtomyrma) of Australia, Borneo, New Guinea, Solomon Is, keys); Kohout, 2007a: 189 (P. Aulacomyrma) diagnosis, all species revision, key); Kohout, 2007c: 8 (P. (Campomyrma) Borneo, key); Kohout, 2008a: 255 (Sulawesi species, keys); Kohout, 2008b: 298 (P. vestita group key); Kohout, 2008d: 12 (flavoflagellata group key); Kohout, 2009: 40, 44 (modifications to Kohout 2006b key); Terayama, 2009: 218 (Taiwan species key); Heterick, 2009: 106 (south-western Australia species key); Kohout, 2010: 169, 171, 200 (Australia subgenera key, Australia P. (Myrmhopla) species key, Australia P. (Hirtomyrma) species key).

Ponerinae Ponerini Anochetus TRUE TRUE FALSE

Extant: 94 species

Fossil: 8 species

Anochetus Mayr, 1861: 53. Type-species: Odontomachus ghilianii, by monotypy.

Taxonomic history

Anochetus in Formicidae, Odontomachidae: Mayr, 1862: 712; Mayr, 1865: 11.

Anochetus in Odontomachidae: Ashmead, 1905b: 382.

Anochetus in Ponerinae: Mayr, 1861: 53 [Poneridae]; Emery & Forel, 1879a: 455 [Poneridae]; Dalla Torre, 1893: 47.

Anochetus in Ponerinae, Odontomachini: Forel, 1893a: 163; Emery, 1895j: 768; Forel, 1899c: 18; Wheeler, W.M. 1910g: 136; Emery, 1911d: 106; Arnold, 1915: 103; Forel, 1917: 238; Gallardo, 1918b: 89; Wheeler, W.M. 1922a: 653; Donisthorpe, 1943f: 623; Chapman & Capco, 1951: 31; Wilson, 1959a: 483; Wheeler, G.C. & Wheeler, J. 1985: 256; Dlussky & Fedoseeva, 1988: 78; Hölldobler & Wilson, 1990: 11; Jaffe, 1993: 8.

Anochetus in Ponerinae, Ponerini: Brown, 1976a: 71; Brown, 1978c: 550 [subtribe Odontomachiti]; Bolton, 1994: 164; Bolton, 2003: 159.

Junior synonyms of ANOCHETUS

Stenomyrmex Mayr, 1862: 711. Type-species: Myrmecia emarginata, by subsequent designation of Wheeler, W.M. 1911f: 173.

Taxonomic history

Stenomyrmex in Formicidae, Odontomachidae: Mayr, 1862: 711; Mayr, 1865: 11.

Stenomyrmex in Odontomachidae: Ashmead, 1905b: 382.

Stenomyrmex as subgenus of Anochetus: Dalla Torre, 1893: 47; Forel, 1893a: 163; Dalla Torre, 1893: 47; Wheeler, W.M. 1910g: 136; Emery, 1911d: 110; Forel, 1917: 238; Wheeler, W.M. 1922a: 653; Borgmeier, 1923: 76; Donisthorpe, 1943g: 728; Kempf, 1972a: 21.

Stenomyrmex as junior synonym of Anochetus: Forel, 1887: 382; Brown, 1978c: 552; all subsequent authors.

Myrmapatetes Wheeler, W.M. 1929b: 6. Type-species: Myrmapatetes filicornis, by original designation.

Taxonomic history

Myrmapatetes in Dolichoderinae: Wheeler, W.M. 1929b: 7; Chapman & Capco, 1951: 186.

Myrmapatetes as junior synonym of Anochetus: Brown, 1953h: 2.

Genus Anochetus references

Roger, 1863b: 21, 22 (catalogue); Mayr, 1863: 394, 454 (catalogue); Mayr, 1865: 11 (Anochetus, Stenomyrmex diagnoses); André, 1882b: 230 (Europe & Algeria species); Emery, 1884a: 378 (all species key); Dalla Torre, 1893: 47 (catalogue); Emery, 1894c: 185 (Neotropical species key); Forel, 1900c: 58 (India & Sri Lanka species key); Bingham, 1903: 39 (India, Sri Lanka & Burma species key); Emery, 1911d: 107 (diagnosis, catalogue); Arnold, 1915: 103 (South Africa species key); Gallardo, 1918b: 90 (Argentina species key); Mann, 1919: 303 (New Guinea & Solomon Is species key); Wheeler, W.M. 1922a: 96, 790, 1012 (diagnosis, Afrotropical, Malagasy catalogues); Wheeler, W.M. 1925a: 9 (Stenomyrmex species key); Chapman & Capco, 1951: 39 (Asia checklist); Wilson, 1959a: 503 (Melanesia & Moluccas species, revision, key); Kempf, 1964f: 244 (A. (Stenomyrmex) species key); Kempf, 1972a: 20 (Neotropical catalogue); Brown, 1978c: 552, 565, 569, 571 (diagnosis & review of genus, Malesian species, Austral, Palaearctic, Afrotropical & Malagasy species, Neotropical species keys); Taylor & Brown, D.R. 1985: 20 (Australia catalogue); Taylor, 1987a: 7 (Australia checklist); Terayama, 1989a: 26 (Taiwan species key); Brandão, 1991: 324 (Neotropical catalogue); Wang, M. 1993a: 226 (China species key); De Andrade, 1994: 24 (†Dominican Amber species key); Bolton, 1995a: 1048 (census); Bolton, 1995b: 63 (catalogue); Gronenberg & Ehmer, 1996: 183 (mandible morphology); Collingwood & Agosti, 1996: 309 (Saudi Arabia species key); Shattuck, 1999: 181 (Australia synopsis); Zhou, 2001b: 29 (China, Guangxi species key); Kugler, J. & Ionescu, 2007: 298 (Mediterraean & Middle East species key); Fisher & Smith, 2008: 4 (Malagasy species revision, key); González-Campero & Elizalde, 2008: 98 (Argentina & Paraguay species key); Zabala, in Fernández, 2008: 125 (Neotropical species key); Terayama, 2009: 117 (Taiwan species key).

