subfamily tribe genus species species author date country valid available current valid name original combination fossil taxonomic history Ponerinae Ponerini Anochetus splendidulus TRUE TRUE FALSE

splendidulus. Anochetus splendidulus Yasumatsu, 1940b: 313, fig. 2 (w.q.) CAROLINA IS.

Dolichoderinae TRUE TRUE FALSE

Extant: 28 genera, 692 species, 130 subspecies

Fossil: 16 genera, 103 species

Dolichoderidae Forel, 1878: 364. Type-genus: Dolichoderus.

Taxonomic history

Dolichoderinae as family: Emery, 1894g: 378 [Dolichoderidae]; Ashmead, 1905b: 384 [Dolichoderidae]; Novák & Sadil, 1941: 94 [Dolichoderidae]; Bernard, 1951: 1071 [Dolichoderidae]; Bernard, 1953b: 255 [Dolichoderidae].

Dolichoderinae as tribe of Formicidae: André, 1882a: 127 [Dolichoderidae].

Dolichoderinae as subfamily of Formicidae: Forel, 1878: 364 [Dolichoderidae]; Emery & Forel, 1879a: 454 [Dolichoderidae]; Nasonov, 1889: 26 [Dolichoderidae]; Forel, 1892j: 220 [Dolichoderidae]; Forel, 1893a: 165; Dalla Torre, 1893: 156; Forel, 1895b: 107 [Dolichoderidae]; Emery, 1895j: 771 [subfamily spelled Dolichoderini]; Emery, 1896e: 186; Forel, 1899c: 98; Bingham, 1903: 288; Wheeler, W.M. 1910g: 142; Emery, 1913a: 2; Arnold, 1915: 143; Wheeler, W.M. 1915h: 71; Wheeler, 1915g: 811 [Dolichoderides]; Arnold, 1915: 143; Donisthorpe, 1915d: 178; Forel, 1917: 247; Escherich, 1917: 2 [Dolichoderini]; Bondroit, 1918: 86 [Dolichoderitae]; Wheeler, W.M. 1920: 53; Wheeler, W.M. 1922a: 199; Borgmeier, 1923: 80; Karavaiev, 1936: 163; Smith, M.R. 1951a: 833; Clark, 1951: 16; Brown, 1954e: 29; Kempf, 1972a: 266; Wheeler, G.C. & Wheeler, J. 1972a: 41; Brown, 1973b: 169; all subsequent authors.

Dolichoderinae as formicomorph subfamily of Formicidae: Bolton, 2003: 18, 80.

Dolichoderinae as formicoid subfamily of Formicidae: Brady, Schultz, et al. 2006: 18173; Moreau, Bell, et al. 2006: 102.

Dolichoderinae as formicoid dolichoderomorph subfamily of Formicidae: Ward, 2007a: 556.

Tribes (extant) of Dolichoderinae: Bothriomyrmecini, Dolichoderini, Leptomyrmecini, Tapinomini.

Tribes (extinct) incertae sedis in Dolichoderinae: †Miomyrmecini, †Zherichiniini.

Genera (extinct) incertae sedis in Dolichoderinae: †Alloiomma, †Asymphylomyrmex, †Elaeomyrmex, †Elaphrodites, †Eldermyrmex, †Emplastus, †Eotapinoma, †Eurymyrmex, †Kotshkorkia, †Leptomyrmula, †Miomyrmex, †Petraeomyrmex, †Proiridomyrmex, †Protazteca, †Zherichinius.

Subfamily Dolichoderinae and tribes references, world

Forel, 1878: 364, 380 (diagnosis, genera); Dalla Torre, 1893: 156 (catalogue); Emery, 1895j: 771 (synoptic classification); Emery, 1896e: 186 (genera key); Handlirsch, 1907: 869 (†fossil taxa catalogue); Wheeler, W.M. 1910g: 142 (diagnosis); Emery, 1913a: 2, 6 (diagnosis, tribe key, catalogue); Emery, 1913a: 17 (Tapinomini diagnosis, genera key, catalogue); Arnold, 1915: 144, (diagnosis); Gallardo, 1916a: 3 (diagnosis); Forel, 1917: 247 (synoptic classification); Forel, 1921c: 136 (diagnosis); Wheeler, W.M. 1922a: 199, 687, 688 (diagnosis, tribe key, Tapinomini genera key); Brown & Nutting, 1950: 127 (venation, phylogeny); Brown, 1954e: 29 (phylogeny); Pavan, 1955: 135 (gastral organs); Eisner, 1957: 453 (proventriculus morphology); Bernard, 1967: 246 (diagnosis); Gotwald, 1969: 118 (mouthparts morphology); Wheeler, G.C. & Wheeler, J. 1972a: 41 (diagnosis); Brown, 1973b: 169 (genera & distribution); Wheeler, G.C. & Wheeler, J. 1976b: 61 (larvae, review & synthesis); Snelling, R.R. 1981: 401 (synoptic classification); Dazzini Valcurone & Fanfani, 1985: 1 (gastral glands); Wheeler, G.C. & Wheeler, J. 1985: 258 (synoptic classification); Billen, 1986: 173 (Dufour's gland); Billen, 1987: 278 (abdominal glands); Baroni Urbani & Wilson, 1987: 1 (†fossil Leptomyrmecini); Dlussky & Fedoseeva, 1988: 77 (synoptic classification); Hölldobler & Wilson, 1990: 9 (synoptic classification, genera keys); Baroni Urbani, Bolton & Ward, 1992: 316 (phylogeny); Shattuck, 1992b: 199 (higher classification, phylogeny); Shattuck, 1992c: 20 (revision of subfamily, genera key); Shattuck, 1994: 3 (catalogue); Bolton, 1994: 22 (diagnosis, synoptic classification, genera keys); Shattuck, 1995: 217 (phylogeny, genera); Bolton, 1995a: 1038 (census); Bolton, 1995b: 10 (catalogue); Wenseleers, Schoeters, et al., 1998: 121 (cloacal gland); Brandão, Baroni Urbani, et al. 1999: 411 (phylogeny, genera); Chiotis, Jermiin & Crozier, 2000: 108 (phylogeny, genera); Dlussky & Rasnitsyn, 2002: 416 (diagnosis for impression fossils); Bolton, 2003: 18, 80 (diagnosis, synopsis); Dubovikov, 2005a: 91 (tribes of Dolichoderinae); Brady, Schultz, et al. 2006: 18173 (phylogeny); Moreau, Bell, et al. 2006: 102 (phylogeny); Ward, Brady, et al. 2010: 342 (phylogeny, tribe-rank classification).

Regional and national faunas with keys

André, 1882a: 127 (Europe & Algeria); Nasonov, 1889: 50 (Russia); Forel, 1891b: 9 (Madagascar genera); Forel, 1895e: 460 (India & Sri Lanka); Bingham, 1903: 288 (India, Sri Lanka & Burma); Ruzsky, 1905b: 102 (Russian Empire); Bondroit, 1910: 488 (Belgium); Wheeler, W.M. 1910g: 560 (North America genera); Stitz, 1914: 77 (Central Europe); Gallardo, 1915: 34 (Argentina genera); Forel, 1915d: 40 (Switzerland); Arnold, 1915: 145 (South Africa); Donisthorpe, 1915d: 178 (Britain); Gallardo, 1916a: 12 (Argentina); Emery, 1916b: 208 (Italy); Wheeler, W.M. 1916m: 589 (U.S.A., Connecticut); Bondroit, 1918: 86 (France & Belgium); Gallardo, 1919b: 253 (Argentina genera); Soudek, 1922: 58 (Czechoslovakia); Stärcke, 1926: 117 (Netherlands); Karavaiev, 1927c: 271 (Ukraine); Donisthorpe, 1927b: 199 (Britain); Menozzi & Russo, 1930: 172 (Dominican Republic); Arnol'di, 1933b: 600 (Russia); Menozzi, 1933b: 90 (Israel genera); Karavaiev, 1936: 164 (Ukraine); Smith, M.R. 1937: 861 (Puerto Rico); Stitz, 1939: 209 (Germany); Kratochvíl, 1941: 94 (Central Europe); Novák & Sadil, 1941: 94 (Central Europe); Cole, 1942: 370 (U.S.A., Utah); Smith, M.R. 1943f: 309 (U.S.A. males); Buren, 1944a: 290 (U.S.A., Iowa); Smith, M.R. 1947f: 592 (U.S.A. genera); Creighton, 1950a: 330 (North America); Kusnezov, 1956: 27 (Argentina); Brown, 1958h: 29 (New Zealand); Kusnezov, 1959: 40 (Neotropical genera); Gregg, 1963: 341 (U.S.A., Colorado); Wheeler, G.C. & Wheeler, J. 1963: 149 (U.S.A., North Dakota); Bernard, 1967: 246 (Western Europe); Wilson & Taylor, 1967: 17 (Polynesia); Bolton, 1973a: 329 (West Africa genera); Bolton & Collingwood, 1975: 3 (Britain); Boven, 1970b: 24 (Netherlands); Snelling, R.R. & Hunt, 1976: 90 (Chile); Tarbinsky, 1976: 118 (Kyrghyzstan); Boven, 1977: 123 (Belgium); Kutter, 1977c: 170 (Switzerland); Arnol'di & Dlussky, 1978: 546 (former European U.S.S.R.); Collingwood, 1978: 80 (Iberian Peninsula); Collingwood, 1979: 32 (Fennoscandia & Denmark); Greenslade, 1979: 28 (South Australia genera); Schembri & Collingwood, 1981: 420 (Malta); Allred, 1982: 443 (U.S.A., Utah); Baroni Urbani, 1984: 80 (Neotropical genera); Gösswald, 1985: 314 (Germany); Collingwood, 1985: 242 (Saudi Arabia); Wheeler, G.C. & Wheeler, J. 1986g: 55 (U.S.A., Nevada); Agosti & Collingwood, 1987b: 278 (Balkans); Dlussky, Soyunov & Zabelin, 1990: 168 (Turkmenistan); Kupyanskaya, 1990: 152 (Far Eastern Russia); Morisita, Kubota, Onoyama, et al., 1991: 2 (Japan); Atanasov & Dlussky, 1992: 49 (Bulgaria); Lattke, in Jaffe, 1993: 147 (Neotropical genera); Arakelian, 1994: 70 (Armenia); Wu, J. & Wang, 1995: 115 (China genera); Kupyanskaya, 1995: 331 (Far Eastern Russia); Collingwood & Agosti, 1996: 359 (Saudi Arabia); Seifert, 1996b: 161 (Central Europe); Skinner & Allen, 1996: 40 (Britain); Collingwood & Prince, 1998: 20 (Portugal); Kim & Kim, 1999: 18 (Korea); Shattuck, 1999: 26, 64 (Australia genera, synopsis); Andersen, 2000: 59 (northern Australia genera); Zhou, 2001b: 149 (China, Guangxi); Czechowski, Radchenko & Czechowska, 2002: 134 (Poland); Aktaç & Radchenko, 2002: 54 (Turkey genera); Yoshimura & Onoyama, 2002: 425 (Japan genera, males); Mackay & Mackay, 2002: 221 (U.S.A., New Mexico); Palacio & Fernández, in Fernández, 2003d: 239 (Neotropical genera); Radchenko, 2005b: 186 (North Korea); Coovert, 2005: 102 (U.S.A., Ohio); Clouse, 2007b: 190 (Micronesia); Seifert, 2007: 147 (North and Central Europe); Fisher, 2009: 51 (Malagasy genera); Terayama, 2009: 196 (Taiwan); Heterick, 2009: 28 (south-western Australia genera); Boer, 2010: 14 (Benelux); Yoshimura & Fisher, 2011: 7 (Malagasy males key).

Ecitoninae TRUE TRUE FALSE

Extant: 5 genera, 149 species, 21 subspecies

Fossil: 1 species

Ecitonii Forel, 1893a: 163. Type-genus: Eciton.

Taxonomic history

Ecitoninae as subfamily of Dorylidae: Ashmead, 1905b: 381; Ashmead, 1906: 23.

Ecitoninae as subfamily of Formicidae: Brown, 1973b: 166; Snelling, R.R. 1981: 392; Bolton, 1990c: 1357; Hölldobler & Wilson, 1990: 11; all subsequent authors.

Ecitoninae as dorylomorph subfamily of Formicidae: Bolton, 2003: 34, 142; Brady & Ward, 2005: 593.

Ecitoninae as formicoid subfamily of Formicidae: Moreau, Bell, et al. 2006: 102.

Ecitoninae as formicoid dorylomorph subfamily of Formicidae: Brady, Schultz, et al. 2006: 18173; Ward, 2007a: 555.

Tribes of Ecitoninae: Cheliomyrmecini, Ecitonini.

Subfamily and tribe Ecitonini references

Emery, 1895j: 765 (diagnosis); Ashmead, 1906: 23, 24 (tribes & genera keys); Emery, 1910b: 15 (diagnosis, genera key, catalogue); Wheeler, W.M. 1910g: 138, 558 (diagnosis, North America genera); Forel, 1917: 240 (synoptic classification); Gallardo, 1920: 312 (Argentina genera, key); Wheeler, W.M. 1922a: 634 (genera key); Borgmeier, 1923: 37 (Brazil catalogue); Smith, M.R. 1943f: 290 (U.S.A. males); Creighton, 1950a: 61 (North America); Borgmeier, 1955: 51, 57 (revision of subfamily, tribes key); Borgmeier, 1955: 79 (revision of tribe, genera key); Brown, 1973b: 166 (genera, distribution); Watkins, 1976: 6 (genera keys); Wheeler, G.C. & Wheeler, J. 1976b: 46 (larvae, review & synthesis); Snelling, R.R. 1981: 392 (synoptic classification); Gotwald & Burdette, 1981: 78 (phylogeny); Watkins, 1982: 210 (Mexico genera, key); Gotwald, 1982: 167 (genera key); Baroni Urbani, 1984: 74 (genera key); Wheeler, G.C. & Wheeler, J. 1986g: 17 (U.S.A., Nevada); Bolton, 1990c: 1357 (diagnosis, morphology, phylogeny); Brandão, 1991: 392 (Neotropical fauna, synoptic classification); Baroni Urbani, Bolton & Ward, 1992: 317 (phylogeny); Jaffe, 1993: 13 (Neotropical genera, synoptic classification); Lattke, in Jaffe, 1993: 149 (genera key); Bolton, 1994: 38 (diagnosis, synoptic classification, genera key); Bolton, 1995a: 1039 (census); Bolton, 1995b: 11 (catalogue); Hölldobler, Obermayer & Peeters, 1996: 158 (metatibial gland); Palacio, 1999: 143 (Colombia genera, key); Perfil’eva, 2002: 1239 (venation); Palacio & Fernández, in Fernández, 2003d: 240 (Neotropical genera key); Brady, 2003: 6575 (phylogeny); Bolton, 2003: 34, 142 (diagnosis, synopsis); Brady & Ward, 2005: 593 (phylogeny); Moreau, Bell, et al. 2006: 102 (phylogeny); Brady, Schultz, et al. 2006: 18173 (phylogeny); Ward, 2007a: 555 (classification).

Formicinae TRUE TRUE FALSE

Extant: 49 genera, 2,831 species, 839 subspecies

Fossil: 29 genera, 193 species, 1 subspecies

Formicariae Latreille, 1809: 124. Type-genus: Formica.

Taxonomic history

Formicinae as group name: Latreille, 1809: 124 [Formicariae]; Lepeletier de Saint-Fargeau, 1835: 197 [Formicites]; Nylander, 1846a: 877 [Formicae].

Formicinae as family: Smith, F. 1858b: 1 [Formicidae]; Smith, F. 1861b: 36 [Formicidae]; Smith, 1871a: 302 [Formicidae]; André, 1882a: 125 [Formicidae]; Saunders, 1896: 18 [Formicidae]; Novák & Sadil, 1941: 97 [Formicidae]; Bernard, 1951: 1073 [Formicidae]; Bernard, 1953b: 256 [Formicidae].

Formicinae as tribe of Formicidae: André, 1874: 167 [Formicidae].

Formicinae as subfamily of Formicidae: Mayr, 1855: 286, 299 [Formicidae]; Mayr, 1861: 21 [Formicidae]; Mayr, 1862: 651 [Formicidae]; Mayr, 1865: 6 [Formicidae]; Mayr, 1868c: 24 [Formicidae]; Forel, 1870: 307 [Formicidae]; Forel, 1874: 21 [Formicidae]; Emery, 1877a: 70 [Formicidae]; André, 1881b: 54 [Formicidae]; Ashmead, 1905b: 384; Bondroit, 1918: 17 [Formicitae]; Wheeler, W.M. 1920: 53; Wheeler, W.M. 1922a: 210; Emery, 1925b: 2; Karavaiev, 1936: 172; Clark, 1951: 16; Brown, 1954e: 29; Kempf, 1972a: 266; Brown, 1973b: 169; all subsequent authors.

Formicinae as formicomorph subfamily of Formicidae: Bolton, 2003: 20, 93.

Formicinae as formicoid subfamily of Formicidae: Brady, Schultz, et al. 2006: 18173; Moreau, Bell, et al. 2006: 102.

Formicinae as formicoid formicomorph subfamily of Formicidae: Ward, 2007a: 556.

Tribes of Formicinae: Camponotini, Formicini, Gesomyrmecini, Gigantiopini, Lasiini, Melophorini, Myrmecorhynchini, Myrmoteratini, Notostigmatini, Oecophyllini, Plagiolepidini.

Genera (extinct) incertae sedis in Formicinae: †Camponotites, †Drymomyrmex, †Imhoffia, †Kyromyrma, †Leucotaphus, †Protrechina, †Zhangidris.

Hong (2002) genera (extinct) incertae sedis in Formicinae: †Curtipalpulus, †Eoleptocerites, †Eurytarsites, †Fushuniformica, †Huaxiaformica, †Leptogasteritus, †Liaoformica, †Longiformica, †Magnogasterites, †Orbicapitia, †Ovalicapito, †Ovaligastrula, †Sinoformica, †Sinotenuicapito, †Wilsonia (unresolved junior homonym).

Subfamily Formicinae references, world

Mayr, 1862: 651 (genera key); Mayr, 1865: 6 (diagnosis); Handlirsch, 1907: 859 (†fossil taxa catalogue); Dalla Torre, 1893: 171 (catalogue); Emery, 1895j: 772 (synoptic classification); Emery, 1896e: 187 (genera key); Wheeler, W.M. 1910g: 143 (diagnosis); Forel, 1912i: 88 (tribes key); Forel, 1917: 248 (synoptic classification); Arnold, 1920a: 551 (diagnosis); Forel, 1921c: 139 (diagnosis); Wheeler, W.M. 1922a: 210, 691 (diagnosis, tribes key); Emery, 1925b: 2 (diagnosis, tribe key, catalogue); Brown & Nutting, 1950: 127 (venation, phylogeny); Eisner, 1957: 465 (proventriculus morphology); Hung & Brown, 1966: 198 (gastric apex, structure); Bernard, 1967: 267 (diagnosis); Gotwald, 1969: 120 (mouthparts morphology); Wheeler, G.C. & Wheeler, J. 1972a: 41 (diagnosis); Brown, 1973b: 169 (genera, distribution); Wheeler, G.C. & Wheeler, J. 1976b: 62 (larvae, review & synthesis); Snelling, R.R. 1981: 402 (synoptic classification); Wheeler, G.C. & Wheeler, J. 1985: 258 (synoptic classification); Billen, 1986: 173 (Dufour's gland); Dlussky & Fedoseeva, 1988: 77 (synoptic classification); Hölldobler & Wilson, 1990: 9 (synoptic classification, genera keys); Agosti, 1991: 295 (genus group diagnoses); Shattuck, 1992b: 201 (phylogeny); Baroni Urbani, Bolton & Ward, 1992: 317 (phylogeny); Bolton, 1994: 42 (diagnosis, synoptic classification, genera keys); Bolton, 1995a: 1039 (census); Bolton, 1995b: 11 (catalogue); Wenseleers, Schoeters, et al. 1998: 121 (cloacal gland); Dlussky & Rasnitsyn, 2002: 417 (diagnosis for impression fossils); Bolton, 2003: 20, 93 (diagnosis, synopsis); Brady, Schultz, et al. 2006: 18173 (phylogeny); Moreau, Bell, et al. 2006: 102 (phylogeny).

Regional and national faunas with keys

Mayr, 1855: 299 (Austria); Mayr, 1861: 25 (Europe); Mayr, 1868c: 25 (†Baltic Amber); André, 1874: 167 (Europe); Forel, 1874: 22 (Switzerland); Saunders, E. 1880: 203 (Britain); André, 1882a: 126 (Europe & Algeria); Provancher, 1887: 225 (Canada); Cresson, 1887: 94 (U.S.A. genera); Nasonov, 1889: 50 (Russia); Forel, 1891b: 8 (Madagascar genera); Lameere, 1892: 62 (Belgium); Forel, 1892j: 220 (India & Sri Lanka); Bingham, 1903: 308 (India, Sri Lanka & Burma); Ruzsky, 1905b: 100 (Russian Empire); Wasmann, 1906: 7 (Luxemburg); Bondroit, 1910: 481 (Belgium); Wheeler, W.M. 1910g: 560 (North America genera); Stitz, 1914: 80 (Central Europe); Gallardo, 1915: 35 (Argentina genera); Forel, 1915d: 45 (Switzerland); Donisthorpe, 1915d: 184 (Britain); Emery, 1916b: 216 (Italy); Wheeler, W.M. 1916m: 590 (U.S.A., Connecticut); Bondroit, 1918: 17 (France & Belgium); Arnold, 1920a: 552 (South Africa); Kutter, 1920b: 134 (Switzerland); Soudek, 1922: 61 (Czechoslovakia); Lomnicki, 1925a: 160 (Poland); Stärcke, 1926: 118, 146 (Netherlands); Karavaiev, 1927c: 273 (Ukraine); Donisthorpe, 1927b: 205 (Britain); Menozzi & Russo, 1930: 172 (Dominican Republic); Arnol'di, 1933b: 601 (Russia); Menozzi, 1933b: 90 (Israel genera); Karavaiev, 1936: 173 (Ukraine); Smith, M.R. 1937: 865 (Puerto Rico); Stitz, 1939: 230 (Germany); Kratochvíl, 1941: 97 (Central Europe); Novák & Sadil, 1941: 97 (Central Europe); Cole, 1942: 373 (U.S.A., Utah); Smith, M.R. 1943f: 309 (U.S.A., males); Buren, 1944a: 292 (U.S.A., Iowa); Holgersen, 1943b: 173 (Norway); Holgersen, 1944: 199 (Norway); Smith, M.R. 1947f: 599 (U.S.A. genera); Boven, 1947: 181 (Belgium); Creighton, 1950a: 355 (North America); Kusnezov, 1956: 31 (Argentina); Brown, 1958h: 42 (New Zealand); Boven, 1959: 11 (Netherlands); Gregg, 1963: 447 (U.S.A., Colorado); Wheeler, G.C. & Wheeler, J. 1963: 160 (U.S.A., North Dakota); Collingwood, 1964: 104 (Britain); Bernard, 1967: 268 (Western Europe); Wilson & Taylor, 1967: 17 (Polynesia); Boven, 1970b: 26 (Netherlands); Kempf, 1972a: 266 (Neotropical, synoptic classification); Bolton, 1973a: 329 (West Africa genera); Bolton & Collingwood, 1975: 3 (Britain); Snelling & Hunt, 1976: 104 (Chile); Tarbinsky, 1976: 126 (Kyrghyzstan); Boven, 1977: 126 (Belgium); Kutter, 1977c: 183 (Switzerland); Arnol'di & Dlussky, 1978: 548 (former European U.S.S.R.); Collingwood, 1978: 88 (Iberian Peninsula); Collingwood, 1979: 85 (Fennoscandia & Denmark); Greenslade, 1979: 32 (South Australia genera); Schembri & Collingwood, 1981: 436 (Malta); Prins, 1983: 8 (Southern Africa genera); Allred, 1982: 444 (U.S.A., Utah); Verhaeghe, Deligne, et al., 1984: 106 (Belgium genera); Baroni Urbani, 1984: 81 (Neotropical genera); Gösswald, 1985: 263 (Germany); Collingwood, 1985: 273 (Saudi Arabia); Wheeler, G.C. & Wheeler, J. 1986g: 58 (U.S.A., Nevada); Nilsson & Douwes, 1987: 68 (Norway); Agosti & Collingwood, 1987b: 279 (Balkans); Dlussky, Soyunov & Zabelin, 1990: 124 (Turkmenistan); Kupyanskaya, 1990: 162 (Far Eastern Russia); Morisita, Kubota, Onoyama, et al., 1991: 10 (Japan); Atanasov & Dlussky, 1992: 49 (Bulgaria); Shattuck, 1992b: 199 (higher classification, phylogeny); Lattke, in Jaffe, 1993: 150 (Neotropical genera); Arakelian, 1994: 76 (Armenia); Wu, J. & Wang, 1995: 125 (China genera); Kupyanskaya, 1995: 332 (Far Eastern Russia); Collingwood & Agosti, 1996: 361 (Saudi Arabia); Seifert, 1996b: 166 (Central Europe); Skinner & Allen, 1996: 41 (Britain); Collingwood & Prince, 1998: 21 (Portugal); Shattuck, 1999: 25, 86 (Australia genera, synopsis); Andersen, 2000: 68 (northern Australia genera); Zhou, 2001b: 165 (China, Guangxi); Czechowski, Radchenko & Czechowska, 2002: 147 (Poland); Aktaç & Radchenko, 2002: 54 (Turkey genera); Yoshimura & Onoyama, 2002: 425 (Japan genera, males); Mackay & Mackay, 2002: 236 (U.S.A., New Mexico); Palacio & Fernández, in Fernández, 2003d: 242 (Neotropical genera); Radchenko, 2005b: 187 (North Korea); Coovert, 2005: 113 (U.S.A., Ohio); Clouse, 2007b: 190 (Micronesia); Seifert, 2007: 150 (North and Central Europe); Terayama, 2009: 202 (Taiwan); Heterick, 2009: 30 (south-western Australia genera); Boer, 2010: 17 (Benelux).

Myrmicinae TRUE TRUE FALSE

Extant: 141 genera, 6,087 species, 864 subspecies

Fossil: 31 genera, 130 species

Myrmicites Lepeletier de Saint-Fargeau, 1835: 169. Type-genus: Myrmica.

Taxonomic history

Myrmicinae as group name: Lepeletier de Saint-Fargeau, 1835: 169 [Myrmicites]; Nylander, 1846a: 877 [Myrmicae].

Myrmicinae as family: Smith, F. 1851: 4 [Myrmicidae]; Smith, F. 1861b: 45 [Myrmicidae]; Smith, F. 1871a: 324 [Myrmicidae]; André, 1882a: 125 [Myrmicidae]; Cresson, 1887: 93 [Myrmicidae]; Emery, 1894g: 383 [Myrmicidae]; Saunders, 1896: 18 [Myrmicidae]; Ashmead, 1905b: 383 [Myrmicidae]; Novák & Sadil, 1941: 71 [Myrmicidae]; Bernard, 1951: 1058 [Myrmicidae]; Bernard, 1953b: 222 [Myrmicidae].

Myrmicinae as subfamily of Poneridae: Smith, F. 1858b: 114 [Myrmicidae].

Myrmicinae as tribe of Formicidae: André, 1874: 167 [Myrmicidae].

Myrmicinae as subfamily of Myrmicidae: Ashmead, 1905b: 383.

Myrmicinae as subfamily of Formicidae: Mayr, 1855: 290, 299 [Myrmicidae]; Smith, F. 1857a: 70 [Myrmicidae]; Mayr, 1861: 21 [Myrmicidae]; Smith, F. 1862b: 33 [Myrmicidae]; Mayr, 1862: 738 [Myrmicidae]; Mayr, 1865: 17 [Myrmicidae]; Mayr, 1868c: 24 [Myrmicidae]; Forel, 1870: 307 [Myrmicidae]; Forel, 1874: 22 [Myrmicidae]; Emery, 1877a: 70 [Myrmicidae]; Forel, 1878: 367 [Myrmicidae]; Emery & Forel, 1879a: 456 [Myrmicidae]; André, 1881b: 64 [Myrmicidae]; Nasonov, 1889: 28 [Myrmicidae]; Forel, 1891b: 11 [Myrmicidae]; Forel, 1892j: 220 [Myrmicidae]; Forel, 1893a: 163; Dalla Torre, 1893: 53; Emery, 1895j: 768 [subfamily spelled Myrmicini]; Emery, 1896e: 179; Forel, 1899c: 30; Forel, 1902f: 520; Bingham, 1903: 105; Wheeler, W.M. 1910g: 138; Emery, 1914a: 29; Wheeler, 1915g: 806 [Myrmicides]; Wheeler, W.M. 1915h: 40; Donisthorpe, 1915d: 74; Arnold, 1916: 166; Escherich, 1917: 2 [Myrmicini]; Forel, 1917: 240 [subfamily spelled Myrmicini]; Bondroit, 1918: 14 [Myrmicitae]; Wheeler, W.M. 1920: 53; Wheeler, W.M. 1922a: 124; Emery, 1921f: 3; Karavaiev, 1934: 59; Clark, 1951: 16; Brown, 1954e: 28; Wheeler, G.C. & Wheeler, J. 1972a: 40; Brown, 1973b: 166; all subsequent authors.

Myrmicinae as myrmicomorph subfamily of Formicidae: Bolton, 2003: 52, 182.

Myrmicinae as formicoid subfamily of Formicidae: Moreau, Bell, et al. 2006: 102; Brady, Schultz, et al. 2006: 18173.

Myrmicinae as formicoid myrmicomorph subfamily of Formicidae: Ward, 2007a: 556.

Tribes of Myrmicinae: Adelomyrmecini, Ankylomyrmini, Attini, Basicerotini, Blepharidattini, Cataulacini, Cephalotini, Crematogastrini, Dacetini, Formicoxenini, Lenomyrmecini, Liomyrmecini, Melissotarsini, Meranoplini, Metaponini, Myrmecinini, Myrmicariini, Myrmicini, Paratopulini, Phalacromyrmecini, Pheidolini, Solenopsidini, Stegomyrmecini, Stenammini, Tetramoriini.

Genera (extant) incertae sedis in Myrmicinae: Diaphoromyrma, Tyrannomyrmex.

Genera (extinct) incertae sedis in Myrmicinae: †Afromyrma, †Attopsis, †Cephalomyrmex, †Electromyrmex, †Eocenidris, †Eomyrmex, †Fallomyrma, †Miosolenopsis, †Myrmecites, †Solenopsites.

Hong (2002) genera (extinct) incertae sedis in Myrmicinae: †Brachytarsites, †Clavipetiola, †Fushunomyrmex, †Orbigastrula, †Quadrulicapito, †Quineangulicapito, †Sinomyrmex, †Sphaerogasterites, †Wumyrmex.

Collective group name in Myrmicinae: †Myrmicites.

Subfamily Myrmicinae references, world

Mayr, 1865: 17 (diagnosis); Mayr, 1867a: 91 (diagnosis); Forel, 1878: 367 (diagnosis); Handlirsch, 1907: 872 (†fossil taxa catalogue); Dalla Torre, 1893: 53 (catalogue); Emery, 1895j: 768 (diagnosis); Emery, 1896e: 179 (genera key); Wheeler, W.M. 1910g: 138 (diagnosis); Emery, 1912b: 101 (phylogeny); Emery, 1914a: 34 (phylogeny, tribe key); Arnold, 1916: 164 (diagnosis); Forel, 1917: 240 (synoptic classification); Forel, 1921c: 139 (diagnosis); Emery, 1921f: 3 (diagnosis, tribes & genera key, catalogue); Wheeler, W.M. 1922a: 124, 655 (diagnosis, tribes key); Brown & Nutting, 1950: 126 (venation, phylogeny); Brown, 1954e: 28 (phylogeny); Eisner, 1957: 477 (proventriculus morphology); Bernard, 1967: 93 (diagnosis); Gotwald, 1969: 99 (mouthparts morphology); Wheeler, G.C. & Wheeler, J. 1972a: 40 (diagnosis); Brown, 1973b: 166 (genera, distribution); Wheeler, G.C. & Wheeler, J. 1976b: 52 (larvae, review & synthesis); Kugler, C. 1978a: 413 (sting structure); Kugler, C. 1978b: 267 (pygidial glands); Kugler, C. 1979c: 117 (sting, evolution); Snelling, R.R. 1981: 393 (synoptic classification); Caetano, F.H. 1984: 257 (digestive tract, morphology); Wheeler, G.C. & Wheeler, J. 1985: 257 (synoptic classification); Billen, 1986: 167 (Dufour's gland); Dlussky & Fedoseeva, 1988: 79 (synoptic classification); Hölldobler & Wilson, 1990: 9 onward (synoptic classification, genera keys); Baroni Urbani, Bolton & Ward, 1992: 317 (phylogeny); Bolton, 1994: 75 (diagnosis, synoptic classification, genera keys); Bolton, 1995a: 1040 (census); Bolton, 1995b: 13 (catalogue); Hashimoto, 1996: 354 (phylogenetic position); Baroni Urbani, 2000: 480 (phylogeny); Dlussky & Rasnitsyn, 2002: 422 (diagnosis for wingless fossils); Bolton, 2003: 52, 182 (diagnosis, synopsis); Moreau, Bell, et al. 2006: 102 (phylogeny); Brady, Schultz, et al. 2006: 18173 (phylogeny); Ward, 2007a: 556 (classification).

Regional and national faunas with keys

Mayr, 1855: 391 (Austria); Mayr, 1861: 29 (Europe); Mayr, 1868c: 79 (†Baltic Amber); André, 1874: 171 (Europe); Forel, 1874: 29 (Switzerland); Saunders, E. 1880: 213 (Britain); André, 1882c: 256 (Europe & Algeria); Cresson, 1887: 98 (U.S.A. genera); Provancher, 1887: 243 (Canada); Nasonov, 1889: 54 (Russia); Forel, 1891b: 11 (Madagascar genera); Lameere, 1892: 66 (Belgium); Forel, 1902f: 520 (India & Sri Lanka genera); Bingham, 1903: 105 (India, Sri Lanka & Burma); Ruzsky, 1905b: 103 (Russian Empire); Wasmann, 1906: 13 (Luxemburg); Bondroit, 1910: 490 (Belgium); Wheeler, W.M. 1910g: 558 (North America genera); Stitz, 1914: 55 (Central Europe); Gallardo, 1915: 32 (Argentina genera); Forel, 1915d: 8 (Switzerland); Donisthorpe, 1915d: 74 (Britain); Arnold, 1916: 166, 170 (South Africa tribes, genera); Emery, 1916b: 112 (Italy); Wheeler, W.M. 1916m: 581 (U.S.A., Connecticut); Bondroit, 1918: 90 (France & Belgium); Kutter, 1920b: 144 (Switzerland); Soudek, 1922: 20 (Czechoslovakia); Stärcke, 1926: 84 (Netherlands); Karavaiev, 1927c: 256 (Ukraine); Donisthorpe, 1927b: 77 (Britain); Menozzi & Russo, 1930: 170 (Dominican Republic); Gallardo, 1932b: 91 (Argentina, tribes); Arnol'di, 1933b: 596 (Russia); Menozzi, 1933b: 88 (Israel genera); Karavaiev, 1934: 60 (Ukraine); Smith, M.R. 1937: 829 (Puerto Rico); Stitz, 1939: 63 (Germany); Kratochvíl, 1941: 71 (Central Europe); Novák & Sadil, 1941: 71 (Central Europe); Cole, 1942: 360 (U.S.A., Utah); Smith, M.R. 1943f: 291 (U.S.A., males); Holgersen, 1943b: 166 (Norway); Holgersen, 1944: 198 (Norway); Buren, 1944a: 281 (U.S.A., Iowa); Smith, M.R. 1947f: 543 (U.S.A. genera); Boven, 1947: 170 (Belgium); Creighton, 1950a: 83 (North America); Kusnezov, 1956: 15 (Argentina); Brown, 1958h: 25 (New Zealand); Boven, 1959: 7 (Netherlands); Gregg, 1963: 288 (U.S.A., Colorado); Wheeler, G.C. & Wheeler, J. 1963: 92 (U.S.A., North Dakota); Collingwood, 1964: 94 (Britain); Bernard, 1967: 95 (Western Europe); Wilson & Taylor, 1967: 13 (Polynesia); Boven, 1970b: 9 (Netherlands); Kempf, 1972a: 263 (Neotropical, synoptic classification); Bolton, 1973a: 325 (West Africa genera); Bolton & Collingwood, 1975: 3 (Britain); Snelling, R.R. & Hunt, 1976: 70 (Chile); Tarbinsky, 1976: 19 (Kyrghyzstan); Boven, 1977: 69 (Belgium); Kutter, 1977c: 31 (Switzerland); Arnol'di & Dlussky, 1978: 524 (former European U.S.S.R.); Collingwood, 1978: 75 (Iberian Peninsula); Collingwood, 1979: 36 (Fennoscandia & Denmark); Greenslade, 1979: 20 (South Australia genera); Schembri & Collingwood, 1981: 423 (Malta); Allred, 1982: 438 (U.S.A., Utah); Baroni Urbani, 1984: 76 (Neotropical genera); Verhaeghe, Deligne, et al. 1984: 112 (Belgium genera); Gösswald, 1985: 289 (Germany); Collingwood, 1985: 245 (Saudi Arabia); Wheeler, G.C. & Wheeler, J. 1986g: 20 (U.S.A., Nevada); Nilsson & Douwes, 1987: 57 (Norway); Agosti & Collingwood, 1987b: 265 (Balkans); Dlussky, Soyunov & Zabelin, 1990: 181 (Turkmenistan); Kupyanskaya, 1990: 89 (Far Eastern Russia); Ogata, 1991b: 61 (Japan genera); Morisita, Kubota, et al. 1992: 1 (Japan); Atanasov & Dlussky, 1992: 51 (Bulgaria); Lattke, in Jaffe, 1993: 153 (Neotropical genera); Arakelian, 1994: 15 (Armenia); Wu, J. & Wang, 1995: 57 (China genera); Kupyanskaya, 1995: 327 (Far Eastern Russia); Collingwood & Agosti, 1996: 308 (Saudi Arabia); Seifert, 1996b: 108 (Central Europe); Skinner & Allen, 1996: 43 (Britain); Collingwood & Prince, 1998: 10 (Portugal); Shattuck, 1999: 39, 122 (Australia genera, synopsis); Andersen, 2000: 37 (northern Australia genera); Zhou, 2001b: 69 (China, Guangxi); Czechowski, Radchenko & Czechowska, 2002: 135 (Poland); Aktaç & Radchenko, 2002: 55 (Turkey genera); Yoshimura & Onoyama, 2002: 424 (Japan genera, males); Mackay & Mackay, 2002: 58 (U.S.A., New Mexico); Palacio & Fernández, in Fernández, 2003d: 244 (Neotropical genera and synopsis); Coovert, 2005: 32 (U.S.A., Ohio); Radchenko, 2005b: 184 (North Korea); Clouse, 2007b: 190 (Micronesia); Seifert, 2007: 110 (North and Central Europe); Terayama, 2009: 131 (Taiwan); Heterick, 2009: 36 (south-western Australia genera); Boer, 2010: 46 (Benelux); Eguchi, Bui & Yamane, 2011: 8 (Vietnam genera).

Ponerinae TRUE TRUE FALSE

Extant: 27 genera, 1,033 species, 155 subspecies

Fossil: 9 genera, 68 species, 2 subspecies

Ponerites Lepeletier de Saint-Fargeau, 1835: 185. Type-genus: Ponera.

Taxonomic history

Ponerinae as group name: Lepeletier de Saint-Fargeau, 1835: 185 [Ponérites].

Ponerinae as family: Smith, F. 1851: 6 [Poneridae]; Smith, F. 1858b: 76 [Poneridae]; Smith, F. 1861b: 44 [Poneridae]; Smith, F. 1871a: 320 [Poneridae]; André, 1882a: 125 [Poneridae]; Cresson, 1887: 93 [Poneridae]; Emery, 1894g: 379 [Poneridae]; Saunders, 1896: 18 [Poneridae]; Ashmead, 1905b: 382 [Poneridae]; Novák & Sadil, 1941: 70 [Poneridae]; Bernard, 1951: 1042 [Poneridae]; Bernard, 1953b: 185 [Poneridae].

Ponerinae as tribe of Formicidae: André, 1874: 167 [Poneridae].

Ponerinae as subfamily of Poneridae: Ashmead, 1905b: 382.

Ponerinae as subfamily of Formicidae: Mayr, 1855: 289, 299 [Poneridae]; Mayr, 1861: 21 [Poneridae]; Mayr, 1862: 712 [Poneridae]; Smith, F. 1857a: 64 [Poneridae]; Smith, F. 1862b: 31 [Poneridae]; Mayr, 1865: 11 [Poneridae]; Mayr, 1868c: 24 [Poneridae]; Forel, 1870: 307 [Poneridae]; Forel, 1874: 21 [Poneridae]; Emery, 1877a: 70 [Poneridae]; Forel, 1878: 366 [Poneridae]; Emery & Forel, 1879a: 455 [Poneridae]; André, 1881b: 64 [Poneridae]; Forel, 1892j: 220 [Poneridae]; Dalla Torre, 1893: 13; Forel, 1893a: 162; Forel, 1895b: 111 [Poneridae]; Nasonov, 1889: 27 [Poneridae]; Emery, 1895j: 766 [subfamily spelled Ponerini]; Emery, 1896e: 176; Forel, 1899c: 2; Emery, 1901a: 36; Bingham, 1903: 23; Wheeler, W.M. 1910g: 134; Emery, 1911d: 2; Wheeler, 1915g: 805 [Ponerides]; Donisthorpe, 1915d: 65; Wheeler, W.M. 1915h: 25; Arnold, 1915: 9; Escherich, 1917: 2 [Ponerini]; Forel, 1917: 235; Bondroit, 1918: 78 [Poneritae]; Wheeler, W.M. 1920: 53; Wheeler, W.M. 1922a: 56, 632, 640; Borgmeier, 1923: 37; Karavaiev, 1934: 49; Clark, 1951: 15; Brown, 1954e: 24; Wheeler, G.C. & Wheeler, J. 1972a: 39; Brown, 1973b: 165; all subsequent authors.

Ponerinae as poneromorph subfamily of Formicidae: Bolton, 2003: 42, 156.

Ponerinae as poneroid subfamily of Formicidae: Ouellette, Fisher, et al. 2006: 365; Brady, Schultz, et al. 2006: 18173; Moreau, Bell et al. 2006: 102; Ward, 2007a: 555.

Tribes of Ponerinae: Platythyreini, Ponerini, Thaumatomyrmecini.

Genera (extinct) incertae sedis in Ponerinae: †Afropone, †Eogorgites, †Eoponerites, †Furcisutura, †Longicapitia.

Subfamily Ponerinae references, world

Smith, F. 1858b: 76 (diagnosis); Mayr, 1862: 712 (genera key); Mayr, 1865: 11 (Odontomachidae, Poneridae diagnoses), Mayr, 1867a: 79, 81 (Odontomachidae, Poneridae diagnoses); Forel, 1878: 366 (diagnosis); Dalla Torre, 1893: 13 (world catalogue); Emery, 1895j: 766 (diagnosis); Emery, 1896e: 176 (genera key); Handlirsch, 1907: 879 (†fossil taxa catalogue); Wheeler, W.M. 1910g: 134 (diagnosis); Emery, 1911d: 3 (diagnosis, tribes key); Forel, 1917: 235 (synoptic classification); Forel, 1921c: 133 (diagnosis); Wheeler, W.M. 1922a: 56, 636 (diagnosis, tribes key); Brown & Nutting, 1950: 124 (venation, phylogeny); Brown, 1954e: 24 (phylogeny); Eisner, 1957: 475 (proventriculus morphology); Bernard, 1967: 79 (diagnosis); Gotwald, 1969: 25 (mouthparts morphology); Wheeler, G.C. & Wheeler, J. 1972a: 39 (diagnosis); Brown, 1973b: 165 (genera, distribution); Wheeler, G.C. & Wheeler, J. 1976b: 48 (larvae, review & synthesis); Snelling, R.R. 1981: 387 (synoptic classification); Wheeler, G.C. & Wheeler, J. 1985: 256 (synoptic classification); Billen, 1986: 168 (Dufour's gland); Fanfani & Dazzini Valcurone, 1986: 115 (exocrine glands synopsis); Dlussky & Fedoseeva, 1988: 78 (synoptic classification); Hölldobler & Wilson, 1990: 9 onward (synoptic classification, genera keys); Baroni Urbani, Bolton & Ward, 1992: 317 (phylogeny); Bolton, 1994: 153 (diagnosis, synoptic classification, genera keys); Bolton, 1995a: 1042 (census); Bolton, 1995b: 14 (catalogue); Grimaldi, Agosti & Carpenter, 1997: 20 (†Cretaceous genera); Dlussky & Rasnitsyn, 2002: 422 (diagnosis for wingless fossils); Bolton, 2003: 42, 156 (diagnosis, synopsis); Ouellette, Fisher et al. 2006: 359 (phylogeny); Brady, Schultz, et al. 2006: 18173 (phylogeny); Moreau, Bell et al. 2006: 102 (phylogeny); Ward, 2007a: 555 (classification).

.

Regional and national faunas with keys

Mayr, 1861: 28 (Europe); Mayr, 1868c: 69 (†Baltic Amber); André, 1874: 170 (Europe); Forel, 1874: 29 (Switzerland); André, 1882b: 227 (Europe & Algeria); Cresson, 1887: 97 (U.S.A., genera); Provancher, 1887: 238 (Canada); Nasonov, 1889: 54 (Russia); Forel, 1891b: 9 (Madagascar genera); Forel, 1900c: 52 (India & Sri Lanka); Bingham, 1903: 23 (India, Sri Lanka & Burma); Ruzsky, 1905b: 108 (Russian Empire); Wheeler, W.M. 1910g: 557 (North America genera); Bondroit, 1910: 489 (Belgium); Stitz, 1914: 54 (Central Europe); Gallardo, 1915: 31 (Argentina genera); Forel, 1915d: 6 (Switzerland); Arnold, 1915: 10 (South Africa); Donisthorpe, 1915d: 65 (Britain); Emery, 1916b: 98 (Italy); Wheeler, W.M. 1916m: 580 (U.S.A., Connecticut); Bondroit, 1918: 79 (France & Belgium); Gallardo, 1918b: 4 (Argentina); Soudek, 1922: 18 (Czechoslovakia); Stärcke, 1926: 82 (Netherlands); Donisthorpe, 1927b: 68 (Britain); Menozzi & Russo, 1930: 169 (Dominican Republic); Arnol'di, 1933b: 596 (Russia); Karavaiev, 1934: 50 (Ukraine); Smith, M.R. 1937: 823 (Puerto Rico); Stitz, 1939: 58 (Germany); Smith, M.R. 1943f: 278 (U.S.A. males); Buren, 1944a: 279 (U.S.A., Iowa); Smith, M.R. 1947f: 530 (U.S.A. genera); Creighton, 1950a: 31 (North America); Kusnezov, 1956: 11 (Argentina); Brown, 1958h: 11 (New Zealand); Gregg, 1963: 280 (U.S.A., Colorado); Bernard, 1967: 80 (Western Europe); Wilson & Taylor, 1967: 10 (Polynesia); Kempf, 1972a: 262 (Neotropical synoptic classification); Bolton, 1973a: 323 (West Africa genera); Bolton & Collingwood, 1975: 3 (Britain); Boven, 1977: 66 (Belgium); Kutter, 1977c: 21 (Switzerland); Arnol'di & Dlussky, 1978: 522 (former European U.S.S.R.); Collingwood, 1978: 74 (Iberian Peninsula); Collingwood, 1979: 29 (Fennoscandia & Denmark); Greenslade, 1979: 14 (South Australia genera); Francoeur, 1979: 30 (Canada, Québec); Schembri & Collingwood, 1981: 417 (Malta); Allred, 1982: 438 (U.S.A., Utah); Baroni Urbani, 1984: 75 (Neotropical genera); Collingwood, 1985: 236 (Saudi Arabia); Wheeler, G.C. & Wheeler, J. 1986g: 17 (U.S.A., Nevada); Agosti & Collingwood, 1987b: 264 (Balkans); Ogata, 1987: 101 (Japan genera); Morisita, Kubota, et al. 1989: 8 (Japan); Dlussky, Soyunov & Zabelin, 1990: 173 (Turkmenistan); Kupyanskaya, 1990: 85 (Far Eastern Russia); Atanasov & Dlussky, 1992: 51 (Bulgaria); Lattke, in Jaffe, 1993: 166 (Neotropical genera); Arakelian, 1994: 10 (Armenia); Wu, J. & Wang, 1995: 31 (China genera); Collingwood & Agosti, 1996: 308 (Saudi Arabia); Seifert, 1996b: 107 (Central Europe); Collingwood & Prince, 1998: 10 (Portugal); Kim, Kim & Kim, 1998: 145 (Korea); Shattuck, 1999: 52, 179 (Australia genera synopsis); Andersen, 2000: 19 (northern Australia genera); Zhou, 2001b: 22 (China, Guangxi); Czechowski, Radchenko & Czechowska, 2002: 133 (Poland); Aktaç & Radchenko, 2002: 53 (Turkey genera); Yoshimura & Onoyama, 2002: 436 (Japan genera, males); Mackay & Mackay, 2002: 27 (U.S.A., New Mexico); Csösz, 2003: 147 (Carpathian Basin); Palacio & Fernández, in Fernández, 2003d: 256 (Neotropical genera); Radchenko, 2005b: 184 (North Korea); Coovert, 2005: 23 (U.S.A., Ohio); Yoshimura & Fisher, 2007: 27 (Malagasy genera, males); Clouse, 2007b: 190 (Micronesia); Seifert, 2007: 108 (North and Central Europe); Bolton & Fisher, 2008c: 32 (Afrotropical genera); Fernández & Arias-Penna, 2008: 32 (Neotropical genera); Dlussky, 2009: 1071 (Eocene amber); Terayama, 2009: 101 (Taiwan genera); Heterick, 2009: 34 (south-western Australia genera); Boer, 2010: 68 (Benelux).

Dolichoderinae Dolichoderini Dolichoderus TRUE TRUE FALSE

Extant: 122 species, 21 subspecies

Fossil: 53 species

Dolichoderus Lund, 1831a: 130. Type-species: Formica attelaboides, by monotypy.

Taxonomic history

Dolichoderus in Formicidae: Smith, F. 1858b: 75.

Dolichoderus in Formicinae: Mayr, 1862: 652 (in key) [Formicidae]; Mayr, 1865: 9 [Formicidae].

Dolichoderus in Dolichoderidae: Ashmead, 1905b: 384.

Dolichoderus in Dolichoderinae: Forel, 1878: 386 [Dolichoderidae]; Emery & Forel, 1879a: 455 [Dolichoderidae]; Dalla Torre, 1893: 156; Forel, 1895b: 107 [Dolichoderidae]; Emery, 1895j: 771; Forel, 1895e: 462; Forel, 1899c: 98; Wheeler, W.M. 1910g: 142.

Dolichoderus in Dolichoderinae, Dolichoderini: Emery, 1913a: 7; Wheeler, W.M. 1915h: 77; Forel, 1917: 247; Wheeler, W.M. 1922a: 688; all subsequent authors.

Junior synonyms of DOLICHODERUS

Hypoclinea Mayr, 1855: 377 (attributed to Foerster). Type-species: Formica quadripunctata, by subsequent designation of Wheeler, W.M. 1911f: 165.

Taxonomic history

Hypoclinea in Formicidae: Smith, F. 1858b: 57.

Hypoclinea in Formicinae: Mayr, 1855: 377 [Formicidae]; Mayr, 1861: 40 [Formicidae]; Mayr, 1862: 652 (in key) [Formicidae]; Mayr, 1865: 9 [Formicidae]; Mayr, 1868c: 53 [Formicidae]; André, 1874: 168 [Formicidae].

Hypoclinea in Dolichoderidae: Ashmead, 1905b: 384.

Hypoclinea in Dolichoderinae, Dolichoderini: Emery, 1913a: 18; Forel, 1917: 247; all subsequent authors.

Hypoclinea as genus: Mayr, 1855: 377; Mayr, 1865: 9; Mayr, 1868c: 53; Mayr, 1870b: 953; Smith, 1871a: 310; Ashmead, 1905b: 384; Kempf, 1972a: 118; Lattke, 1987b: 259; Dlussky & Fedoseeva, 1988: 78; Mackay & Vinson, 1989b: 17; Hölldobler & Wilson, 1990: 17; Wu, J. & Wang, 1995: 120.

Hypoclinea as subgenus of Dolichoderus: Emery, 1894c: 234; Wheeler, W.M. 1910g: 142; Emery, 1913a: 10; Wheeler, W.M. 1915h: 77; Forel, 1917: 247; Borgmeier, 1923: 84; Clark, 1930b: 252; McAreavey, 1949: 17; Creighton, 1950a: 331; Chapman & Capco, 1951: 182; Kusnezov, 1956: 28; Kusnezov, 1959: 50; Smith, D.R. 1979: 1415; Kupyanskaya, 1990: 154.

Hypoclinea as junior synonym of Dolichoderus: Forel, 1878: 386; Emery & Forel, 1879a: 455; Dalla Torre, 1893: 156; Bingham, 1903: 291; Brown, 1973b: 181 [provisional]; Shattuck, 1992c: 66; Mackay, 1993b: 6; Shattuck, 1994: 39; Bolton, 1994: 26; Bolton, 1995b: 31; Bolton, 2003: 85.

Monacis Roger, 1862a: 233. Type-species: Formica bispinosa, by subsequent designation of Wheeler, W.M. 1911f: 167.

Taxonomic history

Monacis in Dolichoderidae: Ashmead, 1905b: 384.

Monacis in Dolichoderinae, Dolichoderini: Emery, 1913a: 9; Forel, 1917: 247; all subsequent authors.

Monacis as genus: Roger, 1862a: 233; Ashmead, 1905b: 384; Brown, 1950e: 249; Kempf, 1959b: 227; Kempf, 1972a: 141; Kempf, 1972c: 253; Dlussky & Fedoseeva, 1988: 78; Mackay & Vinson, 1989b: 20; Hölldobler & Wilson, 1990: 17.

Monacis as subgenus of Dolichoderus: Emery, 1894c: 228; Wheeler, W.M. 1910g: 142; Emery, 1913a: 9; Forel, 1917: 247; Wheeler, W.M. 1922a: 688; Borgmeier, 1923: 81; Kusnezov, 1956: 27; Kusnezov, 1959: 50.

Monacis as junior synonym of Hypoclinea: Mayr, 1862: 704.

Monacis as junior synonym of Dolichoderus: Forel, 1878: 386; Emery & Forel, 1879a: 455; Dalla Torre, 1893: 156; Brown, 1973b: 182 [provisional]; Shattuck, 1992c: 66; Mackay, 1993b: 6; Shattuck, 1994: 39; Bolton, 1994: 26; Bolton, 1995b: 35; Bolton, 2003: 85.

Monoceratoclinea Wheeler, W.M. 1935c: 68 [as subgenus of Dolichoderus]. Type-species: Dolichoderus (Hypoclinea) monoceros, by original designation.

Taxonomic history

Monoceratoclinea as subgenus of Dolichoderus: Wheeler, W.M. 1935c: 68.

Monoceratoclinea as genus: Brown, 1950e: 249; Dlussky & Fedoseeva, 1988: 78; Hölldobler & Wilson, 1990: 17.

Monoceratoclinea as junior synonym of Dolichoderus: Shattuck, 1992c: 66; Shattuck, 1994: 40; Bolton, 1994: 26; Bolton, 1995b: 35; Bolton, 2003: 85.

Diceratoclinea Wheeler, W.M. 1935c: 69 [as subgenus of Dolichoderus]. Type-species: Dolichoderus scabridus, by original designation.

Taxonomic history

Diceratoclinea as subgenus of Dolichoderus: Wheeler, W.M. 1935c: 69; Chapman & Capco, 1951: 181.

Diceratoclinea as genus: Brown, 1950e: 249.

Diceratoclinea as junior synonym of Dolichoderus: Brown, 1973b: 180 [provisional]; Taylor & Brown, D.R. 1985: 93; Shattuck, 1992c: 66; Shattuck, 1994: 40; Bolton, 1994: 26; Bolton, 1995b: 26; Bolton, 2003: 85.

Acanthoclinea Wheeler, W.M. 1935c: 69 [as subgenus of Dolichoderus]. Type-species: Dolichoderus doriae, by original designation.

Taxonomic history

Acanthoclinea in Dolichoderinae, Dolichoderini: Donisthorpe, 1943f: 618.

Acanthoclinea as subgenus of Dolichoderus: Wheeler, W.M. 1935c: 69.

Acanthoclinea as genus: Brown, 1950e: 249.

Acanthoclinea as junior synonym of Dolichoderus: Taylor & Brown, D.R. 1985: 93; Shattuck, 1992c: 66; Shattuck, 1994: 40; Bolton, 1994: 26; Bolton, 1995b: 18; Bolton, 2003: 85.

Karawajewella Donisthorpe, 1944b: 59.

Taxonomic history

[Replacement name for Diabolus Karavaiev, 1926d: 424; junior homonym of Diabolus Gray, J.E. 1841: 400 (Mammalia).]

Karawajewella as junior synonym of Dolichoderus: Brown, 1973b: 181 [provisional]; Shattuck, 1992c: 66; Shattuck, 1994: 39, 40; Bolton, 1994: 26; Bolton, 1995b: 32; Bolton, 2003: 85.

Homonym replaced by Karawajewella

Diabolus Karavaiev, 1926d: 424 [as subgenus of Dolichoderus]. Type-species: Dolichoderus (Diabolus) bifurcatus (junior synonym of Dolichoderus cuspidatus), by original designation.

Taxonomic history

[Diabolus Karavaiev junior homonym of Diabolus Gray, J.E. 1841: 400 (Mammalia).]

Diabolus as subgenus of Dolichoderus: Wheeler, W.M. 1935c: 68.

Genus Dolichoderus references

Roger, 1863b: 10, 14, 15 (Dolichoderus, Hypoclinea, Monacis catalogues); Mayr, 1863: 407, 423 (Dolichoderus, Hypoclinea catalogues); Mayr, 1865: 9 (Dolichoderus, Hypoclinea diagnoses); Mayr, 1867a: 75 (Hypoclinea diagnosis); Mayr, 1868c: 54 (†Baltic Amber species key); Mayr, 1870b: 955 (Hypoclinea species key); Mayr, 1876: 79 (Australia species key); André, 1882b: 225 (Europe & Algeria species); Mayr, 1886d: 435 (U.S.A. species key); Cresson, 1887: 258 (U.S.A. catalogue); Dalla Torre, 1893: 156 (catalogue); Forel, 1895e: 462 (India & Sri Lanka species key); Bingham, 1903: 292 (India, Sri Lanka & Burma species key); Wheeler, W.M. 1905d: 306 (North America species key); Emery, 1913a: 7 (diagnosis, subgenera key, catalogue); Emery, 1913a: 9, 10 (D. (Monacis), D. (Hypoclinea), catalogues); Mann, 1916: 460 (Brazil D. (Dolichoderus) species key); Wheeler, W.M. 1922a: 688 (subgenera key); Clark, 1930b: 253 (Australia D. (Hypoclinea) species key); Creighton, 1950a: 333 (North America D. (Hypoclinea) species key); Chapman & Capco, 1951: 181 (Asia checklist); Kempf, 1959b: 230 (Monacis, all species revision, key); Wheeler, J. 1963: 149 (U.S.A., North Dakota species key); Bernard, 1967: 249 (diagnosis); Kempf, 1969: 292 (Neotropical D. (Dolichoderus) species key); Kempf, 1972a: 98, 118, 141 (Neotropical Dolichoderus, Hypoclinea, Monacis, catalogues); Kempf, 1972c: 253 (Monacis species, additions to 1959a key); Smith, D.R. 1979: 1415 (North America catalogue); Taylor & Brown, D.R. 1985: 93 (Australia catalogue); Taylor, 1987a: 24 (Australia, New Caledonia checklist); Harada, 1987: 602 (Monacis species key); Lattke, 1987b: 263 (Neotropical D. bispinosus group, key); Dlussky & Fedoseeva, 1988: 78 (synoptic classification); Johnson, 1989a: 5 (U.S.A. species key); Hölldobler & Wilson, 1990: 17 (synoptic classification); Brandão, 1991: 340, 347, 356 (Neotropical catalogue); Shattuck, 1992c: 66 (diagnosis, review of genus); Mackay, 1993b: 17 (New World, all species revision, key); Radchenko, 1994b: 112 (South Siberia species key); Shattuck, 1994: 39 (catalogue); Bolton, 1995a: 1049 (census); Bolton, 1995b: 172 (catalogue); Wu, J. & Wang, 1995: 120 (China species key); Xu, 1995a: 34 (China species key); Shattuck, 1999: 68 (Australia synopsis); Xu, 2001b: 356 (China species key); Zhou, 2001b: 153 (China, Guangxi species key); Dill, 2002: 21, 30 (southeast Asia species groups key, D. cuspidatus group revision, key); Dlussky & Rasnitsyn, 2002: 416 (diagnosis for impression fossils); Dlussky, 2002b: 54 (Baltic & Rovno amber species); Dlussky, 2008b: 509 (Europe Eocene amber species key); Heterick, 2009: 45 (south-western Australia species key).

Dolichoderinae Leptomyrmecini Dorymyrmex TRUE TRUE FALSE

57 species, 26 subspecies

Dorymyrmex Mayr, 1866a: 494. Type-species: Dorymyrmex flavescens, by monotypy.

Taxonomic history

Dorymyrmex in Dolichoderinae: Forel, 1878: 383 [Dolichoderidae]; Dalla Torre, 1893: 167; Emery, 1895j: 771; Forel, 1899c: 103; Wheeler, W.M. 1910g: 142; all subsequent authors.

Dorymyrmex in Dolichoderidae: Ashmead, 1905b: 384.

Dorymyrmex in Dolichoderinae, Tapinomini: Emery, 1913a: 36; Forel, 1917: 248; Wheeler, W.M. 1922a: 689; Donisthorpe, 1943f: 640; subsequent authors and Jaffe, 1993: 9.

Dorymyrmex in Dolichoderinae, Dolichoderini: Shattuck, 1992c: 77; Bolton, 1994: 26; Bolton, 2003: 86.

Dorymyrmex in Dolichoderinae, Leptomyrmecini: Ward, Brady, et al. 2010: 361.

Junior synonyms of DORYMYRMEX

Psammomyrma Forel, 1912h: 43 [as subgenus of Dorymyrmex]. Type-species: Dorymyrmex planidens, by subsequent designation of Wheeler, W.M. 1913a: 82.

Taxonomic history

Psammomyrma as subgenus of Dorymyrmex: Forel, 1912h: 43; Kempf, 1972a: 100.

Psammomyrma as junior synonym of Dorymyrmex: Forel, 1913b: 350 (in text); Santschi, 1922b: 365; Wheeler, W.M. 1922a: 689; Snelling, R.R. & Hunt, 1976: 93; Shattuck, 1992c: 77; Shattuck, 1994: 73; Bolton, 1994: 26; Bolton, 2003: 86.

Conomyrma Forel, 1913b: 350 (in text) [as subgenus of Dorymyrmex]. Type-species: Prenolepis pyramica, by subsequent designation of Santschi, 1922b: 365.

Taxonomic history

Conomyrma in Dolichoderinae, Tapinomini: Forel, 1917: 248; Donisthorpe, 1943f: 634; Jaffe, 1993: 9.

Conomyrma as subgenus of Dorymyrmex: Forel, 1913b: 350; Forel, 1917: 248; Santschi, 1922b: 365; Wheeler, W.M. 1922a: 689; Gallardo, 1930: 147; Smith, M.R. 1951a: 837; Smith, M.R. 1958c: 140.

Conomyrma as genus: Kusnezov, 1952g: 429; Kusnezov, 1959: 51; Kusnezov, 1964: 66; Kempf, 1972a: 78; Snelling, R.R. 1973b: 1; Smith, D.R. 1979: 1419; Hölldobler & Wilson, 1990: 17; Jaffe, 1993: 9.

Conomyrma as junior synonym of Dorymyrmex: Brown, 1973b: 179 [provisional]; Shattuck, 1992c: 77; Shattuck, 1994: 73; Bolton, 1994: 26; Bolton, 2003: 86.

Araucomyrmex Gallardo, 1919b: 249. Type-species: Dorymyrmex tener, by original designation.

Taxonomic history

Araucomyrmex in Dolichoderinae, Tapinomini: Wheeler, W.M. 1922a: 689; Donisthorpe, 1943f: 625; Kempf, 1972a: 25; Dlussky & Fedoseeva, 1988: 77.

Araucomyrmex as genus: Gallardo, 1919b: 249; Wheeler, W.M. 1922a: 689; Kusnezov, 1956: 28; Kusnezov, 1959: 51; Kusnezov, 1964: 66; Kempf, 1972a: 25; Snelling, R.R. 1975: 9; Snelling, R.R. & Hunt, 1976: 93; Dlussky & Fedoseeva, 1988: 77.

Araucomyrmex as subgenus of Dorymyrmex: Santschi, 1922b: 365; Gallardo, 1930: 147; Kusnezov, 1952g: 428.

Araucomyrmex as junior synonym of Conomyrma: Snelling, R.R. 1981: 402.

Araucomyrmex as junior synonym of Dorymyrmex: Brown, 1973b: 178 [provisional]; Shattuck, 1992c: 77; Shattuck, 1994: 73; Bolton, 1994: 26; Bolton, 2003: 86.

Ammomyrma Santschi, 1922b: 365 [as subgenus of Dorymyrmex]. Type-species: Dorymyrmex exsanguis, by original designation.

Taxonomic history

Ammomyrma as subgenus of Dorymyrmex: Santschi, 1922b: 365; Gallardo, 1930: 147; Kempf, 1972a: 100.

Ammomyrma as junior synonym of Araucomyrmex: Snelling, R.R. & Hunt, 1976: 93.

Ammomyrma as junior synonym of Dorymyrmex: Shattuck, 1992c: 77; Shattuck, 1994: 73; Bolton, 1994: 26; Bolton, 2003: 87.

Biconomyrma Kusnezov, 1952g: 429 [as subgenus of Conomyrma]. Type-species: Dorymyrmex pyramicus var. brunneus (now Dorymyrmex brunneus), by subsequent designation of Kusnezov, 1959: 51.

Taxonomic history

Biconomyrma as subgenus of Conomyrma: Kusnezov, 1952g: 429.

Biconomyrma as genus: Kusnezov, 1959: 51; Kusnezov, 1964: 67.

Biconomyrma as junior synonym of Conomyrma: Smith, M.R. 1958c: 140; Kempf, 1972a: 78.

Biconomyrma as junior synonym of Dorymyrmex: Shattuck, 1992c: 78; Shattuck, 1994: 73; Bolton, 1994: 26; Bolton, 2003: 87.

Spinomyrma Kusnezov, 1952g: 429 (diagnosis in key) [as subgenus of Dorymyrmex]. Type-species: Dorymyrmex alboniger, by subsequent designation of Kusnezov, 1959: 51.

Taxonomic history

Spinomyrma as subgenus of Dorymyrmex: Kusnezov, 1952g: 429; Kempf, 1972a: 100.

Spinomyrma as genus: Kusnezov, 1956: 30 (in key); Kusnezov, 1959: 51; Kusnezov, 1964: 66.

Spinomyrma as junior synonym of Dorymyrmex: Snelling, R.R. & Hunt, 1976: 93; Shattuck, 1992c: 78; Shattuck, 1994: 74; Bolton, 1994: 26; Bolton, 2003: 87.

Genus Dorymyrmex references

Mayr, G. 1868a: 165 (diagnosis); Forel, 1878: 383 (diagnosis); Cresson, 1887: 257 (U.S.A. catalogue); Dalla Torre, 1893: 167 (catalogue); André, 1903: 364 (species key); Emery, 1913a: 36 (diagnosis, catalogue); Gallardo, 1916a: 21 (Argentina species key); Wheeler, W.M. 1922a: 689 (subgenera key); Cole, 1942: 371 (U.S.A., Utah species key); Creighton, 1950a: 348 (North America species key); Kusnezov, 1952g: 428 (subgenera key); Kempf, 1972a: 25, 78, 99 (Neotropical Araucomyrmex, Conomyrma, Dorymyrmex catalogues); Snelling, R.R. 1973b: 1 (U.S.A. Conomyrma species, revision); Snelling, R.R. & Hunt, 1976: 96 (Chile Araucomyrmex species key); Smith, D.R. 1979: 1419 (North America catalogue); Allred, 1982: 444 (U.S.A., Utah Conomyrma species key); Wheeler, G.C. & Wheeler, J. 1986g: 57 (U.S.A., Nevada Conomyrma species key); Trager, 1988: 14 (Southeastern U.S.A. Conomyrma species revision, key): Johnson, 1989b: 191 (U.S.A. Conomyrma species revision, key); Brandão, 1991: 327, 338, 340 (catalogue); Shattuck, 1992c: 77 (diagnosis, review of genus); Shattuck, 1994: 73 (catalogue); Snelling, R.R. 1995: 9 (U.S.A. species revision, key); Bolton, 1995a: 1049 (census); Bolton, 1995b: 181 (catalogue); Mackay & Mackay, 2002: 222 (U.S.A., New Mexico species key).

Dolichoderinae Leptomyrmecini Forelius TRUE TRUE FALSE

18 species

Forelius Emery, 1888d: 389. Type-species: Iridomyrmex mccooki, by monotypy.

Taxonomic history

Forelius in Dolichoderinae: Dalla Torre, 1893: 168; Emery, 1895j: 771; Forel, 1899c: 102; Wheeler, W.M. 1910g: 142.

Forelius in Dolichoderidae: Ashmead, 1905b: 384.

Forelius in Dolichoderinae, Tapinomini: Emery, 1913a: 35; Forel, 1917: 248; Wheeler, W.M. 1922a: 690; Donisthorpe, 1943f: 645; all subsequent authors and Jaffe, 1993: 9.

Forelius in Dolichoderinae, Dolichoderini: Shattuck, 1992c: 87; Bolton, 1994: 26; Bolton, 2003: 88.

Forelius in Dolichoderinae, Leptomyrmecini: Ward, Brady, et al. 2010: 361.

Junior synonym of FORELIUS

Neoforelius Kusnezov, 1953b: 327. Type-species: Neoforelius tucumanus, by monotypy.

Taxonomic history

Neoforelius in Dolichoderinae, Tapinomini: Kusnezov, 1953b: 327.

Neoforelius as genus: Kusnezov, 1953b: 327; Kusnezov, 1959: 51; Kusnezov, 1964: 66; Kempf, 1972a: 160; Jaffe, 1993: 9.

Neoforelius as junior synonym of Forelius: Shattuck, 1992c: 87; Shattuck, 1994: 91; Bolton, 1994: 26; Cuezzo, 2000: 204; Bolton, 2003: 88.

Genus Forelius references

Dalla Torre, 1893: 168 (catalogue); Emery, 1913a: 35 (diagnosis, catalogue); Gallardo, 1916a: 74 (Argentina species key); Creighton, 1950a: 343 (North America species, review); Kusnezov, 1957b: 16 (Neotropical species key); Kempf, 1972a: 108, 160 (Neotropical Forelius, Neoforelius catalogues); Smith, D.R. 1979: 1419 (North America catalogue); Wheeler, G.C. & Wheeler, J. 1986g: 56 (U.S.A., Nevada species key); Brandão, 1991: 344 (catalogue); Shattuck, 1992c: 87 (diagnosis, review of genus); Shattuck, 1994: 91 (catalogue); Bolton, 1995a: 1049 (census); Bolton, 1995b: 190 (catalogue); Cuezzo, 2000: 197 (diagnosis, all species revision, key); Mackay & Mackay, 2002: 227 (U.S.A., New Mexico species key); Guerrero & Fernández, 2008: 58 (addition to key).

Dolichoderinae Leptomyrmecini Iridomyrmex TRUE TRUE FALSE

Extant: 79 species

Fossil:

Iridomyrmex Mayr, 1862: 653 (diagnosis in key), 702. Type-species: Formica detecta (junior synonym of Formica purpurea), by subsequent designation of Bingham, 1903: 297.

Taxonomic history

Iridomyrmex in Formicinae: Mayr, 1862: 653 (in key) [Formicidae]; Mayr, 1865: 10 [Formicidae].

Iridomyrmex in Dolichoderinae: Forel, 1878: 381 [Dolichoderidae]; Dalla Torre, 1893: 168; Emery, 1895j: 771; Forel, 1899c: 102; Wheeler, W.M. 1910g: 142.

Iridomyrmex in Dolichoderidae: Ashmead, 1905b: 384.

Iridomyrmex in Dolichoderinae, Tapinomini: Emery, 1913a: 21; Wheeler, W.M. 1915h: 86; Arnold, 1915: 145; Forel, 1917: 248; Wheeler, W.M. 1922a: 689; Carpenter, 1930: 50; all subsequent authors; Jaffe, 1993: 9 (anachronism).

Iridomyrmex in Dolichoderinae, Dolichoderini: Shattuck, 1992c: 101; Bolton, 1994: 26; Bolton, 2003: 88.

Iridomyrmex in Dolichoderinae, Iridomyrmecini, Iridomyrmecina: Dubovikov, 2005a: 92.

Iridomyrmex in Dolichoderinae, Leptomyrmecini: Ward, Brady, et al. 2010: 362.

Genus Iridomyrmex references

Roger, 1863b: 13 (catalogue); Mayr, 1863: 424 (catalogue); Mayr, 1865: 10 (diagnosis); Mayr, 1867a: 76 (diagnosis); Forel, 1878: 381 (diagnosis); Cresson, 1887: 257 (U.S.A. catalogue); Dalla Torre, 1893: 168 (catalogue); Forel, 1895e: 467 (India & Sri Lanka species key); Bingham, 1903: 298 (India, Sri Lanka & Burma species key); Emery, 1913a: 21 (diagnosis, catalogue); Arnold, 1915: 145 (diagnosis); Creighton, 1950a: 341 (North America species key); Chapman & Capco, 1951: 188 (Asia checklist); Bernard, 1967: 250 (diagnosis); Kempf, 1972a: 124 (Neotropical catalogue); Smith, D.R. 1979: 1417 (North America catalogue); Taylor & Brown, D.R. 1985: 96 (Australia catalogue); Taylor, 1987a: 30 (Australia, New Caledonia & New Zealand checklist); Shattuck, 1992a: 14 (diagnosis); Shattuck, 1992c: 101 (diagnosis, review of genus); Shattuck, 1993a: 118 (I. purpureus group, key); Shattuck, 1993b: 1306 (I. calvus group, key); Shattuck, 1994: 99 (catalogue); Bolton, 1995a: 1050 (census); Bolton, 1995b: 217 (catalogue); Shattuck, 1996a: 37 (I. discors group); Shattuck, 1998: 303 (I. conifer group, key); Shattuck, 1999: 72 (Australia synopsis); Heterick, 2009: 48 (south-western Australia species key); Heterick & Shattuck, 2011: 22 (diagnosis, all species revision, key).

Dolichoderinae Tapinomini Liometopum TRUE TRUE FALSE

Extant: 7 species

Fossil: 9 species

Liometopum Mayr, 1861: 38. Type-species: Formica microcephala, by monotypy.

Taxonomic history

Liometopum in Formicinae: Mayr, 1861: 38 [Formicidae]; Mayr, 1862: 653 (in key) [Formicidae]; Mayr, 1865: 9 [Formicidae]; André, 1874: 168 [Formicidae].

Liometopum in Dolichoderinae: Forel, 1878: 383 [Dolichoderidae]; Emery & Forel, 1879a: 454 [Dolichoderidae]; Dalla Torre, 1893: 163; Emery, 1895j: 771; Forel, 1899c: 104; Wheeler, W.M. 1910g: 142.

Liometopum in Dolichoderidae: Ashmead, 1905b: 384.

Liometopum in Dolichoderinae, Tapinomini: Emery, 1913a: 19; Wheeler, W.M. 1915h: 95; Forel, 1917: 248; Bondroit, 1918: 87; Wheeler, W.M. 1922a: 688; Carpenter, 1930: 46; subsequent authors except those below; then Jaffe, 1993: 9; Ward, Brady, et al. 2010: 361.

Liometopum in Dolichoderinae, Liometopini: Dlussky & Fedoseeva, 1988: 77.

Liometopum in Dolichoderinae, Dolichoderini: Shattuck, 1992c: 121; Bolton, 1994: 26; Bolton, 2003: 89.

Genus Liometopum references

Roger, 1863b: 14 (catalogue); Mayr, 1863: 427 (catalogue); Mayr, 1865: 9 (diagnosis); Forel, 1878: 383 (diagnosis); André, 1882b: 220 (Europe & Algeria species); Cresson, 1887: 258 (U.S.A. catalogue); Dalla Torre, 1893: 162 (catalogue); Bingham, 1903: 289 (diagnosis); Wheeler, W.M. 1905e: 322 (North America species); Emery, 1913a: 19 (diagnosis, catalogue); Creighton, 1950a: 338 (North America species key); Chapman & Capco, 1951: 192 (Asia checklist); Gregg, 1963: 438 (U.S.A., Colorado species key); Bernard, 1967: 253 (diagnosis); Smith, D.R. 1979: 1417 (North America catalogue); Wheeler, G.C. & Wheeler, J. 1986g: 55 (U.S.A., Nevada species key); Shattuck, 1992c: 121 (diagnosis, review of genus); Shattuck, 1994: 128 (catalogue); Bolton, 1995a: 1050 (census); Bolton, 1995b: 247 (catalogue); Mackay & Mackay, 2002: 232 (U.S.A., New Mexico species key); Del Toro, Pacheco & Mackay, 2009: 306 (all species revision, key).

Dolichoderinae Tapinomini Tapinoma TRUE TRUE FALSE

Extant: 64 species, 26 subspecies

Fossil: 4 species

Tapinoma Foerster, 1850a: 43. Type-species: Tapinoma collina (junior synonym of Formica erratica), by monotypy.

Taxonomic history

Tapinoma in Formicidae: Smith, F. 1858b: 55.

Tapinoma in Formicinae: Mayr, 1855: 372 [Formicidae]; Mayr, 1861: 41 [Formicidae]; Mayr, 1862: 653 (in key) [Formicidae]; Mayr, 1865: 10 [Formicidae]; André, 1874: 168 [Formicidae]; André, 1881b: 63 [Formicidae].

Tapinoma in Dolichoderinae: Forel, 1878: 385 [Dolichoderidae]; Emery & Forel, 1879a: 454 [Dolichoderidae]; Forel, 1895b: 109 [Dolichoderidae]; Dalla Torre, 1893: 164; Emery, 1895j: 771; Forel, 1899c: 101; Wheeler, W.M. 1910g: 142.

Tapinoma in Dolichoderidae: Ashmead, 1905b: 384.

Tapinoma in Dolichoderinae, Tapinomini: Emery, 1913a: 38; Arnold, 1915: 152; Forel, 1917: 248; Bondroit, 1918: 89; Wheeler, W.M. 1922a: 690; subsequent authors except those below; then Jaffe, 1993: 9; Dubovikov, 2005a: 91 (implied); Ward, Brady, et al. 2010: 361.

Tapinoma in Dolichoderinae, Dolichoderini: Shattuck, 1992c: 146; Bolton, 1994: 26; Bolton, 2003: 91.

Junior synonyms of TAPINOMA

Micromyrma Dufour, 1857: 60. Type-species: Micromyrma pygmaea, by monotypy.

Taxonomic history

[Type-species not Formica erratica, unjustified subsequent designation by Wheeler, W.M. 1911f: 167.]

Micromyrma as subgenus of Tapinoma: Menozzi, 1925a: 19; Santschi, 1928e: 472; Chapman & Capco, 1951: 192.

Micromyrma as junior synonym of Tapinoma: Mayr, 1863: 455; Forel, 1878: 385; Emery & Forel, 1879a: 454; Dalla Torre, 1893: 164; Emery, 1913a: 38; Wheeler, W.M. 1922a: 923; Smith, M.R. 1951a: 837; Shattuck, 1992c: 146; Shattuck, 1994: 140; Bolton, 1994: 26; Bolton, 2003: 91; Fisher & Bolton, 2007: 65.

Semonius Forel, 1910f: 21. Type-species: Semonius schultzei, by monotypy.

Taxonomic history

Semonius in Dolichoderinae: Wheeler, W.M. 1910g: 142.

Semonius in Dolichoderinae, Tapinomini: Emery, 1913a: 44; Arnold, 1915: 157; Forel, 1917: 247; Wheeler, W.M. 1922a: 688; all subsequent authors.

Semonius as junior synonym of Tapinoma: Shattuck, 1992c: 146; Shattuck, 1994: 140; Bolton, 1994: 26; Bolton, 2003: 91; Fisher & Bolton, 2007: 66.

Pseudaphomomyrmex Wheeler, W.M. 1920: 53. Type-species: Aphomomyrmex emeryi, by original designation.

Taxonomic history

Pseudaphomomyrmex in Formicinae, Dimorphomyrmecini: Donisthorpe, 1943g: 721.

Pseudaphomomyrmex incertae sedis in Formicinae: Dlussky & Fedoseeva, 1988: 77.

Pseudaphomomyrmex in Formicinae, Myrmelachistini: Hölldobler & Wilson, 1990: 18.

Pseudaphomomyrmex in Formicinae, Brachymyrmecini: Bolton, 1994: 50.

Pseudaphomomyrmex in Formicinae, Plagiolepidini: Wheeler, W.M. 1922a: 695; Chapman & Capco, 1951: 214; Wheeler, G.C. & Wheeler, J. 1985: 258; Bolton, 2003: 106.

Pseudaphomomyrmex in Dolichoderinae: LaPolla & Longino, 2006: 305.

Pseudaphomomyrmex as junior synonym of Aphomomyrmex: Emery, 1925b: 44; Donisthorpe, 1943g: 721.

Pseudaphomomyrmex as genus: Wheeler, W.M. 1920: 53; Chapman & Capco, 1951: 214; Bolton, 1994: 50; Bolton, 1995b: 369; Bolton, 2003: 106.

Pseudaphomomyrmex as junior synonym of Tapinoma: Fisher & Bolton, 2007: 66.

Zatapinoma Wheeler, W.M. 1928a: 20. Type-species: Zatapinoma annandalei, by original designation.

Taxonomic history

Zatapinoma in Dolichoderinae, Tapinomini: Donisthorpe, 1943g: 737; all subsequent authors.

Zatapinoma as junior synonym of Tapinoma: Shattuck, 1992c: 146; Shattuck, 1994: 140; Bolton, 1994: 26; Bolton, 2003: 91; Fisher & Bolton, 2007: 66.

Neoclystopsenella Kurian, 1955: 133. Type-species: Neoclystopsenella luffae, by monotypy.

Taxonomic history

Neoclystopsenella in Bethylidae: Kurian, 1955: 133.

Neoclystopsenella in Dolichoderinae: Brown, 1988a: 337.

Neoclystopsenella as junior synonym of Tapinoma: Brown, 1988a: 337; Shattuck, 1992c: 146; Shattuck, 1994: 140; Bolton, 1994: 26; Bolton, 2003: 91; Fisher & Bolton, 2007: 66.

Genus Tapinoma references

Mayr, 1855: 372 (diagnosis); Smith, F. 1858b: 55 (diagnosis); Roger, 1863b: 13, 14 (Tapinoma, Micromyrma catalogues); Mayr, 1863: 455 (catalogue); Mayr, 1865: 10 (diagnosis); Mayr, 1867a: 78 (diagnosis); André, 1882b: 222 (Europe & Algeria species key); Cresson, 1887: 258 (U.S.A. catalogue); Dalla Torre, 1893: 164 (catalogue); Bingham, 1903: 304 (India, Sri Lanka & Burma species key); Emery, 1913a: 38 (diagnosis, catalogue); Emery, 1913a: 44 (Semonius diagnosis, catalogue); Arnold, 1915: 152, 157 (diagnosis, South Africa species key, Semonius diagnosis); Wheeler, W.M. 1922a: 695 (Pseudaphomomyrmex diagnosis, in key). Wheeler, W.M. 1922a: 923, 927 (Afrotropical Tapinoma, Semonius catalogues); Wheeler, W.M. 1922a: 1034 (Malagasy catalogue); Emery, 1925d: 63 (Palaearctic species key); Kuznetsov-Ugamsky, 1927d: 33 (Turkestan species key); Smith, M.R. 1928a: 311 (North America species key); Kratochvíl, Novák & Snoflák, 1944: 79 (Czechoslovakia species key); Creighton, 1950a: 352 (North America species key); Chapman & Capco, 1951: 192, 196 (Asia Semonius, Tapinoma, Zatapinoma checklists); Bernard, 1967: 254 (diagnosis, Western Europe species key); Boven, 1970b: 24 (Netherlands species key); Kempf, 1972a: 246 (Neotropical catalogue); Tarbinsky, 1976: 118 (Kyrghyzstan species key); Kutter, 1977c: 178 (Switzerland species key); Arnol'di & Dlussky, 1978: 547 (former European U.S.S.R. species key); Collingwood, 1978: 87 (Iberian Peninsula species key); Smith, D.R. 1979: 1421 (North America catalogue); Schembri & Collingwood, 1981: 421 (Malta species key); Gösswald, 1985: 317 (Germany species key); Taylor & Brown, D.R. 1985: 106 (Australia catalogue); Agosti & Collingwood, 1987b: 279 (Balkans species key); Taylor, 1987a: 77 (Australia, New Caledonia & New Zealand checklist); Dlussky, Soyunov & Zabelin, 1990: 168 (Turkmenistan species key); Brandão, 1991: 380 (Neotropical catalogue); Shattuck, 1992c: 146 (diagnosis, review of genus); Arakelian, 1994: 72 (Armenia species key); Shattuck, 1994: 140 (catalogue); Bolton, 1995a: 1053 (census); Bolton, 1995b: 399 (catalogue); Wu, J. & Wang, 1995: 116 (China species key); Collingwood & Agosti, 1996: 360 (Saudi Arabia species key); Seifert, 1996b: 165 (Central Europe species key); Skinner & Allen, 1996: 40 (Britain species key); Collingwood & Prince, 1998: 20 (Portugal species key); Shattuck, 1999: 81 (Australia synopsis); Zhou, 2001b: 151 (China, Guangxi species key); Czechowski, Radchenko & Czechowska, 2002: 135 (Poland species key); Seifert, 2007: 149 (North and Central Europe species key); Heterick, 2009: 58 (south-western Australia species key); Yoshimura & Fisher, 2011: 13 (Malagasy males).

Formicinae Lasiini Acropyga TRUE TRUE FALSE

Extant: 40 species

Fossil: 1 species

Acropyga Roger, 1862a: 242. Type-species: Acropyga acutiventris,

by monotypy.

Taxonomic history

Acropyga in Camponotinae: Forel, 1878: 379 [Camponotidae]; Dalla Torre, 1893: 174.

Acropyga in Camponotinae, Plagiolepidini: Forel, 1886f: 212; Forel, 1893a: 165; Forel, 1895b: 107; Emery, 1895j: 771; Wheeler, W.M. 1910g: 143; Forel, 1912i: 88; Forel, 1917: 249.

Acropyga in Formicinae, Plagiolepidini: Ashmead, 1905b: 384; Wheeler, W.M. 1922a: 695; Emery, 1925b: 27; all subsequent authors except the two following.

Acropyga incertae sedis in Formicinae: Dlussky & Fedoseeva, 1988: 77.

Acropyga in Formicinae, Lasius genus group: Agosti, 1991: 296.

Acropyga in Formicinae, Lasiini: Bolton, 2003: 22, 94.

Acropyga as junior synonym of Plagiolepis: Mayr, 1863: 394.

Acropyga as genus: Roger, 1862a: 242; Smith, F. 1871a: 319; Forel, 1878: 379; all subsequent authors.

Junior synonyms of ACROPYGA.

Rhizomyrma Forel, 1893g: 347 [as subgenus of Acropyga]. Type-species: Acropyga (Rhizomyrma) goeldii, by subsequent designation of Wheeler, W.M. 1911f: 172.

Taxonomic history

Rhizomyrma in Camponotinae, Plagiolepidini: Forel, 1893g: 347; Wheeler, W.M. 1910g: 143; Forel, 1912i: 89; Forel, 1917: 249.