Formicinae Lasiini Acropyga lauta TRUE TRUE FALSE

lauta. Acropyga (Rhizomyrma) lauta Mann, 1919: 365 (w.q.) SOLOMON IS. See also: LaPolla, 2004a: 65.

Amblyoponinae Amblyoponini Amblyopone celata TRUE TRUE FALSE

celata. Stigmatomma (Fulakora) celata Mann, 1919: 279, fig. 2 (w.) SOLOMON IS. Combination in Amblyopone: Wilson, 1958a: 143. See also: Taylor, 1979: 830.

Amblyoponinae Amblyoponini Amblyopone gnoma TRUE TRUE FALSE

gnoma. Amblyopone gnoma Taylor, 1979: 829, figs. 13-16 (w.q.) SOLOMON IS.

Ponerinae Ponerini Anochetus isolatus TRUE TRUE FALSE

isolatus. Anochetus cato subsp. isolatus Mann, 1919: 302, fig. 11 (w.m.) SOLOMON IS. Raised to species: Wilson, 1959a: 508. Senior synonym of rossi, splendens: Brown, 1978c: 557 (see also p. 585).

Formicinae Camponotini Camponotus elysii TRUE TRUE FALSE

elysii. Camponotus (Colobopsis) elysii Mann, 1919: 372 (s.w.) SOLOMON IS.

Formicinae Camponotini Camponotus guppyi TRUE TRUE FALSE

guppyi. Camponotus (Colobopsis) guppyi Mann, 1920: 370, figs. 43, 44 (s.w.) SOLOMON IS.

Formicinae Camponotini Camponotus loa TRUE TRUE FALSE

1 subspecies

loa. Camponotus (Colobopsis) loa Mann, 1919: 373, figs. 45, 46 (s.w.) SOLOMON IS. Current subspecies: nominal plus belli.

Formicinae Camponotini Camponotus loa belli TRUE TRUE FALSE

belli. Camponotus (Colobopsis) loa var. belli Mann, 1919: 375 (s.w.) SOLOMON IS.

Myrmicinae Formicoxenini Cardiocondyla nivalis TRUE TRUE FALSE

nivalis. Cardiocondyla nivalis Mann, 1919: 317 (w.) SOLOMON IS.

Myrmicinae Solenopsidini Carebara viehmeyeri TRUE TRUE FALSE

viehmeyeri. Oligomyrmex viehmeyeri Mann, 1919: 331, fig. 25 (s.w.q.m.) SOLOMON IS. Combination in Carebara: Fernández, 2004a: 235.

Cerapachyinae Cerapachyini Cerapachys pawa TRUE TRUE FALSE

pawa. Cerapachys (Syscia) pusilla subsp. pawa Mann, 1919: 277 (w.) SOLOMON IS. Raised to species: Wilson, 1959b: 51.

Cerapachyinae Cerapachyini Cerapachys terricola TRUE TRUE FALSE

terricola. Cerapachys (Cerapachys) terricola Mann, 1919: 277, fig. 1 (w.m.) SOLOMON IS. Junior synonym of inconspicuus: Wilson, 1959b: 47. Revived from synonymy and senior synonym of tulagi: Brown, 1975: 23.

Myrmicinae Crematogastrini Crematogaster abrupta TRUE TRUE FALSE

abrupta. Crematogaster abrupta Mann, 1919: 320 (w.) SOLOMON IS. Combination in C. (Crematogaster): Bolton, 1995b: 166

Myrmicinae Crematogastrini Crematogaster elysii TRUE TRUE FALSE

elysii. Crematogaster elysii Mann, 1919: 319, fig. 22 (w.) SOLOMON IS. Combination in C. (Crematogaster): Bolton, 1995b: 166

Myrmicinae Crematogastrini Crematogaster foxi TRUE TRUE FALSE

foxi. Crematogaster foxi Mann, 1919: 321, fig. 23 (w.q.) SOLOMON IS. Combination in C. (Crematogaster): Bolton, 1995b: 166

Myrmicinae Crematogastrini Crematogaster nesiotis TRUE TRUE FALSE

nesiotis. Crematogaster nesiotis Mann, 1919: 322, fig. 24 (w.) SOLOMON IS. Combination in C. (Crematogaster): Bolton, 1995b: 166

Myrmicinae Crematogastrini Crematogaster obnigra TRUE TRUE FALSE

obnigra. Crematogaster obnigra Mann, 1919: 323 (w.) SOLOMON IS. Combination in C. (Crematogaster): Bolton, 1995b: 166

Myrmicinae Crematogastrini Crematogaster wheeleri TRUE TRUE FALSE

wheeleri. Crematogaster (Rhachiocrema) wheeleri Mann, 1919: 318, fig. 21 (w.) SOLOMON IS.

Myrmicinae Basicerotini Eurhopalothrix greensladei TRUE TRUE FALSE

greensladei. Eurhopalothrix greensladei Taylor, 1968b: 342, figs. 6, 7 (w.) SOLOMON IS.

Myrmicinae Basicerotini Eurhopalothrix isabellae TRUE TRUE FALSE

isabellae. Rhopalothrix isabellae Mann, 1919: 357, fig. 35 (w.) SOLOMON IS. Combination in Eurhopalothrix: Brown & Kempf, 1960: 225. See also: Taylor, 1968b: 344.

Ectatomminae Ectatommini Gnamptogenys albiclava TRUE TRUE FALSE

albiclava. Wheeleripone albiclava Mann, 1919: 283, fig. 3 (w.) SOLOMON IS. Lattke, 2004: 58 (q.). Combination in Gnamptogenys: Brown, 1958g: 227. See also: Lattke, 2004: 56.

Ectatomminae Ectatommini Gnamptogenys crenaticeps TRUE TRUE FALSE

crenaticeps. Wheeleripone crenaticeps Mann, 1919: 285, fig. 4 (w.) SOLOMON IS. Combination in Gnamptogenys: Brown, 1958g: 228. See also: Lattke, 2004: 62.

Ectatomminae Ectatommini Gnamptogenys lucida TRUE TRUE FALSE

lucida. Wheeleripone lucida Mann, 1919: 285 (w.) SOLOMON IS. Combination in Gnamptogenys: Brown, 1958g: 228. See also: Lattke, 2004: 65.

Ectatomminae Ectatommini Gnamptogenys malaensis TRUE TRUE FALSE

malaensis. Rhopalopone malaensis Mann, 1919: 281 (w.) SOLOMON IS. Lattke, 2004: 176 (q.). Combination in Gnamptogenys: Brown, 1958g: 228.

Ectatomminae Ectatommini Gnamptogenys preciosa TRUE TRUE FALSE

preciosa. Gnamptogenys preciosa Lattke, 2004: 66, fig. 14 (w.q.) SOLOMON IS.

Ectatomminae Ectatommini Gnamptogenys solomonensis TRUE TRUE FALSE

solomonensis. Gnamptogenys solomonensis Lattke, 2004: 69, fig. 15 (w.q.) SOLOMON IS.

Ponerinae Ponerini Leptogenys foreli TRUE TRUE FALSE

foreli. Leptogenys (Leptogenys) foreli Mann, 1919: 297, fig. 9 (not fig. 10 as given in the original description) (w.m.) SOLOMON IS. Senior synonym of walkeri: Wilson, 1958a: 126.

Ponerinae Ponerini Leptogenys oresbia TRUE TRUE FALSE

oresbia. Leptogenys oresbia Wilson, 1958a: 131 (w.) SOLOMON IS.

Ponerinae Ponerini Leptogenys truncata TRUE TRUE FALSE

truncata. Leptogenys (Leptogenys) truncatus Mann, 1919: 296, fig. 10 (not fig. 9 as indicated in the original description) (w.) SOLOMON IS.

Myrmicinae Stenammini Lordomyrma epinotalis TRUE TRUE FALSE

epinotalis. Rogeria epinotalis Mann, 1919: 343 (w.q.) SOLOMON IS. Combination in Lordomyrma: Kugler, C. 1994: 26.

Myrmicinae Myrmecinini Myrmecina modesta TRUE TRUE FALSE

1 subspecies

modesta. Myrmecina modesta Mann, 1919: 335 (w.m.) SOLOMON IS. Current subspecies: nominal plus subarmata.

Myrmicinae Myrmecinini Myrmecina modesta subarmata TRUE TRUE FALSE

subarmata. Myrmecina modesta subsp. subarmata Mann, 1919: 337 (w.) SOLOMON IS.

Formicinae Plagiolepidini Nylanderia dichroa TRUE TRUE FALSE

dichroa. Nylanderia dichroa Wheeler, W.M. 1934a: 181 (w.) SOLOMON IS. Unnecessary replacement name: consuta Wheeler, 1935c: 72.

Formicinae Plagiolepidini Nylanderia manni TRUE TRUE FALSE

manni. Paratrechina (Euprenolepis) manni Donisthorpe, 1941b: 41 (w.) SOLOMON IS. Combination in Nylanderia: LaPolla, Brady & Shattuck, 2010a: 127.

Formicinae Plagiolepidini Nylanderia stigmatica TRUE TRUE FALSE

stigmatica. Prenolepis (Nylanderia) stigmaticus Mann, 1919: 367, fig. 41 (w.) SOLOMON IS. Combination in Paratrechina (Nylanderia): Emery, 1925b: 221; in Euprenolepis: Donisthorpe, 1941c: 42; in Paratrechina: LaPolla, 2009: 5; in Nylanderia: LaPolla, Brady & Shattuck, 2010a: 127.

Formicinae Camponotini Opisthopsis manni TRUE TRUE FALSE

1 subspecies

manni. Opisthopsis manni Wheeler, W.M. 1918d: 361, pl. 3, figs. 25, 26 (w.q.) SOLOMON IS. Current subspecies: nominal plus aruanus.