Rhizomyrma in Formicinae, Plagiolepidini: Wheeler, W.M. 1922a: 695.

Rhizomyrma as genus: Emery, 1906c: 182; Forel, 1912i: 89; Wheeler, W.M. 1910g: 143; Bruch, 1914: 227; Wheeler, W.M. & Mann, 1914: 46; Forel, 1917: 249; Wheeler, W.M. 1922a: 695; Kusnezov, 1956: 33; Kusnezov, 1964: 68.

Rhizomyrma as subgenus of Acropyga: Forel, 1893g: 347; Forel, 1912m: 771; Emery, 1925b: 29; subsequent authors except below; Bolton, 1994: 51; Bolton, 2003: 95.

Rhizomyrma as junior synonym of Acropyga: Brown, 1973b: 184 [provisional]; Hölldobler & Wilson, 1990: 18; LaPolla, 2004a: 16.

Atopodon Forel, 1912m: 771 [as subgenus of Acropyga]. Type-species: Acropyga (Atopodon) inezae, by subsequent designation of Wheeler, W.M. 1913a: 79.

Taxonomic history

[Atopodon also described as new by Forel, 1913k: 100.]

Atopodon in Camponotinae, Plagiolepidini: Forel, 1917: 249.

Atopodon in Formicinae, Plagiolepidini: Wheeler, W.M. 1922a: 695.

Atopodon as subgenus of Rhizomyrma: Forel, 1917: 249.

Atopodon as genus: Wheeler, W.M. 1922a: 695.

Atopodon as subgenus of Acropyga: Forel, 1912m: 771; Forel, 1913k: 100; Emery, 1925b: 30; subsequent authors except below; Bolton, 1994: 51; Bolton, 2003: 95.

Atopodon as junior synonym of Acropyga: Brown, 1973b: 178 [provisional]; Hölldobler & Wilson, 1990: 18; LaPolla, 2004a: 16.

Malacomyrma Emery, 1922d: 109 [as subgenus of Acropyga]. Type-species: Acropyga silvestrii, by monotypy.

Taxonomic history

Malacomyrma in Formicinae, Pseudolasius genus group: Agosti, 1991: 296.

Malacomyrma as subgenus of Acropyga: Emery, 1922d: 109; Emery, 1925b: 30; subsequent authors except below; Bolton, 1994: 51; Bolton, 2003: 95.

Malacomyrma as junior synonym of Acropyga: Brown, 1973b: 182 [provisional]; Hölldobler & Wilson, 1990: 18; LaPolla, 2004a: 16.

Genus Acropyga references

Roger, 1863b: 11 (catalogue); Forel, 1878: 379 (diagnosis); Dalla Torre, 1893: 174 (catalogue); Bingham, 1903: 332 (diagnosis); Wheeler, W.M. 1922a: 928 (Afrotropical catalogue); Emery, 1925b: 27 (diagnosis, subgenera key, catalogue); Emery, 1925b: 29, 30 (A. (Rhizomyrma, Atopodon & Malacomyrma) diagnoses, catalogues); Weber, 1944: 91 (Neotropical A. (Rhizomyrma) species key); Chapman & Capco, 1951: 210 (Asia checklist); Kempf, 1972a: 16 (Neotropical catalogue); Smith, D.R. 1979: 1423 (North America catalogue); Taylor & Brown, D.R. 1985: 107 (Australia catalogue); Taylor, 1987a: 5 (Australia checklist); Morisita, Kubota, et al., 1991: 14 (Japan species key); Wang, C. & Wu, 1992a: 227 (China species key); Bolton, 1994: 51 (synoptic classification); Bolton, 1995a: 1047 (census); Bolton, 1995b: 57 (catalogue); Shattuck, 1999: 86 (Australia synopsis); Terayama, Fellowes & Zhou, 2002: 22 (diagnosis, East Asian species key); LaPolla, 2004a: 16 (diagnosis, all species revision, New World species key: 23, Old World species key: 27); LaPolla & Fisher, 2005: 601 (Afrotropical species, key); LaPolla, Schultz, et al. 2006: 197 (phylogeny); Terayama, 2009: 204 (Taiwan species key); Heterick, 2009: 60 (south-western Australia species key).

Formicinae Lasiini Lasius TRUE TRUE FALSE

Extant: 100 species, 2 subspecies

Fossil: 22 species, 1 subspecies

Lasius Fabricius, 1804: 415. Type-species: Formica nigra, by subsequent designation of Bingham, 1903: 338.

Taxonomic history

[Lasius Jurine, 1801: 164 (Apidae), resurrected by Morice & Durrant, 1915: 421, later suppressed by Act of Commission.]

Lasius in Formicinae: Mayr, 1861: 49 [Formicidae]; Mayr, 1862: 652 (in key) [Formicidae]; Mayr, 1865: 8 [Formicidae]; Mayr, 1868c: 42 [Formicidae].

Lasius in Camponotinae: Forel, 1878: 374 [Camponotidae]; Emery & Forel, 1879a: 452 [Camponotidae]; Forel, 1886f: 206; Dalla Torre, 1893: 181.

Lasius in Camponotinae, Camponotini: Forel, 1886f: 206.

Lasius in Camponotinae, Formicini: Forel, 1893a: 165; Forel, 1895b: 103; Emery, 1895j: 772; Forel, 1899c: 127; Wheeler, W.M. 1910g: 143; Forel, 1912i: 89; Wheeler, W.M. 1915h: 120.

Lasius in Formicinae, Formicini: Wheeler, W.M. 1922a: 698; Carpenter, 1930: 58; Chapman & Capco, 1951: 201; Wheeler, G.C. & Wheeler, J. 1970: 651; Wheeler, G.C. & Wheeler, J. 1976b: 101; Wheeler, G.C. & Wheeler, J. 1985: 258.

Lasius in Formicinae, Acanthomyopsini: Donisthorpe, 1943f: 655.

Lasius in Formicinae, Lasius genus group: Agosti, 1991: 296.

Lasius in Formicinae, Lasiini: Ashmead, 1905b: 384; Emery, 1925b: 226; all subsequent authors except the entries above; Bolton, 1994: 50; Bolton, 2003: 22, 96.

Lasius as junior synonym of Acanthomyops: Forel, 1916: 460; Donisthorpe, 1916c: 276; Morice & Durrant, 1917: 442. Donisthorpe, 1927b: 206; Donisthorpe, 1937d: 132; Donisthorpe, 1943f: 655; Donisthorpe, 1946g: 91.

Lasius as junior synonym of Formicina: Emery, 1916a: 61; Emery, 1916b: 239; Emery, 1917a: 96; Wheeler, W.M. 1916k: 170; Bondroit, 1918: 19.

Lasius as genus: Fabricius, 1804: 415; Wheeler, W.M. 1916k: 172; Emery, 1925b: 226; Wilson, 1955a: 11; all subsequent authors.

Junior synonyms of LASIUS

Donisthorpea Morice & Durrant, 1915: 423. Type-species: Formica nigra, by original designation.

Taxonomic history

[Unnecessary replacement name for Lasius Fabricius.]

Donisthorpea in Formicinae, Acanthomyopsini: Donisthorpe, 1943f: 640; Donisthorpe, 1946g: 91.

Donisthorpea as genus: Morice & Durrant, 1915: 423; Donisthorpe, 1915d: 184.

Donisthorpea as subgenus of Formicina: Emery, 1916a: 11; Emery, 1916b: 240.

Donisthorpea as subgenus of Acanthomyops: Donisthorpe, 1927b: 229; Donisthorpe, 1943f: 640; Donisthorpe, 1946g: 91.

Donisthorpea as junior synonym of Acanthomyops: Forel, 1916: 460; Morice & Durrant, 1917: 442; Donisthorpe, 1927b: 209.

Donisthorpea as junior synonym of Lasius: Wheeler, W.M. 1916k: 172; Emery, 1925b: 226; Wilson, 1955a: 11. [Lasius and Donisthorpea share the same type-species; synonymy is therefore absolute.]

†Tylolasius Zhang, 1989: 295. Type-species: †Tylolasius inflatus, by original designation.

Taxonomic history

†Tylolasius incertae sedis in Formicinae: Bolton, 2003: 28, 131.

†Tylolasius as junior synonym of Lasius: Hong, Wu & Ren, 2001: 348.

Subgenera of LASIUS include the nominal plus the following.

Subgenus LASIUS (ACANTHOMYOPS)

Acanthomyops Mayr, 1862: 652 (diagnosis in key), 699. Type-species: Formica clavigera, by monotypy.

Taxonomic history

Acanthomyops in Formicinae: Mayr, 1862: 652 (in key) [Formicidae]; Mayr, 1865: 8 [Formicidae].

Acanthomyops in Camponotinae, Formicini: Wheeler, W.M. 1910g: 143; Forel, 1917: 249.

Acanthomyops in Formicinae, Formicini: Wheeler, W.M. 1922a: 698; Wheeler, G.C. & Wheeler, J. 1970: 651; Wheeler, G.C. & Wheeler, J. 1976b: 101; Wheeler, G.C. & Wheeler, J. 1985: 258; Jaffe, 1993: 14.

Acanthomyops in Formicinae, Acanthomyopsini: Donisthorpe, 1943f: 618.

Acanthomyops in Formicinae, Lasius genus group: Agosti, 1991: 296.

Acanthomyops in Formicinae, Lasiini: Ashmead, 1905b: 384 (misspelled as Acanthomyrmex); Emery, 1925b: 236; all subsequent authors except for those above.

Acanthomyops as junior synonym of Lasius: Mayr, 1866b: 888; Dalla Torre, 1893: 181.

Acanthomyops as subgenus of Formicina: Emery, 1916a: 11; Forel, 1917: 249.

Acanthomyops as genus: Mayr, 1862: 699; Donisthorpe, 1927b: 209; Donisthorpe, 1943f: 618; Creighton, 1950a: 426; Smith, M.R. 1958c: 150; Wing, 1968: 47; Bolton, 1994: 50; Bolton, 2003: 94.

Acanthomyops as subgenus of Lasius: Emery, 1893i: 642; Wheeler, W.M. 1910g: 143; Wheeler, W.M. 1916k: 172; Wheeler, W.M. 1922a: 698; Emery, 1925b: 236; Buren, 1950: 188; Smith, M.R. 1951a: 852; Gregg, 1963: 453; Ward, 2005: 13.

Subgenus LASIUS (DENDROLASIUS)

Dendrolasius Ruzsky, 1912: 630 [as subgenus of Lasius]. Type-species: Formica fuliginosa, by monotypy.

Taxonomic history

[Dendrolasius also described as new by Ruzsky, 1914a: 59.]

Dendrolasius as subgenus of Formicina: Forel, 1917: 249.

Dendrolasius as subgenus of Acanthomyops: Donisthorpe, 1927b: 211; Donisthorpe, 1943f: 637.

Dendrolasius as subgenus of Lasius: Ruzsky, 1912: 630; Wheeler, W.M. 1922a: 698; Emery, 1925b: 235; Wilson, 1955a: 14; all subsequent authors except the following.

Dendrolasius as junior synonym of Lasius: Brown, 1973b: 180 [provisional].

Subgenus LASIUS (CHTHONOLASIUS)

Chtonolasius Ruzsky, 1912: 630 [as subgenus of Lasius]. Type-species: Formica umbrata, by subsequent designation of Emery, 1925b: 232.

Taxonomic history

[Chtonolasius also described as new by Ruzsky, 1914a: 59. The claim by some authors (e.g. Emery, 1916a: 11; Donisthorpe, 1941f: 37; Donisthorpe, 1943f: 633) that Ruzsky nominated Lasius flavus as type-species is incorrect. Ruzsky (1912: 630) included those species "with the appearance of the yellow Lasius (Las. flavus D.G.) and its closely related species, and races (Las. carniolicus, Las. umbratus, Las. myops, Las. mixtus)". This is not a direct nomination of a type-species, nor is De Geer (= D.G.) the author of flavus: Bolton, 1995b: 24.]

Chthonolasius justified emendation of spelling: Wheeler, W.M. 1916k: 170.

Chthonolasius as junior synonym of Formicina: Wheeler, W.M. 1916k: 172; Emery, 1916a: 11.

Chthonolasius as subgenus of Acanthomyops: Donisthorpe, 1927b: 254.

Chthonolasius as subgenus of Lasius: Ruzsky, 1912: 630; Ruzsky, 1914a: 59; Wheeler, W.M. 1922a: 698; Emery, 1925b: 231; Wilson, 1955a: 13; all subsequent authors except the following.

Chthonolasius as junior synonym of Lasius: Brown, 1973b: 179 [provisional].

Subgenus LASIUS (CAUTOLASIUS)

Cautolasius Wilson, 1955a: 13 [as subgenus of Lasius]. Type-species: Formica flava, by original designation.

Taxonomic history

Cautolasius as junior synonym of Lasius: Brown, 1973b: 179 [provisional].

Cautolasius as subgenus of Lasius: Bolton, 2003: 97.

Subgenus LASIUS (AUSTROLASIUS)

Austrolasius Faber, 1967: 74 [as subgenus of Lasius]. Type-species: Lasius carniolicus, by original designation.

Taxonomic history

Austrolasius as junior synonym of Lasius: Brown, 1973b: 179 [provisional].

Autrolasius as subgenus of Lasius: Bolton, 2003: 97.

Genus Lasius references

Mayr, 1861: 49 (Europe species key); Roger, 1863b: 11 (catalogue); Mayr, 1863: 394, 425 (catalogue); Mayr, 1865: 8 (diagnosis); Mayr, 1867a: 74 (diagnosis); André, 1874: 179 (Europe species key); Forel, 1874: 46 (Switzerland species key); Forel, 1878: 374 (diagnosis); André, 1882b: 191 (Europe & Algeria species key); Cresson, 1887: 257 (U.S.A. catalogue); Provancher, 1887: 236 (Canada species key); Nasonov, 1889: 64 (Russia species key); Dalla Torre, 1893: 181 (catalogue); Emery, 1893i: 637 (North America species key); Bingham, 1903: 339 (India species key); Ruzsky, 1905b: 264 (Russian Empire species key); Wasmann, 1906: 10 (Luxemburg species key); Bondroit, 1910: 484 (Belgium species key); Wheeler, W.M. 1910e: 237 (North America L. umbratus forms, key); Stitz, 1914: 82 (Central Europe species key); Donisthorpe, 1915d: 188 (Britain species key); Emery, 1916b: 243 (Italy species key); Wheeler, W.M. 1916m: 591 (U.S.A., Connecticut species key); Bondroit, 1918: 19 (France & Belgium species key); Wheeler, W.M. 1922a: 698 (subgenera key); Emery, 1925b: 226 (diagnosis, subgenera key, catalogue); Emery, 1925b: 231 (L. (Chthonolasius) diagnosis, catalogue); Emery, 1925b: 235 (L. (Dendrolasius) diagnosis, catalogue); Emery, 1925b: 236 (L. (Acanthomyops) diagnosis, catalogue); Kuznetsov-Ugamsky, 1927e: 186 (Turkestan species key); Karavaiev, 1927c: 277 (Ukraine species key); Donisthorpe, 1927b: 206 (Britain species key); Arnol'di, 1933b: 602 (Russia species key); Karavaiev, 1936: 193 (Ukraine species key); Stärcke, 1937: 57 (Europe L. umbratus group, key); Menozzi, 1939a: 313 (Himalaya & Tibet species key); Stitz, 1939: 267 (Germany species key); Kratochvíl, 1941: 100 (Central Europe species key); Novák & Sadil, 1941: 100 (Central Europe species key); Cole, 1942: 373 (U.S.A., Utah species key); Holgersen, 1943b: 174 (Norway species key); Holgersen, 1944: 199 (Norway species key); Buren, 1944a: 296 (U.S.A., Iowa species key); Stärcke, 1944a: 157 (Europe L. niger group, key); Boven, 1947: 184 (Belgium species key); Creighton, 1950a: 418 (North America Lasius species key); Creighton, 1950a: 428 (North America Acanthomyops species key); Buren, 1950: 188 (Acanthomyops queens, key); Chapman & Capco, 1951: 201 (Asia checklist); Wilson, 1955a: 13 (subgenera review); Wilson, 1955a: 26 (Nearctic Lasius species revision, key); Wilson, 1955a: 28 (Palaearctic species revision, key); Boven, 1959: 11 (Netherlands species key); Wheeler, G.C. & Wheeler, J. 1963: 177 (U.S.A., North Dakota Lasius species key); Wheeler, G.C. & Wheeler, J. 1963: 195 (U.S.A., North Dakota Acanthomyops species key); Gregg, 1963: 453 (U.S.A., Colorado species key); Collingwood, 1963: 155 (Europe L. umbratus group species key); Collingwood, 1964: 107 (Britain species key); Bernard, 1967: 349 (diagnosis, Western Europe species key); Wing, 1968: 53 (Acanthomyops diagnosis, all species revision, key); Bourne, 1973: 19 (Britain species key); Boven, 1970b: 28 (Netherlands species key); Bolton & Collingwood, 1975: 7 (Britain species key); Tarbinsky, 1976: 133 (Kyrghyzstan species key); Boven, 1977: 135 (Belgium species key); Kutter, 1977c: 208 (Switzerland species key); Arnol'di & Dlussky, 1978: 555 (former European U.S.S.R. species key); Collingwood, 1978: 89 (Iberian Peninsula species key); Collingwood, 1979: 92 (Fennoscandia & Denmark species key); Smith, D.R. 1979: 1435 (North America Lasius catalogue); Smith, D.R. 1979: 1440 (North America Acanthomyops catalogue); Yamauchi, 1979: 147 (Japan species keys); Allred, 1982: 444 (U.S.A., Utah Acanthomyops species key); Allred, 1982: 450 (U.S.A., Utah Lasius species key); Collingwood, 1982: 283 (Himalayan species key); Gösswald, 1985: 268 (Germany species key); Wheeler, G.C. & Wheeler, J. 1986g: 64 (U.S.A., Nevada Lasius species key); Wheeler, G.C. & Wheeler, J. 1986g: 68 (U.S.A., Nevada Acanthomyops species key); Nilsson & Douwes, 1987: 70 (Norway species key); Agosti & Collingwood, 1987b: 281 (Balkans species key); Dlussky & Fedoseeva, 1988: 77 (synoptic classification); Seifert, 1988a: 143 (Europe L. (Chthonolasius) species key); Kupyanskaya, 1989: 781 (East Palaearctic L. (Dendrolasius) species key); Dlussky, Soyunov & Zabelin, 1990: 158 (Turkmenistan species key); Seifert, 1990: 1 (supplement to European L. (Chthonolasius)); Agosti & Bolton, 1990b: 149 (characters); Kupyanskaya, 1990: 211 (Far Eastern Russia species key); Morisita, Kubota, Onoyama, et al., 1991: 24 (Japan species key); Agosti, 1991: 295 (Lasius genus group diagnosis); Seifert, 1992b: 48 (Palaearctic L. (Lasius) species key); Atanasov & Dlussky, 1992: 232 (Bulgaria species key); Arakelian, 1994: 111 (Armenia species key); Bolton, 1994: 50 (synoptic classification); Radchenko, 1994b: 114 (South Siberia species key); Bolton, 1995a: 1050 (census); Bolton, 1995b: 53 (Acanthomyops catalogue); Bolton, 1995b: 221 (Lasius catalogue); Douwes, 1995: 92 (Sweden species key); Kupyanskaya, 1995: 361 (Far Eastern Russia species key); Wu, J. & Wang, 1995: 153 (China species key); Seifert, 1996b: 179 (Central Europe species key); Skinner & Allen, 1996: 42 (Britain species key); Collingwood & Prince, 1998: 22 (Portugal species key); Zhou, 2001b: 187 (China, Guangxi species key); Blacker & Collingwood, 2002: 44 (British species key); Czechowski, Radchenko & Czechowska, 2002: 154 (Poland species key); Mackay & Mackay, 2002: 237 (U.S.A., New Mexico Acanthomyops species key); Mackay & Mackay, 2002: 331 (U.S.A., New Mexico Lasius species key); Radchenko, 2005a: 93 (East Palaearctic L. (Dendrolasius) species key); Radchenko, 2005b: 190 (North Korea species key); Seifert, 2007: 159 (North and Central Europe species key); Terayama, 2009: 207 (Taiwan species key).

Formicinae Lasiini Myrmecocystus TRUE TRUE FALSE

29 species

Myrmecocystus Wesmael, 1838: 770. Type-species: Myrmecocystus mexicanus, by monotypy.

Taxonomic history

Myrmecocystus in Camponotinae: Forel, 1878: 372 [Camponotidae]; Emery & Forel, 1879a: 449 [Camponotidae]; Dalla Torre, 1893: 216.

Myrmecocystus in Camponotinae, Camponotini: Forel, 1886f: 201.

Myrmecocystus in Camponotinae, Formicini: Forel, 1893a: 165; Emery, 1895j: 772; Forel, 1899c: 129; Wheeler, W.M. 1910g: 144; Forel, 1912i: 89; Forel, 1917: 250.

Myrmecocystus in Formicinae, Formicini: Wheeler, W.M. 1922a: 699; Chapman & Capco, 1951: 203; Wheeler, G.C. & Wheeler, J. 1970: 651; Wheeler, G.C. & Wheeler, J. 1976b: 101; Wheeler, G.C. & Wheeler, J. 1985: 258; Jaffe, 1993: 14.

Myrmecocystus in Formicinae, Acanthomyopsini: Donisthorpe, 1943f: 665.

Myrmecocystus in Formicinae, Lasius genus group: Agosti, 1991: 296.

Myrmecocystus in Formicinae, Lasiini: Ashmead, 1905b: 384; Emery, 1925b: 238; subsequent authors except the above; Smith, D.R. 1979: 1445; Bolton, 1994: 50; Bolton, 2003: 98.

Myrmecocystus as junior synonym of Cataglyphis: Roger, 1863b: 12; Mayr, 1863: 431

Myrmecocystus as genus: Forel, 1878: 372; all subsequent authors.

Junior synonyms of MYRMECOCYSTUS

Endiodioctes Snelling, R.R. 1976: 25 [as subgenus of Myrmecocystus]. Type-species: Myrmecocystus melliger, by original designation.

Taxonomic history

Endiodioctes as junior synonym of Myrmecocystus: Snelling, R.R. 1981: 403.

Eremnocystus Snelling, R.R. 1976: 92 [as subgenus of Myrmecocystus]. Type-species: Myrmecocystus creightoni, by original designation.

Taxonomic history

Eremnocystus as junior synonym of Myrmecocystus: Snelling, R.R. 1981: 403.

Genus Myrmecocystus references

Forel, 1878: 372 (diagnosis); Cresson, 1887: 255 (U.S.A. catalogue); Dalla Torre, 1893: 216 (catalogue); Wheeler, W.M. 1912d: 173 (all species key); Emery, 1925b: 238 (diagnosis, catalogue); Cole, 1942: 385 (U.S.A., Utah species key); Creighton, 1950a: 440 (North America species key); Gregg, 1963: 643 (U.S.A., Colorado species key); Snelling, R.R. 1969a: 6 (M. melliger complex, key); Snelling, R.R. 1976: 22, 25, 27, 93, 115 (diagnosis, all species revision, keys); Smith, D.R. 1979: 1445 (North America catalogue); Snelling, R.R. 1982b: 77, 83 (supplements to 1976 keys); Allred, 1982: 451 (U.S.A., Utah species key); Wheeler, G.C. & Wheeler, J. 1986g: 69 (U.S.A., Nevada species key); Dlussky & Fedoseeva, 1988: 77 (synoptic classification); Bolton, 1995a: 1050 (census); Bolton, 1995b: 274 (catalogue); Mackay & Mackay, 2002: 344 (U.S.A., New Mexico species key).

Formicinae Plagiolepidini Brachymyrmex TRUE TRUE FALSE

41 species, 17 subspecies

Brachymyrmex Mayr, 1868b: 163. Type-species: Brachymyrmex patagonicus, by monotypy.

Taxonomic history

Brachymyrmex in Camponotinae: Forel, 1878: 375 [Camponotidae]; Dalla Torre, 1893: 174.

Brachymyrmex in Camponotinae, Plagiolepidini: Forel, 1893a: 165; Forel, 1895b: 106; Emery, 1895j: 771; Forel, 1899c: 123; Wheeler, W.M. 1910g: 143.

Brachymyrmex in Formicinae, Plagiolepidini: Ashmead, 1905b: 384.

Brachymyrmex in Camponotinae, Myrmelachistini: Forel, 1912i: 89.

Brachymyrmex in Formicinae, Dimorphomyrmecini: Emery, 1925b: 40 [subtribe Brachymyrmecini]; Donisthorpe, 1943f: 628.

Brachymyrmex incertae sedis in Formicinae: Dlussky & Fedoseeva, 1988: 77.

Brachymyrmex in Formicinae, Myrmelachistini: Wheeler, W.M. 1922a: 697; Hölldobler & Wilson, 1990: 18.

Brachymyrmex in Formicinae, Pseudolasius genus group: Agosti, 1991: 296.

Brachymyrmex in Formicinae, Brachymyrmecini: Wheeler, W.M. 1929a: 12; Kempf, 1972a: 37; Wheeler, G.C. & Wheeler, J. 1985: 258; Jaffe, 1993: 13; Bolton, 1994: 50.

Brachymyrmex in Formicinae, Plagiolepidini: Bolton, 2003: 23, 101.

Junior synonym of BRACHYMYRMEX

Bryscha Santschi, 1923b: 652 [as subgenus of Brachymyrmex]. Type-species: Brachymyrmex pilipes, by original designation.

Taxonomic history

Bryscha as junior synonym of Brachymyrmex: Brown, 1973b: 179 [provisional]; Smith, D.R. 1979: 1424; all subsequent authors.

Genus Brachymyrmex references

Forel, 1878: 375 (diagnosis); André, 1882b: 214 (Europe & Algeria species); Cresson, 1887: 255 (U.S.A. catalogue); Dalla Torre, 1893: 174 (catalogue); Wheeler, W.M. 1922a: 1036 (Malagasy catalogue); Santschi, 1923b: 652 (diagnosis, all species key); Emery, 1925b: 40 (diagnosis, catalogue); Emery, 1925b: 43 (B. (Bryscha) diagnosis, catalogue); Creighton, 1950a: 356 (North America species, review); Bernard, 1967: 279 (diagnosis); Kempf, 1972a: 37 (Neotropical catalogue); Alayo, 1974: 26 (Cuba species key); Smith, D.R. 1979: 1424 (North America catalogue); Brandão, 1991: 331 (catalogue); Bolton, 1995a: 1048 (census); Bolton, 1995b: 81 (catalogue); Quirán, et al. 2004: 275 (diagnosis).

Formicinae Plagiolepidini Myrmelachista TRUE TRUE FALSE

56 species, 13 subspecies

Myrmelachista Roger, 1863a: 162. Type-species: Myrmelachista kraatzii, by monotypy.

Taxonomic history

Myrmelachista in Formicinae: Mayr, 1865: 10 [Formicidae].

Myrmelachista in Camponotinae: Forel, 1878: 367 [Camponotidae]; Dalla Torre, 1893: 174.

Myrmelachista in Camponotinae, Plagiolepidini: Forel, 1886f: 214; Forel, 1893a: 165; Forel, 1895b: 106; Emery, 1895j: 771; Forel, 1899c: 124; Wheeler, W.M. 1910g: 143.

Myrmelachista in Camponotinae, Myrmelachistini: Forel, 1912i: 89; Forel, 1917: 249.

Myrmelachista in Formicinae, Myrmelachistini: Wheeler, W.M. 1922a: 696; Emery, 1925b: 32; all subsequent authors except the following.

Myrmelachista in Formicinae, Pseudolasius genus group: Agosti, 1991: 296.

Myrmelachista in Formicinae, Plagiolepidini: Ashmead, 1905b: 384; Bolton, 2003: 23, 103.

Junior synonyms of MYRMELACHISTA

Neaphomus Menozzi, 1935c: 324 [as subgenus of Aphomomyrmex]. Type-species: Aphomomyrmex (Neaphomus) goetschi, by monotypy.

Taxonomic history

Neaphomus in Formicinae, Dimorphomyrmecini: Donisthorpe, 1943f: 673.

Neaphomus in Formicinae, Myrmelachistini: Kempf, 1972a: 152; Dlussky & Fedoseeva, 1988: 77.

Neaphomus as genus: Kempf, 1972a: 152; Dlussky & Fedoseeva, 1988: 77.

Neaphomus as junior synonym of Myrmelachista: Brown, 1973b: 183 [provisional]; Snelling & Hunt, 1976: 110; all subsequent authors except the above.

Hincksidris Donisthorpe, 1944b: 59 [as subgenus of Myrmelachista].

Taxonomic history

[Replacement name for Decamera Roger, 1863a: 166; junior homonym of Decamera Mulsant, 1842: 503 (Coleoptera).]

Hincksidris as junior synonym of Myrmelachista: Brown, 1973b: 181 [provisional]; Snelling & Hunt, 1976: 110; all subsequent authors.

Homonym replaced by Hincksidris

Decamera Roger, 1863a: 166. Type-species: Decamera nigella, by monotypy.

Taxonomic history

[Decamera Roger junior homonym of Decamera Mulsant, 1842: 503 (Coleoptera).]

Decamera in Formicinae: Mayr, 1865: 10 [Formicidae].

Decamera in Camponotinae, Myrmelachistini: Forel, 1917: 249.

Decamera as junior synonym of Myrmelachista: Forel, 1878: 376; Mayr, 1887: 525; Dalla Torre, 1893: 174.

Decamera as subgenus of Myrmelachista: Forel, 1886f: 214; Forel, 1917: 249; Wheeler, W.M. 1922a: 696; Emery, 1925b: 32; Donisthorpe, 1943f: 637.

Genus Myrmelachista references

Roger, 1863b: 11 (catalogue); Mayr, 1863: 429 (catalogue); Mayr, 1865: 10 (Myrmelachista, Decamera diagnoses); Forel, 1878: 376 (diagnosis); Mayr, 1887: 526 (species key); Dalla Torre, 1893: 174 (catalogue); Forel, 1917: 249 (synoptic classification); Wheeler, W.M. 1922a: 696 (genera, subgenera key); Emery, 1925b: 31 (diagnosis, catalogue); Emery, 1925b: 32 (M. (Decamera) diagnosis, catalogue); Wheeler, W.M. 1934g: 188 (M. (Myrmelachista) species key); Kempf, 1972a: 147, 152 (Neotropical Myrmelachista, Neaphomus catalogues); Alayo, 1974: 26 (Cuba species key); Snelling & Hunt, 1976: 110 (Chile species key); Smith, D.R. 1979: 1423 (North America catalogue); Dlussky & Fedoseeva, 1988: 77 (synoptic classification); Emery, 1925b: 31 (diagnosis, catalogue); Wheeler, G.C. & Wheeler, J. 1976b: 65 (larvae, review & synthesis); Hölldobler & Wilson, 1990: 18 (synoptic classification); Bolton, 1994: 50 (synoptic classification); Bolton, 1995a: 1051 (census); Bolton, 1995b: 275 (catalogue); Longino, 2006a: 10 (Costa Rica species, keys).

Formicinae Plagiolepidini Paratrechina TRUE TRUE FALSE

1 species

Paratrechina Motschoulsky, 1863: 13. Type-species: Paratrechina currens (junior synonym of Formica longicornis), by subsequent designation of Wheeler, W.M. 1911f: 170.

Taxonomic history

Paratrechina in Formicinae, Acanthomyopsini: Donisthorpe, 1943f: 682.

Paratrechina in Formicinae, Prenolepidini: Chapman & Capco, 1951: 218; Hölldobler & Wilson, 1990: 18.

Paratrechina in Formicinae, Myrmelachistinini: Wheeler, G.C. & Wheeler, J. 1970: 652; Wheeler, G.C. & Wheeler, J. 1976b: 101.

Paratrechina in Formicinae, Brachymyrmecini: Wheeler, G.C. & Wheeler, J. 1985: 258.

Paratrechina in Formicinae, Pseudolasius genus group: Agosti, 1991: 296.

Paratrechina in Formicinae, Lasiini: Emery, 1925b: 216; Stitz, 1939: 379; Bernard, 1953b: 257; Kempf, 1972a: 182; Smith, D.R. 1979: 1442; Dlussky & Fedoseeva, 1988: 77; Jaffe, 1993: 14; Bolton, 1994: 50.

Paratrechina in Formicinae, Plagiolepidini: Bolton, 2003: 23, 104.

Paratrechina as junior synonym of Prenolepis: Dalla Torre, 1893: 177; Wheeler, W.M. 1922a: 940.

Paratrechina as genus: Emery, 1925b: 216; all subsequent authors.

Genus Paratrechina references

References prior to 2010 also include Nylanderia and Paraparatrechina.

André, 1882b: 203 (Europe & Algeria species key); Forel, 1894c: 406 (India & Sri Lanka species key); Wheeler, W.M. 1922a: 941, 1037 (Afrotropical, Malagasy P. (Nylanderia) catalogues); Emery, 1925b: 216 (diagnosis, catalogue); Emery, 1925b: 217 (P. (Nylanderia) diagnosis, catalogue); Emery, 1925b: 224 (Andragnathus diagnosis, catalogue); Finzi, 1936: 191 (Egypt species key); Buren, 1944a: 295 (U.S.A., Iowa species key); Creighton, 1950a: 404 (North America species key); Chapman & Capco, 1951: 197, 214, 218 (Asia Andragnathus, Nylanderia, Paratrechina checklists); Kusnezov, 1956: 32 (Argentina species key); Bernard, 1967: 347 (diagnosis, Western Europe species key); Kempf, 1972a: 166, 182 (Neotropical Nylanderia, Paratrechina catalogues); Alayo, 1974: 26 (Cuba species key); Arnol'di & Dlussky, 1978: 556 (former European U.S.S.R. species key); Smith, D.R. 1979: 1442 (North America, catalogue); Trager, 1984b: 65 (Nearctic species revision, key); Wheeler, G.C. & Wheeler, J. 1986g: 64 (U.S.A., Nevada species key); Collingwood, 1985: 298 (Saudi Arabia species key); Taylor & Brown, D.R. 1985: 129 (Australia catalogue); Taylor, 1987a: 52 (Australia, New Caledonia & New Zealand checklist); Brandão, 1991: 365 (Neotropical catalogue); Morisita, Kubota, et al. 1991: 20 (Japan species key); Bolton, 1995a: 1051 (census); Bolton, 1995b: 312 (catalogue); Wu, J. & Wang, 1995: 150 (China species key); Collingwood & Agosti, 1996: 369 (Saudi Arabia species key); Terayama, 1999c: 50 (Japan species key); Shattuck, 1999: 104 (Australia synopsis); Zhou, 2001b: 177 (China, Guangxi species key); Mackay & Mackay, 2002: 354 (U.S.A., New Mexico species key); Terayama, 2009: 210 (Taiwan species key); Heterick, 2009: 104 (south-western Australia species key); LaPolla, Brady & Shattuck, 2010a: 127 (diagnosis).

Formicinae Plagiolepidini Plagiolepis TRUE TRUE FALSE

Extant: 57 species, 19 subspecies

Fossil: 10 species

Plagiolepis Mayr, 1861: 42. Type-species: Formica pygmaea, by monotypy.

Taxonomic history

Plagiolepis in Formicinae: Mayr, 1861: 42 [Formicinae]; Mayr, 1862: 652 [Formicidae]; Mayr, 1865: 7 [Formicidae]; Mayr, 1868c: 36 [Formicidae].

Plagiolepis in Camponotinae: Forel, 1878: 378 [Camponotidae]; Emery & Forel, 1879a: 453 [Camponotidae]; Dalla Torre, 1893: 172.

Plagiolepis in Camponotinae, Plagiolepidini: Forel, 1886f: 212; Forel, 1893a: 165; Emery, 1895j: 771; Wheeler, W.M. 1910g: 143; Forel, 1912i: 88; Wheeler, W.M. 1915h: 100; Forel, 1917: 249; Arnold, 1920a: 578.

Plagiolepis in Formicinae, Plagiolepidini: Ashmead, 1905b: 384; Bondroit, 1918: 18; Wheeler, W.M. 1922a: 696; Emery, 1925b: 19; Santschi, 1926a: 14; all subsequent authors except the one below; Bolton, 2003: 23, 105.

Plagiolepis in Formicinae, Pseudolasius genus group: Agosti, 1991: 296.

Junior synonyms of PLAGIOLEPIS

†Rhopalomyrmex Mayr, 1868c: 41. Type-species: †Rhopalomyrmex pygmaeus, by monotypy.

Taxonomic history

†Rhopalomyrmex in Formicinae: Mayr, 1868c: 41 [Formicidae].

†Rhopalomyrmex in Camponotinae: Forel, 1878: 376 [Camponotidae]; Dalla Torre, 1893: 175.

†Rhopalomyrmex in Camponotinae, Myrmelachistini: Forel, 1912i: 89.

†Rhopalomyrmex in Camponotinae, Plagiolepidini: Wheeler, W.M. 1910g: 143; Wheeler, W.M. 1915h: 103.

†Rhopalomyrmex in Formicinae, Plagiolepidini: Donisthorpe, 1943g: 724; Bolton, 1994: 51.

†Rhopalomyrmex as junior synonym of Plagiolepis: Dlussky, 1997: 624; Bolton, 2003: 105.

Anacantholepis Santschi, 1914e: 36 [as subgenus of Plagiolepis]. Type-species: Plagiolepis (Anacantholepis) decora, by original designation.

Taxonomic history

Anacantholepis as genus: Chapman & Capco, 1951: 212.

Anacantholepis as subgenus of Plagiolepis: Santschi, 1914e: 36 Forel, 1917: 249; Arnold, 1922: 581; Wheeler, W.M. 1922a: 696; Emery, 1925b: 22; Bolton, 1994: 51.

Anacantholepis as junior synonym of Plagiolepis: Brown, 1973b: 178 [provisional]; Bolton, 2003: 105, 268.

Aporomyrmex Faber, 1969: 52. Type-species: Aporomyrmex ampeloni, by original designation.

Taxonomic history

Aporomyrmex in Formicinae, Plagiolepidini: Faber, 1969: 52 (by implication).

Aporomyrmex as junior synonym of Plagiolepis: Brown, 1973b: 178 [provisional]; Bolton, 1994: 51; all subsequent authors.

Paraplagiolepis Faber, 1969: 65 [as subgenus of Plagiolepis]. Type-species: Plagiolepis xene, by monotypy.

Taxonomic history

Paraplagiolepis as junior synonym of Plagiolepis: Brown, 1973b: 183 [provisional]; Bolton, 1994: 51; all subsequent authors.

Genus Plagiolepis references

Roger, 1863b: 11 (catalogue); Mayr, 1863: 442 (catalogue); Mayr, 1865: 7 (diagnosis); Mayr, 1867a: 73 (diagnosis); Forel, 1878: 376, 378 (†Rhopalomyrmex, Plagiolepis diagnoses); André, 1882b: 208 (Europe & Algeria species key); Dalla Torre, 1893: 172, 175 (Plagiolepis, †Rhopalomyrmex, catalogue); Forel, 1894c: 414 (India & Sri Lanka species key); Bingham, 1903: 320 (India, Sri Lanka & Burma species key); Arnold, 1920a: 578 (diagnosis, subgenera key); Arnold, 1922: 579 (South Africa species key); Wheeler, W.M. 1922a: 211, 696, 928, 1035  (diagnosis, subgenera key, Afrotropical, Malagasy catalogues); Emery, 1925b: 19 (diagnosis, catalogue); Emery, 1925b: 22 (P. (Anacantholepis) diagnosis, catalogue); Stitz, 1939: 231 (Germany species key); Kratochvíl, 1941: 98 (Central Europe species key); Novák & Sadil, 1941: 98 (Central Europe species key); Chapman & Capco, 1951: 212, 213 (Asia Anacantholepis, Plagiolepis checklists); Bernard, 1967: 272 (diagnosis, Western Europe species key); Kempf, 1972a: 205 (Neotropical catalogue); Kutter, 1977c: 185 (Switzerland species key); Collingwood, 1978: 89 (Iberian Peninsula species key); Smith, D.R. 1979: 1422 (North America, catalogue); Gösswald, 1985: 264 (Germany species key); Collingwood, 1985: 297 (Saudi Arabia species key); Taylor & Brown, D.R. 1985: 130 (Australia catalogue); Taylor, 1987a: 55 (Australia, New Caledonia checklist); Agosti & Collingwood, 1987b: 280 (Balkans species key); Radchenko, 1989b: 153 (European former U.S.S.R. species key); Dlussky, Soyunov & Zabelin, 1990: 161 (Turkmenistan species key); Morisita, Kubota, et al. 1991: 16 (Japan species key); Atanasov & Dlussky, 1992: 201 (Bulgaria species key); Arakelian, 1994: 78 (Armenia species key); Bolton, 1994: 51 (synoptic classification); Bolton, 1995a: 1052 (census); Bolton, 1995b: 334 (catalogue); Wu, J. & Wang, 1995: 130 (China species key); Radchenko, 1996a: 178 (Central & Southern Palaearctic species key); Collingwood & Agosti, 1996: 362 (Saudi Arabia species key); Seifert, 1996b: 178 (Central Europe species key); Collingwood & Prince, 1998: 22 (Portugal species key); Shattuck, 1999: 106 (Australia synopsis); Seifert, 2007: 158 (North and Central Europe species key); Terayama, 2009: 213 (Taiwan species key); Heterick, 2009: 106 (south-western Australia species key); Dlussky, 2010: 72 (Eocene European amber species key).

Formicinae Plagiolepidini Prenolepis TRUE TRUE FALSE

Extant: 19 species, 6 subspecies

Fossil: 1 species

Prenolepis Mayr, 1861: 52. Type-species: Tapinoma nitens, by subsequent designation of Bingham, 1903: 325.

Taxonomic history

[Type-species not Formica imparis, unjustified subsequent designation by Emery, 1906b: 134, repeated in Wheeler, W.M. 1911f: 171 and Wheeler, W.M. 1922a: 940.]

Prenolepis in Formicinae: Mayr, 1861: 52 [Formicidae]; Mayr, 1862: 652 [Formicidae]; Mayr, 1865: 7 [Formicidae]; Mayr, 1868c: 32 [Formicidae].

Prenolepis in Camponotinae: Forel, 1878: 377 [Camponotidae]; Emery & Forel, 1879a: 453 [Camponotidae]; Dalla Torre, 1893: 177.

Prenolepis in Camponotinae, Camponotini: Forel, 1886f: 209.

Prenolepis in Camponotinae, Formicini: Forel, 1893a: 165; Forel, 1895b: 103; Emery, 1895j: 772; Forel, 1899c: 125; Wheeler, W.M. 1910g: 143.

Prenolepis in Formicinae, Plagiolepidini: Ashmead, 1905b: 384.

Prenolepis in Camponotinae, Prenolepidini: Forel, 1912i: 89; Wheeler, W.M. 1915h: 117; Forel, 1917: 249; Arnold, 1922: 605.

Prenolepis in Formicinae, Prenolepidini: Wheeler, W.M. 1922a: 697; Chapman & Capco, 1951: 219; Hölldobler & Wilson, 1990: 18.

Prenolepis in Formicinae, Acanthomyopsini: Donisthorpe, 1943f: 685.

Prenolepis in Formicinae, Brachymyrmecini: Wheeler, G.C. & Wheeler, J. 1970: 651; Wheeler, G.C. & Wheeler, J. 1976b: 101; Wheeler, G.C. & Wheeler, J. 1985: 258.

Prenolepis in Formicinae, Lasiini: Emery, 1925b: 224; Stitz, 1939: 379, Kempf, 1972a: 209; Smith, D.R. 1979: 1444; Dlussky & Fedoseeva, 1988: 77; Jaffe, 1993: 14; Bolton, 1994: 50.

Prenolepis in Formicinae, Lasius genus group: Agosti, 1991: 296.

Prenolepis in Formicinae, Plagiolepidini: Bolton, 2003: 23, 105.

Genus Prenolepis references

Roger, 1863b: 10 (catalogue); Mayr, 1863: 451 (catalogue); Mayr, 1865: 7 (diagnosis); Mayr, 1867a: 71 (diagnosis); Mayr, 1870b: 947 (all species key); Forel, 1878: 377 (diagnosis); André, 1882b: 203 (Europe & Algeria species key); Cresson, 1887: 257 (U.S.A. catalogue); Dalla Torre, 1893: 177 (catalogue); Bingham, 1903: 326 (India, Sri Lanka & Burma species key); Emery, 1910a: 127 (Palaearctic species key); Arnold, 1922: 605 (diagnosis); Wheeler, W.M. 1922a: 216, 697, 940 (diagnosis, subgenera key, Afrotropical catalogue); Emery, 1925b: 224 (diagnosis, catalogue); Creighton, 1950a: 410 (North America species review); Chapman & Capco, 1951: 219 (Asia checklist); Kempf, 1972a: 209 (Neotropical catalogue); Smith, D.R. 1979: 1442 (North America catalogue); Bolton, 1994: 50 (synoptic classification); Bolton, 1995a: 1052 (census); Bolton, 1995b: 363 (catalogue); Wu, J. & Wang, 1995: 137 (China species key); Xu, 1995c: 338 (Southern & Eastern Asia species key); Fontenla Rizo, 2000: 81 (Antilles Is species key); Zhou, 2001b: 170 (China, Guangxi species key); Wang, W. & Wu, 2007: 721 (China species key); LaPolla, Brady & Shattuck, 2010a: 129 (diagnosis).

Formicinae Camponotini Camponotus TRUE TRUE FALSE

Extant: 1,058 species, 495 subspecies

Fossil: 31 species

Camponotus Mayr, 1861: 35. Type-species: Formica ligniperda, by subsequent designation of Bingham, 1903: 347.

Taxonomic history

[Type-species not Formica herculeana, unjustified subsequent designation by Forel, 1914a: 259, repeated in Arnold, 1922: 612.]

Camponotus in Formicinae: Mayr, 1861: 35 [Formicidae]; Mayr, 1862: 652 (in key) [Formicidae]; Mayr, 1865: 6 [Formicidae]; Mayr, 1868c: 26 [Formicidae].

Camponotus in Camponotinae: Forel, 1878: 367 [Camponotidae]; Emery & Forel, 1879a: 447 [Camponotidae]; Forel, 1892j: 223 [Camponotidae]; Dalla Torre, 1893: 171.

Camponotus in Camponotinae, Camponotini: Forel, 1886f: 141; Forel, 1893a: 165; Forel, 1895b: 101; Emery, 1895j: 772; Forel, 1899c: 130; Ashmead, 1905b: 384; Wheeler, W.M. 1910g: 144; Forel, 1912i: 89; Wheeler, W.M. 1915h: 138; Forel, 1917: 250; Arnold, 1922: 610.

Camponotus in Formicinae, Camponotini: Bondroit, 1918: 65; Wheeler, W.M. 1922a: 703; Emery, 1925b: 59; all subsequent authors except the following.

Camponotus in Formicinae, Formica genus group: Agosti, 1991: 295.

Camponotus as subgenus of Formica: Smith, F. 1871a: 306.

Camponotus as genus: all authors except the entry above.

Junior synonym of CAMPONOTUS

†Paleosminthurus Pierce & Gibron, 1962: 146. Type-species: †Paleosminthurus juliae, by monotypy.

Taxonomic history

†Paleosminthurus in Collembola, family †Paleosminthuridae: Pierce & Gibron, 1962: 146.

†Paleosminthurus incertae sedis in Formicidae: Najt, 1987: 152.

†Paleosminthurus incertae sedis in Formicinae: Hölldobler & Wilson, 1990: 18.

†Paleosminthurus in †Paleosmithurinae: Bolton, 1994: 187 [provisional subfamily status]; Bolton, 1995b: 14, 311; Bolton, 2003: 76, 263.

†Paleosminthurus as junior synonym of Camponotus: Snelling (pers. comm., 2004). [Synonymy remained unpublished at his death in 2008.]

†Shanwangella Zhang, 1989: 307. Type-species: †Shanwangella palaeoptera, by original designation.

Taxonomic history

†Shanwangella as junior synonym of Camponotus: Hong & Wu, 2000: 20.

Subgenera of CAMPONOTUS include the nominal plus the following.

All the subgenera were given as provisional junior synonyms of Camponotus by Brown, 1973b: 179 - 185. Brown's list was repeated in Hölldobler & Wilson, 1990: 18 but with all subgenera listed as junior synonyms; subgeneric status was reinstated in Bolton, 1994: 50. These consecutive entries are not repeated individually below.

Subgenus CAMPONOTUS (COLOBOPSIS)

Colobopsis Mayr, 1861: 38. Type-species: Formica truncata, by subsequent designation of Bingham, 1903: 342.

Taxonomic history

Colobopsis in Formicinae: Mayr, 1861: 38 [Formicidae]; Mayr, 1862: 652 (in key) [Formicidae]; Mayr, 1865: 7 [Formicidae].

Colobopsis in Camponotinae: Forel, 1878: 368 [Camponotidae]; Emery & Forel, 1897: 449 [Camponotidae].

Colobopsis in Camponotinae, Camponotini: Forel, 1886f: 193; Emery, 1895j: 772; Ashmead, 1905b: 384.

Colobopsis in Formicinae, Camponotini: Bondroit, 1918: 65; Emery, 1925b: 144; all subsequent authors.

Colobopsis as genus: Mayr, 1861: 38; Mayr, 1862: 652; Mayr, 1865: 7; Mayr, 1870b: 940; Forel, 1878: 368; Forel, 1879a: 125; Emery & Forel, 1879a: 449; André, 1882b: 159; Bingham, 1903: 342; Forel, 1886f: 193; Ashmead, 1905b: 384; Bondroit, 1918: 66; Jaffe, 1993: 14.

Colobopsis as subgenus of Camponotus: Emery, 1889b: 517; Dalla Torre, 1893: 219; Forel, 1893a: 165; Forel, 1893b: 435; Emery, 1895j: 772; Wheeler, W.M. 1904b: 139; Wheeler, W.M. 1910g: 144; Forel, 1912i: 90; Forel, 1914a: 263; Forel, 1917: 251; Emery, 1920b: 247; Arnold, 1922: 613; Wheeler, W.M. 1922a: 708; Emery, 1925b: 144; Donisthorpe, 1943f: 634; Creighton, 1950a: 390; Chapman & Capco, 1951: 222; Kempf, 1972a: 42; Smith, D.R., 1979: 1433; Bolton, 1994: 50.

Junior synonyms of CAMPONOTUS (COLOBOPSIS)

Condylomyrma Santschi, 1928c: 72 [as subgenus of Camponotus]. Type-species: Camponotus (Condylomyrma) bryani, by monotypy.

Taxonomic history

Condylomyrma as subgenus of Camponotus: Santschi, 1928c: 72.

Condylomyrma as junior synonym of Colobopsis: Wheeler, W.M. 1934e: 422 (in text).

[Campylomyrma Wheeler, W.M. 1934e: 421 (in text), incorrect subsequent spelling.]

Dolophra Wu, J. & Wang, 1994: 35. Type-species: Dolophra politae, by original designation.

Taxonomic history

Dolophra as genus: Wu, J. & Wang, 1994: 35; Wu, J. & Wang, 1995: 158.

Dolophra as junior synonym of Camponotus: Bolton, 1995b: 27.

Dolophra as junior synonym of Camponotus (Colobopsis): Bolton, 2003: 113, 268.

Subgenus CAMPONOTUS (MAYRIA)

Mayria Forel, 1878: 369. Type-species: Mayria madagascarensis (junior secondary homonym in Camponotus, replaced by Camponotus repens), by monotypy.

Taxonomic history

Mayria in Camponotinae: Forel, 1878: 369 [Camponotidae]; Dalla Torre, 1893: 219.

Mayria in Camponotinae, Camponotini: Forel, 1893a: 165; Emery, 1895j: 772; Ashmead, 1905b: 384; Wheeler, W.M. 1910g: 144.

Mayria as subgenus of Camponotus: Forel, 1894e: 227; Forel, 1914a: 262; Forel, 1917: 251; Wheeler, W.M. 1922a: 706; Emery, 1925b: 121; all subsequent authors.

Junior synonym of CAMPONOTUS (MAYRIA)

Myrmosaga Forel, 1912i: 92 [as subgenus of Camponotus]. Type-species: Camponotus kelleri, by subsequent designation of Wheeler, W.M. 1913a: 81.

Taxonomic history

[Type-species not Camponotus quadrimaculatus, unjustified subsequent designation by Forel, 1914a: 260; repeated in Arnold, 1922: 613.]

Myrmosaga as subgenus of Camponotus Forel, 1912i: 92; Forel, 1914a: 260; Forel, 1917: 250; Emery, 1920b: 241; Arnold, 1922: 613; Wheeler, W.M. 1922a: 706; all subsequent authors.

Myrmosaga as junior synonym of Mayria: Emery, 1925b: 121.

Subgenus CAMPONOTUS (RHINOMYRMEX)

Rhinomyrmex Forel, 1886f: 192. Type-species: Rhinomyrmex klaesii, by monotypy.

Taxonomic history

Rhinomyrmex in Camponotinae: Forel, 1886f: 192 [Camponotii]; Dalla Torre, 1893: 219.

Rhinomyrmex in Camponotinae, Camponotini: Forel, 1893a: 165; Emery, 1895j: 772; Ashmead, 1905b: 384; Wheeler, W.M. 1910g: 144.

Rhinomyrmex as genus: Forel, 1886f: 192; Forel, 1893a: 165; Dalla Torre, 1893: 219; Emery, 1895j: 772; Wheeler, W.M. 1910g: 144.

Rhinomyrmex as subgenus of Camponotus: Emery, 1896d: 374; Forel, 1914a: 264; Forel, 1917: 251; Emery, 1920b: 247; Wheeler, W.M. 1922a: 708; Emery, 1925b: 142; all subsequent authors.

Subgenus CAMPONOTUS (DENDROMYRMEX)

Dendromyrmex Emery, 1895j: 772. Type-species: Formica chartifex, by subsequent designation of Wheeler, W.M. 1911f: 161.

Taxonomic history

Dendromyrmex in Camponotinae, Camponotini: Emery, 1895j: 772; Ashmead, 1905b: 384; Forel, 1912i: 89; Wheeler, W.M. 1910g: 144; Forel, 1917: 250.

Dendromyrmex in Formicinae, Camponotini: Wheeler, W.M. 1922a: 700; Emery, 1925b: 172; all subsequent authors except the following.

Dendromyrmex in Formicinae, Formica genus group: Agosti, 1991: 295.

Dendromyrmex as subgenus of Camponotus: Fernández, 2002: 51.

Subgenus CAMPONOTUS (DINOMYRMEX)

Dinomyrmex Ashmead, 1905b: 384. Type-species: Formica gigas, by original designation.

Taxonomic history

Dinomyrmex in Camponotinae, Camponotini: Ashmead, 1905b: 384.

Dinomyrmex as genus: Ashmead, 1905b: 384.

Dinomyrmex as subgenus of Camponotus: Forel, 1913b: 350; Forel, 1914a: 259; Forel, 1917: 250; Emery, 1920b: 236; Arnold, 1922: 612; Wheeler, W.M. 1922a: 704; Emery, 1925b: 69; all subsequent authors.

Junior synonym of CAMPONOTUS (DINOMYRMEX)

Myrmogigas Forel, 1912i: 91 [as subgenus of Camponotus]. Type-species: Formica gigas, by subsequent designation of Wheeler, W.M. 1913a: 81.

Taxonomic history

Myrmogigas as subgenus of Camponotus: Forel, 1912i: 91.

Myrmogigas as junior synonym of Dinomyrmex: Forel, 1913b: 350 (in text); Forel, 1914a: 259; Wheeler, W.M. 1922a: 704. [Dinomyrmex and Myrmogigas share the same type-species, synonymy is therefore absolute.]

Subgenus CAMPONOTUS (TANAEMYRMEX)

Tanaemyrmex Ashmead, 1905b: 384. Type-species: Formica longipes (junior primary homonym in Formica, replaced by Camponotus etiolipes), by original designation.

Taxonomic history

Tanaemyrmex in Camponotinae, Camponotini: Ashmead, 1905b: 384.

Tanaemyrmex as genus: Ashmead, 1905b: 384.

Tanaemyrmex as subgenus of Camponotus: Emery, 1925b: 75; all subsequent authors.

Junior synonym of CAMPONOTUS (TANAEMYRMEX)

Myrmoturba Forel, 1912i: 91 [as subgenus of Camponotus]. Type-species: Formica maculata, by subsequent designation of Wheeler, W.M. 1913a: 82.

Taxonomic history

Myrmoturba as subgenus of Camponotus: Forel, 1912i: 91; Forel, 1914a: 259; Forel, 1917: 250; Emery, 1920b: 235; Arnold, 1922: 612; Wheeler, W.M. 1922a: 704.

Myrmoturba as junior synonym of Tanaemyrmex: Emery, 1925b: 75; all subsequent authors.

Subgenus CAMPONOTUS (ORTHONOTOMYRMEX)

Orthonotomyrmex Ashmead, 1906: 31.

Taxonomic history

[Replacement name for Orthonotus Ashmead, 1905b: 384; junior homonym of Orthonotus Westwood, in Stephens, 1829: 344 (Hemiptera). Type-species not Formica lateralis, unjustified subsequent designation by Forel, 1914a: 264; repeated in Arnold, 1922: 613.]

Orthonotomyrmex as subgenus of Camponotus: Forel, 1913i: 350 (in text); Forel, 1914a: 264; Forel, 1917: 251; Emery, 1920b: 244; Arnold, 1922: 613; Wheeler, W.M. 1922a: 707; Emery, 1925b: 124; all subsequent authors.

Homonym replaced by ORTHONOTOMYRMEX

Orthonotus Ashmead, 1905b: 384. Type-species: Formica sericea, by original designation.

Taxonomic history

[Junior homonym of Orthonotus Westwood, in Stephens, 1829: 344 (Hemiptera).]

Orthonotus in Camponotinae, Camponotini: Ashmead, 1905b: 384.

Subgenus CAMPONOTUS (MYRMAMBLYS)

Myrmamblys Forel, 1912i: 90 [as subgenus of Camponotus]. Type-species: Camponotus reticulatus, by subsequent designation of Wheeler, W.M. 1913a: 80.

Taxonomic history

[Type-species not Camponotus fastigatus, unjustified subsequent designation by Forel, 1914a: 263; repeated in Arnold, 1922: 613.]

Myrmamblys as subgenus of Camponotus: Forel, 1912i: 90; Forel, 1914a: 263; Forel, 1917: 251; Emery, 1920b: 248; Arnold, 1922: 613; Wheeler, W.M. 1922a: 708; Emery, 1925b: 136; Santschi, 1926c: 601; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOTHRIX)

Myrmothrix Forel, 1912i: 91 [as subgenus of Camponotus]. Type-species: Formica abdominalis (junior primary homonym; Formica atriceps is first available replacement name), by subsequent designation of Wheeler, W.M. 1913a: 81.

Taxonomic history

[Type-species not Formica rufipes, unjustified subsequent designation by Forel, 1914a: 260.]

Myrmothrix as subgenus of Camponotus: Forel, 1912i: 91; Forel, 1914a: 260; Forel, 1917: 250; Emery, 1920b: 236; Emery, 1925b: 107; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOSERICUS)

Myrmosericus Forel, 1912i: 91 [as subgenus of Camponotus]. Type-species: Formica rufoglauca, by subsequent designation of Wheeler, W.M. 1913a: 81.

Taxonomic history

Myrmosericus as subgenus of Camponotus: Forel, 1912i: 91; Forel, 1914a: 259; Forel, 1917: 250; Emery, 1920b: 236; Arnold, 1922: 613; Wheeler, W.M. 1922a: 704; Emery, 1925b: 104; all subsequent authors.

[See note under Myrmosaulus.]

Subgenus CAMPONOTUS (MYRMOPHYMA)

Myrmophyma Forel, 1912i: 91 [as subgenus of Camponotus]. Type-species: Camponotus capito, by subsequent designation of Wheeler, W.M. 1913a: 81.

Taxonomic history

[Type-species not Camponotus quadrisectus, unjustified subsequent designation by Forel, 1914a: 261; repeated in Arnold, 1922: 612.]

Myrmophyma as subgenus of Camponotus: Forel, 1912i: 91; Forel, 1914a: 261; Forel, 1917: 250; Emery, 1920b: 239; Arnold, 1922: 612; Wheeler, W.M. 1922a: 706; Emery, 1925b: 109; all subsequent authors.

Junior synonym of CAMPONOTUS (MYRMOPHYMA)

Myrmocamelus Forel, 1914a: 261 [as subgenus of Camponotus]. Type-species: Formica ephippium, by original designation.

Taxonomic history

[Type-species not Camponotus gambeyi, unjustified subsequent designation by Forel, 1922: 101. Myrmocamelus also described as new by Forel, 1915b: 102.]

Myrmocamelus as subgenus of Camponotus: Forel, 1914a: 261; Forel, 1915b: 102; Forel, 1917: 250; Forel, 1922: 101.

Myrmocamelus as junior synonym of Myrmophyma: Emery, 1920b: 257; Wheeler, W.M. 1921a: 18; Emery, 1925b: 109.

Subgenus CAMPONOTUS (MYRMOTREMA)

Myrmotrema Forel, 1912i: 91 [as subgenus of Camponotus]. Type-species: Camponotus foraminosus, by subsequent designation of Wheeler, W.M. 1913a: 81.

Taxonomic history

Myrmotrema as subgenus of Camponotus: Forel, 1912i: 91; Forel, 1914a: 262; Forel, 1917: 250; Emery, 1920b: 245; Arnold, 1922: 613; Wheeler, W.M. 1922a: 707; Emery, 1925b: 130; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOBRACHYS)

Myrmobrachys Forel, 1912i: 91 [as subgenus of Camponotus]. Type-species: Formica senex, by subsequent designation of Wheeler, W.M. 1913a: 81.

Taxonomic history

Myrmobrachys as subgenus of Camponotus: Forel, 1912i: 91; Forel, 1914a: 262; Forel, 1917: 251; Emery, 1920b: 251; Wheeler, W.M. 1922a: 709; Emery, 1925b: 161; all subsequent authors.

Subgenus CAMPONOTUS (MYRMENTOMA)

Myrmentoma Forel, 1912i: 92 [as subgenus of Camponotus]. Type-species: Formica lateralis, by subsequent designation of Wheeler, W.M. 1913a: 80.

Taxonomic history

Myrmentoma as junior synonym of Orthonotomyrmex: Forel, 1913b: 350 (in text); Forel, 1914a: 264; Wheeler, W.M. 1922a: 970.

Myrmentoma as subgenus of Camponotus: Forel, 1912i: 92; Emery, 1920b: 243; Wheeler, W.M. 1922a: 707; Emery, 1925a: 62; Emery, 1925b: 116; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOSPHINCTA)

Myrmosphincta Forel, 1912i: 92 [as subgenus of Camponotus]. Type-species: Formica sexguttata, by subsequent designation of Wheeler, W.M. 1913a: 81.

Taxonomic history

[Type-species not Formica cinerascens, unjustified subsequent designation by Forel, 1914a: 265; repeated in Arnold, 1922: 614.]

Myrmosphincta as subgenus of Camponotus: Forel, 1912i: 92; Forel, 1914a: 265; Forel, 1917: 251; Emery, 1920b: 238; Arnold, 1922: 614; Wheeler, W.M. 1922a: 708; Emery, 1925b: 151; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOTARSUS)

Myrmotarsus Forel, 1912i: 92 [as subgenus of Camponotus]. Type-species: Formica mistura, by subsequent designation of Wheeler, W.M. 1913a: 81.

Taxonomic history

[Type-species not Formica irritabilis, unjustified subsequent designation by Forel, 1914a: 260.]

Myrmotarsus as subgenus of Camponotus: Forel, 1912i: 92; Forel, 1914a: 260; Forel, 1917: 250; Emery, 1920b: 238; Wheeler, W.M. 1922a: 705; Emery, 1925b: 134; all subsequent authors.

Subgenus CAMPONOTUS (MYRMEPOMIS)

Myrmepomis Forel, 1912i: 92 [as subgenus of Camponotus]. Type-species: Formica sericeiventris, by subsequent designation of Wheeler, W.M. 1913a: 81.

Taxonomic history

[Type-species not Formica fulvopilosa, unjustified subsequent designation by Forel, 1914a: 265; repeated in Arnold, 1922: 614.]

Myrmepomis as subgenus of Camponotus: Forel, 1912i: 92; Forel, 1914a: 265; Forel, 1917: 251; Emery, 1920b: 245; Arnold, 1922: 614; Wheeler, W.M. 1922a: 705; Emery, 1925b: 170; all subsequent authors.

Junior synonym of CAMPONOTUS (MYRMEPOMIS)

Myrmolophus Emery, 1920b: 237 [as subgenus of Camponotus]. Type-species: Formica sericeiventris, by original designation.

Taxonomic history

Myrmolophus as subgenus of Camponotus: Emery, 1920b: 237.

Myrmolophus as junior synonym of Myrmepomis: Wheeler, W.M. 1921a: 17; Wheeler, W.M. 1922a: 705; Emery, 1925b: 170. [Myrmepomis and Myrmolophus share the same type-species, synonymy is therefore absolute.]

Subgenus CAMPONOTUS (MYRMOGONIA)

Myrmogonia Forel, 1912i: 92 [as subgenus of Camponotus]. Type-species: Camponotus laminatus, by subsequent designation of Wheeler, W.M. 1913a: 81.

Taxonomic history

Myrmogonia as subgenus of Camponotus: Forel, 1912i: 92; Forel, 1914a: 261; Forel, 1917: 250; Emery, 1920b: 240; Wheeler, W.M. 1922a: 706; Emery, 1925b: 143; all subsequent authors.

Subgenus CAMPONOTUS (MYRMEURYNOTA)

Myrmeurynota Forel, 1912i: 92 [as subgenus of Camponotus]. Type-species: Camponotus eurynotus, by subsequent designation of Wheeler, W.M. 1913a: 81.

Taxonomic history

[Type-species not Camponotus gilviventris, unjustified subsequent designation by Forel, 1914a: 266.]

Myrmeurynota as subgenus of Camponotus: Forel, 1912i: 92; Forel, 1914a: 266; Forel, 1917: 251; Emery, 1920b: 251; Wheeler, W.M. 1922a: 710; Emery, 1925b: 167; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOPSAMMA)

Myrmopsamma Forel, 1914a: 261 [as subgenus of Camponotus]. Type-species: Camponotus mystaceus, by original designation.

Taxonomic history

Myrmopsamma as subgenus of Camponotus: Forel, 1914a: 261; Forel, 1917: 250; Emery, 1920b: 246; Arnold, 1922: 612; Wheeler, W.M. 1922a: 708; Emery, 1925b: 70; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOMALIS)

Myrmomalis Forel, 1914a: 263 [as subgenus of Camponotus]. Type-species: Camponotus depressus, by original designation.

Taxonomic history

Myrmomalis as subgenus of Camponotus: Forel, 1914a: 263; Forel, 1917: 251; Emery, 1920b: 251; Wheeler, W.M. 1922a: 710; Emery, 1925b: 169; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOPLATYS)

Myrmoplatys Forel, 1916: 460 [as subgenus of Camponotus]. Type-species: Camponotus korthalsiae, by original designation.

Taxonomic history

Myrmoplatys as genus: Wheeler, W.M. 1936e: 219 (in text); Donisthorpe, 1943f: 670.

Myrmoplatys as subgenus of Camponotus: Forel, 1916: 460; Forel, 1917: 251; Emery, 1920b: 238; Wheeler, W.M. 1922a: 705; Emery, 1925b: 135; Bolton, 1994: 50.

Subgenus CAMPONOTUS (MYRMAPHAENUS)

Myrmaphaenus Emery, 1920b: 237 [as subgenus of Camponotus]. Type-species: Camponotus leydigi, by original designation.

Taxonomic history

Myrmaphaenus as subgenus of Camponotus: Emery, 1920b: 237; Wheeler, W.M. 1922a: 705; Emery, 1925b: 152; all subsequent authors.

Junior synonyms of CAMPONOTUS (MYRMAPHAENUS)

Paracolobopsis Emery, 1920b: 249 [as subgenus of Camponotus]. Type-species: Camponotus salvini, by original designation.

Taxonomic history

Paracolobopsis as subgenus of Camponotus: Emery, 1920b: 249; Wheeler, W.M. 1922a: 709.

Paracolobopsis as junior synonym of Myrmaphaenus: Emery, 1925b: 152; Kempf, 1972a: 44.

Neomyrmamblys Wheeler, W.M. 1921a: 19 [as subgenus of Camponotus]. Type-species: Camponotus fastigatus, by subsequent designation of Santschi, 1921f: 311.

Taxonomic history

Neomyrmamblys as subgenus of Camponotus: Wheeler, W.M. 1921a: 19; Wheeler, W.M. 1922a: 708.

Neomyrmamblys as junior synonym of Myrmaphaenus: Emery, 1925b: 152; Kempf, 1972a: 44.

Subgenus CAMPONOTUS (MYRMONESITES)

Myrmonesites Emery, 1920b: 242 [as subgenus of Camponotus]. Type-species: Camponotus putatus, by original designation.

Taxonomic history

Myrmonesites as subgenus of Camponotus: Emery, 1920b: 242; Wheeler, W.M. 1922a: 706; Emery, 1925b: 123.

[Myrmensites Donisthorpe, 1943f: 666, incorrect subsequent spelling.]

Subgenus CAMPONOTUS (MYRMOPYTIA)

Myrmopytia Emery, 1920b: 243 [as subgenus of Camponotus]. Type-species: Camponotus imitator, by original designation.

Taxonomic history

Myrmopytia as subgenus of Camponotus: Emery, 1920b: 243; Wheeler, W.M. 1922a: 707; Emery, 1925b: 114; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOTEMNUS)

Myrmotemnus Emery, 1920b: 246 [as subgenus of Camponotus]. Type-species: Camponotus moeschi, by original designation.

Taxonomic history

Myrmotemnus as junior synonym of Myrmamblys: Wheeler, W.M. 1921a: 19; Wheeler, W.M. 1922a: 708; Emery, 1925b: 137; Donisthorpe, 1943f: 671 (anachronism).

Myrmotemnus as subgenus of Camponotus: Emery, 1920b: 246; Santschi, 1926c: 601.

Subgenus CAMPONOTUS (PSEUDOCOLOBOPSIS)

Pseudocolobopsis Emery, 1920b: 249 [as subgenus of Camponotus]. Type-species: Camponotus macrocephalus, by original designation.

Taxonomic history

Pseudocolobopsis as subgenus of Camponotus: Emery, 1920b: 249; Wheeler, W.M. 1922a: 709; Emery, 1925b: 157; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOSTENUS)

Myrmostenus Emery, 1920b: 250 [as subgenus of Camponotus]. Type-species: Camponotus mirabilis, by original designation.

Taxonomic history

Myrmostenus as subgenus of Camponotus: Emery, 1920b: 250; Wheeler, W.M. 1922a: 709; Emery, 1925b: 161; all subsequent authors.

Subgenus CAMPONOTUS (HYPERCOLOBOPSIS)

Hypercolobopsis Emery, 1920b: 250 [as subgenus of Camponotus]. Type-species: Colobopsis paradoxa, by original designation.

Taxonomic history

Hypercolobopsis as subgenus of Camponotus: Emery, 1920b: 250; Wheeler, W.M. 1922a: 709; Emery, 1925b: 160; all subsequent authors.

Junior synonym of CAMPONOTUS (HYPERCOLOBOPSIS)

Neocolobopsis Borgmeier, 1928b: 65 [as subgenus of Camponotus]. Type-species: Camponotus (Neocolobopsis) scrobifer (junior synonym of Camponotus coriolanus), by original designation.

Taxonomic history

Neocolobopsis as subgenus of Camponotus Borgmeier, 1928b: 65.

Neocolobopsis as junior synonym of Hypercolobopsis: Kempf, 1968b: 411; Kempf, 1972a: 43.

Subgenus CAMPONOTUS (MYRMOPIROMIS)

Myrmopiromis Wheeler, W.M. 1921a: 17 [as subgenus of Camponotus]. Type-species: Formica fulvopilosa, by subsequent designation of Wheeler, W.M. 1922a: 707.

Taxonomic history

Myrmopiromis as subgenus of Camponotus: Wheeler, W.M. 1921a: 17; Wheeler, W.M. 1922a: 707; Emery, 1925b: 1127.

Subgenus CAMPONOTUS (MYRMOSAULUS)

Myrmosaulus Wheeler, W.M. 1921a: 18 [as subgenus of Camponotus]. Type-species: Formica cinerascens, by subsequent designation of Wheeler, W.M. 1922a: 705.

Taxonomic history

[Type-species not Formica singularis, unjustified subsequent designation by Emery, 1925b: 113, repeated in Donisthorpe, 1943f: 670. Note that Donisthorpe, 1932c: 445, states that C. cinerascens belongs in subgenus Myrmosericus. If so then Myrmosaulus would fall as a junior synonym of Myrmosericus.]

Myrmosaulus as subgenus of Camponotus: Wheeler, W.M. 1921a: 18; Wheeler, W.M. 1922a: 705; Emery, 1925b: 112; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOCLADOECUS)

Myrmocladoecus Wheeler, W.M. 1921a: 19 [as subgenus of Camponotus]. Type-species: Camponotus latangulus, by subsequent designation of Wheeler, W.M. 1922a: 709.

Taxonomic history

Myrmocladoecus as subgenus of Camponotus: Wheeler, W.M. 1921a: 19; Wheeler, W.M. 1922a: 709; Emery, 1925b: 166; all subsequent authors.

Subgenus CAMPONOTUS (MANNIELLA)

Manniella Wheeler, W.M. 1921a: 19 [as subgenus of Camponotus]. Type-species: Camponotus sphaericus, by original designation.

Taxonomic history

Manniella subgenus of Camponotus: Wheeler, W.M. 1921a: 19; Wheeler, W.M. 1922a: 710; Emery, 1925b: 159; all subsequent authors.

Subgenus CAMPONOTUS (MYRMISOLEPIS)

Myrmisolepis Santschi, 1921f: 310 [as subgenus of Camponotus]. Type-species: Camponotus epinotalis, by original designation.

Taxonomic history

Myrmisolepis as subgenus of Camponotus: Santschi, 1921f: 310; Emery, 1925b: 133; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOPELTA)

Myrmopelta Santschi, 1921f: 310 [as subgenus of Camponotus]. Type-species: Camponotus arminius, by original designation.

Taxonomic history

Myrmopelta as junior synonym of Myrmamblys: Emery, 1925b: 137.

Myrmopelta as subgenus of Camponotus: Santschi, 1921f: 310; Wheeler, W.M. 1922a: 704 (footnote); Santschi, 1926a: 16; Santschi, 1926c: 603; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOPLATYPUS)

Myrmoplatypus Santschi, 1921f: 311 [as subgenus of Camponotus]. Type-species: Camponotus platytarsus, by original designation.

Taxonomic history

Myrmoplatypus as subgenus of Camponotus: Santschi, 1921f: 311; Emery, 1925b: 169; all subsequent authors.

Subgenus CAMPONOTUS (MYRMEPINOTUS)

Myrmepinotus Santschi, 1921f: 312 [as subgenus of Camponotus]. Type-species: Camponotus echinoploides, by original designation.

Taxonomic history

Myrmepinotus as subgenus of Camponotus: Santschi, 1921f: 312; Emery, 1925b: 126; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOXYGENYS)

Myrmoxygenys Emery, 1925b: 70 [as subgenus of Camponotus]. Type-species: Camponotus caesar, by monotypy.

Taxonomic history

Myrmoxygenys as subgenus of Camponotus: Emery, 1925b: 70; all subsequent authors.

Subgenus CAMPONOTUS (KARAVAIEVIA)

Karavaievia Emery, 1925b: 115 [as subgenus of Camponotus]. Type-species: Camponotus exsectus, by original designation.

Taxonomic history

Karavaievia as subgenus of Camponotus: Emery, 1925b: 115; all subsequent authors.

Subgenus CAMPONOTUS (MYRMODIRACHIS)

Myrmodirachis Emery, 1925b: 168 [as subgenus of Camponotus]. Type-species: Camponotus heathi, by original designation.

Taxonomic history

Myrmodirachis as subgenus of Camponotus: Emery, 1925b: 168; all subsequent authors.

Subgenus CAMPONOTUS (MYRMESPERA)

Myrmespera Santschi, 1926b: 247 [as subgenus of Camponotus]. Type-species: Camponotus (Myrmespera) debellator, by subsequent designation of Santschi, 1926c: 604.

Taxonomic history

Myrmespera as subgenus of Camponotus: Santschi, 1926b: 247; Santschi, 1926c: 603; all subsequent authors.

Subgenus CAMPONOTUS (PARAMYRMAMBLYS)

Paramyrmamblys Santschi, 1926c: 604 [as subgenus of Camponotus].Type-species: Camponotus (Myrmamblys) ostiarius, by original designation.

Taxonomic history

Paramyrmamblys as subgenus of Camponotus: Santschi, 1926c: 604; all subsequent authors.

Subgenus CAMPONOTUS (MYRMACRHAPHE)

Myrmacrhaphe Santschi, 1926c: 607 [as subgenus of Camponotus]. Type-species: Camponotus conradti, by original designation.

Taxonomic history

Myrmacrhaphe as subgenus of Camponotus: Santschi, 1926c: 607; all subsequent authors.

Subgenus CAMPONOTUS (THLIPSEPINOTUS)

Thlipsepinotus Santschi, 1928e: 483 [as subgenus of Camponotus]. Type-species: Camponotus claripes, by original designation.

Taxonomic history

Thlipsepinotus as subgenus of Camponotus: Santschi, 1928e: 483; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOPALPELLA)

Myrmopalpella Stärcke, 1934: 30 [as subgenus of Camponotus]. Type-species: Camponotus megalonyx, by monotypy.

Taxonomic history

Myrmopalpella as genus: Wheeler, W.M. 1936e: 219 (in text); Donisthorpe, 1943f: 669.

Myrmopalpella as subgenus of Camponotus: Bolton, 1995b: 37.

Genus Camponotus references

Mayr, 1861: 35 (Europe species key); Roger, 1863b: 1, 9 (Camponotus, Colobopsis catalogues); Mayr, 1863: 397, 403 (Camponotus, Colobopsis catalogues); Mayr, 1865: 6, 7 (Camponotus, Colobopsis diagnosis); Mayr, 1867a: 67 (Colobopsis diagnois); Mayr, 1868c: 27 (†Baltic Amber species key); Mayr, 1870a: 373 (Colombia + Panama (= New Grenada) species key); Mayr, 1870b: 941 (Colobopsis species key); André, 1874: 175 (Europe species key); Forel, 1874: 38 (Switzerland species key); Mayr, 1876: 58 (Australia species key); Mayr, 1878: 869 (all Dendromyrmex species key (as part of Camponotus)); Forel, 1878: 367, 368 (Camponotus, Colobopsis diagnoses); André, 1882a: 138 (Europe & Algeria species key); André, 1882b: 159 (Europe & Algeria Colobopsis species key); Cresson, 1887: 255 (U.S.A. Camponotus, Colobopsis catalogue); Provancher, 1887: 228 (Canada species key); Nasonov, 1889: 57 (Russia species key); Forel, 1891b: 71, 216 (Madagascar species key); Forel, 1892j: 223 (India & Sri Lanka species key); Dalla Torre, 1893: 219 (Rhinomyrmex, Mayria, Camponotus catalogues); Emery, 1893i: 667 (North America species key); Forel, 1893b: 435 (India & Sri Lanka C. (Colobopsis) species key); Emery, 1895j: 772 (synoptic classification); Emery, 1896d: 370, 378 (species groups, catalogue); Emery, 1898a: 225 (additions to 1896a catalogue); Emery, 1903 (South America species key); Bingham, 1903: 343, 348 (India, Sri Lanka & Burma Colobopsis, Camponotus species keys); Ruzsky, 1905b: 185 (Russian Empire species key); Emery, 1908a: 183 (Palaearctic C. herculeanus & C. maculatus groups species key); Bondroit, 1910: 486 (Belgium species key); Wheeler, W.M. 1910d: 297 (North America species key); Forel, 1914a: 259, 266 (subgenera diagnoses, species listed by subgenus); Stitz, 1914: 94 (Central Europe species key); Santschi, 1915c: 266 (C. (Myrmotrema) species key); Emery, 1916b: 227 (Italy species key); Wheeler, W.M. 1916m: 600 (U.S.A., Connecticut species key); Mann, 1916: 489 (Dendromyrmex, all species key); Bondroit, 1918 (France & Belgium species key); Emery, 1920b: 235 (subgenera, diagoses); Emery, 1920c: 8 (Old World C. maculatus group, checklist & key); Wheeler, W.M. 1921a: 16 (subgenera, notes); Santschi, 1921f: 310 (subgenera, notes); Mann, 1921: 489, 499 (Fiji Is C. (Myrmogonia) & C. (Colobopsis) species keys); Wheeler, W.M. 1922a: 704 (subgenera synopsis); Wheeler, W.M. 1922a: 232, 948, 1039 (diagnosis, Afrotropical, Malagasy catalogues); Arnold, 1922: 610, 612, 619, 662, 673 (diagnosis, South Africa subgenera synopsis, South Africa C. (Tanaemyrmex), C. (Myrmophyma), C. (Myrmopsamma) species keys); Arnold, 1924: 683, 700, 709, 724, 735 (South Africa C. (Myrmosericus), C. (Myrmamblys), C. (Orthonotomyrmex), C. (Myrmotrema), C. (Myrmepomis) species keys); Emery, 1925a: 62 (Palaearctic C. lateralis group, key); Emery, 1925b: 59 (diagnosis, subgenera key, catalogue); Emery, 1925b: 69 (C. (Dinomyrmex) diagnosis, catalogue); Emery, 1925b: 70 (C. (Myrmoxygenys) & C. (Myrmopsamma) diagnoses, catalogues); Emery, 1925b: 71 (C. (Camponotus) diagnosis, catalogue); Emery, 1925b: 75 (C. (Tanaemyrmex) diagnosis, catalogue); Emery, 1925b: 104 (C. (Myrmosericus) diagnosis, catalogue); Emery, 1925b: 107 (C. (Myrmothrix) diagnosis, catalogue); Emery, 1925b: 109 (C. (Myrmophyma) diagnosis, catalogue); Emery, 1925b: 112 (C. (Myrmosaulus) diagnosis, catalogue); Emery, 1925b: 114 (C. (Myrmopytia) diagnosis, catalogue); Emery, 1925b: 115 (C. (Karavaievia) diagnosis, catalogue); Emery, 1925b: 116 (C. (Myrmentoma) diagnosis, catalogue); Emery, 1925b: 121 (C. (Mayria) diagnosis, catalogue); Emery, 1925b: 123 (C. (Myrmonesites) diagnosis, catalogue); Emery, 1925b: 124 (C. (Orthonotomyrmex) diagnosis, catalogue); Emery, 1925b: 126 (C. (Myrmepinotus) diagnosis, catalogue); Emery, 1925b: 127 (C. (Myrmopiromis) diagnosis, catalogue); Emery, 1925b: 130 (C. (Myrmotrema) diagnosis, catalogue); Emery, 1925b: 133 (C. (Myrmisolepis) diagnosis, catalogue); Emery, 1925b: 134 (C. (Myrmotarsus) diagnosis, catalogue); Emery, 1925b: 135 (C. (Myrmoplatys) diagnosis, catalogue); Emery, 1925b: 136 (C. (Myrmamblys) diagnosis, catalogue); Emery, 1925b: 142 (C. (Rhinomyrmex) diagnosis, catalogue); Emery, 1925b: 143 (C. (Myrmogonia) diagnosis, catalogue); Emery, 1925b: 144 (C. (Colobopsis) diagnosis, catalogue); Emery, 1925b: 151 (C. (Myrmosphincta) diagnosis, catalogue); Emery, 1925b: 152 (C. (Myrmaphaenus) diagnosis, catalogue); Emery, 1925b: 157 (C. (Pseudocolobopsis) diagnosis, catalogue); Emery, 1925b: 159 (C. (Manniella) diagnosis, catalogue); Emery, 1925b: 160 (C. (Hypercolobopsis) diagnosis, catalogue); Emery, 1925b: 161 (C. (Myrmostenus) & C. (Myrmobrachys) diagnoses, catalogues); Emery, 1925b: 166 (C. (Myrmocladoecus) diagnosis, catalogue); Emery, 1925b: 167 (C. (Myrmeurynota) diagnosis, catalogue); Emery, 1925b: 168 (C. (Myrmodirachis) diagnosis, catalogue); Emery, 1925b: 169 (C. (Myrmomalis) & C. (Myrmoplatypus) diagnoses, catalogues); Emery, 1925b: 170 (C. (Myrmepomis) diagnosis, catalogue); Emery, 1925b: 172 (Dendromyrmex diagnosis, catalogue); Santschi, 1926a: 16, 21 (C. (Myrmopelta) diagnosis, species key); Karavaiev, 1927c: 275 (Ukraine species key); Arnol'di, 1933b: 601 (Russia species key); Menozzi, 1933b: 81 (Israel species key); Menozzi, 1935c: 330 (Chile species key); Karavaiev, 1936: 175 (Ukraine species key); Finzi, 1936: 190 (Egypt species key); Menozzi, 1939a: 316 (Himalaya & Tibet species key); Stitz, 1939: 236 (Germany species key); Kratochvíl, 1941: 109 (Central Europe species key); Novák & Sadil, 1941: 109 (Central Europe species key); Cole, 1942: 387 (U.S.A., Utah species key); Donisthorpe, 1932g: 248 (C. (Orthonotomyrmex) notes); Holgersen, 1943b: 174 (Norway species key); Holgersen, 1944: 199 (Norway species key); Buren, 1944a: 293 (U.S.A., Iowa species key); Creighton, 1950a: 362 (North America species keys); Chapman & Capco, 1951: 220 (Asia checklist); Yasumatsu & Brown, 1951: 29 (C. herculeanus complex); Kusnezov, 1952d: 192 (Argentina species key); Yasumatsu & Brown, 1957: 45 (C. herculeanus complex); Pisarski, 1961a: 175 (Poland species key); Gregg, 1963: 655 (U.S.A., Colorado species key); Wheeler, G.C. & Wheeler, J. 1963: 163 (U.S.A., North Dakota species key); Arnol'di, 1967: 1817 (former U.S.S.R. C. (Camponotus) species key); Bernard, 1967: 330 (diagnosis, Western Europe species key); Kempf, 1972a: 42, 95 (Neotropical catalogue); Hashmi, 1973: 1 (C. (Myrmothrix) all species revision, key); Alayo, 1974: 27 (Cuba species key); Snelling & Hunt, 1976: 117 (Chile species key); Tarbinsky, 1976: 144 (Kyrghyzstan species key); Boven, 1977: 130 (Belgium species key); Kutter, 1977c: 191 (Switzerland species key); Arnol'di & Dlussky, 1978: 551 (former European U.S.S.R. species key); Collingwood, 1978: 90 (Iberian Peninsula species key); Collingwood, 1979: 87 (Fennoscandia & Denmark species key); Smith, D.R. 1979: 1424 (North America catalogue); Allred, 1982: 445 (U.S.A., Utah species key); Gösswald, 1985: 264 (Germany species key); Collingwood, 1985: 275 (Saudi Arabia species key); Taylor & Brown, D.R. 1985: 109 (Australia catalogue); Dumpert, 1986: 571 (C. (Karavaievia) species key); Wheeler, G.C. & Wheeler, J. 1986g: 60 (U.S.A., Nevada species key); Agosti & Collingwood, 1987b: 282 (Balkans species key); Taylor, 1987a: 10, 16 (Australia, New Caledonia, New Zealand checklists); Snelling, R.R. 1988: 57 (Nearctic C. (Myrmentoma) species key); Wang, C., Xiao & Wu, 1989a: 222 (China species key); Dlussky, Soyunov & Zabelin, 1990: 125 (Turkmenistan species key); Kupyanskaya, 1990: 164 (Far Eastern Russia species key); Brandão, 1991: 332 (Neotropical catalogue); Morisita, Kubota, Onoyama, et al. 1991: 38 (Japan species key); Terayama, 1991: 165 (Japan C. (Paramyrmamblys) species); Atanasov & Dlussky, 1992: 207 (Bulgaria species key); Wang, C. & Wu, 1994: 28 (China species key); Radchenko, 1994b: 116 (South Siberia species key); Arakelian, 1994: 82 (Armenia species key); Bolton, 1994: 50 (synoptic classification); Bolton, 1995a: 1048 (census); Bolton, 1995b: 83, 169 (catalogue); Douwes, 1995: 92 (Sweden species key); Dumpert, Maschwitz, Weissflog et al. 1995: 104 (C. (Karavaievia) species key); Kupyanskaya, 1995: 352 (Far Eastern Russia species key); Wu, J. & Wang, 1995: 168 (China species key); Collingwood & Agosti, 1996: 370 (Saudi Arabia species key); Seifert, 1996b: 170 (Central Europe species key); Radchenko, 1996b: 1195 (Asiatic Palaearctic species key); Cagniant, 1996b: 87 (Morocco species key); McArthur & Adams, 1996: 18 (Australia C. nigriceps group, key); Mackay, 1997: 194 (C. (Myrmostenus) species key); Mackay & Mackay, 1997b: 319 and Snelling, R.R. 2000: 607 (C. montivagus complex, keys); Robertson & Zachariades, 1997: 2 (C. fulvopilosus group, key); Snelling & Torres, 1998: 9 (Puerto Rico & Virgin Is species key); Collingwood & Prince, 1998: 24 (Portugal species key); Terayama, 1999b: 27 (Japan species key); Shattuck, 1999: 91 (Australia synopsis); Zhou, 2001b: 200 (China, Guangxi species key); McArthur & Shattuck, 2001: 27 (Australia C. macrocephalus group, key); Czechowski, Radchenko & Czechowska, 2002: 153 (Poland species key); Fernández, 2002: 58 (C. (Dendromyrmex) diagnosis, all species revision, key); Mackay & Mackay, 2002: 248 (U.S.A., New Mexico species key); Shattuck & McArthur, 2002: 65 (Australia C. wiederkehri group key); Bolton, 2003: 112 (synopsis); Shattuck, 2005: 3 (C. aureopilus group key); Radchenko, 2005b: 188 (North Korea species key); Hansen & Klotz, 2005: 75 (North American “Carpenter Ants” key); Snelling, R.R. 2006: 96 (U.S.A. C. festinatus complex, key); Karmaly & Narendran, 2006: 29 (India species, key); Dumpert, Maschwitz & Weissflog, 2006: 77 (C. (Karavaievia) species, keys); Seifert, 2007: 152 (North and Central Europe species key); McArthur, 2007a: 290 (Australia species key); Shattuck & Janda, 2009: 252 (C. aureopilus group key); Mackay & Delsinne, 2009: 496 (New World C. maculatus complex, key); Terayama, 2009: 216 (Taiwan species key); Heterick, 2009: 60 (south-western Australia species key); Ionescu-Hirsch, 2009: 59 (Israel species key); McArthur, 2010: 18 (south Australia species key).