Ponerinae Ponerini Pachycondyla aequalis TRUE TRUE FALSE

aequalis. Ectomomyrmex exarata subsp. aequalis Mann, 1919: 289, fig. 7 (w.) SOLOMON IS. Combination in Pachycondyla: Brown, in Bolton, 1995b: 302. Raised to species: Wilson, 1958d: 364.

Ponerinae Ponerini Pachycondyla manni TRUE TRUE FALSE

manni. Euponera (Mesoponera) melanaria subsp. manni Viehmeyer, 1924a: 228 (w.) SOLOMON IS. Combination in Mesoponera: Wilson, 1958d: 349; in Pachycondyla: Brown, in Bolton, 1995b: 307. Raised to species and senior synonym of robiginosa: Wilson, 1958d: 349.

Ponerinae Ponerini Pachycondyla sheldoni TRUE TRUE FALSE

sheldoni. Euponera (Trachymesopus) sheldoni Mann, 1919: 292 (w.) SOLOMON IS. Combination in Trachymesopus: Wilson, 1958d: 354; in Pachycondyla: Brown, in Bolton, 1995b: 309.

Myrmicinae Pheidolini Pheidole belli TRUE TRUE FALSE

belli. Pheidole (Pheidolacanthinus) belli Mann, 1919: 306, fig. 14 (w.) SOLOMON IS.

Myrmicinae Pheidolini Pheidole erato TRUE TRUE FALSE

erato. Pheidole (Pheidolacanthinus) erato Mann, 1919: 307 (s.w.q.) SOLOMON IS.

Myrmicinae Pheidolini Pheidole isis TRUE TRUE FALSE

1 subspecies

isis. Pheidole isis Mann, 1919: 311, figs. 16-18 (s.w.q.) SOLOMON IS. Current subspecies: nominal plus taki.

Myrmicinae Pheidolini Pheidole mendanai TRUE TRUE FALSE

mendanai. Pheidole mendanai Mann, 1919: 309, fig. 15 (s.w.q.) SOLOMON IS.

Myrmicinae Pheidolini Pheidole nindi TRUE TRUE FALSE

nindi. Pheidole nindi Mann, 1919: 314, figs. 19, 20 (s.w.q.) SOLOMON IS.

Myrmicinae Pheidolini Pheidole philemon TRUE TRUE FALSE

philemon. Pheidole philemon Forel, 1910b: 44 (s.w.m.) SOLOMON IS.

Myrmicinae Pheidolini Pheidole isis taki TRUE TRUE FALSE

taki. Pheidole isis var. taki Mann, 1919: 314 (s.) SOLOMON IS.

Myrmicinae Formicoxenini Podomyrma basalis woodfordi TRUE TRUE FALSE

woodfordi. Podomyrma basalis subsp. woodfordi Mann, 1919: 334, fig. 26 (w.q.) SOLOMON IS.

Myrmicinae Formicoxenini Podomyrma basalis salomo TRUE TRUE FALSE

salomo. Podomyrma basalis subsp. salomo Mann, 1919: 333 (w.) SOLOMON IS.

Formicinae Camponotini Polyrhachis annae TRUE TRUE FALSE

annae. Polyrhachis (Hedomyrma) annae Mann, 1919: 377, fig. 48 (w.) SOLOMON IS. Wheeler, W.M. 1934a: 180 (q.).

Formicinae Camponotini Polyrhachis campbelli TRUE TRUE FALSE

campbelli. Polyrhachis (Hedomyrma) santschii subsp. campbelli Mann, 1919: 376 (w.) SOLOMON IS. Raised to species: Kohout, 1998: 513.

Formicinae Camponotini Polyrhachis emeryana TRUE TRUE FALSE

emeryana. Polyrhachis (Cyrtomyrma) emeryana Mann, 1919: 390, fig. 59 (w.) SOLOMON IS. See also: Kohout, 2006b: 139.

Formicinae Camponotini Polyrhachis fulakora TRUE TRUE FALSE

fulakora. Polyrhachis (Cyrtomyrma) rastellata var. fulakora Mann, 1919: 389 (w.) SOLOMON IS. Raised to species: Kohout, 2006b: 139.

Formicinae Camponotini Polyrhachis geminata TRUE TRUE FALSE

geminata. Polyrhachis (Hedomyrma) geminata Mann, 1919: 376, fig. 47 (w.) SOLOMON IS.

Formicinae Camponotini Polyrhachis greensladei TRUE TRUE FALSE

greensladei. Polyrhachis greensladei Kohout, 1990: 503 (w.q.) SOLOMON IS.

Formicinae Camponotini Polyrhachis johnsoni TRUE TRUE FALSE

johnsoni. Polyrhachis (Cyrtomyrma) rastellata var. johnsoni Mann, 1919: 390 (w.) SOLOMON IS. Subspecies of debilis: Donisthorpe, 1938c: 266. Raised to species: Kohout, 2006b: 140.

Formicinae Camponotini Polyrhachis kaipi TRUE TRUE FALSE

kaipi. Polyrhachis (Chariomyrma) kaipi Mann, 1919: 382, fig. 52 (not 62 as given in text) (w.) SOLOMON IS .

Formicinae Camponotini Polyrhachis malaensis TRUE TRUE FALSE

malaensis. Polyrhachis (Dolichorhachis) malaensis Mann, 1919: 386, fig. 57 (w.q.) SOLOMON IS. Combination in P. (Hedomyrma): Dorow, 1995: 28.

Formicinae Camponotini Polyrhachis nofra TRUE TRUE FALSE

nofra. Polyrhachis nofra Bolton, 1975c: 9, fig. 8 (w.) SOLOMON IS.

Formicinae Camponotini Polyrhachis obliqua TRUE TRUE FALSE

obliqua. Polyrhachis labella var. obliqua Stitz, 1911a: 375, fig. 21 (w.) SOLOMON IS. Combination in P. (Myrma): Emery, 1925b: 201. Raised to species and material of the unavailable name aloseana referred here: Kohout, 1998: 523.

Formicinae Camponotini Polyrhachis osae TRUE TRUE FALSE

osae. Polyrhachis (Myrmatopa) osae Mann, 1919: 384, figs. 54-56 (w.q.) SOLOMON IS.

Formicinae Camponotini Polyrhachis pacifica TRUE TRUE FALSE

pacifica. Polyrhachis pacifica Kohout, 2006b: 140, figs. 12C-D (w.) SOLOMON IS.

Formicinae Camponotini Polyrhachis rere TRUE TRUE FALSE

rere. Polyrhachis (Chariomyrma) rere Mann, 1919: 381, fig. 51 (w.) SOLOMON IS.

Formicinae Camponotini Polyrhachis salomo TRUE TRUE FALSE

1 subspecies

salomo. Polyrhachis salomo Forel, 1910b: 87 (w.) SOLOMON IS. Combination in P. (Myrma): Mann, 1919: 379. Current subspecies: nominal plus hiram.

Formicinae Camponotini Polyrhachis santschii TRUE TRUE FALSE

santschii. Polyrhachis (Hedomyrma) santschi Mann, 1919: 375 (w.) SOLOMON IS.

Formicinae Camponotini Polyrhachis setosa TRUE TRUE FALSE

setosa. Polyrhachis setosa Kohout, 2006b: 141, figs. 12E-F (w.q.) SOLOMON IS.

Formicinae Camponotini Polyrhachis ugiensis TRUE TRUE FALSE

ugiensis. Polyrhachis (Cyrtomyrma) rastellata subsp. ugiensis Mann, 1919: 389 (w.) SOLOMON IS. Raised to species: Donisthorpe, 1938c: 260. See also: Kohout, 2006b: 142.

Formicinae Camponotini Polyrhachis ulysses TRUE TRUE FALSE

ulysses. Polyrhachis ulysses Forel, 1910b: 91 (w.q.m.) SOLOMON IS. Mann, 1919: 383 (q.). Combination in P. (Myrmatopa): Mann, 1919: 383.

Formicinae Camponotini Polyrhachis undulata TRUE TRUE FALSE

undulata. Polyrhachis undulata Kohout, 2006b: 142, figs. 12G-H (w.) SOLOMON IS.

Formicinae Camponotini Polyrhachis wheeleri TRUE TRUE FALSE

wheeleri. Polyrhachis (Myrmhopla) wheeleri Mann, 1919: 387, fig. 58 (w.) SOLOMON IS. Donisthorpe, 1941h: 63 (q.); Wheeler, G.C. & Wheeler, J. 1970: 649 (l.).

Formicinae Camponotini Polyrhachis labella brunneipes TRUE TRUE FALSE

brunneipes. Polyrhachis (Myrma) labella var. brunneipes Wheeler, W.M. 1934a: 179 (w.) SOLOMON IS.

Formicinae Camponotini Polyrhachis salomo hiram TRUE TRUE FALSE

hiram. Polyrhachis salomo subsp. hiram Forel, 1912a: 80 (w.q.) SOLOMON IS. Combination in P. (Myrma): Emery, 1925b: 202.

Myrmicinae Myrmecinini Pristomyrmex obesus TRUE TRUE FALSE

obesus. Pristomyrmex obesus Mann, 1919: 339 (w.q.m.) SOLOMON IS. Senior synonym of melanoticus, pegasus: Wang, M. 2003: 509.

Proceratiinae Probolomyrmecini Probolomyrmex salomonis TRUE TRUE FALSE

salomonis. Probolomyrmex salomonis Taylor, 1965d: 358, figs. 20, 21 (w.) SOLOMON IS.

Ectatomminae Ectatommini Rhytidoponera araneoides TRUE TRUE FALSE

araneoides. Ponera araneoides Le Guillou, 1842: 317 (w.) SOLOMON IS. Combination in Ectatomma: Mayr, 1876: 92; in Rhytidoponera: Emery, 1897d: 548. Senior synonym of rugosa: Mayr, 1876: 92; Donisthorpe, 1932c: 454; of ceranensis, froggatti, impressinodis: Wilson, in Brown, 1958g: 202; Wilson, 1958d: 307; of arcuata: Brown, 1958g: 202.

Myrmicinae Stenammini Rogeria megastigmatica TRUE TRUE FALSE

megastigmatica. Rogeria megastigmatica Kugler, C. 1994: 35, fig. 13 (w.) SOLOMON IS.