Formicinae Formicini Formica TRUE TRUE FALSE

Extant: 176 species, 24 subspecies

Fossil: 64 species

Formica Linnaeus, 1758: 579. Type-species: Formica rufa, by subsequent designation of Curtis, 1839: 752.

Taxonomic history

Formica in Formicites: Lepeletier de Saint-Fargeau, 1835: 199.

Formica in Formicidae: Smith, F. 1858b: 1.

Formica in Formicinae: Mayr, 1861: 45 [Formicidae]; Mayr, 1862: 653 (in key) [Formicidae]; Mayr, 1865: 8 [Formicidae]; Mayr, 1868c: 47 [Formicidae].

Formica in Camponotinae: Forel, 1878: 373 [Camponotidae]; Emery & Forel, 1879a: 450 [Camponotidae]; Dalla Torre, 1893: 192.

Formica in Camponotinae, Camponotini: Forel, 1886f: 205.

Formica in Camponotinae, Formicini: Forel, 1893a: 165; Emery, 1895j: 772; Forel, 1899c: 127; Wheeler, W.M. 1910g: 143; Forel, 1912i: 89; Wheeler, W.M. 1915h: 120; Forel, 1917: 250.

Formica in Formicinae, Formicini: Ashmead, 1905b: 384; Bondroit, 1918: 41; Wheeler, W.M. 1922a: 699; Emery, 1925b: 241; all subsequent authors except the following.

Formica in Formicinae, Formica genus group: Agosti, 1991: 295.

Junior synonyms of FORMICA

Formicina Shuckard, in Swainson & Shuckard, 1840: 172. Type-species: Formica rufa, by subsequent designation of Wheeler, W.M. 1911f: 164.

Taxonomic history

[Type-species not Formica flava, unjustified subsequent designation by Emery, 1916a: 61 (footnote); repeated by Emery, 1916b: 239 and Wheeler, W.M. 1916k: 170.]

Formicina in Camponotinae, Formicini: Forel, 1917: 249.

Formicina as genus: Emery, 1916a: 61; Emery, 1916b: 239; Forel, 1917: 249; Bondroit, 1918: 19.

Formicina as subgenus of Lasius: Wheeler, W.M. 1916k: 172.

Formicina as junior synonym of Lasius: Emery, 1925b: 226; Brown, 1973b: 180.

Formicina as junior synonym of Acanthomyops: Donisthorpe, 1927b: 209.

Formicina as junior synonym of Formica: Wheeler, W.M. 1911c: 860; Wheeler, W.M. 1911f: 164; Donisthorpe, 1943f: 646; Agosti, 1994a: 106; Bolton, 1994: 50. [Formica and Formicina share the same type-species, synonymy is therefore absolute.]

Neoformica Wheeler, W.M. 1913a: 82 [as subgenus of Formica]. Type-species: Formica pallidefulva, by original designation.

Taxonomic history

[Neoformica also described as new by Wheeler, W.M. 1913f: 548.]

Neoformica as subgenus of Formica: Wheeler, W.M. 1913a: 82; Wheeler, W.M. 1913f: 388 (in key); Forel, 1917: 250; Wheeler, W.M. 1922a: 700; Emery, 1925b: 244; Donisthorpe, 1943f: 673; Creighton, 1950a: 543.

Neoformica as junior synonym of Formica: Buren, 1968a: 39; Agosti, 1994a: 107.

Raptiformica Forel, 1913i: 361 [as subgenus of Formica]. Type-species: Formica sanguinea, by original designation.

Taxonomic history

Raptiformica as subgenus of Formica: Forel, 1917: 250; Emery, 1925b: 258; Donisthorpe, 1943g: 723; Creighton, 1950a: 460.

Raptiformica as junior synonym of Formica: Wheeler, W.M. 1922a: 699 (footnote); Smith, M.R. 1951a: 860; Smith, D.R. 1979: 1448; Agosti, 1994a: 107.

Serviformica Forel, 1913i: 361 [as subgenus of Formica]. Type-species: Formica fusca, by original designation.

Taxonomic history

Serviformica as subgenus of Formica: Forel, 1913i: 361; Forel, 1917: 250; Emery, 1925b: 245; Donisthorpe, 1943g: 725.

Serviformica as junior synonym of Formica: Wheeler, W.M. 1922a: 699 (footnote); Smith, M.R. 1951a: 860; Francoeur, 1973: 34; Smith, D.R. 1979: 1448; Agosti, 1994a: 106.

Coptoformica Müller, 1923: 146 [as subgenus of Formica]. Type-species: Formica exsecta, by subsequent designation of Donisthorpe, 1941f: 37.

Taxonomic history

Coptoformica as junior synonym of Formica: Smith, M.R. 1951a: 860; Smith, D.R. 1979: 1448; Agosti, 1994a: 107.

Adformica Lomnicki, 1925a: 164 [as subgenus of Formica]. Type-species: Formica exsecta, by subsequent designation of Donisthorpe, 1927b: 316.

Taxonomic history

Adformica as junior synonym of Coptoformica: Stitz, 1939: 306; Donisthorpe, 1943f: 620 [Adformica and Coptoformica share the same type-species, synonymy is therefore absolute.]

Iberoformica Tinaut, 1990a: 282 [as subgenus of Formica]. Type-species: Formica subrufa, by original designation.

Taxonomic history

Iberoformica as junior synonym of Formica: Agosti, 1994a: 107.

Genus Formica references

Mayr, 1855: 300 (diagnosis, Austria species key); Smith, F. 1858b: 1 (diagnosis); Mayr, 1861: 46 (Europe species key); Roger, 1863b: 12 (catalogue); Mayr, 1863: 410 (catalogue); Mayr, 1865: 8 (diagnosis); André, 1874: 182 (Europe species key); Forel, 1874: 51 (Switzerland species key); Forel, 1878: 373 (diagnosis); André, 1882b: 176 (Europe & Algeria species key); Cresson, 1887: 255 (U.S.A. catalogue); Provancher, 1887: 232 (Canada species key); Nasonov, 1889: 61 (Russia species key); Dalla Torre, 1893: 192 (catalogue); Emery, 1893i: 643 (North America species key); Forel, 1894c: 402 (India species key); Bingham, 1903: 334 (India species key); Ruzsky, 1905b: 318 (Russian Empire species key); Wasmann, 1906: 8 (Luxemburg species key); Emery, 1909b: 180 (Palaearctic species key); Bondroit, 1910: 482 (Belgium species key); Wheeler, W.M. 1913f: 387 (subgenera, species groups, all species keys); Stitz, 1914: 88 (Central Europe species key); Donisthorpe, 1915d: 244 (Britain species key); Emery, 1916b: 257 (Italy species key); Wheeler, W.M. 1916m: 594 (U.S.A., Connecticut species key); Bondroit, 1917a: 174 (France F. rufa group, key); Bondroit, 1918: 41 (France & Belgium species key); Wheeler, W.M. 1922a: 699 (subgenera key); Emery, 1925b: 241 (diagnosis, subgenera key, catalogue); Emery, 1925b: 244 (F. (Neoformica) diagnosis, catalogue; Emery, 1925b: 245 (F. (Serviformica) diagnosis, catalogue; Emery, 1925b: 251 (F. (Formica) diagnosis, catalogue; Emery, 1925b: 258 (F. (Raptiformica) diagnosis, catalogue; Lomnicki, 1925a: 171 (Poland species key); Kuznetsov-Ugamsky, 1926b: 93 (Turkestan species key); Karavaiev, 1927c: 282 (Ukraine species key); Donisthorpe, 1927b: 284 (Britain species key); Arnol'di, 1933b: 603 (Russia species key); Karavaiev, 1936: 220 (Ukraine species key); Menozzi, 1939a: 321 (Himalaya & Tibet species key); Stitz, 1939: 306 (Germany species key); Creighton, 1940a: 1 (North America F. rufa subspecies key); Kratochvíl, 1941: 103 (Central Europe species key); Novák & Sadil, 1941: 103 (Central Europe species key); Cole, 1942: 375 (U.S.A., Utah species key); Holgersen, 1943b: 175 (Norway species key); Holgersen, 1944: 200 (Norway species key); Buren, 1944a: 299 (U.S.A., Iowa species key); Boven, 1947: 187 (Belgium species key); Creighton, 1950a: 456 (North America species key); Chapman & Capco, 1951: 197 (Asia checklist); Yarrow, 1954a: 229 (Britain F. fusca group); Yarrow, 1955a: 29 (Britain F. rufa group, key); Wilson & Brown, 1955: 108 (F. sanguinea & F. neogagates groups, synopsis); Boven, 1959: 12 (Netherlands species key); Betrem, 1960b: 75 (F. rufa group, key); Gregg, 1963: 498 (U.S.A., Colorado species key); Wheeler, G.C. & Wheeler, J. 1963: 205 (U.S.A., North Dakota species key); Cotti, 1963: 1 (F. rufa group, bibliography 1930-61); Dlussky, 1964: 1026 (former U.S.S.R. F. (Coptoformica) species key); Collingwood, 1964: 109 (Britain species key); Brown, 1965d: 181 (F. integra subgroup); Dlussky, 1965a: 15 (Mongolia and Tibet species key); Dlussky, 1967a: 39 (Palaearctic species key); Dlussky, 1967b: 80 (†Baltic Amber species key); Bernard, 1967: 285 (diagnosis, Western Europe species key); Buren, 1968a: 36 (Nearctic F. sanguinea group species key); Boven, 1970b: 34 (Netherlands species key); Dlussky & Pisarski, 1971: 145 (Poland species key); Francoeur, 1973: 272 (Nearctic F. fusca group, revision, key); Bolton & Collingwood, 1975: 6 (Britain species key); Tarbinsky, 1976: 177 (Kyrghyzstan species key); Boven, 1977: 94 (Belgium species key); Kutter, 1977c: 236 (Switzerland species key); Arnol'di & Dlussky, 1978: 552 (former European U.S.S.R. species key); Collingwood, 1978: 93 (Iberian Peninsula species key); Collingwood, 1979: 112 (Fennoscandia & Denmark species key); Smith, D.R. 1979: 1448 (North America catalogue); Douwes, 1979: 187 (F. rufa group systematics); Kupyanskaya, 1980: 95 (Far Eastern Russia species key); Allred, 1982: 446 (U.S.A., Utah species key); Kupyanskaya, 1984: 103 (Far Eastern Russia F. truncorum complex, key); Snelling & Buren, 1985: 71 (Nearctic F. sanguinea group, key); Gösswald, 1985: 274 (Germany species key); Wheeler, G.C. & Wheeler, J. 1986g: 76 (U.S.A., Nevada species key); Nilsson & Douwes, 1987: 74 (Norway species key); Agosti & Collingwood, 1987b: 285 (Balkans species key); Gösswald, 1989: 49 (F. rufa group taxonomy); Agosti & Bolton, 1990b: 149 (characters); Wu, J. 1990: 4 (China species key); Dlussky, Soyunov & Zabelin, 1990: 138 (Turkmenistan species key); Kupyanskaya, 1990: 174 (Far Eastern Russia species key); Morisita, Kubota, Onoyama, et al. 1991: 31 (Japan species key); Atanasov & Dlussky, 1992: 255 (Bulgaria species key); Agosti, 1994a: 106 (diagnosis, review of genus); Radchenko, 1994b: 112 (South Siberia species key); Arakelian, 1994: 89 (Armenia species key); Bolton, 1994: 50 (synoptic classification); Bolton, 1995a: 1049 (census); Bolton, 1995b: 190 (catalogue); Douwes, 1995: 94 (Sweden species key); Kupyanskaya, 1995: 353 (Far Eastern Russia species key); Wu, J. & Wang, 1995: 138 (China species key); Seifert, 1996b: 201 (Central Europe species key); Skinner & Allen, 1996: 41 (Britain species key); Collingwood & Prince, 1998: 25 (Portugal species key); Tinaut, A. & Martínez-Ibánez, 1998: 36 (Iberian Peninsula F. rufa, F. sanguinea & F. exsecta groups, key); Seifert, 2000a: 563 (Palaearctic F. (Coptoformica) species key); Czechowski, Radchenko & Czechowska, 2002: 148 (Poland species key); Chang & He, 2002a: 51 (northwest China species key); Seifert, 2002b: 267 (F. cinerea group revision key); Mackay & Mackay, 2002: 277 (U.S.A., New Mexico species key); Bolton, 2003: 127 (synopsis); Radchenko, 2005b: 188 (North Korea species key); Seifert, 2007: 176 (North and Central Europe species key); Trager, MacGown & Trager, 2007: 627 (F. pallidefulva group key); Dlussky, 2008a: 57 (Europe Eocene amber species key); Terayama, 2009: 224 (Taiwan species key); Seifert & Schultz, 2009: 258 (F. rufibarbis group key).

Formicinae Formicini Polyergus TRUE TRUE FALSE

5 species, 5 subspecies

Polyergus Latreille, 1804: 179. Type-species: Formica rufescens, by monotypy.

Taxonomic history

Polyergus in Formicites: Lepeletier de Saint-Fargeau, 1835: 198.

Polyergus in Formicidae: Smith, F. 1858b: 1.

Polyergus in Formicinae: Mayr, 1855: 383 [Formicidae]; Mayr, 1861: 43 [Formicinae]; Mayr, 1862: 653 [Formicidae]; Mayr, 1865: 8 [Formicidae].

Polyergus in Camponotinae: Forel, 1878: 367 [Camponotidae]; Dalla Torre, 1893: 214.

Polyergus in Camponotinae, Camponotini: Forel, 1886f: 200.

Polyergus in Camponotinae, Formicini: Forel, 1893a: 165; Emery, 1895j: 772; Forel, 1899c: 128; Wheeler, W.M. 1910g: 143; Forel, 1912i: 89; Forel, 1917: 250.

Polyergus in Formicinae, Formicini: Ashmead, 1905b: 384; Bondroit, 1918: 41; Wheeler, W.M. 1922a: 699; Emery, 1925b: 267; all subsequent authors except the following.

Polyergus in Formicinae, Formica genus group: Agosti, 1991: 295.

Genus Polyergus references

Mayr, 1855: 383 (diagnosis); Roger, 1863b: 12 (catalogue); Mayr, 1863: 443 (catalogue); Mayr, 1865: 8 (diagnosis); Forel, 1878: 373 (diagnosis); André, 1882b: 162 (Europe & Algeria species); Cresson, 1887: 255 (U.S.A. catalogue); Dalla Torre, 1893: 214 (catalogue); Emery, 1925b: 267 (diagnosis, catalogue); Buren, 1944a: 310 (U.S.A., Iowa species key); Smith, M.R. 1947g: 152 (U.S.A. species key); Creighton, 1950a: 556 (North America species key); Chapman & Capco, 1951: 203 (Asia checklist); Gregg, 1963: 632 (U.S.A., Colorado species key); Wheeler, G.C. & Wheeler, J. 1963: 274 (U.S.A., North Dakota species key); Bernard, 1967: 329 (diagnosis); Wheeler, J. 1968: 156 (Nearctic species, male genitalia); Smith, D.R. 1979: 1466 (North America catalogue); Wheeler, G.C. & Wheeler, J. 1986g: 95 (U.S.A., Nevada species key); Kupyanskaya, 1990: 207 (Far Eastern Russia species key); Agosti, 1994a: 109 (diagnosis, review of genus); Bolton, 1994: 50 (synoptic classification); Bolton, 1995a: 1052 (census); Bolton, 1995b: 342 (catalogue); Kupyanskaya, 1995: 361 (Far Eastern Russia species key); Mackay & Mackay, 2002: 359 (U.S.A., New Mexico species key); Bolton, 2003: 129 (synopsis).

Pseudomyrmecinae Pseudomyrmecini Pseudomyrmex TRUE TRUE FALSE

Extant: 131 species, 10 subspecies

Fossil: 12 species

Pseudomyrmex Lund, 1831b: 106. Type-species: Formica gracilis, by subsequent designation of Smith, M.R. 1952a: 98.

Taxonomic history

Pseudomyrmex in Pseudomyrmecinae: Smith, M.R. 1952a: 98; all subsequent authors.

Junior synonyms of PSEUDOMYRMEX

Leptalea Erichson, 1839: 309. Type-species: Formica gracilis, by subsequent designation of Wheeler, W.M. 1911f: 166.

Taxonomic history

Leptalea in Pseudomyrmecinae: Donisthorpe, 1943f: 656.

Leptalea as junior synonym of Pseudomyrma: Smith, F. 1858b: 153; Roger, 1863b: 24; Smith, F. 1877b: 58; Dalla Torre, 1893: 55; Emery, 1921f: 28.

[Leptalaea Spinola, 1851b: 52 and Spinola, 1853: 68, incorrect subsequent spellings.]

[Pseudomyrmex and Leptalea share the same type-species; synonymy is therefore absolute.]

Pseudomyrma Guérin-Méneville, 1844a: 427 [as subgenus of Formica]. Type-species: Pseudomyrma bicolor, by monotypy.

Taxonomic history

Pseudomyrma in Poneridae, Myrmicidae: Smith, F. 1858b: 153.

Pseudomyrma in Myrmicidae: Smith, F. 1871a: 328; Smith, F. 1877b: 58; Cresson, 1887: 262.

Pseudomyrma in Myrmicidae, Myrmeciidae: Emery, 1877a: 81.

Pseudomyrma in Myrmicinae: Mayr, 1865: 24 [Myrmicidae]; Dalla Torre, 1893: 55.

Pseudomyrma in Myrmicidae, Pseudomyrminae: Ashmead, 1905b: 383.

Pseudomyrma in Myrmicinae, Pseudomyrmini: Forel, 1893a: 164; Emery, 1895j: 768; Forel, 1899c: 86; Wheeler, W.M. 1910g: 139; Emery, 1914a: 40; Forel, 1917: 240; Emery, 1921f: 28.

Pseudomyrma in Pseudomyrminae: Wheeler, W.M. 1922a: 654; Donisthorpe, 1941d: 27; Donisthorpe, 1943g: 722.

Pseudomyrma as junior synonym of Pseudomyrmex: Smith, M.R. 1952a: 98; Kempf, 1972a: 214; Ward, 1990: 469; Bolton, 1994: 184.

Myrmex Guérin-Méneville, 1844a: 427. Type-species: Myrmex perboscii, by monotypy.

Taxonomic history

[Junior homonym of Myrmex Sturm, 1826: 32 (Coleoptera).]

Myrmex Guérin-Méneville junior synonym of Pseudomyrma: Smith, F. 1858b: 153; Roger, 1863b: 24; Smith, F. 1877b: 58; Dalla Torre, 1893: 55; Emery, 1921f: 28.

Clavanoda Enzmann, E.V. 1944: 61 [as "branch" of Pseudomyrma]. Type-species: Formica gracilis, by original designation.

Taxonomic history

Clavanoda as junior synonym of Pseudomyrmex: Ward, 1990: 469. [Pseudomyrmex and Clavanoda share the same type-species, synonymy is therefore absolute.]

Ornatinoda Enzmann, E.V. 1944: 61 [as "branch" of Pseudomyrma]. Type-species: Formica tenuis, by original designation.

Taxonomic history

Ornatinoda as junior synonym of Pseudomyrmex: Ward, 1990: 469.

Triangulinoda Enzmann, E.V. 1944: 61 [as "branch" of Pseudomyrma]. Type-species: Pseudomyrma spinicola, by original designation.

Taxonomic history

Triangulinoda as junior synonym of Pseudomyrmex: Ward, 1990: 469.

Apedunculata Enzmann, E.V. 1944: 62 [as "branch" of Pseudomyrma]. Type-species: Pseudomyrma sericea, by original designation.

Taxonomic history

Apedunculata as junior synonym of Pseudomyrmex: Ward, 1990: 469.

Latinoda Enzmann, E.V. 1944: 62 [as "branch" of Pseudomyrma]. Type-species: Pseudomyrma latinoda, by original designation.

Taxonomic history

Latinoda as junior synonym of Pseudomyrmex: Ward, 1990: 469.

Genus Pseudomyrmex references

Smith, F. 1858b: 153 (diagnosis); Roger, 1863b: 24 (catalogue); Mayr, 1863: 426, 451 (Leptalea, Pseudomyrma catalogues); Mayr, 1865: 24 (diagnosis); Mayr, 1870a: 407 (Colombia + Panama (= New Grenada) species key); Smith, 1877b: 58 (checklist); Cresson, 1887: 262 (U.S.A. catalogue); Dalla Torre, 1893: 55 (catalogue); Emery, 1921f: 28 (diagnosis, catalogue); Gallardo, 1932a: 44 (Argentina species key); Enzmann, E.V. 1944: 62 (all species keys); Creighton, 1950a: 79 (North America species key); Kempf, 1961a: 373 (P. gracilis group, partial key); Kempf, 1972a: 215 (Neotropical catalogue); Alayo, 1974: 9 (Cuba species key); Smith, D.R. 1979: 1345 (North America catalogue); Ward, 1985b: 215 (Nearctic species key); Billen, 1986: 173 (Dufour's gland); Ward, 1989: 407 (P. oculatus group, key); Ward, 1989: 430 (P. subtilissimus group, key); Ward, 1990: 469 (diagnosis, review of genus); Brandão, 1991: 373 (catalogue); Ward, 1992: 76 (†Dominican Amber species key); Ward, 1993: 130 (swollen-thorn Acacia-associated species key); Bolton, 1994: 184 (synoptic classification); Bolton, 1995a: 1052 (census); Bolton, 1995b: 370 (catalogue); Ward, 1999b: 473 (P. viduus group, and other Tachigali- and Triplaris-associated species key); Mackay & Mackay, 2002: 38 (U.S.A., New Mexico species key).

Cerapachyinae Cerapachyini Cerapachys TRUE TRUE FALSE

146 species

Cerapachys Smith, F. 1857a: 74. Type-species: Cerapachys antennatus, by subsequent designation of Bingham, 1903: 28.

Taxonomic history

Cerapachys in Myrmicinae: Smith, F. 1857a: 74 [Myrmicidae]; Smith, F. 1861b: 47 [Myrmicidae]; Smith, F. 1871a: 329 [Myrmicidae].

Cerapachys in Ponerinae: Dalla Torre, 1893: 17.

Cerapachys in Dorylinae, Cerapachyini: Emery, 1895j: 765; Emery, 1901a: 34.

Cerapachys in Ponerinae, Cerapachyini: Forel, 1893a: 162 (also misspelled Orapachys, same page); Forel, 1900d: 331; Ashmead, 1905b: 382; Wheeler, W.M. 1910g: 137; Emery, 1911d: 8; Arnold, 1915: 11; Forel, 1917: 239; Brown, 1975: 18; Snelling, R.R. 1981: 389; Dlussky & Fedoseeva, 1988: 79; Hölldobler & Wilson, 1990: 10.

Cerapachys in Cerapachyinae, Cerapachyini: Wheeler, W.M. 1902d: 185; Wheeler, W.M. 1922a: 639; Donisthorpe, 1943f: 632; Chapman & Capco, 1951: 17; Kempf, 1972a: 76; Wheeler, G.C. & Wheeler, J. 1985: 256; Bolton, 1990a: 67; Bolton, 1990c: 1357; Jaffe, 1993: 9; Bolton, 1994: 19; Bolton, 2003: 34.

Junior synonyms of CERAPACHYS

Syscia Roger, 1861a: 19. Type-species: Syscia typhla, by monotypy.

Taxonomic history

Syscia in Ponerinae: Mayr, 1862: 714 (in key) [Poneridae]; Mayr, 1865: 15 [Poneridae]; Dalla Torre, 1893: 17.

Syscia in Dorylinae, Cerapachyini: Emery, 1895j: 765.

Syscia in Ponerinae, Cerapachyini: Forel, 1893a: 162; Forel, 1900d: 329; Ashmead, 1905b: 382.

Syscia as genus: Dalla Torre, 1893: 17; Forel, 1900d: 329; Bingham, 1903: 31.

Syscia as subgenus of Cerapachys: Wheeler, W.M. 1902d: 185; Emery, 1902c: 24; Wheeler, W.M. 1910g: 137; Emery, 1911d: 10; Forel, 1917: 239; Wheeler, W.M. 1922a: 639; Donisthorpe, 1943g: 730.

Syscia as junior synonym of Cerapachys: Kempf, 1972a: 76; Brown, 1975: 18; all subsequent authors.

Ooceraea Roger, 1862a: 248. Type-species: Ooceraea fragosa, by monotypy.

Taxonomic history

Ooceraea in Myrmicinae: Mayr, 1865: 24 [Myrmicidae].

Ooceraea in Myrmicidae, Pheidolidae: Emery, 1877a: 81.

Ooceraea in Poneridae: Smith, F. 1871a: 324.

Ooceraea in Ponerinae: Dalla Torre, 1893: 17.

Ooceraea in Ponerinae, Cerapachyini: Forel, 1893a: 162.

Ooceraea in Dorylinae, Cerapachyini: Emery, 1895j: 765; Emery, 1901a: 34.

Ooceraea in Ecitoninae, Ecitonini: Ashmead, 1905b: 381; Ashmead, 1906: 25.

Ooceraea as genus: Roger, 1862a: 248; Smith, F. 1871a: 324; Dalla Torre, 1893: 17; Forel, 1900d: 329; Bingham, 1903: 31.

Ooceraea as subgenus of Cerapachys: Emery, 1902c: 24; Wheeler, W.M. 1902d: 185; Wheeler, W.M. 1910g: 137; Emery, 1911d: 10; Forel, 1917: 239; Wheeler, W.M. 1922a: 639; Donisthorpe, 1943f: 678.

Ooceraea as junior synonym of Cerapachys: Brown, 1973b: 183 [provisional]; Brown, 1975: 19; all subsequent authors.

Lioponera Mayr, 1879: 666. Type-species: Lioponera longitarsus, by monotypy.

Taxonomic history

Lioponera in Ponerinae: Dalla Torre, 1893: 17.

Lioponera in Pachycondylinae, Lioponerini: Ashmead, 1905b: 382.

Lioponera in Ponerinae, Cerapachyini: Forel, 1893a: 162; Forel, 1900d: 329; Wheeler, W.M. 1910g: 137; Emery, 1911d: 11; Forel, 1917: 239.

Lioponera in Dorylinae, Cerapachyini: Emery, 1895j: 765.

Lioponera in Cerapachyinae, Cerapachyini: Wheeler, W.M. 1902d: 185; Wheeler, W.M. 1922a: 639; Donisthorpe, 1943f: 657.

Lioponera as subgenus of Cerapachys: Forel, 1892l: 244.

Lioponera as genus: Mayr, 1879: 666; Dalla Torre, 1893: 17; Forel, 1900d: 329; Bingham, 1903: 26; Emery, 1911d: 11; Wheeler, W.M. 1922a: 639; Donisthorpe, 1943f: 657.

Lioponera as junior synonym of Cerapachys: Brown, 1975: 19; all subsequent authors.

Parasyscia Emery, in André, 1882c: 235. Type-species: Parasyscia piochardi, by monotypy.

Taxonomic history

Parasyscia in Ponerinae: Dalla Torre, 1893: 17.

Parasyscia in Dorylinae, Cerapachyini: Emery, 1895j: 765.

Parasyscia in Ponerinae, Cerapachyini: Forel, 1893a: 162 [Cerapachysii]; Ashmead, 1905b: 382.

Parasyscia as genus: Dalla Torre, 1893: 17.

Parasyscia as subgenus of Cerapachys: Forel, 1892l: 243; Wheeler, W.M. 1902d: 185; Wheeler, W.M. 1910g: 137; Emery, 1911d: 9; Forel, 1917: 239; Wheeler, W.M. 1922a: 639; Donisthorpe, 1943f: 681; Creighton, 1950a: 57.

Parasyscia as junior synonym of Cerapachys: Kempf, 1972a: 76; Brown, 1975: 18; all subsequent authors.

Phyracaces Emery, 1902c: 23. Type-species: Cerapachys mayri, by original designation.

Taxonomic history

Phyracaces in Ponerinae, Cerapachyini: Ashmead, 1905b: 382; Wheeler, W.M. 1910g: 137; Emery, 1911d: 10; Arnold, 1915: 17; Forel, 1917: 239.

Phyracaces in Cerapachyinae, Cerapachyini: Wheeler, W.M. 1902d: 185; Wheeler, W.M. 1922a: 639; Donisthorpe, 1943f: 683.

Phyracaces as junior synonym of Cerapachys: Brown, 1973b: 183 [provisional]; Brown, 1975: 19; all subsequent authors.

Cysias Emery, 1902c: 24 [as subgenus of Cerapachys]. Type-species: Ooceraea papuana, by original designation.

Taxonomic history

Cysias in Ponerinae, Cerapachyini: Ashmead, 1905b: 382.

Cysias as genus: Ashmead, 1905b: 382.

Cysias as subgenus of Cerapachys: Emery, 1902c: 24; Wheeler, W.M. 1910g: 137; Donisthorpe, 1943f: 637.

Cysias as junior synonym of Syscia: Emery, 1911d: 10; Brown, 1975: 19; all subsequent authors.

Ceratopachys Schulz, W.A. 1906: 155, unjustified emendation of Cerapachys.

Taxonomic history

Ceratopachys as junior synonym of Cerapachys: Wheeler, W.M. 1922a: 755.

Chrysapace Crawley, 1924: 380. Type-species: Chrysapace jacobsoni (junior secondary homonym in Cerapachys, replaced by Cerapachys crawleyi), by monotypy.

Taxonomic history

Chrysapace in Ponerinae: Crawley, 1924: 380.

Chrysapace in Cerapachyinae, Cerapachyini: Donisthorpe, 1943f: 633.

Chrysapace as junior synonym of Cerapachys: Brown, 1975: 19; all subsequent authors.

Neophyracaces Clark, 1941: 76. Type-species: Phyracaces clarus, by original designation.

Taxonomic history

Neophyracaces in Cerapachyinae: Clark, 1941: 76.

Neophyracaces as junior synonym of Cerapachys: Brown, 1973b: 183 [provisional]; Brown, 1975: 19; all subsequent authors.

Yunodorylus Xu, 2000b: 297. Type-species: Yunodorylus sexspinus, by original designation.

Taxonomic history

Yunodorylus in Dorylinae: Xu, 2000b: 297.

Yunodorylus in Cerapachyinae, Cerapachyini: Bolton, 2003: 141, 268.

Yunodorylus as junior synonym of Cerapachys: Bolton, 2003: 141, 268; Borowiec, 2009: 57.

Genus Cerapachys references

Roger, 1863b: 21, 25 (catalogue); Mayr, 1863: 403, 439, 455 (catalogue); Mayr, 1865: 15, 24 (Syscia, Ooceraea diagnoses); André, 1882c: 236 (Europe & Algeria species); Dalla Torre, 1893: 17 (Lioponera, Cerapachys, Parasyscia, Ooceraea, Syscia catalogues); Forel, 1900d: 331 (India & Sri Lanka species key); Bingham, 1903: 27, 28 (India, Sri Lanka & Burma Lioponera, Cerapachys species keys); Emery, 1911d: 8 (diagnosis, subgenera key, catalogue); Emery, 1911d: 9 (C. (Parasyscia) diagnosis, catalogue); Emery, 1911d: 10 (C. (Ooceraea) & C. (Syscia) diagnoses, catalogues); Emery, 1911d: 10 (Phyracaces diagnosis, catalogue); Emery, 1911d: 11 (Lioponera diagnosis, catalogue); Arnold, 1915: 11 (diagnosis, South Africa species key); Wheeler, W.M. 1918a: 239 (Australia Phyracaces species key); Mann, 1921: 408 (Fiji species key); Wheeler, W.M. 1922a: 52, 53, 639 (Cerapachys & Phyracaces diagnoses, subgenera key); Wheeler, W.M. 1924d: 224 (Chrysapace, comments); Arnold, 1926: 193 (South Africa species, revised key); Donisthorpe, 1939a: 253 (Lioponera diagnosis); Chapman & Capco, 1951: 17, 21, 22 (Asia Cerapachys, Lioponera, Phyracaces checklists); Wilson, 1959b: 44, 52, 55, 56 (Melanesia, Fiji Is, New Guinea & New Caledonia species keys); Kempf, 1972a: 76 (Neotropical catalogue); Brown, 1975: 18, 26 (diagnosis, keys); Smith, D.R. 1979: 1333 (North America catalogue); Taylor & Brown, D.R. 1985: 23 (Australia catalogue); Taylor, 1987a: 16 (Australia, New Caledonia checklist); Terayama, Kubota, Sakai & Kawazoe, 1988: 35 (Taiwan species key); Morisita, Kubota, Onoyama, et al. 1989: 30 (Japan species key); Bolton, 1995a: 1048 (census); Bolton, 1995b: 142 (catalogue); Wu, J. & Wang, 1995: 47 (China species key); Collingwood & Agosti, 1996: 312 (Saudi Arabia species key); Terayama, 1996: 18 (Japan species key); Shattuck, 1999: 60 (Australia synopsis); Mackay & Mackay, 2002: 36 (U.S.A., New Mexico species key); Borowiec, 2009: 46 (C. sexspinus group key); Terayama, 2009: 122 (Taiwan species key); Heterick, 2009: 126 (south-western Australia species key).

Ecitoninae Ecitonini Labidus TRUE TRUE FALSE

7 species, 1 subspecies

Labidus Jurine, 1807: 282. Type-species: Labidus latreillii (junior synonym of Formica coeca), by monotypy.

Taxonomic history

Labidus in Dorylidae: Leach, 1815: 147 [Dorylida]; Haliday, 1836: 331; Shuckard, 1840a: 196; Smith, F. 1859b: 1; Cresson, 1887: 259.

Labidus in Dorylinae: Mayr, 1865: 16 [Dorylidae]; Forel, 1878: 365 (footnote) [Dorylidae].

Labidus in Dorylinae, Ecitonini: Emery, 1910b: 21 [Ecitini]; Forel, 1917: 240; Donisthorpe, 1943f: 654; Borgmeier, 1955: 79; Kempf, 1972a: 126; Dlussky & Fedoseeva, 1988: 79.

Labidus in Ecitoninae: Brown, 1973b: 166; Snelling, R.R. 1981: 392.

Labidus in Ecitoninae, Ecitonini: Bolton, 1990c: 1357; Hölldobler & Wilson, 1990: 12; Jaffe, 1993: 13; Bolton, 1994: 39; all subsequent authors.

Labidus as junior synonym of Eciton: Dalla Torre, 1893: 1.

Labidus as subgenus of Eciton: Emery, 1910b: 21; Forel, 1917: 240; Gallardo, 1920: 312; Wheeler, W.M. 1922a: 635; Borgmeier, 1923: 41; Donisthorpe, 1943f: 654; Creighton, 1950a: 61; Smith, M.R. 1951a: 779; Kusnezov, 1956: 9.

Labidus as genus: Jurine, 1807: 282; Shuckard, 1840a: 196; Borgmeier, 1953: 4; Borgmeier, 1955: 80; Smith, M.R. 1958c: 108; all subsequent authors.

Junior synonyms of LABIDUS

Nycteresia Roger, 1861a: 21. Type-species: Formica coeca, by monotypy.

Taxonomic history

Nycteresia in Ponerinae: Mayr, 1862: 714 [Poneridae].

Nycteresia as junior synonym of Eciton: Mayr, 1865: 76; Dalla Torre, 1893: 1.

Nycteresia as junior synonym of Labidus: Emery, 1910b: 21; Donisthorpe, 1943f: 675; Borgmeier, 1955: 80; all subsequent authors.

Pseudodichthadia André, 1885: 838. Type-species: Pseudodichthadia incerta (junior synonym of Formica coeca), by monotypy.

Taxonomic history

Pseudodichthadia in Dorylidae: André, 1885: 838.

Pseudodichthadia in Dorylinae: Dalla Torre, 1893: 7.

Pseudodichthadia in Dorylinae, Ecitonini: Forel, 1893a: 163; Emery, 1895j: 765 [Ecitonii].

Pseudodichthadia as junior synonym of Eciton: André, in Forel, 1899c: 160 (footnote).

Pseudodichthadia as junior synonym of Labidus: Emery, 1910b: 21; Borgmeier, 1955: 80; all subsequent authors.

Genus Labidus references

Roger, 1863b: 21, 41 (catalogue); Mayr, 1863: 424, 436 (catalogue); Mayr, 1865: 16 (diagnosis); Cresson, 1887: 259 (U.S.A. catalogue); Emery, 1910b: 21 (diagnosis, catalogue); Gallardo, 1920: 312 (Argentina species key); Creighton, 1950a: 62 (North America species key); Borgmeier, 1955: 80, 84 (diagnosis, all species revision, key); Kempf, 1972a: 126 (Neotropical catalogue); Watkins, 1976: 8 (all species key); Smith, D.R. 1979: 1327 (North America catalogue); Watkins, 1982: 209 (Mexico species key); Bolton, 1995a: 1050 (census); Bolton, 1995b: 219 (catalogue); Palacio, 1999: 153 (Colombia species key).

Ecitoninae Ecitonini Neivamyrmex TRUE TRUE FALSE

Extant: 124 species, 1 subspecies

Fossil: 1 species

Neivamyrmex Borgmeier, 1940: 606 [as subgenus of Eciton].

Taxonomic history

[Replacement name for Acamatus Emery, 1894c: 181; junior homonym of Acamatus Schoenherr, 1833: 20 (Coleoptera.]

Neivamyrmex in Dorylinae, Ecitonini: Donisthorpe, 1943f: 673; Borgmeier, 1955: 79; Kempf, 1972a: 152; Dlussky & Fedoseeva, 1988: 79.

Neivamyrmex in Ecitoninae: Brown, 1973b: 166; Snelling, R.R. 1981: 392.

Neivamyrmex in Ecitoninae, Ecitonini: Bolton, 1990c: 1357; Hölldobler & Wilson, 1990: 12; Jaffe, 1993: 13; Bolton, 1994: 39; all subsequent authors.

Neivamyrmex as subgenus of Eciton: Creighton, 1950a: 64; Kusnezov, 1956: 8.

Neivamyrmex as genus: Borgmeier, 1950c: 624; Borgmeier, 1955: 277; Smith, M.R. 1958c: 108; all subsequent authors except the above entry.

Junior synonym of NEIVAMYRMEX

Woitkowskia Enzmann, E.V. 1952: 443. Type-species: Woitkowskia connectens (junior synonym of Labidus walkerii), by original designation.

Taxonomic history

Woitkowskia as junior synonym of Neivamyrmex: Borgmeier, 1955: 277; all subsequent authors.

Genus Neivamyrmex references

Emery, 1894c: 183 (species key); Emery, 1910b: 23 (diagnosis, catalogue); Gallardo, 1920: 348 (Argentina species key); Smith, M.R. 1942c: 542 (U.S.A. species key); Buren, 1944a: 280 (U.S.A., Iowa species key); Creighton, 1950a: 66 (North America species key); Borgmeier, 1955: 277, 290 (diagnosis, all species revision, key); Watkins, 1971: 93 (U.S.A. species key); Kempf, 1972a: 152 (Neotropical catalogue); Watkins, 1972: 348 (U.S.A. species key); Watkins, 1976: 11 (all species key); Smith, D.R. 1979: 1329 (North America catalogue); Watkins, 1982: 210 (Mexico species key); Watkins, 1985: 481 (U.S.A. species key); Wheeler, G.C. & Wheeler, J. 1986g: 19 (U.S.A., Nevada species key); Brandão, 1991: 359 (Neotropical catalogue); Bolton, 1995a: 1051 (census); Bolton, 1995b: 287 (catalogue); Ward, 1999a: 90 (modification of Watkins, 1985 key); Palacio, 1999: 154 (Colombia species key); Mackay & Mackay, 2002: 40 (U.S.A., New Mexico species key); Coovert, 2005: 30 (U.S.A., Ohio species key); Snelling, G.C & Snelling, R.R., 2007: 462 (U.S.A. species, keys).

Homonym replaced by NEIVAMYRMEX

Acamatus Emery, 1894c: 181 [as subgenus of Eciton]. Type-species: Eciton (Acamatus) schmitti (junior synonym of Labidus nigrescens), by subsequent designation of Ashmead, 1906: 24; Wheeler, W.M. 1911f: 157.

Taxonomic history

[Junior homonym of Acamatus Schoenherr, 1833: 20 (Coleoptera).]

Acamatus in Ecitoninae, Ecitonini: Ashmead, 1905b: 381; Ashmead, 1906: 24.

Acamatus in Dorylinae, Ecitonini: Emery, 1910b: 23 [Ecitini].

Acamatus as genus: Ashmead, 1905b: 381; Ashmead, 1906: 24.

Acamatus as subgenus of Eciton: Emery, 1894c: 181; Emery, 1895j: 765; Emery, 1910b: 23; Wheeler, W.M. 1910g: 138; Forel, 1917: 240; Gallardo, 1920: 345; Wheeler, W.M. 1922a: 635; Borgmeier, 1923: 44; Donisthorpe, 1943f: 618.

Ecitoninae Ecitonini Nomamyrmex TRUE TRUE FALSE

2 species, 2 subspecies

Nomamyrmex Borgmeier, 1936: 55 [as subgenus of Eciton]. Type-species: Eciton crassicornis, by original designation.

Taxonomic history

Nomamyrmex in Dorylinae, Ecitonini: Donisthorpe, 1943f: 674; Borgmeier, 1955: 79; Kempf, 1972a: 164; Dlussky & Fedoseeva, 1988: 79.

Nomamyrmex in Ecitoninae: Brown, 1973b: 166; Snelling, R.R. 1981: 392.

Nomamyrmex in Ecitoninae, Ecitonini: Bolton, 1990c: 1357; Hölldobler & Wilson, 1990: 12; Jaffe, 1993: 13; Bolton, 1994: 39.

Nomamyrmex as subgenus of Eciton: Borgmeier, 1936: 55; Kusnezov, 1956: 9.

Nomamyrmex as genus: Borgmeier, 1953: 4; Borgmeier, 1955: 135; Smith, M.R. 1958c: 108; all subsequent authors except the entry above.

Genus Nomamyrmex references

Borgmeier, 1955: 135 (diagnosis, all species revision, key); Kempf, 1972a: 164 (catalogue); Watkins, 1976: 7 (all species key); Watkins, 1977a: 207 (all species key); Smith, D.R. 1979: 1329 (North America catalogue); Watkins, 1982: 215 (Mexico species key); Brandão, 1991: 360 (catalogue); Bolton, 1995a: 1051 (census); Bolton, 1995b: 292 (catalogue); Palacio, 1999: 161 (Colombia species key).

Ectatomminae Ectatommini Ectatomma TRUE TRUE FALSE

Extant: 15 species

Fossil: 1 species

Ectatomma Smith, F. 1858b: 102. Type-species: Formica tuberculata, by subsequent designation of Bingham, 1903: 82.

Taxonomic history

Ectatomma in Poneridae: Smith, F. 1858b: 102; Smith, F. 1871a: 324.

Ectatomma in Ponerinae: Mayr, 1862: 714 (in key) [Poneridae]; Mayr, 1865: 14 [Poneridae]; Mayr, 1868c: 75 [Poneridae]; Dalla Torre, 1893: 23.

Ectatomma in Ponerinae, Ponerini: Forel, 1895b: 112; Forel, 1899c: 5; Forel, 1900d: 316.

Ectatomma in Pachycondylinae, Ectatommini: Ashmead, 1905b: 382.

Ectatomma in Ponerinae, Ectatommini: Emery, 1895j: 767; Wheeler, W.M. 1910g: 135; Emery, 1911d: 41; Wheeler, W.M. 1915h: 33; Forel, 1917: 236; Gallardo, 1918b: 27; Wheeler, W.M. 1922a: 643; all subsequent authors to the following.

Ectatomma in Ectatomminae, Ectatommini: Bolton, 2003: 46.

Genus Ectatomma references

Roger, 1863b: 17 (catalogue); Mayr, 1863: 409 (catalogue); Mayr, 1865: 14 (diagnosis); Mayr, 1867a: 87 (diagnosis); Dalla Torre, 1893: 23 (catalogue); Emery, 1911d: 41 (diagnosis, subgenera key, catalogue); Gallardo, 1918b: 30 (Argentina species key); Kusnezov, 1957b: 13 (partial key); Brown, 1958g: 206 (review of genus, key); Kempf, 1972a: 104 (Neotropical catalogue); Smith, D.R. 1979: 1336 (North America catalogue); Kugler, C. & Brown, 1982: 6 (all species key); Brandão, 1991: 341 (catalogue); Fernández, 1991: 551 (Colombia species key); Lattke, 1994: 109 (phylogeny); Bolton, 1995a: 1049 (census); Bolton, 1995b: 186 (catalogue) ); Arias-Penna, 2008b: 56 (all species key).

Ectatomminae Ectatommini Gnamptogenys TRUE TRUE FALSE

Extant: 137 species

Fossil: 6 species

Gnamptogenys Roger, 1863a: 174. Type-species: Ponera tornata, by subsequent designation of Emery, 1911d: 44.

Taxonomic history

[Type-species not Ectatomma concinna, unjustified subsequent designation by Wheeler, W.M. 1911f: 164; see Emery, 1912d: 271 and Wheeler, W.M. 1913a: 79.]

Gnamptogenys in Ponerinae: Mayr, 1865: 15 [Poneridae].

Gnamptogenys in Pachycondylinae, Ectatommini: Ashmead, 1905b: 382.

Gnamptogenys in Ponerinae, Ectatommini: Emery, 1895j: 767; Emery, 1911d: 44; Forel, 1917: 236; Wheeler, W.M. 1922a: 643; all subsequent authors to the following.

Gnamptogenys in Ectatomminae, Ectatommini: Bolton, 2003: 46.

Gnamptogenys as subgenus of Ectatomma: Mayr, 1887: 540; Dalla Torre, 1893: 23; Emery, 1895j: 767; Forel, 1895b: 113; Forel, 1899c: 7; Wheeler, W.M. 1910g: 135; Emery, 1911d: 44; Gallardo, 1918b: 42; Wheeler, W.M. 1922a: 644; Borgmeier, 1923: 60; Kusnezov, 1956: 14.

Gnamptogenys as genus: Roger, 1863a: 174; Forel, 1917: 236; Brown, 1958g: 211; all subsequent authors.

Junior synonyms of GNAMPTOGENYS

Stictoponera Mayr, 1887: 539 [as subgenus of Ectatomma]. Type-species: Ponera coxalis, by subsequent designation of Bingham, 1903: 82.

Taxonomic history

Stictoponera in Pachycondylinae, Ectatommini: Ashmead, 1905b: 382.

Stictoponera in Ponerinae, Ectatommini: Emery, 1895j: 767; Emery, 1911d: 47; Forel, 1917: 236; Wheeler, W.M. 1922a: 642; Donisthorpe, 1943g: 728.

Stictoponera as subgenus of Ectatomma: Mayr, 1887: 539; Dalla Torre, 1893: 23; Emery, 1895j: 767; Forel, 1900d: 316; Bingham, 1903: 82; Wheeler, W.M. 1910g: 135.

Stictoponera as genus: Emery, 1911d: 47; Forel, 1917: 236; Wheeler, W.M. 1922a: 642; Donisthorpe, 1943g: 728.

Stictoponera as junior synonym of Gnamptogenys: Brown, 1958g: 211; all subsequent authors.

Holcoponera Mayr, 1887: 540 [as subgenus of Ectatomma]. Type-species: Gnamptogenys striatula, by subsequent designation of Emery, 1911d: 40.

Taxonomic history

Holcoponera in Pachycondylinae, Ectatommini: Ashmead, 1905b: 382.

Holcoponera in Ponerinae, Ectatommini: Emery, 1895j: 767; Emery, 1911d: 40; Forel, 1917: 236; Gallardo, 1918b: 21; Wheeler, W.M. 1922a: 643; all subsequent authors.

Holcoponera as subgenus of Ectatomma: Mayr, 1887: 540; Dalla Torre, 1893: 23; Emery, 1895j: 767; Wheeler, W.M. 1910g: 135.

Holcoponera as genus: Emery, 1902a: 181; Emery, 1911d: 40; Forel, 1917: 236; Gallardo, 1918b: 21; Wheeler, W.M. 1922a: 643; Borgmeier, 1923: 55; Santschi, 1929h: 437; Kusnezov, 1956: 14.

Holcoponera as junior synonym of Gnamptogenys: Brown, 1958g: 211; all subsequent authors.

Alfaria Emery, 1896e: 177 (diagnosis in key). Type-species: Alfaria simulans, by original designation.

Taxonomic history

[Alfaria also described as new by Emery, 1896g: 41.]

Alfaria in Ponerinae, Ponerini: Forel, 1899c: 3.

Alfaria in Pachycondylinae, Ectatommini: Ashmead, 1905b: 382.

Alfaria in Ponerinae, Ectatommini: Wheeler, W.M. 1910g: 135; Emery, 1911d: 46; Forel, 1917: 236; Wheeler, W.M. 1922a: 642; all subsequent authors.

Alfaria as junior synonym of Gnamptogenys: Brown, 1958g: 211; all subsequent authors.

Poneracantha Emery, 1897d: 547 [as subgenus of Ectatomma]. Type-species: Ectatomma (Holcoponera) bispinosum, by original designation.

Taxonomic history

Poneracantha as subgenus of Ectatomma: Emery, 1911d: 43; Forel, 1917: 236; Wheeler, W.M. 1922a: 643.

Poneracantha as junior synonym of Gnamptogenys: Brown, 1958g: 211; all subsequent authors.

Rhopalopone Emery, 1897d: 549. Type-species: Rhopalopone epinotalis, by monotypy.

Taxonomic history

Rhopalopone in Ponerinae, Ponerini: Ashmead, 1905b: 382; Wheeler, W.M. 1910g: 135.

Rhopalopone in Ponerinae, Ectatommini: Emery, 1911d: 34 [subtribe Typhlomyrmecini]; Forel, 1917: 236; Wheeler, W.M. 1922a: 642; all subsequent authors.

Rhopalopone as genus: Emery, 1911d: 34; Wheeler, W.M. 1922a: 642; Borgmeier, 1923: 54.

Rhopalopone as junior synonym of Gnamptogenys: Brown, 1958g: 211; all subsequent authors.

Emeryella Forel, 1901e: 334. Type-species: Emeryella schmitti, by monotypy.

Taxonomic history

Emeryella in Pachycondylinae, Amblyoponini: Ashmead, 1905b: 283.

Emeryella in Ponerinae, Ectatommini: Wheeler, W.M. 1910g: 135; Emery, 1911d: 46; Forel, 1917: 236; Wheeler, W.M. 1922a: 643; Wheeler, W.M. 1930f: 13; all subsequent authors.

Emeryella as junior synonym of Gnamptogenys: Brown, 1958g: 211; all subsequent authors.

Mictoponera Forel, 1901e: 372 [as subgenus of Ectatomma]. Type-species: Ectatomma (Mictoponera) diehlii, by monotypy.

Taxonomic history

Mictoponera as subgenus of Ectatomma: Forel, 1901e: 372; Wheeler, W.M. 1910g: 135.

Mictoponera as junior synonym of Rhopalopone: Emery, 1911d: 34.

Mictoponera as junior synonym of Gnamptogenys: Brown, 1958g: 212; all subsequent authors.

Parectatomma Emery, 1911d: 44 [as subgenus of Ectatomma]. Type-species: Ectatomma triangulare, by original designation.

Taxonomic history

Parectatomma as subgenus of Ectatomma: Emery, 1911d: 44; Creighton, 1950a: 35; Kusnezov, 1956: 14.

Parectatomma as subgenus of Gnamptogenys: Forel, 1917: 236; Wheeler, W.M. 1922a: 643.

Parectatomma as junior synonym of Gnamptogenys: Brown, 1958g: 212; all subsequent authors.

Spaniopone Wheeler, W.M. & Mann, 1914: 11. Type-species: Spaniopone haytiana, by monotypy.

Taxonomic history

Spaniopone in Ponerinae, Proceratiini: Wheeler, W.M. & Mann, 1914: 13; Wheeler, W.M. 1916i: 36; Forel, 1917: 236; Wheeler, W.M. 1922a: 645; Donisthorpe, 1943g: 727.

Spaniopone in Ponerinae, Ectatommini: Emery, 1919c: 107.

Spaniopone as junior synonym of Gnamptogenys: Brown, 1958g: 212; all subsequent authors.

Wheeleripone Mann, 1919: 282. Type-species: Wheeleripone albiclava, by original designation.

Taxonomic history

Wheeleripone in Ponerinae, Ectatommini: Wheeler, W.M. 1922a: 642; Donisthorpe, 1943g: 736.

Wheeleripone as junior synonym of Gnamptogenys: Brown, 1958g: 212; all subsequent authors.

Opisthoscyphus Mann, 1922: 4. Type-species: Opisthoscyphus scabrosus (junior synonym of Alfaria minuta), by original designation.

Taxonomic history

Opisthoscyphus in Ponerinae, Ectatommini: Wheeler, W.M. 1930f: 13; Donisthorpe, 1943f: 678.

Opisthoscyphus as junior synonym of Alfaria: Brown, in Borgmeier, 1957: 115.

Opisthoscyphus as junior synonym of Gnamptogenys: Brown, 1958g: 212; all subsequent authors.

Commateta Santschi, 1929h: 476 [as subgenus of Ectatomma]. Type-species: Ectatomma (Parectatomma) bruchi, by original designation.

Taxonomic history

Commateta as junior synonym of Gnamptogenys: Brown, 1958g: 212; all subsequent authors.

Tammoteca Santschi, 1929h: 476 [as subgenus of Ectatomma]. Type-species: Ectatomma concinnum, by original designation.

Taxonomic history

Tammoteca as junior synonym of Gnamptogenys: Brown, 1958g: 212; all subsequent authors.

Barbourella Wheeler, W.M. 1930f: 10 [as subgenus of Emeryella]. Type-species: Emeryella (Barbourella) banksi, by original designation.

Taxonomic history

Barbourella as junior synonym of Gnamptogenys: Brown, 1958g: 212; all subsequent authors.

Genus Gnamptogenys references

Roger, 1863b: 19 (catalogue); Mayr, 1863: 422 (catalogue); Mayr, 1865: 15 (diagnosis); Mayr, 1870b: 963 (all species key); Mayr, 1887: 540 (diagnosis); Emery, 1889b: 494 (Ectatomma (Stictoponera) species key); Emery, 1896g: 43, 44 (G. (Holcoponera), G. (Gnamptogenys) species keys; Forel, 1900d: 316 (India & Sri Lanka species key); Bingham, 1903: 82 (India, Sri Lanka & Burma species key); Emery, 1911d: 34 (Rhopalopone diagnosis, catalogue); Emery, 1911d: 40 (Holcoponera diagnosis, catalogue); Emery, 1911d: 43 (Ectatomma (Poneracantha) diagnosis, catalogue); Emery, 1911d: 44 (Ectatomma (Parectatomma) & E. (Gnamptogenys) diagnoses, catalogues); Emery, 1911d: 46 (Emeryella diagnosis, catalogue); Emery, 1911d: 46 (Alfaria diagnosis, catalogue); Emery, 1911d: 47 (Stictoponera diagnosis, catalogue); Gallardo, 1918b: 22, 42 (Argentina species key); Mann, 1919: 286 (Wheeleripone species key); Wheeler, W.M. 1924b: 240 (Rhopalopone species key); Santschi, 1929h: 471 (Holcoponera species key); Chapman & Capco, 1951: 26, 29 (Asia Rhopalopone, Stictoponera checklists); Brown, 1954h: 1 (G. coxalis group species); Brown, 1956: 489 (Holcoponera species notes); Brown, 1958g: 211 (revision of genus, all species keys); Brown, 1958g: 230 (New World species revision, key); Brown, 1958g: 237 (Old World species revision, key); Kempf, 1972a: 111 (Neotropical catalogue); Smith, D.R. 1979: 1337 (North America catalogue); Taylor & Brown, D.R. 1985: 30 (Australia catalogue); Taylor, 1987a: 28 (Australia checklist); Brandão & Lattke, 1990: 489 (Alfaria group, status); Lattke, 1990b: 22 (Venezuela species key); Hölldobler & Wilson, 1990: 10 (synoptic classification); Brandão, 1991: 344 (Neotropical catalogue); Lattke, 1992a: 123 (G. minuta group, key); Bolton, 1995a: 1049 (census); Bolton, 1995b: 208 (catalogue); Lattke, 1995: 149 (New World species revision, key); Wu, J. & Wang, 1995: 36 (China species key); Xu & Zhang, 1996: 56 (China species key); Shattuck, 1999: 185 (Australia synopsis); Zhou, 2001b: 31 (China, Guangxi species key); Lattke, 2004: 51 (Old World species revision, key); Lattke, Fernández & Palacio, 2007: 257 (New World species key); Lattke, Fernández, et al. 2008: 68 (Colombia species key).

Myrmicinae Basicerotini Eurhopalothrix TRUE TRUE FALSE

38 species

Eurhopalothrix Brown & Kempf, 1961: 44. Type-species: Rhopalothrix bolaui, by original designation.

Taxonomic history

[Eurhopalothrix Brown & Kempf, 1960: 202; unavailable name, proposed without designation of type-species.]

Eurhopalothrix in Myrmicinae, Basicerotini: Brown & Kempf, 1961: 44; all subsequent authors except the entries below; Bolton, 1998: 67; Bolton, 2003: 184.

Eurhopalothrix in Myrmicinae, Dacetini: Baroni Urbani & De Andrade, 1994: 31; Baroni Urbani & De Andrade, 2007: 88 (as junior synonym of Basiceros).

Genus Eurhopalothrix references

Brown & Kempf, 1960: 202, 245 (diagnosis, all species revision, key); Snelling, R.R. 1968c: 3 (supplement to Brown & Kempf, 1960 key); Taylor, 1968b: 334 (Indo-Australian species key); Taylor, 1970a: 51 (supplement to Taylor, 1968b key); Kempf, 1972a: 107 (Neotropical catalogue); Smith, D.R. 1979: 1409 (North America catalogue); Taylor & Brown, D.R. 1985: 64 (Australia catalogue); Taylor, 1987a: 26 (Australia, New Caledonia checklist); Taylor, 1990b: 401 (Old World species key); Brandão, 1991: 343 (Neotropical catalogue); Bolton, 1995a: 1049 (census); Bolton, 1995b: 189 (catalogue); Shattuck, 1999: 135 (Australia synopsis); Ketterl, Verhaag & Dietz, 2004: 48 (Neotropical species key); Mezger & Pfeiffer, 2010: 135 (platisquama group key).

Myrmicinae Dacetini Pyramica TRUE TRUE FALSE

Extant: 344 species

Fossil: 4 species

Pyramica Roger, 1862a: 251. Type-species: Pyramica gundlachi, by monotypy.

Taxonomic history

Pyramica in Myrmicinae, Dacetini: Emery, 1924d: 320; all subsequent authors.

Pyramica as junior synonym of Strumigenys: Roger, 1863b: 40; Emery & Forel, 1879a: 464; Dalla Torre, 1893: 145; Bingham, 1903: 147; Emery, 1924d: 320; Donisthorpe, 1943g: 723; Brown & Wilson, 1959b: 281; Brown, 1960b: 37; Kempf, 1972a: 242; Bolton, 1983: 358; Bolton, 1994: 105; Baroni Urbani & De Andrade, 2007: 101.

Pyramica as subgenus of Strumigenys: Brown, 1948e: 110.

Pyramica as genus: Mayr, 1863: 453; Bolton, 1999: 1667; Bolton, 2000: 90; Eguchi, Bui & Yamane, 2011: 26.

Junior synonyms of PYRAMICA

Epitritus Emery, 1869a: 136. Type-species: Epitritus argiolus, by monotypy.

Taxonomic history

Epitritus in Myrmicidae, Cryptoceridae: Emery, 1877a: 81.

Epitritus in Myrmicinae: Emery & Forel, 1879a: 465 [Myrmicidae]; Dalla Torre, 1893: 148.

Epitritus in Cryptoceridae, Dacetonini: Ashmead, 1905b: 384.

Epitritus in Myrmicinae, Dacetini: Forel, 1892d: 344; Forel, 1893a: 164; Emery, 1895j: 770; Wheeler, W.M. 1910g: 141; Emery, 1914a: 42; Forel, 1917: 246; Arnold, 1917: 384; Wheeler, W.M. 1922a: 666; Emery, 1924d: 326; Donisthorpe, 1943f: 643; Brown, 1948e: 122; Brown, 1949b: 44; all subsequent authors.

Epitritus as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 12; Baroni Urbani & De Andrade, 2007: 101.

Epitritus as junior synonym of Pyramica: Bolton, 1999: 1667; Bolton, 2000: 90.

Trichoscapa Emery, 1869b: 24 [as subgenus of Strumigenys]. Type-species: Strumigenys (Trichoscapa) membranifera, by monotypy.

Taxonomic history

Trichoscapa in Myrmicinae: Emery & Forel, 1879a: 464 [Myrmicidae]; Dalla Torre, 1893: 145.

Trichoscapa in Myrmicinae, Dacetini: Forel, 1917: 246; Wheeler, W.M. 1922a: 668; Emery, 1924d: 323; Donisthorpe, 1943g: 734; Brown, 1948e: 112; all subsequent authors.

Trichoscapa as subgenus of Strumigenys: Emery & Forel, 1879a: 464; Forel, 1917: 246; Creighton, 1950a: 304.

Trichoscapa as junior synonym of Strumigenys: Dalla Torre, 1893: 145; Baroni Urbani & De Andrade, 1994: 13; Baroni Urbani & De Andrade, 2007: 101.

Trichoscapa as junior synonym of Cephaloxys: Wheeler, W.M. 1922a: 668; Emery, 1924d: 323; Donisthorpe, 1943g: 734.

Trichoscapa as genus: Brown, 1948e: 112; Kempf, 1972a: 255; Bolton, 1983: 319.

Trichoscapa as junior synonym of Pyramica: Bolton, 1999: 1667; Bolton, 2000: 90.

Pentastruma Forel, 1912a: 50. Type-species: Pentastruma sauteri, by monotypy.

Taxonomic history

Pentastruma in Myrmicinae, Dacetini: Emery, 1914a: 42; Forel, 1917: 246; Wheeler, W.M. 1922a: 666; Emery, 1924d: 326; Donisthorpe, 1943f: 682; Brown, 1948e: 102.

Pentastruma as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 15; Baroni Urbani & De Andrade, 2007: 101.

Pentastruma as junior synonym of Pyramica: Bolton, 1999: 1667; Bolton, 2000: 91.

Glamyromyrmex Wheeler, W.M. 1915e: 487. Type-species: Glamyromyrmex beebei, by monotypy.

Taxonomic history

Glamyromyrmex in Myrmicinae, Dacetini: Wheeler, W.M. 1915e: 491; Forel, 1917: 246; Wheeler, W.M. 1922a: 667; Emery, 1924d: 326; Donisthorpe, 1943f: 647; Brown, 1948e: 116; all subsequent authors.

Glamyromyrmex as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 14; Baroni Urbani & De Andrade, 2007: 101.

Glamyromyrmex as junior synonym of Pyramica: Bolton, 1999: 1667; Bolton, 2000: 91.

Codiomyrmex Wheeler, W.M. 1916d: 326. Type-species: Codiomyrmex thaxteri, by monotypy.

Taxonomic history

Codiomyrmex in Myrmicinae, Dacetini: Wheeler, W.M. 1922a: 667; Emery, 1924d: 325; Donisthorpe, 1943f: 633; Brown, 1948e: 114; all subsequent authors.

Codiomyrmex as subgenus of Strumigenys: Emery, 1924d: 325; Donisthorpe, 1943f: 633.

Codiomyrmex as genus: Wheeler, W.M. 1916d: 326; Brown, 1948e: 114.

Codiomyrmex as junior synonym of Glamyromyrmex: Brown, 1973b: 179 [provisional].

Codiomyrmex as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 14; Baroni Urbani & De Andrade, 2007: 101.

Codiomyrmex as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Tingimyrmex Mann, 1926: 104 [as subgenus of Strumigenys]. Type-species: Strumigenys (Tingimyrmex) mirabilis, by monotypy.

Taxonomic history

Tingimyrmex in Myrmicinae, Dacetini: Donisthorpe, 1943g: 732; Brown, 1948e: 111; all subsequent authors.

Tingimyrmex as subgenus of Strumigenys: Mann, 1926: 104; Donisthorpe, 1943g: 732.

Tingimyrmex as genus: Brown, 1948e: 111; Kempf, 1972a: 250.

Tingimyrmex as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 15; Baroni Urbani & De Andrade, 2007: 101.

Tingimyrmex as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Codioxenus Santschi, 1931c: 278 [as subgenus of Epitritus]. Type-species: Epitritus (Codioxenus) simulans, by monotypy.

Taxonomic history

Codioxenus in Myrmicinae, Dacetini: Donisthorpe, 1943f: 633; Brown, 1948e: 123; all subsequent authors.

Codioxenus as genus: Brown, 1948e: 123.

Codioxenus as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 14; Baroni Urbani & De Andrade, 2007: 101.

Codioxenus as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Smithistruma Brown, 1948e: 104.

Taxonomic history

[Replacement name for Cephaloxys Smith, F. 1865: 76; junior homonym of Cephaloxys Signoret, 1847: 294 (Hemiptera). Type-species not Strumigenys pulchella, unjustified subsequent designation by Brown, 1948e: 104, repeated in Kempf, 1972a: 230, Bolton, 1983: 274 and Bolton, 1995b: 46.]

Smithistruma in Myrmicinae, Dacetini: Brown, 1948e: 104.

Smithistruma as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 14; Baroni Urbani & De Andrade, 2007: 102.

Smithistruma as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Homonym replaced by Smithistruma

Cephaloxys Smith, F. 1865: 76. Type-species: Cephaloxys capitata, by monotypy.

Taxonomic history

[Cephaloxys Smith, F. junior homonym of Cephaloxys Signoret, 1847: 294 (Hemiptera).]

Cephaloxys in Cryptoceridae: Smith, F. 1871a: 335.

Cephaloxys in Myrmicinae, Dacetini: Wheeler, W.M. 1922a: 668.

Cephaloxys as junior synonym of Strumigenys: Dalla Torre, 1893: 15.

Cephaloxys as subgenus of Strumigenys: Wheeler, W.M. 1922a: 668; Emery, 1924d: 323; Bernard, 1953b: 254 (anachronism).

Weberistruma Brown, 1948e: 106 [as subgenus of Smithistruma]. Type-species: Strumigenys (Cephaloxys) leptothrix, by original designation.

Taxonomic history

Weberistruma in Myrmicinae, Dacetini: Brown, 1948e: 106.

Weberistruma as genus: Brown, 1949d: 7; Brown, 1953g: 24.

Weberistruma as junior synonym of Smithistruma: Brown, 1973a: 35.

Weberistruma as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Weberistruma as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 2007: 102.

Wessonistruma Brown, 1948e: 106 [as subgenus of Smithistruma]. Type-species: Strumigenys pergandei, by original designation.

Taxonomic history

Wessonistruma in Myrmicinae, Dacetini: Brown, 1948e: 106.

Wessonistruma junior synonym of Smithistruma: Brown, 1973a: 35.

Wessonistruma as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Wessonistruma as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 2007: 102.

Serrastruma Brown, 1948e: 107 [as subgenus of Smithistruma]. Type-species: Strumigenys simoni, by original designation.

Taxonomic history

Serrastruma in Myrmicinae, Dacetini: Brown, 1948e: 107, and all subsequent authors.

Serrastruma as genus: Brown, 1949d: 6; Brown, 1952e: 70; Bolton, 1983: 335; Bolton, 1994: 105.

Serrastruma as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 14; Baroni Urbani & De Andrade, 2007: 102.

Serrastruma as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Neostruma Brown, 1948e: 111. Type-species: Strumigenys crassicornis, by original designation.

Taxonomic history

Neostruma in Myrmicinae, Dacetini: Brown, 1948e: 111; all subsequent authors.

Neostruma as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 12; Baroni Urbani & De Andrade, 2007: 102.

Neostruma as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Dorisidris Brown, 1948e: 116. Type-species: Strumigenys (Codiomyrmex) nitens, by original designation.

Taxonomic history

Dorisidris in Myrmicinae, Dacetini: Brown, 1948e: 116; all subsequent authors.

Dorisidris as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 14; Baroni Urbani & De Andrade, 2007: 102.

Dorisidris as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Miccostruma Brown, 1948e: 123. Type-species: Epitritus mandibularis, by original designation.

Taxonomic history

Miccostruma in Myrmicinae, Dacetini: Brown, 1948e: 123.

Miccostruma as junior synonym of Smithistruma: Bolton, 1983: 274.

Miccostruma as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Miccostruma as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 2007: 102.

Kyidris Brown, 1949d: 3. Type-species: Kyidris mutica, by original designation.

Taxonomic history

Kyidris in Myrmicinae, Dacetini: Brown, 1949d: 3.

Kyidris as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 15; Baroni Urbani & De Andrade, 2007: 103.

Kyidris as junior synonymof Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Chelystruma Brown, 1950a: 33 [as subgenus of Glamyromyrmex]. Type-species: Glamyromyrmex (Chelystruma) lilloana, by monotypy.

Taxonomic history

Chelystruma in Myrmicinae, Dacetini: Brown, 1950a: 33; all subsequent authors.

Chelystruma as subgenus of Glamyromyrmex: Brown, 1953g: 16.

Chelystruma as genus: Kempf, 1959d: 338; Kempf, 1972a: 77.

Chelystruma as junior synonym of Glamyromyrmex: Brown, 1973b: 179 [provisional].

Chelystruma as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 14; Baroni Urbani & De Andrade, 2007: 103.

Chelystruma as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Polyhomoa Azuma, 1950: 36. Type-species: Polyhomoa itoi (junior synonym of Kyidris mutica), by monotypy.

Taxonomic history

Polyhomoa as junior synonym of Kyidris: Creighton, 1950b: 93; Brown & Yasumatsu, 1951: 93.

Polyhomoa as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Polyhomoa as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 2007: 103.

Borgmeierita Brown, 1953g: 23. Type-species: Codiomyrmex excisus, by original designation.

Taxonomic history

Borgmeierita in Myrmicinae, Dacetini: Brown, 1953g: 23.

Borgmeierita as junior synonym of Glamyromyrmex: Brown, 1973a: 35.

Borgmeierita as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Borgmeierita as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 2007: 103.

Platystruma Brown, 1953g: 112 [as subgenus of Smithistruma]. Type-species: Strumigenys (Cephaloxys) depressiceps, by original designation.

Taxonomic history

Platystruma in Myrmicinae, Dacetini: Brown, 1953g: 112; all subsequent authors.

Platystruma as junior synonym of Smithistruma: Brown, 1973a: 35.

Platystruma as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Platystruma as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 2007: 103.

Gymnomyrmex Borgmeier, 1954b: 279. Type-species: Gymnomyrmex splendens, by original designation.

Taxonomic history

Gymnomyrmex in Myrmicinae, Dacetini: Borgmeier, 1954b: 279; all subsequent authors.

Gymnomyrmex as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 14; Baroni Urbani & De Andrade, 2007: 103.

Gymnomyrmex as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Dysedrognathus Taylor, 1968d: 132. Type-species: Dysedrognathus extemenus, by original designation.

Taxonomic history

Dysedrognathus in Myrmicinae, Dacetini: Taylor, 1968d: 132.

Dysedrognathus as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 13; Baroni Urbani & De Andrade, 2007: 103.

Dysedrognathus as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Asketogenys Brown, 1972: 23. Type-species: Asketogenys acubecca, by original designation.

Taxonomic history

Asketogenys in Myrmicinae, Dacetini: Brown, 1972: 23.

Asketogenys as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 15; Baroni Urbani & De Andrade, 2007: 103.

Asketogenys as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Cladarogenys Brown, 1976b: 33. Type-species: Cladarogenys lasia, by original designation.

Taxonomic history

Cladarogenys in Myrmicinae, Dacetini: Brown, 1976b: 33.

Cladarogenys as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 14; Baroni Urbani & De Andrade, 2007: 103.

Cladarogenys as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Genus Pyramica references

Mayr, 1863: 453 (catalogue); André, 1883b: 398, 402 (Europe & Algeria Strumigenys (Trichoscapa), Epitritus species); Dalla Torre, 1893: 145, 148 (Strumigenys, Epitritus catalogues); Emery, 1895c: 325 (North America species key); Wheeler, W.M. 1908a: 146 (New World species key); Santschi, 1913b: 257 (Afrotropical species key); Emery, 1916b: 205 (Italy species key); Arnold, 1917: 373 (South Africa species key); Wheeler, W.M. 1922a: 918, 920 (Afrotropical Strumigenys (Cephaloxys), Epitritus catalogues); Emery, 1924d: 323, 325, 326 (Strumigenys (Cephaloxys), Strumigenys (Codiomyrmex), Glamyromyrmex, Pentastruma, Epitritus diagnoses, catalogues); Smith, M.R. 1931c: 691 (North America Strumigenys (Cephaloxys) species key); Buren, 1944a: 290 (U.S.A., Iowa species key); Brown, 1948e: 112 - 124 (minor genera reviews); Brown, 1949b: 43 (Epitritus diagnosis, review); Brown, 1949d: 8 (Weberistruma species, review); Brown, 1949d: 21 (Japan, China & Taiwan species key); Creighton, 1950a: 301 (North America Strumigenys (Trichoscapa) species key); Chapman & Capco, 1951: 104, 105, 106 (Asia Epitritus, Kyidris, Pentastruma, Smithistruma, Strumigenys (Cephaloxys) checklists); Brown, 1950a: 27 (Glamyromyrmex review); Brown, 1952e: 86 (Serrastruma diagnosis, species revision, key); Brown, 1953g: 20, 24 (Codiomyrmex, Weberistruma reviews); Brown, 1953g: 31, 46, 92, 114, 125, 130 (Smithistruma diagnosis, Nearctic, Neotropical, Malesian, Afrotropical, Palaearctic species revisions, keys); Wilson & Brown, 1956b: 439 (Kyidris species review); Brown, 1959b: 1 (Neostruma species review); Kempf, 1959d: 339 (Gymnomyrmex species key); Brown, 1960b: 37 (Strumigenys gundlachi group revision, key); Kempf, 1960f: 447 (Neotropical Glamyromyrmex species key); Kempf, 1960f: 451 (Gymnomyrmex species key); Brown, 1962b: 79 (Epitritus species key); Brown, 1964a: 183 (supplement to 1953e Smithistruma revision); Bernard, 1967: 243, 245 (Smithistruma, Epitritus diagnoses); Kempf, 1972a: 37, 77, 98, 110, 116, 163, 230, 250, 255 (Neotropical Borgmeierita, Chelystruma, Codiomyrmex, Codioxenus, Dorisidris, Glamyromyrmex, Gymnomyrmex, Neostruma, Smithistruma, Tingimyrmex, Trichoscapa catalogues); Bolton, 1972: 208 (Epitritus species key); Alayo, 1974: 19 (Cuba species key); Smith, D.R. 1979: 1405, 1408 (North America Smithistruma, Trichoscapa catalogues); Taylor, 1987a: 28 (Australia checklist); Brown & Boisvert, 1979: 201 (Pentastruma species review); Bolton, 1983: 274, 319, 320, 335, 353, 354 (Afrotropical Smithistruma, Trichoscapa, Glamyromyrmex, Serrastruma, Cladarogenys, Epitritus species revisions, keys); Taylor & Brown, D.R. 1985: 64 (Australia Glamyromyrmex catalogue); Perrault, 1986a: 3 (Gymnomyrmex species key); Ward, 1988: 122 (Western Nearctic Smithistruma species key); Terayama & Kubota, 1989: 787 (Taiwan Smithistruma species key); Hölldobler & Wilson, 1990: 15 (synoptic classification); Brandão, 1991: 344, 346, 360, 378 (Neotropical catalogue); Arakelian & Dlussky, 1991: 150 (former U.S.S.R. species key); Morisita, Kubota, et al. 1992: 68 (Japan Smithistruma species key); Morisita, Kubota, et al. 1992: 76 (Japan Epitritus species key); Bolton, 1994: 105 (synoptic classification); Bolton, 1995a: 1048 - 1053 (census); Bolton, 1995b: 75, 145, 146, 177, 183, 188, 207, 211, 219, 292, 316, 382, 383, 420, 421 (catalogue); Terayama, Lin & Wu, 1996: 328 (Taiwan Smithistruma species key); Ogata & Onoyama, 1998: 277 (Japan Smithistruma species key); Shattuck, 1999: 136, 170 (Australia Glamyromyrmex, Trichoscapa synopses); Bolton, 1999: 1657, 1667 (morphology, diagnosis, review of genus, phylogeny); Xu, 2000e: 297 (China Epitritus species key); Bolton, 2000: 95, 137, 272, 285, 342, 372, 472 (Nearctic, Neotropical, Afrotropical, West Palaearctic, Malagasy, Malesian-Oriental-East Palaearctic, Austral species revisions, keys); Xu & Zhou, 2004: 441 (China species, key); Longino, 2006b: 139 (Costa Rica species, additions to key); Terayama, 2009: 138 (Taiwan species key).

Myrmicinae Dacetini Strumigenys TRUE TRUE FALSE

Extant: 487 species

Fossil:

Strumigenys Smith, F. 1860c: 72. Type-species: Strumigenys mandibularis, by monotypy.

Taxonomic history

Strumigenys in Myrmicidae: Smith, F. 1871a: 334; Cresson, 1887: 262.

Strumigenys in Myrmicidae, Cryptoceridae: Emery, 1877a: 81.

Strumigenys in Myrmicinae: Mayr, 1865: 26 [Myrmicidae]; Emery & Forel, 1879a: 464 [Myrmicidae]; Dalla Torre, 1893: 145.

Strumigenys in Cryptoceridae, Dacetini: Ashmead, 1905b: 384.

Strumigenys in Myrmicinae, Dacetini: Forel, 1892d: 344; Forel, 1893a: 164; Forel, 1895b: 133; Emery, 1895j: 770; Forel, 1899c: 42; Wheeler, W.M. 1910g: 141; Emery, 1914a: 42; Forel, 1917: 246; Arnold, 1917: 372; Wheeler, W.M. 1922a: 668; Emery, 1924d: 319; Brown, 1948e: 108; all subsequent authors.

Junior synonyms of STRUMIGENYS

Labidogenys Roger, 1862a: 249. Type-species: Labidogenys lyroessa, by monotypy.

Taxonomic history

Labidogenys in Myrmicinae, Dacetini: Donisthorpe, 1943f: 654; Brown, 1948e: 102.

Labidogenys as genus: Roger, 1862a: 249; Brown, 1948e: 102.

Labidogenys as junior synonym of Strumigenys: Roger, 1863b: 40; Emery & Forel, 1879a: 464; Dalla Torre, 1893: 145; Bingham, 1903: 147; Emery, 1924d: 320; Brown & Wilson, 1959b: 281; Brown, 1960b: 38; Bolton, 1983: 358; Bolton, 1999: 1673; Bolton, 2000: 492.

Proscopomyrmex Patrizi, 1946: 294. Type-species: Proscopomyrmex londianensis, by monotypy.

Taxonomic history

Proscopomyrmex in Myrmicinae, Dacetini: Patrizi, 1946: 294 [Dacetonini].

Proscopomyrmex as junior synonym of Strumigenys: Brown, 1949d: 15; Bolton, 1999: 1673; Bolton, 2000: 492.

Eneria Donisthorpe, 1948d: 598. Type-species: Eneria excisa (junior synonym of Strumigenys loriae), by original designation.

Taxonomic history

Eneria as genus: Chapman & Capco, 1951: 104 (anachronism).

Eneria as junior synonym of Strumigenys: Brown, 1949d: 15; Bolton, 1999: 1673; Bolton, 2000: 492.

Quadristruma Brown, 1949b: 47. Type-species: Epitritus emmae, by original designation.

Taxonomic history

Quadristruma in Myrmicinae, Dacetini: Brown, 1949b: 47; all subsequent authors.

Quadristruma as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 12; Bolton, 1999: 1673; Bolton, 2000: 492.

Genus Strumigenys references

Roger, 1863b: 40 (catalogue); Mayr, 1863: 424, 455 (catalogue); Mayr, 1865: 26 (diagnosis); Forel, 1886a: 216 (all species key); Mayr, 1887: 568 (all species key); Dalla Torre, 1893: 145 (catalogue); Emery, 1895c: 325 (North America species key); Emery, 1897c: 574 (New Guinea species key); Forel, 1903a: 707 (India & Sri Lanka species key); Bingham, 1903: 147 (India, Sri Lanka & Burma species key); Wheeler, W.M. 1908a: 146 (New World species key); Santschi, 1913b: 257 (Afrotropical species key); Arnold, 1917: 373 (diagnosis, South Africa species key); Mann, 1921: 467 (Fiji Is species key); Wheeler, W.M. 1922a: 917, 1034 (Afrotropical, Malagasy catalogues); Emery, 1924d: 319 (diagnosis, catalogue); Brown, 1948e: 108 (diagnosis, review of genus); Brown, 1949d: 21 (Japan, China & Taiwan species key); Creighton, 1950a: 301 (North America species key); Chapman & Capco, 1951: 104, 107 (Asia Eneria, Strumigenys (Strumigenys) checklists); Brown, 1954k: 10 (Afrotropical species revision, key); Brown, 1958h: 38 (New Zealand species); Brown, 1962c: 238 (Neotropical species synopsis, key); Kempf, 1972a: 226, 243 (Neotropical Quadristruma, Strumigenys catalogues); Alayo, 1974: 19 (Cuba species key); Kempf, 1976b: 43 (additions to Brown, 1962c key); Smith, D.R. 1979: 1403, 1409 (North America Strumigenys, Quadristruma catalogues); Bolton, 1983: 358, 400 (Afrotropical Strumigenys, Quadristruma species revisions, keys); Taylor & Brown, D.R. 1985: 85, 87 (Australia Quadristruma, Strumigenys catalogues); Taylor, 1987a: 66, 76 (Australia, New Caledonia & New Zealand checklists); Terayama & Kubota, 1989: 780 (Taiwan species key); Brandão, 1991: 380 (Neotropical catalogue); Arakelian & Dlussky, 1991: 150 (former U.S.S.R. species key); Morisita, Kubota, et al. 1992: 63 (Japan species key); Dlussky, 1993: 53 (Fiji Is, Tonga and Samoa species key); Bolton, 1994: 105 (synoptic classification); Bolton, 1995a: 1053 (census); Bolton, 1995b: 377, 395 (catalogue); Gronenberg, 1996: 2021 (mandible morphology); Lin & Wu, 1996: 139 (Taiwan species key); Lattke & Goitía, 1997: 370 (Venezuela species key); Shattuck, 1999: 160, 167 (Australia synopsis); Bolton, 1999: 1657, 1673 (morphology, diagnosis, review of genus, phylogeny); Bolton, 2000: 496, 583, 716, 942 (Neotropical-Nearctic, Afrotropical, Malesian-Oriental-East Palaearctic, Austral species revisions, keys); Fisher, 2000: 614 (Malagasy species revision, key); Zhou, 2001b: 78 (China, Guangxi species key); Lin & Wu, 2001: 166 (Taiwan species key); Zhou & Xu, 2003: 737 (China species key); Yoshimura & Onoyama, 2007: 676 (Japan species key); Terayama, 2009: 142 (Taiwan species key); Heterick, 2009: 176 (south-western Australia species key).

Myrmicinae Cephalotini Cephalotes TRUE TRUE FALSE

Extant: 115 species

Fossil: 16 species

Cephalotes Latreille, 1802a: 357. Type-species: Formica atrata, by monotypy.

Taxonomic history

[Type-species not Formica cephalotes, unjustified subsequent designation by Wheeler, W.M. 1911f: 160; corrected by Wheeler, W.M. 1913a: 78.]

Cephalotes in Myrmicinae, Cryptocerini: Emery, 1914a: 42; Forel, 1917: 246; Wheeler, W.M. 1922a: 665; Emery, 1924d: 303; all subsequent authors to 1949, and Dlussky & Fedoseeva, 1988: 79 (anachronism).

Cephalotes in Myrmicinae, Cephalotini: Smith, M.R. 1949c: 19; Kempf, 1951: 105; all subsequent authors except the above.

Cephalotes as junior synonym of Cryptocerus: Fabricius, 1804: 419.

Junior synonyms of CEPHALOTES

Cryptocerus Latreille, 1803: 311. Type-species: Formica atrata, by subsequent designation of Latreille, 1810: 437.

Taxonomic history

[Type-species not Cryptocerus umbraculatus, unjustified subsequent designation by Emery, 1914c: 38, repeated in Emery, 1924d: 305.]

Cryptocerus in Myrmicites: Lepeletier de Saint-Fargeau, 1835: 170.

Cryptocerus in Myrmicidae, Cryptoceridae: Smith, F. 1853: 214; Emery, 1877a: 81.

Cryptocerus in Poneridae, Cryptoceridae: Smith, F. 1858b: 187.

Cryptocerus in Cryptoceridae: Smith, F. 1876d: 605.

Cryptocerus in Cryptoceridae, Cryptocerinae: Ashmead, 1905b: 384.

Cryptocerus in Myrmicinae: Mayr, 1865: 25 [Myrmicidae]; Mayr, 1866b: 907 [Myrmicidae]; Dalla Torre, 1893: 140.

Cryptocerus in Myrmicinae, Cryptocerini: Forel, 1892d: 345; Forel, 1893a: 164; Forel, 1895b: 133; Emery, 1895j: 771; Forel, 1899c: 48; Wheeler, W.M. 1910g: 141; Emery, 1914a: 42; Forel, 1917: 246; Wheeler, W.M. 1922a: 665; Emery, 1924d: 305; Wheeler, G.C. & Wheeler, J. 1976b: 59.

Cryptocerus as junior synonym of Cephalotes: Wheeler, W.M. 1913a: 78; Donisthorpe, 1943f: 635; Smith, M.R. 1949c: 19; Kempf, 1951: 105; all subsequent authors except the above. [Cephalotes and Cryptocerus share the same type-species, synonymy is therefore absolute.]