Myrmicinae Formicoxenini Romblonella elysii TRUE TRUE FALSE

elysii. Tetramorium scrobiferum var. elysii Mann, 1919: 346, fig. 29 (w.) SOLOMON IS. Combination in Romblonella: Smith, M.R. 1956b: 18. Raised to species: Smith, M.R. 1956b: 18.

Myrmicinae Solenopsidini Solenopsis pawaensis TRUE TRUE FALSE

pawaensis. Solenopsis pawaensis Mann, 1919: 329 (w.) SOLOMON IS.

Myrmicinae Formicoxenini Stereomyrmex dispar TRUE TRUE FALSE

dispar. Willowsiella dispar Wheeler, W.M. 1934a: 175, fig. 1 (w.) SOLOMON IS. Combination in Stereomyrmex: Bolton, 2003: 273.

Myrmicinae Dacetini Strumigenys decollata TRUE TRUE FALSE

decollata. Strumigenys decollata Mann, 1919: 353, figs. 32, 33 (w.m.) SOLOMON IS. See also: Brown, 1958i: 109; Bolton, 2000: 772.

Myrmicinae Tetramoriini Tetramorium antennatum TRUE TRUE FALSE

antennatum. Triglyphothrix fulviceps subsp. antennatus Mann, 1919: 350, fig. 30 (w.) SOLOMON IS. Combination in Tetramorium: Bolton, 1985: 247. Raised to species: Bolton, 1976: 344.

Myrmicinae Tetramoriini Tetramorium mayri TRUE TRUE FALSE

mayri. Triglyphothrix mayri Mann, 1919: 351 (w.) SOLOMON IS. Combination in Tetramorium: Bolton, 1985: 247. See also: Bolton, 1976: 352.

Myrmicinae Tetramoriini Tetramorium melanogyna TRUE TRUE FALSE

melanogyna. Tetramorium melanogyna Mann, 1919: 345, fig. 28 (w.q.) SOLOMON IS. Junior synonym of insolens: Bolton, 1977: 99. Revived from synonymy: Bolton, 1979: 173.

Myrmicinae Tetramoriini Tetramorium salomo TRUE TRUE FALSE

salomo. Tetramorium salomo Mann, 1919: 344 (w.) SOLOMON IS. See also: Bolton, 1977: 109.

Myrmicinae Tetramoriini Tetramorium vombis TRUE TRUE FALSE

vombis. Triglyphothrix vombis Bolton, 1976: 358, fig. 73 (w.q.) SOLOMON IS. Mann, 1919: 349 (m.). Combination in Tetramorium: Bolton, 1985: 247.

Dolichoderinae Leptomyrmecini Turneria pacifica TRUE TRUE FALSE

pacifica. Turneria pacifica Mann, 1919: 361, figs. 39, 40 (w.) SOLOMON IS. Wilson, 1962c: 17 (q.); Shattuck, 1990: 112 (m.).

Myrmicinae Stenammini Vollenhovia dentata TRUE TRUE FALSE

1 subspecies

dentata. Vollenhovia dentata Mann, 1919: 325 (w.m.) SOLOMON IS. Current subspecies: nominal plus marginata.

Myrmicinae Stenammini Vollenhovia elysii TRUE TRUE FALSE

elysii. Vollenhovia elysii Mann, 1919: 327 (w.) SOLOMON IS.

Myrmicinae Stenammini Vollenhovia foveaceps TRUE TRUE FALSE

foveaceps. Vollenhovia foveaceps Mann, 1919: 328 (w.q.) SOLOMON IS.

Myrmicinae Stenammini Vollenhovia loboii TRUE TRUE FALSE

loboii. Vollenhovia loboii Mann, 1919: 324 (w.) SOLOMON IS.

Myrmicinae Stenammini Vollenhovia dentata marginata TRUE TRUE FALSE

marginata. Vollenhovia dentata subsp. marginata Mann, 1919: 327 (w.) SOLOMON IS.

Myrmicinae Solenopsidini Carebara tahitiensis TRUE TRUE FALSE

tahitiensis. Oligomyrmex tahitiensis Wheeler, W.M. 1936f: 11 (q.m.) TAHITIAN IS. Perrault, 1976: 304 (s.w.). Combination in Carebara: Fernández, 2004a: 235.

Myrmicinae Pheidolini Pheidole fervens TRUE TRUE FALSE

1 subspecies

fervens. Pheidole fervens Smith, F. 1858b: 176 (s.) SINGAPORE. Ogata, 1982: 197 (m.). Senior synonym of cavannae, javana, nigriscapa and material of the unavailable name tahitiana referred here: Wilson & Taylor, 1967: 45; senior synonym of desucta: Eguchi, 2001b: 53; of amia, azumai, dharmsalana, dolenda, pungens, soror: Eguchi, 2004: 197. Current subspecies: nominal plus pectinata.

Myrmicinae Dacetini Pyramica insula TRUE TRUE FALSE

insula. Pyramica insula Bolton, 2000: 400 (w.) FRENCH POLYNESIA (Tahiti).

Myrmicinae Solenopsidini Carebara tahitiensis TRUE TRUE FALSE

tahitiensis. Oligomyrmex tahitiensis Wheeler, W.M. 1936f: 11 (q.m.) TAHITIAN IS. Perrault, 1976: 304 (s.w.). Combination in Carebara: Fernández, 2004a: 235.