[Cryptocephalus Lowne, 1865b: 336, incorrect subsequent spelling of Cryptocerus in the combination Cryptocephalus pubescens; current combination Meranoplus pubescens: Bolton, 1981a: 47.]

Zacryptocerus Wheeler, W.M. 1911f: 175 (see also footnote). Type-species: Cryptocerus clypeatus, by original designation.

Taxonomic history

[Zacryptocerus Ashmead, 1905b: 384, nomen nudum. (Based on a non-existent type-species: Cryptocerus multistrigus, nomen nudum, attributed to Smith, F.).]

Zacryptocerus in Myrmicinae, Cryptocerini: Forel, 1917: 246; Wheeler, W.M. 1922a: 665; Emery, 1924d: 304; all subsequent authors except the following; Dlussky & Fedoseeva, 1988: 79 (anachronism).

Zacryptocerus in Myrmicinae, Cephalotini: Kempf, 1951: 133; all subsequent authors.

Zacryptocerus as junior synonym of Cephalotes: De Andrade & Baroni Urbani, 1999: 59.

Paracryptocerus Emery, 1915i: 192 [as subgenus of Cryptocerus]. Type-species: Cryptocerus spinosus, by original designation.

Taxonomic history

Paracryptocerus in Myrmicinae, Cryptocerini: Forel, 1917: 246; Wheeler, W.M. 1922a: 665; Emery, 1924d: 306; Donisthorpe, 1943f: 680.

Paracryptocerus in Myrmicinae, Cephalotini: Smith, M.R. 1949c: 20; Kempf, 1951: 153; all subsequent authors.

Paracryptocerus as subgenus of Cryptocerus: Emery, 1915i: 192; Forel, 1917: 246; Wheeler, W.M. 1922a: 665; Emery, 1924d: 306.

Paracryptocerus as genus: Smith, M.R. 1949c: 20; Kempf, 1951: 153; Kempf, 1972a: 175.

Paracryptocerus as junior synonym of Zacryptocerus: Kempf, 1973c: 460; Smith, D.R. 1979: 1402.

Hypocryptocerus Wheeler, W.M. 1920: 53 [as subgenus of Cryptocerus]. Type-species: Formica haemorrhoidalis, by original designation.

Taxonomic history

Hypocryptocerus in Myrmicinae, Cryptocerini: Wheeler, W.M. 1922a: 665; Wheeler, W.M. 1936b: 200; Donisthorpe, 1943f: 652.

Hypocryptocerus in Myrmicinae, Cephalotini: Kempf, 1951: 143.

Hypocryptocerus as subgenus of Cryptocerus: Wheeler, W.M. 1920: 53; Wheeler, W.M. 1922a: 665.

Hypocryptocerus as genus: Wheeler, W.M. 1936b: 200; Kempf, 1951: 143; Kempf, 1972a: 120.

Hypocryptocerus as junior synonym of Zacryptocerus: Kempf, 1973c: 460; Smith, D.R. 1979: 1402.

Cyathomyrmex Creighton, 1933: 100 [as subgenus of Cryptocerus].

Taxonomic history

[Replacement name for Cyathocephalus Emery; junior homonym of Cyathocephalus Kessler, 1868: 135 (Cestoda).]

Cyathomyrmex in Myrmicinae, Cephalotini: Smith, M.R. 1949c: 21.

Cyathomyrmex as subgenus of Cryptocerus: Creighton, 1933: 100; Creighton, 1950a: 298.

Cyathomyrmex as subgenus of Paracryptocerus: Smith, M.R. 1949c: 21; Kempf, 1951: 156.

Cyathomyrmex as junior synonym of Paracryptocerus: Kempf, 1972a: 175.

Homonym replaced by Cyathomyrmex

Cyathocephalus Emery, 1915i: 192 [as subgenus of Cryptocerus]. Type-species: Cryptocerus pallens, by original designation.

Taxonomic history

[Cyathocephalus Emery junior homonym of Cyathocephalus Kessler, 1868: 135 (Cestoda).]

Cyathocephalus as subgenus of Cryptocerus: Emery, 1915i: 192; Forel, 1917: 246; Wheeler, W.M. 1922a: 665; Emery, 1924d: 311.

Harnedia Smith, M.R. 1949c: 20 [as subgenus of Paracryptocerus]. Type-species: Cryptocerus umbraculatus, by original designation.

Taxonomic history

Harnedia in Myrmicinae, Cephalotini: Smith, M.R. 1949c: 20; Kempf, 1951: 233.

Harnedia as subgenus of Paracryptocerus: Smith, M.R. 1949c: 20; Kempf, 1951: 233.

Harnedia as junior synonym of Paracryptocerus: Kempf, 1972a: 175.

Eucryptocerus Kempf, 1951: 127. Type-species: Cryptocerus oculatus, by original designation.

Taxonomic history

Eucryptocerus in Myrmicinae, Cephalotini: Kempf, 1951: 127; all subsequent authors except the following.

Eucryptocerus in Myrmicinae, Cryptocerini: Dlussky & Fedoseeva, 1988: 79 (anachronism).

Eucryptocerus as genus: Kempf, 1972a: 107; Kempf, 1973c: 460; Bolton, 1994: 105.

Eucryptocerus as junior synonym of Cephalotes: Brown, 1973b: 180 [provisional]; De Andrade & Baroni Urbani, 1999: 59.

†Exocryptocerus Vierbergen & Scheven, 1995: 159. Type-species: †Exocryptocerus serratus, by original designation.

Taxonomic history

†Exocryptocerus as junior synonym of Cephalotes: De Andrade & Baroni Urbani, 1999: 59.

Genus Cephalotes references

Smith, F. 1853: 215 (diagnosis); Smith, F. 1858b: 187 (diagnosis); Roger, 1863b: 38 (catalogue); Mayr, 1863: 405 (catalogue); Mayr, 1865: 25 (diagnosis); Dalla Torre, 1893: 140 (catalogue); Santschi, 1920f: 149 (species key); Wheeler, W.M. 1922a: 665 (Cryptocerus subgenera, key); Emery, 1924d: 303 (diagnosis, catalogue); Emery, 1924d: 304 (Zacryptocerus diagnosis, catalogue); Emery, 1924d: 305 (Cryptocerus diagnosis, subgenera key, catalogue); Emery, 1924d: 306 (C. (Paracryptocerus) diagnosis, catalogue); Emery, 1924d: 308 (C. (Cryptocerus) diagnosis, catalogue); Emery, 1924d: 311 (C. (Cyathocephalus) diagnosis, catalogue); Smith, M.R. 1947a: 29 (U.S.A. species key); Creighton, 1950a: 296 (North America species key); Kempf, 1951: 107 (diagnosis, revision, key); Kempf, 1951: 127 (Eucryptocerus species revision, key); Kempf, 1951: 134 (Zacryptocerus species revision, key); Kempf, 1951: 143 (Hypocryptocerus species revision, key); Kempf, 1951: 153 (Paracryptocerus species revision, key); Kempf, 1952: 5 (Paracryptocerus (Harnedia) pinellii complex revision, key); Kempf, 1958a: 10 (Argentina Paracryptocerus species key); Kempf, 1958a: 64 (Paracryptocerus subgenera, key); Kempf, 1958a: 66 (Paracryptocerus (Harnedia) species revision, key); Kempf, 1958a: 145 (Paracryptocerus (Cyathomyrmex) species revision, key); Kempf, 1960f: 443 (addition to 1958 key); Snelling, R.R. 1968a: 9 (C. wheeleri complex, key); Kempf, 1972a: 75, 107, 120, 175, 259 (Neotropical Cephalotes, Eucryptocerus, Hypocryptocerus, Paracryptocerus, Zacryptocerus catalogues); Kempf, 1974a: 73 (Paracryptocerus pinelii group, key); Smith, D.R. 1979: 1402 (North America catalogue); Hölldobler & Wilson, 1990: 13 (synoptic classification); Brandão, 1991: 337, 343, 383 (catalogue); Bolton, 1994: 105 (synoptic classification); Bolton, 1995a: 1048 (census); Bolton, 1995b: 140, 189, 424 (catalogue); De Andrade & Baroni Urbani, 1999: 736 (diagnosis, all species revision, key).

Myrmicinae Attini Acromyrmex TRUE TRUE FALSE

30 species, 30 subspecies

Acromyrmex Mayr, 1865: 83 [as subgenus of Atta]. Type-species: Formica hystrix, by monotypy.

Taxonomic history

Acromyrmex in Cryptoceridae, Attinae: Ashmead, 1905b: 384.

Acromyrmex in Myrmicinae, Attini: Emery, 1895j: 770; Emery, 1913b: 251; Emery, 1914a: 42; Forel, 1917: 247; Wheeler, W.M. 1922a: 669; Emery, 1924d: 347; all subsequent authors.

Acromyrmex as subgenus of Atta: Mayr, 1865: 83; Forel, 1885a: 354; Dalla Torre, 1893: 151; Forel, 1893a: 163; Emery, 1895j: 770; Forel, 1899c: 30; Emery, 1905c: 39; Wheeler, W.M. 1907c: 670; Wheeler, W.M. 1910g: 141; Wheeler, W.M. 1922a: 669.

Acromyrmex as genus: Emery, 1913b: 251; Forel, 1917: 247; Emery, 1924d: 347; Santschi, 1925a: 355; all subsequent authors.

Subgenera of ACROMYRMEX include the nominal plus the following.

Subgenus ACROMYRMEX (MOELLERIUS)

Moellerius Forel, 1893e: 589 [as subgenus of Atta]. Type-species: Atta (Acromyrmex) landolti, by subsequent designation of Wheeler, W.M. 1911f: 167.

Taxonomic history

Moellerius as subgenus of Atta: Forel, 1893e: 589; Forel, 1899c: 30; Emery, 1905c: 39; Wheeler, W.M. 1907c: 669; Wheeler, W.M. 1910g: 141; Wheeler, W.M. 1922a: 669.

Moellerius as junior synonym of Acromyrmex: Brown, 1973b: 182 [provisional].

Moellerius as subgenus of Acromyrmex: Emery, 1913b: 251; Forel, 1917: 247; Emery, 1924d: 350; all subsequent authors except the above.

Genus Acromyrmex references

Forel, 1885a: 361 (all species key); Emery, 1905c: 40 (all species key); Gallardo, 1916d: 325 (Argentina species key); Emery, 1924d: 347 (diagnosis, catalogue); Emery, 1924d: 350 (A. (Moellerius) diagnosis, catalogue); Santschi, 1925a: 388 (all species revision, key); Creighton, 1950a: 326 (North America species key); Kusnezov, 1956: 33 (Argentina species key); Gonçalves, 1961: 118 (Brazil species revision, key); Kempf, 1972a: 10 (Neotropical catalogue); Alayo, 1974: 42 (Cuba checklist); Zolessi & Abenante, 1977: 73 (Uruguay species, male genitalia); Smith, D.R. 1979: 1412 (North America catalogue); Fowler, 1988: 284 (A. (Moellerius) species key); Cherrett & Cherrett, 1989: 1 (bibliography); Bolton, 1994: 105 (synoptic classification); Bolton, 1995a: 1047 (census); Bolton, 1995b: 54 (catalogue).

Myrmicinae Attini Atta TRUE TRUE FALSE

15 species

Atta Fabricius, 1804: 421. Type-species: Formica cephalotes, by subsequent designation of Wheeler, W.M. 1911f: 159.

Taxonomic history

Atta in Myrmicites: Lepeletier de Saint-Fargeau, 1835: 172.

Atta in Poneridae, Attidae: Smith, F. 1858b: 161.

Atta in Myrmicinae: Mayr, 1855: 459 [Myrmicidae]; Smith, F. 1857a: 77 [Myrmicidae]; Mayr, 1861: 65 [Myrmicidae]; Mayr, 1865: 18 [Myrmicidae]; Smith, F. 1871a: 333 [Myrmicidae]; Cresson, 1887: 259 [Myrmicidae]; Dalla Torre, 1893: 150.

Atta in Cryptoceridae, Attinae: Ashmead, 1905b: 384.

Atta in Myrmicinae, Attini: Emery, 1877a: 81 [Myrmicidae, Attidae]; Forel, 1893a: 163; Emery, 1895j: 770; Forel, 1899c: 30; Wheeler, W.M. 1910g: 141; Emery, 1914a: 42; Forel, 1917: 247; Wheeler, W.M. 1922a: 669; Emery, 1924d: 352; all subsequent authors.

[Myrmegis (nomen nudum) referable to Atta: Brown, 1973b: 182 (see under Myrmegis).]

Junior synonyms of ATTA

Oecodoma Latreille, 1818: 222. Type-species: Formica cephalotes, by subsequent designation of Shuckard, in Swainson & Shuckard, 1840: 174.

Taxonomic history

Oecodoma in Myrmicites: Lepeletier de Saint-Fargeau, 1835: 176.

Oecodoma in Poneridae, Attidae: Smith, F. 1858b: 180.

Oecodoma as junior synonym of Atta: Roger, 1863b: 35. [Atta and Oecodoma share the same type-species, synonymy is therefore absolute.]

Archeatta Gonçalves, 1942: 342 [as subgenus of Atta]. Type-species: Oecodoma mexicana, by original designation.

Taxonomic history

Archeatta as subgenus of Atta: Gonçalves, 1942: 342; Borgmeier, 1959b: 350; Kempf, 1972a: 27.

Archeatta as junior synonym of Atta: Smith, M.R. 1951a: 832; Weber, 1958a: 8; Brown, 1973b: 178 [provisional]; Bolton, 1994: 105.

Neoatta Gonçalves, 1942: 346 [as subgenus of Atta]. Type-species: Formica sexdens, by original designation.

Taxonomic history

Neoatta as subgenus of Atta: Gonçalves, 1942: 346; Borgmeier, 1959b: 358; Kempf, 1972a: 27.

Neoatta as junior synonym of Atta: Weber, 1958a: 8; Brown, 1973b: 183 [provisional]; Bolton, 1994: 105.

Palaeatta Borgmeier, 1950d: 244 [as subgenus of Atta]. Type-species: Atta bisphaerica, by original designation.

Taxonomic history

Palaeatta as subgenus of Atta: Borgmeier, 1950d: 244; Borgmeier, 1959b: 377; Kempf, 1972a: 28.

Palaeatta as junior synonym of Atta: Weber, 1958a: 8; Brown, 1973b: 183 [provisional]; Bolton, 1994: 105.

Epiatta Borgmeier, 1950d: 246 [as subgenus of Atta]. Type-species: Oecodoma laevigata, by original designation.

Taxonomic history

Epiatta as junior synonym of Atta: Weber, 1958a: 8.

Epiatta as junior synonym of Neoatta: Borgmeier, 1959b: 358.

Genus Atta references

Mayr, 1855: 459 (diagnosis); Smith, F. 1858b: 161, 180 (Atta, Oecodoma diagnoses); Mayr, 1861: 66 (Europe species key); Roger, 1863b: 35 (catalogue); Mayr, 1863: 395, 437 (catalogue); Mayr, 1865: 18 (diagnosis); Cresson, 1887: 259 (U.S.A. catalogue); Dalla Torre, 1893: 150 (catalogue); Gallardo, 1916d: 340 (Argentina species key); Emery, 1924d: 352 (diagnosis, catalogue); Gonçalves, 1942: 334 (all species key); Gonçalves, 1947a: 186 (south & central Brazil species key); Chapman & Capco, 1951: 82 (Asia checklist, anachronisms); Borgmeier, 1959b: 335 (subgenera key); Borgmeier, 1959b: 337 (diagnosis, all species revision, key); Kempf, 1972a: 26 (Neotropical catalogue); Smith, D.R. 1979: 1413 (North America catalogue); Mackay & Mackay, 1988: 25 (Colombia species key); Cherrett & Cherrett, 1989: 1 (bibliography); Brandão, 1991: 328 (catalogue); Bolton, 1994: 105 (synoptic classification); Bolton, 1995a: 1048 (census); Bolton, 1995b: 75 (catalogue).

Myrmicinae Attini Cyphomyrmex TRUE TRUE FALSE

Extant: 39 species

Fossil: 2 species

Cyphomyrmex Mayr, 1862: 651 (diagnosis in key), 690. Type-species: Cyphomyrmex minutus, by monotypy.

Taxonomic history

[Type-species not Cryptocerus rimosus, unjustified subsequent designation by Wheeler, W.M. 1911f: 161. Type-species not Meranoplus difformis, unjustified subsequent designation by Emery, 1924d: 340.]

Cyphomyrmex in Formicinae: Mayr, 1862: 651 (in key) [Formicidae].

Cyphomyrmex in Myrmicidae, Cryptoceridae: Emery, 1877a: 81.

Cyphomyrmex in Myrmicinae: Mayr, 1865: 25 [Myrmicidae]; Dalla Torre, 1893: 149.

Cyphomyrmex in Cryptoceridae, Dacetonini: Ashmead, 1905b: 384.

Cyphomyrmex in Myrmicinae, Attini: Forel, 1893a: 164; Emery, 1895j: 770; Forel, 1899c: 40; Wheeler, W.M. 1907c: 670; Wheeler, W.M. 1910g: 141; Emery, 1913b: 251; Emery, 1914a: 42; Forel, 1917: 247; Wheeler, W.M. 1922a: 669; Emery, 1924d; 339; all subsequent authors.

Cyphomyrmex as junior synonym of Cataulacus: Roger, 1863b: 39.

Cyphomyrmex as subgenus of Atta: Forel, 1912e: 188.

Cyphomyrmex as genus: Mayr, 1862: 690; Mayr, 1863: 406; Emery, 1913b: 251; Bruch, 1914: 217; Emery, 1914a: 42; Wheeler, W.M. 1922a: 669; Emery, 1924d; 339; all subsequent authors.

Junior synonym of CYPHOMYRMEX

Cyphomannia Weber, 1938b: 183 [as subgenus of Cyphomyrmex]. Type-species: Cyphomyrmex (Cyphomannia) laevigatus, by original designation.

Taxonomic history

Cyphomannia as subgenus of Cyphomyrmex: Weber, 1938b: 183; Weber, 1966: 166.

Cyphomannia as junior synonym of Cyphomyrmex: Kempf, 1962b: 29; Kempf, 1968a: 39; Kempf, 1972a: 92; Bolton, 1994: 105.

Genus Cyphomyrmex references

Mayr, 1863: 406 (catalogue); Mayr, 1865: 25 (diagnosis); Mayr, 1887: 556 (all species key); Dalla Torre, 1893: 149 (catalogue); Emery, 1924d; 340 (diagnosis, subgenera key, catalogue); Kusnezov, 1949d: 436 (Argentina species key); Weber, 1940a: 408 (all species key); Creighton, 1950a: 316 (North America species key); Kusnezov, 1957b: 9 (Neotropical species, partial key); Kempf, 1962b: 30 (species groups); Kempf, 1964d: 5 (C. strigatus group revision, key); Kempf, 1966: 161 (C. rimosus group revision, key); Kempf, 1972a: 92 (Neotropical catalogue); Smith, D.R. 1979: 1410 (North America catalogue); Brandão, 1991: 339 (catalogue); Snelling, R.R. & Longino, 1992: 480 (C. rimosus group revision, key); Bolton, 1994: 105 (synoptic classification); Bolton, 1995a: 1049 (census); Bolton, 1995b: 167 (catalogue); Mackay & Mackay, 2002: 38 (U.S.A., New Mexico species key).

Myrmicinae Attini Mycetosoritis TRUE TRUE FALSE

5 species

Mycetosoritis Wheeler, W.M. 1907c: 714 [as subgenus of Atta]. Type-species: Atta (Mycetosoritis) hartmanni, by monotypy.

Taxonomic history

Mycetosoritis in Myrmicinae, Attini: Emery, 1913b: 251; Wheeler, W.M. 1922a: 669; Emery, 1924d: 343; all subsequent authors.

Mycetosoritis as subgenus of Atta: Wheeler, W.M. 1907c: 670; Wheeler, W.M. 1910g: 141.

Mycetosoritis as subgenus of Cyphomyrmex: Emery, 1913b: 251; Forel, 1917: 247; Emery, 1924d: 343; Donisthorpe, 1943f: 663; Kusnezov, 1949d: 444; Kusnezov, 1956: 23.

Mycetosoritis as subgenus of Trachymyrmex: Wheeler, W.M. 1922a: 669.

Mycetosoritis as genus: Creighton, 1950a: 317; Kempf, 1972a: 146.

Genus Mycetosoritis references

Kempf, 1972a: 146 (catalogue); Smith, D.R. 1979: 1410 (North America catalogue); Bolton, 1994: 105 (synoptic classification); Bolton, 1995a: 1050 (census); Bolton, 1995b: 269 (catalogue).

Myrmicinae Attini Trachymyrmex TRUE TRUE FALSE

Extant: 47 species, 1 subspecies

Fossil: 1 species

Trachymyrmex Forel, 1893e: 600 [as subgenus of Atta]. Type-species: Atta septentrionalis, by subsequent designation of Wheeler, W.M. 1911f: 174.

Taxonomic history

Trachymyrmex in Cryptoceridae, Attinae: Ashmead, 1905b: 384.

Trachymyrmex in Myrmicinae, Attini: Emery, 1895j: 770; Emery, 1913b: 251; Forel, 1917: 247; Wheeler, W.M. 1922a: 669; Emery, 1924d: 344; all subsequent authors.

Trachymyrmex as subgenus of Atta: Forel, 1893e: 600 Emery, 1895j: 770; Forel, 1893e: 600; Forel, 1899c: 30; Wheeler, W.M. 1907c: 670; Wheeler, W.M. 1910g: 141.

Trachymyrmex as subgenus of Cyphomyrmex: Emery, 1913b: 251; Emery, 1924d: 344; Donisthorpe, 1943g: 733.

Trachymyrmex as subgenus of Acromyrmex: Forel, 1917: 247.

Trachymyrmex as genus: Gallardo, 1916b: 242; Gallardo, 1916d: 318; Wheeler, W.M. 1916c: 11; Wheeler, W.M. 1922a: 669; Borgmeier, 1927c: 128; Borgmeier, 1950b: 384; Kusnezov, 1964: 63; Kempf, 1972a: 252.

Genus Trachymyrmex references

Wheeler, W.M. 1911g: 248 (all species key); Emery, 1924d: 344 (diagnosis, catalogue); Creighton, 1950a: 320 (North America species key); Kempf, 1972a: 252 (Neotropical catalogue); Smith, D.R. 1979: 1411 (North America catalogue); Brandão, 1991: 382 (catalogue); Bolton, 1994: 105 (synoptic classification); Bolton, 1995a: 1053 (census); Bolton, 1995b: 420 (catalogue); Mackay & Mackay, 2002: 38 (U.S.A., New Mexico species key); Mayhé-Nunes & Brandão, 2002: 672 (diagnosis, T. opulentus group key); Mayhé-Nunes & Brandão, 2005: 274 (T. iheringi group, key); Brandão& Mayhé-Nunes, 2007: 72 (species group phylogeny). Mayhé-Nunes & Brandão, 2007: 4 (T. jamaicensis group, key); Rabeling, Cover, et al. 2007: 4 (North America species, key).

Ichnotaxon attached to Attini

Ichnogenus †ATTAICHNUS

†Attaichnus Laza, 1982: 112. Included ichnospecies: †Attaichnus kuenzelii. [Ichnofossil, purportedly fossil traces of workings attributable to attine ants.]

Myrmicinae Stenammini Rogeria TRUE TRUE FALSE

40 species

Rogeria Emery, 1894c: 188. Type-species: Rogeria curvipubens, by subsequent designation of Wheeler, W.M. 1911f: 172.

Taxonomic history

Rogeria in Myrmicinae, Myrmicini: Emery, 1895j: 769; Forel, 1899c: 53; Wheeler, W.M. 1910g: 139; Kusnezov, 1958a: 44; Kusnezov, 1964: 57.

Rogeria in Myrmicinae, Tetramoriini: Ashmead, 1905b: 383.

Rogeria in Myrmicinae, Leptothoracini: Emery, 1914a: 42; Forel, 1917: 245; Wheeler, W.M. 1922a: 664; Emery, 1924d: 266; Donisthorpe, 1943g: 724; Chapman & Capco, 1951: 111; Kempf, 1972a: 227; Smith, D.R. 1979: 1391; Dlussky & Fedoseeva, 1988: 79; Jaffe, 1993: 11; Kugler, C. 1994: 24.

Rogeria in Myrmicinae, Pheidolini: Hölldobler & Wilson, 1990: 16.

Rogeria in Myrmicinae, Stenammini: Bolton, 1994: 106.

Junior synonym of ROGERIA

Irogera Emery, 1915i: 191 [as subgenus of Rogeria]. Type-species: Rogeria procera, by original designation.

Taxonomic history

Irogera in Myrmicinae, Leptothoracini: Forel, 1917: 245; Emery, 1924d: 267; Donisthorpe, 1943f: 653.

Irogera as subgenus of Rogeria: Emery, 1915i: 191; Forel, 1917: 245; Emery, 1924d: 267; Wheeler, W.M. 1922a: 677; Donisthorpe, 1943f: 653.

Irogera as genus: Brown, 1953h: 4 (in text); Kempf, 1961d: 436; Kempf, 1964e: 66.

Irogera as junior synonym of Rogeria: Kempf, 1965: 185; Kugler, C. 1994: 23.

Genus Rogeria references

Mann, 1921: 456 (Melanesia species key); Wheeler, W.M. 1922a: 677 (subgenera key); Emery, 1924d: 266 (diagnosis, catalogue); Chapman & Capco, 1951: 111 (Asia checklist); Kempf, 1963a: 195 (Neotropical species key); Kempf, 1964e: 66 (Irogera species key); Kempf, 1972a: 227 (Neotropical catalogue); Smith, D.R. 1979: 1391 (North America catalogue); Taylor, 1987a: 72 (New Caledonia checklist); Brandão, 1991: 377 (catalogue); Kugler, C. 1994: 17 (diagnosis, all species revision, key); Bolton, 1995a: 1052 (census); Bolton, 1995b: 381 (catalogue); LaPolla & Sosa-Calvo, 2006: 67 (Guyana species key); Heterick, 2009: 175 (south-western Australia species key).

Myrmicinae Stenammini Stenamma TRUE TRUE FALSE

Extant: 44 species

Fossil: 1 species

Stenamma Westwood, 1839: 219. Type-species: Stenamma westwoodii, by monotypy.

Taxonomic history

Stenamma in Myrmicidae, Myrmicidae: Emery, 1877a: 81.

Stenamma in Myrmicinae: Mayr, 1861: 55 [Myrmicidae]; Mayr, 1865: 23 [Myrmicidae]; Emery & Forel, 1879a: 456 [Myrmicidae]; Dalla Torre, 1893: 121.

Stenamma in Myrmicinae, Myrmicini: Emery, 1895j: 769; Wheeler, W.M. 1910g: 140; Wheeler, W.M. 1915h: 53; Kempf, 1972a: 242 (anachronism).

Stenamma in Myrmicinae, Pheidolini: Emery, 1914a: 40; Forel, 1917: 241; Emery, 1921f: 52 [subtribe Stenammini]; Wheeler, W.M. 1922a: 661; Karavaiev, 1934: 96; Kugler, C. 1994: 25.

Stenamma in Myrmicinae, Stenammini: Ashmead, 1905b: 383; Bolton, 1994: 106.

Junior synonyms of STENAMMA

Asemorhoptrum Mayr, 1861: 76. Type-species: Myrmica lippula, by monotypy.

Taxonomic history

Asemorhoptrum in Myrmicinae: Mayr, 1865: 20 [Myrmicidae]; Emery, 1877a: 81 [Myrmicidae]; Emery & Forel, 1879a: 460; [Myrmicidae].

Asemorhoptrum as junior synonym of Stenamma: André, 1883a: 310; all subsequent authors except the following.

[Asemorhoptrum as junior synonym of Proatta: Snelling, R.R. 1981: 395 (error).]

Theryella Santschi, 1921d: 68. Type-species: Theryella myops (provisional junior synonym of Stenamma punctiventre), by monotypy.

Taxonomic history

Theryella as junior synonym of Stenamma: Santschi, 1923a: 136; all subsequent authors except the following.

[Theryella as junior synonym of Proatta: Snelling, R.R. 1981: 395 (error).]

Genus Stenamma references

Roger, 1863b: 25, 27 (catalogue); Mayr, 1863: 395, 454 (catalogue); Mayr, 1865: 20, 23 (Asemorhoptrum, Stenamma diagnoses); André, 1883a: 311 (Europe & Algeria species key); Cresson, 1887: 261 (U.S.A. catalogue); Dalla Torre, 1893: 121 (catalogue); Forel, 1903a: 693 (India & Sri Lanka species key); Emery, 1908c: 306 (Palaearctic species key); Emery, 1916b: 127 (Italy species key); Emery, 1921f: 52 (diagnosis, catalogue); Arnol'di, 1928b: 215 (former U.S.S.R. species key); Arnol'di, 1933b: 599 (Russia species key); Buren, 1944a: 284 (U.S.A., Iowa species key); Creighton, 1950a: 135 (North America species key); Chapman & Capco, 1951: 153 (Asia checklist); Smith, M.R. 1957b: 141 (U.S.A. species key); Yasumatsu & Murakami, 1960: 28 (Japan species key); Smith, M.R. 1962a: 38 (Mexico and Central America species key); Gregg, 1963: 347 (U.S.A., Colorado species key); Wheeler, G.C. & Wheeler, J. 1963: 120 (U.S.A., North Dakota species key); Bernard, 1967: 126 (diagnosis, Western Europe species key); Kempf, 1972a: 242 (Neotropical catalogue); Snelling, R.R. 1973c: 5 (Western U.S.A. species key); Arnol'di, 1975: 1826 (former U.S.S.R. species key); Kutter, 1977c: 74 (Switzerland species key); Arnol'di & Dlussky, 1978: 535 (former European U.S.S.R. species key); Collingwood, 1978: 80 (Iberian Peninsula species key); Smith, D.R. 1979: 1357 (North America, catalogue); Allred, 1982: 443 (U.S.A., Utah species key); Gösswald, 1985: 297 (Germany species key); Wheeler, G.C. & Wheeler, J. 1986g: 33 (U.S.A., Nevada species key); Agosti & Collingwood, 1987b: 268 (Balkans species key); Kupyanskaya, 1990: 116 (Far Eastern Russia species key); Morisita, Kubota, et al. 1992: 14 (Japan species key); Arakelian, 1994: 29 (Armenia species key); Kupyanskaya, 1995: 344 (Far Eastern Russia species key); Bolton, 1995a: 1052 (census); Bolton, 1995b: 393 (catalogue); Seifert, 1996b: 155 (Central Europe species key); Skinner & Allen, 1996: 44 (Britain species key); Fernández, Baena & Palacio, 1996: 9 (Colombia species key); DuBois, 1998b: 215 (Palaearctic & Oriental species revision, keys); Mackay & Mackay, 2002: 209 (U.S.A., New Mexico species key); Lyu & Cho, 2004: 263 (Korea species key); Seifert, 2007: 143 (North and Central Europe species key); Branstetter, 2009: 43 (diagnosis).

Myrmicinae Solenopsidini Carebara TRUE TRUE FALSE

Extant: 160 species, 13 subspecies

Fossil: 6 species

Carebara Westwood, 1840b: 86. Type-species: Carebara lignata, by monotypy.

Taxonomic history

Carebara in Poneridae, Attidae: Smith, F. 1858b: 178.

Carebara in Myrmicinae: Mayr, 1865: 23 [Myrmicidae]; Smith, F. 1871a: 334 [Myrmicidae]; Dalla Torre, 1893: 74.

Carebara in Myrmicinae, Pheidolini: Emery, 1877a: 81 [Myrmicidae, Pheidolidae].

Carebara in Myrmicinae, Myrmicariini: Ashmead, 1905b: 383.

Carebara in Myrmicinae, Solenopsidini: Forel, 1893a: 164; Emery, 1895j: 770; Wheeler, W.M. 1910g: 140; Wheeler, W.M. 1922a: 663; Kusnezov, 1964: 61; Kempf, 1972a: 74; Jaffe, 1993: 10; Bolton, 2003: 61.

Carebara in Myrmicinae, Pheidologeton genus group: Ettershank, 1966: 81; Bolton, 1987: 265.

Carebara in Myrmicinae, Pheidologetini: Emery, 1914a: 41; Arnold, 1916: 248; Forel, 1917: 244; Emery, 1924d: 219; Dlussky & Fedoseeva, 1988: 80; Hölldobler & Wilson, 1990: 16; Bolton, 1994: 106 [Pheidologetonini].

Junior synonyms of CAREBARA

Oligomyrmex Mayr, 1867a: 110. Type-species: Oligomyrmex concinnus, by monotypy.

Taxonomic history

Oligomyrmex in Myrmicinae: Dalla Torre, 1893: 74.

Oligomyrmex in Myrmicinae, Stenammini: Ashmead, 1905b: 383.

Oligomyrmex in Myrmicinae, Solenopsidini: Forel, 1893a: 164; Emery, 1895j: 770; Wheeler, W.M. 1910g: 140; Wheeler, W.M. 1922a: 663; Kempf, 1972a: 172; Smith, D.R. 1979: 1389; Jaffe, 1993: 10.

Oligomyrmex in Myrmicinae, Pheidologeton genus group: Ettershank, 1966: 81; Bolton, 1987: 265.

Oligomyrmex in Myrmicinae, Pheidologetini: Emery, 1914a: 41; Forel, 1917: 243; Emery, 1924d: 215; Dlussky & Fedoseeva, 1988: 80; Hölldobler & Wilson, 1990: 16; Bolton, 1994: 106 [Pheidologetonini].

Oligomyrmex in Myrmicinae, Solenopsidini: Bolton, 2003: 61.

Oligomyrmex as junior synonym of Carebara: Fernández, 2004a: 194.

Aeromyrma Forel, 1891b: 198. Type-species: Aeromyrma nosindambo, by monotypy.

Taxonomic history

[Aeromyrma also described as new by Forel, 1891a: cccvii; no species-rank taxa named.]

Aeromyrma in Myrmicinae: Dalla Torre, 1893: 78.

Aeromyrma in Myrmicinae, Solenopsidini: Forel, 1893a: 164; Emery, 1895j: 770; Ashmead, 1905b: 383; Wheeler, W.M. 1910g: 140; Wheeler, W.M. 1922a: 663.

Aeromyrma in Myrmicinae, Pheidologetini: Emery, 1914a: 41; Arnold, 1916: 256; Forel, 1917: 243; Emery, 1924d: 215; all subsequent authors.

Aeromyrma as genus: Forel, 1891b: 198; Arnold, 1916: 256; Forel, 1917: 243; Wheeler, W.M. 1922a: 663, 882.

Aeromyrma as subgenus of Oligomyrmex: Emery, 1915c: 59 (footnote); Emery, 1924d: 215; Donisthorpe, 1943f: 620.

Aeromyrma as junior synonym of Oligomyrmex: Ettershank, 1966: 119; all subsequent authors to the following.

Aeromyrma as junior synonym of Carebara: Fernández, 2004a: 194.

Aneleus Emery, 1900c: 327 [as subgenus of Pheidologeton]. Type-species: Solenopsis similis, by subsequent designation of Wheeler, W.M. 1911f: 158.

Taxonomic history

[Type-species not Pheidologeton pygmaeus, unjustified subsequent designation by Wheeler, W.M. 1913a: 77; repeated in Emery, 1924d: 214.]

Aneleus in Myrmicinae, Solenopsidini: Wheeler, W.M. 1910g: 140; Wheeler, W.M. 1922a: 663.

Aneleus in Myrmicinae, Pheidologetini: Emery, 1914a: 41; Arnold, 1916: 254; Forel, 1917: 243; Emery, 1924d: 213; all subsequent authors.

Aneleus as genus: Emery, 1914a: 41; Arnold, 1916: 254; Forel, 1917: 243; Wheeler, W.M. 1922a: 663; Emery, 1924d: 213.

Aneleus as junior synonym of Oligomyrmex: Ettershank, 1966: 119; all subsequent authors to the following.

Aneleus as junior synonym of Carebara: Fernández, 2004a: 194.

Erebomyrma Wheeler, W.M. 1903a: 138. Type-species: Erebomyrma longii, by monotypy.

Taxonomic history

Erebomyrma in Myrmicinae, Solenopsidini: Wheeler, W.M. 1903a: 145; Wheeler, W.M. 1910g: 140; Wheeler, W.M. 1915h: 46; Wheeler, W.M. 1922a: 663.

Erebomyrma in Myrmicinae, Stenammini: Ashmead, 1905b: 383.

Erebomyrma in Myrmicinae, Pheidologetini: Emery, 1914a: 41; Forel, 1917: 244; Emery, 1924d: 218; all subsequent authors.

Erebomyrma as genus: Wheeler, W.M. 1922a: 663; Emery, 1924d: 218; Creighton, 1950a: 245; Kusnezov, 1964: 62; Wilson, 1986b: 61; Hölldobler & Wilson, 1990: 13.

Erebomyrma as junior synonym of Oligomyrmex: Ettershank, 1966: 119; Kempf, 1972a: 172; Smith, D.R. 1979: 1389; Bolton, 1994: 106; Bolton, 1995b: 298.

Erebomyrma as junior synonym of Carebara: Fernández, 2004a: 194.

Paedalgus Forel, 1911i: 217. Type-species: Paedalgus escherichi, by monotypy.

Taxonomic history

Paedalgus in Myrmicinae, Solenopsidini: Forel, 1911i: 218; Wheeler, W.M. 1922a: 663.

Paedalgus in Myrmicinae, Pheidologeton genus group: Ettershank, 1966: 81; Bolton, 1987: 265.

Paedalgus in Myrmicinae, Pheidologetini: Emery, 1914a: 41; Forel, 1917: 244; Emery, 1924d: 221; Dlussky & Fedoseeva, 1988: 80; Bolton, 1994: 106 [Pheidologetonini].

Paedalgus in Myrmicinae, Solenopsidini: Bolton, 2003: 61.

Paedalgus as junior synonym of Carebara: Fernández, 2004a: 194.

Lecanomyrma Forel, 1913k: 56 [as subgenus of Pheidologeton]. Type-species: Pheidologeton (Lecanomyrma) butteli, by monotypy.

Taxonomic history

Lecanomyrma as subgenus of Pheidologeton: Forel, 1913k: 56; Forel, 1917: 243.

Lecanomyrma as subgenus of Aneleus: Emery, 1924d: 215.

Lecanomyrma as junior synonym of Oligomyrmex: Ettershank, 1966: 119; all subsequent authors to the following.

Lecanomyrma as junior synonym of Carebara: Fernández, 2004a: 194.

Spelaeomyrmex Wheeler, W.M. 1922c: 9. Type-species: Spelaeomyrmex urichi, by original designation.

Taxonomic history

Spelaeomyrmex in Myrmicinae, Pheidologitonini: Wheeler, W.M. 1922c: 11; Donisthorpe, 1944b: 59.

Spelaeomyrmex in Myrmicinae, Solenopsidini: Donisthorpe, 1943g: 727; Kusnezov, 1964: 61.

Spelaeomyrmex as junior synonym of Erebomyrma: Wilson, 1962a: 63.

Spelaeomyrmex as junior synonym of Carebara: Fernández, 2004a: 194.

Hendecatella Wheeler, W.M. 1927h: 93 [as subgenus of Oligomyrmex]. Type-species: Oligomyrmex (Hendecatella) capreolus, by monotypy.

Taxonomic history

Hendecatella as junior synonym of Oligomyrmex: Ettershank, 1966: 119; all subsequent authors.

Hendecatella as junior synonym of Carebara: Fernández, 2004a: 194.

Solenops Karavaiev, 1930a: 207 [as subgenus of Solenopsis]. Type-species: Solenopsis (Solenops) weyeri, by monotypy.

Taxonomic history

Solenops as junior synonym of Oligomyrmex: Ettershank, 1966: 119; all subsequent authors.

Solenops as junior synonym of Carebara: Fernández, 2004a: 194.

Crateropsis Patrizi, 1948: 174 [as subgenus of Solenopsis]. Type-species: Solenopsis (Crateropsis) elmenteitae, by original designation.

Taxonomic history

Crateropsis as junior synonym of Oligomyrmex: Ettershank, 1966: 120; all subsequent authors.

Crateropsis as junior synonym of Carebara: Fernández, 2004a: 194.

Sporocleptes Arnold, 1948: 219. Type-species: Sporocleptes nicotianae, by original designation.

Taxonomic history

Sporocleptes as junior synonym of Aneleus: Consani, 1951: 169; Arnold, 1952a: 460.

Sporocleptes as junior synonym of Carebara: Fernández, 2004a: 194.

Nimbamyrma Bernard, 1953b: 240. Type-species: Nimbamyrma villiersi, by monotypy.

Taxonomic history

Nimbamyrma in Myrmicinae, Pheidologetini: Bernard, 1953b: 240.

Nimbamyrma as junior synonym of Oligomyrmex: Ettershank, 1966: 120; all subsequent authors.

Nimbamyrma as junior synonym of Carebara: Fernández, 2004a: 194; Fernández, 2010: 201.

Afroxyidris Belshaw & Bolton, 1994: 631. Type-species: Afroxyidris crigensis, by original designation.

Taxonomic history

Afroxyidris in Myrmicinae, Pheidologetonini: Belshaw & Bolton, 1994: 632; Bolton, 1994: 106.

Afroxyidris in Myrmicinae, Solenopsidini: Bolton, 2003: 61.

Afroxyidris as junior synonym of Carebara: Fernández, 2004a: 194.

Neoblepharidatta Sheela & Narendran, 1997: 88. Type-species: Neoblepharidatta nayana, by original designation.

Taxonomic history

Neoblepharidatta incertae sedis in Myrmicinae: Sheela & Narendran, 1997: 89.

Neoblepharidatta as junior synonym of Oligomyrmex: Bolton, 2003: 216, 273.

Neoblepharidatta as junior synonym of Carebara: Fernández, 2004a: 194.

Parvimyrma Eguchi & Bui, 2007: 40. Type-species: Parvimyrma sangi, by original designation.

Taxonomic history

Parvimyrma in Myrmicinae, Solenopsis genus group: Eguchi & Bui, 2007: 40.

Parvimyrma as junior synonym of Carebara: Fernández, 2010: 195.

Genus Carebara references

Smith, F. 1858b: 178 (Carebara diagnosis); Roger, 1863b: 32 (Carebara catalogue); Mayr, 1863: 402 (Carebara catalogue); Mayr, 1865: 23 (Carebara diagnosis); Mayr, 1867a: 106 (Carebara diagnosis); Dalla Torre, 1893: 74, 78 (Carebara, Oligomyrmex, Aeromyrma catalogues); Bingham, 1903: 150 (Carebara diagnosis); Bingham, 1903: 154 (Oligomyrmex India, Sri Lanka & Burma species key); Santschi, 1913e: 459 (African Oligomyrmex, key); Santschi, 1914d: 363 (Afrotropical Carebara species key); Arnold, 1916: 248, 254, 256 (Carebara, Aneleus, Aeromyrma diagnoses); Wheeler, W.M. 1922a: 165, 687 (Aeromyrma diagnosis, Oligomyrmex subgenera key); Wheeler, W.M. 1922a: 168, 172, 883 (Carebara diagnosis, Afrotropical species key, Afrotropical catalogue); Wheeler, W.M. 1922a: 177, 884 (Paedalgus diagnosis, Afrotropical catalogue); Wheeler, W.M. 1922a: 880, 881, 882 (Afrotropical Aneleus, Oligomyrmex, Aeromyrma catalogues); Wheeler, W.M. 1922a: 1028 (Malagasy Oligomyrmex, Aeromyrma catalogues); Emery, 1924d: 213 (Aneleus diagnosis, catalogue); Emery, 1924d: 215 (Aneleus (Lecanomyrma) diagnosis, catalogue); Emery, 1924d: 215 (Oligomyrmex (Aeromyrma) diagnosis, catalogue); Emery, 1924d: 217 (O. (Oligomyrex) diagnosis, catalogue); Emery, 1924d: 218 (Erebomyrma diagnosis, catalogue); Emery, 1924d: 219 (Carebara diagnosis, catalogue); Emery, 1924d: 221 (Paedalgus diagnosis, catalogue); Weber, 1950a: 1 (Africa Oligomyrmex species, synopsis); Chapman & Capco, 1951: 151, 154, 156, 158 (Asia Carebara, Aeromyrma, Aneleus, Oligomyrmex, Paedalgus checklists); Weber, 1952: 11 (Africa O. (Aeromyrma) species key); Ettershank, 1966: 119 (Oligomyrmex diagnosis, review of genus, checklist); Ettershank, 1966: 125 (Carebara diagnosis, review of genus, checklist); Ettershank, 1966: 128 (Paedalgus diagnosis, review of genus, checklist); Kempf, 1972a: 74 (Carebara Neotropical catalogue); Kempf, 1972a: 172 (Oligomyrmex Neotropical catalogue); Smith, D.R. 1979: 1389 (Oligomyrmex North America catalogue); Taylor & Brown, D.R. 1985: 71 (Oligomyrmex Australia catalogue); Taylor, 1987a: 50 (Oligomyrmex Australia, New Caledonia checklist); Taylor, 1991db: 605 (Oligomyrmex Australia species, partial key); Brandão, 1991: 342 (Oligomyrmex Neotropical catalogue); Morisita, Kubota, et al. 1992: 45 (Oligomyrmex Japan species key); Bolton & Belshaw, 1993: 181 (Paedalgus diagnosis, all species revision, key); Bolton, 1995a: 1048, 1051 (Carebara, Oligomyrmex, Paedalgus census); Bolton, 1995b: 133, 298, 311 (Carebara, Oligomyrmex, Paedalgus catalogues); Wu, J. & Wang, 1995: 74 (Oligomyrmex China species key); Terayama, 1996: 24 (Oligomyrmex Japan species key); Shattuck, 1999: 148 (Oligomyrmex Australia synopsis); Xu, 2003: 311 (Oligomyrmex China species key); Terayama, 2009: 151 (Taiwan species key).

Myrmicinae Solenopsidini Monomorium TRUE TRUE FALSE

Extant: 386 species, 30 subspecies

Fossil: 3 species

Monomorium Mayr, 1855: 452. Type-species: Monomorium monomorium (replacement name for Monomorium minutum, junior secondary homonym in Monomorium), by monotypy.

Taxonomic history

Monomorium in Myrmicidae, Myrmicidae: Emery, 1877a: 81.

Monomorium in Myrmicinae: Mayr, 1855: 452 [Myrmicidae]; Mayr, 1861: 71 [Myrmicidae]; Mayr, 1865: 22 [Myrmicidae]; Emery & Forel, 1879a: 456 [Myrmicidae]; Dalla Torre, 1893: 65.

Monomorium in Myrmicinae, Myrmicini: Forel, 1895b: 124; Emery, 1895j: 769; Forel, 1899c: 78; Wheeler, W.M. 1910g: 139.

Monomorium in Myrmicinae, Tetramoriini: Ashmead, 1905b: 383.

Monomorium in Myrmicinae, Monomoriini: Wheeler, W.M. 1915h: 45; Dlussky & Fedoseeva, 1988: 80.

Monomorium in Myrmicinae, Monomorium genus group: Ettershank, 1966: 81; in Solenopsis genus group: Bolton, 1987: 271.

Monomorium in Myrmicinae, Solenopsidini: Emery, 1914a: 41 [subtribe Monomoriini]; Arnold, 1916: 203; Forel, 1917: 242; Emery, 1922e: 166; Wheeler, W.M. 1922a: 662; Hölldobler & Wilson, 1990: 16; Jaffe, 1993: 10; Bolton, 1994: 106.

Junior synonyms of MONOMORIUM

Phacota Roger, 1862b: 260. Type-species: Phacota sichelii, by monotypy.

Taxonomic history

Phacota in Myrmicidae, Pheidolidae: Emery, 1877a: 81.

Phacota in Myrmicinae: Mayr, 1865: 22 [Myrmicidae]; Emery & Forel, 1879a: 464 [Myrmicidae]; Dalla Torre, 1893: 71.

Phacota in Myrmicinae, Myrmicini: Emery, 1895j: 769; Wheeler, W.M. 1910g: 139.

Phacota in Myrmicinae, Stenammini: Ashmead, 1905b: 383.

Phacota in Myrmicinae, Solenopsidini: Emery, 1914a: 41 [subtribe Monomoriini]; Forel, 1917: 243; Emery, 1922e: 187; Wheeler, W.M. 1922a: 662; all subsequent authors except the entry below; Hölldobler & Wilson, 1990: 16; Bolton, 1994: 106.

Phacota in Myrmicinae, Solenopsis genus group: Bolton, 1987: 271.

Phacota as genus: Bolton, 1987: 281.

Phacota as junior synonym of Monomorium: Ettershank, 1966: 82; Fernández, in Heterick, 2006: 79; Fernández, 2007b: 130.

Trichomyrmex Mayr, 1865: 19. Type-species: Trichomyrmex rogeri, by monotypy.

Taxonomic history

Trichomyrmex in Myrmicidae, Myrmicidae: Emery, 1877a: 81.

Trichomyrmex in Myrmicinae: Mayr, 1865: 19 [Myrmicidae]; Dalla Torre, 1893: 72.

Trichomyrmex in Myrmicinae, Myrmicini: Wheeler, W.M. 1910g: 140.

Trichomyrmex in Myrmicinae, Solenopsidini: Emery, 1914a: 41 [subtribe Monomoriini]; Forel, 1917: 243; Emery, 1922e: 185; Wheeler, W.M. 1922a: 662; Donisthorpe, 1943g: 733.

Trichomyrmex as junior synonym of Monomorium: Ettershank, 1966: 82; Smith, D.R. 1979: 1381; Bolton, 1987: 296.

†Lampromyrmex Mayr, 1868c: 93. Type-species: †Monomorium mayrianum (replacement name for †Lampromyrmex gracillimus, junior secondary homonym in Monomorium), by monotypy.

Taxonomic history

†Lampromyrmex in Myrmicidae, Pheidolidae: Emery, 1877a: 81.

†Lampromyrmex in Myrmicinae: Mayr, 1868c: 93 [Myrmicidae]; Dalla Torre, 1893: 78.

†Lampromyrmex in Myrmicinae, Solenopsidini: Emery, 1922e: 183 [subtribe Monomoriini]; Donisthorpe, 1943f: 655.

†Lampromyrmex as junior synonym of Monomorium: Wheeler, W.M. 1915h: 45.

†Lampromyrmex as subgenus of Monomorium: Emery, 1922e: 183; Wheeler, W.M. 1922a: 676.

†Lampromyrmex as junior synonym of Monomorium: Wheeler, W.M. 1915h: 45; Ettershank, 1966: 82; Smith, D.R. 1979: 1381; Bolton, 1987: 299.

Holcomyrmex Mayr, 1879: 671. Type-species: Holcomyrmex scabriceps, by subsequent designation of Bingham, 1903: 280.

Taxonomic history

[Holcomyrmex Smith, F. 1873: ix, nomen nudum, attributed to Mayr.]

Holcomyrmex in Myrmicinae, Myrmicini: Emery, 1895j: 769; Ashmead, 1905b: 383.

Holcomyrmex in Myrmicinae, Solenopsidini: Emery, 1922e: 181 [subtribe Monomoriini]; Donisthorpe, 1943f: 650.

Holcomyrmex as genus: Dalla Torre, 1893: 65; Forel, 1903a: 692; Bingham, 1903: 280.

Holcomyrmex as subgenus of Monomorium: Wheeler, W.M. 1910g: 139; Emery, 1915i: 190; Forel, 1917: 242; Emery, 1922e: 181; Wheeler, W.M. 1922a: 676.

Holcomyrmex as junior synonym of Monomorium: Emery, 1908h: 667; Ettershank, 1966: 82; Smith, D.R. 1979: 1381; Bolton, 1987: 295.

Epoecus Emery, 1893a: cclxxvi. Type-species: Epoecus pergandei, by monotypy.

Taxonomic history

Epoecus in Myrmicinae, Myrmicini: Emery, 1895j: 769; Wheeler, W.M. 1910g: 139.

Epoecus in Myrmicinae, Stenammini: Ashmead, 1905b: 383.

Epoecus in Myrmicinae, Solenopsidini: Emery, 1914a: 41 [subtribe Anergatini]; Forel, 1917: 243; Emery, 1922e: 204; Wheeler, W.M. 1922a: 663; Donisthorpe, 1943f: 643.

Epoecus as junior synonym of Monomorium: Ettershank, 1966: 82; Smith, D.R. 1979: 1381; Bolton, 1987: 287.

Wheeleriella Forel, 1907f: 145.

Taxonomic history

[Replacement name for Wheeleria Forel, 1905b: 171; junior homonym of Wheeleria Tutt, 1905: 37 (Lepidoptera).]

Wheeleriella in Myrmicinae, Myrmicini: Wheeler, W.M. 1910g: 139.

Wheeleriella in Myrmicinae, Solenopsidini: Emery, 1914a: 41 [subtribe Monomoriini]; Forel, 1917: 243; Emery, 1922e: 186; Wheeler, W.M. 1922a: 662; Donisthorpe, 1943g: 735.

Wheeleriella as junior synonym of Monomorium: Ettershank, 1966: 82; Smith, D.R. 1979: 1381; Bolton, 1987: 291.

Homonym replaced by Wheeleriella

Wheeleria Forel, 1905b: 171. Type-species: Wheeleria santschii, by monotypy.

Taxonomic history

[Junior homonym of Wheeleria Tutt, 1905: 37 (Lepidoptera).]

Epixenus Emery, 1908f: 556. Type-species: Epixenus andrei (junior secondary homonym in Monomorium, replaced by Monomorium advena), by subsequent designation of Wheeler, W.M. 1911f: 163.

Taxonomic history

Epixenus in Myrmicinae, Solenopsidini: Emery, 1914a: 41 [subtribe Monomoriini]; Forel, 1917: 243; Emery, 1922e: 184; Wheeler, W.M. 1922a: 662; Donisthorpe, 1943f: 643.

Epixenus as junior synonym of Monomorium: Brown & Wilson, 1957bb: 244; Ettershank, 1966: 82; Smith, D.R. 1979: 1381; Bolton, 1987: 287; Tinaut & Ortiz, 1988: 167.

Mitara Emery, 1913b: 261 [as subgenus of Monomorium]. Type-species: Monomorium laeve, by original designation.

Taxonomic history

Mitara in Myrmicinae, Solenopsidini: Arnold, 1916: 238; Forel, 1917: 242 [subtribe Monomoriini]; all subsequent authors.

Mitara as subgenus of Monomorium: Emery, 1913b: 261; Emery, 1915i: 190; Arnold, 1916: 238; Forel, 1917: 242.

Mitara as junior synonym of †Lampromyrmex: Emery, 1922e: 183; Wheeler, W.M. 1922a: 162, 676.

Mitara as genus: Donisthorpe, 1943f: 662.

Mitara as junior synoynm of Monomorium: Smith, D.R. 1979: 1381; Bolton, 1987: 299.

Chelaner Emery, 1914f: 410 [as subgenus of Monomorium]. Type-species: Monomorium (Chelaner) forcipatum, by subsequent designation of Emery, 1922e: 168.

Taxonomic history

Chelaner in Myrmicinae, Solenopsidini: Forel, 1917: 242 [subtribe Monomoriini]; Emery, 1922e: 168; all subsequent authors except the following.

Chelaner in Myrmicinae, Monomorium genus group: Ettershank, 1966: 81.

Chelaner in Myrmicinae, Monomoriini: Dlussky & Fedoseeva, 1988: 80.

Chelaner as subgenus of Monomorium: Emery, 1914f: 410; Forel, 1917: 242; Emery, 1922e: 168; Wheeler, W.M. 1922a: 676.

Chelaner as genus: Ettershank, 1966: 93; Taylor & Brown, D.R. 1985: 55; Wheeler, G.C. & Wheeler, J. 1985: 257; Dlussky & Fedoseeva, 1988: 80 (anachronism).

Chelaner (and its junior synonyms Notomyrmex, Protholcomyrmex, Schizopelta) as junior synonym of Monomorium: Bolton, 1987: 300; Heterick, 2001: 354.

Notomyrmex Emery, 1915i: 190 [as subgenus of Monomorium]. Type-species: Atta antarctica, by original designation.

Taxonomic history

Notomyrmex as subgenus of Monomorium: Emery, 1915i: 190; Forel, 1917: 242; Emery, 1922e: 168; Wheeler, W.M. 1922a: 676; Kusnezov, 1956: 20.

Notomyrmex as genus: Kusnezov, 1957a: 269 (in key); Kusnezov, 1960b: 343; Kusnezov, 1962b: 160; Kusnezov, 1964: 60.

Notomyrmex as junior synonym of Chelaner: Ettershank, 1966: 93.

Parholcomyrmex Emery, 1915i: 190 [as subgenus of Monomorium]. Type-species: Myrmica gracillima (junior synonym of Atta destructor), by original designation.

Taxonomic history

Parholcomyrmex as subgenus of Monomorium: Emery, 1915i: 190; Forel, 1917: 242; Emery, 1922e: 179; Wheeler, W.M. 1922a: 676; subsequent authors to the following.

Parholcomyrmex as junior synonym of Monomorium: Ettershank, 1966: 82; Smith, D.R. 1979: 1381; Bolton, 1987: 296.

[Paraholcomyrmex Emery, 1915i: 191, Forel, 1917: 242, incorrect subsequent spellings.]

Xeromyrmex Emery, 1915i: 190 [as subgenus of Monomorium]. Type-species: Formica salomonis, by original designation.

Taxonomic history

Xeromyrmex as subgenus of Monomorium: Forel, 1917: 242; Emery, 1922e: 175; Wheeler, W.M. 1922a: 677.

Xeromyrmex as junior synonym of Monomorium: Ettershank, 1966: 82; Smith, D.R. 1979: 1381; Bolton, 1987: 294.

Syllophopsis Santschi, 1915c: 259 [as subgenus of Monomorium.] Type-species: Monomorium modestum, by monotypy.

Taxonomic history

Syllophopsis in Myrmicinae, Solenopsidini: Forel, 1917: 242 [subtribe Monomoriini]; Emery, 1922e: 175; all subsequent authors to the following.

Syllophopsis in Myrmicinae, Monomorium genus group: Ettershank, 1966: 81.

Syllophopsis in Myrmicinae, Monomoriini: Dlussky & Fedoseeva, 1988: 80.

Syllophopsis as subgenus of Monomorium: Santschi, 1915c: 259; Forel, 1917: 242; Emery, 1922e: 175; Wheeler, W.M. 1922a: 676; Donisthorpe, 1943g: 729; Arnold, 1952a: 465.

Syllophopsis as genus: Santschi, 1921c: 119; Santschi, 1936a: 32; Ettershank, 1966: 100.

Syllophopsis as junior synonym of Monomorium: Bolton, 1987: 287.

[Syllopsis Santschi, 1921c: 120, incorrect subsequent spelling.]

Corynomyrmex Viehmeyer, 1916a: 133 [as subgenus of Monomorium]. Type-species: Monomorium (Corynomyrmex) hospitum, by monotypy.

Taxonomic history

Corynomyrmex as subgenus of Monomorium: Viehmeyer, 1916a: 133; Forel, 1917: 242; Emery, 1922e: 174.

Corynomyrmex as junior synonym of Monomorium: Ettershank, 1966: 82 [provisional]; Smith, D.R. 1979: 1381; Bolton, 1987: 287.

Isolcomyrmex Santschi, 1917b: 296 [as subgenus of Monomorium]. Type-species: Holcomyrmex santschii (junior secondary homonym in Monomorium, replaced by Monomorium santschianum), by monotypy.

Taxonomic history

Isolcomyrmex as subgenus of Monomorium: Santschi, 1917b: 296; Wheeler, W.M. 1922a: 676.

Isolcomyrmex as junior synonym of Monomorium: Ettershank, 1966: 82; Smith, D.R. 1979: 1381; Bolton, 1987: 296.

[Isholcomyrmex Santschi, 1936a: 32, incorrect subsequent spelling.]

Paraphacota Santschi, 1919d: 90. Type-species: Paraphacota surcoufi, by monotypy.

Taxonomic history

[Type-species not Phacota noualhieri, unjustified subsequent designation by Santschi, 1927d: 245; repeated in Ettershank, 1966: 82 and Bolton, 1973a: 352.]

Paraphacota in Myrmicinae, Solenopsidini: Wheeler, W.M. 1922a: 663; Donisthorpe, 1943f: 681.

Paraphacota as junior synonym of Monomorium: Santschi, 1927d: 243; Ettershank, 1966: 82; Smith, D.R. 1979: 1381; Bolton, 1987: 293.

Equestrimessor Santschi, 1919d: 92 [as subgenus of Monomorium]. Type-species: Holcomyrmex chobauti, by subsequent designation of Donisthorpe, 1943f: 644.

Taxonomic history

Equestrimessor as subgenus of Monomorium: Santschi, 1919d: 92;Wheeler, W.M. 1922a: 677; Santschi, 1936a: 32.

Equestrimessor as junior synonym of Monomorium: Ettershank, 1966: 82; Smith, D.R. 1979: 1381; Bolton, 1987: 287, 297.

[Equessimessor Santschi, 1936a: 32, incorrect subsequent spelling.]

Xenhyboma Santschi, 1919g: 405. Type-species: Xenhyboma mystes (junior synonym of Monomorium medinae), by monotypy.

Taxonomic history

Xenhyboma in Myrmicinae, Solenopsidini: Donisthorpe, 1943g: 736.

Xenhyboma as junior synonym of Monomorium: Ettershank, 1966: 82 [provisional]; Smith, D.R. 1979: 1381; Espadaler, 1982b: 112; Bolton, 1987: 293.

Protholcomyrmex Wheeler, W.M. 1922a: 162 [as subgenus of Monomorium]. Type-species: Monomorium rothsteini, by original designation.

Taxonomic history

Protholcomyrmex as junior synonym of Chelaner: Ettershank, 1966: 93.

Ireneidris Donisthorpe, 1943c: 81. Type-species: Ireneidris myops (junior synonym of Monomorium talpa), by original designation.

Taxonomic history

Ireneidris in Myrmicinae, Solenopsidini [subtribe Monomoriini]: Donisthorpe, 1943c: 82.

Ireneidris as junior synonym of Monomorium: Ettershank, 1966: 82; Smith, D.R. 1979: 1381; Bolton, 1987: 298.

Schizopelta McAreavey, 1949: 14. Type-species: Schizopelta falcata, by original designation.

Taxonomic history

Schizopelta as junior synonym of Chelaner: Ettershank, 1966: 93.

Nothidris Ettershank, 1966: 105. Type-species: Monomorium latastei, by original designation.

Taxonomic history

Nothidris in Myrmicinae, Megalomyrmex genus group: Ettershank, 1966: 81; in Solenopsis genus group: Bolton, 1987: 271.

Nothidris in Myrmicinae, Megalomyrmecini: Dlussky & Fedoseeva, 1988: 80.

Nothidris in Myrmicinae, Solenopsidini: Kempf, 1972a: 165; Hölldobler & Wilson, 1990: 16; Jaffe, 1993: 10; Bolton, 1994: 106.

Nothidris as junior synonym of Monomorium: Fernández, in Heterick, 2006: 79; Fernández, 2007b: 130.

Pharaophanes Bernard, 1967: 168 (attributed to Santschi) [as subgenus of Monomorium]. Type-species: Formica pharaonis, by monotypy.

Taxonomic history

[Pharaophanes Bernard, 1953b: 238 (attributed to Santschi) [as subgenus of Monomorium]. Unavailable name; proposed without designation of type-species. Species included by Bernard (1953) are all referable to Monomorium: Bolton, 1987: 288; Bolton, 1995b: 42 (see also note in Baroni Urbani, 1971c: 90).]

Pharaophanes as junior synonym of Monomorium: Bolton, 2003: 213.

Antichthonidris Snelling, R.R. 1975: 5. Type-species: Monomorium denticulatum, by original designation.

Taxonomic history

Antichthonidris in Myrmicinae, Solenopsis genus group: Bolton, 1987: 271.

Antichthonidris in Myrmicinae, Solenopsidini: Hölldobler & Wilson, 1990: 16; Jaffe, 1993: 10; Bolton, 1994: 106.

Antichthonidris as junior synonym of Monomorium: Heterick, 2001: 361.

Epelysidris Bolton, 1987: 279. Type-species: Epelysidris brocha, by original designation.

Taxonomic history

Epelysidris in Myrmicinae, Solenopsis genus group: Bolton, 1987: 271.

Epelysidris in Myrmicinae, Solenopsidini: Hölldobler & Wilson, 1990: 16; Bolton, 1994: 106; Bolton, 1995a: 1049; Bolton, 1995b: 188.

Epelysidris as junior synonym of Monomorium: Fernández, in Heterick, 2006: 79; Fernández, 2007b: 130.

Genus Monomorium references

Roger, 1863b: 30, 31 (Phacota, Monomorium catalogues); Mayr, 1863: 429, 440 (Monomorium, Phacota catalogues); Mayr, 1865: 19, 22 (Trichomyrmex, Monomorium, Phacota diagnoses); Mayr, 1867a: 95 (Monomorium diagnosis); Mayr, 1876: 100 (Australia Monomorium species key); Emery, 1881b: 530 (Mediterranean & Red Sea Monomorium species key); André, 1883a: 331, 343 (Europe & Algeria Monomorium, Holcomyrmex species keys); André, 1883b: 397 (Phacota); Cresson, 1887: 262 (U.S.A. Monomorium catalogue); Nasonov, 1889: 69 (Russia Monomorium species key); Dalla Torre, 1893: 65, 71, 72, 78 (Holcomyrmex, Monomorium, Phacota, Trichomyrmex, †Lampromyrmex catalogues); Forel, 1903a: 685, 692 (India & Sri Lanka Monomorium, Holcomyrmex species keys); Bingham, 1903: 200, 282 (India, Sri Lanka & Burma Monomorium, Holcomyrmex species keys); Ruzsky, 1905b: 633 (Russian Empire Monomorium species key); Emery, 1908h: 664 (Palaearctic Monomorium species key); Emery, 1915i: 190 (Monomorium subgenera key); Arnold, 1916: 203, 204 (Monomorium diagnosis, South Africa species key); Emery, 1916b: 161 (Italy Monomorium species key); Santschi, 1921c: 121 (Syllophopsis species key); Emery, 1922e: 166 (Monomorium diagnosis, subgenera key, catalogue); Emery, 1922e: 168 (M. (Chelaner) & M. (Notomyrmex) diagnoses, catalogues); Emery, 1922e: 170 (M. (Monomorium) diagnosis, catalogue); Emery, 1922e: 174 (M. (Corynomyrmex) diagnosis, catalogue); Emery, 1922e: 175 (M. (Syllophopsis) & M. (Xeromyrmex) diagnoses, catalogues); Emery, 1922e: 179 (M. (Parholcomyrmex) diagnosis, catalogue); Emery, 1922e: 181 (M. (Holcomyrmex) diagnosis, catalogue); Emery, 1922e: 183 (M. (Lampromyrmex) diagnosis, catalogue); Emery, 1922e: 184 (Epixenus diagnosis, catalogue); Emery, 1922e: 185 (Trichomyrmex diagnosis, catalogue); Emery, 1922e: 186 (Wheeleriella diagnosis, catalogue); Emery, 1922e: 187 (Phacota diagnosis, catalogue); Emery, 1922e: 204 (Epoecus diagnosis, catalogue); Wheeler, W.M. 1922a: 161, 675, 862, 1026 (Monomorium diagnosis, subgenera key, Afrotropical, Malagasy catalogues); Menozzi, 1933b: 64 (Israel Monomorium species key); Santschi, 1936a: 33 (Mediterranean & Africa M. (Xeromyrmex) key); Buren, 1944a: 289 (U.S.A., Iowa Monomorium species key); Kusnezov, 1949a: 431 (Argentina Monomorium species); Creighton, 1950a: 217 (North America Monomorium species key); Creighton, 1950a: 239 (Epoecus, review of genus); Chapman & Capco, 1951: 162, 169 (Asia Ireneidris, Monomorium, Trichomyrmex, Wheeleriella checklists); Bernard, 1955b: 282 (Epixenus species key); Brown, 1958h: 28 (New Zealand Monomorium species key); Gregg, 1963: 367 (U.S.A., Colorado Monomorium species key); Ettershank, 1966: 82, 93, 100 (Monomorium, Chelaner, Syllophopsis diagnoses, reviews of genera, checklists); Bernard, 1967: 165 (Monomorium diagnosis, Western Europe species key); Kempf, 1972a: 143 (Neotropical Monomorium catalogue); Kempf, 1972a: 165 (Nothidris catalogue); Alayo, 1974: 14 (Cuba Monomorium species key); Snelling, R.R. 1975: 5 (Nothidris all species key); Snelling, R.R. & Hunt, 1976: 79 (Nothidris all species key); Arnol'di & Dlussky, 1978: 538 (former European U.S.S.R. Monomorium species key); Collingwood, 1978: 82 (Iberian Peninsula Monomorium species key); Smith, D.R. 1979: 1381 (North America Monomorium catalogue); Tohmé & Tohmé, 1980a: 1087 (Lebanon & Syria Epixenus species key); Collingwood, 1985: 267 (Saudi Arabia Monomorium species key); Taylor & Brown, D.R. 1985: 55, 70 (Australia Chelaner, Monomorium catalogues); DuBois, 1986: 74 (Nearctic Monomorium species revision, key); Bolton, 1987: 281 (Phacota diagnosis, review of genus); Bolton, 1987: 283 (Antichthonidris, review of genus); Bolton, 1987: 284 (Nothidris review of genus); Bolton, 1987: 287 (Monomorium diagnosis, review of genus, Afrotropical species revision, key); Taylor, 1987a: 19, 40 and Taylor, 1987b: 2 (Australia, New Caledonia & New Zealand Monomorium checklist); Agosti & Collingwood, 1987b: 272 (Balkans Monomorium species key); Hölldobler & Wilson, 1990: 14 (synoptic classification); Dlussky, Soyunov & Zabelin, 1990: 231 (Turkmenistan Monomorium species key); Brandão, 1991: 326, 357 (Neotropical Monomorium catalogue); Brandão, 1991: 361 (Nothidris catalogue); Morisita, Kubota, et al. 1992: 38 (Japan Monomorium species key); Arakelian, 1994: 46 (Armenia Monomorium species key); Bolton, 1995a: 1048, 1050, 1051 (census); Bolton, 1995b: 67, 258, 292, 316 (Antichthonidris, Monomorium, Nothidris, Phacota catalogues); Wu, J. & Wang, 1995: 88 (China Monomorium species key); Collingwood & Agosti, 1996: 337 (Saudi Arabia Monomorium species key); Radchenko, 1997f: 211 (M. scabriceps group revision, key); Collingwood & Prince, 1998: 19 (Portugal Monomorium species key); Shattuck, 1999: 145 (Australia synopsis); Heterick, 2001: 367 (Australia Monomorium species revision, key); Zhou, 2001b: 112 (China, Guangxi Monomorium species key); Mackay & Mackay, 2002: 127 (U.S.A., New Mexico Monomorium species key); Heterick, 2006: 84 (Malagasy Monomorium species revision, key); Fernández, 2007b: 139 (Neotropical Monomorium species key); Terayama, 2009: 154 (Taiwan species key); Heterick, 2009: 158 (south-western Australia species key); Aldawood & Sharaf, 2011: 52 (Saudi Arabia M. monomorium group key).

Myrmicinae Solenopsidini Solenopsis TRUE TRUE FALSE

Extant: 183 species, 75 subspecies

Fossil: 8 species

Solenopsis Westwood, 1840b: 86. Type-species: Solenopsis mandibularis (junior synonym of Atta geminata), by monotypy.

Taxonomic history

Solenopsis in Poneridae, Attidae: Smith, F. 1858b: 177.

Solenopsis in Attidae: Smith, F. 1860a: 74; Smith, F. 1861b: 48.

Solenopsis in Myrmicidae: Smith, F. 1871a: 333; Cresson, 1887: 262.

Solenopsis in Myrmicidae, Pheidolidae: Emery, 1877a: 81.

Solenopsis in Myrmicinae: Mayr, 1865: 24 [Myrmicidae]; Emery & Forel, 1879a: 464 [Myrmicidae]; Dalla Torre, 1893: 75.

Solenopsis in Myrmicinae, Solenopsis genus group: Ettershank, 1966: 81; Bolton, 1987: 271.

Solenopsis in Myrmicinae, Solenopsidini: Forel, 1893a: 164; Forel, 1895b: 130; Emery, 1895j: 770; Forel, 1899c: 79; Ashmead, 1905b: 383; Wheeler, W.M. 1910g: 140; Emery, 1914a: 41; Arnold, 1916: 242; Forel, 1917: 243; Emery, 1922e: 195; Wheeler, W.M. 1922a: 663; Kempf, 1972a: 232; Smith, D.R. 1979: 1384; Dlussky & Fedoseeva, 1988: 80; Hölldobler & Wilson, 1990: 16; Jaffe, 1993: 10; Bolton, 1994: 106.

Junior synonyms of SOLENOPSIS

Diplorhoptrum Mayr, 1855: 449. Type-species: Formica fugax, by monotypy.

Taxonomic history

Diplorhoptrum in Myrmicinae: Mayr, 1855: 449 [Myrmicidae]; Mayr, 1861: 72 [Myrmicidae].

Diplorhoptrum in Myrmicinae, Solenopsidini: Emery, 1922e: 195; all subsequent authors.

Diplorhoptrum as genus: Mayr, 1855: 449; Baroni Urbani, 1968b: 68; Kutter, 1977c: 98; Bernard, 1978a: 543; Arnol'di & Dlussky, 1978: 539; Collingwood, 1979: 64; Dlussky & Fedoseeva, 1988: 80; Kupyanskaya, 1990: 132; Atanasov & Dlussky, 1992: 167; Dlussky & Radchenko, 1994: 102; Baroni Urbani, 1995: 20 (footnote).

Diplorhoptrum as subgenus of Solenopsis: Creighton, 1930b: 43; Santschi, 1934e: 566 (in text), 568; Creighton, 1950a: 233; Smith, M.R. 1951a: 813; Kusnezov, 1956: 27; Kusnezov, 1957a: 273; Kusnezov, 1962b: 160; Smith, M.R. 1967: 358.

Diplorhoptrum as junior synonym of Solenopsis: Mayr, 1862: 751; Roger, 1863b: 32; Mayr, 1863: 453; Emery & Forel, 1879a: 464; Dalla Torre, 1893: 75; Forel, 1915d: 9; Emery, 1922e: 195; Wheeler, W.M. 1922a: 877; Donisthorpe, 1943f: 639; Ettershank, 1966: 134; Kempf, 1972a: 232; Smith, D.R. 1979: 1384; Bolton, 1987: 285; Bolton, 1994: 106; Bolton, 1995b: 27.

Octella Forel, 1915b: 70 [as subgenus of Oligomyrmex]. Type-species: Oligomyrmex (Octella) pachycerus, by original designation.

Taxonomic history

Octella as subgenus of Oligomyrmex: Forel, 1917: 243; Wheeler, W.M. 1922a: 687; Emery, 1924d: 218.

Octella as junior synonym of Oligomyrmex: Ettershank, 1966: 119.

Octella as junior synonym of Solenopsis: Taylor, 1991db: 611.

Synsolenopsis Forel, 1918b: 155 [as subgenus of Solenopsis]. Type-species: Solenopsis (Synsolenopsis) bruchi (junior primary homonym in Solenopsis, replaced by Solenopsis bruchiella), by monotypy.

Taxonomic history

Synsolenopsis as subgenus of Solenopsis: Forel, 1918b: 155; Emery, 1922e: 196.

Synsolenopsis as genus: Creighton, 1930b: 41 (in text); Kusnezov, 1953c: 341; Kusnezov, 1957a: 272; Kusnezov, 1962b: 160; Kusnezov, 1964: 61.

Synsolenopsis as junior synonym of Solenopsis: Ettershank, 1966: 134.

Diagyne Santschi, 1923c: 268 [as subgenus of Solenopsis]. Type-species: Solenopsis succinea by monotypy.

Taxonomic history

Diagyne in Myrmicinae, Solenopsidini: Donisthorpe, 1943f: 638.

Diagyne as subgenus of Solenopsis: Santschi, 1923c: 268; Creighton, 1930b: 42; Kusnezov, 1962b: 160.

Diagyne junior synonym of Solenopsis: Ettershank, 1966: 134.

Labauchena Santschi, 1930d: 81. Type-species: Labauchena daguerrei, by monotypy.

Taxonomic history

Labauchena in Myrmicinae, Solenopsidini: Santschi, 1930d: 81; all subsequent authors.

Labauchena as junior synonym of Solenopsis: Ettershank, 1966: 134.

Euophthalma Creighton, 1930b: 43 [as subgenus of Solenopsis]. Type-species: Myrmica globularia, by original designation.

Taxonomic history

Euophthalma as junior synonym of Solenopsis: Ettershank, 1966: 134.

Oedaleocerus Creighton, 1930b: 43 [as subgenus of Solenopsis]. Type-species: Solenopsis angulata, by original designation.

Taxonomic history

Oedaleocerus in Myrmicinae, Solenopsidini: Donisthorpe, 1943f: 677.

Oedaleocerus as subgenus of Solenopsis: Creighton, 1930b: 43.

Oedaleocerus as genus: Kusnezov, 1957a: 272; Kusnezov, 1962b: 160; Kusnezov, 1964: 61.

Oedaleocerus as junior synonym of Solenopsis: Ettershank, 1966: 134.

Bisolenopsis Kusnezov, 1953d: 9. Type-species: Bisolenopsis sea, by monotypy.

Taxonomic history

Bisolenopsis in Myrmicinae, Solenopsidini: Kusnezov, 1953d: 9; Kusnezov, 1957a: 272; Kusnezov, 1962b: 159; Kusnezov, 1964: 61.

Bisolenopsis as junior synonym of Solenopsis: Ettershank, 1966: 134.

[Disolenopsis Snelling, R.R. 1981: 397, incorrect subsequent spelling.]

Paranamyrma Kusnezov, 1953d: 17. Type-species: Paranamyrma solenopsidis, by monotypy.

Taxonomic history

[Paranamyrma also described as new by Kusnezov, 1954a: 9.]

Paranamyrma in Myrmicinae, Solenopsidini: Kusnezov, 1957a: 267 (in key); Kusnezov, 1962b: 160; Kusnezov, 1964: 61.

Paranamyrma as junior synonym of Solenopsis: Ettershank, 1966: 134.

Granisolenopsis Kusnezov, 1956: 26 [as subgenus of Solenopsis] (diagnosis in key). Type-species: Solenopsis (Granisolenopsis) granivora, by original designation.

Taxonomic history

[Granisolenopsis also described as new by Kusnezov, 1957a: 277.]

Granisolenopsis as junior synonym of Solenopsis: Ettershank, 1966: 134.

Lilidris Kusnezov, 1956: 26 (diagnosis in key). Type-species: Lilidris metatarsalis, by original designation.

Taxonomic history

[Lilidris also described as new by Kusnezov, 1957a: 274 and Kusnezov, 1958b: 189.]

Lilidris in Myrmicinae, Solenopsidini: Kusnezov, 1957a: 274; Kusnezov, 1958b: 189; Kusnezov, 1962b: 159; Kusnezov, 1964: 61.

Lilidris as junior synonym of Solenopsis: Ettershank, 1966: 134.

Genus Solenopsis references

Smith, F. 1858b: 177 (diagnosis); Roger, 1863b: 32 (catalogue); Mayr, 1863: 453 (catalogue); Mayr, 1865: 24 (diagnosis); Mayr, 1867a: 109 (diagnosis); Mayr, 1870b: 996 (all species key); André, 1883b: 387 (Europe & Algeria species key); Cresson, 1887: 262 (U.S.A. catalogue); Dalla Torre, 1893: 75 (catalogue); Emery, 1896g: 82 (New World species key); Forel, 1903a: 689 (India & Sri Lanka species key); Bingham, 1903: 158 (India, Sri Lanka & Burma species key); Emery, 1909a: 29 (Palaearctic species key); Emery, 1916b: 165 (Italy species key); Arnold, 1916: 242 (diagnosis); Emery, 1922e: 195 (diagnosis, catalogue); Wheeler, W.M. 1922a: 163, 877, 1028 (diagnosis, Afrotropical, Malagasy catalogues); Emery, 1924d: 218 (Oligomyrmex (Octella) diagnosis, catalogue); Creighton, 1930b: 42 (subgenera key); Creighton, 1930b: 45 (S. (Solenopsis) species key); Creighton, 1930b: 106 (S. (Euophthalma) species key); Cole, 1942: 361 (U.S.A., Utah species key); Bernard, 1950a: 5 (France species key); Creighton, 1950a: 228 (North America species key); Chapman & Capco, 1951: 167 (Asia checklist); Wilson, 1952b: 50 (South America S. saevissima complex, key); Kusnezov, 1953c: 347 (Synsolenopsis species key); Gregg, 1963: 370 (U.S.A., Colorado species key); Snelling, R.R. 1963: 1 (U.S.A S. geminata group species); Ettershank, 1966: 134 (diagnosis, review of genus, checklist); Bernard, 1967: 170 (diagnosis, Western Europe species key); Kusnezov, 1969: 37 (Bisolenopsis species key); Buren, 1972: 1 (S. geminata group, partial revision); Kempf, 1972a: 232 (Neotropical catalogue); Snelling, R.R. & Hunt, 1976: 81 (Chile species key); Kutter, 1977c: 99 (Switzerland species key); Arnol'di & Dlussky, 1978: 539 (former European U.S.S.R. species key); Collingwood, 1978: 82 (Iberian Peninsula species key); Bernard, 1978a: 573 (France species key); Smith, D.R. 1979: 1385 (North America catalogue); Allred, 1982: 443 (U.S.A., Utah species key); Gösswald, 1985: 302 (Germany species key); Taylor & Brown, D.R. 1985: 87 (Australia catalogue); Wheeler, G.C. & Wheeler, J. 1986g: 50 (U.S.A., Nevada species key); Taylor, 1987a: 72 (Australia, New Caledonia checklist); Agosti & Collingwood, 1987b: 273 (Balkans species key); Bolton, 1987: 285 (synonymy notes); Thompson & Johnson, 1989: 697 (U.S.A., Florida species key); Dlussky, Soyunov & Zabelin, 1990: 240 (Turkmenistan species key); Brandão, 1991: 378 (Neotropical catalogue); Trager, 1991: 141 (S. geminata group revision, key); Morisita, Kubota, et al. 1992: 41 (Japan species key); Arakelian, 1994: 48 (Armenia species key); Dlussky & Radchenko, 1994: 102 (Central Palaearctic species key); Bolton, 1995a: 1052 (census); Bolton, 1995b: 386 (catalogue); Wu, J. & Wang, 1995: 70 (China species key); Collingwood & Agosti, 1996: 358 (Saudi Arabia species key); Collingwood & Prince, 1998: 18 (Portugal species key); Shattuck, 1999: 165 (Australia synopsis); Taber, 2000: 234 (U.S.A. S. geminata group, key); Snelling, R.R. 2001: 522 (Puerto Rico S. fugax group, key); Mackay & Mackay, 2002: 196 (U.S.A., New Mexico species key); Terayama, 2009: 157 (Taiwan species key); Heterick, 2009: 176 (south-western Australia species key).

Myrmicinae Myrmicini Manica TRUE TRUE FALSE

6 species

Manica Jurine, 1807: 276. Type-species: Formica rubida, by subsequent designation of Wheeler, W.M. 1911f: 166.

Taxonomic history

Manica in Myrmicinae, Myrmicini: Wheeler, W.M. 1922a: 660; all subsequent authors.

Manica as junior synonym of Myrmica: Roger, 1863b: 28; Mayr, 1863: 431; Emery & Forel, 1879a: 460; Dalla Torre, 1893: 108; Forel, 1915d: 9.

Manica as subgenus of Myrmica: Emery, 1921f: 42; Wheeler, W.M. 1922a: 660; Smith, M.R. 1951a: 791.

Manica as genus: Jurine, 1807: 276; Weber, 1947: 439; Creighton, 1950a: 105; Smith, M.R. 1958c: 113; all subsequent authors.

Junior synonyms of MANICA

Neomyrma Forel, 1914a: 275 [as subgenus of Aphaenogaster]. Type-species: Aphaenogaster (Neomyrma) calderoni (junior synonym of Myrmica bradleyi), by monotypy.

Taxonomic history

Neomyrma as subgenus of Aphaenogaster: Forel, 1914a: 275.

Neomyrma as subgenus of Myrmica: Forel, 1915c: 364 (in text); Emery, 1915d: 69; Emery, 1916b: 120; Forel, 1917: 241.

Neomyrma as genus: Bondroit, 1918: 97.

Neomyrma as junior synonym of Myrmica: Donisthorpe, 1916b: 242.

Neomyrma as junior synonym of Manica: Emery, 1921f: 42; Wheeler, W.M. 1922a: 660.

Oreomyrma Wheeler, W.M. 1914d: 118 (in text) [as subgenus of Myrmica]. Type-species: Formica rubida (quoted by Wheeler as Myrmica rubida), by original designation.

Taxonomic history

Oreomyrma as junior synonym of Neomyrma: Wheeler, W.M. 1915a: 50; Forel, 1915c: 364; Forel, 1915d: 9; Donisthorpe, 1916b: 242. [Manica and Oreomyrma share the same type-species, synonymy is therefore absolute.]

Genus Manica references

Wheeler, W.M. 1914d: 119 (U.S.A. species key); Emery, 1921f: 42 (diagnosis, catalogue); Creighton, 1950a: 108 (North America species key); Bernard, 1967: 125 (diagnosis); Smith, D.R. 1979: 1352 (North America catalogue); Allred, 1982: 440 (U.S.A., Utah species key); Wheeler, G.C. & Wheeler, J. 1986g: 25 (U.S.A., Nevada species key); Morisita, Kubota, et al. 1992: 13 (Japan species key); Bolton, 1994: 106 (synoptic classification); Bolton, 1995a: 1050 (census); Bolton, 1995b: 249 (catalogue).

Myrmicinae Myrmicini Myrmica TRUE TRUE FALSE

Extant: 171 species, 2 subspecies

Fossil: 5 species

Myrmica Latreille, 1804: 179. Type-species: Formica rubra, by subsequent designation of Latreille, 1810: 437; see also Yarrow, 1955b: 113.

Taxonomic history

Myrmica in Myrmicites: Lepeletier de Saint-Fargeau, 1835: 180.

Myrmica in Poneridae, Myrmicidae: Smith, F. 1858b: 114.

Myrmica in Myrmicidae: Smith, F. 1871a: 324; Cresson, 1887: 260.

Myrmica in Myrmicinae: Mayr, 1855: 396 [Myrmicidae]; Smith, F. 1857a: 70 [Myrmicidae]; Mayr, 1861: 62 [Myrmicidae]; Mayr, 1865: 19 [Myrmicidae]; Emery & Forel, 1879a: 460 [Myrmicidae]; Dalla Torre, 1893: 108.

Myrmica in Myrmicinae, Myrmicini: Forel, 1895b: 125; Emery, 1895j: 769; Forel, 1899c: 63; Ashmead, 1905b: 383; Wheeler, W.M. 1910g: 140; Emery, 1914a: 40; Wheeler, W.M. 1915h: 59; Forel, 1917: 241; Bondroit, 1918: 96; Emery, 1921f: 36; Wheeler, W.M. 1922a: 660; all subsequent authors.

Junior synonyms of MYRMICA

Sifolinia Emery, 1907: 49. Type-species: Sifolinia laurae, by monotypy.

Taxonomic history

Sifolinia in Myrmicinae, Pheidolini: Emery, 1914a: 40; Forel, 1917: 241; Emery, 1922e: 117; Wheeler, W.M. 1922a: 661.

Sifolinia in Myrmicinae, Myrmicini: Wheeler, W.M. 1910g: 139; Collingwood, 1979: 58; Dlussky & Fedoseeva, 1988: 79.

Sifolinia as junior synonym of Myrmica: Brown, 1973b: 184 [provisional]; Bolton, 1988a: 3; Radchenko & Elmes, 2003a: 219.

†Nothomyrmica Wheeler, W.M. 1915h: 60. Type-species: †Macromischa rudis, by original designation.

Taxonomic history

[†Nothomyrmica Wheeler, W.M. 1908g: 413, and Wheeler, W.M. 1910g: 167, nomina nuda.]

†Nothomyrmica in Myrmicinae, Myrmicini: Wheeler, W.M. 1915h: 60; all subsequent authors except the following.

†Nothomyrmica in Myrmicinae, Tetramoriini: Donisthorpe, 1943f: 675 (error).

†Nothomyrmica in Myrmicinae, Leptothoracini: Dlussky & Fedoseeva, 1988: 79; Bolton, 1994: 106; Bolton, 1995b: 292.

†Nothomyrmica incertae sedis in Myrmicinae, Myrmicini: Bolton, 2003: 62.

†Nothomyrmica as junior synonym of Myrmica: Radchenko, Dlussky & Elmes, 2007: 1495.

Sommimyrma Menozzi, 1925d: 25. Type-species: Sommimyrma symbiotica, by original designation.

Taxonomic history

Sommimyrma in Myrmicinae, Myrmicini: Donisthorpe, 1943g: 726.

Sommimyrma as junior synonym of Myrmica: Brown, 1973b: 184 [provisional]; Bolton, 1988a: 3; Radchenko & Elmes, 2003a: 224.

Symbiomyrma Arnol'di, 1930c: 267. Type-species: Symbiomyrma karavajevi, by monotypy.

Taxonomic history

[Symbiomyrma also described as new by Arnol'di, 1933a: 41.]

Symbiomyrma in Myrmicinae, Pheidolini: Karavaiev, 1934: 102.

Symbiomyrma in Myrmicinae, Myrmicini: Donisthorpe, 1943g: 729.

Symbiomyrma as genus: Karavaiev, 1934: 102; Seifert, 1994: 15; Seifert, 1996b: 236.

Symbiomyrma as junior synonym of Sifolinia: Samsinak, 1964: 156.

Symbiomyrma as junior synonym of Myrmica: Bolton, 1988a: 3; Bolton, 1994: 106; Radchenko & Elmes, 2003a: 219.

Paramyrmica Cole, 1957a: 37. Type-species: Paramyrmica colax, by original designation.

Taxonomic history

Paramyrmica in Myrmicinae, Myrmicini: Wheeler, G.C. & Wheeler, J. 1985: 257.

Paramyrmica as genus: Gregg, 1961: 215; Smith, M.R. 1967: 350; Smith, D.R. 1979: 1351.

Paramyrmica as junior synonym of Myrmica: Brown, 1973b: 183 [provisional]; Bolton, 1988a: 3.

Dodecamyrmica Arnol'di, 1968: 1803 [as subgenus of Myrmica]. Type-species: Myrmica arnoldii, by original designation.

Taxonomic history

Dodecamyrmica as junior synonym of Myrmica: Brown, 1973b: 180 [provisional]; Francoeur, 1981: 759; Bolton, 1988a: 4.

Genus Myrmica references

Mayr, 1855: 396, 397 (diagnosis, Austria species key); Smith, F. 1858b: 114 (diagnosis); Mayr, 1861: 62 (Europe species key); Roger, 1863b: 28 (catalogue); Mayr, 1863: 429 (catalogue); Mayr, 1865: 19 (diagnosis); André, 1874: 192 (Europe species key); Forel, 1874: 75 (Switzerland species key); André, 1883a: 315 (Europe & Algeria species key); Cresson, 1887: 260 (U.S.A. catalogue); Provancher, 1887: 246 (Canada species key); Nasonov, 1889: 71 (Russia species key); Lameere, 1892: 68 (Belgium species key); Dalla Torre, 1893: 108 (catalogue); Forel, 1903a: 696 (India species key); Bingham, 1903: 267 (India, Sri Lanka & Burma species key); Ruzsky, 1905b: 652 (Russian Empire species key); Wasmann, 1906: 15 (Luxemburg species key); Emery, 1908a: 165 (Palaearctic species key); Bondroit, 1910: 493 (Belgium species key); Stitz, 1914: 70 (Central Europe species key); Donisthorpe, 1915d: 110 (Britain species key); Emery, 1916b: 121 (Italy species key); Wheeler, W.M. 1916m: 581 (U.S.A., Connecticut species key); Bondroit, 1918: 98 (France & Belgium species key); Emery, 1921f: 36 (diagnosis, catalogue); Emery, 1922e: 117 (Sifolinia diagnosis, catalogue); Finzi, 1926: 77 (diagnosis, European species key); Kuznetsov-Ugamsky, 1927e: 190 (Turkestan species key); Karavaiev, 1927c: 258 (Ukraine species key); Donisthorpe, 1927b: 118 (Britain species key); Arnol'di, 1933b: 599 (Russia species key); Arnol'di, 1934: 151 (former European U.S.S.R. species, biometrics); Karavaiev, 1934: 63 (Ukraine species key); Menozzi, 1939a: 293 (Himalaya & Tibet species key); Stitz, 1939: 74 (Germany species key); Kratochvíl, 1941: 75 (Central Europe species key); Novák & Sadil, 1941: 75 (Central Europe species key); Cole, 1942: 367 (U.S.A., Utah species key); Holgersen, 1943b: 168 (Norway species key); Holgersen, 1944: 199 (Norway species key); Buren, 1944a: 281 (U.S.A., Iowa species key); Boven, 1947: 175 (Belgium species key); Weber, 1947: 444 (North America species key); Creighton, 1950a: 92 (North America species key); Chapman & Capco, 1951: 124 (Asia checklist); Sadil, 1952: 264 (Czechia & Slovakia species key); Collingwood, 1958b: 65 (Britain species key); Boven, 1959: 7 (Netherlands species key); Gregg, 1963: 290 (U.S.A., Colorado species key); Wheeler, G.C. & Wheeler, J. 1963: 96 (U.S.A., North Dakota species key); Collingwood, 1964: 101 (Britain species key); Bernard, 1967: 107 (diagnosis, Western Europe species key); Bernard, 1967: 156 (Sifolinia diagnosis); Boven, 1970b: 16 (Netherlands species key); Arnol'di, 1970b: 1832 (former European U.S.S.R. species key); Kutter, 1973c: 253 (morphology, satellite genera, Sifolinia, Sommimyrma, Symbiomyrma, key to first); Bolton & Collingwood, 1975: 5 (Britain species key); Arnol'di, 1976a: 554 (central Asia & Kazakhstan species key); Tarbinsky, 1976: 20 (Kyrghyzstan species key); Boven, 1977: 94 (Belgium species key); Kutter, 1977c: 43 (Switzerland species key); Arnol'di & Dlussky, 1978: 530 (former European U.S.S.R. species key); Collingwood, 1978: 78 (Iberian Peninsula species key); Collingwood, 1979: 41 (Fennoscandia & Denmark species key); Smith, D.R. 1979: 1347, 1351 (North America Myrmica, Paramyrmica catalogues); Francoeur, 1981: 755 (M. lampra group); Allred, 1982: 440 (U.S.A., Utah species key); Gösswald, 1985: 294 (Germany species key); Wheeler, G.C. & Wheeler, J. 1986g: 22 (U.S.A., Nevada species key); Kupyanskaya, 1986a: 83 (Far Eastern Russia M. lobicornis group, key); Nilsson & Douwes, 1987: 62 (Norway species key); Agosti & Collingwood, 1987b: 267 (Balkans species key); Seifert, 1988b: 43 (Europe, Asia Minor & Caucasus species key); Bolton, 1988a: 3 (diagnosis, review of genus, venation); Dlussky, Soyunov & Zabelin, 1990: 181 (Turkmenistan species key); Kupyanskaya, 1990: 92 (Far Eastern Russia species key); Morisita, Kubota, et al. 1992: 8 (Japan species key); Atanasov & Dlussky, 1992: 80 (Bulgaria species key); Arakelian, 1994: 20 (Armenia species key); Radchenko, 1994b: 105 (South Siberia species key); Radchenko, 1994c: 130 (Central & Eastern Palaearctic species key); Douwes, 1995: 86 (Sweden species key); Bolton, 1994: 106 (synoptic classification); Kupyanskaya, 1995: 337 (Far Eastern Russia species key); Bolton, 1995a: 1051 (census); Bolton, 1995b: 277 (catalogue); Wu, J. & Wang, 1995: 91 (China species key); Seifert, 1996b: 140 (Central Europe species key); Skinner & Allen, 1996: 45 (Britain species key); Radchenko, Czechowski & Czechowsa, 1997: 481 (Poland species key); Collingwood & Prince, 1998: 12 (Portugal species key); Radchenko & Elmes, 1998: 3 (M. ritae group, key); Elmes & Radchenko, 1998: 218 (Taiwan species key); Wei, Zhou, He & Liu, 2001: 561 (China species key); Radchenko & Elmes, 2001: 262 (Himalaya species key); Czechowski, Radchenko & Czechowska, 2002: 141 (Poland species key); Radchenko & Elmes, 2003a: 239 (Palaearctic socially parasitic species key); Mackay & Mackay, 2002: 133 (U.S.A., New Mexico species key); Radchenko, 2005b: 192 (North Korea species key); Radchenko, Elmes & Bui, 2006: 43 (Vietnam species key); Radchenko, Elmes & Alicata, 2006: 531 (Palaearctic schencki group key); Seifert, 2007: 131 (North and Central Europe species key); Francoeur, 2007: 156 (punctiventris group key); Radchenko, Dlussky & Elmes, 2007: 1497 (late Eocene amber species key); Terayama, 2009: 160 (Taiwan species key); Radchenko & Elmes, 2009a: 85 (pachei group key); Radchenko & Elmes, 2010: 713 (Palaearctic, Oriental, all species revision, keys).

Myrmicinae Myrmicini Pogonomyrmex TRUE TRUE FALSE

Extant: 67 species, 9 subspecies

Fossil: 1 species

Pogonomyrmex Mayr, 1868b: 169. Type-species: Formica badia, by subsequent designation of Wheeler, W.M. 1911f: 170.

Taxonomic history

Pogonomyrmex in Myrmicidae, Myrmicidae: Emery, 1877a: 81.

Pogonomyrmex in Myrmicidae: Cresson, 1887: 260.

Pogonomyrmex in Myrmicinae: Dalla Torre, 1893: 118.

Pogonomyrmex in Myrmicinae, Myrmicini: Forel, 1895b: 125; Emery, 1895j: 769; Forel, 1899c: 61; Ashmead, 1905b: 383; Wheeler, W.M. 1910g: 140; Emery, 1914a: 40; Forel, 1917: 240; Emery, 1921f: 44; Wheeler, W.M. 1922a: 660; all subsequent authors.

Junior synonyms of POGONOMYRMEX

Ephebomyrmex Wheeler, W.M. 1902c: 390 [as subgenus of Pogonomyrmex]. Type-species: Pogonomyrmex naegelii, by subsequent designation of Wheeler, W.M. 1911f: 163.

Taxonomic history

Ephebomyrmex in Myrmicinae, Myrmicini: Ashmead, 1905b: 383; Forel, 1917: 240; Emery, 1921f: 48; Wheeler, W.M. 1922a: 660; all subsequent authors.

Ephebomyrmex as subgenus of Pogonomyrmex: Wheeler, W.M. 1902c: 390; Wheeler, W.M. 1910g: 140; Forel, 1917: 240; Emery, 1921f: 48; Wheeler, W.M. 1922a: 660; Gallardo, 1932b: 94; Donisthorpe, 1943f: 643; Kusnezov, 1949c: 293; Creighton, 1950a: 132; Kusnezov, 1951a: 245; Smith, M.R. 1951a: 794; Smith, M.R. 1958c: 115; Kusnezov, 1964: 56; Kusnezov, 1956: 118; Smith, M.R. 1967: 351; Cole, 1968: 22; Smith, D.R. 1979: 1357; Snelling, R.R. 1982a: 108; Mackay, Mackay, Dominguez, et al. 1985: 42.

Ephebomyrmex as genus: Creighton, 1957a: 54; Kusnezov, 1960b: 350; Kempf, 1972a: 106; Brown, 1973b: 180; Snelling, R.R. 1981: 395; Wheeler, G.C. & Wheeler, J. 1985: 257; Dlussky & Fedoseeva, 1988: 79; Mackay & Vinson, 1989b: 14; Hölldobler & Wilson, 1990: 13; Taber, 1998: 146.

Ephebomyrmex as junior synonym of Pogonomyrmex: Lattke, 1991a: 305; Bolton, 1994: 106; Bolton, 1995b: 28; Fernández & Palacio, 1998: 1649; Bolton, 2003: 223.

Forelomyrmex Wheeler, W.M. 1913a: 80 [as subgenus of Pogonomyrmex].

Taxonomic history

[Replacement name for Janetia Forel, 1899c: 61 (footnote); junior homonym of Janetia Kieffer, 1896: 236 (Diptera).]

Forelomyrmex as subgenus of Pogonomyrmex: Wheeler, W.M. 1913a: 80; Forel, 1917: 240; Emery, 1921f: 48; Wheeler, W.M. 1922a: 660; Donisthorpe, 1943f: 646; Kusnezov, 1951a: 245; Kempf, 1972a: 209.

Forelomyrmex as junior synonym of Pogonomyrmex: Brown, 1973b: 180 [provisional]; Snelling, R.R. 1981: 395; Lattke, 1991a: 305; Bolton, 1994: 106; Bolton, 1995b: 29; Fernández & Palacio, 1998: 1649.

Homonym replaced by Forelomyrmex

Janetia Forel, 1899c: 61 (footnote) [as subgenus of Pogonomyrmex]. Type-species: Pogonomyrmex (Janetia) mayri, by monotypy.

Taxonomic history

[Junior homonym of Janetia Kieffer, 1896: 236 (Diptera).]

Genus Pogonomyrmex references

Mayr, 1870b: 970 (all species key); Mayr, 1887: 608 (all species key); Cresson, 1887: 260 (U.S.A. catalogue); Dalla Torre, 1893: 118 (catalogue); Wheeler, W.M. 1902a: 97 (North America species key); Emery, 1921f: 44 (diagnosis, subgenera key, catalogue); Emery, 1921f: 48 (P. (Ephebomyrmex) & P. (Forelomyrmex) diagnoses, catalogues); Gallardo, 1932b: 95 (Argentina species key); Cole, 1942: 364 (U.S.A., Utah species key); Kusnezov, 1949c: 301 (Argentina P. (Ephebomyrmex) species key); Creighton, 1950a: 113 (North America species key); Kusnezov, 1951a: 254 (Argentina species key); Kusnezov, 1954c: 85 (Bolivia species key); Creighton, 1957a: 54 (North America Ephebomyrmex); Gregg, 1963: 317 (U.S.A., Colorado species key); Cole, 1968: 21, 38, 155 (North America Pogonomyrmex & P. (Ephebomyrmex) diagnoses, species revision, keys); Kempf, 1972a: 106, 206 (Neotropical Ephebomyrmex, Pogonomyrmex catalogues); Snelling, R.R. & Hunt, 1976: 72 (Chile species key); Smith, D.R. 1979: 1353 (North America catalogue); Allred, 1982: 442 (U.S.A., Utah species key); Snelling, R.R. 1982a: 104 (supplement to Cole, 1968 key); Snelling, R.R. 1982a: 108 (North America P. (Ephebomyrmex) species key); Mackay, Mackay, Dominguez, et al. 1985: 42 (Mexico species key); Wheeler, G.C. & Wheeler, J. 1986g: 26 (U.S.A., Nevada species key); Wheeler, G.C. & Wheeler, J. 1986g: 33 (U.S.A., Nevada Ephebomyrmex species key); Shattuck, 1987: 175 (P. occidentalis complex, key); Taber, 1990: 307 (U.S.A. species complexes, phylogeny); Brandão, 1991: 342, 372 (Neotropical catalogue); Bolton, 1994: 106 (synoptic classification); Bolton, 1995a: 1052 (census); Bolton, 1995b: 339 (catalogue); Taber, 1998: 146, 147 (Ephebomyrmex, Pogonomyrmex species key); Fernández & Palacio, 1998: 1657 (northern South America species key); Mackay & Mackay, 2002: 180 (U.S.A., New Mexico species key); Lattke, 2006: 56 (northern South America species key); Johnson, R.A. & Overson, 2009: 313 (P. californicus group key); Cuezzo & Claver, 2009: 99 (Argentina species key).

Myrmicinae Tetramoriini Anergates TRUE TRUE FALSE

1 species

Anergates Forel, 1874: 67 (see also p. 93). Type-species: Myrmica atratula, by monotypy.

Taxonomic history

Anergates in Myrmicinae: Emery, 1877a: 81 [Myrmicidae]; Emery & Forel, 1879a: 457 [Myrmicidae]; Dalla Torre, 1893: 64.

Anergates in Myrmicinae, Formicoxenini: Forel, 1893a: 165.

Anergates in Myrmicinae, Myrmicini: Emery, 1895j: 769; Wheeler, W.M. 1910g: 139.

Anergates in Myrmicinae, Solenopsidini: Emery, 1914a: 41 [subtribe Anergatini]; Forel, 1917: 243; Emery, 1922e: 205; Wheeler, W.M. 1922a: 663; all subsequent authors to the following; Wheeler, G.C. & Wheeler, J. 1985: 257 (anachronism).

Anergates incertae sedis in Myrmicinae: Ettershank, 1966: 81.

Anergates in Myrmicinae, Tetramoriini: Bolton, 1976: 296; Smith, D.R. 1979: 1401; Dlussky & Fedoseeva, 1988: 80; Bolton, 1994: 106; Sanetra & Buschinger, 2000: 108.

Genus Anergates references

André, 1882c: 278 (Europe & Algeria); Dalla Torre, 1893: 64 (catalogue); Emery, 1922e: 205 (diagnosis, catalogue); Creighton, 1950a: 241 (North America, review); Ettershank, 1966: 157 (diagnosis, review of genus, checklist); Bernard, 1967: 241 (diagnosis); Bolton, 1976: 296 (diagnosis, review of genus); Smith, D.R. 1979: 1401 (North America catalogue); Bolton, 1995a: 1047 (census); Bolton, 1995b: 63 (catalogue); Sanetra & Buschinger, 2000: 95 (phylogeny).

Myrmicinae Tetramoriini Tetramorium TRUE TRUE FALSE

Extant: 461 species, 16 subspecies

Fossil:

Tetramorium Mayr, 1855: 423. Type-species: Formica caespitum, by subsequent designation of Girard, 1879: 1016.

Taxonomic history

Tetramorium in Myrmicidae, Myrmicidae: Emery, 1877a: 81.

Tetramorium in Myrmicinae: Mayr, 1855: 423 [Myrmicidae]; Mayr, 1861: 61 [Myrmicidae]; Mayr, 1865: 20 [Myrmicidae]; Emery & Forel, 1879a: 457 [Myrmicidae]; Dalla Torre, 1893: 130.

Tetramorium in Myrmicinae, Myrmicini: Forel, 1895b: 125; Forel, 1899c: 52; Kusnezov, 1964: 57 (anachronism).

Tetramorium in Myrmicinae, Tetramoriini: Emery, 1895j: 770; Ashmead, 1905b: 383; Wheeler, W.M. 1910g: 141; Emery, 1914a: 42; Arnold, 1917: 271; Forel, 1917: 245; Wheeler, W.M. 1922a: 664; Emery, 1924d: 275; all subsequent authors.

Junior synonyms of TETRAMORIUM

Tetrogmus Roger, 1857: 10. Type-species: Tetrogmus caldarius, by monotypy.

Taxonomic history

Tetrogmus as subgenus of Tetramorium: Wheeler, W.M. 1910g: 141.

Tetrogmus as junior synonym of Tetramorium: Roger, 1862c: 297; all subsequent authors except the above; Bolton, 1976: 359; Bolton, 1980: 195.

Xiphomyrmex Forel, 1887: 385 [as subgenus of Tetramorium]. Type-species: Tetramorium (Xiphomyrmex) kelleri, by subsequent designation of Wheeler, W.M. 1911f: 175.

Taxonomic history

Xiphomyrmex in Myrmicinae: Dalla Torre, 1893: 130.

Xiphomyrmex in Myrmicinae, Myrmicini: Ashmead, 1905b: 383.

Xiphomyrmex in Myrmicinae, Tetramoriini: Emery, 1895j: 770; Emery, 1914a: 42; Forel, 1917: 245; Wheeler, W.M. 1922a: 664; Emery, 1924d: 286; all subsequent authors.

Xiphomyrmex as subgenus of Tetramorium: Dalla Torre, 1893: 130; Forel, 1899c: 53; Forel, 1903a: 700; Wheeler, W.M. 1910g: 141; Arnold, 1917: 274; Arnold, 1926: 277.

Xiphomyrmex as genus: Emery, 1895j: 770; Emery, 1896e: 183; Emery, 1914a: 42; Forel, 1917: 245; Wheeler, W.M. 1922a: 193; Emery, 1924d: 286; Smith, M.R. 1938a: 126; Donisthorpe, 1943g: 736; Creighton, 1950a: 293; Smith, M.R. 1951a: 824; Chapman & Capco, 1951: 179; Bernard, 1953b: 250; Wheeler, G.C. & Wheeler, J. 1985: 257 (anachronism); Dlussky & Fedoseeva, 1988: 80 (anachronism).

Xiphomyrmex as junior synonym of Tetramorium: Bingham, 1903: 175; Bolton, 1976: 359; Bolton, 1980: 195; Bolton, 1994: 106.

Triglyphothrix Forel, 1890b: cvi. Type-species: Triglyphothrix walshi, by monotypy.

Taxonomic history

Triglyphothrix in Myrmicinae: Dalla Torre, 1893: 135.

Triglyphothrix in Myrmicinae, Tetramoriini: Emery, 1895j: 770; Ashmead, 1905b: 383; Wheeler, W.M. 1910g: 141; Emery, 1914a: 42; Forel, 1917: 245; Wheeler, W.M. 1922a: 664; Emery, 1924d: 272; all subsequent authors.

Triglyphothrix as subgenus of Tetramorium: Arnold, 1917: 274; Arnold, 1926: 272.

Triglyphothrix as genus: Forel, 1890b: cvi. Dalla Torre, 1893: 135; Emery, 1895j: 770; Forel, 1903a: 703; Bingham, 1903: 171; Wheeler, W.M. 1910g: 141; Emery, 1914a: 42; Forel, 1917: 245; Wheeler, W.M. 1922a: 196; Emery, 1924d: 272; Donisthorpe, 1943g: 734; Creighton, 1950a: 285; Smith, M.R. 1951a: 823; Chapman & Capco, 1951: 178; Bernard, 1953b: 248; Kempf, 1972a: 255; Bolton, 1976: 310; Smith, D.R. 1979: 1399; Wheeler, G.C. & Wheeler, J. 1985: 257; Taylor & Brown, D.R. 1985: 91.

Triglyphothrix as junior synonym of Tetramorium: Bolton, 1985: 247; Bolton, 1994: 106.

Atopula Emery, 1912b: 104. Type-species: Atopomyrmex nodifer, by original designation.

Taxonomic history

Atopula in Myrmicinae, Myrmecinini: Emery, 1912b: 105; Emery, 1914a: 41; Forel, 1917: 244; Wheeler, W.M. 1922a: 663; Emery, 1924d: 242 [subtribe Podomyrmini]; all subsequent authors to Bolton, 1976: 359.

Atopula in Myrmicinae, Podomyrmini: Dlussky & Fedoseeva, 1988: 79.

Atopula as subgenus of Vollenhovia: Emery, 1912d: 273.

Atopula as subgenus of Terataner: Arnold, 1952b: 129.

Atopula as genus: Emery, 1912b: 104; Emery, 1914a: 41; Wheeler, W.M. 1922a: 663; Emery, 1924d: 242; Bernard, 1948: 177; Wheeler, G.C. & Wheeler, J. 1985: 257 (anachronism); Dlussky & Fedoseeva, 1988: 79 (anachronism).

Atopula as junior synonym of Tetramorium: Bolton, 1976: 359; Bolton, 1980: 195; Bolton, 1994: 106.

Macromischoides Wheeler, W.M. 1920: 53. Type-species: Macromischa aculeata, by original designation.

Taxonomic history

Macromischoides in Myrmicinae, Leptothoracini: Wheeler, W.M. 1922a: 664; Wheeler, G.C. & Wheeler, J. 1985: 257 (anachronism).

Macromischoides in Myrmicinae, Tetramoriini: Santschi, 1924b: 207; all subsequent authors.

Macromischoides as genus: Wheeler, W.M. 1922a: 187; Santschi, 1924b: 206; Bernard, 1953b: 248; Dlussky & Fedoseeva, 1988: 80 (anachronism).

Macromischoides as junior synonym of Tetramorium: Bolton, 1976: 359; Bolton, 1980: 196.

[Macromichoides Santschi, 1924b: 206, incorrect subsequent spelling.]

Lobomyrmex Kratochvíl, 1941: 84 [as subgenus of Tetramorium]. Type-species: Tetramorium (Lobomyrmex) ferox silhavyi (junior synonym of Tetramorium ferox), by monotypy.

Taxonomic history

Lobomyrmex as junior synonym of Tetramorium: Brown, 1973b: 181 [provisional]; Bolton, 1976: 359; Bolton, 1980: 196.

Sulcomyrmex Kratochvíl, 1941: 84 [as subgenus of Tetramorium]. Unavailable name. Proposed without designation of type-species and therefore unavailable. Species included by Kratochvíl (1941) are all referable to Tetramorium: Bolton, 1976: 359.]

Apomyrmex Calilung, 2000: 66. Type-species: Apomyrmex manobo, by original designation.

Taxonomic history

Apomyrmex incertae sedis in Myrmicinae: Calilung, 2000: 66.

Apomyrmex as junior synonym of Tetramorium: Bolton, 2003: 227, 269.

Genus Tetramorium references

Roger, 1863b: 26, 27 (catalogue); Mayr, 1863: 456 (catalogue); Mayr, 1865: 20 (diagnosis); Mayr, 1870b: 972 (all species key); André, 1883a: 285 (Europe & Algeria species key); Cresson, 1887: 261 (U.S.A. catalogue); Emery, 1893e: 214 (Triglyphothrix species key); Dalla Torre, 1893: 130, 135 (Tetramorium, Triglyphothrix catalogues); Forel, 1903a: 700, 703 (India & Sri Lanka Tetramorium, Triglyphothrix species keys); Bingham, 1903: 172, 175 (India, Sri Lanka & Burma Triglyphothrix, Tetramorium species keys); Ruzsky, 1905b: 517 (Russian Empire species key); Emery, 1909d: 695 (Palaearctic species key); Emery, 1916b: 195 (Italy species key); Arnold, 1917: 271, 275, 334, 345 (diagnosis, South Africa Tetramorium (Tetramorium), T. (Triglyphothrix), T. (Xiphomyrmex) species keys); Bondroit, 1918: 106 (France & Belgium species key); Mann, 1919: 353 (Papuasia Triglyphothrix species key); Wheeler, W.M. 1922a: 187, 190, 193, 196 (Macromischoides, Tetramorium, Xiphomyrmex, Triglyphothrix diagnoses); Wheeler, W.M. 1922a: 886, 889, 893, 906, 909 (Afrotropical Atopula, Macromischoides, Tetramorium, Xiphomyrmex, Triglyphothrix catalogues); Wheeler, W.M. 1922a: 1030, 1031, 1032 (Malagasy Tetramorium, Xiphomyrmex, Triglyphothrix catalogues); Emery, 1924d: 242 (Atopula diagnosis, catalogue); Emery, 1924d: 272 (Triglyphothrix diagnosis, catalogue); Emery, 1924d: 275 (Tetramorium diagnosis, catalogue); Emery, 1924d: 286 (Xiphomyrmex diagnosis, catalogue); Santschi, 1924b: 210 (Macromischoides species key); Arnold, 1926: 241 (South Africa Tetramorium (Tetramorium) species key); Menozzi, 1933b: 74 (Israel species key); Finzi, 1936: 183 (Egypt species key); Smith, M.R. 1938a: 127 (North America Xiphomyrmex species key); Kratochvíl, 1941: 84 (Central Europe species key); Novák & Sadil, 1941: 84 (Central Europe species key); Smith, M.R. 1943b: 2 (North America species key); Kratochvíl, Novák & Snoflák, 1944: 76, 95 (Central Europe species key); Creighton, 1950a: 290, 293 (North America Tetramorium, Xiphomyrmex species key); Chapman & Capco, 1951: 174, 178, 179 (Asia Tetramorium, Triglyphothrix, Xiphomyrmex checklists); Brown, 1958h: 27 (New Zealand species); Bernard, 1967: 227 (diagnosis, Western Europe species key); Kempf, 1972a: 249, 255 (Neotropical Tetramorium, Triglyphothrix catalogues); Alayo, 1974: 18 (Cuba species key); Tarbinsky, 1976: 105 (Kyrghyzstan species key); Bolton, 1976: 310, 314, 341 (Triglyphothrix diagnosis, Afrotropical, Oriental & Malesian species revisions, keys); Bolton, 1976: 359 (Tetramorium diagnosis, review of genus, synonymy); Bolton, 1977: 72, 133 (Oriental & Malesian, Australia species revisions, keys); Kutter, 1977c: 150 (Switzerland species key); Arnol'di & Dlussky, 1978: 544 (former European U.S.S.R. species key); Collingwood, 1978: 86 (Iberian Peninsula species key); Collingwood, 1979: 82 (Fennoscandia & Denmark species key); Smith, D.R. 1979: 1399, 1400 (North America Triglyphothrix, Tetramorium catalogues); Bolton, 1979: 132, 159 (Malagasy, New World species revisions, keys); Bolton, 1980: 196, 205 (diagnosis, review of genus, Afrotropical species revision, key); Collingwood, 1985: 262 (Saudi Arabia species key); Gösswald, 1985: 311 (Germany species key); Taylor & Brown, D.R. 1985: 88 (Australia catalogue); Taylor, 1987a: 78 (Australia, New Caledonia & New Zealand checklist); Agosti & Collingwood, 1987b: 277 (Balkans species key); Wang, M., Xiao & Wu, 1988: 264 (China species key); Radchenko & Arakelian, 1990: 371 (Crimea & Caucasus T. ferox complex, key); Dlussky, Soyunov & Zabelin, 1990: 197 (Turkmenistan species key); Atanasov & Dlussky, 1992: 149 (Bulgaria species key); Radchenko, 1992a: 44 (former U.S.S.R. species key); Morisita, Kubota, et al. 1992: 33 (Japan species key); Arakelian, 1994: 60 (Armenia species key); Radchenko, 1994b: 109 (South Siberia species key); Bolton, 1994: 106 (synoptic classification); Bolton, 1995a: 1053 (census); Bolton, 1995b: 403 (catalogue); Wu, J. & Wang, 1995: 79 (China species key); Collingwood & Agosti, 1996: 332 (Saudi Arabia species key); Seifert, 1996b: 158 (Central Europe species key); Cagniant, 1997: 89 (Morocco species key); Radchenko, Czechowski & Czechowsa, 1998: 107 (Poland species key); Collingwood & Prince, 1998: 17 (Portugal species key); Sanetra, Güsten & Schulz, 1999: 317 (Italy, species); Shattuck, 1999: 168 (Australia synopsis); Chang & He, 2001a: 1 (northwest China species key); Zhou, 2001b: 100 (China, Guangxi species key); Czechowski, Radchenko & Czechowska, 2002: 145 (Poland species key); Mackay & Mackay, 2002: 214 (U.S.A., New Mexico species key); Seifert, 2007: 145 (North and Central Europe species key); Csösz, Radchenko & Schulz, 2007: 8 (T. chefketi complex key); Terayama, 2009: 162 (Taiwan species key); Heterick, 2009: 177 (south-western Australia species key); Csösz & Schulz, 2010: 12 (T. ferox complex key); Hita Garcia, Fischer & Peters, 2010b: 12 (Afrotropical T. weitzeckeri species group key).

Myrmicinae Pheidolini Aphaenogaster TRUE TRUE FALSE

Extant: 176 species, 39 subspecies

Fossil: 19 species

Aphaenogaster Mayr, 1853b: 107. Type-species: Aphaenogaster sardoa, by subsequent designation of Bingham, 1903: 270.

Taxonomic history

Aphaenogaster in Myrmicinae: Mayr, 1855: 466 [Myrmicidae]; Mayr, 1865: 19 [Myrmicidae]; Emery & Forel, 1879a: 461 [Myrmicidae]; Cresson, 1887: 260 [Myrmicidae]; Dalla Torre, 1893: 98.

Aphaenogaster in Myrmicinae, Myrmicini: Forel, 1891b: 166; Forel, 1895b: 129; Emery, 1895j: 769; Forel, 1899c: 58; Wheeler, W.M. 1910g: 140; Wheeler, W.M. 1915h: 53; Santschi, 1932b: 13; Kempf, 1972a: 22 (anachronism).

Aphaenogaster in Myrmicinae, Aphaenogastrini: Enzmann, J. 1947b: 147.

Aphaenogaster in Myrmicinae, Pheidolini: Emery, 1877a: 81 [Pheidolidae]; Emery, 1914a: 40; Forel, 1917: 241; Emery, 1921f: 55 [subtribe Stenammini]; Wheeler, W.M. 1922a: 661; Brown, 1949a: 48; Chapman & Capco, 1951: 131; all subsequent authors.

Aphaenogaster as junior synonym of Atta: Mayr, 1863: 395.

Aphaenogaster as subgenus of Stenamma: Emery, 1895c: 298; Emery, 1895j: 769; Forel, 1903a: 693.

Aphaenogaster as genus: Emery, 1908c: 309; all subsequent authors.

Junior synonyms of APHAENOGASTER

Deromyrma Forel, 1913b: 350 [as subgenus of Aphaenogaster]. Type-species: Aphaenogaster (Ischnomyrmex) swammerdami, by original designation.

Taxonomic history

[Deromyrma also described as new by Forel, 1913k: 49.]

Deromyrma as subgenus of Aphaenogaster: Forel, 1913b: 350; Forel, 1913k: 49; Forel, 1917: 241; Emery, 1921f: 64; Wheeler, W.M. 1922a: 680; all subsequent authors to the following.

Deromyrma as junior synonym of Aphaenogaster: Brown, 1973b: 180 [provisional]; Smith, D.R. 1979: 1359; Bolton, 1982: 364.

Planimyrma Viehmeyer, 1914d: 604 [as subgenus of Aphaenogaster]. Type-species: Stenamma (Ischnomyrmex) loriai, by original designation.

Taxonomic history

Planimyrma as subgenus of Aphaenogaster: Viehmeyer, 1914d: 604; Forel, 1917: 241; Emery, 1921f: 65; Wheeler, W.M. 1922a: 680; all subsequent authors to the following.

Planimyrma as junior synonym of Aphaenogaster: Brown, 1973b: 184 [provisional]; Smith, D.R. 1979: 1359; Bolton, 1982: 364.

Attomyrma Emery, 1915d: 70 [as subgenus of Aphaenogaster]. Type-species: Formica subterranea, by original designation.

Taxonomic history

Attomyrma as subgenus of Aphaenogaster: Emery, 1915d: 70; Emery, 1916b: 129; Emery, 1921f: 56; all subsequent authors to the following.

Attomyrma as junior synonym of Aphaenogaster: Brown, 1973b: 178 [provisional]; Smith, D.R. 1979: 1359; Bolton, 1982: 364.

Novomessor Emery, 1915d: 73. Type-species: Aphaenogaster (Ischnomyrmex) cockerelli, by original designation.

Taxonomic history

Novomessor in Myrmicinae, Pheidolini: Forel, 1917: 241; Emery, 1921f: 66 [subtribe Stenammini]; Wheeler, W.M. 1922a: 661; all subsequent authors except the following.

Novomessor in Myrmicinae, Myrmicini: Donisthorpe, 1943f: 675; Kempf, 1972a: 166.

Novomessor as genus: Wheeler, W.M. & Creighton, 1934: 343; Creighton, 1950a: 155; Hölldobler, Stanton & Engel, 1976: 32.

Novomessor as junior synonym of Aphaenogaster: Brown, 1974b: 47; Bolton, 1982: 364 (discussion pp. 339-341).

Nystalomyrma Wheeler, W.M. 1916j: 215 [as subgenus of Aphaenogaster]. Type-species: Myrmica longiceps, by original designation.

Taxonomic history

Nystalomyrma as subgenus of Aphaenogaster: Wheeler, W.M. 1916j: 215; Emery, 1921f: 61; Wheeler, W.M. 1922a: 680; Donisthorpe, 1943f: 676.

Nystalomyrma as junior synonym of Aphaenogaster: Brown, 1973b: 183 [provisional]; Smith, D.R. 1979: 1359; Bolton, 1982: 364.

Brunella Forel, 1917: 234. Type-species: Aphaenogaster belti, by monotypy.

Taxonomic history

Brunella in Myrmicinae, Leptothoracini: Forel, 1917: 245.

Brunella in Myrmicinae, Myrmecinini: Wheeler, W.M. 1922a: 663; Emery, 1924d: 242; Donisthorpe, 1943f: 629.

Brunella incertae sedis in Formicidae: Wheeler, G.C. & Wheeler, J. 1985: 259 (anachronism, incomprehensible entry).

Brunella incertae sedis in Myrmicinae: Dlussky & Fedoseeva, 1988: 80 (anachronism).

Brunella as junior synonym of Atopula: Emery, 1924d: 242; Donisthorpe, 1943f: 629.

Brunella as junior synonym of Aphaenogaster: Bolton, 1982: 364; Bolton, 1994: 106.

†Sinaphaenogaster Zhang, 1989: 266 [as subgenus of Aphaenogaster]. Type-species: †Paraphaenogaster shanwangensis, by original designation.

Taxonomic history

†Sinaphaenogaster as junior synonym of Aphaenogaster: Bolton, 2003: 230, 273.

Genus Aphaenogaster references

Mayr, 1855: 466 (diagnosis); Roger, 1863b: 29 (catalogue); Mayr, 1865: 19 (diagnosis); Mayr, 1867a: 92 (diagnosis); André, 1874: 195 (Europe species key); Forel, 1874: 74 (Switzerland species key); André, 1883b: 348 (Europe & Algeria species key); Mayr, 1886d: 443 (U.S.A. species key); Cresson, 1887: 260 (U.S.A. catalogue); Emery, 1888a: 531 (A. (Ischnomyrmex) species key); Nasonov, 1889: 74 (Russia species key); Dalla Torre, 1893: 98 (catalogue); Forel, 1903a: 693 (India & Sri Lanka species key); Bingham, 1903: 270 (India, Sri Lanka & Burma species key); Ruzsky, 1905b: 714 (Russian Empire species key); Emery, 1908c: 310 (Palaearctic species key); Bondroit, 1910: 495 (Belgium species key); Emery, 1916b: 132 (Italy species key); Wheeler, W.M. 1916m: 585 (U.S.A., Connecticut species key); Wheeler, W.M. 1916j: 213 (Australia species); Bondroit, 1918: 156 (France & Belgium species key); Emery, 1921f: 55 (diagnosis, subgenera key, catalogue); Emery, 1921f: 56, 61, 64, 65 (A. (Attomyrma, Nystalomyrma, Deromyrma, Planimyrma) diagnoses, catalogues); Emery, 1921f: 66 (Novomessor diagnosis, catalogue); Wheeler, W.M. 1922a: 680 (subgenera key); Wheeler, W.M. 1922a: 1016, 1029 (Aphaenogaster, Brunella catalogues); Wheeler, W.M. & Creighton, 1934: 348 (Novomessor species key); Menozzi, 1939a: 296 (Himalaya & Tibet species key); Stitz, 1939: 112 (Germany species key); Kratochvíl, 1941: 81 (Central Europe species key); Novák & Sadil, 1941: 81 (Central Europe species key); Cole, 1942: 363 (U.S.A., Utah species key); Buren, 1944a: 284 (U.S.A., Iowa species key); Creighton, 1950a: 140, 155 (North America Aphaenogaster, Novomessor species keys); Chapman & Capco, 1951: 131 (Asia checklist); Smith, M.R. 1961b: 218 (New Guinea species key); Gregg, 1963: 336 (U.S.A., Colorado species key); Bernard, 1967: 128 (diagnosis, Western Europe species key); Kempf, 1972a: 22, 166 (Neotropical Aphaenogaster (Deromyrma), Novomessor catalogues); Arnol'di, 1976b: 1023 (former U.S.S.R. species key); Kutter, 1977c: 77 (Switzerland species key); Arnol'di & Dlussky, 1978: 536 (former European U.S.S.R. species key); Collingwood, 1978: 79 (Iberian Peninsula species key); Smith, D.R. 1979: 1359 (North America catalogue); Allred, 1982: 439 (U.S.A., Utah species key); Taylor & Brown, D.R. 1985: 54 (Australia catalogue); Gösswald, 1985: 298 (Germany species key); Wheeler, G.C. & Wheeler, J. 1986g: 35 (U.S.A., Nevada species key); Taylor, 1987a: 8 (Australia checklist); Agosti & Collingwood, 1987b: 269 (Balkans species key); Dlussky & Fedoseeva, 1988: 79 (synoptic classification); Dlussky, Soyunov & Zabelin, 1990: 211 (Turkmenistan species key); Brandão, 1991: 326 (Neotropical catalogue); Morisita, Kubota, et al. 1992: 15 (Japan species key); Atanasov & Dlussky, 1992: 107 (Bulgaria species key); Schulz, A. 1994a: 425 (A. (Attomyrma), partial key); Arakelian, 1994: 29 (Armenia species key); Bolton, 1994: 106 (synoptic classification); Bolton, 1995a: 1048 (census); Bolton, 1995b: 68 (catalogue); Wu, J. & Wang, 1995: 110 (China species key); Umphrey, 1996: 557 (North America A. fulva complex key); Cagniant, 1996a: 67 (Morocco species key); Seifert, 1996b: 156 (Central Europe species key); Collingwood & Prince, 1998: 13 (Portugal species key); Shattuck, 1999: 126 (Australia synopsis); Park & Kim, 2000: 108 (Korea species key); Zhou, 2001b: 142 (China, Guangxi species key); Mackay & Mackay, 2002: 60 (U.S.A., New Mexico species key); Longino & Cover, 2004: 1 (A. phalangium complex revision); Radchenko, 2005b: 192 (North Korea species key); Seifert, 2007: 144 (North and Central Europe species key); Shattuck, 2008a: 27 (Australia species revision, key); Terayama, 2009: 164 (Taiwan species key); Heterick, 2009: 144 (south-western Australia species key).

Myrmicinae Pheidolini Messor TRUE TRUE FALSE

Extant: 112 species, 49 subspecies

Fossil:

Messor Forel, 1890a: lxviii [as subgenus of Aphaenogaster]. Type-species: Formica barbara, by subsequent designation of Bingham, 1903: 277.

Taxonomic history

Messor in Myrmicinae: Dalla Torre, 1893: 98.

Messor in Myrmicinae, Myrmicini: Emery, 1895j: 769; Forel, 1899c: 59; Ashmead, 1905b: 383; Wheeler, W.M. 1910g: 140.

Messor in Myrmicinae, Pheidolini: Emery, 1914a: 40; Forel, 1917: 241; Arnold, 1920a: 404; Emery, 1921f: 68 [subtribe Stenammini]; Wheeler, W.M. 1922a: 661; all subsequent authors.

Messor as subgenus of Aphaenogaster: Forel, 1890a: lxviii; Dalla Torre, 1893: 98; Forel, 1899c: 59.

Messor as subgenus of Stenamma: Emery, 1895c: 298; Emery, 1895j: 769; Forel, 1903a: 693.

Messor as genus: Bingham, 1903: 277; Ruzsky, 1905b: 726; Emery, 1908e: 437; Wheeler, W.M. 1910g: 140; Forel, 1917: 241; Bondroit, 1918: 149; Arnold, 1920a: 404; Emery, 1921f: 68; Wheeler, W.M. 1922a: 661; all subsequent authors.

Junior synonyms of MESSOR

Cratomyrmex Emery, 1892d: 572. Type-species: Cratomyrmex regalis, by monotypy.

Taxonomic history

Cratomyrmex in Myrmicinae, Myrmicini: Emery, 1895j: 769; Ashmead, 1905b: 383; Wheeler, W.M. 1910g: 140; Emery, 1914a: 40; Forel, 1917: 240; Emery, 1921f: 43; Wheeler, W.M. 1922a: 660; Donisthorpe, 1943f: 635.

Cratomyrmex as subgenus of Messor: Santschi, 1920d: 378.

Cratomyrmex as genus: Emery, 1892d: 572; Wheeler, W.M. 1910g: 140; Forel, 1917: 240; Emery, 1921f: 43; Wheeler, W.M. 1922a: 660, 802; Bernard, 1971: 6.

Cratomyrmex as junior synonym of Messor: Emery, 1924d: 357; Donisthorpe, 1943f: 635; Bolton, 1982: 338; Hölldobler & Wilson, 1990: 14; Bolton, 1994: 106.

Veromessor Forel, 1917: 235 [as subgenus of Novomessor]. Type-species: Aphaenogaster andrei, by subsequent designation of Emery, 1921f: 67.

Taxonomic history

Veromessor in Myrmicinae, Pheidolini: Wheeler, W.M. 1922a: 661; all subsequent authors.

Veromessor as subgenus of Novomessor: Forel, 1917: 235; Emery, 1921f: 67; Donisthorpe, 1943g: 735.

Veromessor as genus: Wheeler, W.M. 1922a: 661; Wheeler, W.M. & Creighton, 1934: 354; Creighton, 1950a: 157; Smith, M.R. 1951a: 799; Kusnezov, 1952b: 10 (in key); Kempf, 1972a: 257; Smith, D.R. 1979: 1364.

Veromessor as junior synonym of Messor: Bolton, 1982: 338.

Lobognathus Enzmann, J. 1947b: 152 [as subgenus of Veromessor]. Erroneous entry for Veromessor lobognathus and hence junior synonym of Veromessor: Brown, 1949a: 49.

[Sphaeromessor Bernard, 1985: 48. Unavailable name. Proposed without designation of type-species and therefore unavailable. Species included by Bernard (1985) are all referable to Messor: Bolton, 1995b: 46.]

Genus Messor references

Forel, 1903a: 693 (India & Sri Lanka species key); Bingham, 1903: 278 (India, Sri Lanka & Burma species key); Ruzsky, 1905b: 726 (Russian Empire species key); Emery, 1908e: 437 (Palaearctic species key); Emery, 1916b: 143 (Italy species key); Bondroit, 1918: 149 (France & Belgium species key); Arnold, 1920a: 404 (diagnosis); Emery, 1921f: 43 (Cratomyrmex diagnosis, catalogue); Emery, 1921f: 67 (Veromessor diagnosis, catalogue); Emery, 1921f: 68 (diagnosis, catalogue); Wheeler, W.M. 1922a: 802 (Afrotropical Cratomyrmex, Messor catalogues); Santschi, 1927c: 247 (M. instabilis group key); Kuznetsov-Ugamsky, 1927a: 89 (Central Asia species key); Kuznetsov-Ugamsky, 1929d: 4 (Central Asia species key); Finzi, 1929: 77 (Italy species key); Menozzi, 1933b: 56 (Israel species key); Wheeler, W.M. & Creighton, 1934: 360 (Veromessor species key); Finzi, 1936: 160 (Egypt species key); Novák & Sadil, 1941: 82 (Central Europe species key); Creighton, 1950a: 158 (North America Veromesor species key); Chapman & Capco, 1951: 136 (Asia checklist); Bernard, 1955a: 360 (Mediterranean M. structor group key); Smith, M.R. 1956a: 36 (U.S.A. Veromessor species key); Bernard, 1967: 137 (diagnosis, Western Europe species key); Kempf, 1972a: 257 (Neotropical Veromessor catalogue); Tarbinsky, 1976: 50 (Kyrghyzstan species key); Arnol'di, 1977b: 1640 (former U.S.S.R. species key); Arnol'di & Dlussky, 1978: 537 (former European U.S.S.R. species key); Collingwood, 1978: 80 (Iberian Peninsula species key); Smith, D.R. 1979: 1364 (North America Veromessor catalogue); Bolton, 1982: 338 (diagnosis, review of genus, Afrotropical species revision, key); Collingwood, 1985: 248 (Saudi Arabia species key); Wheeler, G.C. & Wheeler, J. 1986g: 38 (U.S.A., Nevada Veromessor species key); Agosti & Collingwood, 1987b: 270 (Balkans species key); Dlussky, Soyunov & Zabelin, 1990: 215 (Turkmenistan species key); Casevitz-Weulersse, 1990b: 156 (Corsica species key); Atanasov & Dlussky, 1992: 113 (Bulgaria species key); Arakelian, 1994: 33 (Armenia species key); Bolton, 1994: 106 (synoptic classification); Bolton, 1995a: 1050 (census); Bolton, 1995b: 252 (catalogue); Collingwood & Agosti, 1996: 314 (Saudi Arabia species key); Cagniant & Espadaler, 1998: 419 (Morocco species key); Collingwood & Prince, 1998: 14 (Portugal species key).

Myrmicinae Pheidolini Pheidole TRUE TRUE FALSE

Extant: 970 species, 142 subspecies

Fossil: 4 species

Pheidole Westwood, 1839: 219. Type-species: Atta providens, by monotypy.

Taxonomic history

Pheidole in Poneridae, Attidae: Smith, F. 1858b: 172.

Pheidole in Attidae: Smith, F. 1860a: 74; Smith, F. 1862a: 49.

Pheidole in Myrmicidae: Smith, F. 1871a: 331; Cresson, 1887: 262.

Pheidole in Myrmicidae, Pheidolidae: Emery, 1877a: 81.

Pheidole in Myrmicinae: Mayr, 1861: 69 [Myrmicidae]; Mayr, 1865: 22 [Myrmicidae]; Emery & Forel, 1879a: 463 [Myrmicidae]; Dalla Torre, 1893: 88.

Pheidole in Myrmicinae, Myrmicini: Forel, 1895b: 127; Emery, 1895j: 769; Forel, 1899c: 64; Ashmead, 1905b: 383; Wheeler, W.M. 1910g: 139; Kempf, 1972a: 183 (anachronism).

Pheidole in Myrmicinae, Pheidolini: Forel, 1893a: 165; Emery, 1914a: 40; Forel, 1917: 241; Arnold, 1920a: 414; Emery, 1921f: 77; Wheeler, W.M. 1922a: 661; all subsequent authors except Kempf, 1972a: 183, above.

Junior synonyms of PHEIDOLE

Oecophthora Heer, 1852: 15. Type-species: Oecophthora pusilla (junior synonym of Formica megacephala), by monotypy.

Taxonomic history

Oecophthora in Myrmicinae: Mayr, 1855: 453 [Myrmicidae].

Oecophthora as junior synonym of Pheidole: Smith, F. 1858a: 282; Smith, F. 1858b: 172; Roger, 1863b: 30.

Ischnomyrmex Mayr, 1862: 738. Type-species: Myrmica longipes, by monotypy.

Taxonomic history

Ischnomyrmex in Myrmicinae: Mayr, 1865: 18 [Myrmicidae]; Dalla Torre, 1893: 98.

Ischnomyrmex in Myrmicinae, Myrmicini: Emery, 1895j: 769; Ashmead, 1905b: 383; Wheeler, W.M. 1910g: 140.

Ischnomyrmex in Myrmicinae, Pheidolini: Emery, 1914a: 40; Forel, 1917: 241; Emery, 1922e: 113; Wheeler, W.M. 1922a: 661; all subsequent authors.

Ischnomyrmex as junior synonym of Aphaenogaster: Emery & Forel, 1879a: 461; Bingham, 1903: 270.

Ischnomyrmex as subgenus of Aphaenogaster: Forel, 1891b: 166; Dalla Torre, 1893: 98; Forel, 1899c: 59; Emery, 1908c: 313; Wheeler, W.M. 1910g: 140.

Ischnomyrmex as subgenus of Stenamma: Emery, 1895c: 298; Emery, 1895j: 769; Forel, 1902h: 439; Forel, 1903a: 693.

Ischnomyrmex as subgenus of Pheidole: Forel, 1913k: 49.

Ischnomyrmex as genus: Mayr, 1862: 738; Emery, 1914a: 40; Viehmeyer, 1914d: 601; Forel, 1917: 241; Wheeler, W.M. 1922a: 661; Emery, 1922e: 113. Stitz, 1911a: 367; Kusnezov, 1952b: 10 (in key).

Ischnomyrmex as junior synonym of Pheidole: Brown, 1973b: 181 [provisional]; Smith, D.R. 1979: 1365; Snelling, R.R. 1981: 395.

Leptomyrma Motschoulsky, 1863: 17. Type-species: Leptomyrma gracilipes, by monotypy.

Taxonomic history

Leptomyrma in Myrmicidae: Smith, F. 1871a: 329.

Leptomyrma as junior synonym of Pheidole: Emery, 1892b: 166; all subsequent authors.

Pheidolacanthinus Smith, F. 1865: 75. Type-species: Pheidolacanthinus armatus (junior synonym of Myrmica quadrispinosa), by monotypy.

Taxonomic history

Pheidolacanthinus in Myrmicidae: Smith, F. 1871a: 333.

Pheidolacanthinus in Myrmicinae: Dalla Torre, 1893: 98.

Pheidolacanthinus as subgenus of Pheidole: Emery, 1915i: 190; Forel, 1917: 241; Emery, 1921f: 81; Wheeler, W.M. 1922a: 673; all subsequent authors to the following.

Pheidolacanthinus as junior synonym of Pheidole: Brown, 1973b: 183 [provisional]; Smith, D.R. 1979: 1365; Snelling, R.R. 1981: 395.

Ceratopheidole Pergande, 1896: 889 [as subgenus of Pheidole]. Type-species: Pheidole (Ceratopheidole) granulata, by monotypy.

Taxonomic history

Ceratopheidole in Myrmicinae, Pheidolini: Emery, 1914a: 40; Forel, 1917: 241; Emery, 1922e: 112; Wheeler, W.M. 1922a: 661; all subsequent authors.

Ceratopheidole as genus: Emery, 1914a: 40; Forel, 1917: 241; Emery, 1922e: 112; Wheeler, W.M. 1922a: 661; Chapman & Capco, 1951: 135; Wu, J. & Wang, 1995: 104 (anachronism).

Ceratopheidole as subgenus of Pheidole: Pergande, 1896: 889; Kempf, 1972a: 183.

Ceratopheidole as junior synonym of Pheidole: Brown, 1973b: 179 [provisional]; Smith, D.R. 1979: 1365; Snelling, R.R. 1981: 395; Wilson, 2003: 6.

Epipheidole Wheeler, W.M. 1903f: 664. Type-species: Epipheidole inquilina, by monotypy.

Taxonomic history

[Epipheidole also described as new by Wheeler, W.M. 1904a: 14.]

Epipheidole in Myrmicinae, Stenammini: Ashmead, 1905b: 383.

Epipheidole in Myrmicinae, Myrmicini: Wheeler, W.M. 1910g: 140.

Epipheidole in Myrmicinae, Pheidolini: Emery, 1914a: 40; Forel, 1917: 241; Emery, 1922e: 114; Wheeler, W.M. 1922a: 661; Creighton, 1950a: 192.

Epipheidole as junior synonym of Pheidole: Cole, 1965: 174; Smith, D.R. 1979: 1366.

Phidole Bingham, 1903: 220, unjustified emendation of Pheidole.

Taxonomic history

Phidole as junior synonym of Pheidole: Wheeler, 1922a: 806.

Sympheidole Wheeler, W.M. 1904a: 7. Type-species: Sympheidole elecebra, by monotypy.

Taxonomic history

Sympheidole in Myrmicinae, Myrmicini: Wheeler, W.M. 1910g: 140.

Sympheidole in Myrmicinae, Stenammini: Ashmead, 1905b: 383.

Sympheidole in Myrmicinae, Pheidolini: Emery, 1914a: 40; Forel, 1917: 241; Emery, 1922e: 115; Wheeler, W.M. 1922a: 661; all subsequent authors to the following.

Sympheidole as junior synonym of Pheidole: Brown, 1973b: 185 [provisional]; Smith, D.R. 1979: 1365; Snelling, R.R. 1981: 395; Wilson, 2003: 6.

Phidola Schulz, W.A. 1906: 155, unjustified emendation of Pheidole.

Taxonomic history

Phidola as junior synonym of Pheidole: Smith, D.R. 1979: 1365.

Allopheidole Forel, 1912f: 237 [as subgenus of Pheidole]. Type-species: Pheidole kingi (junior synonym of Pheidole tepicana), by subsequent designation of Wheeler, W.M. 1913a: 79.

Taxonomic history

Allopheidole as subgenus of Pheidole: Forel, 1912f: 237; Forel, 1917: 241.

Allopheidole as junior synonym of Pheidole: Emery, 1915i: 190; Emery, 1921f: 84; Smith, D.R. 1979: 1366.

Decapheidole Forel, 1912f: 237 [as subgenus of Pheidole]. Type-species: Pheidole perpusilla, by subsequent designation of Wheeler, W.M. 1913a: 80.

Taxonomic history

Decapheidole in Myrmicinae, Pheidolini: Wheeler, W.M. 1922a: 661.

Decapheidole as genus: Wheeler, W.M. 1922a: 661; Borgmeier, 1927c: 78.

Decapheidole as subgenus of Pheidole: Forel, 1912f: 237, Forel, 1917: 241; Emery, 1922e: 112; Kempf, 1972a: 183.

Decapheidole as junior synonym of Pheidole: Brown, 1973b: 180 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395; Wilson, 2003: 6.

Isopheidole Forel, 1912m: 765 [as subgenus of Pheidole]. Type-species: Myrmica longipes, by monotypy.

Taxonomic history

Isopheidole as junior synonym of Ischnomyrmex: Forel, 1913b: 350 (in text); Forel, 1913k: 49; Emery, 1922e: 113; Wheeler, W.M. 1922a: 680 (in key). [Ischnomyrmex and Isopheidole share the same type-species, synonymy is therefore absolute.]

Elasmopheidole Forel, 1913k: 43 [as subgenus of Pheidole]. Type-species: Pheidole aberrans, by subsequent designation of Emery, 1915i: 190.

Taxonomic history

Elasmopheidole as subgenus of Pheidole: Forel, 1913k: 43; Emery, 1915i: 190; Forel, 1917: 241; Emery, 1922e: 110; Wheeler, W.M. 1922a: 673; all subsequent authors to the following.

Elasmopheidole as junior synonym of Pheidole: Brown, 1973b: 180 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395.

Cardiopheidole Wheeler, W.M. 1914b: 48 [as subgenus of Pheidole]. Type-species: Pheidole vaslitii, by original designation.

Taxonomic history

Cardiopheidole as subgenus of Pheidole: Wheeler, W.M. 1914b: 48; Forel, 1917: 241.

Cardiopheidole as junior synonym of Pheidole: Emery, 1915i: 190; Emery, 1921f: 84; Smith, D.R. 1979: 1366.

Anergatides Wasmann, 1915b: 281. Type-species: Anergatides kohli (junior secondary homonym in Pheidole, replaced by Pheidole neokohli), by monotypy.

Taxonomic history

Anergatides in Myrmicinae, Pheidolini: Forel, 1917: 243 [subtribe Anergatini]; Emery, 1922e: 116 [subtribe Anergatidini]; Donisthorpe, 1943f: 623.

Anergatides in Myrmicinae, Solenopsidini: Wheeler, W.M. 1922a: 663; Wheeler, G.C. & Wheeler, J. 1985: 257 (anachronism).

Anergatides as junior synonym of Pheidole: Brown, 1973b: 178 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395; Wilson, 1984: 328; Bolton, 1994: 106.

Parapheidole Emery, 1915d: 68. Type-species: Aphaenogaster oculata, by monotypy.

Taxonomic history

Parapheidole in Myrmicinae, Pheidolini: Forel, 1917: 241; Emery, 1922e: 115; Wheeler, W.M. 1922a: 661; all subsequent authors.

Parapheidole as junior synonym of Pheidole: Brown, 1973b: 183 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395.

Macropheidole Emery, 1915i: 190 [as subgenus of Pheidole]. Type-species: Pheidole fimbriata, by monotypy.

Taxonomic history

Macropheidole as subgenus of Pheidole: Emery, 1915i: 190; Forel, 1917: 241; Emery, 1921f: 81; Wheeler, W.M. 1922a: 673; all subsequent authors to the following.

Macropheidole as junior synonym of Pheidole: Smith, M.R. 1951a: 800; Brown, 1973b: 181 [provisional]; Smith, D.R. 1979: 1366.

Scrobopheidole Emery, 1915i: 190 [as subgenus of Pheidole]. Type-species: Pheidole scrobifera, by monotypy.

Taxonomic history

Scrobopheidole as subgenus of Pheidole: Forel, 1917: 241; Emery, 1922e: 112; Wheeler, W.M. 1922a: 673; all subsequent authors to the following.

Scrobopheidole as junior synonym of Pheidole: Brown, 1973b: 184 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395; Wilson, 2003: 6.

Stegopheidole Emery, 1915i: 190 [as subgenus of Pheidole]. Type-species: Pheidole (Elasmopheidole) upeneci, by monotypy.

Taxonomic history

Stegopheidole as subgenus of Pheidole: Emery, 1921f: 83; Wheeler, W.M. 1922a: 673; all subsequent authors to the following.

Stegopheidole as junior synonym of Pheidole: Brown, 1973b: 185 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395.

Trachypheidole Emery, 1915i: 190 [as subgenus of Pheidole]. Type-species: Pheidole bicornis, by original designation.

Taxonomic history

Trachypheidole as subgenus of Pheidole: Forel, 1917: 241; Emery, 1922e: 111; Wheeler, W.M. 1922a: 673; all subsequent authors to the following.

Trachypheidole as junior synonym of Pheidole: Brown, 1973b: 185 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395; Wilson, 2003: 6.

Electropheidole Mann, 1921: 438 [as subgenus of Pheidole]. Type-species: Pheidole (Electropheidole) roosevelti, by subsequent designation of Donisthorpe, 1943f: 642.

Taxonomic history

Electropheidole as subgenus of Pheidole: Mann, 1921: 438; Wheeler, W.M. 1922a: 672; all subsequent authors to the following.

Electropheidole as junior synonym of Pheidole: Brown, 1973b: 180 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395.

Bruchomyrma Santschi, 1922d: 248. Type-species: Bruchomyrma acutidens, by monotypy.

Taxonomic history

Bruchomyrma in Myrmicinae, Pheidolini: Donisthorpe, 1943f: 629.

Bruchomyrma in Myrmicinae, Myrmicini: Kempf, 1972a: 42.

Bruchomyrma as junior synonym of Pheidole: Brown, 1973b: 179 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395; Wilson, 1984: 327; Bolton, 1994: 106; Wilson, 2003: 6.

Bruchomyrma as genus: Dlussky & Fedoseeva, 1988: 80 (anachronism).

Cephalomorium Forel, 1922: 91 [as subgenus of Tetramorium]. Type-species: Tetramorium (Cephalomorium) bahai, by monotypy.

Taxonomic history

Cephalomorium as junior synonym of Hendecapheidole: Santschi, 1925d: 228.

Hendecapheidole Wheeler, W.M. 1922d: 3 [as subgenus of Pheidole]. Type-species: Pheidole tachigaliae, by original designation.

Taxonomic history

Hendecapheidole as junior synonym of Pheidole: Brown, 1973b: 181 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395; Wilson, 2003: 6.

Gallardomyrma Bruch, 1932: 271. Type-species: Gallardomyrma argentina, by original designation.

Taxonomic history

Gallardomyrma in Myrmicinae, Pheidolini: Donisthorpe, 1943f: 646.

Gallardomyrma in Myrmicinae, Myrmicini: Kempf, 1972a: 110.

Gallardomyrma as junior synonym of Pheidole: Brown, 1973b: 180 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395; Wilson, 1984: 327; Wilson, 2003: 6.

Gallardomyrma as genus: Kempf, 1972a: 110; Dlussky & Fedoseeva, 1988: 80 (anachronism).

Conothoracoides Strand, 1935: 176.

Taxonomic history

[Replacement name for Conothorax Karavaiev; junior homonym of Conothorax Jekel, 1854: 9 bis (foldout pages) (Coleoptera).]

Conothoracoides as junior synonym of Pheidole: Brown, 1973b: 179 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395.

Homonym replaced by Conothoracoides

Conothorax Karavaiev, 1935a: 75. Type-species: Conothorax bilobum, by monotypy.

Taxonomic history

[Conothorax Karavaiev junior homonym of Conothorax Jekel, 1854: 9 bis (foldout pages) (Coleoptera).]

Conothorax in Myrmicinae, Pheidolini: Karavaiev, 1935a: 75.

Eriopheidole Kusnezov, 1952a: 10. Type-species: Eriopheidole symbiotica (unresolved junior secondary homonym in Pheidole), by monotypy.

Taxonomic history

Eriopheidole in Myrmicinae, Pheidolini: Kusnezov, 1952a: 10.

Eriopheidole in Myrmicinae, Myrmicini: Kempf, 1972a: 107.

Eriopheidole junior synonym of Pheidole: Brown, 1973b: 180 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395; Bolton, 1994: 106.

Eriopheidole as genus: Kempf, 1972a: 107; Dlussky & Fedoseeva, 1988: 80 (anachronism).

Xenoaphaenogaster Baroni Urbani, 1964b: 50. Type-species: Xenoaphaenogaster inquilina (junior synonym of Myrmica pallidula), by original designation.

Taxonomic history

Xenoaphaenogaster as junior synonym of Monomorium: Brown, 1973b: 185 [provisional].

Xenoaphaenogaster as junior synonym of Pheidole: Bolton, 1987: 291.

Genus Pheidole references

Smith, F. 1858b: 172 (diagnosis); Mayr, 1861: 70 (Europe species key); Roger, 1863b: 27, 30 (catalogue); Mayr, 1863: 424, 440 (Ischnomyrmex, Pheidole catalogues); Mayr, 1865: 18, 22 (Ischnomyrmex, Pheidole diagnoses); Mayr, 1867a: 92, 96 (Ischnomyrmex, Pheidole diagnoses); Mayr, 1870b: 977, 979 (Australia & New World species keys); Mayr, 1876: 102 (Australia species key); André, 1883b: 382 (Europe & Algeria species key); Mayr, 1879: 674 (Asia species key); Mayr, 1887: 582 (New World species key); Cresson, 1887: 262 (U.S.A., catalogue); Nasonov, 1889: 75 (Russia species key); Dalla Torre, 1893: 88, 98 (Pheidole, Pheidolacanthinus catalogues); Emery, 1895c: 288 (North America species key); Mayr, 1896: 241 (Africa species, partial key); Emery, 1896g: 80 (P. flavens group, key); Forel, 1902c: 165 (India species key); Forel, 1902f: 523 (India & Sri Lanka species key); Bingham, 1903: 223 (India, Sri Lanka & Burma species key); Wheeler, W.M. 1908a: 133 (Puerto Rico species key); Arnold, 1920a: 414 (diagnosis, South Africa species key); Emery, 1921f: 78 (diagnosis, subgenera key, catalogue); Emery, 1921f: 81 (P. (Macropheidole) & P. (Pheidolacanthinus) diagnoses, catalogues); Emery, 1921f: 83 (P. (Stegopheidole) diagnosis, catalogue); Mann, 1921: 442 (Fiji Is species key); Emery, 1922e: 110 (P. (Elasmopheidole) diagnosis, catalogue); Emery, 1922e: 111 (P. (Trachypheidole) diagnosis, catalogue); Emery, 1922e: 112 (P. (Scrobopheidole), P. (Decapheidole) & P. (Ceratopheidole) diagnoses, catalogues); Emery, 1922e: 113 (Ischnomyrmex diagnosis, catalogue); Emery, 1922e: 114 (Epipheidole diagnosis, catalogue); Emery, 1922e: 115 (Sympheidole, Parapheidole diagnoses, catalogues); Emery, 1922e: 116 (Anergatides diagnosis, catalogue); Wheeler, W.M. 1922a: 126, 672 (diagnosis, subgenera key); Wheeler, W.M. 1922a: 806, 879 (Afrotropical Pheidole, Anergatides catalogues); Wheeler, W.M. 1922a: 1017, 1021 (Malagasy Pheidole, Parapheidole catalogues); Gallardo, 1932c: 179 (Argentina Elasmopheidole species key); Finzi, 1936: 165 (Egypt species key); Menozzi, 1939a: 298 (Himalaya species key); Cole, 1942: 362 (U.S.A., Utah species key); Buren, 1944a: 285 (U.S.A., Iowa species key); Creighton, 1950a: 163 (North America species key); Creighton, 1950a: 192, 194 (North America Epipheidole, Sympheidole species); Chapman & Capco, 1951: 135, 137 (Asia Ceratopheidole, Conothorax, Ischnomyrmex, Pheidole checklists); Kusnezov, 1952b: 61 (Argentina species key); Smith, M.R. 1955c: 302 (P. (Hendecapheidole) species key); Gregg, 1959: 9 (U.S.A. species key); Gregg, 1963: 407 (U.S.A., Colorado species key); Wheeler, G.C. & Wheeler, J. 1963: 126 (U.S.A., North Dakota species key); Bernard, 1967: 149 (diagnosis, Western Europe species key); Gregg, 1969: 95 (addition to Gregg, 1959 U.S.A. key); Kempf, 1972a: 183 (Neotropical catalogue); Alayo, 1974: 11 (Cuba species key); Smith, D.R. 1979: 1365 (North America catalogue); Allred, 1982: 441 (U.S.A., Utah species key); Ogata, 1982: 191 (Japan species key); Collingwood, 1985: 253 (Saudi Arabia species key); Naves, 1985: 54 (U.S.A., Florida species key); Taylor & Brown, D.R. 1985: 74 (Australia catalogue); Wheeler, G.C. & Wheeler, J. 1986g: 41 (U.S.A., Nevada species key); Taylor, 1987a: 53 (Australia, New Caledonia & New Zealand checklists); Agosti & Collingwood, 1987b: 271 (Balkans species key); Hölldobler & Wilson, 1990: 15 (synoptic classification); Kupyanskaya, 1990: 123 (Far Eastern Russia species key); Brandão, 1991: 368 (Neotropical catalogue); Morisita, Kubota, et al. 1992: 22 (Japan species key); Bolton, 1994: 106 (synoptic classification); Kupyanskaya, 1995: 345 (Far Eastern Russia species key); Bolton, 1995a: 1051 (census); Bolton, 1995b: 316 (catalogue); Wu, J. & Wang, 1995: 99 (China species key); Collingwood & Agosti, 1996: 322 (Saudi Arabia species key); Park & Kim, 2000: 109 (Korea species key); Shattuck, 1999: 152 (Australia synopsis); Eguchi, 1999: 103 (Borneo P. longipes group, key); Ward, 2000: 96 (U.S.A. P. hyatti complex, key); Zhou, 2001b: 121 (China, Guangxi species key); Eguchi, 2001b: 10 (Borneo species key); ; Mackay & Mackay, 2002: 149 (U.S.A., New Mexico species key)Wilson, 2003: 27 (New World, all species revision, keys); Sarnat, 2008: 8 (Fiji Is P. roosevelti group, key); Eguchi, 2008: 8 (North Vietnam species key); Terayama, 2009: 171 (Taiwan species key); Heterick, 2009: 169 (south-western Australia species key); Sarnat & Moreau, 2011: 114 (P. roosevelti group phylogeny and radiation).

Genera incertae sedis in Pheidolini

Myrmicinae Formicoxenini Cardiocondyla TRUE TRUE FALSE

67 species, 2 subspecies

Cardiocondyla Emery, 1869b: 20. Type-species: Cardiocondyla elegans, by monotypy.

Taxonomic history

Cardiocondyla in Myrmicinae: Emery, 1877a: 81 [Myrmicidae]; Emery & Forel, 1879a: 456 [Myrmicidae]; Dalla Torre, 1893: 70.

Cardiocondyla in Myrmicinae, Myrmicini: Emery, 1895j: 769; Wheeler, W.M. 1910g: 139; Kusnezov, 1964: 57 (anachronism).

Cardiocondyla in Myrmicinae, Stenammini: Ashmead, 1905b: 383.

Cardiocondyla in Myrmicinae, Cardiocondylini: Emery, 1914a: 40; Arnold, 1916: 200; Forel, 1917: 242; Emery, 1922e: 124; Wheeler, W.M. 1922a: 661; all subsequent authors; Dlussky & Fedoseeva, 1988: 80 (anachronism); Jaffe, 1993: 10 (anachronism).

Cardiocondyla in Myrmicinae, Leptothoracini: Bolton, 1982: 311 (in text).

Cardiocondyla in Myrmicinae, Formicoxenini: Bolton, 1994: 105.

Junior synonyms of CARDIOCONDYLA

Emeryia Forel, 1890b: cx. Type-species: Emeryia wroughtonii, by monotypy.

Taxonomic history

Emeryia as junior synonym of Cardiocondyla: Forel, 1892h: 461; Forel, 1892i: 313; Dalla Torre, 1893: 70.

Xenometra Emery, 1917a: 96. Type-species: Xenometra monilicornis (junior synonym of Cardiocondyla emeryi), by monotypy.

Taxonomic history

Xenometra in Myrmicinae, Cardiocondylini: Emery, 1922e: 126; Wheeler, W.M. 1922a: 661; subsequent authors.

Xenometra as junior synonym of Cardiocondyla: Baroni Urbani, 1973: 199; Marikovsky & Yakushin, 1974: 60.

Dyclona Santschi, 1930b: 70 (footnote) [as subgenus of Cardiocondyla]. Type-species: Monomorium cristatum, by original designation.

Taxonomic history

Dyclona as junior synonym of Cardiocondyla: Brown, 1973b: 180 [provisional]; Smith, D.R. 1979: 1375; Bolton, 1982: 309.

Loncyda Santschi, 1930b: 70 [as subgenus of Cardiocondyla]. Type-species: Cardiocondyla (Loncyda) monardi, by monotypy.

Taxonomic history

Loncyda as junior synonym of Cardiocondyla: Brown, 1973b: 181 [provisional]; Smith, D.R. 1979: 1375; Bolton, 1982: 309.

Prosopidris Wheeler, W.M. 1935b: 40 [as subgenus of Cardiocondyla]. Type-species: Cardiocondyla (Prosopidris) sima, by original designation.

Taxonomic history

Prosopidris in Myrmicinae, Cardiocondylini: Donisthorpe, 1943f: 688.

Prosopidris as genus: Reiskind, 1965: 80.

Prosopidris as junior synonym of Cardiocondyla: Brown, 1973b: 184 [provisional]; Smith, D.R. 1979: 1375; Bolton, 1982: 309.

Genus Cardiocondyla references

André, 1883a: 327 (Europe & Algeria species key); Dalla Torre, 1893: 70 (catalogue); Forel, 1903a: 688 (India & Sri Lanka species key); Bingham, 1903: 287 (India, Sri Lanka & Burma species key); Ruzsky, 1905b: 623 (Russian Empire species key); Emery, 1909a: 20 (Palaearctic species key); Arnold, 1916: 200 (diagnosis); Emery, 1922e: 124 (diagnosis, catalogue); Emery, 1922e: 126 (Xenometra diagnosis, catalogue); Wheeler, W.M. 1922a: 149, 827, 1021 (diagnosis, Afrotropical, Malagasy catalogues); Kuznetsov-Ugamsky, 1927d: 37 (Turkestan species key); Karavaiev, 1934: 114 (Ukraine species key); Finzi, 1936: 172 (Egypt species key); Smith, M.R. 1944a: 32 (U.S.A. species key); Creighton, 1950a: 197 (North America species key); Chapman & Capco, 1951: 83 (Asia checklist); Bernard, 1956c: 301 (Palaearctic species key); Bernard, 1967: 156 (diagnosis, Western Europe species key); Bernard, 1967: 158 (Xenometra diagnosis); Kempf, 1972a: 73, 259 (Neotropical Cardiocondyla, Xenometra catalogues); Alayo, 1974: 12 (Cuba species key); Tarbinsky, 1976: 71 (Kyrghyzstan species key); Arnol'di & Dlussky, 1978: 538 (former European U.S.S.R. species key); Collingwood, 1978: 86 (Iberian Peninsula species key); Smith, D.R. 1979: 1375 (North America catalogue); Bolton, 1982: 309 (diagnosis, Afrotropical species key); Kugler, J. 1984: 17 (males, key); Collingwood, 1985: 256 (Saudi Arabia species key); Agosti & Collingwood, 1987b: 276 (Balkans species key); Dlussky, Soyunov & Zabelin, 1990: 193 (Turkmenistan species key); Morisita, Kubota, et al. 1992: 31 (Japan species key); Atanasov & Dlussky, 1992: 172 (Bulgaria species key); Arakelian, 1994: 42 (Armenia species key); Bolton, 1995a: 1048 (census); Bolton, 1995b: 132 (catalogue); Radchenko, 1995b: 452 (Palaearctic species key); Mackay, 1995: 170 (New World species key); Wu, J. & Wang, 1995: 68 (China species key); Collingwood & Agosti, 1996: 326 (Saudi Arabia species key); Collingwood & Prince, 1998: 16 (Portugal species key); Terayama, 1999d: 99 (Japan species, review); Shattuck, 1999: 129 (Australia synopsis); Zhou, 2001b: 85 (China, Guangxi species key); Rigato, 2002: 172 (Afrotropical species key); Seifert, 2003a: 292 (C. elegans, C. bulgarica, C. batesii, C. nuda, C. shuckardi, C. stambuloffii, C. wroughtonii, C. emeryi, C. minutior groups revision, key); Terayama, 2009: 179 (Taiwan species key).

Myrmicinae Formicoxenini Harpagoxenus TRUE TRUE FALSE

3 species, 1 subspecies

Harpagoxenus Forel, 1893a: 167.

Taxonomic history

[Replacement name for Tomognathus Mayr, 1861: 56; junior homonym of Tomognathus Agassiz, 1850: 376 (Pisces).]

Harpagoxenus in Myrmicinae, Stenammini: Ashmead, 1905b: 383.

Harpagoxenus in Myrmicinae, Myrmicini: Wheeler, W.M. 1910g: 139.

Harpagoxenus in Myrmicinae, Leptothoracini: Emery, 1914a: 42; Forel, 1917: 245; Wheeler, W.M. 1922a: 664; Emery, 1924d: 265; all subsequent authors to the following.

Harpagoxenus in Myrmicinae, Formicoxenini: Bolton, 1994: 105.

Genus Harpagoxenus references

Mayr, 1863: 456 (catalogue); Mayr, 1865: 23 (diagnosis); André, 1882c: 279 (Europe & Algeria species); Dalla Torre, 1893: 63 (catalogue); Emery, 1924d: 265 (diagnosis, catalogue); Creighton, 1950a: 284 (North America species key); Bernard, 1967: 223 (diagnosis); Smith, D.R. 1979: 1398 (North America catalogue); Radchenko, 1994b: 112 (South Siberia species key); Bolton, 1995a: 1049 (census); Bolton, 1995b: 211 (catalogue).

Homonym replaced by HARPAGOXENUS

Tomognathus Mayr, 1861: 56. Type-species: Myrmica sublaevis, by monotypy.

Taxonomic history

[Junior homonym of Tomognathus Agassiz, 1850: 376 (Pisces).]

Tomognathus in Myrmicinae: Mayr, 1861: 56 [Myrmicidae]; Mayr, 1865: 23 [Myrmicidae]; Emery & Forel, 1879a: 457 [Myrmicidae]; Dalla Torre, 1893: 64.

Tomognathus in Myrmicinae, Formicoxenini: Forel, 1893a: 165.

Tomognathus in Myrmicidae, Myrmicidae: Emery, 1877a: 81.

Tomognathus in Myrmicinae, Myrmicini: Emery, 1895j: 769.

Myrmicinae Formicoxenini Leptothorax TRUE TRUE FALSE

Extant: 18 species, 2 subspecies

Fossil:

Leptothorax Mayr, 1855: 431. Type-species: Formica acervorum, by subsequent designation of Bingham, 1903: 214.

Taxonomic history

[Type-species not Myrmica clypeata, unjustified subsequent designation by Emery, 1912d: 271; repeated in Wheeler, W.M. 1913a: 79 and Emery, 1924d: 248.]

Leptothorax in Myrmicidae, Myrmicidae: Emery, 1877a: 81.

Leptothorax in Myrmicinae: Mayr, 1855: 431 [Myrmicidae]; Mayr, 1861: 57 [Myrmicidae]; Mayr, 1865: 20 [Myrmicidae]; Emery & Forel, 1879a: 458 [Myrmicidae]; Dalla Torre, 1893: 122.

Leptothorax in Myrmicinae, Myrmicini: Forel, 1895b: 125; Emery, 1895j: 769; Forel, 1899c: 54; Wheeler, W.M. 1910g: 139; Wheeler, W.M. 1915h: 63; Kusnezov, 1964: 57 (anachronism).

Leptothorax in Myrmicinae, Stenammini: Ashmead, 1905b: 383.

Leptothorax in Myrmicinae, Leptothoracini: Emery, 1914a: 42; Arnold, 1916: 257; Forel, 1917: 244; Wheeler, W.M. 1922a: 664; Emery, 1924d: 247; subsequent authors to the following.

Leptothorax in Myrmicinae, Formicoxenini: Bolton, 1994: 105; Mackay, 2000: 267.

Junior synonyms of LEPTOTHORAX

Mychothorax Ruzsky, 1904a: 288. Type-species: Formica acervorum, by original designation.

Taxonomic history

Mychothorax as subgenus of Leptothorax: Ruzsky, 1905b: 609; Emery, 1915g: 24; Emery, 1916b: 176; Forel, 1917: 245; Wheeler, W.M. 1922a: 679; Emery, 1924d: 260; Creighton, 1950a: 274.

Mychothorax as junior synonym of Leptothorax: Smith, M.R. 1950: 29; Brown, 1973b: 182 [provisional]; Bolton, 1982: 319. [Leptothorax and Mychothorax share the same type-species, synonymy is therefore absolute.]

Doronomyrmex Kutter, 1945: 485. Type-species: Doronomyrmex pacis, by monotypy.

Taxonomic history

Doronomyrmex in Myrmicinae, Leptothoracini:Dlussky & Fedoseeva, 1988: 79.

Doronomyrmex in Myrmicinae, Formicoxenini: Bolton, 1994: 105.

Doronomyrmex as junior synonym of Leptothorax: Brown, 1973b: 180 [provisional]; Heinze, 1998: 195; Bolton, 2003: 270.

Genus Leptothorax references

Mayr, 1855: 433 (Austria species key); Mayr, 1861: 58 (Europe species key); Roger, 1863b: 26, 28, 30 (catalogue); Mayr, 1863: 426, 428, 456 (Leptothorax, Macromischa, Temnothorax catalogues); Mayr, 1865: 19, 20, 21 (Macromischa, Leptothorax, Temnothorax diagnoses); Mayr, 1868c: 83 (†Baltic Amber species key); André, 1874: 188 (Europe species key); Forel, 1874: 84 (Switzerland species key); André, 1883a: 293 (Europe & Algeria species key); Mayr, 1886d: 451 (U.S.A. species key); Cresson, 1887: 261 (U.S.A. catalogue); Nasonov, 1889: 70 (Russia species key); Emery, 1891b: 5 (North Africa species key); Dalla Torre, 1893: 120, 122 (Macromischa, Leptothorax catalogues); Emery, 1895c: 317 (North America species key); Emery, 1896g: 58 (Neotropical species key); Bingham, 1903: 215 (India, Sri Lanka & Burma species key); Wheeler, W.M. 1903c: 223 (North America species key); Ruzsky, 1905b: 570 (Russian Empire species key); Wasmann, 1906: 16 (Luxemburg species key); Wheeler, W.M. 1908a: 141 (Macromischa species key); Santschi, 1909b: 460 (L. rottenbergi group key); Bondroit, 1910: 496 (Belgium species key); Stitz, 1914: 60 (Central Europe species key); Crawley, 1914: 91 (Britain species key); Donisthorpe, 1915d: 147 (Britain species key); Emery, 1916b: 176 (Italy species key); Wheeler, W.M. 1916m: 581 (U.S.A., Connecticut species key); Arnold, 1916: 257 (diagnosis, South Africa species key); Bondroit, 1918: 117 (France & Belgium species key); Mann, 1920: 408 (Macromischa species key); Wheeler, W.M. 1922a: 677 (Macromischa subgenera key); Wheeler, W.M. 1922a: 679 (Leptothorax subgenera key); Wheeler, W.M. 1922a: 890 (Afrotropical catalogue); Wheeler, W.M. 1922a: 1029 (Malagasy catalogue); Emery, 1924d: 245 (Macromischa catalogue); Emery, 1924d: 247 (diagnosis, subgenera key, catalogue); Emery, 1924d: 249 (L. (Goniothorax) diagnosis, catalogue); Emery, 1924d: 251 (L. (Leptothorax) diagnosis, catalogue); Emery, 1924d: 259 (L. (Temnothorax) diagnosis, catalogue); Emery, 1924d: 260 (L. (Dichothorax) & L. (Mychothorax) diagnoses, catalogues); Emery, 1924d: 264 (Symmyrmica review, catalogue); Karavaiev, 1927c: 266 (Ukraine species key); Donisthorpe, 1927b: 163 (Britain species key); Kuznetsov-Ugamsky, 1927d: 38 (Turkestan species key); Wheeler, W.M. 1931b: 31 (Macromischa, Croesomyrmex, Antillaemyrmex checklists); Arnol'di, 1933b: 598 (Russia species key); Menozzi, 1933b: 68 (Israel species key); Karavaiev, 1934: 134 (Ukraine species key); Wheeler, W.M. 1937b: 463 (Macromischa checklist); Menozzi, 1939a: 307 (Himalaya & Tibet species key); Stitz, 1939: 158 (Germany species key); Smith, M.R. 1939e: 503 (U.S.A. Macromischa species key); Kratochvíl, 1941: 89 (Central Europe species key); Novák & Sadil, 1941: 89 (Central Europe species key); Cole, 1942: 369 (U.S.A., Utah species key); Holgersen, 1943b: 172 (Norway species key); Holgersen, 1944: 198 (Norway species key); Kratochvíl, Novák & Snoflák, 1944: 123 (Czechoslovakia males, key); Buren, 1944a: 286 (U.S.A., Iowa species key); Boven, 1947: 178 (Belgium species key); Creighton, 1950a: 251, 256 (North America Macromischa, Leptothorax species keys); Chapman & Capco, 1951: 110 (Asia checklist); Smith, M.R. 1952b: 97 (U.S.A. L. tricarinatus complex, key); Bernard, 1956a: 151 (Western Europe species groups); Kusnezov, 1958c: 266 (subgenera key); Boven, 1959: 8 (Netherlands species key); Gregg, 1963: 380 (U.S.A., Colorado species key); Wheeler, G.C. & Wheeler, J. 1963: 139 (U.S.A., North Dakota species key); Collingwood, 1964: 99 (Britain species key); Bernard, 1967: 185 (diagnosis, Western Europe species key); Bernard, 1967: 187 (diagnosis, Western Europe species key); Bernard, 1967: 224 (Doronomyrmex diagnosis); Boven, 1970b: 20 (Netherlands species key); Arnol'di, 1971: 1824 (Kazakhstan species key); Kempf, 1972a: 132, 135 (Neotropical Leptothorax, Macromischa catalogues); Alayo, 1974: 15 (Macromischa Cuba species key); Bolton & Collingwood, 1975: 5 (Britain species key); Tarbinsky, 1976: 83 (Kyrghyzstan species key); Boven, 1977: 94 (Belgium species key); Kutter, 1977c: 103 (Switzerland species key); Arnol'di & Dlussky, 1978: 540 (former European U.S.S.R. species key); Collingwood, 1978: 83 (Iberian Peninsula species key); Collingwood, 1979: 68 (Fennoscandia & Denmark species key); Baroni Urbani, 1978b: 533 (L. (Macromischa) all species revision, key); Smith, D.R. 1979: 1391 (North America catalogue); Buschinger, 1981: 211 (Doronomyrmex review); Allred, 1982: 440 (U.S.A., Utah species key); Bolton, 1982: 319 (diagnosis, review of genus, Afrotropical species key); Gösswald, 1985: 303 (Germany species key); Wheeler, G.C. & Wheeler, J. 1986g: 51 (U.S.A., Nevada species key); Nilsson & Douwes, 1987: 60 (Norway species key); Agosti & Collingwood, 1987b: 273 (Balkans species key); Dlussky & Soyunov, 1988: 29 (former U.S.S.R. Temnothorax species); Hölldobler & Wilson, 1990: 13 (synoptic classification); Dlussky, Soyunov & Zabelin, 1990: 184, 188 (Turkmenistan species key); Kupyanskaya, 1990: 136 (Far Eastern Russia species key); Brandão, 1991: 349 (Neotropical catalogue); Morisita, Kubota, et al. 1992: 26 (Japan species key); Atanasov & Dlussky, 1992: 129 (Bulgaria species key); Arakelian, 1994: 52 (Armenia species key); Radchenko, 1994b: 111 (South Siberia species key); Radchenko, 1994d: 146 (Central & Eastern Palaearctic species key); Bolton, 1995a: 1050 (census); Bolton, 1995b: 235 (catalogue); Douwes, 1995: 89 (Sweden species key); Kupyanskaya, 1995: 347 (Far Eastern Russia species key); Wu, J. & Wang, 1995: 108 (China species key); Collingwood & Agosti, 1996: 325 (Saudi Arabia species key); Seifert, 1996b: 13 (Central Europe Doronomyrmex species key); Seifert, 1996b: 119 (Central Europe species key); Skinner & Allen, 1996: 45 (Britain species key); Cagniant & Espadaler, 1997: 260 (Morocco species key); Collingwood & Prince, 1998: 16 (Portugal species key); Terayama & Onoyama, 1999: 73 (Japan species key); Mackay, 2000: 265, 288, 297 (New World L. (Myrafant) species revision key, Mexico & Guatemala species key, species complexes key); Fontenla Rizo, 2001: 15 (Cuba Macromischa species key); Chang & He, 2001b: 1 (northwest China species key); Czechowski, Radchenko & Czechowska, 2002: 143 (Poland species key); Mackay & Mackay, 2002: 93 (U.S.A., New Mexico species key); Bolton, 2003: 270 (genus diagnosis); Lyu & Cho, 2003b: 269 (Korea species key); Radchenko, 2004: 131 (Eastern Palaearctic species, key); Deyrup & Cover, 2004a: 56 (southeast U.S.A. species key); Radchenko, 2005b: 192 (North Korea species key); Seifert, 2007: 118 (North and Central Europe species key).