Myrmicinae Dacetini Strumigenys TRUE TRUE FALSE

Extant: 487 species

Fossil:

Strumigenys Smith, F. 1860c: 72. Type-species: Strumigenys mandibularis, by monotypy.

Taxonomic history

Strumigenys in Myrmicidae: Smith, F. 1871a: 334; Cresson, 1887: 262.

Strumigenys in Myrmicidae, Cryptoceridae: Emery, 1877a: 81.

Strumigenys in Myrmicinae: Mayr, 1865: 26 [Myrmicidae]; Emery & Forel, 1879a: 464 [Myrmicidae]; Dalla Torre, 1893: 145.

Strumigenys in Cryptoceridae, Dacetini: Ashmead, 1905b: 384.

Strumigenys in Myrmicinae, Dacetini: Forel, 1892d: 344; Forel, 1893a: 164; Forel, 1895b: 133; Emery, 1895j: 770; Forel, 1899c: 42; Wheeler, W.M. 1910g: 141; Emery, 1914a: 42; Forel, 1917: 246; Arnold, 1917: 372; Wheeler, W.M. 1922a: 668; Emery, 1924d: 319; Brown, 1948e: 108; all subsequent authors.

Junior synonyms of STRUMIGENYS

Labidogenys Roger, 1862a: 249. Type-species: Labidogenys lyroessa, by monotypy.

Taxonomic history

Labidogenys in Myrmicinae, Dacetini: Donisthorpe, 1943f: 654; Brown, 1948e: 102.

Labidogenys as genus: Roger, 1862a: 249; Brown, 1948e: 102.

Labidogenys as junior synonym of Strumigenys: Roger, 1863b: 40; Emery & Forel, 1879a: 464; Dalla Torre, 1893: 145; Bingham, 1903: 147; Emery, 1924d: 320; Brown & Wilson, 1959b: 281; Brown, 1960b: 38; Bolton, 1983: 358; Bolton, 1999: 1673; Bolton, 2000: 492.

Proscopomyrmex Patrizi, 1946: 294. Type-species: Proscopomyrmex londianensis, by monotypy.

Taxonomic history

Proscopomyrmex in Myrmicinae, Dacetini: Patrizi, 1946: 294 [Dacetonini].

Proscopomyrmex as junior synonym of Strumigenys: Brown, 1949d: 15; Bolton, 1999: 1673; Bolton, 2000: 492.

Eneria Donisthorpe, 1948d: 598. Type-species: Eneria excisa (junior synonym of Strumigenys loriae), by original designation.

Taxonomic history

Eneria as genus: Chapman & Capco, 1951: 104 (anachronism).

Eneria as junior synonym of Strumigenys: Brown, 1949d: 15; Bolton, 1999: 1673; Bolton, 2000: 492.

Quadristruma Brown, 1949b: 47. Type-species: Epitritus emmae, by original designation.

Taxonomic history

Quadristruma in Myrmicinae, Dacetini: Brown, 1949b: 47; all subsequent authors.

Quadristruma as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 12; Bolton, 1999: 1673; Bolton, 2000: 492.

Genus Strumigenys references

Roger, 1863b: 40 (catalogue); Mayr, 1863: 424, 455 (catalogue); Mayr, 1865: 26 (diagnosis); Forel, 1886a: 216 (all species key); Mayr, 1887: 568 (all species key); Dalla Torre, 1893: 145 (catalogue); Emery, 1895c: 325 (North America species key); Emery, 1897c: 574 (New Guinea species key); Forel, 1903a: 707 (India & Sri Lanka species key); Bingham, 1903: 147 (India, Sri Lanka & Burma species key); Wheeler, W.M. 1908a: 146 (New World species key); Santschi, 1913b: 257 (Afrotropical species key); Arnold, 1917: 373 (diagnosis, South Africa species key); Mann, 1921: 467 (Fiji Is species key); Wheeler, W.M. 1922a: 917, 1034 (Afrotropical, Malagasy catalogues); Emery, 1924d: 319 (diagnosis, catalogue); Brown, 1948e: 108 (diagnosis, review of genus); Brown, 1949d: 21 (Japan, China & Taiwan species key); Creighton, 1950a: 301 (North America species key); Chapman & Capco, 1951: 104, 107 (Asia Eneria, Strumigenys (Strumigenys) checklists); Brown, 1954k: 10 (Afrotropical species revision, key); Brown, 1958h: 38 (New Zealand species); Brown, 1962c: 238 (Neotropical species synopsis, key); Kempf, 1972a: 226, 243 (Neotropical Quadristruma, Strumigenys catalogues); Alayo, 1974: 19 (Cuba species key); Kempf, 1976b: 43 (additions to Brown, 1962c key); Smith, D.R. 1979: 1403, 1409 (North America Strumigenys, Quadristruma catalogues); Bolton, 1983: 358, 400 (Afrotropical Strumigenys, Quadristruma species revisions, keys); Taylor & Brown, D.R. 1985: 85, 87 (Australia Quadristruma, Strumigenys catalogues); Taylor, 1987a: 66, 76 (Australia, New Caledonia & New Zealand checklists); Terayama & Kubota, 1989: 780 (Taiwan species key); Brandão, 1991: 380 (Neotropical catalogue); Arakelian & Dlussky, 1991: 150 (former U.S.S.R. species key); Morisita, Kubota, et al. 1992: 63 (Japan species key); Dlussky, 1993: 53 (Fiji Is, Tonga and Samoa species key); Bolton, 1994: 105 (synoptic classification); Bolton, 1995a: 1053 (census); Bolton, 1995b: 377, 395 (catalogue); Gronenberg, 1996: 2021 (mandible morphology); Lin & Wu, 1996: 139 (Taiwan species key); Lattke & Goitía, 1997: 370 (Venezuela species key); Shattuck, 1999: 160, 167 (Australia synopsis); Bolton, 1999: 1657, 1673 (morphology, diagnosis, review of genus, phylogeny); Bolton, 2000: 496, 583, 716, 942 (Neotropical-Nearctic, Afrotropical, Malesian-Oriental-East Palaearctic, Austral species revisions, keys); Fisher, 2000: 614 (Malagasy species revision, key); Zhou, 2001b: 78 (China, Guangxi species key); Lin & Wu, 2001: 166 (Taiwan species key); Zhou & Xu, 2003: 737 (China species key); Yoshimura & Onoyama, 2007: 676 (Japan species key); Terayama, 2009: 142 (Taiwan species key); Heterick, 2009: 176 (south-western Australia species key).