Myrmicinae Formicoxenini Xenomyrmex TRUE TRUE FALSE

3 species, 1 subspecies

Xenomyrmex Forel, 1885a: 369. Type-species: Xenomyrmex stollii, by monotypy.

Taxonomic history

Xenomyrmex in Myrmicinae: Dalla Torre, 1893: 64.

Xenomyrmex in Myrmicinae, Formicoxenini: Forel, 1893a: 165.

Xenomyrmex in Myrmicinae, Myrmicini: Emery, 1895j: 769; Forel, 1899c: 52; Wheeler, W.M. 1910g: 139.

Xenomyrmex in Myrmicinae, Stenammini: Ashmead, 1905b: 383.

Xenomyrmex in Myrmicinae, Solenopsidini: Emery, 1914a: 41 [subtribe Monomoriini]; Forel, 1917: 243; Emery, 1922e: 188; Wheeler, W.M. 1922a: 663; Wheeler, W.M. 1931a: 131; all subsequent authors to the following.

Xenomyrmex incertae sedis in Myrmicinae: Ettershank, 1966: 81.

Xenomyrmex in Myrmicinae, Solenopsidini: Smith, D.R. 1979: 1384.

Xenomyrmex in Myrmicinae, Archaeomyrmecini?: Brandão, 1991: 391.

Xenomyrmex in Myrmicinae, Myrmecinini: Kempf, 1972a: 259; Dlussky & Fedoseeva, 1988: 79; Jaffe, 1993: 11.

Xenomyrmex in Myrmicinae, Metaponini?: Hölldobler & Wilson, 1990: 16; Bolton, 1994: 105.

Xenomyrmex in Myrmicinae, Formicoxenini: Bolton, 2003: 70.

Junior synonym of XENOMYRMEX

Myrmecinella Wheeler, W.M. 1922d: 1. Type-species: Myrmecinella panamana, by original designation.

Taxonomic history

Myrmecinella in Myrmicinae, Myrmecinini: Wheeler, W.M. 1922d: 3; Donisthorpe, 1943f: 665.

Myrmecinella as junior synonym of Xenomyrmex: Wheeler, W.M. 1931a: 129.

Genus Xenomyrmex references

Dalla Torre, 1893: 64 (catalogue); Emery, 1922e: 188 (diagnosis, catalogue); Wheeler, W.M. 1931a: 129 (diagnosis, all species key); Creighton, 1957c: 6 (all species key); Ettershank, 1966: 150 (diagnosis, review of genus, checklist); Smith, D.R. 1979: 1384 (North America catalogue); Bolton, 1995a: 1053 (census); Bolton, 1995b: 424 (catalogue).

Genera incertae sedis in Formicoxenini

Myrmicinae Myrmecinini Myrmecina TRUE TRUE FALSE

51 species, 1 subspecies

Myrmecina Curtis, 1829: 265. Type-species: Myrmecina latreillii (junior synonym of Formica graminicola), by monotypy.

Taxonomic history

Myrmecina in Poneridae, Myrmicidae: Smith, F. 1858b: 132.

Myrmecina in Myrmicidae: Smith, F. 1871a: 327; Cresson, 1887: 261.

Myrmecina in Myrmicidae, Myrmicidae: Emery, 1877a: 81.

Myrmecina in Myrmicinae: Mayr, 1855: 420 [Myrmicidae]; Mayr, 1861: 73 [Myrmicidae]; Mayr, 1865: 21 [Myrmicidae]; Emery & Forel, 1879a: 457 [Myrmicidae]; Dalla Torre, 1893: 61.

Myrmecina in Myrmicinae, Myrmicini: Emery, 1895j: 769; Wheeler, W.M. 1910g: 139.

Myrmecina in Myrmicinae, Myrmecinini: Ashmead, 1905b: 383; Emery, 1912b: 105; Emery, 1914a: 41; Forel, 1917: 244; Wheeler, W.M. 1922a: 663; Emery, 1924d: 230; all subsequent authors.

Junior synonym of MYRMECINA

Archaeomyrmex Mann, 1921: 448. Type-species: Archaeomyrmex cacabau, by original designation.

Taxonomic history

Archaeomyrmex in Myrmicinae, Archaeomyrmecini: Mann, 1921: 449; Wheeler, W.M. 1922a: 663.

Archaeomyrmex as junior synonym of Myrmecina: Brown, 1971a: 1.

Genus Myrmecina references

Smith, F. 1858b: 132 (diagnosis); Roger, 1863b: 33 (catalogue); Mayr, 1863: 429 (catalogue); Mayr, 1865: 21 (diagnosis); André, 1882c: 275 (Europe & Algeria species); Cresson, 1887: 261 (U.S.A. catalogue); Dalla Torre, 1893: 61 (catalogue); Bingham, 1903: 197 (diagnosis); Emery, 1916b: 171 (Italy species key); Mann, 1919: 337 (Papuasian species key); Emery, 1924d: 230 (diagnosis, catalogue); Smith, M.R. 1948: 239 (U.S.A. species key); Creighton, 1950a: 246 (North America review); Chapman & Capco, 1951: 117 (Asia checklist); Bernard, 1967: 226 (diagnosis); Smith, D.R. 1979: 1399 (North America catalogue); Taylor & Brown, D.R. 1985: 71 (Australia catalogue); Taylor, 1987a: 47 (Australia checklist); Morisita, Kubota, et al. 1992: 54 (Japan species key); Bolton, 1994: 105 (synoptic classification); Bolton, 1995a: 1050 (census); Bolton, 1995b: 274 (catalogue); Terayama, 1996: 31 (Japan species key); Lin & Wu, 1998: 89 (Taiwan species key); Rigato, 1999: 83 (West Palaearctic species key); Shattuck, 1999: 147 (Australia synopsis); Zhou, 2001b: 138 (China, Guangxi species key); Radchenko, 2005b: 192 (North Korea species key); Zhou, Huang & Ma, 2008: 285 (China species key); Shattuck, 2009: 3 (Australia species key); Terayama, 2009: 187 (Taiwan species key).

Amblyoponinae Amblyoponini Amblyopone TRUE TRUE FALSE

Extant: 72 species

Fossil: 2 species

Amblyopone Erichson, 1842: 260. Type-species: Amblyopone australis, by monotypy.

Taxonomic history

Amblyopone in Poneridae: Smith, F. 1858b: 108; Smith, F. 1871a: 324.

Amblyopone in Ponerinae: Mayr, 1862: 714 (in key) [Poneridae]; Mayr, 1865: 16 [Poneridae]; Emery & Forel, 1879a: 455 [Poneridae]; Dalla Torre, 1893: 13.

Amblyopone in Amblyoponinae: Forel, 1893a: 162; Clark, 1951: 15 (in key).

Amblyopone in Pachycondylinae, Amblyoponini: Ashmead, 1905b: 382.

Amblyopone in Ponerinae, Amblyoponini: Emery, 1895c: 261; Emery, 1895j: 766; Emery, 1901a: 34; Wheeler, W.M. 1910g: 134; Emery, 1911d: 25; Forel, 1917: 235; Wheeler, W.M. 1922a: 641; Donisthorpe, 1943f: 621; Brown, 1953c: 11; all subsequent authors to the following.

Amblyopone in Amblyoponinae, Amblyoponini: Bolton, 2003: 42, 154.

Junior synonyms of AMBLYOPONE

Stigmatomma Roger, 1859: 250. Type-species: Stigmatomma denticulatum, by subsequent designation of Bingham, 1903: 36.

Taxonomic history

Stigmatomma in Ponerinae: Mayr, 1861: 53 [Poneridae]; Mayr, 1862: 714 [Poneridae]; Mayr, 1865: 16 [Poneridae]; Dalla Torre, 1893: 14.

Stigmatomma in Amblyoponinae: Forel, 1893a: 162; Forel, 1895b: 110 [Amblyoponeridae]

Stigmatomma in Pachycondylinae, Amblyoponini: Ashmead, 1905b: 283.

Stigmatomma in Ponerinae, Amblyoponini: Emery, 1895j: 766; Emery, 1901a: 34; Wheeler, W.M. 1910g: 134; Emery, 1911d: 23; Forel, 1917: 235; Wheeler, W.M. 1922a: 641.

Stigmatomma as subgenus of Amblyopone: Forel, 1900c: 55; Clark, 1934b: 27; Brown, 1949c: 87.

Stigmatomma as genus: Roger, 1859: 250; Dalla Torre, 1893: 14; Emery, 1895j: 766; Bingham, 1903: 36; Emery, 1911d: 23; Forel, 1917: 235; Wheeler, W.M. 1922a: 641; Borgmeier, 1923: 52; Donisthorpe, 1943g: 728; Creighton, 1950a: 31; Kusnezov, 1956: 12.

Stigmatomma as junior synonym of Amblyopone: Emery & Forel, 1879a: 455; Mayr, 1887: 546; Brown, 1960a: 155; all subsequent authors.

Arotropus Provancher, 1881a: 205. Type-species: Arotropus binodosus (junior synonym of Typhlopone pallipes), by monotypy.

Taxonomic history

Arotropus as junior synonym of Amblyopone: Provancher, 1887: 240; Brown, 1960a: 155.

Amblyopopone Dalla Torre, 1893: 13, unjustified emendation of Amblyopone.

Taxonomic history

Amblyopopone as junior synonym of Amblyopone: Forel, 1893a: 166; Emery, 1911d: 25.

Amblyopopona Schulz, W.A. 1906: 154, unjustified emendation of Amblyopone.

Taxonomic history

Amblyopopona as junior synonym of Amblyopone: Bolton, 1994: 164.

Xymmer Santschi, 1914d: 311 [as subgenus of Stigmatomma]. Type-species: Stigmatomma (Xymmer) muticum, by monotypy.

Taxonomic history

Xymmer as subgenus of Stigmatomma: Santschi, 1914d: 311; Forel, 1917: 235; Emery, 1919c: 106; Donisthorpe, 1943g: 737.

Xymmer as genus: Wheeler, W.M. 1922a: 641 (in key).

Xymmer as subgenus of Amblyopone: Clark, 1934b: 27.

Xymmer as junior synonym of Stigmatomma: Brown, 1949c: 87.

Xymmer as junior synonym of Amblyopone: Brown, 1960a: 155.

Fulakora Mann, 1919: 279 [as subgenus of Stigmatomma]. Type-species: Stigmatomma (Fulakora) celata, by original designation.

Taxonomic history

Fulakora as junior synonym of Stigmatomma: Brown, 1949c: 88.

Fulakora as junior synonym of Amblyopone: Brown, 1960a: 155.

Neoamblyopone Wheeler, W.M. 1927c: 1 [as subgenus of Amblyopone]. Type-species: Amblyopone (Neoamblyopone) clarki, by monotypy.

Taxonomic history

Neoamblyopone as junior synonym of Amblyopone: Brown, 1949c: 87; Brown, 1960a: 155.

Protamblyopone Wheeler, W.M. 1927c: 1 [as subgenus of Amblyopone]. Type-species: Amblyopone (Protamblyopone) aberrans, by monotypy.

Taxonomic history

Protamblyopone as junior synonym of Amblyopone: Brown, 1949c: 87; Brown, 1960a: 155.

Lithomyrmex Clark, 1928a: 30. Type-species: Lithomyrmex glauerti, by original designation.

Taxonomic history

Lithomyrmex in Ponerinae, Amblyoponini: Clark, 1928a: 30.

Lithomyrmex as junior synonym of Amblyopone: Brown, 1960a: 156.

Ericapelta Kusnezov, 1955: 273. Type-species: Ericapelta egregia, by monotypy.

Taxonomic history

Ericapelta as junior synonym of Amblyopone: Brown, 1960a: 156.

Genus Amblyopone references

Roger, 1863b: 20 (catalogue); Mayr, 1863: 394, 454 (catalogue); Mayr, 1865: 16 (Amblyopone, Stigmatomma diagnoses); André, 1882c: 233 (Europe & Algeria species key); Cresson, 1887: 259 (U.S.A. catalogue); Dalla Torre, 1893: 13, 14 (Amblyopone, Stigmatomma catalogues); Forel, 1900c: 55 (India & Sri Lanka species key); Bingham, 1903: 37 (India, Sri Lanka & Burma Stigmatomma species key); Emery, 1911d: 23 (Stigmatomma diagnosis, catalogue); Emery, 1911d: 25 (diagnosis, catalogue); Santschi, 1915a: 55 (Mediterranean species key); Emery, 1916b: 100 (Italy species key); Bondroit, 1918: 80 (France & Belgium species key); Wheeler, W.M. 1922a: 758 (Afrotropical catalogue); Wheeler, W.M. 1927c: 3 (Austral & New Guinea species key); Chapman & Capco, 1951: 22, 24 (Asia Amblyopone, Stigmatomma checklists); Brown, 1958h: 11 (New Zealand species); Brown, 1960a: 155, 191 (diagnosis & review of genus, New World species key); Kempf, 1972a: 19 (Neotropical catalogue); Baroni Urbani, 1978a: 41 (Mediterranean species key); Smith, D.R. 1979: 1334 (North America catalogue); Taylor, 1979: 824 (Melanesia species key); Taylor & Brown, D.R. 1985: 18 (Australia catalogue); Taylor, 1987a: 6 (Australia, New Caledonia & New Zealand checklist); Agosti & Collingwood, 1987b: 264 (Balkans species key); Dlussky & Fedoseeva, 1988: 78 (synoptic classification); Morisita, Kubota, et al. 1989: 8 (Japan species key); Terayama, 1989b: 345 (Taiwan species key); Tinaut, 1990d: 189 (Iberian Peninsula species); Dlussky, Soyunov & Zabelin, 1990: 173 (Turkmenistan species key); Lattke, 1991c: 6 (New World species key); Atanasov & Dlussky, 1992: 61 (Bulgaria species key); Bolton, 1995a: 1047 (census); Bolton, 1995b: 61 (catalogue); Onoyama, 1999: 190 (Japan species key); Shattuck, 1999: 179 (Australia synopsis); Xu, 2001d: 552 (East & South Asia species key); Lacau & Delabie, 2002: 40 (Neotropical species key); Xu, 2006: 152 (China species key); Arias-Penna, 2008a: 43 (Neotropical species key); Terayama, 2009: 93 (Taiwan species key); Heterick, 2009: 131 (south-western Australia species key).

Amblyoponinae Amblyoponini Prionopelta TRUE TRUE FALSE

15 species

Prionopelta Mayr, 1866a: 503. Type-species: Prionopelta punctulata, by monotypy.

Taxonomic history

Prionopelta in Ponerinae: Dalla Torre, 1893: 15.

Prionopelta in Amblyoponinae: Forel, 1893a: 162; Forel, 1895b: 110 [Amblyoponeridae].

Prionopelta in Ponerinae, Proceratiini: Ashmead, 1905b: 382.

Prionopelta in Ponerinae, Ectatommini: Emery, 1911d: 32 [subtribe Typhlomyrmecini]; Forel, 1917: 236; Wheeler, W.M. 1922a: 642.

Prionopelta in Ponerinae, Amblyoponini: Emery, 1895j: 766; Wheeler, W.M. 1910g: 134; Brown, 1953c: 11; Brown, 1960a: 173; all subsequent authors to the following.

Prionopelta in Amblyoponinae, Amblyoponini: Bolton, 2003: 42, 156.

Junior synonyms of PRIONOPELTA

Renea Donisthorpe, 1947d: 183. Type-species: Renea testacea (junior synonym of Prionopelta majuscula), by original designation.

Taxonomic history

Renea as genus: Donisthorpe, 1947d: 183; Chapman & Capco, 1951: 77.

Renea as junior synonym of Prionopelta: Brown, 1953c: 11.

Examblyopone Donisthorpe, 1949g: 401. Type-species: Examblyopone churchilli (junior synonym of Prionopelta majuscula), by original designation.

Taxonomic history

Examblyopone as junior synonym of Prionopelta: Brown, 1951: 102.

Genus Prionopelta references

Dalla Torre, 1893: 15 (catalogue); Forel, 1909a: 242 (New World species key); Emery, 1911d: 32 (diagnosis, catalogue); Chapman & Capco, 1951: 26 (Asia checklist); Wilson, 1958a: 146 (Melanesia species key); Brown, 1960a: 173, 218, 221 (review of genus, Neotropical species key, Indo-Australian species key); Kempf, 1972a: 210 (Neotropical catalogue); Smith, D.R. 1979: 1335 (North America catalogue); Taylor & Brown, D.R. 1985: 39 (Australia catalogue); Taylor, 1987a: 64 (Australia & New Caledonia checklist); Bolton, 1995a: 1052 (census); Bolton, 1995b: 364 (catalogue); Shattuck, 1999: 201 (Australia synopsis); Shattuck, 2008b: 22 (Indo-Pacific species revision, key) ); Arias-Penna, 2008a: 48 (Neotropical species key).

Ponerinae Ponerini Hypoponera TRUE TRUE FALSE

Extant: 142 species, 26 subspecies

Fossil: 1 species

Hypoponera Santschi, 1938b: 79 [as subgenus of Ponera]. Type-species: Ponera abeillei, by original designation.

Taxonomic history

Hypoponera in Ponerinae, Ponerini: Donisthorpe, 1943f: 652; all subsequent authors.

Hypoponera as subgenus of Ponera: Santschi, 1938b: 79.

Hypoponera as genus: Taylor, 1967a: 9; all subsequent authors.

Genus Hypoponera references

[References prior to 1967, see under PONERA.]

Taylor, 1967a: 9 (Hypoponera diagnosis, discussion); Kempf, 1972a: 121 (Neotropical catalogue); Alayo, 1974: 7 (Cuba species key); Kutter, 1977c: 24 (Switzerland species key); Arnol'di & Dlussky, 1978: 524 (former European U.S.S.R. species key); Smith, D.R. 1979: 1342 (North America catalogue); Taylor & Brown, D.R. 1985: 31 (Australia catalogue); Taylor, 1987a: 29 (Australia, New Caledonia & New Zealand checklists); Agosti & Collingwood, 1987b: 265 (Balkans species key); Morisita, Kubota, et al. 1989: 25 (Japan species key); Dlussky, Soyunov & Zabelin, 1990: 176 (Turkmenistan species key); Brandão, 1991: 347 (Neotropical catalogue); Atanasov & Dlussky, 1992: 69 (Bulgaria species key); Bolton, 1994: 164 (synoptic classification); Bolton, 1995a: 1050 (census); Bolton, 1995b: 213 (catalogue); Wu, J. & Wang, 1995: 40 (China species key); Collingwood & Agosti, 1996: 311 (Saudi Arabia species key); Kim, Kim & Kim, 1998: 148 (Korea species key); Shattuck, 1999: 187 (Australia synopsis); Bolton, 2003: 162 (synopsis); Terayama, 2009: 113 (Taiwan species key); Heterick, 2009: 133 (south-western Australia species key); Bolton & Fisher, 2011: 17, 23 (Afrotropical and West Palaearctic, all species revision, keys).

Ponerinae Ponerini Leptogenys TRUE TRUE FALSE

Extant: 216 species, 31 subspecies

Fossil: 1 species

Leptogenys Roger, 1861a: 41. Type-species: Leptogenys falcigera, by subsequent designation of Bingham, 1903: 52.

Taxonomic history

Leptogenys in Ponerinae: Dalla Torre, 1893: 46; Mayr, 1862: 714 (in key) [Poneridae]; Mayr, 1865: 14 [Poneridae].

Leptogenys in Ponerinae, Leptogenyini: Forel, 1893a: 162; Forel, 1899c: 17; Ashmead, 1905b: 382; Emery, 1911d: 97; Arnold, 1915: 89; Forel, 1917: 238; Gallardo, 1918b: 84; Wheeler, W.M. 1922a: 652; Donisthorpe, 1943f: 656; Chapman & Capco, 1951: 31; Dlussky & Fedoseeva, 1988: 78.

Leptogenys in Ponerinae, Ponerini: Emery, 1895j: 767; Forel, 1900d: 304; Wheeler, W.M. 1910g: 135; Brown, 1963: 3; Kempf, 1972a: 129; Jaffe, 1993: 8; Bolton, 1994: 164; Bolton, 2003: 163.

Junior synonyms of LEPTOGENYS

Lobopelta Mayr, 1862: 714 (diagnosis in key), 733. Type-species: Ponera diminuta, by subsequent designation of Bingham, 1903: 54.

Taxonomic history

Lobopelta in Ponerinae: Mayr, 1862: 714 (in key) [Poneridae]; Mayr, 1865: 15 [Poneridae]; Dalla Torre, 1893: 43.

Lobopelta in Ponerinae, Leptogenyini: Forel, 1893a: 162; Ashmead, 1905b: 382; Emery, 1911d: 101; Wheeler, W.M. 1922a: 653; Donisthorpe, 1943f: 658.

Lobopelta as genus: Mayr, 1862: 733; Cresson, 1887: 258; Dalla Torre, 1893: 43; Bingham, 1903: 54.

Lobopelta as subgenus of Leptogenys: Forel, 1892k: 520; Forel, 1899c: 18; Forel, 1900d: 304; Wheeler, W.M. 1910g: 135; Emery, 1911d: 101; Arnold, 1915: 95; Forel, 1917: 238; Gallardo, 1918b: 85; Wheeler, W.M. 1922a: 653; Wheeler, W.M. 1923d: 1; Borgmeier, 1923: 75; Creighton, 1950a: 50; Kempf, 1972a: 131; Smith, D.R. 1979: 1343 (anachronism).

Lobopelta as junior synonym of Leptogenys: Emery, 1896e: 177 (footnote); Brown, 1973b: 181 [provisional]; Bolton, 1975a: 240; Bolton, 2003: 163.

Prionogenys Emery, 1895g: 348. Type-species: Prionogenys podenzanai, by monotypy.

Taxonomic history

Prionogenys in Ponerinae, Ponerini: Emery, 1895j: 767; Wheeler, W.M. 1910g: 135;

Prionogenys in Ponerinae, Leptogenyini: Ashmead, 1905b: 382; Emery, 1911d: 106; Forel, 1917: 238; Wheeler, W.M. 1922a: 652; Donisthorpe, 1943f: 685; Dlussky & Fedoseeva, 1988: 78.

Prionogenys as junior synonym of Leptogenys: Taylor, 1988: 33.

Machaerogenys Emery, 1911d: 100 [as subgenus of Leptogenys]. Type-species: Leptogenys truncatirostris, by original designation.

Taxonomic history

Machaerogenys as subgenus of Leptogenys: Emery, 1911d: 100; Forel, 1917: 238; Wheeler, W.M. 1922a: 653; subsequent authors to the following.

Machaerogenys as junior synonym of Leptogenys: Brown, 1973b: 181 [provisional]; Bolton, 1975a: 240.

Odontopelta Emery, 1911d: 101 [as subgenus of Leptogenys]. Type-species: Leptogenys turneri, by monotypy.

Taxonomic history

Odontopelta as subgenus of Leptogenys: Emery, 1911d: 101; Forel, 1917: 238; Wheeler, W.M. 1922a: 653; Donisthorpe, 1943f: 677.

Odontopelta as junior synonym of Leptogenys: Brown, 1973b: 183 [provisional]; Snelling, R.R. 1981: 390; Taylor & Brown, D.R. 1985: 32.

Dorylozelus Forel, 1915b: 24. Type-species: Dorylozelus mjobergi (junior secondary homonym in Leptogenys, replaced by Leptogenys tricosa), by monotypy.

Taxonomic history

Dorylozelus in Ponerinae, Ponerini: Forel, 1917: 238.

Dorylozelus in Ponerinae, Dorylozelini: Wheeler, W.M. 1922a: 646; Donisthorpe, 1943f: 640.

Dorylozelus incertae sedis in Ponerinae, Amblyoponini: Brown, 1960a: 181.

Dorylozelus as junior synonym of Leptogenys: Taylor, 1969: 132; Bolton, 1975a: 240.

Microbolbos Donisthorpe, 1948f: 170. Type-species: Microbolbos testaceus, by original designation.

Taxonomic history

Microbolbos as junior synonym of Leptogenys: Wilson, 1955c: 136; Bolton, 1975a: 240.

Genus Leptogenys references

Roger, 1863b: 19 (Lobopelta, Leptogenys catalogues); Mayr, 1863: 426, 427 (Leptogenys, Lobopelta catalogues); Mayr, 1865: 14, 15 (Leptogenys, Lobopelta diagnoses); Mayr, 1867a: 89 (diagnosis); Mayr, 1879: 664 (Lobopelta species key); Cresson, 1887: 258 (U.S.A. catalogue); Dalla Torre, 1893: 43, 46 (Lobopelta, Leptogenys catalogues); Forel, 1900d: 304 (India & Sri Lanka species key); Bingham, 1903: 52, 55 (India, Sri Lanka & Burma Leptogenys, Lobopelta species keys); Emery, 1911d: 97 (diagnosis, subgenera key, catalogue); Arnold, 1915: 89, 95 (diagnosis, South Africa L. (Leptogenys) & L. (Lobopelta) species keys); Mann, 1919: 298 (Papuasia L. (Leptogenys) species key); Mann, 1921: 426 (Fiji L. (Lobopelta) species key); Wheeler, W.M. 1922a: 93, 786, 1010 (diagnosis, Afrotropical, Malagasy catalogues); Wheeler, W.M. 1923d: 14 (New World L. (Lobopelta) species key); Creighton, 1950a: 51 (North America L. (Lobopelta) species key); Chapman & Capco, 1951: 31 (Asia checklist); Wilson, 1958a: 113 (Melanesia & Moluccas species revision, key); Wilson, 1958a: 136 (New Caledonia species key); Kempf, 1972a: 129 (Neotropical catalogue); Alayo, 1974: 8 (Cuba species key); Bolton, 1975a: 239 (diagnosis, review of genus); Bolton, 1975a: 244 (Afrotropical species revision, key); Bolton, 1975a: 293 (Malagasy species key); Smith, D.R. 1979: 1343 (North America catalogue); Taylor & Brown, D.R. 1985: 32, 39 (Australia catalogue); Taylor, 1987a: 34, 64 (Australia, New Caledonia checklists); Trager & Johnson, 1988: 30 (U.S.A. species key); Brandão, 1991: 349 (Neotropical catalogue); Bolton, 1995a: 1050 (census); Bolton, 1995b: 229 (catalogue); Wu, J. & Wang, 1995: 41 (China species key); Shattuck, 1999: 189 (Australia synopsis); Xu, 2000a: 118 (China species key); Zhou, 2001b: 39 (China, Guangxi species key); Bolton, 2003: 163 (synopsis); Lattke, in Fernández, 2008: 145 (Colombia species, review); Terayama, 2009: 115 (Taiwan species key); Heterick, 2009: 133 (south-western Australia species key).

Ponerinae Ponerini Odontomachus TRUE TRUE FALSE

Extant: 66 species

Fossil: 2 species

Odontomachus Latreille, 1804: 179. Type-species: Formica haematoda, by monotypy.

Taxonomic history

Odontomachus in Ponérites: Lepeletier de Saint-Fargeau, 1835: 185.

Odontomachus in Poneridae: Smith, F. 1857a: 64; Smith, F. 1858b: 76.

Odontomachus in Formicidae, Odontomachidae: Mayr, 1862: 711; Mayr, 1865: 11; Smith, F. 1871a: 319.

Odontomachus in Odontomachidae: Ashmead, 1905b: 382.

Odontomachus in Ponerinae: Dalla Torre, 1893: 49.

Odontomachus in Ponerinae, Odontomachini: Forel, 1893a: 163; Emery, 1895j: 768; Forel, 1895b: 118; Forel, 1899c: 19; Wheeler, W.M. 1910g: 136; Emery, 1911d: 111; Arnold, 1915: 108; Forel, 1917: 238; Gallardo, 1918b: 93; Wheeler, W.M. 1922a: 653; Donisthorpe, 1943f: 676; Chapman & Capco, 1951: 42; Kempf, 1972a: 170; Wheeler, G.C. & Wheeler, J. 1985: 256; Dlussky & Fedoseeva, 1988: 78; Jaffe, 1993: 8.

Odontomachus in Ponerinae, Ponerini: Brown, 1976a: 70 [subtribe Odontomachiti]; Bolton, 1994: 164.

Junior synonyms of ODONTOMACHUS

Pedetes Bernstein, 1861: 7. Type-species: Pedetes macrorhynchus, by monotypy.

Taxonomic history

Pedetes as junior synonym of Odontomachus: Dalla Torre, 1893: 51; all subsequent authors.

Champsomyrmex Emery, 1892d: 558 (footnote). Type-species: Odontomachus coquereli, by monotypy.

Taxonomic history

Champsomyrmex in Ponerinae: Dalla Torre, 1893: 49.

Champsomyrmex in Odontomachidae: Ashmead, 1905b: 382.

Champsomyrmex in Ponerinae, Odontomachini: Emery, 1895j: 768; Wheeler, W.M. 1910g: 136; Emery, 1911d: 111; Wheeler, W.M. 1922a: 653; Donisthorpe, 1943f: 632.

Champsomyrmex as subgenus of Odontomachus: Forel, 1893a: 163 (misspelled as Thempsomyrmex); Forel, 1917: 238.

Champsomyrmex as genus: Emery, 1892d: 558 (footnote); Dalla Torre, 1893: 49; Emery, 1911d: 111; Wheeler, W.M. 1922a: 653.

Champsomyrmex as junior synonym of Odontomachus: Brown, 1973b: 179 [provisional]; Brown, 1976a: 96.

Myrtoteras Matsumura, 1912: 191. Type-species: Myrtoteras kuroiwae, by monotypy.

Taxonomic history

Myrtoteras in Ponerinae, Odontomachini: Donisthorpe, 1943f: 672.

Myrtoteras as junior synonym of Odontomachus: Brown, 1973b: 182 [provisional]; Brown, 1976a: 96.

Genus Odontomachus references

Roger, 1863b: 21 (catalogue); Mayr, 1863: 436 (catalogue); Mayr, 1865: 11 (diagnosis); Cresson, 1887: 258 (U.S.A. catalogue); Emery, 1892d: 559 (all species key); Dalla Torre, 1893: 49 (catalogue); Forel, 1900c: 57 (India & Sri Lanka species key); Bingham, 1903: 47 (India, Sri Lanka & Burma species key); Emery, 1911d: 111 (diagnosis, catalogue); Gallardo, 1918b: 95 (Argentina species key); Wheeler, W.M. 1922a: 99, 792, 1013 (diagnosis, Afrotropical, Malagasy catalogues); Creighton, 1950a: 55 (North America species key); Chapman & Capco, 1951: 42 (Asia checklist); Wilson, 1959a: 485 (Melanesia & Moluccas species revision, key); Kempf, 1962b: 17 (Neotropical species key); Kempf, 1972a: 170 (Neotropical catalogue); Brown, 1976a: 111, 114, 117 (New World species, Malesian & Austral species, Afrotropical & Malagasy species keys); Smith, D.R. 1979: 1344 (North America catalogue); Deyrup, Trager & Carlin, 1985: 188 (Southeastern U.S.A. species, review); Brown, 1978b: 281 (supplement to Brown, 1976a); Taylor & Brown, D.R. 1985: 36 (Australia catalogue); Taylor, 1987a: 48 (Australia, New Caledonia checklist); Wang, M. 1993: 220 (China species key); De Andrade, 1994: 24 (†Dominican Amber species key); Bolton, 1995a: 1051 (census); Bolton, 1995b: 294 (catalogue); Gronenberg, 1995: 391 (mandible morphology); Shattuck, 1999: 194 (Australia synopsis); Zhou, 2001b: 25 (China, Guangxi species key); Bolton, 2003: 164 (synopsis); Deyrup & Cover, 2004b: 141 (U.S.A. species key); Yoshimura, Onoyama & Ogata, 2007: 89 (Japan species key); Fisher & Smith, 2008: 13 (Malagasy species revision, key); Rodríguez, in Fernández, 2008: 151 (Neotropical species key); Sorger & Zettel, 2011: 143 (Philippines species key).

Ponerinae Ponerini Pachycondyla TRUE TRUE FALSE

Extant: 252 species, 62 subspecies

Fossil: 12 species

Pachycondyla Smith, F. 1858b: 105. Type-species: Formica crassinoda, by subsequent designation of Emery, 1901a: 42.

Taxonomic history

Pachycondyla in Poneridae: Smith, F. 1858b: 105; Smith, 1871a: 324.

Pachycondyla in Ponerinae: Mayr, 1862: 713 [Poneridae]; Mayr, 1865: 13 [Poneridae]; Dalla Torre, 1893: 32.

Pachycondyla in Pachycondylinae, Pachycondylini: Ashmead, 1905b: 382.

Pachycondyla in Ponerinae, Ponerini: Emery, 1895j: 767; Forel, 1895b: 114; Forel, 1899c: 10; Wheeler, W.M. 1910g: 135; Emery, 1911d: 73 [subtribe Pachycondylini]; Arnold, 1915: 53; Forel, 1917: 237; Wheeler, W.M. 1922a: 648; all subsequent authors.

Junior synonyms of PACHYCONDYLA

Bothroponera Mayr, 1862: 713 (diagnosis in key), 717. Type-species: Ponera pumicosa, by subsequent designation of Emery, 1901a: 42.

Taxonomic history

Bothroponera in Ponerinae: Mayr, 1862: 713 (in key) [Poneridae]; Mayr, 1865: 13 [Poneridae]; Dalla Torre, 1893: 35.

Bothroponera in Pachycondylinae, Pachycondylini: Ashmead, 1905b: 382.

Bothroponera in Ponerinae, Ponerini: Emery, 1895j: 767; Emery, 1911d: 75 [subtribe Pachycondylini]; Wheeler, W.M. 1922a: 648; all subsequent authors.

Bothroponera as subgenus of Ponera: Emery, 1895j: 767; Forel, 1900d: 325.

Bothroponera as subgenus of Pachycondyla: Emery, 1901a: 42; Wheeler, W.M. 1910g: 135; Emery, 1911d: 75; Arnold, 1915: 55; Forel, 1917: 237; Donisthorpe, 1943f: 628.

Bothroponera as genus: Mayr, 1862: 717; Dalla Torre, 1893: 35; Bingham, 1903: 95; Ashmead, 1905b: 382; Wheeler, W.M. 1918c: 299 (footnote); Wheeler, W.M. 1922a: 648; Wheeler, W.M. & Chapman, 1925: 67; Chapman & Capco, 1951: 49; Bernard, 1953b: 187; Taylor, 1987a: 9; Taylor & Brown, D.R. 1985: 21; Wheeler, G.C. & Wheeler, J. 1985: 256; Dlussky & Fedoseeva, 1988: 78.

Bothroponera as junior synonym of Pachycondyla: Brown, 1973b: 179 [provisional]; Snelling, R.R. 1981: 389; Hölldobler & Wilson, 1990: 11; Brown, in Bolton, 1994: 164; Bolton, 2003: 166.

Megaponera Mayr, 1862: 714 (diagnosis in key), 734. Type-species: Formica foetens (junior primary homonym in Formica, replaced by Formica analis), by monotypy.

Taxonomic history

Megaponera in Ponerinae: Dalla Torre, 1893: 30; Mayr, 1862: 714 [Poneridae]; Mayr, 1865: 15 [Poneridae].

Megaponera in Pachycondylinae, Ectatommini: Ashmead, 1905b: 382.

Megaponera in Ponerinae, Ponerini: Emery, 1895j: 767; Wheeler, W.M. 1910g: 135; Emery, 1911d: 67 [subtribe Pachycondylini]; Arnold, 1915: 46; Wheeler, W.M. 1922a: 647; all subsequent authors to the following.

Megaponera as junior synonym of Pachycondyla: Brown, in Bolton, 1994: 164; Bolton, 2003: 166.

[Megaloponera Roger, 1863b: 17, incorrect subsequent spelling; misspelling repeated by several authors, for example Emery, 1877b: 368; Forel, 1917: 237.]

Paltothyreus Mayr, 1862: 714 (diagnosis in key), 735. Type-species: Formica tarsata, by monotypy.

Taxonomic history

Paltothyreus in Ponerinae: Mayr, 1862: 714 [Poneridae]; Mayr, 1865: 15 [Poneridae]; Dalla Torre, 1893: 32.

Paltothyreus in Pachycondylinae, Ectatommini: Ashmead, 1905b: 382.

Paltothyreus in Ponerinae, Ponerini: Emery, 1895j: 767; Wheeler, W.M. 1910g: 135; Emery, 1911d: 62 [subtribe Pachycondylini]; Arnold, 1915: 43; Forel, 1917: 237; Wheeler, W.M. 1922a: 647; all subsequent authors.

Paltothyreus as junior synonym of Pachycondyla: Brown, in Bolton, 1994: 164; Bolton, 2003: 166.

Ectomomyrmex Mayr, 1867a: 83. Type-species: Ectomomyrmex javanus, by subsequent designation of Bingham, 1903: 85.

Taxonomic history

Ectomomyrmex in Ponerinae: Dalla Torre, 1893: 30.

Ectomomyrmex in Pachycondylinae, Pachycondylini: Ashmead, 1905b: 382.

Ectomomyrmex in Ponerinae, Ponerini: Emery, 1895j: 767; Forel, 1900d: 321; Emery, 1911d: 78 [subtribe Pachycondylini]; Wheeler, W.M. 1922a: 648; all subsequent authors.

Ectomomyrmex as subgenus of Pachycondyla: Emery, 1901a: 42; Wheeler, W.M. 1910g: 135; Emery, 1911d: 78; Arnold, 1915: 53; Forel, 1917: 237.

Ectomomyrmex as genus: Mayr, 1867a: 83; Dalla Torre, 1893: 30; Ashmead, 1905b: 382; Forel, 1900d: 321; Bingham, 1903: 85; Wheeler, W.M. 1922a: 648; Chapman & Capco, 1951: 60; Brown, 1963: 9; Taylor & Brown, D.R. 1985: 29; Taylor, 1987a: 26; Ogata, 1987: 112.

Ectomomyrmex as junior synonym of Pachycondyla: Brown, 1973b: 180 [provisional]; Snelling, R.R. 1981: 389; Hölldobler & Wilson, 1990: 11; Brown, in Bolton, 1994: 164; Bolton, 2003: 166.

[Ectomyrmex Donisthorpe, 1943f: 641, incorrect subsequent spelling.]

Ophthalmopone Forel, 1890b: cxii. Type-species: Ophthalmopone berthoudi, by monotypy.

Taxonomic history

Ophthalmopone in Ponerinae: Dalla Torre, 1893: 31.

Ophthalmopone in Pachycondylinae, Pachycondylini: Ashmead, 1905b: 382.

Ophthalmopone in Ponerinae, Ponerini: Emery, 1895j: 767; Wheeler, W.M. 1910g: 135; Emery, 1911d: 69 [subtribe Pachycondylini]; Arnold, 1915: 49; Forel, 1917: 237; Wheeler, W.M. 1922a: 647; all subsequent authors.

Ophthalmopone as junior synonym of Pachycondyla: Brown, in Bolton, 1994: 164; Bolton, 2003: 166.

Euponera Forel, 1891b: 126 [as subgenus of Ponera]. Type-species: Ponera (Euponera) sikorae, by monotypy.

Taxonomic history

Euponera in Ponerinae, Ponerini: Emery, 1895j: 767; Emery, 1911d: 79 [subtribe Pachycondylini]; Wheeler, W.M. 1915h: 37; Arnold, 1915: 63; Forel, 1917: 237; Wheeler, W.M. 1922a: 648; all subsequent authors.

Euponera as subgenus of Ponera: Forel, 1891b: 126; Emery, 1895j: 767.

Euponera as genus: Emery, 1901a: 46; Emery, 1909c: 364; Wheeler, W.M. 1910g: 135; Emery, 1911d: 79; Arnold, 1915: 63; Forel, 1917: 237; Gallardo, 1918b: 64; Wheeler, W.M. 1922a: 648; Borgmeier, 1923: 70; Clark, 1934b: 30; Creighton, 1950a: 44; Chapman & Capco, 1951: 63; Bernard, 1953b: 189; Wheeler, G.C. & Wheeler, J. 1985: 256; Dlussky & Fedoseeva, 1988: 78; Hölldobler & Wilson, 1990: 11.

Euponera as junior synonym of Pachycondyla: Brown, in Bolton, 1994: 164; Bolton, 2003: 166.

Cryptopone Emery, 1893a: cclxxv. Type-species: Cryptopone testacea, by monotypy.

Taxonomic history

[Cryptopone also described as new by Emery, 1893f: 240. Type-species not Amblyopone testacea, unjustified subsequent designation by Wheeler, W.M. 1911f: 161, repeated in Emery, 1911d: 88, Wheeler, W.M. 1922a: 780, Donisthorpe, 1943f: 636, Kempf, 1972a: 90 and Taylor & Brown, D.R. 1985: 28; see discussion in Wilson, 1958d: 360.]

Cryptopone in Ponerinae, Ponerini: Emery, 1895j: 767; Ashmead, 1905b: 382; Wheeler, W.M. 1910g: 135; Emery, 1911d: 88 [subtribe Ponerini]; Forel, 1917: 238; Wheeler, W.M. 1922a: 650; all subsequent authors.

Cryptopone as junior synonym of Pachycondyla: Mackay & Mackay, 2010: 3.

Pseudoponera Emery, 1900c: 314 [as subgenus of Pachycondyla]. Type-species: Ponera quadridentata (junior synonym of Formica stigma), by monotypy.

Taxonomic history

[Pseudoponera also described as new by Emery, 1901a: 42. Type-species not Ponera amblyops, unjustified subsequent designation by Emery, 1901a: 42; repeated in Wheeler, W.M. 1911f: 171, Wheeler, W.M. 1922a: 779; Donisthorpe, 1943g: 723.]

Pseudoponera in Pachycondylinae, Pachycondylini: Ashmead, 1905b: 382.

Pseudoponera in Ponerinae, Ponerini: Emery, 1911d: 86 [subtribe Pachycondylini]; Forel, 1917: 238; all subsequent authors.

Pseudoponera as subgenus of Pachycondyla: Emery, 1900c: 314; Emery, 1901a: 42.

Pseudoponera as subgenus of Euponera: Forel, 1900c: 141; Forel, 1900e: 398; Emery, 1909c: 364; Wheeler, W.M. 1910g: 135.

Pseudoponera as genus: Bingham, 1903: 91; Ashmead, 1905b: 382; Emery, 1911d: 86; Forel, 1917: 238; Wheeler, W.M. 1922a: 649; Donisthorpe, 1943g: 722; Chapman & Capco, 1951: 74.

Pseudoponera as junior synonym of Pachycondyla: Brown, 1973b: 184 [provisional]; Snelling, R.R. 1981: 389; Hölldobler & Wilson, 1990: 11; Brown, in Bolton, 1994: 164; Bolton, 2003: 167.

Brachyponera Emery, 1900c: 315 [as subgenus of Euponera]. Type-species: Euponera (Brachyponera) croceicornis, by monotypy.

Taxonomic history

[Brachyponera also described as new by Emery, 1901a: 43. Type-species not Ponera sennaarensis, unjustified subsequent designation by Emery, 1901a: 43, repeated in Wheeler, W.M. 1911f: 160, Emery, 1911d: 84, Wheeler, W.M. 1922a: 777, Donisthorpe, 1943f: 628, Wilson, 1958d: 346; Bolton, 1973a: 335; Taylor & Brown, D.R. 1985: 23.]

Brachyponera in Pachycondylinae, Pachycondylini: Ashmead, 1905b: 382.

Brachyponera in Ponerinae, Ponerini: Emery, 1911d: 83; all subsequent authors.

Brachyponera as subgenus of Euponera: Emery, 1900c: 315; Emery, 1909c: 366; Wheeler, W.M. 1910g: 135; Emery, 1911d: 83; Arnold, 1915: 72; Forel, 1917: 237; Wheeler, W.M. 1922a: 649; subsequent authors to the following.

Brachyponera as genus: Bingham, 1903: 101; Ashmead, 1905b: 382; Wilson, 1958d: 346; Taylor & Brown, D.R. 1985: 23; Wheeler, G.C. & Wheeler, J. 1985: 256; Taylor, 1987a: 9; Ogata, 1987: 116; Dlussky & Fedoseeva, 1988: 78; Hölldobler & Wilson, 1990: 10.

Brachyponera as junior synonym of Pachycondyla: Snelling, R.R. 1981: 389 [provisional]; Brown, in Bolton, 1994: 164; Bolton, 2003: 167.

Mesoponera Emery, 1900d: 668 [as subgenus of Euponera]. Type-species: Ponera melanaria, by monotypy.

Taxonomic history

[Mesoponera also described as new by Emery, 1901a: 43. Type-species not Ponera caffraria, unjustified subsequent designation by Emery, 1901a: 43, repeated by Emery, 1911d: 81, Wheeler, W.M. 1911f: 167, Wheeler, W.M. 1922a: 775, Donisthorpe, 1943f: 661, Wilson, 1958d: 349; Kempf, 1972a: 141, Bolton, 1973a: 338; Taylor & Brown, D.R. 1985: 35.]

Mesoponera in Pachycondylinae, Pachycondylini: Ashmead, 1905b: 382.

Mesoponera in Ponerinae, Ponerini: Emery, 1911d: 80; all subsequent authors.

Mesoponera as subgenus of Euponera: Emery, 1900d: 668; Wheeler, W.M. 1910g: 135; Emery, 1911d: 80; Arnold, 1915: 64; Forel, 1917: 237; Gallardo, 1918b: 65; Wheeler, W.M. 1922a: 649; Borgmeier, 1923: 71; Donisthorpe, 1943f: 661; Chapman & Capco, 1951: 64; Bernard, 1953b: 190.

Mesoponera as genus: Bingham, 1903: 99; Ashmead, 1905b: 382; Wilson, 1958d: 349; Kempf, 1972a: 141; Taylor & Brown, D.R. 1985: 34; Taylor, 1987a: 39; Dlussky & Fedoseeva, 1988: 78.

Mesoponera as junior synonym of Pachycondyla: Brown, 1973b: 182 [provisional]; Snelling, R.R. 1981: 389; Hölldobler & Wilson, 1990: 11; Brown, in Bolton, 1994: 164; Bolton, 2003: 167.

Eumecopone Forel, 1901e: 335 [as subgenus of Neoponera]. Type-species: Neoponera (Eumecopone) agilis, by monotypy.

Taxonomic history

Eumecopone as subgenus of Neoponera: Forel, 1901e: 335; Wheeler, W.M. 1910g: 135; Emery, 1911d: 71; Forel, 1917: 237; Wheeler, W.M. 1922a: 648; subsequent authors to the following.

Eumecopone junior synonym of Pachycondyla: Brown, 1973b: 180 [provisional]; Snelling, R.R. 1981: 389; Hölldobler & Wilson, 1990: 11; Brown, in Bolton, 1994: 164; Bolton, 2003: 167.

Hagensia Forel, 1901f: 333 [as subgenus of Megaloponera (sic)]. Type-species: Megaloponera (Hagensia) havilandi, by monotypy.

Taxonomic history

Hagensia in Ponerinae, Ponerini: Emery, 1911d: 69 [subtribe Pachycondylini]; all subsequent authors.

Hagensia as subgenus of Megaponera: Forel, 1901f: 333; Wheeler, W.M. 1910g: 135; Emery, 1911d: 61.

Hagensia as subgenus of Euponera: Forel, 1917: 237.

Hagensia as genus: Arnold, 1926: 202; Donisthorpe, 1943f: 648; Wheeler, G.C. & Wheeler, J. 1985: 256; Dlussky & Fedoseeva, 1988: 78; Hölldobler & Wilson, 1990: 11.

Hagensia as junior synonym of Pachycondyla: Brown, in Bolton, 1994: 164; Bolton, 2003: 167.

Neoponera Emery, 1901a: 43. Type-species: Formica villosa, by original designation.

Taxonomic history

Neoponera in Pachycondylinae, Pachycondylini: Ashmead, 1905b: 382.

Neoponera in Ponerinae, Ponerini: Wheeler, W.M. 1910g: 135; Emery, 1911d: 70 [subtribe Pachycondylini]; Forel, 1917: 237; Wheeler, W.M. 1922a: 648; all subsequent authors.

Neoponera as junior synonym of Pachycondyla: Brown, 1973b: 183 [provisional]; Snelling, R.R. 1981: 389; Hölldobler & Wilson, 1990: 11; Brown, in Bolton, 1994: 164; Bolton, 2003: 168.

Trachymesopus Emery, 1911d: 84 [as subgenus of Euponera]. Type-species: Formica stigma, by original designation.

Taxonomic history

Trachymesopus in Ponerinae, Ponerini: Emery, 1911d: 84; all subsequent authors.

Trachymesopus as subgenus of Euponera: Emery, 1911d: 84; Arnold, 1915: 74; Forel, 1917: 237; Wheeler, W.M. 1922a: 649; Borgmeier, 1923: 72; Donisthorpe, 1943g: 733; Creighton, 1950a: 46; Chapman & Capco, 1951: 65; Smith, M.R. 1951a: 786; Bernard, 1953b: 195.

Trachymesopus as genus: Wilson, 1958d: 352; Kempf, 1960f: 423; Brown, 1963: 6; Kempf, 1972a: 251; Taylor & Brown, D.R. 1985: 52; Ogata, 1987: 114; Dlussky & Fedoseeva, 1988: 78.

Trachymesopus as junior synonym of Pachycondyla: Brown, 1973b: 185 [provisional]; Snelling, R.R. 1981: 389; Hölldobler & Wilson, 1990: 11; Brown, in Bolton, 1994: 164.

Trachymesopus as junior synonym of Pseudoponera: Bolton, 1995b: 48 (because of synonymous type-species); Bolton, 2003: 168.

[Trachyponera Santschi, 1928a: 43, incorrect subsequent spelling.]

Xiphopelta Forel, 1913a: 108 [as subgenus of Ponera]. Type-species: Ponera (Xiphopelta) arnoldi, by monotypy.

Taxonomic history

Xiphopelta as subgenus of Ponera: Forel, 1913a: 108; Donisthorpe, 1943g: 737.

Xiphopelta as subgenus of Euponera: Forel, 1917: 237; Emery, 1919c: 106; Bernard, 1953b: 191.

Xiphopelta as junior synonym of Mesoponera: Wheeler, W.M. 1922a: 775; all subsequent authors to the following.

Xiphopelta as junior synonym of Pachycondyla: Brown, 1973b: 185 [provisional]; Snelling, R.R. 1981: 389; Hölldobler & Wilson, 1990: 11; Brown, in Bolton, 1994: 164; Bolton, 2003: 168.

[Hiphopelta Forel, 1913a: 108, incorrect subsequent spelling.]

Termitopone Wheeler, W.M. 1936d: 159. Type-species: Ponera laevigata, by original designation.

Taxonomic history

Termitopone in Ponerinae, Ponerini: Donisthorpe, 1943g: 731.

Termitopone as junior synonym of Pachycondyla: Brown, 1973b: 185 [provisional]; Snelling, R.R. 1981: 389; Hölldobler & Wilson, 1990: 11; Brown, in Bolton, 1994: 164; Bolton, 2003: 168.

Syntermitopone Wheeler, W.M. 1936d: 169 [as subgenus of Termitopone.] Type-species: Ponera commutata, by original designation.

Taxonomic history

Syntermitopone as subgenus of Termitopone: Wheeler, W.M. 1936d: 169.

Syntermitopone as genus: Kusnezov, 1956: 15.

Syntermitopone as junior synonym of Termitopone: Borgmeier, 1959a: 312.

Syntermitopone as junior synonym of Pachycondyla: Brown, 1973b: 185 [provisional]; Snelling, R.R. 1981: 389; Hölldobler & Wilson, 1990: 11; Brown, in Bolton, 1994: 164; Bolton, 2003: 168.

Wadeura Weber, 1939a: 102. Type-species: Wadeura guianensis, by original designation.

Taxonomic history

Wadeura in Ponerinae, Ponerini: Donisthorpe, 1943g: 735.

Wadeura as junior synonym of Pachycondyla: Brown, 1973b: 185 [provisional]; Snelling, R.R. 1981: 389; Hölldobler & Wilson, 1990: 11; Brown, in Bolton, 1994: 164; Bolton, 2003: 168.

Pseudoneoponera Donisthorpe, 1943d: 439. Type-species: Pseudoneoponera verecundae, by original designation.

Taxonomic history

Pseudoneoponera in Ponerinae, Pseudoneoponerini: Chapman & Capco, 1951: 77.

Pseudoneoponera as junior synonym of Bothroponera: Wilson, 1958d: 361.

Pseudoneoponera as junior synonym of Pachycondyla: Brown, 1973b: 184 [provisional]; Snelling, R.R. 1981: 389; Hölldobler & Wilson, 1990: 11; Brown, in Bolton, 1994: 164; Bolton, 2003: 168.

Genus Pachycondyla references

Mayr, 1862: 719 (diagnosis); Roger, 1863b: 16, 17, 18 (catalogue); Mayr, 1863: 397, 428, 439, 440 (catalogue); Mayr, 1865: 13, 15 (Bothroponera, Pachycondyla, Megaponera, Paltothyreus diagnoses); Mayr, 1867a: 82 (Bothroponera diagnosis); Mayr, 1870a: 396 (Colombia + Panama (= New Grenada) species key); Emery, 1887b: 18 (footnote) (Asia & Australia Bothroponera species key); Emery, 1890a: 71 (Neotropical species key); Dalla Torre, 1893: 30, 31, 32, 35 (Ectomomyrmex, Megaponera, Ophthalmopone, Paltothyreus, Pachycondyla, Bothroponera catalogues); Forel, 1900d: 321 (India & Sri Lanka Ectomomyrmex species key); Bingham, 1903: 86, 91, 96, 101 (India, Sri Lanka & Burma Ectomomyrmex, Pseudoponera, Bothroponera, Brachyponera species keys); Emery, 1911d: 62 (Paltothyreus diagnosis, catalogue); Emery, 1911d: 67 (Megaponera diagnosis, catalogue); Emery, 1911d: 69 (Megaponera (Hagensia) and Ophthalmopone diagnoses, catalogues); Emery, 1911d: 70 (Neoponera diagnosis, catalogue); Emery, 1911d: 73 (diagnosis, subgenera key, catalogue); Emery, 1911d: 75 (P. (Bothroponera) diagnosis, catalogue); Emery, 1911d: 78 (P. (Ectomomyrmex) diagnosis, catalogue); Emery, 1911d: 80 (Euponera diagnosis, subgenera key, catalogue); Emery, 1911d: 80 (Euponera (Mesoponera) diagnosis, catalogue); Emery, 1911d: 83 (Euponera (Brachyponera) diagnosis, catalogue); Emery, 1911d: 84 (Euponera (Trachymesopus) diagnosis, catalogue); Emery, 1911d: 86 (Pseudoponera diagnosis, catalogue); Emery, 1911d: 88 (Cryptopone diagnosis, catalogue); Arnold, 1915: 43, 46, 49, 53, 63 (Paltothyreus, Megaponera, Ophthalmopone, Pachycondyla, Euponera diagnoses); Arnold, 1915: 50, 55, 64 (South Africa Ophthalmopone, Bothroponera, Euponera (Mesoponera) species keys); Gallardo, 1918b: 56, 66 (Argentina Neoponera, Mesoponera, keys); Wheeler, W.M. 1922a: 72 (Afrotropical Bothroponera species key); Wheeler, W.M. 1922a: 60, 63, 69, 81 (Paltothyreus, Megaponera, Bothroponera, Euponera diagnoses); Wheeler, W.M. 1922a: 763, 766, 768, 769, 774, 780 (Afrotropical Paltothyreus, Megaponera, Ophthalmopone, Bothroponera, Ectomomyrmex, Euponera, Pseudoponera, Cryptopone catalogues); Wheeler, W.M. 1922a: 1007, 1008 (Malagasy Bothroponera, Euponera catalogues); Arnold, 1926: 203 (South Africa Hagensia species key); Wheeler, W.M. 1933g: 5 (Cryptopone all species key); Santschi, 1935b: 259 (Africa Mesoponera, key); Creighton, 1950a: 45 (North America Euponera species key); Arnold, 1951: 53 (Hagensia species key); Chapman & Capco, 1951: 49, 52, 60, 63, 74 (Asia Bothroponera, Cryptopone, Ectmomomyrmex, Euponera, Pseudoponera checklists); Arnold, 1952a: 461 (South Africa Bothroponera, key); Bernard, 1953b: 191 (Guinea Euponera (Xiphopelta) species key); Brown, 1958h: 20 (New Zealand species); Wilson, 1958d: 346, 357, 362 (Melanesia Brachyponera, Cryptopone, Ectomomyrmex species keys); Kempf, 1961b: 192 (Brazil species key); Brown, 1963: 6, 9 (reviews of Cryptopone, Trachymesopus & Ectomomyrmex); Kempf, 1964e: 51 (Neotropical Pachycondyla species key); Kusnezov, 1969: 36 (Argentina Neoponera species key); Kempf, 1972a: 90, 141, 160, 174, 248, 251, 257 (Neotropical Cryptopone, Mesoponera, Neoponera, Pachycondyla, Termitopone, Trachymesopus, Wadeura catalogues); Alayo, 1974: 7 (Cuba species key); Smith, D.R. 1979: 1339, 1341 (North America, Pachycondyla, Brachyponera, Cryptopone catalogues); Taylor & Brown, D.R. 1985: 21, 23, 28, 29, 34, 34, 52 (Australia, catalogue); Taylor, 1987a: 9, 23, 26, 39, 80 (Australia, New Caledonia & New Zealand checklists); Morisita, Kubota, et al. 1989: 18 (Japan Ectomomyrmex, Cryptopone, Brachyponera, Trachymesopus species key); Brandão, 1991: 354, 360, 365, 381, 382 (Neotropical catalogue); Xu, 1994b: 182 (China Brachyponera species key); Bolton, 1994: 164 (synoptic classification); Bolton, 1995a: 1051 (census); Bolton, 1995b: 166, 302 (Cryptopone, Pachycondyla catalogues); Wu, J. & Wang, 1995: 45 (China species key); Xu, 1995b: 104 (China species key); Collingwood & Agosti, 1996: 310 (Cryptopone, Pachycondyla Saudi Arabia species key); Kim, Kim & Kim, 1998: 147 (Korea species key); Shattuck, 1999: 182, 196 (Australia Cryptopone, Pachycondyla fauna synopsis); Zhou, 2001b: 35, 49 (China, Guangxi Cryptopone, Pachycondyla species keys); Dlussky & Rasnitsyn, 2002: 424 (wingless female fossils, diagnosis); Wild, 2002: 4 (Paraguay species key); Bolton, 2003: 161, 165 (Cryptopone, Pachycondyla synopses); Wild, 2005: 5 (P. apicalis complex key); Yamane, 2007: 652 (P. nigrita species group key); Mackay, Mackay, Fernández & Arias-Penna, in Fernández, 2008: 172 (Neotropical species key); Terayama, 2009: 105, 108 (Taiwan Pachycondyla, Cryptopone species keys); Heterick, 2009: 134 (south-western Australia species key); Mackay & Mackay, 2010: 26, 107 (New World Pachycondyla species revision, species complexes key and species key).

Ponerinae Ponerini Ponera TRUE TRUE FALSE

Extant: 52 species

Fossil: 5 species

Ponera Latreille, 1804: 179. Type-species: Formica coarctata, by subsequent designation of Westwood, 1840a: 83 (see discussion in Taylor, 1967a: 6).

Taxonomic history

Ponera in Ponérites: Lepeletier de Saint-Fargeau, 1835: 188.

Ponera in Poneridae: Smith, F. 1858b: 83; Smith, F. 1871a: 320.

Ponera in Ponerinae: Mayr, 1855: 386 [Poneridae]; Smith, F. 1857a: 65 [Poneridae]; Mayr, 1861: 54 [Poneridae]; Mayr, 1862: 713 [Poneridae]; Mayr, 1865: 13 [Poneridae]; Emery & Forel, 1879a: 455 [Poneridae]; Dalla Torre, 1893: 37.

Ponera in Ponerinae, Ponerini: Emery, 1895j: 767; Forel, 1899c: 15; Ashmead, 1905b: 382; Wheeler, W.M. 1910g: 135; Emery, 1911d: 88 [subtribe Ponerini]; Wheeler, W.M. 1915h: 38; Arnold, 1915: 77; Forel, 1917: 238; Wheeler, W.M. 1922a: 650; all subsequent authors.

Junior synonyms of PONERA

Pseudocryptopone Wheeler, W.M. 1933g: 12. Type-species: Cryptopone tenuis, by original designation.

Taxonomic history

Pseudocryptopone in Ponerinae, Ponerini: Donisthorpe, 1943g: 722.

Pseudocryptopone as junior synonym of Ponera: Wilson, 1957b: 356.

Selenopone Wheeler, W.M. 1933g: 19. Type-species: Ponera selenophora, by original designation.

Taxonomic history

Selenopone in Ponerinae, Ponerini: Donisthorpe, 1943g: 725.

Selenopone as junior synonym of Ponera: Wilson, 1957b: 356.

Pteroponera Bernard, 1950a: 3. Type-species: Pteroponera sysphinctoides, by monotypy.

Taxonomic history

Pteroponera as junior synonym of Ponera: Brown, 1973b: 184 [provisional]; Brown, in Bolton, 1994: 164.

Genus Ponera references

[References before Taylor, 1967a refer to both Ponera and Hypoponera.]

Mayr, 1855: 386 (diagnosis); Smith, F. 1858b: 83 (diagnosis); Mayr, 1862: 721 (diagnosis); Roger, 1863b: 16 (catalogue); Mayr, 1863: 447 (catalogue); Mayr, 1865: 13 (diagnosis); Mayr, 1867a: 85 (diagnosis); Forel, 1874: 64 (Switzerland species key); Mayr, 1879: 661 (all species key); André, 1882c: 239 (Europe & Algeria species key); Cresson, 1887: 258 (U.S.A. catalogue); Dalla Torre, 1893: 37 (catalogue); Emery, 1895b: 60 (Mediterranean species key); Emery, 1896g: 53 (New World species key); Emery, 1900c: 316 (Australia, Papuasia & Oceania species key); Forel, 1900d: 322 (India & Sri Lanka species key); Bingham, 1903: 89 (India, Sri Lanka & Burma species key); Emery, 1909c: 367 (Palaearctic species key); Emery, 1911d: 88 (diagnosis, catalogue); Arnold, 1915: 76 (diagnosis, South Africa species key); Emery, 1916b: 107 (Italy species key); Bondroit, 1918: 81 (France & Belgium species key); Gallardo, 1918b: 73 (Argentina species key); Mann, 1921: 419 (Fiji species key); Wheeler, W.M. 1922a: 780, 1009 (Afrotropical, Malagasy catalogues); Menozzi, 1931b: 266 (Costa Rica species key); Wheeler, W.M. 1933g: 13 (Pseudocryptopone species key); Smith, M.R. 1936: 420 (North America species key); Stitz, 1939: 58 (Germany species key); Creighton, 1950a: 47 (North America species key); Chapman & Capco, 1951: 68, 73, 74 (Asia Ponera, Pseudocryptopone, Selenopone checklists); Bernard, 1953b: 197 (West Africa species key); Wilson, 1957b: 359, 376 (P. tenuis, P. selenophora groups revisions, keys); Wilson, 1958d: 323, 343 (Melanesia & Moluccas species, Fiji species revisions, keys); Brown, 1958h: 22 (New Zealand species); Taylor, 1960: 180 (P. leae complex, key); Taylor, 1964: 139 (†fossil species, review); Taylor, 1967a: 18 (diagnosis, all species revision, key); Bernard, 1967: 83 (diagnosis, Western Europe species key); Smith, D.R. 1979: 1341 (North America catalogue); Taylor & Brown, D.R. 1985: 38 (Australia catalogue); Taylor, 1987a: 63 (Australia, New Caledonia & New Zealand, checklist); Morisita, Kubota, et al. 1989: 22 (Japan species key); Perrault, 1993: 337 (addition to Taylor, 1967a key); Bolton, 1994: 164 (synoptic classification); Bolton, 1995a: 1052 (census); Bolton, 1995b: 360 (catalogue); Terayama, 1996: 14 (Japan species key); Kim, Kim & Kim, 1998: 149 (Korea species key); Shattuck, 1999: 199 (Australia synopsis); Xu, 2001a: 52 and Xu, 2001c: 218 (China species keys); Zhou, 2001b: 37 (China, Guangxi species key); Bolton, 2003: 170 (synopsis); Yoshimura, Hosoishi, et al. 2009: 200 (Japan species key); Terayama, 2009: 109 (Taiwan species key).

Proceratiinae Proceratiini Discothyrea TRUE TRUE FALSE

Extant: 32 species

Fossil: 2 species

Discothyrea Roger, 1863a: 176. Type-species: Discothyrea testacea, by monotypy.

Taxonomic history

Discothyrea in Ponerinae: Mayr, 1865: 13 [Poneridae]; Dalla Torre, 1893: 18.

Discothyrea in Dorylinae, Proceratiini: Emery, 1895j: 765.

Discothyrea in Ponerinae, Proceratiini: Ashmead, 1905b: 382; Wheeler, W.M. 1910g: 136; Emery, 1911d: 51; Forel, 1917: 236; Wheeler, W.M. 1922a: 645; Donisthorpe, 1943f: 640; Chapman & Capco, 1951: 76; Kusnezov, 1964: 51; Wheeler, G.C. & Wheeler, J. 1985: 256; Jaffe, 1993: 8; Lattke, 1994: 112.

Discothyrea in Ponerinae, Ectatommini: Brown, 1958g: 248; Kempf, 1972a: 97; Smith, D.R. 1979: 1339; Dlussky & Fedoseeva, 1988: 78; Hölldobler & Wilson, 1990: 10; Bolton, 1994: 164.

Discothyrea in Proceratiinae, Proceratiini: Bolton, 2003: 49, 179.

Junior synonyms of DISCOTHYREA

Prodiscothyrea Wheeler, W.M. 1916i: 33. Type-species: Prodiscothyrea velutina, by monotypy.

Taxonomic history

Prodiscothyrea in Ponerinae, Proceratiini: Wheeler, W.M. 1916i: 36; Wheeler, W.M. 1922a: 645; Donisthorpe, 1943f: 687; Chapman & Capco, 1951: 77.

Prodiscothyrea as junior synonym of Discothyrea: Brown, 1958g: 248.

Pseudosysphincta Arnold, 1916: 161. Type-species: Pseudosysphincta poweri, by original designation.

Taxonomic history

Pseudosysphincta in Ponerinae, Proceratiini: Forel, 1917: 236; Wheeler, W.M. 1922a: 645; Donisthorpe, 1943g: 723.

Pseudosysphincta as junior synonym of Discothyrea: Brown, 1958g: 248.

[Pseudosphincta Wheeler, W.M. 1922a: 645, incorrect subsequent spelling.]

Genus Discothyrea references

Roger, 1863b: 21 (catalogue); Mayr, 1865: 13 (diagnosis); Cresson, 1887: 259 (U.S.A., catalogue); Dalla Torre, 1893: 18 (catalogue); Emery, 1911d: 51 (diagnosis, catalogue); Arnold, 1916: 159 (diagnosis); Wheeler, W.M. 1922a: 761, 762 (Afrotropical Discothyrea, Pseudosysphincta catalogues); Weber, 1939a: 99 (Neotropical species key); Chapman & Capco, 1951: 76, 77 (Asia Discothyrea, Prodiscothyrea checklists); Brown, 1958g: 248 (review of genus); Brown, 1958h: 19 (New Zealand species); Kempf, 1972a: 97 (Neotropical catalogue); Smith, D.R. 1979: 1339 (North America catalogue); Taylor & Brown, D.R. 1985: 29 (Australia catalogue); Taylor, 1987a: 24 (Australia, New Caledonia & New Zealand checklists); Morisita, Kubota, et al. 1989: 16 (Japan species key); Lattke, 1994: 113 (Neotropical species); Bolton, 1995a: 1049 (census); Bolton, 1995b: 171 (catalogue); Shattuck, 1999: 184 (Australia synopsis); Bolton, 2003: 179 (synopsis); Sosa-Calvo & Longino, 2008: 223 (Colombia species key); Yoshimura & Fisher, 2009: 9 (Malagasy males diagnosis); Terayama, 2009: 98 (Taiwan species key); Heterick, 2009: 143 (south-western Australia species key).

Proceratiinae Proceratiini Proceratium TRUE TRUE FALSE

Extant: 78 species

Fossil: 5 species

Proceratium Roger, 1863a: 171. Type-species: Proceratium silaceum, by monotypy.

Taxonomic history

Proceratium in Ponerinae: Mayr, 1865: 12 [Poneridae]; Dalla Torre, 1893: 18.

Proceratium in Ponerinae, Ponerini: Forel, 1895b: 111.

Proceratium in Dorylinae, Proceratiini: Emery, 1895j: 765.

Proceratium in Ponerinae, Ectatommini: Brown, 1958g: 241; Kempf, 1972a: 211; Smith, D.R. 1979: 1338; Dlussky & Fedoseeva, 1988: 79; Hölldobler & Wilson, 1990: 10; Bolton, 1994: 164.

Proceratium in Ponerinae, Proceratiini: Ashmead, 1905b: 382; Wheeler, W.M. 1910g: 136; Emery, 1911d: 50; Forel, 1917: 236; Wheeler, W.M. 1922a: 645; Wheeler, G.C. & Wheeler, J. 1985: 256; Jaffe, 1993: 8; Lattke, 1994: 112.

Proceratium in Proceratiinae, Proceratiini: Bolton, 2003: 49, 179.

Junior synonym of PROCERATIUM

Sysphingta Roger, 1863a: 175. Type-species: Sysphingta micrommata, by monotypy.

Taxonomic history

Sysphingta in Ponerinae: Mayr, 1865: 12 [Poneridae]; Dalla Torre, 1893: 18.

Sysphingta in Dorylinae, Proceratiini: Emery, 1895j: 765.

Sysphingta in Ponerinae, Proceratiini: Ashmead, 1905b: 382; Wheeler, W.M. 1910g: 136; Emery, 1911d: 50; Arnold, 1915: 34; Forel, 1917: 236; Wheeler, W.M. 1922a: 645; Donisthorpe, 1943g: 730.

Sysphingta as genus: Emery, 1895c: 262; Wheeler, W.M. 1910g: 136; Forel, 1917: 236; Wheeler, W.M. 1922a: 645; Creighton, 1950a: 40.

Sysphingta as junior synonym of Proceratium: Mayr, 1886d: 437; Dalla Torre, 1893: 18; Brown, in Borgmeier, 1957: 118; Brown, 1958g: 241; all subsequent authors.

[Sysphincta Mayr, 1865: 12 (and most subsequent authors), incorrect subsequent spelling.]

Genus Proceratium references

Roger, 1863b: 16, 21 (catalogue); Mayr, 1863: 451, 455 (catalogue); Mayr, 1865: 12 (Sysphingta, Proceratium diagnoses); Cresson, 1887: 258 (U.S.A. catalogue); Dalla Torre, 1893: 18 (catalogue); Emery, 1911d: 50 (diagnosis, catalogue); Emery, 1911d: 50 (Sysphingta diagnosis, catalogue); Arnold, 1915: 34 (Sysphingta diagnosis); Emery, 1916b: 103 (Italy species key); Wheeler, W.M. 1922a: 761 (Afrotropical catalogue); Kratochvíl, Novák & Snoflák, 1944: 58, 88 (West Palaearctic species key); Creighton, 1950a: 39, 41 (North America Proceratium, Sysphingta species keys); Chapman & Capco, 1951: 77 (Asia Proceratium, Sysphingta checklists); Brown, 1958g: 241 (review of genus); Bernard, 1967: 81 (diagnosis); Snelling, R.R. 1967: 8 (New World species key); Kempf, 1972a: 211 (Neotropical catalogue); Brown, 1974a: 81 (species notes); Baroni Urbani, 1977d: 91 (European species, synopsis); Smith, D.R. 1979: 1338 (North America catalogue); Brown, 1980b: 342 (New World species key); Terron, 1981: 102 (Afrotropical species notes); Taylor & Brown, D.R. 1985: 39 (Australia catalogue); Taylor, 1987a: 65 (Australia checklist); Agosti & Collingwood, 1987b: 264 (Balkans species key); Ward, 1988: 116 (New World species key); Morisita, Kubota, et al. 1989: 14 (Japan species key); Brandão, 1991: 373 (Neotropical catalogue); Lattke, 1994: 113 (phylogeny); Bolton, 1995a: 1052 (census); Bolton, 1995b: 366 (catalogue); Shattuck, 1999: 203 (Australia synopsis); Xu, 2000c: 434 (China species key); Onoyama & Yoshimura, 2002: 31 (Japan species key); Bolton, 2003: 179 (synopsis); Baroni Urbani & De Andrade, 2003b: 40 (diagnosis, all species revision, keys; Palaearctic species key: 445; Indomalayan species key: 447; Oceanean species key: 449; Afrotropical species key: 451; Nearctic and Neotropical species key: 452; world species key: 456); Xu, 2006: 153 (China species key); Sosa-Calvo & Longino, 2008: 228 (Neotropical species key); Yoshimura & Fisher, 2009: 13 (Malagasy males diagnosis); Terayama, 2009: 97 (Taiwan species key).

Cerapachyinae TRUE TRUE FALSE

Extant: 5 genera, 217 species

Fossil: 1 genus, 7 species

Cerapachysii Forel, 1893a: 162. Type-genus: Cerapachys.

Taxonomic history

Cerapachyinae as family: Bernard, 1951: 1046 [Cerapachyidae]; Bernard, 1953b: 215 [Cerapachyidae].

Cerapachyinae as junior synonym of Ponerinae: Brown, 1975: 14; Snelling, R.R. 1981: 387; Hölldobler & Wilson, 1990: 10.

Cerapachyinae as group of Dorylinae: Emery, 1901a: 36 [Cerapachinae].

Cerapachyinae as subfamily of Formicidae: Wheeler, W.M. 1902d: 185; Wheeler, W.M. 1920: 53; Wheeler, W.M. 1922a: 51, 632; Donisthorpe, 1922: xlv; Borgmeier, 1923: 50; Clark, 1924b: 76; Clark, 1934c: 49; Donisthorpe, 1943f: 620; Creighton, 1950a: 56; Clark, 1951: 15 (in key); Chapman & Capco, 1951: 17; Smith, M.R. 1951a: 781; Brown, 1954e: 26; Kusnezov, 1956: 11; Kusnezov, 1964: 48; Wheeler, G.C. & Wheeler, J. 1972a: 37; Kempf, 1972a: 263; Wheeler, G.C. & Wheeler, J. 1976b: 46; Smith, D.R. 1979: 1333; Wheeler, G.C. & Wheeler, J. 1985: 256; Ogata, 1987: 127; Bolton, 1990a: 66; Bolton, 1990c: 1356; Baroni Urbani, Bolton & Ward, 1992: 316; Jaffe, 1993: 9; Bolton, 1994: 18; Wu, J. & Wang, 1995: 47.

Cerapachyinae as dorylomorph subfamily of Formicidae: Bolton, 2003: 32, 137; Brady & Ward, 2005: 593.

Cerapachyinae as formicoid subfamily of Formicidae: Moreau, Bell, et al. 2006: 102.

Cerapachyinae as formicoid dorylomorph subfamily of Formicidae: Brady, Schultz, et al. 2006: 18173; Ward, 2007a: 555.

Tribes of Cerapachyinae: Acanthostichini, Cerapachyini, Cylindromyrmecini.

Subfamily Cerapachyinae references

Emery, 1895j: 765 (diagnosis); Wheeler, W.M. 1910g: 136 (diagnosis); Emery, 1911d: 5 (tribes key); Wheeler, W.M. 1922a: 51, 636 (diagnosis, tribes key); Morley, 1939: 114 (phylogeny); Smith, M.R. 1947f: 528 (U.S.A. diagnosis, genera); Brown, 1954e: 26 (phylogeny, notes); Eisner, 1957: 476 (proventriculus morphology); Gotwald, 1969: 43 (mouthparts morphology); Wheeler, G.C. & Wheeler, J. 1972a: 37 (diagnosis); Kempf, 1972a: 263 (Neotropical, synoptic classification); Brown, 1975: 11 (revision of tribes and genera, diagnoses); Wheeler, G.C. & Wheeler, J. 1976b: 46 (larvae, review & synthesis); Wheeler, G.C. & Wheeler, J. 1985: 261 (diagnosis); Ogata, 1987: 129 (Japan genera); Bolton, 1990a: 53 (abdominal morphology, diagnosis, synoptic classification, zoogeography); Bolton, 1990c: 1356 (diagnosis, morphology, phylogeny); Brandão, 1991: 390 (Neotropical fauna, synoptic classification, genera); Baroni Urbani, Bolton & Ward, 1992: 316 (phylogeny); Jaffe, 1993: 7 (Neotropical genera, synoptic classification); Lattke, in Jaffe, 1993: 165 (Neotropical genera); Bolton, 1994: 18 (diagnosis, synoptic classification, key to genera); Bolton, 1995a: 1038 (census); Bolton, 1995b: 10 (catalogue); Hölldobler, Obermayer & Peeters, 1996: 158 (metatibial gland); Perfil’eva, 2002: 1239 (venation); Palacio & Fernández, in Fernández, 2003d: 238 (Neotropical genera keys); Brady, 2003: 6575 (phylogeny); Bolton, 2003: 32, 137 (diagnosis, synopsis); Brady & Ward, 2005: 593 (phylogeny); Moreau, Bell, et al. 2006: 102 (phylogeny); Brady, Schultz, et al. 2006: 18173 (phylogeny); Ward, 2007a: 555 (classification).

Dolichoderinae TRUE TRUE FALSE

Extant: 28 genera, 692 species, 130 subspecies

Fossil: 16 genera, 103 species

Dolichoderidae Forel, 1878: 364. Type-genus: Dolichoderus.

Taxonomic history

Dolichoderinae as family: Emery, 1894g: 378 [Dolichoderidae]; Ashmead, 1905b: 384 [Dolichoderidae]; Novák & Sadil, 1941: 94 [Dolichoderidae]; Bernard, 1951: 1071 [Dolichoderidae]; Bernard, 1953b: 255 [Dolichoderidae].

Dolichoderinae as tribe of Formicidae: André, 1882a: 127 [Dolichoderidae].

Dolichoderinae as subfamily of Formicidae: Forel, 1878: 364 [Dolichoderidae]; Emery & Forel, 1879a: 454 [Dolichoderidae]; Nasonov, 1889: 26 [Dolichoderidae]; Forel, 1892j: 220 [Dolichoderidae]; Forel, 1893a: 165; Dalla Torre, 1893: 156; Forel, 1895b: 107 [Dolichoderidae]; Emery, 1895j: 771 [subfamily spelled Dolichoderini]; Emery, 1896e: 186; Forel, 1899c: 98; Bingham, 1903: 288; Wheeler, W.M. 1910g: 142; Emery, 1913a: 2; Arnold, 1915: 143; Wheeler, W.M. 1915h: 71; Wheeler, 1915g: 811 [Dolichoderides]; Arnold, 1915: 143; Donisthorpe, 1915d: 178; Forel, 1917: 247; Escherich, 1917: 2 [Dolichoderini]; Bondroit, 1918: 86 [Dolichoderitae]; Wheeler, W.M. 1920: 53; Wheeler, W.M. 1922a: 199; Borgmeier, 1923: 80; Karavaiev, 1936: 163; Smith, M.R. 1951a: 833; Clark, 1951: 16; Brown, 1954e: 29; Kempf, 1972a: 266; Wheeler, G.C. & Wheeler, J. 1972a: 41; Brown, 1973b: 169; all subsequent authors.

Dolichoderinae as formicomorph subfamily of Formicidae: Bolton, 2003: 18, 80.

Dolichoderinae as formicoid subfamily of Formicidae: Brady, Schultz, et al. 2006: 18173; Moreau, Bell, et al. 2006: 102.

Dolichoderinae as formicoid dolichoderomorph subfamily of Formicidae: Ward, 2007a: 556.

Tribes (extant) of Dolichoderinae: Bothriomyrmecini, Dolichoderini, Leptomyrmecini, Tapinomini.

Tribes (extinct) incertae sedis in Dolichoderinae: †Miomyrmecini, †Zherichiniini.

Genera (extinct) incertae sedis in Dolichoderinae: †Alloiomma, †Asymphylomyrmex, †Elaeomyrmex, †Elaphrodites, †Eldermyrmex, †Emplastus, †Eotapinoma, †Eurymyrmex, †Kotshkorkia, †Leptomyrmula, †Miomyrmex, †Petraeomyrmex, †Proiridomyrmex, †Protazteca, †Zherichinius.

Subfamily Dolichoderinae and tribes references, world

Forel, 1878: 364, 380 (diagnosis, genera); Dalla Torre, 1893: 156 (catalogue); Emery, 1895j: 771 (synoptic classification); Emery, 1896e: 186 (genera key); Handlirsch, 1907: 869 (†fossil taxa catalogue); Wheeler, W.M. 1910g: 142 (diagnosis); Emery, 1913a: 2, 6 (diagnosis, tribe key, catalogue); Emery, 1913a: 17 (Tapinomini diagnosis, genera key, catalogue); Arnold, 1915: 144, (diagnosis); Gallardo, 1916a: 3 (diagnosis); Forel, 1917: 247 (synoptic classification); Forel, 1921c: 136 (diagnosis); Wheeler, W.M. 1922a: 199, 687, 688 (diagnosis, tribe key, Tapinomini genera key); Brown & Nutting, 1950: 127 (venation, phylogeny); Brown, 1954e: 29 (phylogeny); Pavan, 1955: 135 (gastral organs); Eisner, 1957: 453 (proventriculus morphology); Bernard, 1967: 246 (diagnosis); Gotwald, 1969: 118 (mouthparts morphology); Wheeler, G.C. & Wheeler, J. 1972a: 41 (diagnosis); Brown, 1973b: 169 (genera & distribution); Wheeler, G.C. & Wheeler, J. 1976b: 61 (larvae, review & synthesis); Snelling, R.R. 1981: 401 (synoptic classification); Dazzini Valcurone & Fanfani, 1985: 1 (gastral glands); Wheeler, G.C. & Wheeler, J. 1985: 258 (synoptic classification); Billen, 1986: 173 (Dufour's gland); Billen, 1987: 278 (abdominal glands); Baroni Urbani & Wilson, 1987: 1 (†fossil Leptomyrmecini); Dlussky & Fedoseeva, 1988: 77 (synoptic classification); Hölldobler & Wilson, 1990: 9 (synoptic classification, genera keys); Baroni Urbani, Bolton & Ward, 1992: 316 (phylogeny); Shattuck, 1992b: 199 (higher classification, phylogeny); Shattuck, 1992c: 20 (revision of subfamily, genera key); Shattuck, 1994: 3 (catalogue); Bolton, 1994: 22 (diagnosis, synoptic classification, genera keys); Shattuck, 1995: 217 (phylogeny, genera); Bolton, 1995a: 1038 (census); Bolton, 1995b: 10 (catalogue); Wenseleers, Schoeters, et al., 1998: 121 (cloacal gland); Brandão, Baroni Urbani, et al. 1999: 411 (phylogeny, genera); Chiotis, Jermiin & Crozier, 2000: 108 (phylogeny, genera); Dlussky & Rasnitsyn, 2002: 416 (diagnosis for impression fossils); Bolton, 2003: 18, 80 (diagnosis, synopsis); Dubovikov, 2005a: 91 (tribes of Dolichoderinae); Brady, Schultz, et al. 2006: 18173 (phylogeny); Moreau, Bell, et al. 2006: 102 (phylogeny); Ward, Brady, et al. 2010: 342 (phylogeny, tribe-rank classification).

Regional and national faunas with keys

André, 1882a: 127 (Europe & Algeria); Nasonov, 1889: 50 (Russia); Forel, 1891b: 9 (Madagascar genera); Forel, 1895e: 460 (India & Sri Lanka); Bingham, 1903: 288 (India, Sri Lanka & Burma); Ruzsky, 1905b: 102 (Russian Empire); Bondroit, 1910: 488 (Belgium); Wheeler, W.M. 1910g: 560 (North America genera); Stitz, 1914: 77 (Central Europe); Gallardo, 1915: 34 (Argentina genera); Forel, 1915d: 40 (Switzerland); Arnold, 1915: 145 (South Africa); Donisthorpe, 1915d: 178 (Britain); Gallardo, 1916a: 12 (Argentina); Emery, 1916b: 208 (Italy); Wheeler, W.M. 1916m: 589 (U.S.A., Connecticut); Bondroit, 1918: 86 (France & Belgium); Gallardo, 1919b: 253 (Argentina genera); Soudek, 1922: 58 (Czechoslovakia); Stärcke, 1926: 117 (Netherlands); Karavaiev, 1927c: 271 (Ukraine); Donisthorpe, 1927b: 199 (Britain); Menozzi & Russo, 1930: 172 (Dominican Republic); Arnol'di, 1933b: 600 (Russia); Menozzi, 1933b: 90 (Israel genera); Karavaiev, 1936: 164 (Ukraine); Smith, M.R. 1937: 861 (Puerto Rico); Stitz, 1939: 209 (Germany); Kratochvíl, 1941: 94 (Central Europe); Novák & Sadil, 1941: 94 (Central Europe); Cole, 1942: 370 (U.S.A., Utah); Smith, M.R. 1943f: 309 (U.S.A. males); Buren, 1944a: 290 (U.S.A., Iowa); Smith, M.R. 1947f: 592 (U.S.A. genera); Creighton, 1950a: 330 (North America); Kusnezov, 1956: 27 (Argentina); Brown, 1958h: 29 (New Zealand); Kusnezov, 1959: 40 (Neotropical genera); Gregg, 1963: 341 (U.S.A., Colorado); Wheeler, G.C. & Wheeler, J. 1963: 149 (U.S.A., North Dakota); Bernard, 1967: 246 (Western Europe); Wilson & Taylor, 1967: 17 (Polynesia); Bolton, 1973a: 329 (West Africa genera); Bolton & Collingwood, 1975: 3 (Britain); Boven, 1970b: 24 (Netherlands); Snelling, R.R. & Hunt, 1976: 90 (Chile); Tarbinsky, 1976: 118 (Kyrghyzstan); Boven, 1977: 123 (Belgium); Kutter, 1977c: 170 (Switzerland); Arnol'di & Dlussky, 1978: 546 (former European U.S.S.R.); Collingwood, 1978: 80 (Iberian Peninsula); Collingwood, 1979: 32 (Fennoscandia & Denmark); Greenslade, 1979: 28 (South Australia genera); Schembri & Collingwood, 1981: 420 (Malta); Allred, 1982: 443 (U.S.A., Utah); Baroni Urbani, 1984: 80 (Neotropical genera); Gösswald, 1985: 314 (Germany); Collingwood, 1985: 242 (Saudi Arabia); Wheeler, G.C. & Wheeler, J. 1986g: 55 (U.S.A., Nevada); Agosti & Collingwood, 1987b: 278 (Balkans); Dlussky, Soyunov & Zabelin, 1990: 168 (Turkmenistan); Kupyanskaya, 1990: 152 (Far Eastern Russia); Morisita, Kubota, Onoyama, et al., 1991: 2 (Japan); Atanasov & Dlussky, 1992: 49 (Bulgaria); Lattke, in Jaffe, 1993: 147 (Neotropical genera); Arakelian, 1994: 70 (Armenia); Wu, J. & Wang, 1995: 115 (China genera); Kupyanskaya, 1995: 331 (Far Eastern Russia); Collingwood & Agosti, 1996: 359 (Saudi Arabia); Seifert, 1996b: 161 (Central Europe); Skinner & Allen, 1996: 40 (Britain); Collingwood & Prince, 1998: 20 (Portugal); Kim & Kim, 1999: 18 (Korea); Shattuck, 1999: 26, 64 (Australia genera, synopsis); Andersen, 2000: 59 (northern Australia genera); Zhou, 2001b: 149 (China, Guangxi); Czechowski, Radchenko & Czechowska, 2002: 134 (Poland); Aktaç & Radchenko, 2002: 54 (Turkey genera); Yoshimura & Onoyama, 2002: 425 (Japan genera, males); Mackay & Mackay, 2002: 221 (U.S.A., New Mexico); Palacio & Fernández, in Fernández, 2003d: 239 (Neotropical genera); Radchenko, 2005b: 186 (North Korea); Coovert, 2005: 102 (U.S.A., Ohio); Clouse, 2007b: 190 (Micronesia); Seifert, 2007: 147 (North and Central Europe); Fisher, 2009: 51 (Malagasy genera); Terayama, 2009: 196 (Taiwan); Heterick, 2009: 28 (south-western Australia genera); Boer, 2010: 14 (Benelux); Yoshimura & Fisher, 2011: 7 (Malagasy males key).

Ecitoninae TRUE TRUE FALSE

Extant: 5 genera, 149 species, 21 subspecies

Fossil: 1 species

Ecitonii Forel, 1893a: 163. Type-genus: Eciton.

Taxonomic history

Ecitoninae as subfamily of Dorylidae: Ashmead, 1905b: 381; Ashmead, 1906: 23.

Ecitoninae as subfamily of Formicidae: Brown, 1973b: 166; Snelling, R.R. 1981: 392; Bolton, 1990c: 1357; Hölldobler & Wilson, 1990: 11; all subsequent authors.

Ecitoninae as dorylomorph subfamily of Formicidae: Bolton, 2003: 34, 142; Brady & Ward, 2005: 593.

Ecitoninae as formicoid subfamily of Formicidae: Moreau, Bell, et al. 2006: 102.

Ecitoninae as formicoid dorylomorph subfamily of Formicidae: Brady, Schultz, et al. 2006: 18173; Ward, 2007a: 555.

Tribes of Ecitoninae: Cheliomyrmecini, Ecitonini.

Subfamily and tribe Ecitonini references

Emery, 1895j: 765 (diagnosis); Ashmead, 1906: 23, 24 (tribes & genera keys); Emery, 1910b: 15 (diagnosis, genera key, catalogue); Wheeler, W.M. 1910g: 138, 558 (diagnosis, North America genera); Forel, 1917: 240 (synoptic classification); Gallardo, 1920: 312 (Argentina genera, key); Wheeler, W.M. 1922a: 634 (genera key); Borgmeier, 1923: 37 (Brazil catalogue); Smith, M.R. 1943f: 290 (U.S.A. males); Creighton, 1950a: 61 (North America); Borgmeier, 1955: 51, 57 (revision of subfamily, tribes key); Borgmeier, 1955: 79 (revision of tribe, genera key); Brown, 1973b: 166 (genera, distribution); Watkins, 1976: 6 (genera keys); Wheeler, G.C. & Wheeler, J. 1976b: 46 (larvae, review & synthesis); Snelling, R.R. 1981: 392 (synoptic classification); Gotwald & Burdette, 1981: 78 (phylogeny); Watkins, 1982: 210 (Mexico genera, key); Gotwald, 1982: 167 (genera key); Baroni Urbani, 1984: 74 (genera key); Wheeler, G.C. & Wheeler, J. 1986g: 17 (U.S.A., Nevada); Bolton, 1990c: 1357 (diagnosis, morphology, phylogeny); Brandão, 1991: 392 (Neotropical fauna, synoptic classification); Baroni Urbani, Bolton & Ward, 1992: 317 (phylogeny); Jaffe, 1993: 13 (Neotropical genera, synoptic classification); Lattke, in Jaffe, 1993: 149 (genera key); Bolton, 1994: 38 (diagnosis, synoptic classification, genera key); Bolton, 1995a: 1039 (census); Bolton, 1995b: 11 (catalogue); Hölldobler, Obermayer & Peeters, 1996: 158 (metatibial gland); Palacio, 1999: 143 (Colombia genera, key); Perfil’eva, 2002: 1239 (venation); Palacio & Fernández, in Fernández, 2003d: 240 (Neotropical genera key); Brady, 2003: 6575 (phylogeny); Bolton, 2003: 34, 142 (diagnosis, synopsis); Brady & Ward, 2005: 593 (phylogeny); Moreau, Bell, et al. 2006: 102 (phylogeny); Brady, Schultz, et al. 2006: 18173 (phylogeny); Ward, 2007a: 555 (classification).

Formicinae TRUE TRUE FALSE

Extant: 49 genera, 2,831 species, 839 subspecies

Fossil: 29 genera, 193 species, 1 subspecies

Formicariae Latreille, 1809: 124. Type-genus: Formica.

Taxonomic history

Formicinae as group name: Latreille, 1809: 124 [Formicariae]; Lepeletier de Saint-Fargeau, 1835: 197 [Formicites]; Nylander, 1846a: 877 [Formicae].

Formicinae as family: Smith, F. 1858b: 1 [Formicidae]; Smith, F. 1861b: 36 [Formicidae]; Smith, 1871a: 302 [Formicidae]; André, 1882a: 125 [Formicidae]; Saunders, 1896: 18 [Formicidae]; Novák & Sadil, 1941: 97 [Formicidae]; Bernard, 1951: 1073 [Formicidae]; Bernard, 1953b: 256 [Formicidae].

Formicinae as tribe of Formicidae: André, 1874: 167 [Formicidae].

Formicinae as subfamily of Formicidae: Mayr, 1855: 286, 299 [Formicidae]; Mayr, 1861: 21 [Formicidae]; Mayr, 1862: 651 [Formicidae]; Mayr, 1865: 6 [Formicidae]; Mayr, 1868c: 24 [Formicidae]; Forel, 1870: 307 [Formicidae]; Forel, 1874: 21 [Formicidae]; Emery, 1877a: 70 [Formicidae]; André, 1881b: 54 [Formicidae]; Ashmead, 1905b: 384; Bondroit, 1918: 17 [Formicitae]; Wheeler, W.M. 1920: 53; Wheeler, W.M. 1922a: 210; Emery, 1925b: 2; Karavaiev, 1936: 172; Clark, 1951: 16; Brown, 1954e: 29; Kempf, 1972a: 266; Brown, 1973b: 169; all subsequent authors.

Formicinae as formicomorph subfamily of Formicidae: Bolton, 2003: 20, 93.

Formicinae as formicoid subfamily of Formicidae: Brady, Schultz, et al. 2006: 18173; Moreau, Bell, et al. 2006: 102.

Formicinae as formicoid formicomorph subfamily of Formicidae: Ward, 2007a: 556.

Tribes of Formicinae: Camponotini, Formicini, Gesomyrmecini, Gigantiopini, Lasiini, Melophorini, Myrmecorhynchini, Myrmoteratini, Notostigmatini, Oecophyllini, Plagiolepidini.

Genera (extinct) incertae sedis in Formicinae: †Camponotites, †Drymomyrmex, †Imhoffia, †Kyromyrma, †Leucotaphus, †Protrechina, †Zhangidris.

Hong (2002) genera (extinct) incertae sedis in Formicinae: †Curtipalpulus, †Eoleptocerites, †Eurytarsites, †Fushuniformica, †Huaxiaformica, †Leptogasteritus, †Liaoformica, †Longiformica, †Magnogasterites, †Orbicapitia, †Ovalicapito, †Ovaligastrula, †Sinoformica, †Sinotenuicapito, †Wilsonia (unresolved junior homonym).

Subfamily Formicinae references, world

Mayr, 1862: 651 (genera key); Mayr, 1865: 6 (diagnosis); Handlirsch, 1907: 859 (†fossil taxa catalogue); Dalla Torre, 1893: 171 (catalogue); Emery, 1895j: 772 (synoptic classification); Emery, 1896e: 187 (genera key); Wheeler, W.M. 1910g: 143 (diagnosis); Forel, 1912i: 88 (tribes key); Forel, 1917: 248 (synoptic classification); Arnold, 1920a: 551 (diagnosis); Forel, 1921c: 139 (diagnosis); Wheeler, W.M. 1922a: 210, 691 (diagnosis, tribes key); Emery, 1925b: 2 (diagnosis, tribe key, catalogue); Brown & Nutting, 1950: 127 (venation, phylogeny); Eisner, 1957: 465 (proventriculus morphology); Hung & Brown, 1966: 198 (gastric apex, structure); Bernard, 1967: 267 (diagnosis); Gotwald, 1969: 120 (mouthparts morphology); Wheeler, G.C. & Wheeler, J. 1972a: 41 (diagnosis); Brown, 1973b: 169 (genera, distribution); Wheeler, G.C. & Wheeler, J. 1976b: 62 (larvae, review & synthesis); Snelling, R.R. 1981: 402 (synoptic classification); Wheeler, G.C. & Wheeler, J. 1985: 258 (synoptic classification); Billen, 1986: 173 (Dufour's gland); Dlussky & Fedoseeva, 1988: 77 (synoptic classification); Hölldobler & Wilson, 1990: 9 (synoptic classification, genera keys); Agosti, 1991: 295 (genus group diagnoses); Shattuck, 1992b: 201 (phylogeny); Baroni Urbani, Bolton & Ward, 1992: 317 (phylogeny); Bolton, 1994: 42 (diagnosis, synoptic classification, genera keys); Bolton, 1995a: 1039 (census); Bolton, 1995b: 11 (catalogue); Wenseleers, Schoeters, et al. 1998: 121 (cloacal gland); Dlussky & Rasnitsyn, 2002: 417 (diagnosis for impression fossils); Bolton, 2003: 20, 93 (diagnosis, synopsis); Brady, Schultz, et al. 2006: 18173 (phylogeny); Moreau, Bell, et al. 2006: 102 (phylogeny).

Regional and national faunas with keys

Mayr, 1855: 299 (Austria); Mayr, 1861: 25 (Europe); Mayr, 1868c: 25 (†Baltic Amber); André, 1874: 167 (Europe); Forel, 1874: 22 (Switzerland); Saunders, E. 1880: 203 (Britain); André, 1882a: 126 (Europe & Algeria); Provancher, 1887: 225 (Canada); Cresson, 1887: 94 (U.S.A. genera); Nasonov, 1889: 50 (Russia); Forel, 1891b: 8 (Madagascar genera); Lameere, 1892: 62 (Belgium); Forel, 1892j: 220 (India & Sri Lanka); Bingham, 1903: 308 (India, Sri Lanka & Burma); Ruzsky, 1905b: 100 (Russian Empire); Wasmann, 1906: 7 (Luxemburg); Bondroit, 1910: 481 (Belgium); Wheeler, W.M. 1910g: 560 (North America genera); Stitz, 1914: 80 (Central Europe); Gallardo, 1915: 35 (Argentina genera); Forel, 1915d: 45 (Switzerland); Donisthorpe, 1915d: 184 (Britain); Emery, 1916b: 216 (Italy); Wheeler, W.M. 1916m: 590 (U.S.A., Connecticut); Bondroit, 1918: 17 (France & Belgium); Arnold, 1920a: 552 (South Africa); Kutter, 1920b: 134 (Switzerland); Soudek, 1922: 61 (Czechoslovakia); Lomnicki, 1925a: 160 (Poland); Stärcke, 1926: 118, 146 (Netherlands); Karavaiev, 1927c: 273 (Ukraine); Donisthorpe, 1927b: 205 (Britain); Menozzi & Russo, 1930: 172 (Dominican Republic); Arnol'di, 1933b: 601 (Russia); Menozzi, 1933b: 90 (Israel genera); Karavaiev, 1936: 173 (Ukraine); Smith, M.R. 1937: 865 (Puerto Rico); Stitz, 1939: 230 (Germany); Kratochvíl, 1941: 97 (Central Europe); Novák & Sadil, 1941: 97 (Central Europe); Cole, 1942: 373 (U.S.A., Utah); Smith, M.R. 1943f: 309 (U.S.A., males); Buren, 1944a: 292 (U.S.A., Iowa); Holgersen, 1943b: 173 (Norway); Holgersen, 1944: 199 (Norway); Smith, M.R. 1947f: 599 (U.S.A. genera); Boven, 1947: 181 (Belgium); Creighton, 1950a: 355 (North America); Kusnezov, 1956: 31 (Argentina); Brown, 1958h: 42 (New Zealand); Boven, 1959: 11 (Netherlands); Gregg, 1963: 447 (U.S.A., Colorado); Wheeler, G.C. & Wheeler, J. 1963: 160 (U.S.A., North Dakota); Collingwood, 1964: 104 (Britain); Bernard, 1967: 268 (Western Europe); Wilson & Taylor, 1967: 17 (Polynesia); Boven, 1970b: 26 (Netherlands); Kempf, 1972a: 266 (Neotropical, synoptic classification); Bolton, 1973a: 329 (West Africa genera); Bolton & Collingwood, 1975: 3 (Britain); Snelling & Hunt, 1976: 104 (Chile); Tarbinsky, 1976: 126 (Kyrghyzstan); Boven, 1977: 126 (Belgium); Kutter, 1977c: 183 (Switzerland); Arnol'di & Dlussky, 1978: 548 (former European U.S.S.R.); Collingwood, 1978: 88 (Iberian Peninsula); Collingwood, 1979: 85 (Fennoscandia & Denmark); Greenslade, 1979: 32 (South Australia genera); Schembri & Collingwood, 1981: 436 (Malta); Prins, 1983: 8 (Southern Africa genera); Allred, 1982: 444 (U.S.A., Utah); Verhaeghe, Deligne, et al., 1984: 106 (Belgium genera); Baroni Urbani, 1984: 81 (Neotropical genera); Gösswald, 1985: 263 (Germany); Collingwood, 1985: 273 (Saudi Arabia); Wheeler, G.C. & Wheeler, J. 1986g: 58 (U.S.A., Nevada); Nilsson & Douwes, 1987: 68 (Norway); Agosti & Collingwood, 1987b: 279 (Balkans); Dlussky, Soyunov & Zabelin, 1990: 124 (Turkmenistan); Kupyanskaya, 1990: 162 (Far Eastern Russia); Morisita, Kubota, Onoyama, et al., 1991: 10 (Japan); Atanasov & Dlussky, 1992: 49 (Bulgaria); Shattuck, 1992b: 199 (higher classification, phylogeny); Lattke, in Jaffe, 1993: 150 (Neotropical genera); Arakelian, 1994: 76 (Armenia); Wu, J. & Wang, 1995: 125 (China genera); Kupyanskaya, 1995: 332 (Far Eastern Russia); Collingwood & Agosti, 1996: 361 (Saudi Arabia); Seifert, 1996b: 166 (Central Europe); Skinner & Allen, 1996: 41 (Britain); Collingwood & Prince, 1998: 21 (Portugal); Shattuck, 1999: 25, 86 (Australia genera, synopsis); Andersen, 2000: 68 (northern Australia genera); Zhou, 2001b: 165 (China, Guangxi); Czechowski, Radchenko & Czechowska, 2002: 147 (Poland); Aktaç & Radchenko, 2002: 54 (Turkey genera); Yoshimura & Onoyama, 2002: 425 (Japan genera, males); Mackay & Mackay, 2002: 236 (U.S.A., New Mexico); Palacio & Fernández, in Fernández, 2003d: 242 (Neotropical genera); Radchenko, 2005b: 187 (North Korea); Coovert, 2005: 113 (U.S.A., Ohio); Clouse, 2007b: 190 (Micronesia); Seifert, 2007: 150 (North and Central Europe); Terayama, 2009: 202 (Taiwan); Heterick, 2009: 30 (south-western Australia genera); Boer, 2010: 17 (Benelux).

Myrmicinae TRUE TRUE FALSE

Extant: 141 genera, 6,087 species, 864 subspecies

Fossil: 31 genera, 130 species

Myrmicites Lepeletier de Saint-Fargeau, 1835: 169. Type-genus: Myrmica.

Taxonomic history

Myrmicinae as group name: Lepeletier de Saint-Fargeau, 1835: 169 [Myrmicites]; Nylander, 1846a: 877 [Myrmicae].

Myrmicinae as family: Smith, F. 1851: 4 [Myrmicidae]; Smith, F. 1861b: 45 [Myrmicidae]; Smith, F. 1871a: 324 [Myrmicidae]; André, 1882a: 125 [Myrmicidae]; Cresson, 1887: 93 [Myrmicidae]; Emery, 1894g: 383 [Myrmicidae]; Saunders, 1896: 18 [Myrmicidae]; Ashmead, 1905b: 383 [Myrmicidae]; Novák & Sadil, 1941: 71 [Myrmicidae]; Bernard, 1951: 1058 [Myrmicidae]; Bernard, 1953b: 222 [Myrmicidae].

Myrmicinae as subfamily of Poneridae: Smith, F. 1858b: 114 [Myrmicidae].

Myrmicinae as tribe of Formicidae: André, 1874: 167 [Myrmicidae].

Myrmicinae as subfamily of Myrmicidae: Ashmead, 1905b: 383.

Myrmicinae as subfamily of Formicidae: Mayr, 1855: 290, 299 [Myrmicidae]; Smith, F. 1857a: 70 [Myrmicidae]; Mayr, 1861: 21 [Myrmicidae]; Smith, F. 1862b: 33 [Myrmicidae]; Mayr, 1862: 738 [Myrmicidae]; Mayr, 1865: 17 [Myrmicidae]; Mayr, 1868c: 24 [Myrmicidae]; Forel, 1870: 307 [Myrmicidae]; Forel, 1874: 22 [Myrmicidae]; Emery, 1877a: 70 [Myrmicidae]; Forel, 1878: 367 [Myrmicidae]; Emery & Forel, 1879a: 456 [Myrmicidae]; André, 1881b: 64 [Myrmicidae]; Nasonov, 1889: 28 [Myrmicidae]; Forel, 1891b: 11 [Myrmicidae]; Forel, 1892j: 220 [Myrmicidae]; Forel, 1893a: 163; Dalla Torre, 1893: 53; Emery, 1895j: 768 [subfamily spelled Myrmicini]; Emery, 1896e: 179; Forel, 1899c: 30; Forel, 1902f: 520; Bingham, 1903: 105; Wheeler, W.M. 1910g: 138; Emery, 1914a: 29; Wheeler, 1915g: 806 [Myrmicides]; Wheeler, W.M. 1915h: 40; Donisthorpe, 1915d: 74; Arnold, 1916: 166; Escherich, 1917: 2 [Myrmicini]; Forel, 1917: 240 [subfamily spelled Myrmicini]; Bondroit, 1918: 14 [Myrmicitae]; Wheeler, W.M. 1920: 53; Wheeler, W.M. 1922a: 124; Emery, 1921f: 3; Karavaiev, 1934: 59; Clark, 1951: 16; Brown, 1954e: 28; Wheeler, G.C. & Wheeler, J. 1972a: 40; Brown, 1973b: 166; all subsequent authors.

Myrmicinae as myrmicomorph subfamily of Formicidae: Bolton, 2003: 52, 182.

Myrmicinae as formicoid subfamily of Formicidae: Moreau, Bell, et al. 2006: 102; Brady, Schultz, et al. 2006: 18173.

Myrmicinae as formicoid myrmicomorph subfamily of Formicidae: Ward, 2007a: 556.

Tribes of Myrmicinae: Adelomyrmecini, Ankylomyrmini, Attini, Basicerotini, Blepharidattini, Cataulacini, Cephalotini, Crematogastrini, Dacetini, Formicoxenini, Lenomyrmecini, Liomyrmecini, Melissotarsini, Meranoplini, Metaponini, Myrmecinini, Myrmicariini, Myrmicini, Paratopulini, Phalacromyrmecini, Pheidolini, Solenopsidini, Stegomyrmecini, Stenammini, Tetramoriini.

Genera (extant) incertae sedis in Myrmicinae: Diaphoromyrma, Tyrannomyrmex.

Genera (extinct) incertae sedis in Myrmicinae: †Afromyrma, †Attopsis, †Cephalomyrmex, †Electromyrmex, †Eocenidris, †Eomyrmex, †Fallomyrma, †Miosolenopsis, †Myrmecites, †Solenopsites.

Hong (2002) genera (extinct) incertae sedis in Myrmicinae: †Brachytarsites, †Clavipetiola, †Fushunomyrmex, †Orbigastrula, †Quadrulicapito, †Quineangulicapito, †Sinomyrmex, †Sphaerogasterites, †Wumyrmex.

Collective group name in Myrmicinae: †Myrmicites.

Subfamily Myrmicinae references, world

Mayr, 1865: 17 (diagnosis); Mayr, 1867a: 91 (diagnosis); Forel, 1878: 367 (diagnosis); Handlirsch, 1907: 872 (†fossil taxa catalogue); Dalla Torre, 1893: 53 (catalogue); Emery, 1895j: 768 (diagnosis); Emery, 1896e: 179 (genera key); Wheeler, W.M. 1910g: 138 (diagnosis); Emery, 1912b: 101 (phylogeny); Emery, 1914a: 34 (phylogeny, tribe key); Arnold, 1916: 164 (diagnosis); Forel, 1917: 240 (synoptic classification); Forel, 1921c: 139 (diagnosis); Emery, 1921f: 3 (diagnosis, tribes & genera key, catalogue); Wheeler, W.M. 1922a: 124, 655 (diagnosis, tribes key); Brown & Nutting, 1950: 126 (venation, phylogeny); Brown, 1954e: 28 (phylogeny); Eisner, 1957: 477 (proventriculus morphology); Bernard, 1967: 93 (diagnosis); Gotwald, 1969: 99 (mouthparts morphology); Wheeler, G.C. & Wheeler, J. 1972a: 40 (diagnosis); Brown, 1973b: 166 (genera, distribution); Wheeler, G.C. & Wheeler, J. 1976b: 52 (larvae, review & synthesis); Kugler, C. 1978a: 413 (sting structure); Kugler, C. 1978b: 267 (pygidial glands); Kugler, C. 1979c: 117 (sting, evolution); Snelling, R.R. 1981: 393 (synoptic classification); Caetano, F.H. 1984: 257 (digestive tract, morphology); Wheeler, G.C. & Wheeler, J. 1985: 257 (synoptic classification); Billen, 1986: 167 (Dufour's gland); Dlussky & Fedoseeva, 1988: 79 (synoptic classification); Hölldobler & Wilson, 1990: 9 onward (synoptic classification, genera keys); Baroni Urbani, Bolton & Ward, 1992: 317 (phylogeny); Bolton, 1994: 75 (diagnosis, synoptic classification, genera keys); Bolton, 1995a: 1040 (census); Bolton, 1995b: 13 (catalogue); Hashimoto, 1996: 354 (phylogenetic position); Baroni Urbani, 2000: 480 (phylogeny); Dlussky & Rasnitsyn, 2002: 422 (diagnosis for wingless fossils); Bolton, 2003: 52, 182 (diagnosis, synopsis); Moreau, Bell, et al. 2006: 102 (phylogeny); Brady, Schultz, et al. 2006: 18173 (phylogeny); Ward, 2007a: 556 (classification).

Regional and national faunas with keys

Mayr, 1855: 391 (Austria); Mayr, 1861: 29 (Europe); Mayr, 1868c: 79 (†Baltic Amber); André, 1874: 171 (Europe); Forel, 1874: 29 (Switzerland); Saunders, E. 1880: 213 (Britain); André, 1882c: 256 (Europe & Algeria); Cresson, 1887: 98 (U.S.A. genera); Provancher, 1887: 243 (Canada); Nasonov, 1889: 54 (Russia); Forel, 1891b: 11 (Madagascar genera); Lameere, 1892: 66 (Belgium); Forel, 1902f: 520 (India & Sri Lanka genera); Bingham, 1903: 105 (India, Sri Lanka & Burma); Ruzsky, 1905b: 103 (Russian Empire); Wasmann, 1906: 13 (Luxemburg); Bondroit, 1910: 490 (Belgium); Wheeler, W.M. 1910g: 558 (North America genera); Stitz, 1914: 55 (Central Europe); Gallardo, 1915: 32 (Argentina genera); Forel, 1915d: 8 (Switzerland); Donisthorpe, 1915d: 74 (Britain); Arnold, 1916: 166, 170 (South Africa tribes, genera); Emery, 1916b: 112 (Italy); Wheeler, W.M. 1916m: 581 (U.S.A., Connecticut); Bondroit, 1918: 90 (France & Belgium); Kutter, 1920b: 144 (Switzerland); Soudek, 1922: 20 (Czechoslovakia); Stärcke, 1926: 84 (Netherlands); Karavaiev, 1927c: 256 (Ukraine); Donisthorpe, 1927b: 77 (Britain); Menozzi & Russo, 1930: 170 (Dominican Republic); Gallardo, 1932b: 91 (Argentina, tribes); Arnol'di, 1933b: 596 (Russia); Menozzi, 1933b: 88 (Israel genera); Karavaiev, 1934: 60 (Ukraine); Smith, M.R. 1937: 829 (Puerto Rico); Stitz, 1939: 63 (Germany); Kratochvíl, 1941: 71 (Central Europe); Novák & Sadil, 1941: 71 (Central Europe); Cole, 1942: 360 (U.S.A., Utah); Smith, M.R. 1943f: 291 (U.S.A., males); Holgersen, 1943b: 166 (Norway); Holgersen, 1944: 198 (Norway); Buren, 1944a: 281 (U.S.A., Iowa); Smith, M.R. 1947f: 543 (U.S.A. genera); Boven, 1947: 170 (Belgium); Creighton, 1950a: 83 (North America); Kusnezov, 1956: 15 (Argentina); Brown, 1958h: 25 (New Zealand); Boven, 1959: 7 (Netherlands); Gregg, 1963: 288 (U.S.A., Colorado); Wheeler, G.C. & Wheeler, J. 1963: 92 (U.S.A., North Dakota); Collingwood, 1964: 94 (Britain); Bernard, 1967: 95 (Western Europe); Wilson & Taylor, 1967: 13 (Polynesia); Boven, 1970b: 9 (Netherlands); Kempf, 1972a: 263 (Neotropical, synoptic classification); Bolton, 1973a: 325 (West Africa genera); Bolton & Collingwood, 1975: 3 (Britain); Snelling, R.R. & Hunt, 1976: 70 (Chile); Tarbinsky, 1976: 19 (Kyrghyzstan); Boven, 1977: 69 (Belgium); Kutter, 1977c: 31 (Switzerland); Arnol'di & Dlussky, 1978: 524 (former European U.S.S.R.); Collingwood, 1978: 75 (Iberian Peninsula); Collingwood, 1979: 36 (Fennoscandia & Denmark); Greenslade, 1979: 20 (South Australia genera); Schembri & Collingwood, 1981: 423 (Malta); Allred, 1982: 438 (U.S.A., Utah); Baroni Urbani, 1984: 76 (Neotropical genera); Verhaeghe, Deligne, et al. 1984: 112 (Belgium genera); Gösswald, 1985: 289 (Germany); Collingwood, 1985: 245 (Saudi Arabia); Wheeler, G.C. & Wheeler, J. 1986g: 20 (U.S.A., Nevada); Nilsson & Douwes, 1987: 57 (Norway); Agosti & Collingwood, 1987b: 265 (Balkans); Dlussky, Soyunov & Zabelin, 1990: 181 (Turkmenistan); Kupyanskaya, 1990: 89 (Far Eastern Russia); Ogata, 1991b: 61 (Japan genera); Morisita, Kubota, et al. 1992: 1 (Japan); Atanasov & Dlussky, 1992: 51 (Bulgaria); Lattke, in Jaffe, 1993: 153 (Neotropical genera); Arakelian, 1994: 15 (Armenia); Wu, J. & Wang, 1995: 57 (China genera); Kupyanskaya, 1995: 327 (Far Eastern Russia); Collingwood & Agosti, 1996: 308 (Saudi Arabia); Seifert, 1996b: 108 (Central Europe); Skinner & Allen, 1996: 43 (Britain); Collingwood & Prince, 1998: 10 (Portugal); Shattuck, 1999: 39, 122 (Australia genera, synopsis); Andersen, 2000: 37 (northern Australia genera); Zhou, 2001b: 69 (China, Guangxi); Czechowski, Radchenko & Czechowska, 2002: 135 (Poland); Aktaç & Radchenko, 2002: 55 (Turkey genera); Yoshimura & Onoyama, 2002: 424 (Japan genera, males); Mackay & Mackay, 2002: 58 (U.S.A., New Mexico); Palacio & Fernández, in Fernández, 2003d: 244 (Neotropical genera and synopsis); Coovert, 2005: 32 (U.S.A., Ohio); Radchenko, 2005b: 184 (North Korea); Clouse, 2007b: 190 (Micronesia); Seifert, 2007: 110 (North and Central Europe); Terayama, 2009: 131 (Taiwan); Heterick, 2009: 36 (south-western Australia genera); Boer, 2010: 46 (Benelux); Eguchi, Bui & Yamane, 2011: 8 (Vietnam genera).

Ponerinae TRUE TRUE FALSE

Extant: 27 genera, 1,033 species, 155 subspecies

Fossil: 9 genera, 68 species, 2 subspecies

Ponerites Lepeletier de Saint-Fargeau, 1835: 185. Type-genus: Ponera.

Taxonomic history

Ponerinae as group name: Lepeletier de Saint-Fargeau, 1835: 185 [Ponérites].

Ponerinae as family: Smith, F. 1851: 6 [Poneridae]; Smith, F. 1858b: 76 [Poneridae]; Smith, F. 1861b: 44 [Poneridae]; Smith, F. 1871a: 320 [Poneridae]; André, 1882a: 125 [Poneridae]; Cresson, 1887: 93 [Poneridae]; Emery, 1894g: 379 [Poneridae]; Saunders, 1896: 18 [Poneridae]; Ashmead, 1905b: 382 [Poneridae]; Novák & Sadil, 1941: 70 [Poneridae]; Bernard, 1951: 1042 [Poneridae]; Bernard, 1953b: 185 [Poneridae].

Ponerinae as tribe of Formicidae: André, 1874: 167 [Poneridae].

Ponerinae as subfamily of Poneridae: Ashmead, 1905b: 382.

Ponerinae as subfamily of Formicidae: Mayr, 1855: 289, 299 [Poneridae]; Mayr, 1861: 21 [Poneridae]; Mayr, 1862: 712 [Poneridae]; Smith, F. 1857a: 64 [Poneridae]; Smith, F. 1862b: 31 [Poneridae]; Mayr, 1865: 11 [Poneridae]; Mayr, 1868c: 24 [Poneridae]; Forel, 1870: 307 [Poneridae]; Forel, 1874: 21 [Poneridae]; Emery, 1877a: 70 [Poneridae]; Forel, 1878: 366 [Poneridae]; Emery & Forel, 1879a: 455 [Poneridae]; André, 1881b: 64 [Poneridae]; Forel, 1892j: 220 [Poneridae]; Dalla Torre, 1893: 13; Forel, 1893a: 162; Forel, 1895b: 111 [Poneridae]; Nasonov, 1889: 27 [Poneridae]; Emery, 1895j: 766 [subfamily spelled Ponerini]; Emery, 1896e: 176; Forel, 1899c: 2; Emery, 1901a: 36; Bingham, 1903: 23; Wheeler, W.M. 1910g: 134; Emery, 1911d: 2; Wheeler, 1915g: 805 [Ponerides]; Donisthorpe, 1915d: 65; Wheeler, W.M. 1915h: 25; Arnold, 1915: 9; Escherich, 1917: 2 [Ponerini]; Forel, 1917: 235; Bondroit, 1918: 78 [Poneritae]; Wheeler, W.M. 1920: 53; Wheeler, W.M. 1922a: 56, 632, 640; Borgmeier, 1923: 37; Karavaiev, 1934: 49; Clark, 1951: 15; Brown, 1954e: 24; Wheeler, G.C. & Wheeler, J. 1972a: 39; Brown, 1973b: 165; all subsequent authors.

Ponerinae as poneromorph subfamily of Formicidae: Bolton, 2003: 42, 156.

Ponerinae as poneroid subfamily of Formicidae: Ouellette, Fisher, et al. 2006: 365; Brady, Schultz, et al. 2006: 18173; Moreau, Bell et al. 2006: 102; Ward, 2007a: 555.

Tribes of Ponerinae: Platythyreini, Ponerini, Thaumatomyrmecini.

Genera (extinct) incertae sedis in Ponerinae: †Afropone, †Eogorgites, †Eoponerites, †Furcisutura, †Longicapitia.

Subfamily Ponerinae references, world

Smith, F. 1858b: 76 (diagnosis); Mayr, 1862: 712 (genera key); Mayr, 1865: 11 (Odontomachidae, Poneridae diagnoses), Mayr, 1867a: 79, 81 (Odontomachidae, Poneridae diagnoses); Forel, 1878: 366 (diagnosis); Dalla Torre, 1893: 13 (world catalogue); Emery, 1895j: 766 (diagnosis); Emery, 1896e: 176 (genera key); Handlirsch, 1907: 879 (†fossil taxa catalogue); Wheeler, W.M. 1910g: 134 (diagnosis); Emery, 1911d: 3 (diagnosis, tribes key); Forel, 1917: 235 (synoptic classification); Forel, 1921c: 133 (diagnosis); Wheeler, W.M. 1922a: 56, 636 (diagnosis, tribes key); Brown & Nutting, 1950: 124 (venation, phylogeny); Brown, 1954e: 24 (phylogeny); Eisner, 1957: 475 (proventriculus morphology); Bernard, 1967: 79 (diagnosis); Gotwald, 1969: 25 (mouthparts morphology); Wheeler, G.C. & Wheeler, J. 1972a: 39 (diagnosis); Brown, 1973b: 165 (genera, distribution); Wheeler, G.C. & Wheeler, J. 1976b: 48 (larvae, review & synthesis); Snelling, R.R. 1981: 387 (synoptic classification); Wheeler, G.C. & Wheeler, J. 1985: 256 (synoptic classification); Billen, 1986: 168 (Dufour's gland); Fanfani & Dazzini Valcurone, 1986: 115 (exocrine glands synopsis); Dlussky & Fedoseeva, 1988: 78 (synoptic classification); Hölldobler & Wilson, 1990: 9 onward (synoptic classification, genera keys); Baroni Urbani, Bolton & Ward, 1992: 317 (phylogeny); Bolton, 1994: 153 (diagnosis, synoptic classification, genera keys); Bolton, 1995a: 1042 (census); Bolton, 1995b: 14 (catalogue); Grimaldi, Agosti & Carpenter, 1997: 20 (†Cretaceous genera); Dlussky & Rasnitsyn, 2002: 422 (diagnosis for wingless fossils); Bolton, 2003: 42, 156 (diagnosis, synopsis); Ouellette, Fisher et al. 2006: 359 (phylogeny); Brady, Schultz, et al. 2006: 18173 (phylogeny); Moreau, Bell et al. 2006: 102 (phylogeny); Ward, 2007a: 555 (classification).

.

Regional and national faunas with keys

Mayr, 1861: 28 (Europe); Mayr, 1868c: 69 (†Baltic Amber); André, 1874: 170 (Europe); Forel, 1874: 29 (Switzerland); André, 1882b: 227 (Europe & Algeria); Cresson, 1887: 97 (U.S.A., genera); Provancher, 1887: 238 (Canada); Nasonov, 1889: 54 (Russia); Forel, 1891b: 9 (Madagascar genera); Forel, 1900c: 52 (India & Sri Lanka); Bingham, 1903: 23 (India, Sri Lanka & Burma); Ruzsky, 1905b: 108 (Russian Empire); Wheeler, W.M. 1910g: 557 (North America genera); Bondroit, 1910: 489 (Belgium); Stitz, 1914: 54 (Central Europe); Gallardo, 1915: 31 (Argentina genera); Forel, 1915d: 6 (Switzerland); Arnold, 1915: 10 (South Africa); Donisthorpe, 1915d: 65 (Britain); Emery, 1916b: 98 (Italy); Wheeler, W.M. 1916m: 580 (U.S.A., Connecticut); Bondroit, 1918: 79 (France & Belgium); Gallardo, 1918b: 4 (Argentina); Soudek, 1922: 18 (Czechoslovakia); Stärcke, 1926: 82 (Netherlands); Donisthorpe, 1927b: 68 (Britain); Menozzi & Russo, 1930: 169 (Dominican Republic); Arnol'di, 1933b: 596 (Russia); Karavaiev, 1934: 50 (Ukraine); Smith, M.R. 1937: 823 (Puerto Rico); Stitz, 1939: 58 (Germany); Smith, M.R. 1943f: 278 (U.S.A. males); Buren, 1944a: 279 (U.S.A., Iowa); Smith, M.R. 1947f: 530 (U.S.A. genera); Creighton, 1950a: 31 (North America); Kusnezov, 1956: 11 (Argentina); Brown, 1958h: 11 (New Zealand); Gregg, 1963: 280 (U.S.A., Colorado); Bernard, 1967: 80 (Western Europe); Wilson & Taylor, 1967: 10 (Polynesia); Kempf, 1972a: 262 (Neotropical synoptic classification); Bolton, 1973a: 323 (West Africa genera); Bolton & Collingwood, 1975: 3 (Britain); Boven, 1977: 66 (Belgium); Kutter, 1977c: 21 (Switzerland); Arnol'di & Dlussky, 1978: 522 (former European U.S.S.R.); Collingwood, 1978: 74 (Iberian Peninsula); Collingwood, 1979: 29 (Fennoscandia & Denmark); Greenslade, 1979: 14 (South Australia genera); Francoeur, 1979: 30 (Canada, Québec); Schembri & Collingwood, 1981: 417 (Malta); Allred, 1982: 438 (U.S.A., Utah); Baroni Urbani, 1984: 75 (Neotropical genera); Collingwood, 1985: 236 (Saudi Arabia); Wheeler, G.C. & Wheeler, J. 1986g: 17 (U.S.A., Nevada); Agosti & Collingwood, 1987b: 264 (Balkans); Ogata, 1987: 101 (Japan genera); Morisita, Kubota, et al. 1989: 8 (Japan); Dlussky, Soyunov & Zabelin, 1990: 173 (Turkmenistan); Kupyanskaya, 1990: 85 (Far Eastern Russia); Atanasov & Dlussky, 1992: 51 (Bulgaria); Lattke, in Jaffe, 1993: 166 (Neotropical genera); Arakelian, 1994: 10 (Armenia); Wu, J. & Wang, 1995: 31 (China genera); Collingwood & Agosti, 1996: 308 (Saudi Arabia); Seifert, 1996b: 107 (Central Europe); Collingwood & Prince, 1998: 10 (Portugal); Kim, Kim & Kim, 1998: 145 (Korea); Shattuck, 1999: 52, 179 (Australia genera synopsis); Andersen, 2000: 19 (northern Australia genera); Zhou, 2001b: 22 (China, Guangxi); Czechowski, Radchenko & Czechowska, 2002: 133 (Poland); Aktaç & Radchenko, 2002: 53 (Turkey genera); Yoshimura & Onoyama, 2002: 436 (Japan genera, males); Mackay & Mackay, 2002: 27 (U.S.A., New Mexico); Csösz, 2003: 147 (Carpathian Basin); Palacio & Fernández, in Fernández, 2003d: 256 (Neotropical genera); Radchenko, 2005b: 184 (North Korea); Coovert, 2005: 23 (U.S.A., Ohio); Yoshimura & Fisher, 2007: 27 (Malagasy genera, males); Clouse, 2007b: 190 (Micronesia); Seifert, 2007: 108 (North and Central Europe); Bolton & Fisher, 2008c: 32 (Afrotropical genera); Fernández & Arias-Penna, 2008: 32 (Neotropical genera); Dlussky, 2009: 1071 (Eocene amber); Terayama, 2009: 101 (Taiwan genera); Heterick, 2009: 34 (south-western Australia genera); Boer, 2010: 68 (Benelux).

Dolichoderinae Dolichoderini Dolichoderus TRUE TRUE FALSE

Extant: 122 species, 21 subspecies

Fossil: 53 species

Dolichoderus Lund, 1831a: 130. Type-species: Formica attelaboides, by monotypy.

Taxonomic history

Dolichoderus in Formicidae: Smith, F. 1858b: 75.

Dolichoderus in Formicinae: Mayr, 1862: 652 (in key) [Formicidae]; Mayr, 1865: 9 [Formicidae].

Dolichoderus in Dolichoderidae: Ashmead, 1905b: 384.

Dolichoderus in Dolichoderinae: Forel, 1878: 386 [Dolichoderidae]; Emery & Forel, 1879a: 455 [Dolichoderidae]; Dalla Torre, 1893: 156; Forel, 1895b: 107 [Dolichoderidae]; Emery, 1895j: 771; Forel, 1895e: 462; Forel, 1899c: 98; Wheeler, W.M. 1910g: 142.

Dolichoderus in Dolichoderinae, Dolichoderini: Emery, 1913a: 7; Wheeler, W.M. 1915h: 77; Forel, 1917: 247; Wheeler, W.M. 1922a: 688; all subsequent authors.

Junior synonyms of DOLICHODERUS

Hypoclinea Mayr, 1855: 377 (attributed to Foerster). Type-species: Formica quadripunctata, by subsequent designation of Wheeler, W.M. 1911f: 165.

Taxonomic history

Hypoclinea in Formicidae: Smith, F. 1858b: 57.

Hypoclinea in Formicinae: Mayr, 1855: 377 [Formicidae]; Mayr, 1861: 40 [Formicidae]; Mayr, 1862: 652 (in key) [Formicidae]; Mayr, 1865: 9 [Formicidae]; Mayr, 1868c: 53 [Formicidae]; André, 1874: 168 [Formicidae].

Hypoclinea in Dolichoderidae: Ashmead, 1905b: 384.

Hypoclinea in Dolichoderinae, Dolichoderini: Emery, 1913a: 18; Forel, 1917: 247; all subsequent authors.

Hypoclinea as genus: Mayr, 1855: 377; Mayr, 1865: 9; Mayr, 1868c: 53; Mayr, 1870b: 953; Smith, 1871a: 310; Ashmead, 1905b: 384; Kempf, 1972a: 118; Lattke, 1987b: 259; Dlussky & Fedoseeva, 1988: 78; Mackay & Vinson, 1989b: 17; Hölldobler & Wilson, 1990: 17; Wu, J. & Wang, 1995: 120.

Hypoclinea as subgenus of Dolichoderus: Emery, 1894c: 234; Wheeler, W.M. 1910g: 142; Emery, 1913a: 10; Wheeler, W.M. 1915h: 77; Forel, 1917: 247; Borgmeier, 1923: 84; Clark, 1930b: 252; McAreavey, 1949: 17; Creighton, 1950a: 331; Chapman & Capco, 1951: 182; Kusnezov, 1956: 28; Kusnezov, 1959: 50; Smith, D.R. 1979: 1415; Kupyanskaya, 1990: 154.

Hypoclinea as junior synonym of Dolichoderus: Forel, 1878: 386; Emery & Forel, 1879a: 455; Dalla Torre, 1893: 156; Bingham, 1903: 291; Brown, 1973b: 181 [provisional]; Shattuck, 1992c: 66; Mackay, 1993b: 6; Shattuck, 1994: 39; Bolton, 1994: 26; Bolton, 1995b: 31; Bolton, 2003: 85.

Monacis Roger, 1862a: 233. Type-species: Formica bispinosa, by subsequent designation of Wheeler, W.M. 1911f: 167.

Taxonomic history

Monacis in Dolichoderidae: Ashmead, 1905b: 384.

Monacis in Dolichoderinae, Dolichoderini: Emery, 1913a: 9; Forel, 1917: 247; all subsequent authors.

Monacis as genus: Roger, 1862a: 233; Ashmead, 1905b: 384; Brown, 1950e: 249; Kempf, 1959b: 227; Kempf, 1972a: 141; Kempf, 1972c: 253; Dlussky & Fedoseeva, 1988: 78; Mackay & Vinson, 1989b: 20; Hölldobler & Wilson, 1990: 17.

Monacis as subgenus of Dolichoderus: Emery, 1894c: 228; Wheeler, W.M. 1910g: 142; Emery, 1913a: 9; Forel, 1917: 247; Wheeler, W.M. 1922a: 688; Borgmeier, 1923: 81; Kusnezov, 1956: 27; Kusnezov, 1959: 50.

Monacis as junior synonym of Hypoclinea: Mayr, 1862: 704.

Monacis as junior synonym of Dolichoderus: Forel, 1878: 386; Emery & Forel, 1879a: 455; Dalla Torre, 1893: 156; Brown, 1973b: 182 [provisional]; Shattuck, 1992c: 66; Mackay, 1993b: 6; Shattuck, 1994: 39; Bolton, 1994: 26; Bolton, 1995b: 35; Bolton, 2003: 85.

Monoceratoclinea Wheeler, W.M. 1935c: 68 [as subgenus of Dolichoderus]. Type-species: Dolichoderus (Hypoclinea) monoceros, by original designation.

Taxonomic history

Monoceratoclinea as subgenus of Dolichoderus: Wheeler, W.M. 1935c: 68.

Monoceratoclinea as genus: Brown, 1950e: 249; Dlussky & Fedoseeva, 1988: 78; Hölldobler & Wilson, 1990: 17.

Monoceratoclinea as junior synonym of Dolichoderus: Shattuck, 1992c: 66; Shattuck, 1994: 40; Bolton, 1994: 26; Bolton, 1995b: 35; Bolton, 2003: 85.

Diceratoclinea Wheeler, W.M. 1935c: 69 [as subgenus of Dolichoderus]. Type-species: Dolichoderus scabridus, by original designation.

Taxonomic history

Diceratoclinea as subgenus of Dolichoderus: Wheeler, W.M. 1935c: 69; Chapman & Capco, 1951: 181.

Diceratoclinea as genus: Brown, 1950e: 249.

Diceratoclinea as junior synonym of Dolichoderus: Brown, 1973b: 180 [provisional]; Taylor & Brown, D.R. 1985: 93; Shattuck, 1992c: 66; Shattuck, 1994: 40; Bolton, 1994: 26; Bolton, 1995b: 26; Bolton, 2003: 85.

Acanthoclinea Wheeler, W.M. 1935c: 69 [as subgenus of Dolichoderus]. Type-species: Dolichoderus doriae, by original designation.

Taxonomic history

Acanthoclinea in Dolichoderinae, Dolichoderini: Donisthorpe, 1943f: 618.

Acanthoclinea as subgenus of Dolichoderus: Wheeler, W.M. 1935c: 69.

Acanthoclinea as genus: Brown, 1950e: 249.

Acanthoclinea as junior synonym of Dolichoderus: Taylor & Brown, D.R. 1985: 93; Shattuck, 1992c: 66; Shattuck, 1994: 40; Bolton, 1994: 26; Bolton, 1995b: 18; Bolton, 2003: 85.

Karawajewella Donisthorpe, 1944b: 59.

Taxonomic history

[Replacement name for Diabolus Karavaiev, 1926d: 424; junior homonym of Diabolus Gray, J.E. 1841: 400 (Mammalia).]

Karawajewella as junior synonym of Dolichoderus: Brown, 1973b: 181 [provisional]; Shattuck, 1992c: 66; Shattuck, 1994: 39, 40; Bolton, 1994: 26; Bolton, 1995b: 32; Bolton, 2003: 85.

Homonym replaced by Karawajewella

Diabolus Karavaiev, 1926d: 424 [as subgenus of Dolichoderus]. Type-species: Dolichoderus (Diabolus) bifurcatus (junior synonym of Dolichoderus cuspidatus), by original designation.

Taxonomic history

[Diabolus Karavaiev junior homonym of Diabolus Gray, J.E. 1841: 400 (Mammalia).]

Diabolus as subgenus of Dolichoderus: Wheeler, W.M. 1935c: 68.

Genus Dolichoderus references

Roger, 1863b: 10, 14, 15 (Dolichoderus, Hypoclinea, Monacis catalogues); Mayr, 1863: 407, 423 (Dolichoderus, Hypoclinea catalogues); Mayr, 1865: 9 (Dolichoderus, Hypoclinea diagnoses); Mayr, 1867a: 75 (Hypoclinea diagnosis); Mayr, 1868c: 54 (†Baltic Amber species key); Mayr, 1870b: 955 (Hypoclinea species key); Mayr, 1876: 79 (Australia species key); André, 1882b: 225 (Europe & Algeria species); Mayr, 1886d: 435 (U.S.A. species key); Cresson, 1887: 258 (U.S.A. catalogue); Dalla Torre, 1893: 156 (catalogue); Forel, 1895e: 462 (India & Sri Lanka species key); Bingham, 1903: 292 (India, Sri Lanka & Burma species key); Wheeler, W.M. 1905d: 306 (North America species key); Emery, 1913a: 7 (diagnosis, subgenera key, catalogue); Emery, 1913a: 9, 10 (D. (Monacis), D. (Hypoclinea), catalogues); Mann, 1916: 460 (Brazil D. (Dolichoderus) species key); Wheeler, W.M. 1922a: 688 (subgenera key); Clark, 1930b: 253 (Australia D. (Hypoclinea) species key); Creighton, 1950a: 333 (North America D. (Hypoclinea) species key); Chapman & Capco, 1951: 181 (Asia checklist); Kempf, 1959b: 230 (Monacis, all species revision, key); Wheeler, J. 1963: 149 (U.S.A., North Dakota species key); Bernard, 1967: 249 (diagnosis); Kempf, 1969: 292 (Neotropical D. (Dolichoderus) species key); Kempf, 1972a: 98, 118, 141 (Neotropical Dolichoderus, Hypoclinea, Monacis, catalogues); Kempf, 1972c: 253 (Monacis species, additions to 1959a key); Smith, D.R. 1979: 1415 (North America catalogue); Taylor & Brown, D.R. 1985: 93 (Australia catalogue); Taylor, 1987a: 24 (Australia, New Caledonia checklist); Harada, 1987: 602 (Monacis species key); Lattke, 1987b: 263 (Neotropical D. bispinosus group, key); Dlussky & Fedoseeva, 1988: 78 (synoptic classification); Johnson, 1989a: 5 (U.S.A. species key); Hölldobler & Wilson, 1990: 17 (synoptic classification); Brandão, 1991: 340, 347, 356 (Neotropical catalogue); Shattuck, 1992c: 66 (diagnosis, review of genus); Mackay, 1993b: 17 (New World, all species revision, key); Radchenko, 1994b: 112 (South Siberia species key); Shattuck, 1994: 39 (catalogue); Bolton, 1995a: 1049 (census); Bolton, 1995b: 172 (catalogue); Wu, J. & Wang, 1995: 120 (China species key); Xu, 1995a: 34 (China species key); Shattuck, 1999: 68 (Australia synopsis); Xu, 2001b: 356 (China species key); Zhou, 2001b: 153 (China, Guangxi species key); Dill, 2002: 21, 30 (southeast Asia species groups key, D. cuspidatus group revision, key); Dlussky & Rasnitsyn, 2002: 416 (diagnosis for impression fossils); Dlussky, 2002b: 54 (Baltic & Rovno amber species); Dlussky, 2008b: 509 (Europe Eocene amber species key); Heterick, 2009: 45 (south-western Australia species key).

Dolichoderinae Leptomyrmecini Dorymyrmex TRUE TRUE FALSE

57 species, 26 subspecies

Dorymyrmex Mayr, 1866a: 494. Type-species: Dorymyrmex flavescens, by monotypy.

Taxonomic history

Dorymyrmex in Dolichoderinae: Forel, 1878: 383 [Dolichoderidae]; Dalla Torre, 1893: 167; Emery, 1895j: 771; Forel, 1899c: 103; Wheeler, W.M. 1910g: 142; all subsequent authors.

Dorymyrmex in Dolichoderidae: Ashmead, 1905b: 384.

Dorymyrmex in Dolichoderinae, Tapinomini: Emery, 1913a: 36; Forel, 1917: 248; Wheeler, W.M. 1922a: 689; Donisthorpe, 1943f: 640; subsequent authors and Jaffe, 1993: 9.

Dorymyrmex in Dolichoderinae, Dolichoderini: Shattuck, 1992c: 77; Bolton, 1994: 26; Bolton, 2003: 86.

Dorymyrmex in Dolichoderinae, Leptomyrmecini: Ward, Brady, et al. 2010: 361.

Junior synonyms of DORYMYRMEX

Psammomyrma Forel, 1912h: 43 [as subgenus of Dorymyrmex]. Type-species: Dorymyrmex planidens, by subsequent designation of Wheeler, W.M. 1913a: 82.

Taxonomic history

Psammomyrma as subgenus of Dorymyrmex: Forel, 1912h: 43; Kempf, 1972a: 100.

Psammomyrma as junior synonym of Dorymyrmex: Forel, 1913b: 350 (in text); Santschi, 1922b: 365; Wheeler, W.M. 1922a: 689; Snelling, R.R. & Hunt, 1976: 93; Shattuck, 1992c: 77; Shattuck, 1994: 73; Bolton, 1994: 26; Bolton, 2003: 86.

Conomyrma Forel, 1913b: 350 (in text) [as subgenus of Dorymyrmex]. Type-species: Prenolepis pyramica, by subsequent designation of Santschi, 1922b: 365.

Taxonomic history

Conomyrma in Dolichoderinae, Tapinomini: Forel, 1917: 248; Donisthorpe, 1943f: 634; Jaffe, 1993: 9.

Conomyrma as subgenus of Dorymyrmex: Forel, 1913b: 350; Forel, 1917: 248; Santschi, 1922b: 365; Wheeler, W.M. 1922a: 689; Gallardo, 1930: 147; Smith, M.R. 1951a: 837; Smith, M.R. 1958c: 140.

Conomyrma as genus: Kusnezov, 1952g: 429; Kusnezov, 1959: 51; Kusnezov, 1964: 66; Kempf, 1972a: 78; Snelling, R.R. 1973b: 1; Smith, D.R. 1979: 1419; Hölldobler & Wilson, 1990: 17; Jaffe, 1993: 9.

Conomyrma as junior synonym of Dorymyrmex: Brown, 1973b: 179 [provisional]; Shattuck, 1992c: 77; Shattuck, 1994: 73; Bolton, 1994: 26; Bolton, 2003: 86.

Araucomyrmex Gallardo, 1919b: 249. Type-species: Dorymyrmex tener, by original designation.

Taxonomic history

Araucomyrmex in Dolichoderinae, Tapinomini: Wheeler, W.M. 1922a: 689; Donisthorpe, 1943f: 625; Kempf, 1972a: 25; Dlussky & Fedoseeva, 1988: 77.

Araucomyrmex as genus: Gallardo, 1919b: 249; Wheeler, W.M. 1922a: 689; Kusnezov, 1956: 28; Kusnezov, 1959: 51; Kusnezov, 1964: 66; Kempf, 1972a: 25; Snelling, R.R. 1975: 9; Snelling, R.R. & Hunt, 1976: 93; Dlussky & Fedoseeva, 1988: 77.

Araucomyrmex as subgenus of Dorymyrmex: Santschi, 1922b: 365; Gallardo, 1930: 147; Kusnezov, 1952g: 428.

Araucomyrmex as junior synonym of Conomyrma: Snelling, R.R. 1981: 402.

Araucomyrmex as junior synonym of Dorymyrmex: Brown, 1973b: 178 [provisional]; Shattuck, 1992c: 77; Shattuck, 1994: 73; Bolton, 1994: 26; Bolton, 2003: 86.

Ammomyrma Santschi, 1922b: 365 [as subgenus of Dorymyrmex]. Type-species: Dorymyrmex exsanguis, by original designation.

Taxonomic history

Ammomyrma as subgenus of Dorymyrmex: Santschi, 1922b: 365; Gallardo, 1930: 147; Kempf, 1972a: 100.

Ammomyrma as junior synonym of Araucomyrmex: Snelling, R.R. & Hunt, 1976: 93.

Ammomyrma as junior synonym of Dorymyrmex: Shattuck, 1992c: 77; Shattuck, 1994: 73; Bolton, 1994: 26; Bolton, 2003: 87.

Biconomyrma Kusnezov, 1952g: 429 [as subgenus of Conomyrma]. Type-species: Dorymyrmex pyramicus var. brunneus (now Dorymyrmex brunneus), by subsequent designation of Kusnezov, 1959: 51.

Taxonomic history

Biconomyrma as subgenus of Conomyrma: Kusnezov, 1952g: 429.

Biconomyrma as genus: Kusnezov, 1959: 51; Kusnezov, 1964: 67.

Biconomyrma as junior synonym of Conomyrma: Smith, M.R. 1958c: 140; Kempf, 1972a: 78.

Biconomyrma as junior synonym of Dorymyrmex: Shattuck, 1992c: 78; Shattuck, 1994: 73; Bolton, 1994: 26; Bolton, 2003: 87.

Spinomyrma Kusnezov, 1952g: 429 (diagnosis in key) [as subgenus of Dorymyrmex]. Type-species: Dorymyrmex alboniger, by subsequent designation of Kusnezov, 1959: 51.

Taxonomic history

Spinomyrma as subgenus of Dorymyrmex: Kusnezov, 1952g: 429; Kempf, 1972a: 100.

Spinomyrma as genus: Kusnezov, 1956: 30 (in key); Kusnezov, 1959: 51; Kusnezov, 1964: 66.

Spinomyrma as junior synonym of Dorymyrmex: Snelling, R.R. & Hunt, 1976: 93; Shattuck, 1992c: 78; Shattuck, 1994: 74; Bolton, 1994: 26; Bolton, 2003: 87.

Genus Dorymyrmex references

Mayr, G. 1868a: 165 (diagnosis); Forel, 1878: 383 (diagnosis); Cresson, 1887: 257 (U.S.A. catalogue); Dalla Torre, 1893: 167 (catalogue); André, 1903: 364 (species key); Emery, 1913a: 36 (diagnosis, catalogue); Gallardo, 1916a: 21 (Argentina species key); Wheeler, W.M. 1922a: 689 (subgenera key); Cole, 1942: 371 (U.S.A., Utah species key); Creighton, 1950a: 348 (North America species key); Kusnezov, 1952g: 428 (subgenera key); Kempf, 1972a: 25, 78, 99 (Neotropical Araucomyrmex, Conomyrma, Dorymyrmex catalogues); Snelling, R.R. 1973b: 1 (U.S.A. Conomyrma species, revision); Snelling, R.R. & Hunt, 1976: 96 (Chile Araucomyrmex species key); Smith, D.R. 1979: 1419 (North America catalogue); Allred, 1982: 444 (U.S.A., Utah Conomyrma species key); Wheeler, G.C. & Wheeler, J. 1986g: 57 (U.S.A., Nevada Conomyrma species key); Trager, 1988: 14 (Southeastern U.S.A. Conomyrma species revision, key): Johnson, 1989b: 191 (U.S.A. Conomyrma species revision, key); Brandão, 1991: 327, 338, 340 (catalogue); Shattuck, 1992c: 77 (diagnosis, review of genus); Shattuck, 1994: 73 (catalogue); Snelling, R.R. 1995: 9 (U.S.A. species revision, key); Bolton, 1995a: 1049 (census); Bolton, 1995b: 181 (catalogue); Mackay & Mackay, 2002: 222 (U.S.A., New Mexico species key).

Dolichoderinae Leptomyrmecini Forelius TRUE TRUE FALSE

18 species

Forelius Emery, 1888d: 389. Type-species: Iridomyrmex mccooki, by monotypy.

Taxonomic history

Forelius in Dolichoderinae: Dalla Torre, 1893: 168; Emery, 1895j: 771; Forel, 1899c: 102; Wheeler, W.M. 1910g: 142.

Forelius in Dolichoderidae: Ashmead, 1905b: 384.

Forelius in Dolichoderinae, Tapinomini: Emery, 1913a: 35; Forel, 1917: 248; Wheeler, W.M. 1922a: 690; Donisthorpe, 1943f: 645; all subsequent authors and Jaffe, 1993: 9.

Forelius in Dolichoderinae, Dolichoderini: Shattuck, 1992c: 87; Bolton, 1994: 26; Bolton, 2003: 88.

Forelius in Dolichoderinae, Leptomyrmecini: Ward, Brady, et al. 2010: 361.

Junior synonym of FORELIUS

Neoforelius Kusnezov, 1953b: 327. Type-species: Neoforelius tucumanus, by monotypy.

Taxonomic history

Neoforelius in Dolichoderinae, Tapinomini: Kusnezov, 1953b: 327.

Neoforelius as genus: Kusnezov, 1953b: 327; Kusnezov, 1959: 51; Kusnezov, 1964: 66; Kempf, 1972a: 160; Jaffe, 1993: 9.

Neoforelius as junior synonym of Forelius: Shattuck, 1992c: 87; Shattuck, 1994: 91; Bolton, 1994: 26; Cuezzo, 2000: 204; Bolton, 2003: 88.

Genus Forelius references

Dalla Torre, 1893: 168 (catalogue); Emery, 1913a: 35 (diagnosis, catalogue); Gallardo, 1916a: 74 (Argentina species key); Creighton, 1950a: 343 (North America species, review); Kusnezov, 1957b: 16 (Neotropical species key); Kempf, 1972a: 108, 160 (Neotropical Forelius, Neoforelius catalogues); Smith, D.R. 1979: 1419 (North America catalogue); Wheeler, G.C. & Wheeler, J. 1986g: 56 (U.S.A., Nevada species key); Brandão, 1991: 344 (catalogue); Shattuck, 1992c: 87 (diagnosis, review of genus); Shattuck, 1994: 91 (catalogue); Bolton, 1995a: 1049 (census); Bolton, 1995b: 190 (catalogue); Cuezzo, 2000: 197 (diagnosis, all species revision, key); Mackay & Mackay, 2002: 227 (U.S.A., New Mexico species key); Guerrero & Fernández, 2008: 58 (addition to key).

Dolichoderinae Leptomyrmecini Iridomyrmex TRUE TRUE FALSE

Extant: 79 species

Fossil:

Iridomyrmex Mayr, 1862: 653 (diagnosis in key), 702. Type-species: Formica detecta (junior synonym of Formica purpurea), by subsequent designation of Bingham, 1903: 297.

Taxonomic history

Iridomyrmex in Formicinae: Mayr, 1862: 653 (in key) [Formicidae]; Mayr, 1865: 10 [Formicidae].

Iridomyrmex in Dolichoderinae: Forel, 1878: 381 [Dolichoderidae]; Dalla Torre, 1893: 168; Emery, 1895j: 771; Forel, 1899c: 102; Wheeler, W.M. 1910g: 142.

Iridomyrmex in Dolichoderidae: Ashmead, 1905b: 384.

Iridomyrmex in Dolichoderinae, Tapinomini: Emery, 1913a: 21; Wheeler, W.M. 1915h: 86; Arnold, 1915: 145; Forel, 1917: 248; Wheeler, W.M. 1922a: 689; Carpenter, 1930: 50; all subsequent authors; Jaffe, 1993: 9 (anachronism).

Iridomyrmex in Dolichoderinae, Dolichoderini: Shattuck, 1992c: 101; Bolton, 1994: 26; Bolton, 2003: 88.

Iridomyrmex in Dolichoderinae, Iridomyrmecini, Iridomyrmecina: Dubovikov, 2005a: 92.

Iridomyrmex in Dolichoderinae, Leptomyrmecini: Ward, Brady, et al. 2010: 362.

Genus Iridomyrmex references

Roger, 1863b: 13 (catalogue); Mayr, 1863: 424 (catalogue); Mayr, 1865: 10 (diagnosis); Mayr, 1867a: 76 (diagnosis); Forel, 1878: 381 (diagnosis); Cresson, 1887: 257 (U.S.A. catalogue); Dalla Torre, 1893: 168 (catalogue); Forel, 1895e: 467 (India & Sri Lanka species key); Bingham, 1903: 298 (India, Sri Lanka & Burma species key); Emery, 1913a: 21 (diagnosis, catalogue); Arnold, 1915: 145 (diagnosis); Creighton, 1950a: 341 (North America species key); Chapman & Capco, 1951: 188 (Asia checklist); Bernard, 1967: 250 (diagnosis); Kempf, 1972a: 124 (Neotropical catalogue); Smith, D.R. 1979: 1417 (North America catalogue); Taylor & Brown, D.R. 1985: 96 (Australia catalogue); Taylor, 1987a: 30 (Australia, New Caledonia & New Zealand checklist); Shattuck, 1992a: 14 (diagnosis); Shattuck, 1992c: 101 (diagnosis, review of genus); Shattuck, 1993a: 118 (I. purpureus group, key); Shattuck, 1993b: 1306 (I. calvus group, key); Shattuck, 1994: 99 (catalogue); Bolton, 1995a: 1050 (census); Bolton, 1995b: 217 (catalogue); Shattuck, 1996a: 37 (I. discors group); Shattuck, 1998: 303 (I. conifer group, key); Shattuck, 1999: 72 (Australia synopsis); Heterick, 2009: 48 (south-western Australia species key); Heterick & Shattuck, 2011: 22 (diagnosis, all species revision, key).

Dolichoderinae Tapinomini Liometopum TRUE TRUE FALSE

Extant: 7 species

Fossil: 9 species

Liometopum Mayr, 1861: 38. Type-species: Formica microcephala, by monotypy.

Taxonomic history

Liometopum in Formicinae: Mayr, 1861: 38 [Formicidae]; Mayr, 1862: 653 (in key) [Formicidae]; Mayr, 1865: 9 [Formicidae]; André, 1874: 168 [Formicidae].

Liometopum in Dolichoderinae: Forel, 1878: 383 [Dolichoderidae]; Emery & Forel, 1879a: 454 [Dolichoderidae]; Dalla Torre, 1893: 163; Emery, 1895j: 771; Forel, 1899c: 104; Wheeler, W.M. 1910g: 142.

Liometopum in Dolichoderidae: Ashmead, 1905b: 384.

Liometopum in Dolichoderinae, Tapinomini: Emery, 1913a: 19; Wheeler, W.M. 1915h: 95; Forel, 1917: 248; Bondroit, 1918: 87; Wheeler, W.M. 1922a: 688; Carpenter, 1930: 46; subsequent authors except those below; then Jaffe, 1993: 9; Ward, Brady, et al. 2010: 361.

Liometopum in Dolichoderinae, Liometopini: Dlussky & Fedoseeva, 1988: 77.

Liometopum in Dolichoderinae, Dolichoderini: Shattuck, 1992c: 121; Bolton, 1994: 26; Bolton, 2003: 89.

Genus Liometopum references

Roger, 1863b: 14 (catalogue); Mayr, 1863: 427 (catalogue); Mayr, 1865: 9 (diagnosis); Forel, 1878: 383 (diagnosis); André, 1882b: 220 (Europe & Algeria species); Cresson, 1887: 258 (U.S.A. catalogue); Dalla Torre, 1893: 162 (catalogue); Bingham, 1903: 289 (diagnosis); Wheeler, W.M. 1905e: 322 (North America species); Emery, 1913a: 19 (diagnosis, catalogue); Creighton, 1950a: 338 (North America species key); Chapman & Capco, 1951: 192 (Asia checklist); Gregg, 1963: 438 (U.S.A., Colorado species key); Bernard, 1967: 253 (diagnosis); Smith, D.R. 1979: 1417 (North America catalogue); Wheeler, G.C. & Wheeler, J. 1986g: 55 (U.S.A., Nevada species key); Shattuck, 1992c: 121 (diagnosis, review of genus); Shattuck, 1994: 128 (catalogue); Bolton, 1995a: 1050 (census); Bolton, 1995b: 247 (catalogue); Mackay & Mackay, 2002: 232 (U.S.A., New Mexico species key); Del Toro, Pacheco & Mackay, 2009: 306 (all species revision, key).

Dolichoderinae Tapinomini Tapinoma TRUE TRUE FALSE

Extant: 64 species, 26 subspecies

Fossil: 4 species

Tapinoma Foerster, 1850a: 43. Type-species: Tapinoma collina (junior synonym of Formica erratica), by monotypy.

Taxonomic history

Tapinoma in Formicidae: Smith, F. 1858b: 55.

Tapinoma in Formicinae: Mayr, 1855: 372 [Formicidae]; Mayr, 1861: 41 [Formicidae]; Mayr, 1862: 653 (in key) [Formicidae]; Mayr, 1865: 10 [Formicidae]; André, 1874: 168 [Formicidae]; André, 1881b: 63 [Formicidae].

Tapinoma in Dolichoderinae: Forel, 1878: 385 [Dolichoderidae]; Emery & Forel, 1879a: 454 [Dolichoderidae]; Forel, 1895b: 109 [Dolichoderidae]; Dalla Torre, 1893: 164; Emery, 1895j: 771; Forel, 1899c: 101; Wheeler, W.M. 1910g: 142.

Tapinoma in Dolichoderidae: Ashmead, 1905b: 384.

Tapinoma in Dolichoderinae, Tapinomini: Emery, 1913a: 38; Arnold, 1915: 152; Forel, 1917: 248; Bondroit, 1918: 89; Wheeler, W.M. 1922a: 690; subsequent authors except those below; then Jaffe, 1993: 9; Dubovikov, 2005a: 91 (implied); Ward, Brady, et al. 2010: 361.

Tapinoma in Dolichoderinae, Dolichoderini: Shattuck, 1992c: 146; Bolton, 1994: 26; Bolton, 2003: 91.

Junior synonyms of TAPINOMA

Micromyrma Dufour, 1857: 60. Type-species: Micromyrma pygmaea, by monotypy.

Taxonomic history

[Type-species not Formica erratica, unjustified subsequent designation by Wheeler, W.M. 1911f: 167.]

Micromyrma as subgenus of Tapinoma: Menozzi, 1925a: 19; Santschi, 1928e: 472; Chapman & Capco, 1951: 192.

Micromyrma as junior synonym of Tapinoma: Mayr, 1863: 455; Forel, 1878: 385; Emery & Forel, 1879a: 454; Dalla Torre, 1893: 164; Emery, 1913a: 38; Wheeler, W.M. 1922a: 923; Smith, M.R. 1951a: 837; Shattuck, 1992c: 146; Shattuck, 1994: 140; Bolton, 1994: 26; Bolton, 2003: 91; Fisher & Bolton, 2007: 65.

Semonius Forel, 1910f: 21. Type-species: Semonius schultzei, by monotypy.

Taxonomic history

Semonius in Dolichoderinae: Wheeler, W.M. 1910g: 142.

Semonius in Dolichoderinae, Tapinomini: Emery, 1913a: 44; Arnold, 1915: 157; Forel, 1917: 247; Wheeler, W.M. 1922a: 688; all subsequent authors.

Semonius as junior synonym of Tapinoma: Shattuck, 1992c: 146; Shattuck, 1994: 140; Bolton, 1994: 26; Bolton, 2003: 91; Fisher & Bolton, 2007: 66.

Pseudaphomomyrmex Wheeler, W.M. 1920: 53. Type-species: Aphomomyrmex emeryi, by original designation.

Taxonomic history

Pseudaphomomyrmex in Formicinae, Dimorphomyrmecini: Donisthorpe, 1943g: 721.

Pseudaphomomyrmex incertae sedis in Formicinae: Dlussky & Fedoseeva, 1988: 77.

Pseudaphomomyrmex in Formicinae, Myrmelachistini: Hölldobler & Wilson, 1990: 18.

Pseudaphomomyrmex in Formicinae, Brachymyrmecini: Bolton, 1994: 50.

Pseudaphomomyrmex in Formicinae, Plagiolepidini: Wheeler, W.M. 1922a: 695; Chapman & Capco, 1951: 214; Wheeler, G.C. & Wheeler, J. 1985: 258; Bolton, 2003: 106.

Pseudaphomomyrmex in Dolichoderinae: LaPolla & Longino, 2006: 305.

Pseudaphomomyrmex as junior synonym of Aphomomyrmex: Emery, 1925b: 44; Donisthorpe, 1943g: 721.

Pseudaphomomyrmex as genus: Wheeler, W.M. 1920: 53; Chapman & Capco, 1951: 214; Bolton, 1994: 50; Bolton, 1995b: 369; Bolton, 2003: 106.

Pseudaphomomyrmex as junior synonym of Tapinoma: Fisher & Bolton, 2007: 66.

Zatapinoma Wheeler, W.M. 1928a: 20. Type-species: Zatapinoma annandalei, by original designation.

Taxonomic history

Zatapinoma in Dolichoderinae, Tapinomini: Donisthorpe, 1943g: 737; all subsequent authors.

Zatapinoma as junior synonym of Tapinoma: Shattuck, 1992c: 146; Shattuck, 1994: 140; Bolton, 1994: 26; Bolton, 2003: 91; Fisher & Bolton, 2007: 66.

Neoclystopsenella Kurian, 1955: 133. Type-species: Neoclystopsenella luffae, by monotypy.

Taxonomic history

Neoclystopsenella in Bethylidae: Kurian, 1955: 133.

Neoclystopsenella in Dolichoderinae: Brown, 1988a: 337.

Neoclystopsenella as junior synonym of Tapinoma: Brown, 1988a: 337; Shattuck, 1992c: 146; Shattuck, 1994: 140; Bolton, 1994: 26; Bolton, 2003: 91; Fisher & Bolton, 2007: 66.

Genus Tapinoma references

Mayr, 1855: 372 (diagnosis); Smith, F. 1858b: 55 (diagnosis); Roger, 1863b: 13, 14 (Tapinoma, Micromyrma catalogues); Mayr, 1863: 455 (catalogue); Mayr, 1865: 10 (diagnosis); Mayr, 1867a: 78 (diagnosis); André, 1882b: 222 (Europe & Algeria species key); Cresson, 1887: 258 (U.S.A. catalogue); Dalla Torre, 1893: 164 (catalogue); Bingham, 1903: 304 (India, Sri Lanka & Burma species key); Emery, 1913a: 38 (diagnosis, catalogue); Emery, 1913a: 44 (Semonius diagnosis, catalogue); Arnold, 1915: 152, 157 (diagnosis, South Africa species key, Semonius diagnosis); Wheeler, W.M. 1922a: 695 (Pseudaphomomyrmex diagnosis, in key). Wheeler, W.M. 1922a: 923, 927 (Afrotropical Tapinoma, Semonius catalogues); Wheeler, W.M. 1922a: 1034 (Malagasy catalogue); Emery, 1925d: 63 (Palaearctic species key); Kuznetsov-Ugamsky, 1927d: 33 (Turkestan species key); Smith, M.R. 1928a: 311 (North America species key); Kratochvíl, Novák & Snoflák, 1944: 79 (Czechoslovakia species key); Creighton, 1950a: 352 (North America species key); Chapman & Capco, 1951: 192, 196 (Asia Semonius, Tapinoma, Zatapinoma checklists); Bernard, 1967: 254 (diagnosis, Western Europe species key); Boven, 1970b: 24 (Netherlands species key); Kempf, 1972a: 246 (Neotropical catalogue); Tarbinsky, 1976: 118 (Kyrghyzstan species key); Kutter, 1977c: 178 (Switzerland species key); Arnol'di & Dlussky, 1978: 547 (former European U.S.S.R. species key); Collingwood, 1978: 87 (Iberian Peninsula species key); Smith, D.R. 1979: 1421 (North America catalogue); Schembri & Collingwood, 1981: 421 (Malta species key); Gösswald, 1985: 317 (Germany species key); Taylor & Brown, D.R. 1985: 106 (Australia catalogue); Agosti & Collingwood, 1987b: 279 (Balkans species key); Taylor, 1987a: 77 (Australia, New Caledonia & New Zealand checklist); Dlussky, Soyunov & Zabelin, 1990: 168 (Turkmenistan species key); Brandão, 1991: 380 (Neotropical catalogue); Shattuck, 1992c: 146 (diagnosis, review of genus); Arakelian, 1994: 72 (Armenia species key); Shattuck, 1994: 140 (catalogue); Bolton, 1995a: 1053 (census); Bolton, 1995b: 399 (catalogue); Wu, J. & Wang, 1995: 116 (China species key); Collingwood & Agosti, 1996: 360 (Saudi Arabia species key); Seifert, 1996b: 165 (Central Europe species key); Skinner & Allen, 1996: 40 (Britain species key); Collingwood & Prince, 1998: 20 (Portugal species key); Shattuck, 1999: 81 (Australia synopsis); Zhou, 2001b: 151 (China, Guangxi species key); Czechowski, Radchenko & Czechowska, 2002: 135 (Poland species key); Seifert, 2007: 149 (North and Central Europe species key); Heterick, 2009: 58 (south-western Australia species key); Yoshimura & Fisher, 2011: 13 (Malagasy males).

Formicinae Lasiini Lasius TRUE TRUE FALSE

Extant: 100 species, 2 subspecies

Fossil: 22 species, 1 subspecies

Lasius Fabricius, 1804: 415. Type-species: Formica nigra, by subsequent designation of Bingham, 1903: 338.

Taxonomic history

[Lasius Jurine, 1801: 164 (Apidae), resurrected by Morice & Durrant, 1915: 421, later suppressed by Act of Commission.]

Lasius in Formicinae: Mayr, 1861: 49 [Formicidae]; Mayr, 1862: 652 (in key) [Formicidae]; Mayr, 1865: 8 [Formicidae]; Mayr, 1868c: 42 [Formicidae].

Lasius in Camponotinae: Forel, 1878: 374 [Camponotidae]; Emery & Forel, 1879a: 452 [Camponotidae]; Forel, 1886f: 206; Dalla Torre, 1893: 181.

Lasius in Camponotinae, Camponotini: Forel, 1886f: 206.

Lasius in Camponotinae, Formicini: Forel, 1893a: 165; Forel, 1895b: 103; Emery, 1895j: 772; Forel, 1899c: 127; Wheeler, W.M. 1910g: 143; Forel, 1912i: 89; Wheeler, W.M. 1915h: 120.

Lasius in Formicinae, Formicini: Wheeler, W.M. 1922a: 698; Carpenter, 1930: 58; Chapman & Capco, 1951: 201; Wheeler, G.C. & Wheeler, J. 1970: 651; Wheeler, G.C. & Wheeler, J. 1976b: 101; Wheeler, G.C. & Wheeler, J. 1985: 258.

Lasius in Formicinae, Acanthomyopsini: Donisthorpe, 1943f: 655.

Lasius in Formicinae, Lasius genus group: Agosti, 1991: 296.

Lasius in Formicinae, Lasiini: Ashmead, 1905b: 384; Emery, 1925b: 226; all subsequent authors except the entries above; Bolton, 1994: 50; Bolton, 2003: 22, 96.

Lasius as junior synonym of Acanthomyops: Forel, 1916: 460; Donisthorpe, 1916c: 276; Morice & Durrant, 1917: 442. Donisthorpe, 1927b: 206; Donisthorpe, 1937d: 132; Donisthorpe, 1943f: 655; Donisthorpe, 1946g: 91.

Lasius as junior synonym of Formicina: Emery, 1916a: 61; Emery, 1916b: 239; Emery, 1917a: 96; Wheeler, W.M. 1916k: 170; Bondroit, 1918: 19.

Lasius as genus: Fabricius, 1804: 415; Wheeler, W.M. 1916k: 172; Emery, 1925b: 226; Wilson, 1955a: 11; all subsequent authors.

Junior synonyms of LASIUS

Donisthorpea Morice & Durrant, 1915: 423. Type-species: Formica nigra, by original designation.

Taxonomic history

[Unnecessary replacement name for Lasius Fabricius.]

Donisthorpea in Formicinae, Acanthomyopsini: Donisthorpe, 1943f: 640; Donisthorpe, 1946g: 91.

Donisthorpea as genus: Morice & Durrant, 1915: 423; Donisthorpe, 1915d: 184.

Donisthorpea as subgenus of Formicina: Emery, 1916a: 11; Emery, 1916b: 240.

Donisthorpea as subgenus of Acanthomyops: Donisthorpe, 1927b: 229; Donisthorpe, 1943f: 640; Donisthorpe, 1946g: 91.

Donisthorpea as junior synonym of Acanthomyops: Forel, 1916: 460; Morice & Durrant, 1917: 442; Donisthorpe, 1927b: 209.

Donisthorpea as junior synonym of Lasius: Wheeler, W.M. 1916k: 172; Emery, 1925b: 226; Wilson, 1955a: 11. [Lasius and Donisthorpea share the same type-species; synonymy is therefore absolute.]

†Tylolasius Zhang, 1989: 295. Type-species: †Tylolasius inflatus, by original designation.

Taxonomic history

†Tylolasius incertae sedis in Formicinae: Bolton, 2003: 28, 131.

†Tylolasius as junior synonym of Lasius: Hong, Wu & Ren, 2001: 348.

Subgenera of LASIUS include the nominal plus the following.

Subgenus LASIUS (ACANTHOMYOPS)

Acanthomyops Mayr, 1862: 652 (diagnosis in key), 699. Type-species: Formica clavigera, by monotypy.

Taxonomic history

Acanthomyops in Formicinae: Mayr, 1862: 652 (in key) [Formicidae]; Mayr, 1865: 8 [Formicidae].

Acanthomyops in Camponotinae, Formicini: Wheeler, W.M. 1910g: 143; Forel, 1917: 249.

Acanthomyops in Formicinae, Formicini: Wheeler, W.M. 1922a: 698; Wheeler, G.C. & Wheeler, J. 1970: 651; Wheeler, G.C. & Wheeler, J. 1976b: 101; Wheeler, G.C. & Wheeler, J. 1985: 258; Jaffe, 1993: 14.

Acanthomyops in Formicinae, Acanthomyopsini: Donisthorpe, 1943f: 618.

Acanthomyops in Formicinae, Lasius genus group: Agosti, 1991: 296.

Acanthomyops in Formicinae, Lasiini: Ashmead, 1905b: 384 (misspelled as Acanthomyrmex); Emery, 1925b: 236; all subsequent authors except for those above.

Acanthomyops as junior synonym of Lasius: Mayr, 1866b: 888; Dalla Torre, 1893: 181.

Acanthomyops as subgenus of Formicina: Emery, 1916a: 11; Forel, 1917: 249.

Acanthomyops as genus: Mayr, 1862: 699; Donisthorpe, 1927b: 209; Donisthorpe, 1943f: 618; Creighton, 1950a: 426; Smith, M.R. 1958c: 150; Wing, 1968: 47; Bolton, 1994: 50; Bolton, 2003: 94.

Acanthomyops as subgenus of Lasius: Emery, 1893i: 642; Wheeler, W.M. 1910g: 143; Wheeler, W.M. 1916k: 172; Wheeler, W.M. 1922a: 698; Emery, 1925b: 236; Buren, 1950: 188; Smith, M.R. 1951a: 852; Gregg, 1963: 453; Ward, 2005: 13.

Subgenus LASIUS (DENDROLASIUS)

Dendrolasius Ruzsky, 1912: 630 [as subgenus of Lasius]. Type-species: Formica fuliginosa, by monotypy.

Taxonomic history

[Dendrolasius also described as new by Ruzsky, 1914a: 59.]

Dendrolasius as subgenus of Formicina: Forel, 1917: 249.

Dendrolasius as subgenus of Acanthomyops: Donisthorpe, 1927b: 211; Donisthorpe, 1943f: 637.

Dendrolasius as subgenus of Lasius: Ruzsky, 1912: 630; Wheeler, W.M. 1922a: 698; Emery, 1925b: 235; Wilson, 1955a: 14; all subsequent authors except the following.

Dendrolasius as junior synonym of Lasius: Brown, 1973b: 180 [provisional].

Subgenus LASIUS (CHTHONOLASIUS)

Chtonolasius Ruzsky, 1912: 630 [as subgenus of Lasius]. Type-species: Formica umbrata, by subsequent designation of Emery, 1925b: 232.

Taxonomic history

[Chtonolasius also described as new by Ruzsky, 1914a: 59. The claim by some authors (e.g. Emery, 1916a: 11; Donisthorpe, 1941f: 37; Donisthorpe, 1943f: 633) that Ruzsky nominated Lasius flavus as type-species is incorrect. Ruzsky (1912: 630) included those species "with the appearance of the yellow Lasius (Las. flavus D.G.) and its closely related species, and races (Las. carniolicus, Las. umbratus, Las. myops, Las. mixtus)". This is not a direct nomination of a type-species, nor is De Geer (= D.G.) the author of flavus: Bolton, 1995b: 24.]

Chthonolasius justified emendation of spelling: Wheeler, W.M. 1916k: 170.

Chthonolasius as junior synonym of Formicina: Wheeler, W.M. 1916k: 172; Emery, 1916a: 11.

Chthonolasius as subgenus of Acanthomyops: Donisthorpe, 1927b: 254.

Chthonolasius as subgenus of Lasius: Ruzsky, 1912: 630; Ruzsky, 1914a: 59; Wheeler, W.M. 1922a: 698; Emery, 1925b: 231; Wilson, 1955a: 13; all subsequent authors except the following.

Chthonolasius as junior synonym of Lasius: Brown, 1973b: 179 [provisional].

Subgenus LASIUS (CAUTOLASIUS)

Cautolasius Wilson, 1955a: 13 [as subgenus of Lasius]. Type-species: Formica flava, by original designation.

Taxonomic history

Cautolasius as junior synonym of Lasius: Brown, 1973b: 179 [provisional].

Cautolasius as subgenus of Lasius: Bolton, 2003: 97.

Subgenus LASIUS (AUSTROLASIUS)

Austrolasius Faber, 1967: 74 [as subgenus of Lasius]. Type-species: Lasius carniolicus, by original designation.

Taxonomic history

Austrolasius as junior synonym of Lasius: Brown, 1973b: 179 [provisional].

Autrolasius as subgenus of Lasius: Bolton, 2003: 97.