Formicinae Camponotini Camponotus conicus TRUE TRUE FALSE

conicus. Colobopsis conica Mayr, 1876: 67 (w.) TONGA IS. Combination in Camponotus: Dalla Torre, 1893: 226; in C. (Myrmotemnus): Emery, 1920b: 258; in C. (Colobopsis): Forel, 1914a: 272; Emery, 1925b: 148.

Formicinae Camponotini Camponotus nigrifrons TRUE TRUE FALSE

nigrifrons. Colobopsis nigrifrons Mayr, 1870b: 943 (q.) TONGA. Combination in Camponotus (Colobopsis): Dalla Torre, 1893: 244; in C. (Myrmamblys): Forel, 1914a: 272; in C. (Colobopsis): Emery, 1925b: 146.

Myrmicinae Cataulacini Cataulacus intrudens TRUE TRUE FALSE

intrudens. Meranoplus intrudens Smith, F. 1876d: 609, pl. 11, fig. 7 (w.q.m.) SOUTH AFRICA. Combination in Cataulacus: Mayr, 1886c: 364. Senior synonym of batonga, baumi, densipunctatus, foveosquamosus, gazanus, hararicus, intermedius, johannae, pseudotrema, rugosus, subrugosus, umbilicatus and material of the unavailable names krugeri, tangana referred here: Bolton, 1974a: 43; of bulawayensis: Bolton, 1995b: 138.

Myrmicinae Pheidolini Pheidole umbonata TRUE TRUE FALSE

1 subspecies

umbonata. Pheidole umbonata Mayr, 1870b: 978 (s.w.) TONGA. Mann, 1921: 431 (q.). Senior synonym of zimmermani: Wilson & Taylor, 1967: 50. Current subspecies: nominal plus fusciventris.

Myrmicinae Dacetini Strumigenys zakharovi FALSE FALSE FALSE

zakharovi. Strumigenys zakharovi Dlussky, 1993: 60, fig. 2 (w.) TONGA. Unidentified taxon, incertae sedis in Strumigenys: Bolton, 2000: 995.

Myrmicinae Tetramoriini Tetramorium amatongae TRUE TRUE FALSE

amatongae. Tetramorium amatongae Bolton, 1980: 358, figs. 133, 137 (w.) MOZAMBIQUE.

Myrmicinae Tetramoriini Tetramorium pacificum TRUE TRUE FALSE

pacificum. Tetramorium pacificum Mayr, 1870b: 976 (w.q.) TONGA. Imai, Kubota, et al. 1985: 47 (k.). Senior synonym of subscabrum: Bolton, 1977: 102. See also: Schlick-Steiner, Steiner & Zettel, 2006: 182.

Myrmicinae Tetramoriini Tetramorium tonganum TRUE TRUE FALSE

tonganum. Tetramorium tonganum Mayr, 1870b: 976 (w.) TONGA. Santschi, 1928a: 50 (q.). Senior synonym of magitae: Bolton, 1977: 129.

Formicinae Camponotini Camponotus conicus TRUE TRUE FALSE

conicus. Colobopsis conica Mayr, 1876: 67 (w.) TONGA IS. Combination in Camponotus: Dalla Torre, 1893: 226; in C. (Myrmotemnus): Emery, 1920b: 258; in C. (Colobopsis): Forel, 1914a: 272; Emery, 1925b: 148.

Myrmicinae Metaponini Metapone truki TRUE TRUE FALSE

truki. Metapone truki Smith, M.R. 1953d: 135 (w.) TRUK I.

Myrmicinae Pheidolini Pheidole sexspinosa TRUE TRUE FALSE

2 subspecies

sexspinosa. Pheidole sexspinosa Mayr, 1870b: 977 (s.w.) TUVALU. Combination in P. (Pheidolacanthinus): Mann, 1919: 307. Senior synonym of adamsoni: Wilson & Taylor, 1967: 52. Current subspecies: nominal plus biroi, fuscescens.

Formicinae Camponotini Camponotus pallens TRUE TRUE FALSE

pallens. Formica pallens Le Guillou, 1842: 316 (q.) VAVAO I. Combination in Camponotus: Emery, 1921a: 25; in C. (Tanaemyrmex): Emery, 1925b: 95.