Genus Lasius references

Mayr, 1861: 49 (Europe species key); Roger, 1863b: 11 (catalogue); Mayr, 1863: 394, 425 (catalogue); Mayr, 1865: 8 (diagnosis); Mayr, 1867a: 74 (diagnosis); André, 1874: 179 (Europe species key); Forel, 1874: 46 (Switzerland species key); Forel, 1878: 374 (diagnosis); André, 1882b: 191 (Europe & Algeria species key); Cresson, 1887: 257 (U.S.A. catalogue); Provancher, 1887: 236 (Canada species key); Nasonov, 1889: 64 (Russia species key); Dalla Torre, 1893: 181 (catalogue); Emery, 1893i: 637 (North America species key); Bingham, 1903: 339 (India species key); Ruzsky, 1905b: 264 (Russian Empire species key); Wasmann, 1906: 10 (Luxemburg species key); Bondroit, 1910: 484 (Belgium species key); Wheeler, W.M. 1910e: 237 (North America L. umbratus forms, key); Stitz, 1914: 82 (Central Europe species key); Donisthorpe, 1915d: 188 (Britain species key); Emery, 1916b: 243 (Italy species key); Wheeler, W.M. 1916m: 591 (U.S.A., Connecticut species key); Bondroit, 1918: 19 (France & Belgium species key); Wheeler, W.M. 1922a: 698 (subgenera key); Emery, 1925b: 226 (diagnosis, subgenera key, catalogue); Emery, 1925b: 231 (L. (Chthonolasius) diagnosis, catalogue); Emery, 1925b: 235 (L. (Dendrolasius) diagnosis, catalogue); Emery, 1925b: 236 (L. (Acanthomyops) diagnosis, catalogue); Kuznetsov-Ugamsky, 1927e: 186 (Turkestan species key); Karavaiev, 1927c: 277 (Ukraine species key); Donisthorpe, 1927b: 206 (Britain species key); Arnol'di, 1933b: 602 (Russia species key); Karavaiev, 1936: 193 (Ukraine species key); Stärcke, 1937: 57 (Europe L. umbratus group, key); Menozzi, 1939a: 313 (Himalaya & Tibet species key); Stitz, 1939: 267 (Germany species key); Kratochvíl, 1941: 100 (Central Europe species key); Novák & Sadil, 1941: 100 (Central Europe species key); Cole, 1942: 373 (U.S.A., Utah species key); Holgersen, 1943b: 174 (Norway species key); Holgersen, 1944: 199 (Norway species key); Buren, 1944a: 296 (U.S.A., Iowa species key); Stärcke, 1944a: 157 (Europe L. niger group, key); Boven, 1947: 184 (Belgium species key); Creighton, 1950a: 418 (North America Lasius species key); Creighton, 1950a: 428 (North America Acanthomyops species key); Buren, 1950: 188 (Acanthomyops queens, key); Chapman & Capco, 1951: 201 (Asia checklist); Wilson, 1955a: 13 (subgenera review); Wilson, 1955a: 26 (Nearctic Lasius species revision, key); Wilson, 1955a: 28 (Palaearctic species revision, key); Boven, 1959: 11 (Netherlands species key); Wheeler, G.C. & Wheeler, J. 1963: 177 (U.S.A., North Dakota Lasius species key); Wheeler, G.C. & Wheeler, J. 1963: 195 (U.S.A., North Dakota Acanthomyops species key); Gregg, 1963: 453 (U.S.A., Colorado species key); Collingwood, 1963: 155 (Europe L. umbratus group species key); Collingwood, 1964: 107 (Britain species key); Bernard, 1967: 349 (diagnosis, Western Europe species key); Wing, 1968: 53 (Acanthomyops diagnosis, all species revision, key); Bourne, 1973: 19 (Britain species key); Boven, 1970b: 28 (Netherlands species key); Bolton & Collingwood, 1975: 7 (Britain species key); Tarbinsky, 1976: 133 (Kyrghyzstan species key); Boven, 1977: 135 (Belgium species key); Kutter, 1977c: 208 (Switzerland species key); Arnol'di & Dlussky, 1978: 555 (former European U.S.S.R. species key); Collingwood, 1978: 89 (Iberian Peninsula species key); Collingwood, 1979: 92 (Fennoscandia & Denmark species key); Smith, D.R. 1979: 1435 (North America Lasius catalogue); Smith, D.R. 1979: 1440 (North America Acanthomyops catalogue); Yamauchi, 1979: 147 (Japan species keys); Allred, 1982: 444 (U.S.A., Utah Acanthomyops species key); Allred, 1982: 450 (U.S.A., Utah Lasius species key); Collingwood, 1982: 283 (Himalayan species key); Gösswald, 1985: 268 (Germany species key); Wheeler, G.C. & Wheeler, J. 1986g: 64 (U.S.A., Nevada Lasius species key); Wheeler, G.C. & Wheeler, J. 1986g: 68 (U.S.A., Nevada Acanthomyops species key); Nilsson & Douwes, 1987: 70 (Norway species key); Agosti & Collingwood, 1987b: 281 (Balkans species key); Dlussky & Fedoseeva, 1988: 77 (synoptic classification); Seifert, 1988a: 143 (Europe L. (Chthonolasius) species key); Kupyanskaya, 1989: 781 (East Palaearctic L. (Dendrolasius) species key); Dlussky, Soyunov & Zabelin, 1990: 158 (Turkmenistan species key); Seifert, 1990: 1 (supplement to European L. (Chthonolasius)); Agosti & Bolton, 1990b: 149 (characters); Kupyanskaya, 1990: 211 (Far Eastern Russia species key); Morisita, Kubota, Onoyama, et al., 1991: 24 (Japan species key); Agosti, 1991: 295 (Lasius genus group diagnosis); Seifert, 1992b: 48 (Palaearctic L. (Lasius) species key); Atanasov & Dlussky, 1992: 232 (Bulgaria species key); Arakelian, 1994: 111 (Armenia species key); Bolton, 1994: 50 (synoptic classification); Radchenko, 1994b: 114 (South Siberia species key); Bolton, 1995a: 1050 (census); Bolton, 1995b: 53 (Acanthomyops catalogue); Bolton, 1995b: 221 (Lasius catalogue); Douwes, 1995: 92 (Sweden species key); Kupyanskaya, 1995: 361 (Far Eastern Russia species key); Wu, J. & Wang, 1995: 153 (China species key); Seifert, 1996b: 179 (Central Europe species key); Skinner & Allen, 1996: 42 (Britain species key); Collingwood & Prince, 1998: 22 (Portugal species key); Zhou, 2001b: 187 (China, Guangxi species key); Blacker & Collingwood, 2002: 44 (British species key); Czechowski, Radchenko & Czechowska, 2002: 154 (Poland species key); Mackay & Mackay, 2002: 237 (U.S.A., New Mexico Acanthomyops species key); Mackay & Mackay, 2002: 331 (U.S.A., New Mexico Lasius species key); Radchenko, 2005a: 93 (East Palaearctic L. (Dendrolasius) species key); Radchenko, 2005b: 190 (North Korea species key); Seifert, 2007: 159 (North and Central Europe species key); Terayama, 2009: 207 (Taiwan species key).

Formicinae Lasiini Myrmecocystus TRUE TRUE FALSE

29 species

Myrmecocystus Wesmael, 1838: 770. Type-species: Myrmecocystus mexicanus, by monotypy.

Taxonomic history

Myrmecocystus in Camponotinae: Forel, 1878: 372 [Camponotidae]; Emery & Forel, 1879a: 449 [Camponotidae]; Dalla Torre, 1893: 216.

Myrmecocystus in Camponotinae, Camponotini: Forel, 1886f: 201.

Myrmecocystus in Camponotinae, Formicini: Forel, 1893a: 165; Emery, 1895j: 772; Forel, 1899c: 129; Wheeler, W.M. 1910g: 144; Forel, 1912i: 89; Forel, 1917: 250.

Myrmecocystus in Formicinae, Formicini: Wheeler, W.M. 1922a: 699; Chapman & Capco, 1951: 203; Wheeler, G.C. & Wheeler, J. 1970: 651; Wheeler, G.C. & Wheeler, J. 1976b: 101; Wheeler, G.C. & Wheeler, J. 1985: 258; Jaffe, 1993: 14.

Myrmecocystus in Formicinae, Acanthomyopsini: Donisthorpe, 1943f: 665.

Myrmecocystus in Formicinae, Lasius genus group: Agosti, 1991: 296.

Myrmecocystus in Formicinae, Lasiini: Ashmead, 1905b: 384; Emery, 1925b: 238; subsequent authors except the above; Smith, D.R. 1979: 1445; Bolton, 1994: 50; Bolton, 2003: 98.

Myrmecocystus as junior synonym of Cataglyphis: Roger, 1863b: 12; Mayr, 1863: 431

Myrmecocystus as genus: Forel, 1878: 372; all subsequent authors.

Junior synonyms of MYRMECOCYSTUS

Endiodioctes Snelling, R.R. 1976: 25 [as subgenus of Myrmecocystus]. Type-species: Myrmecocystus melliger, by original designation.

Taxonomic history

Endiodioctes as junior synonym of Myrmecocystus: Snelling, R.R. 1981: 403.

Eremnocystus Snelling, R.R. 1976: 92 [as subgenus of Myrmecocystus]. Type-species: Myrmecocystus creightoni, by original designation.

Taxonomic history

Eremnocystus as junior synonym of Myrmecocystus: Snelling, R.R. 1981: 403.

Genus Myrmecocystus references

Forel, 1878: 372 (diagnosis); Cresson, 1887: 255 (U.S.A. catalogue); Dalla Torre, 1893: 216 (catalogue); Wheeler, W.M. 1912d: 173 (all species key); Emery, 1925b: 238 (diagnosis, catalogue); Cole, 1942: 385 (U.S.A., Utah species key); Creighton, 1950a: 440 (North America species key); Gregg, 1963: 643 (U.S.A., Colorado species key); Snelling, R.R. 1969a: 6 (M. melliger complex, key); Snelling, R.R. 1976: 22, 25, 27, 93, 115 (diagnosis, all species revision, keys); Smith, D.R. 1979: 1445 (North America catalogue); Snelling, R.R. 1982b: 77, 83 (supplements to 1976 keys); Allred, 1982: 451 (U.S.A., Utah species key); Wheeler, G.C. & Wheeler, J. 1986g: 69 (U.S.A., Nevada species key); Dlussky & Fedoseeva, 1988: 77 (synoptic classification); Bolton, 1995a: 1050 (census); Bolton, 1995b: 274 (catalogue); Mackay & Mackay, 2002: 344 (U.S.A., New Mexico species key).

Formicinae Plagiolepidini Brachymyrmex TRUE TRUE FALSE

41 species, 17 subspecies

Brachymyrmex Mayr, 1868b: 163. Type-species: Brachymyrmex patagonicus, by monotypy.

Taxonomic history

Brachymyrmex in Camponotinae: Forel, 1878: 375 [Camponotidae]; Dalla Torre, 1893: 174.

Brachymyrmex in Camponotinae, Plagiolepidini: Forel, 1893a: 165; Forel, 1895b: 106; Emery, 1895j: 771; Forel, 1899c: 123; Wheeler, W.M. 1910g: 143.

Brachymyrmex in Formicinae, Plagiolepidini: Ashmead, 1905b: 384.

Brachymyrmex in Camponotinae, Myrmelachistini: Forel, 1912i: 89.

Brachymyrmex in Formicinae, Dimorphomyrmecini: Emery, 1925b: 40 [subtribe Brachymyrmecini]; Donisthorpe, 1943f: 628.

Brachymyrmex incertae sedis in Formicinae: Dlussky & Fedoseeva, 1988: 77.

Brachymyrmex in Formicinae, Myrmelachistini: Wheeler, W.M. 1922a: 697; Hölldobler & Wilson, 1990: 18.

Brachymyrmex in Formicinae, Pseudolasius genus group: Agosti, 1991: 296.

Brachymyrmex in Formicinae, Brachymyrmecini: Wheeler, W.M. 1929a: 12; Kempf, 1972a: 37; Wheeler, G.C. & Wheeler, J. 1985: 258; Jaffe, 1993: 13; Bolton, 1994: 50.

Brachymyrmex in Formicinae, Plagiolepidini: Bolton, 2003: 23, 101.

Junior synonym of BRACHYMYRMEX

Bryscha Santschi, 1923b: 652 [as subgenus of Brachymyrmex]. Type-species: Brachymyrmex pilipes, by original designation.

Taxonomic history

Bryscha as junior synonym of Brachymyrmex: Brown, 1973b: 179 [provisional]; Smith, D.R. 1979: 1424; all subsequent authors.

Genus Brachymyrmex references

Forel, 1878: 375 (diagnosis); André, 1882b: 214 (Europe & Algeria species); Cresson, 1887: 255 (U.S.A. catalogue); Dalla Torre, 1893: 174 (catalogue); Wheeler, W.M. 1922a: 1036 (Malagasy catalogue); Santschi, 1923b: 652 (diagnosis, all species key); Emery, 1925b: 40 (diagnosis, catalogue); Emery, 1925b: 43 (B. (Bryscha) diagnosis, catalogue); Creighton, 1950a: 356 (North America species, review); Bernard, 1967: 279 (diagnosis); Kempf, 1972a: 37 (Neotropical catalogue); Alayo, 1974: 26 (Cuba species key); Smith, D.R. 1979: 1424 (North America catalogue); Brandão, 1991: 331 (catalogue); Bolton, 1995a: 1048 (census); Bolton, 1995b: 81 (catalogue); Quirán, et al. 2004: 275 (diagnosis).

Formicinae Plagiolepidini Paratrechina TRUE TRUE FALSE

1 species

Paratrechina Motschoulsky, 1863: 13. Type-species: Paratrechina currens (junior synonym of Formica longicornis), by subsequent designation of Wheeler, W.M. 1911f: 170.

Taxonomic history

Paratrechina in Formicinae, Acanthomyopsini: Donisthorpe, 1943f: 682.

Paratrechina in Formicinae, Prenolepidini: Chapman & Capco, 1951: 218; Hölldobler & Wilson, 1990: 18.

Paratrechina in Formicinae, Myrmelachistinini: Wheeler, G.C. & Wheeler, J. 1970: 652; Wheeler, G.C. & Wheeler, J. 1976b: 101.

Paratrechina in Formicinae, Brachymyrmecini: Wheeler, G.C. & Wheeler, J. 1985: 258.

Paratrechina in Formicinae, Pseudolasius genus group: Agosti, 1991: 296.

Paratrechina in Formicinae, Lasiini: Emery, 1925b: 216; Stitz, 1939: 379; Bernard, 1953b: 257; Kempf, 1972a: 182; Smith, D.R. 1979: 1442; Dlussky & Fedoseeva, 1988: 77; Jaffe, 1993: 14; Bolton, 1994: 50.

Paratrechina in Formicinae, Plagiolepidini: Bolton, 2003: 23, 104.

Paratrechina as junior synonym of Prenolepis: Dalla Torre, 1893: 177; Wheeler, W.M. 1922a: 940.

Paratrechina as genus: Emery, 1925b: 216; all subsequent authors.

Genus Paratrechina references

References prior to 2010 also include Nylanderia and Paraparatrechina.

André, 1882b: 203 (Europe & Algeria species key); Forel, 1894c: 406 (India & Sri Lanka species key); Wheeler, W.M. 1922a: 941, 1037 (Afrotropical, Malagasy P. (Nylanderia) catalogues); Emery, 1925b: 216 (diagnosis, catalogue); Emery, 1925b: 217 (P. (Nylanderia) diagnosis, catalogue); Emery, 1925b: 224 (Andragnathus diagnosis, catalogue); Finzi, 1936: 191 (Egypt species key); Buren, 1944a: 295 (U.S.A., Iowa species key); Creighton, 1950a: 404 (North America species key); Chapman & Capco, 1951: 197, 214, 218 (Asia Andragnathus, Nylanderia, Paratrechina checklists); Kusnezov, 1956: 32 (Argentina species key); Bernard, 1967: 347 (diagnosis, Western Europe species key); Kempf, 1972a: 166, 182 (Neotropical Nylanderia, Paratrechina catalogues); Alayo, 1974: 26 (Cuba species key); Arnol'di & Dlussky, 1978: 556 (former European U.S.S.R. species key); Smith, D.R. 1979: 1442 (North America, catalogue); Trager, 1984b: 65 (Nearctic species revision, key); Wheeler, G.C. & Wheeler, J. 1986g: 64 (U.S.A., Nevada species key); Collingwood, 1985: 298 (Saudi Arabia species key); Taylor & Brown, D.R. 1985: 129 (Australia catalogue); Taylor, 1987a: 52 (Australia, New Caledonia & New Zealand checklist); Brandão, 1991: 365 (Neotropical catalogue); Morisita, Kubota, et al. 1991: 20 (Japan species key); Bolton, 1995a: 1051 (census); Bolton, 1995b: 312 (catalogue); Wu, J. & Wang, 1995: 150 (China species key); Collingwood & Agosti, 1996: 369 (Saudi Arabia species key); Terayama, 1999c: 50 (Japan species key); Shattuck, 1999: 104 (Australia synopsis); Zhou, 2001b: 177 (China, Guangxi species key); Mackay & Mackay, 2002: 354 (U.S.A., New Mexico species key); Terayama, 2009: 210 (Taiwan species key); Heterick, 2009: 104 (south-western Australia species key); LaPolla, Brady & Shattuck, 2010a: 127 (diagnosis).

Formicinae Plagiolepidini Prenolepis TRUE TRUE FALSE

Extant: 19 species, 6 subspecies

Fossil: 1 species

Prenolepis Mayr, 1861: 52. Type-species: Tapinoma nitens, by subsequent designation of Bingham, 1903: 325.

Taxonomic history

[Type-species not Formica imparis, unjustified subsequent designation by Emery, 1906b: 134, repeated in Wheeler, W.M. 1911f: 171 and Wheeler, W.M. 1922a: 940.]

Prenolepis in Formicinae: Mayr, 1861: 52 [Formicidae]; Mayr, 1862: 652 [Formicidae]; Mayr, 1865: 7 [Formicidae]; Mayr, 1868c: 32 [Formicidae].

Prenolepis in Camponotinae: Forel, 1878: 377 [Camponotidae]; Emery & Forel, 1879a: 453 [Camponotidae]; Dalla Torre, 1893: 177.

Prenolepis in Camponotinae, Camponotini: Forel, 1886f: 209.

Prenolepis in Camponotinae, Formicini: Forel, 1893a: 165; Forel, 1895b: 103; Emery, 1895j: 772; Forel, 1899c: 125; Wheeler, W.M. 1910g: 143.

Prenolepis in Formicinae, Plagiolepidini: Ashmead, 1905b: 384.

Prenolepis in Camponotinae, Prenolepidini: Forel, 1912i: 89; Wheeler, W.M. 1915h: 117; Forel, 1917: 249; Arnold, 1922: 605.

Prenolepis in Formicinae, Prenolepidini: Wheeler, W.M. 1922a: 697; Chapman & Capco, 1951: 219; Hölldobler & Wilson, 1990: 18.

Prenolepis in Formicinae, Acanthomyopsini: Donisthorpe, 1943f: 685.

Prenolepis in Formicinae, Brachymyrmecini: Wheeler, G.C. & Wheeler, J. 1970: 651; Wheeler, G.C. & Wheeler, J. 1976b: 101; Wheeler, G.C. & Wheeler, J. 1985: 258.

Prenolepis in Formicinae, Lasiini: Emery, 1925b: 224; Stitz, 1939: 379, Kempf, 1972a: 209; Smith, D.R. 1979: 1444; Dlussky & Fedoseeva, 1988: 77; Jaffe, 1993: 14; Bolton, 1994: 50.

Prenolepis in Formicinae, Lasius genus group: Agosti, 1991: 296.

Prenolepis in Formicinae, Plagiolepidini: Bolton, 2003: 23, 105.

Genus Prenolepis references

Roger, 1863b: 10 (catalogue); Mayr, 1863: 451 (catalogue); Mayr, 1865: 7 (diagnosis); Mayr, 1867a: 71 (diagnosis); Mayr, 1870b: 947 (all species key); Forel, 1878: 377 (diagnosis); André, 1882b: 203 (Europe & Algeria species key); Cresson, 1887: 257 (U.S.A. catalogue); Dalla Torre, 1893: 177 (catalogue); Bingham, 1903: 326 (India, Sri Lanka & Burma species key); Emery, 1910a: 127 (Palaearctic species key); Arnold, 1922: 605 (diagnosis); Wheeler, W.M. 1922a: 216, 697, 940 (diagnosis, subgenera key, Afrotropical catalogue); Emery, 1925b: 224 (diagnosis, catalogue); Creighton, 1950a: 410 (North America species review); Chapman & Capco, 1951: 219 (Asia checklist); Kempf, 1972a: 209 (Neotropical catalogue); Smith, D.R. 1979: 1442 (North America catalogue); Bolton, 1994: 50 (synoptic classification); Bolton, 1995a: 1052 (census); Bolton, 1995b: 363 (catalogue); Wu, J. & Wang, 1995: 137 (China species key); Xu, 1995c: 338 (Southern & Eastern Asia species key); Fontenla Rizo, 2000: 81 (Antilles Is species key); Zhou, 2001b: 170 (China, Guangxi species key); Wang, W. & Wu, 2007: 721 (China species key); LaPolla, Brady & Shattuck, 2010a: 129 (diagnosis).

Formicinae Camponotini Camponotus TRUE TRUE FALSE

Extant: 1,058 species, 495 subspecies

Fossil: 31 species

Camponotus Mayr, 1861: 35. Type-species: Formica ligniperda, by subsequent designation of Bingham, 1903: 347.

Taxonomic history

[Type-species not Formica herculeana, unjustified subsequent designation by Forel, 1914a: 259, repeated in Arnold, 1922: 612.]

Camponotus in Formicinae: Mayr, 1861: 35 [Formicidae]; Mayr, 1862: 652 (in key) [Formicidae]; Mayr, 1865: 6 [Formicidae]; Mayr, 1868c: 26 [Formicidae].

Camponotus in Camponotinae: Forel, 1878: 367 [Camponotidae]; Emery & Forel, 1879a: 447 [Camponotidae]; Forel, 1892j: 223 [Camponotidae]; Dalla Torre, 1893: 171.

Camponotus in Camponotinae, Camponotini: Forel, 1886f: 141; Forel, 1893a: 165; Forel, 1895b: 101; Emery, 1895j: 772; Forel, 1899c: 130; Ashmead, 1905b: 384; Wheeler, W.M. 1910g: 144; Forel, 1912i: 89; Wheeler, W.M. 1915h: 138; Forel, 1917: 250; Arnold, 1922: 610.

Camponotus in Formicinae, Camponotini: Bondroit, 1918: 65; Wheeler, W.M. 1922a: 703; Emery, 1925b: 59; all subsequent authors except the following.

Camponotus in Formicinae, Formica genus group: Agosti, 1991: 295.

Camponotus as subgenus of Formica: Smith, F. 1871a: 306.

Camponotus as genus: all authors except the entry above.

Junior synonym of CAMPONOTUS

†Paleosminthurus Pierce & Gibron, 1962: 146. Type-species: †Paleosminthurus juliae, by monotypy.

Taxonomic history

†Paleosminthurus in Collembola, family †Paleosminthuridae: Pierce & Gibron, 1962: 146.

†Paleosminthurus incertae sedis in Formicidae: Najt, 1987: 152.

†Paleosminthurus incertae sedis in Formicinae: Hölldobler & Wilson, 1990: 18.

†Paleosminthurus in †Paleosmithurinae: Bolton, 1994: 187 [provisional subfamily status]; Bolton, 1995b: 14, 311; Bolton, 2003: 76, 263.

†Paleosminthurus as junior synonym of Camponotus: Snelling (pers. comm., 2004). [Synonymy remained unpublished at his death in 2008.]

†Shanwangella Zhang, 1989: 307. Type-species: †Shanwangella palaeoptera, by original designation.

Taxonomic history

†Shanwangella as junior synonym of Camponotus: Hong & Wu, 2000: 20.

Subgenera of CAMPONOTUS include the nominal plus the following.

All the subgenera were given as provisional junior synonyms of Camponotus by Brown, 1973b: 179 - 185. Brown's list was repeated in Hölldobler & Wilson, 1990: 18 but with all subgenera listed as junior synonyms; subgeneric status was reinstated in Bolton, 1994: 50. These consecutive entries are not repeated individually below.

Subgenus CAMPONOTUS (COLOBOPSIS)

Colobopsis Mayr, 1861: 38. Type-species: Formica truncata, by subsequent designation of Bingham, 1903: 342.

Taxonomic history

Colobopsis in Formicinae: Mayr, 1861: 38 [Formicidae]; Mayr, 1862: 652 (in key) [Formicidae]; Mayr, 1865: 7 [Formicidae].

Colobopsis in Camponotinae: Forel, 1878: 368 [Camponotidae]; Emery & Forel, 1897: 449 [Camponotidae].

Colobopsis in Camponotinae, Camponotini: Forel, 1886f: 193; Emery, 1895j: 772; Ashmead, 1905b: 384.

Colobopsis in Formicinae, Camponotini: Bondroit, 1918: 65; Emery, 1925b: 144; all subsequent authors.

Colobopsis as genus: Mayr, 1861: 38; Mayr, 1862: 652; Mayr, 1865: 7; Mayr, 1870b: 940; Forel, 1878: 368; Forel, 1879a: 125; Emery & Forel, 1879a: 449; André, 1882b: 159; Bingham, 1903: 342; Forel, 1886f: 193; Ashmead, 1905b: 384; Bondroit, 1918: 66; Jaffe, 1993: 14.

Colobopsis as subgenus of Camponotus: Emery, 1889b: 517; Dalla Torre, 1893: 219; Forel, 1893a: 165; Forel, 1893b: 435; Emery, 1895j: 772; Wheeler, W.M. 1904b: 139; Wheeler, W.M. 1910g: 144; Forel, 1912i: 90; Forel, 1914a: 263; Forel, 1917: 251; Emery, 1920b: 247; Arnold, 1922: 613; Wheeler, W.M. 1922a: 708; Emery, 1925b: 144; Donisthorpe, 1943f: 634; Creighton, 1950a: 390; Chapman & Capco, 1951: 222; Kempf, 1972a: 42; Smith, D.R., 1979: 1433; Bolton, 1994: 50.

Junior synonyms of CAMPONOTUS (COLOBOPSIS)

Condylomyrma Santschi, 1928c: 72 [as subgenus of Camponotus]. Type-species: Camponotus (Condylomyrma) bryani, by monotypy.

Taxonomic history

Condylomyrma as subgenus of Camponotus: Santschi, 1928c: 72.

Condylomyrma as junior synonym of Colobopsis: Wheeler, W.M. 1934e: 422 (in text).

[Campylomyrma Wheeler, W.M. 1934e: 421 (in text), incorrect subsequent spelling.]

Dolophra Wu, J. & Wang, 1994: 35. Type-species: Dolophra politae, by original designation.

Taxonomic history

Dolophra as genus: Wu, J. & Wang, 1994: 35; Wu, J. & Wang, 1995: 158.

Dolophra as junior synonym of Camponotus: Bolton, 1995b: 27.

Dolophra as junior synonym of Camponotus (Colobopsis): Bolton, 2003: 113, 268.

Subgenus CAMPONOTUS (MAYRIA)

Mayria Forel, 1878: 369. Type-species: Mayria madagascarensis (junior secondary homonym in Camponotus, replaced by Camponotus repens), by monotypy.

Taxonomic history

Mayria in Camponotinae: Forel, 1878: 369 [Camponotidae]; Dalla Torre, 1893: 219.

Mayria in Camponotinae, Camponotini: Forel, 1893a: 165; Emery, 1895j: 772; Ashmead, 1905b: 384; Wheeler, W.M. 1910g: 144.

Mayria as subgenus of Camponotus: Forel, 1894e: 227; Forel, 1914a: 262; Forel, 1917: 251; Wheeler, W.M. 1922a: 706; Emery, 1925b: 121; all subsequent authors.

Junior synonym of CAMPONOTUS (MAYRIA)

Myrmosaga Forel, 1912i: 92 [as subgenus of Camponotus]. Type-species: Camponotus kelleri, by subsequent designation of Wheeler, W.M. 1913a: 81.

Taxonomic history

[Type-species not Camponotus quadrimaculatus, unjustified subsequent designation by Forel, 1914a: 260; repeated in Arnold, 1922: 613.]

Myrmosaga as subgenus of Camponotus Forel, 1912i: 92; Forel, 1914a: 260; Forel, 1917: 250; Emery, 1920b: 241; Arnold, 1922: 613; Wheeler, W.M. 1922a: 706; all subsequent authors.

Myrmosaga as junior synonym of Mayria: Emery, 1925b: 121.

Subgenus CAMPONOTUS (RHINOMYRMEX)

Rhinomyrmex Forel, 1886f: 192. Type-species: Rhinomyrmex klaesii, by monotypy.

Taxonomic history

Rhinomyrmex in Camponotinae: Forel, 1886f: 192 [Camponotii]; Dalla Torre, 1893: 219.

Rhinomyrmex in Camponotinae, Camponotini: Forel, 1893a: 165; Emery, 1895j: 772; Ashmead, 1905b: 384; Wheeler, W.M. 1910g: 144.

Rhinomyrmex as genus: Forel, 1886f: 192; Forel, 1893a: 165; Dalla Torre, 1893: 219; Emery, 1895j: 772; Wheeler, W.M. 1910g: 144.

Rhinomyrmex as subgenus of Camponotus: Emery, 1896d: 374; Forel, 1914a: 264; Forel, 1917: 251; Emery, 1920b: 247; Wheeler, W.M. 1922a: 708; Emery, 1925b: 142; all subsequent authors.

Subgenus CAMPONOTUS (DENDROMYRMEX)

Dendromyrmex Emery, 1895j: 772. Type-species: Formica chartifex, by subsequent designation of Wheeler, W.M. 1911f: 161.

Taxonomic history

Dendromyrmex in Camponotinae, Camponotini: Emery, 1895j: 772; Ashmead, 1905b: 384; Forel, 1912i: 89; Wheeler, W.M. 1910g: 144; Forel, 1917: 250.

Dendromyrmex in Formicinae, Camponotini: Wheeler, W.M. 1922a: 700; Emery, 1925b: 172; all subsequent authors except the following.

Dendromyrmex in Formicinae, Formica genus group: Agosti, 1991: 295.

Dendromyrmex as subgenus of Camponotus: Fernández, 2002: 51.

Subgenus CAMPONOTUS (DINOMYRMEX)

Dinomyrmex Ashmead, 1905b: 384. Type-species: Formica gigas, by original designation.

Taxonomic history

Dinomyrmex in Camponotinae, Camponotini: Ashmead, 1905b: 384.

Dinomyrmex as genus: Ashmead, 1905b: 384.

Dinomyrmex as subgenus of Camponotus: Forel, 1913b: 350; Forel, 1914a: 259; Forel, 1917: 250; Emery, 1920b: 236; Arnold, 1922: 612; Wheeler, W.M. 1922a: 704; Emery, 1925b: 69; all subsequent authors.

Junior synonym of CAMPONOTUS (DINOMYRMEX)

Myrmogigas Forel, 1912i: 91 [as subgenus of Camponotus]. Type-species: Formica gigas, by subsequent designation of Wheeler, W.M. 1913a: 81.

Taxonomic history

Myrmogigas as subgenus of Camponotus: Forel, 1912i: 91.

Myrmogigas as junior synonym of Dinomyrmex: Forel, 1913b: 350 (in text); Forel, 1914a: 259; Wheeler, W.M. 1922a: 704. [Dinomyrmex and Myrmogigas share the same type-species, synonymy is therefore absolute.]

Subgenus CAMPONOTUS (TANAEMYRMEX)

Tanaemyrmex Ashmead, 1905b: 384. Type-species: Formica longipes (junior primary homonym in Formica, replaced by Camponotus etiolipes), by original designation.

Taxonomic history

Tanaemyrmex in Camponotinae, Camponotini: Ashmead, 1905b: 384.

Tanaemyrmex as genus: Ashmead, 1905b: 384.

Tanaemyrmex as subgenus of Camponotus: Emery, 1925b: 75; all subsequent authors.

Junior synonym of CAMPONOTUS (TANAEMYRMEX)

Myrmoturba Forel, 1912i: 91 [as subgenus of Camponotus]. Type-species: Formica maculata, by subsequent designation of Wheeler, W.M. 1913a: 82.

Taxonomic history

Myrmoturba as subgenus of Camponotus: Forel, 1912i: 91; Forel, 1914a: 259; Forel, 1917: 250; Emery, 1920b: 235; Arnold, 1922: 612; Wheeler, W.M. 1922a: 704.

Myrmoturba as junior synonym of Tanaemyrmex: Emery, 1925b: 75; all subsequent authors.

Subgenus CAMPONOTUS (ORTHONOTOMYRMEX)

Orthonotomyrmex Ashmead, 1906: 31.

Taxonomic history

[Replacement name for Orthonotus Ashmead, 1905b: 384; junior homonym of Orthonotus Westwood, in Stephens, 1829: 344 (Hemiptera). Type-species not Formica lateralis, unjustified subsequent designation by Forel, 1914a: 264; repeated in Arnold, 1922: 613.]

Orthonotomyrmex as subgenus of Camponotus: Forel, 1913i: 350 (in text); Forel, 1914a: 264; Forel, 1917: 251; Emery, 1920b: 244; Arnold, 1922: 613; Wheeler, W.M. 1922a: 707; Emery, 1925b: 124; all subsequent authors.

Homonym replaced by ORTHONOTOMYRMEX

Orthonotus Ashmead, 1905b: 384. Type-species: Formica sericea, by original designation.

Taxonomic history

[Junior homonym of Orthonotus Westwood, in Stephens, 1829: 344 (Hemiptera).]

Orthonotus in Camponotinae, Camponotini: Ashmead, 1905b: 384.

Subgenus CAMPONOTUS (MYRMAMBLYS)

Myrmamblys Forel, 1912i: 90 [as subgenus of Camponotus]. Type-species: Camponotus reticulatus, by subsequent designation of Wheeler, W.M. 1913a: 80.

Taxonomic history

[Type-species not Camponotus fastigatus, unjustified subsequent designation by Forel, 1914a: 263; repeated in Arnold, 1922: 613.]

Myrmamblys as subgenus of Camponotus: Forel, 1912i: 90; Forel, 1914a: 263; Forel, 1917: 251; Emery, 1920b: 248; Arnold, 1922: 613; Wheeler, W.M. 1922a: 708; Emery, 1925b: 136; Santschi, 1926c: 601; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOTHRIX)

Myrmothrix Forel, 1912i: 91 [as subgenus of Camponotus]. Type-species: Formica abdominalis (junior primary homonym; Formica atriceps is first available replacement name), by subsequent designation of Wheeler, W.M. 1913a: 81.

Taxonomic history

[Type-species not Formica rufipes, unjustified subsequent designation by Forel, 1914a: 260.]

Myrmothrix as subgenus of Camponotus: Forel, 1912i: 91; Forel, 1914a: 260; Forel, 1917: 250; Emery, 1920b: 236; Emery, 1925b: 107; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOSERICUS)

Myrmosericus Forel, 1912i: 91 [as subgenus of Camponotus]. Type-species: Formica rufoglauca, by subsequent designation of Wheeler, W.M. 1913a: 81.

Taxonomic history

Myrmosericus as subgenus of Camponotus: Forel, 1912i: 91; Forel, 1914a: 259; Forel, 1917: 250; Emery, 1920b: 236; Arnold, 1922: 613; Wheeler, W.M. 1922a: 704; Emery, 1925b: 104; all subsequent authors.

[See note under Myrmosaulus.]

Subgenus CAMPONOTUS (MYRMOPHYMA)

Myrmophyma Forel, 1912i: 91 [as subgenus of Camponotus]. Type-species: Camponotus capito, by subsequent designation of Wheeler, W.M. 1913a: 81.

Taxonomic history

[Type-species not Camponotus quadrisectus, unjustified subsequent designation by Forel, 1914a: 261; repeated in Arnold, 1922: 612.]

Myrmophyma as subgenus of Camponotus: Forel, 1912i: 91; Forel, 1914a: 261; Forel, 1917: 250; Emery, 1920b: 239; Arnold, 1922: 612; Wheeler, W.M. 1922a: 706; Emery, 1925b: 109; all subsequent authors.

Junior synonym of CAMPONOTUS (MYRMOPHYMA)

Myrmocamelus Forel, 1914a: 261 [as subgenus of Camponotus]. Type-species: Formica ephippium, by original designation.

Taxonomic history

[Type-species not Camponotus gambeyi, unjustified subsequent designation by Forel, 1922: 101. Myrmocamelus also described as new by Forel, 1915b: 102.]

Myrmocamelus as subgenus of Camponotus: Forel, 1914a: 261; Forel, 1915b: 102; Forel, 1917: 250; Forel, 1922: 101.

Myrmocamelus as junior synonym of Myrmophyma: Emery, 1920b: 257; Wheeler, W.M. 1921a: 18; Emery, 1925b: 109.

Subgenus CAMPONOTUS (MYRMOTREMA)

Myrmotrema Forel, 1912i: 91 [as subgenus of Camponotus]. Type-species: Camponotus foraminosus, by subsequent designation of Wheeler, W.M. 1913a: 81.

Taxonomic history

Myrmotrema as subgenus of Camponotus: Forel, 1912i: 91; Forel, 1914a: 262; Forel, 1917: 250; Emery, 1920b: 245; Arnold, 1922: 613; Wheeler, W.M. 1922a: 707; Emery, 1925b: 130; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOBRACHYS)

Myrmobrachys Forel, 1912i: 91 [as subgenus of Camponotus]. Type-species: Formica senex, by subsequent designation of Wheeler, W.M. 1913a: 81.

Taxonomic history

Myrmobrachys as subgenus of Camponotus: Forel, 1912i: 91; Forel, 1914a: 262; Forel, 1917: 251; Emery, 1920b: 251; Wheeler, W.M. 1922a: 709; Emery, 1925b: 161; all subsequent authors.

Subgenus CAMPONOTUS (MYRMENTOMA)

Myrmentoma Forel, 1912i: 92 [as subgenus of Camponotus]. Type-species: Formica lateralis, by subsequent designation of Wheeler, W.M. 1913a: 80.

Taxonomic history

Myrmentoma as junior synonym of Orthonotomyrmex: Forel, 1913b: 350 (in text); Forel, 1914a: 264; Wheeler, W.M. 1922a: 970.

Myrmentoma as subgenus of Camponotus: Forel, 1912i: 92; Emery, 1920b: 243; Wheeler, W.M. 1922a: 707; Emery, 1925a: 62; Emery, 1925b: 116; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOSPHINCTA)

Myrmosphincta Forel, 1912i: 92 [as subgenus of Camponotus]. Type-species: Formica sexguttata, by subsequent designation of Wheeler, W.M. 1913a: 81.

Taxonomic history

[Type-species not Formica cinerascens, unjustified subsequent designation by Forel, 1914a: 265; repeated in Arnold, 1922: 614.]

Myrmosphincta as subgenus of Camponotus: Forel, 1912i: 92; Forel, 1914a: 265; Forel, 1917: 251; Emery, 1920b: 238; Arnold, 1922: 614; Wheeler, W.M. 1922a: 708; Emery, 1925b: 151; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOTARSUS)

Myrmotarsus Forel, 1912i: 92 [as subgenus of Camponotus]. Type-species: Formica mistura, by subsequent designation of Wheeler, W.M. 1913a: 81.

Taxonomic history

[Type-species not Formica irritabilis, unjustified subsequent designation by Forel, 1914a: 260.]

Myrmotarsus as subgenus of Camponotus: Forel, 1912i: 92; Forel, 1914a: 260; Forel, 1917: 250; Emery, 1920b: 238; Wheeler, W.M. 1922a: 705; Emery, 1925b: 134; all subsequent authors.

Subgenus CAMPONOTUS (MYRMEPOMIS)

Myrmepomis Forel, 1912i: 92 [as subgenus of Camponotus]. Type-species: Formica sericeiventris, by subsequent designation of Wheeler, W.M. 1913a: 81.

Taxonomic history

[Type-species not Formica fulvopilosa, unjustified subsequent designation by Forel, 1914a: 265; repeated in Arnold, 1922: 614.]

Myrmepomis as subgenus of Camponotus: Forel, 1912i: 92; Forel, 1914a: 265; Forel, 1917: 251; Emery, 1920b: 245; Arnold, 1922: 614; Wheeler, W.M. 1922a: 705; Emery, 1925b: 170; all subsequent authors.

Junior synonym of CAMPONOTUS (MYRMEPOMIS)

Myrmolophus Emery, 1920b: 237 [as subgenus of Camponotus]. Type-species: Formica sericeiventris, by original designation.

Taxonomic history

Myrmolophus as subgenus of Camponotus: Emery, 1920b: 237.

Myrmolophus as junior synonym of Myrmepomis: Wheeler, W.M. 1921a: 17; Wheeler, W.M. 1922a: 705; Emery, 1925b: 170. [Myrmepomis and Myrmolophus share the same type-species, synonymy is therefore absolute.]

Subgenus CAMPONOTUS (MYRMOGONIA)

Myrmogonia Forel, 1912i: 92 [as subgenus of Camponotus]. Type-species: Camponotus laminatus, by subsequent designation of Wheeler, W.M. 1913a: 81.

Taxonomic history

Myrmogonia as subgenus of Camponotus: Forel, 1912i: 92; Forel, 1914a: 261; Forel, 1917: 250; Emery, 1920b: 240; Wheeler, W.M. 1922a: 706; Emery, 1925b: 143; all subsequent authors.

Subgenus CAMPONOTUS (MYRMEURYNOTA)

Myrmeurynota Forel, 1912i: 92 [as subgenus of Camponotus]. Type-species: Camponotus eurynotus, by subsequent designation of Wheeler, W.M. 1913a: 81.

Taxonomic history

[Type-species not Camponotus gilviventris, unjustified subsequent designation by Forel, 1914a: 266.]

Myrmeurynota as subgenus of Camponotus: Forel, 1912i: 92; Forel, 1914a: 266; Forel, 1917: 251; Emery, 1920b: 251; Wheeler, W.M. 1922a: 710; Emery, 1925b: 167; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOPSAMMA)

Myrmopsamma Forel, 1914a: 261 [as subgenus of Camponotus]. Type-species: Camponotus mystaceus, by original designation.

Taxonomic history

Myrmopsamma as subgenus of Camponotus: Forel, 1914a: 261; Forel, 1917: 250; Emery, 1920b: 246; Arnold, 1922: 612; Wheeler, W.M. 1922a: 708; Emery, 1925b: 70; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOMALIS)

Myrmomalis Forel, 1914a: 263 [as subgenus of Camponotus]. Type-species: Camponotus depressus, by original designation.

Taxonomic history

Myrmomalis as subgenus of Camponotus: Forel, 1914a: 263; Forel, 1917: 251; Emery, 1920b: 251; Wheeler, W.M. 1922a: 710; Emery, 1925b: 169; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOPLATYS)

Myrmoplatys Forel, 1916: 460 [as subgenus of Camponotus]. Type-species: Camponotus korthalsiae, by original designation.

Taxonomic history

Myrmoplatys as genus: Wheeler, W.M. 1936e: 219 (in text); Donisthorpe, 1943f: 670.

Myrmoplatys as subgenus of Camponotus: Forel, 1916: 460; Forel, 1917: 251; Emery, 1920b: 238; Wheeler, W.M. 1922a: 705; Emery, 1925b: 135; Bolton, 1994: 50.

Subgenus CAMPONOTUS (MYRMAPHAENUS)

Myrmaphaenus Emery, 1920b: 237 [as subgenus of Camponotus]. Type-species: Camponotus leydigi, by original designation.

Taxonomic history

Myrmaphaenus as subgenus of Camponotus: Emery, 1920b: 237; Wheeler, W.M. 1922a: 705; Emery, 1925b: 152; all subsequent authors.

Junior synonyms of CAMPONOTUS (MYRMAPHAENUS)

Paracolobopsis Emery, 1920b: 249 [as subgenus of Camponotus]. Type-species: Camponotus salvini, by original designation.

Taxonomic history

Paracolobopsis as subgenus of Camponotus: Emery, 1920b: 249; Wheeler, W.M. 1922a: 709.

Paracolobopsis as junior synonym of Myrmaphaenus: Emery, 1925b: 152; Kempf, 1972a: 44.

Neomyrmamblys Wheeler, W.M. 1921a: 19 [as subgenus of Camponotus]. Type-species: Camponotus fastigatus, by subsequent designation of Santschi, 1921f: 311.

Taxonomic history

Neomyrmamblys as subgenus of Camponotus: Wheeler, W.M. 1921a: 19; Wheeler, W.M. 1922a: 708.

Neomyrmamblys as junior synonym of Myrmaphaenus: Emery, 1925b: 152; Kempf, 1972a: 44.

Subgenus CAMPONOTUS (MYRMONESITES)

Myrmonesites Emery, 1920b: 242 [as subgenus of Camponotus]. Type-species: Camponotus putatus, by original designation.

Taxonomic history

Myrmonesites as subgenus of Camponotus: Emery, 1920b: 242; Wheeler, W.M. 1922a: 706; Emery, 1925b: 123.

[Myrmensites Donisthorpe, 1943f: 666, incorrect subsequent spelling.]

Subgenus CAMPONOTUS (MYRMOPYTIA)

Myrmopytia Emery, 1920b: 243 [as subgenus of Camponotus]. Type-species: Camponotus imitator, by original designation.

Taxonomic history

Myrmopytia as subgenus of Camponotus: Emery, 1920b: 243; Wheeler, W.M. 1922a: 707; Emery, 1925b: 114; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOTEMNUS)

Myrmotemnus Emery, 1920b: 246 [as subgenus of Camponotus]. Type-species: Camponotus moeschi, by original designation.

Taxonomic history

Myrmotemnus as junior synonym of Myrmamblys: Wheeler, W.M. 1921a: 19; Wheeler, W.M. 1922a: 708; Emery, 1925b: 137; Donisthorpe, 1943f: 671 (anachronism).

Myrmotemnus as subgenus of Camponotus: Emery, 1920b: 246; Santschi, 1926c: 601.

Subgenus CAMPONOTUS (PSEUDOCOLOBOPSIS)

Pseudocolobopsis Emery, 1920b: 249 [as subgenus of Camponotus]. Type-species: Camponotus macrocephalus, by original designation.

Taxonomic history

Pseudocolobopsis as subgenus of Camponotus: Emery, 1920b: 249; Wheeler, W.M. 1922a: 709; Emery, 1925b: 157; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOSTENUS)

Myrmostenus Emery, 1920b: 250 [as subgenus of Camponotus]. Type-species: Camponotus mirabilis, by original designation.

Taxonomic history

Myrmostenus as subgenus of Camponotus: Emery, 1920b: 250; Wheeler, W.M. 1922a: 709; Emery, 1925b: 161; all subsequent authors.

Subgenus CAMPONOTUS (HYPERCOLOBOPSIS)

Hypercolobopsis Emery, 1920b: 250 [as subgenus of Camponotus]. Type-species: Colobopsis paradoxa, by original designation.

Taxonomic history

Hypercolobopsis as subgenus of Camponotus: Emery, 1920b: 250; Wheeler, W.M. 1922a: 709; Emery, 1925b: 160; all subsequent authors.

Junior synonym of CAMPONOTUS (HYPERCOLOBOPSIS)

Neocolobopsis Borgmeier, 1928b: 65 [as subgenus of Camponotus]. Type-species: Camponotus (Neocolobopsis) scrobifer (junior synonym of Camponotus coriolanus), by original designation.

Taxonomic history

Neocolobopsis as subgenus of Camponotus Borgmeier, 1928b: 65.

Neocolobopsis as junior synonym of Hypercolobopsis: Kempf, 1968b: 411; Kempf, 1972a: 43.

Subgenus CAMPONOTUS (MYRMOPIROMIS)

Myrmopiromis Wheeler, W.M. 1921a: 17 [as subgenus of Camponotus]. Type-species: Formica fulvopilosa, by subsequent designation of Wheeler, W.M. 1922a: 707.

Taxonomic history

Myrmopiromis as subgenus of Camponotus: Wheeler, W.M. 1921a: 17; Wheeler, W.M. 1922a: 707; Emery, 1925b: 1127.

Subgenus CAMPONOTUS (MYRMOSAULUS)

Myrmosaulus Wheeler, W.M. 1921a: 18 [as subgenus of Camponotus]. Type-species: Formica cinerascens, by subsequent designation of Wheeler, W.M. 1922a: 705.

Taxonomic history

[Type-species not Formica singularis, unjustified subsequent designation by Emery, 1925b: 113, repeated in Donisthorpe, 1943f: 670. Note that Donisthorpe, 1932c: 445, states that C. cinerascens belongs in subgenus Myrmosericus. If so then Myrmosaulus would fall as a junior synonym of Myrmosericus.]

Myrmosaulus as subgenus of Camponotus: Wheeler, W.M. 1921a: 18; Wheeler, W.M. 1922a: 705; Emery, 1925b: 112; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOCLADOECUS)

Myrmocladoecus Wheeler, W.M. 1921a: 19 [as subgenus of Camponotus]. Type-species: Camponotus latangulus, by subsequent designation of Wheeler, W.M. 1922a: 709.

Taxonomic history

Myrmocladoecus as subgenus of Camponotus: Wheeler, W.M. 1921a: 19; Wheeler, W.M. 1922a: 709; Emery, 1925b: 166; all subsequent authors.

Subgenus CAMPONOTUS (MANNIELLA)

Manniella Wheeler, W.M. 1921a: 19 [as subgenus of Camponotus]. Type-species: Camponotus sphaericus, by original designation.

Taxonomic history

Manniella subgenus of Camponotus: Wheeler, W.M. 1921a: 19; Wheeler, W.M. 1922a: 710; Emery, 1925b: 159; all subsequent authors.

Subgenus CAMPONOTUS (MYRMISOLEPIS)

Myrmisolepis Santschi, 1921f: 310 [as subgenus of Camponotus]. Type-species: Camponotus epinotalis, by original designation.

Taxonomic history

Myrmisolepis as subgenus of Camponotus: Santschi, 1921f: 310; Emery, 1925b: 133; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOPELTA)

Myrmopelta Santschi, 1921f: 310 [as subgenus of Camponotus]. Type-species: Camponotus arminius, by original designation.

Taxonomic history

Myrmopelta as junior synonym of Myrmamblys: Emery, 1925b: 137.

Myrmopelta as subgenus of Camponotus: Santschi, 1921f: 310; Wheeler, W.M. 1922a: 704 (footnote); Santschi, 1926a: 16; Santschi, 1926c: 603; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOPLATYPUS)

Myrmoplatypus Santschi, 1921f: 311 [as subgenus of Camponotus]. Type-species: Camponotus platytarsus, by original designation.

Taxonomic history

Myrmoplatypus as subgenus of Camponotus: Santschi, 1921f: 311; Emery, 1925b: 169; all subsequent authors.

Subgenus CAMPONOTUS (MYRMEPINOTUS)

Myrmepinotus Santschi, 1921f: 312 [as subgenus of Camponotus]. Type-species: Camponotus echinoploides, by original designation.

Taxonomic history

Myrmepinotus as subgenus of Camponotus: Santschi, 1921f: 312; Emery, 1925b: 126; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOXYGENYS)

Myrmoxygenys Emery, 1925b: 70 [as subgenus of Camponotus]. Type-species: Camponotus caesar, by monotypy.

Taxonomic history

Myrmoxygenys as subgenus of Camponotus: Emery, 1925b: 70; all subsequent authors.

Subgenus CAMPONOTUS (KARAVAIEVIA)

Karavaievia Emery, 1925b: 115 [as subgenus of Camponotus]. Type-species: Camponotus exsectus, by original designation.

Taxonomic history

Karavaievia as subgenus of Camponotus: Emery, 1925b: 115; all subsequent authors.

Subgenus CAMPONOTUS (MYRMODIRACHIS)

Myrmodirachis Emery, 1925b: 168 [as subgenus of Camponotus]. Type-species: Camponotus heathi, by original designation.

Taxonomic history

Myrmodirachis as subgenus of Camponotus: Emery, 1925b: 168; all subsequent authors.

Subgenus CAMPONOTUS (MYRMESPERA)

Myrmespera Santschi, 1926b: 247 [as subgenus of Camponotus]. Type-species: Camponotus (Myrmespera) debellator, by subsequent designation of Santschi, 1926c: 604.

Taxonomic history

Myrmespera as subgenus of Camponotus: Santschi, 1926b: 247; Santschi, 1926c: 603; all subsequent authors.

Subgenus CAMPONOTUS (PARAMYRMAMBLYS)

Paramyrmamblys Santschi, 1926c: 604 [as subgenus of Camponotus].Type-species: Camponotus (Myrmamblys) ostiarius, by original designation.

Taxonomic history

Paramyrmamblys as subgenus of Camponotus: Santschi, 1926c: 604; all subsequent authors.

Subgenus CAMPONOTUS (MYRMACRHAPHE)

Myrmacrhaphe Santschi, 1926c: 607 [as subgenus of Camponotus]. Type-species: Camponotus conradti, by original designation.

Taxonomic history

Myrmacrhaphe as subgenus of Camponotus: Santschi, 1926c: 607; all subsequent authors.

Subgenus CAMPONOTUS (THLIPSEPINOTUS)

Thlipsepinotus Santschi, 1928e: 483 [as subgenus of Camponotus]. Type-species: Camponotus claripes, by original designation.

Taxonomic history

Thlipsepinotus as subgenus of Camponotus: Santschi, 1928e: 483; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOPALPELLA)

Myrmopalpella Stärcke, 1934: 30 [as subgenus of Camponotus]. Type-species: Camponotus megalonyx, by monotypy.

Taxonomic history

Myrmopalpella as genus: Wheeler, W.M. 1936e: 219 (in text); Donisthorpe, 1943f: 669.

Myrmopalpella as subgenus of Camponotus: Bolton, 1995b: 37.

Genus Camponotus references

Mayr, 1861: 35 (Europe species key); Roger, 1863b: 1, 9 (Camponotus, Colobopsis catalogues); Mayr, 1863: 397, 403 (Camponotus, Colobopsis catalogues); Mayr, 1865: 6, 7 (Camponotus, Colobopsis diagnosis); Mayr, 1867a: 67 (Colobopsis diagnois); Mayr, 1868c: 27 (†Baltic Amber species key); Mayr, 1870a: 373 (Colombia + Panama (= New Grenada) species key); Mayr, 1870b: 941 (Colobopsis species key); André, 1874: 175 (Europe species key); Forel, 1874: 38 (Switzerland species key); Mayr, 1876: 58 (Australia species key); Mayr, 1878: 869 (all Dendromyrmex species key (as part of Camponotus)); Forel, 1878: 367, 368 (Camponotus, Colobopsis diagnoses); André, 1882a: 138 (Europe & Algeria species key); André, 1882b: 159 (Europe & Algeria Colobopsis species key); Cresson, 1887: 255 (U.S.A. Camponotus, Colobopsis catalogue); Provancher, 1887: 228 (Canada species key); Nasonov, 1889: 57 (Russia species key); Forel, 1891b: 71, 216 (Madagascar species key); Forel, 1892j: 223 (India & Sri Lanka species key); Dalla Torre, 1893: 219 (Rhinomyrmex, Mayria, Camponotus catalogues); Emery, 1893i: 667 (North America species key); Forel, 1893b: 435 (India & Sri Lanka C. (Colobopsis) species key); Emery, 1895j: 772 (synoptic classification); Emery, 1896d: 370, 378 (species groups, catalogue); Emery, 1898a: 225 (additions to 1896a catalogue); Emery, 1903 (South America species key); Bingham, 1903: 343, 348 (India, Sri Lanka & Burma Colobopsis, Camponotus species keys); Ruzsky, 1905b: 185 (Russian Empire species key); Emery, 1908a: 183 (Palaearctic C. herculeanus & C. maculatus groups species key); Bondroit, 1910: 486 (Belgium species key); Wheeler, W.M. 1910d: 297 (North America species key); Forel, 1914a: 259, 266 (subgenera diagnoses, species listed by subgenus); Stitz, 1914: 94 (Central Europe species key); Santschi, 1915c: 266 (C. (Myrmotrema) species key); Emery, 1916b: 227 (Italy species key); Wheeler, W.M. 1916m: 600 (U.S.A., Connecticut species key); Mann, 1916: 489 (Dendromyrmex, all species key); Bondroit, 1918 (France & Belgium species key); Emery, 1920b: 235 (subgenera, diagoses); Emery, 1920c: 8 (Old World C. maculatus group, checklist & key); Wheeler, W.M. 1921a: 16 (subgenera, notes); Santschi, 1921f: 310 (subgenera, notes); Mann, 1921: 489, 499 (Fiji Is C. (Myrmogonia) & C. (Colobopsis) species keys); Wheeler, W.M. 1922a: 704 (subgenera synopsis); Wheeler, W.M. 1922a: 232, 948, 1039 (diagnosis, Afrotropical, Malagasy catalogues); Arnold, 1922: 610, 612, 619, 662, 673 (diagnosis, South Africa subgenera synopsis, South Africa C. (Tanaemyrmex), C. (Myrmophyma), C. (Myrmopsamma) species keys); Arnold, 1924: 683, 700, 709, 724, 735 (South Africa C. (Myrmosericus), C. (Myrmamblys), C. (Orthonotomyrmex), C. (Myrmotrema), C. (Myrmepomis) species keys); Emery, 1925a: 62 (Palaearctic C. lateralis group, key); Emery, 1925b: 59 (diagnosis, subgenera key, catalogue); Emery, 1925b: 69 (C. (Dinomyrmex) diagnosis, catalogue); Emery, 1925b: 70 (C. (Myrmoxygenys) & C. (Myrmopsamma) diagnoses, catalogues); Emery, 1925b: 71 (C. (Camponotus) diagnosis, catalogue); Emery, 1925b: 75 (C. (Tanaemyrmex) diagnosis, catalogue); Emery, 1925b: 104 (C. (Myrmosericus) diagnosis, catalogue); Emery, 1925b: 107 (C. (Myrmothrix) diagnosis, catalogue); Emery, 1925b: 109 (C. (Myrmophyma) diagnosis, catalogue); Emery, 1925b: 112 (C. (Myrmosaulus) diagnosis, catalogue); Emery, 1925b: 114 (C. (Myrmopytia) diagnosis, catalogue); Emery, 1925b: 115 (C. (Karavaievia) diagnosis, catalogue); Emery, 1925b: 116 (C. (Myrmentoma) diagnosis, catalogue); Emery, 1925b: 121 (C. (Mayria) diagnosis, catalogue); Emery, 1925b: 123 (C. (Myrmonesites) diagnosis, catalogue); Emery, 1925b: 124 (C. (Orthonotomyrmex) diagnosis, catalogue); Emery, 1925b: 126 (C. (Myrmepinotus) diagnosis, catalogue); Emery, 1925b: 127 (C. (Myrmopiromis) diagnosis, catalogue); Emery, 1925b: 130 (C. (Myrmotrema) diagnosis, catalogue); Emery, 1925b: 133 (C. (Myrmisolepis) diagnosis, catalogue); Emery, 1925b: 134 (C. (Myrmotarsus) diagnosis, catalogue); Emery, 1925b: 135 (C. (Myrmoplatys) diagnosis, catalogue); Emery, 1925b: 136 (C. (Myrmamblys) diagnosis, catalogue); Emery, 1925b: 142 (C. (Rhinomyrmex) diagnosis, catalogue); Emery, 1925b: 143 (C. (Myrmogonia) diagnosis, catalogue); Emery, 1925b: 144 (C. (Colobopsis) diagnosis, catalogue); Emery, 1925b: 151 (C. (Myrmosphincta) diagnosis, catalogue); Emery, 1925b: 152 (C. (Myrmaphaenus) diagnosis, catalogue); Emery, 1925b: 157 (C. (Pseudocolobopsis) diagnosis, catalogue); Emery, 1925b: 159 (C. (Manniella) diagnosis, catalogue); Emery, 1925b: 160 (C. (Hypercolobopsis) diagnosis, catalogue); Emery, 1925b: 161 (C. (Myrmostenus) & C. (Myrmobrachys) diagnoses, catalogues); Emery, 1925b: 166 (C. (Myrmocladoecus) diagnosis, catalogue); Emery, 1925b: 167 (C. (Myrmeurynota) diagnosis, catalogue); Emery, 1925b: 168 (C. (Myrmodirachis) diagnosis, catalogue); Emery, 1925b: 169 (C. (Myrmomalis) & C. (Myrmoplatypus) diagnoses, catalogues); Emery, 1925b: 170 (C. (Myrmepomis) diagnosis, catalogue); Emery, 1925b: 172 (Dendromyrmex diagnosis, catalogue); Santschi, 1926a: 16, 21 (C. (Myrmopelta) diagnosis, species key); Karavaiev, 1927c: 275 (Ukraine species key); Arnol'di, 1933b: 601 (Russia species key); Menozzi, 1933b: 81 (Israel species key); Menozzi, 1935c: 330 (Chile species key); Karavaiev, 1936: 175 (Ukraine species key); Finzi, 1936: 190 (Egypt species key); Menozzi, 1939a: 316 (Himalaya & Tibet species key); Stitz, 1939: 236 (Germany species key); Kratochvíl, 1941: 109 (Central Europe species key); Novák & Sadil, 1941: 109 (Central Europe species key); Cole, 1942: 387 (U.S.A., Utah species key); Donisthorpe, 1932g: 248 (C. (Orthonotomyrmex) notes); Holgersen, 1943b: 174 (Norway species key); Holgersen, 1944: 199 (Norway species key); Buren, 1944a: 293 (U.S.A., Iowa species key); Creighton, 1950a: 362 (North America species keys); Chapman & Capco, 1951: 220 (Asia checklist); Yasumatsu & Brown, 1951: 29 (C. herculeanus complex); Kusnezov, 1952d: 192 (Argentina species key); Yasumatsu & Brown, 1957: 45 (C. herculeanus complex); Pisarski, 1961a: 175 (Poland species key); Gregg, 1963: 655 (U.S.A., Colorado species key); Wheeler, G.C. & Wheeler, J. 1963: 163 (U.S.A., North Dakota species key); Arnol'di, 1967: 1817 (former U.S.S.R. C. (Camponotus) species key); Bernard, 1967: 330 (diagnosis, Western Europe species key); Kempf, 1972a: 42, 95 (Neotropical catalogue); Hashmi, 1973: 1 (C. (Myrmothrix) all species revision, key); Alayo, 1974: 27 (Cuba species key); Snelling & Hunt, 1976: 117 (Chile species key); Tarbinsky, 1976: 144 (Kyrghyzstan species key); Boven, 1977: 130 (Belgium species key); Kutter, 1977c: 191 (Switzerland species key); Arnol'di & Dlussky, 1978: 551 (former European U.S.S.R. species key); Collingwood, 1978: 90 (Iberian Peninsula species key); Collingwood, 1979: 87 (Fennoscandia & Denmark species key); Smith, D.R. 1979: 1424 (North America catalogue); Allred, 1982: 445 (U.S.A., Utah species key); Gösswald, 1985: 264 (Germany species key); Collingwood, 1985: 275 (Saudi Arabia species key); Taylor & Brown, D.R. 1985: 109 (Australia catalogue); Dumpert, 1986: 571 (C. (Karavaievia) species key); Wheeler, G.C. & Wheeler, J. 1986g: 60 (U.S.A., Nevada species key); Agosti & Collingwood, 1987b: 282 (Balkans species key); Taylor, 1987a: 10, 16 (Australia, New Caledonia, New Zealand checklists); Snelling, R.R. 1988: 57 (Nearctic C. (Myrmentoma) species key); Wang, C., Xiao & Wu, 1989a: 222 (China species key); Dlussky, Soyunov & Zabelin, 1990: 125 (Turkmenistan species key); Kupyanskaya, 1990: 164 (Far Eastern Russia species key); Brandão, 1991: 332 (Neotropical catalogue); Morisita, Kubota, Onoyama, et al. 1991: 38 (Japan species key); Terayama, 1991: 165 (Japan C. (Paramyrmamblys) species); Atanasov & Dlussky, 1992: 207 (Bulgaria species key); Wang, C. & Wu, 1994: 28 (China species key); Radchenko, 1994b: 116 (South Siberia species key); Arakelian, 1994: 82 (Armenia species key); Bolton, 1994: 50 (synoptic classification); Bolton, 1995a: 1048 (census); Bolton, 1995b: 83, 169 (catalogue); Douwes, 1995: 92 (Sweden species key); Dumpert, Maschwitz, Weissflog et al. 1995: 104 (C. (Karavaievia) species key); Kupyanskaya, 1995: 352 (Far Eastern Russia species key); Wu, J. & Wang, 1995: 168 (China species key); Collingwood & Agosti, 1996: 370 (Saudi Arabia species key); Seifert, 1996b: 170 (Central Europe species key); Radchenko, 1996b: 1195 (Asiatic Palaearctic species key); Cagniant, 1996b: 87 (Morocco species key); McArthur & Adams, 1996: 18 (Australia C. nigriceps group, key); Mackay, 1997: 194 (C. (Myrmostenus) species key); Mackay & Mackay, 1997b: 319 and Snelling, R.R. 2000: 607 (C. montivagus complex, keys); Robertson & Zachariades, 1997: 2 (C. fulvopilosus group, key); Snelling & Torres, 1998: 9 (Puerto Rico & Virgin Is species key); Collingwood & Prince, 1998: 24 (Portugal species key); Terayama, 1999b: 27 (Japan species key); Shattuck, 1999: 91 (Australia synopsis); Zhou, 2001b: 200 (China, Guangxi species key); McArthur & Shattuck, 2001: 27 (Australia C. macrocephalus group, key); Czechowski, Radchenko & Czechowska, 2002: 153 (Poland species key); Fernández, 2002: 58 (C. (Dendromyrmex) diagnosis, all species revision, key); Mackay & Mackay, 2002: 248 (U.S.A., New Mexico species key); Shattuck & McArthur, 2002: 65 (Australia C. wiederkehri group key); Bolton, 2003: 112 (synopsis); Shattuck, 2005: 3 (C. aureopilus group key); Radchenko, 2005b: 188 (North Korea species key); Hansen & Klotz, 2005: 75 (North American “Carpenter Ants” key); Snelling, R.R. 2006: 96 (U.S.A. C. festinatus complex, key); Karmaly & Narendran, 2006: 29 (India species, key); Dumpert, Maschwitz & Weissflog, 2006: 77 (C. (Karavaievia) species, keys); Seifert, 2007: 152 (North and Central Europe species key); McArthur, 2007a: 290 (Australia species key); Shattuck & Janda, 2009: 252 (C. aureopilus group key); Mackay & Delsinne, 2009: 496 (New World C. maculatus complex, key); Terayama, 2009: 216 (Taiwan species key); Heterick, 2009: 60 (south-western Australia species key); Ionescu-Hirsch, 2009: 59 (Israel species key); McArthur, 2010: 18 (south Australia species key).

Formicinae Formicini Cataglyphis TRUE TRUE FALSE

86 species, 20 subspecies

Cataglyphis Foerster, 1850b: 493. Type-species: Cataglyphis fairmairei (junior synonym of Formica bicolor), by monotypy.

Taxonomic history

Cataglyphis in Formicinae: Mayr, 1861: 44 [Formicidae]; Mayr, 1862: 653 (in key) [Formicidae]; Mayr, 1865: 8 [Formicidae].

Cataglyphis in Camponotinae, Formicini: Wheeler, W.M. 1910g: 144; Forel, 1912i: 89; Forel, 1917: 250.

Cataglyphis in Formicinae, Formica genus group: Agosti, 1991: 295.

Cataglyphis in Formicinae, Formicini: Bondroit, 1918: 38; Wheeler, W.M. 1922a: 699; Emery, 1925b: 261; Agosti, 1994a: 93; Bolton, 1994: 50; all subsequent authors.

Cataglyphis as junior synonym of Myrmecocystus: Forel, 1878: 372; Emery & Forel, 1879a: 449; André, 1882b: 165; Forel, 1886f: 201; Mayr, 1886d: 424; Dalla Torre, 1893: 216; Bingham, 1903: 312; Ruzsky, 1905b: 426; Emery, 1906d: 47; Emery, 1908g: 213; Karavaiev, 1924: 301.

Cataglyphis as subgenus of Myrmecocystus: Wheeler, W.M. 1910g: 144.

Cataglyphis as genus: Foerster, 1850b: 493; Roger, 1863b: 12; Mayr, 1863: 402; Wheeler, W.M. 1913a: 78; Emery, 1925b: 261; all subsequent authors.

Junior synonyms of CATAGLYPHIS

Monocombus Mayr, 1855: 381. Type-species: Formica viatica, by monotypy.

Taxonomic history

Monocombus in Formicinae: Mayr, 1855: 381 [Formicidae].

Monocombus as junior synonym of Myrmecocystus: Forel, 1878: 372; Emery & Forel, 1879a: 449; André, 1882b: 165; Mayr, 1886d: 424; Dalla Torre, 1893: 216.

Monocombus as subgenus of Cataglyphis: Santschi, 1929b: 29.

Monocombus as junior synonym of Cataglyphis: Mayr, 1861: 44; Roger, 1863b: 12; Mayr, 1863: 402; Wheeler, W.M. 1922a: 944; Emery, 1925b: 261; Agosti, 1990b: 1462; Agosti, 1994a: 103.

Paraformica Forel, 1915b: 95 (footnote) [as subgenus of Formica]. Type-species: Formica (Paraformica) emmae, by monotypy.

Taxonomic history

Paraformica in Camponotinae, Formicini: Forel, 1917: 250.

Paraformica in Formicinae, Formicini: Wheeler, W.M. 1922a: 699; Emery, 1925b: 260.

Paraformica as subgenus of Formica: Forel, 1915b: 95; Forel, 1917: 250; Wheeler, W.M. 1922a: 699.

Paraformica as genus: Emery, 1925b: 260; Donisthorpe, 1943f: 680.

Paraformica as subgenus of Proformica: Kuznetsov-Ugamsky, 1928a: 7.

Paraformica as subgenus of Cataglyphis: Santschi, 1925g: 353; Santschi, 1929b: 30.

Paraformica as junior synonym of Cataglyphis: Brown, 1973b: 183 [provisional]; Agosti, 1990b: 1462; Agosti, 1994a: 103.

Machaeromyrma Forel, 1916: 441 [as subgenus of Cataglyphis]. Type-species: Formica bombycina, by original designation.

Taxonomic history

Machaeromyrma as subgenus of Cataglyphis: Forel, 1916: 441; Forel, 1917: 250; Wheeler, W.M. 1922a: 699; Emery, 1925b: 266; Santschi, 1929b: 30; subsequent authors to the following.

Machaeromyrma as junior synonym of Cataglyphis: Baroni Urbani, 1969d: 218; Agosti, 1994a: 103.

Eomonocombus Arnol'di, 1968: 1815 [as subgenus of Cataglyphis]. Type-species: Myrmecocystus cinnamomeus, by original designation.

Taxonomic history

Eomonocombus as junior synonym of Cataglyphis: Brown, 1973b: 180 [provisional]; Agosti, 1990b: 1462; Agosti, 1994a: 103.

Genus Cataglyphis references

Mayr, 1861: 44 (Europe species key); Roger, 1863b: 12 (catalogue); Mayr, 1863: 402 (catalogue); Mayr, 1865: 8 (diagnosis); André, 1874: 181 (Europe species key); Forel, 1878: 372 (diagnosis); André, 1882b: 166 (Europe & Algeria species key); Nasonov, 1889: 60 (Russia species key); Bingham, 1903: 312 (diagnosis); Ruzsky, 1905b: 426 (Russian Empire species key); Emery, 1906d: 60 (Palaearctic species (as Myrmecocystus), key); Wheeler, W.M. 1922a: 699 (subgenera key); Wheeler, W.M. 1922a: 945 (Afrotropical catalogue); Emery, 1925b: 260 (Paraformica diagnosis, catalogue); Emery, 1925b: 261 (diagnosis, catalogue); Emery, 1925b: 266 (C. (Machaeromyrma) diagnosis, catalogue); Santschi, 1929b: 56 (all species key); Menozzi, 1933b: 86 (Israel species key); Kratochvíl, 1941: 108 (Central Europe species key); Chapman & Capco, 1951: 197, 203 (Asia checklist); Bernard, 1967: 280 (diagnosis); Tarbinsky, 1976: 197 (Kyrghyzstan species key); Arnol'di & Dlussky, 1978: 554 (former European U.S.S.R. species key); Collingwood, 1978: 92 (Iberian Peninsula species key); Agosti & Collingwood, 1987b: 284 (Balkans species key); Collingwood, 1985: 284 (Saudi Arabia species key); Agosti, 1990b: 1457 (review of genus, diagnosis, species groups key, catalogue); Dlussky, Soyunov & Zabelin, 1990: 146 (Turkmenistan species key); Atanasov & Dlussky, 1992: 292 (Bulgaria species key); Agosti, 1994a: 103 (diagnosis, review of genus); Arakelian, 1994: 103 (Armenia species key); Bolton, 1994: 50 (synoptic classification); Bolton, 1995a: 1048 (census); Bolton, 1995b: 135 (catalogue); Collingwood & Agosti, 1996: 375 (Saudi Arabia species key); Seifert, 1996b: 179 (Central Europe species key); Collingwood & Prince, 1998: 28 (Portugal species key); Radchenko, 1998: 502 (Asia species key); Radchenko, 2001: 885 (phylogeny); Chang & He, 2002b: 62 (northwest China species key); Bolton, 2003: 126 (synopsis); Seifert, 2007: 159 (North and Central Europe species key).

Formicinae Formicini Formica TRUE TRUE FALSE

Extant: 176 species, 24 subspecies

Fossil: 64 species

Formica Linnaeus, 1758: 579. Type-species: Formica rufa, by subsequent designation of Curtis, 1839: 752.

Taxonomic history

Formica in Formicites: Lepeletier de Saint-Fargeau, 1835: 199.

Formica in Formicidae: Smith, F. 1858b: 1.

Formica in Formicinae: Mayr, 1861: 45 [Formicidae]; Mayr, 1862: 653 (in key) [Formicidae]; Mayr, 1865: 8 [Formicidae]; Mayr, 1868c: 47 [Formicidae].

Formica in Camponotinae: Forel, 1878: 373 [Camponotidae]; Emery & Forel, 1879a: 450 [Camponotidae]; Dalla Torre, 1893: 192.

Formica in Camponotinae, Camponotini: Forel, 1886f: 205.

Formica in Camponotinae, Formicini: Forel, 1893a: 165; Emery, 1895j: 772; Forel, 1899c: 127; Wheeler, W.M. 1910g: 143; Forel, 1912i: 89; Wheeler, W.M. 1915h: 120; Forel, 1917: 250.

Formica in Formicinae, Formicini: Ashmead, 1905b: 384; Bondroit, 1918: 41; Wheeler, W.M. 1922a: 699; Emery, 1925b: 241; all subsequent authors except the following.

Formica in Formicinae, Formica genus group: Agosti, 1991: 295.

Junior synonyms of FORMICA

Formicina Shuckard, in Swainson & Shuckard, 1840: 172. Type-species: Formica rufa, by subsequent designation of Wheeler, W.M. 1911f: 164.

Taxonomic history

[Type-species not Formica flava, unjustified subsequent designation by Emery, 1916a: 61 (footnote); repeated by Emery, 1916b: 239 and Wheeler, W.M. 1916k: 170.]

Formicina in Camponotinae, Formicini: Forel, 1917: 249.

Formicina as genus: Emery, 1916a: 61; Emery, 1916b: 239; Forel, 1917: 249; Bondroit, 1918: 19.

Formicina as subgenus of Lasius: Wheeler, W.M. 1916k: 172.

Formicina as junior synonym of Lasius: Emery, 1925b: 226; Brown, 1973b: 180.

Formicina as junior synonym of Acanthomyops: Donisthorpe, 1927b: 209.

Formicina as junior synonym of Formica: Wheeler, W.M. 1911c: 860; Wheeler, W.M. 1911f: 164; Donisthorpe, 1943f: 646; Agosti, 1994a: 106; Bolton, 1994: 50. [Formica and Formicina share the same type-species, synonymy is therefore absolute.]

Neoformica Wheeler, W.M. 1913a: 82 [as subgenus of Formica]. Type-species: Formica pallidefulva, by original designation.

Taxonomic history

[Neoformica also described as new by Wheeler, W.M. 1913f: 548.]

Neoformica as subgenus of Formica: Wheeler, W.M. 1913a: 82; Wheeler, W.M. 1913f: 388 (in key); Forel, 1917: 250; Wheeler, W.M. 1922a: 700; Emery, 1925b: 244; Donisthorpe, 1943f: 673; Creighton, 1950a: 543.

Neoformica as junior synonym of Formica: Buren, 1968a: 39; Agosti, 1994a: 107.

Raptiformica Forel, 1913i: 361 [as subgenus of Formica]. Type-species: Formica sanguinea, by original designation.

Taxonomic history

Raptiformica as subgenus of Formica: Forel, 1917: 250; Emery, 1925b: 258; Donisthorpe, 1943g: 723; Creighton, 1950a: 460.

Raptiformica as junior synonym of Formica: Wheeler, W.M. 1922a: 699 (footnote); Smith, M.R. 1951a: 860; Smith, D.R. 1979: 1448; Agosti, 1994a: 107.

Serviformica Forel, 1913i: 361 [as subgenus of Formica]. Type-species: Formica fusca, by original designation.

Taxonomic history

Serviformica as subgenus of Formica: Forel, 1913i: 361; Forel, 1917: 250; Emery, 1925b: 245; Donisthorpe, 1943g: 725.

Serviformica as junior synonym of Formica: Wheeler, W.M. 1922a: 699 (footnote); Smith, M.R. 1951a: 860; Francoeur, 1973: 34; Smith, D.R. 1979: 1448; Agosti, 1994a: 106.

Coptoformica Müller, 1923: 146 [as subgenus of Formica]. Type-species: Formica exsecta, by subsequent designation of Donisthorpe, 1941f: 37.

Taxonomic history

Coptoformica as junior synonym of Formica: Smith, M.R. 1951a: 860; Smith, D.R. 1979: 1448; Agosti, 1994a: 107.

Adformica Lomnicki, 1925a: 164 [as subgenus of Formica]. Type-species: Formica exsecta, by subsequent designation of Donisthorpe, 1927b: 316.

Taxonomic history

Adformica as junior synonym of Coptoformica: Stitz, 1939: 306; Donisthorpe, 1943f: 620 [Adformica and Coptoformica share the same type-species, synonymy is therefore absolute.]

Iberoformica Tinaut, 1990a: 282 [as subgenus of Formica]. Type-species: Formica subrufa, by original designation.

Taxonomic history

Iberoformica as junior synonym of Formica: Agosti, 1994a: 107.

Genus Formica references

Mayr, 1855: 300 (diagnosis, Austria species key); Smith, F. 1858b: 1 (diagnosis); Mayr, 1861: 46 (Europe species key); Roger, 1863b: 12 (catalogue); Mayr, 1863: 410 (catalogue); Mayr, 1865: 8 (diagnosis); André, 1874: 182 (Europe species key); Forel, 1874: 51 (Switzerland species key); Forel, 1878: 373 (diagnosis); André, 1882b: 176 (Europe & Algeria species key); Cresson, 1887: 255 (U.S.A. catalogue); Provancher, 1887: 232 (Canada species key); Nasonov, 1889: 61 (Russia species key); Dalla Torre, 1893: 192 (catalogue); Emery, 1893i: 643 (North America species key); Forel, 1894c: 402 (India species key); Bingham, 1903: 334 (India species key); Ruzsky, 1905b: 318 (Russian Empire species key); Wasmann, 1906: 8 (Luxemburg species key); Emery, 1909b: 180 (Palaearctic species key); Bondroit, 1910: 482 (Belgium species key); Wheeler, W.M. 1913f: 387 (subgenera, species groups, all species keys); Stitz, 1914: 88 (Central Europe species key); Donisthorpe, 1915d: 244 (Britain species key); Emery, 1916b: 257 (Italy species key); Wheeler, W.M. 1916m: 594 (U.S.A., Connecticut species key); Bondroit, 1917a: 174 (France F. rufa group, key); Bondroit, 1918: 41 (France & Belgium species key); Wheeler, W.M. 1922a: 699 (subgenera key); Emery, 1925b: 241 (diagnosis, subgenera key, catalogue); Emery, 1925b: 244 (F. (Neoformica) diagnosis, catalogue; Emery, 1925b: 245 (F. (Serviformica) diagnosis, catalogue; Emery, 1925b: 251 (F. (Formica) diagnosis, catalogue; Emery, 1925b: 258 (F. (Raptiformica) diagnosis, catalogue; Lomnicki, 1925a: 171 (Poland species key); Kuznetsov-Ugamsky, 1926b: 93 (Turkestan species key); Karavaiev, 1927c: 282 (Ukraine species key); Donisthorpe, 1927b: 284 (Britain species key); Arnol'di, 1933b: 603 (Russia species key); Karavaiev, 1936: 220 (Ukraine species key); Menozzi, 1939a: 321 (Himalaya & Tibet species key); Stitz, 1939: 306 (Germany species key); Creighton, 1940a: 1 (North America F. rufa subspecies key); Kratochvíl, 1941: 103 (Central Europe species key); Novák & Sadil, 1941: 103 (Central Europe species key); Cole, 1942: 375 (U.S.A., Utah species key); Holgersen, 1943b: 175 (Norway species key); Holgersen, 1944: 200 (Norway species key); Buren, 1944a: 299 (U.S.A., Iowa species key); Boven, 1947: 187 (Belgium species key); Creighton, 1950a: 456 (North America species key); Chapman & Capco, 1951: 197 (Asia checklist); Yarrow, 1954a: 229 (Britain F. fusca group); Yarrow, 1955a: 29 (Britain F. rufa group, key); Wilson & Brown, 1955: 108 (F. sanguinea & F. neogagates groups, synopsis); Boven, 1959: 12 (Netherlands species key); Betrem, 1960b: 75 (F. rufa group, key); Gregg, 1963: 498 (U.S.A., Colorado species key); Wheeler, G.C. & Wheeler, J. 1963: 205 (U.S.A., North Dakota species key); Cotti, 1963: 1 (F. rufa group, bibliography 1930-61); Dlussky, 1964: 1026 (former U.S.S.R. F. (Coptoformica) species key); Collingwood, 1964: 109 (Britain species key); Brown, 1965d: 181 (F. integra subgroup); Dlussky, 1965a: 15 (Mongolia and Tibet species key); Dlussky, 1967a: 39 (Palaearctic species key); Dlussky, 1967b: 80 (†Baltic Amber species key); Bernard, 1967: 285 (diagnosis, Western Europe species key); Buren, 1968a: 36 (Nearctic F. sanguinea group species key); Boven, 1970b: 34 (Netherlands species key); Dlussky & Pisarski, 1971: 145 (Poland species key); Francoeur, 1973: 272 (Nearctic F. fusca group, revision, key); Bolton & Collingwood, 1975: 6 (Britain species key); Tarbinsky, 1976: 177 (Kyrghyzstan species key); Boven, 1977: 94 (Belgium species key); Kutter, 1977c: 236 (Switzerland species key); Arnol'di & Dlussky, 1978: 552 (former European U.S.S.R. species key); Collingwood, 1978: 93 (Iberian Peninsula species key); Collingwood, 1979: 112 (Fennoscandia & Denmark species key); Smith, D.R. 1979: 1448 (North America catalogue); Douwes, 1979: 187 (F. rufa group systematics); Kupyanskaya, 1980: 95 (Far Eastern Russia species key); Allred, 1982: 446 (U.S.A., Utah species key); Kupyanskaya, 1984: 103 (Far Eastern Russia F. truncorum complex, key); Snelling & Buren, 1985: 71 (Nearctic F. sanguinea group, key); Gösswald, 1985: 274 (Germany species key); Wheeler, G.C. & Wheeler, J. 1986g: 76 (U.S.A., Nevada species key); Nilsson & Douwes, 1987: 74 (Norway species key); Agosti & Collingwood, 1987b: 285 (Balkans species key); Gösswald, 1989: 49 (F. rufa group taxonomy); Agosti & Bolton, 1990b: 149 (characters); Wu, J. 1990: 4 (China species key); Dlussky, Soyunov & Zabelin, 1990: 138 (Turkmenistan species key); Kupyanskaya, 1990: 174 (Far Eastern Russia species key); Morisita, Kubota, Onoyama, et al. 1991: 31 (Japan species key); Atanasov & Dlussky, 1992: 255 (Bulgaria species key); Agosti, 1994a: 106 (diagnosis, review of genus); Radchenko, 1994b: 112 (South Siberia species key); Arakelian, 1994: 89 (Armenia species key); Bolton, 1994: 50 (synoptic classification); Bolton, 1995a: 1049 (census); Bolton, 1995b: 190 (catalogue); Douwes, 1995: 94 (Sweden species key); Kupyanskaya, 1995: 353 (Far Eastern Russia species key); Wu, J. & Wang, 1995: 138 (China species key); Seifert, 1996b: 201 (Central Europe species key); Skinner & Allen, 1996: 41 (Britain species key); Collingwood & Prince, 1998: 25 (Portugal species key); Tinaut, A. & Martínez-Ibánez, 1998: 36 (Iberian Peninsula F. rufa, F. sanguinea & F. exsecta groups, key); Seifert, 2000a: 563 (Palaearctic F. (Coptoformica) species key); Czechowski, Radchenko & Czechowska, 2002: 148 (Poland species key); Chang & He, 2002a: 51 (northwest China species key); Seifert, 2002b: 267 (F. cinerea group revision key); Mackay & Mackay, 2002: 277 (U.S.A., New Mexico species key); Bolton, 2003: 127 (synopsis); Radchenko, 2005b: 188 (North Korea species key); Seifert, 2007: 176 (North and Central Europe species key); Trager, MacGown & Trager, 2007: 627 (F. pallidefulva group key); Dlussky, 2008a: 57 (Europe Eocene amber species key); Terayama, 2009: 224 (Taiwan species key); Seifert & Schultz, 2009: 258 (F. rufibarbis group key).

Formicinae Formicini Polyergus TRUE TRUE FALSE

5 species, 5 subspecies

Polyergus Latreille, 1804: 179. Type-species: Formica rufescens, by monotypy.

Taxonomic history

Polyergus in Formicites: Lepeletier de Saint-Fargeau, 1835: 198.

Polyergus in Formicidae: Smith, F. 1858b: 1.

Polyergus in Formicinae: Mayr, 1855: 383 [Formicidae]; Mayr, 1861: 43 [Formicinae]; Mayr, 1862: 653 [Formicidae]; Mayr, 1865: 8 [Formicidae].

Polyergus in Camponotinae: Forel, 1878: 367 [Camponotidae]; Dalla Torre, 1893: 214.

Polyergus in Camponotinae, Camponotini: Forel, 1886f: 200.

Polyergus in Camponotinae, Formicini: Forel, 1893a: 165; Emery, 1895j: 772; Forel, 1899c: 128; Wheeler, W.M. 1910g: 143; Forel, 1912i: 89; Forel, 1917: 250.

Polyergus in Formicinae, Formicini: Ashmead, 1905b: 384; Bondroit, 1918: 41; Wheeler, W.M. 1922a: 699; Emery, 1925b: 267; all subsequent authors except the following.

Polyergus in Formicinae, Formica genus group: Agosti, 1991: 295.

Genus Polyergus references

Mayr, 1855: 383 (diagnosis); Roger, 1863b: 12 (catalogue); Mayr, 1863: 443 (catalogue); Mayr, 1865: 8 (diagnosis); Forel, 1878: 373 (diagnosis); André, 1882b: 162 (Europe & Algeria species); Cresson, 1887: 255 (U.S.A. catalogue); Dalla Torre, 1893: 214 (catalogue); Emery, 1925b: 267 (diagnosis, catalogue); Buren, 1944a: 310 (U.S.A., Iowa species key); Smith, M.R. 1947g: 152 (U.S.A. species key); Creighton, 1950a: 556 (North America species key); Chapman & Capco, 1951: 203 (Asia checklist); Gregg, 1963: 632 (U.S.A., Colorado species key); Wheeler, G.C. & Wheeler, J. 1963: 274 (U.S.A., North Dakota species key); Bernard, 1967: 329 (diagnosis); Wheeler, J. 1968: 156 (Nearctic species, male genitalia); Smith, D.R. 1979: 1466 (North America catalogue); Wheeler, G.C. & Wheeler, J. 1986g: 95 (U.S.A., Nevada species key); Kupyanskaya, 1990: 207 (Far Eastern Russia species key); Agosti, 1994a: 109 (diagnosis, review of genus); Bolton, 1994: 50 (synoptic classification); Bolton, 1995a: 1052 (census); Bolton, 1995b: 342 (catalogue); Kupyanskaya, 1995: 361 (Far Eastern Russia species key); Mackay & Mackay, 2002: 359 (U.S.A., New Mexico species key); Bolton, 2003: 129 (synopsis).

Formicinae Formicini Proformica TRUE TRUE FALSE

25 species, 2 subspecies

Proformica Ruzsky, 1902d: 13 [as subgenus of Formica]. Type-species: Formica nasuta, by monotypy.

Taxonomic history

[Proformica also described as new by Ruzsky, 1903bb: 303.]

Proformica in Formicinae, Lasiini: Ashmead, 1905b: 384.

Proformica in Camponotinae, Formicini: Forel, 1917: 250.

Proformica in Formicinae, Formicini: Bondroit, 1918: 40; Wheeler, W.M. 1922a: 699; Emery, 1925b: 242; all subsequent authors except the following.

Proformica in Formicinae, Formica genus group: Agosti, 1991: 295.

Proformica as subgenus of Formica: Ruzsky, 1902d: 13; Wheeler, W.M. 1910g: 143; Forel, 1917: 250; Wheeler, W.M. 1922a: 699; Emery, 1925b: 242; Donisthorpe, 1943f: 687; Creighton, 1950a: 457.

Proformica as genus: Bondroit, 1918: 40; Kuznetsov-Ugamsky, 1927b: 26; Kuznetsov-Ugamsky, 1928a: 7; subsequent authors except the above.

Genus Proformica references

Emery, 1925b: 242 (Formica (Proformica) diagnosis, catalogue); Kuznetsov-Ugamsky, 1927b: 28 (Turkestan species key); Kuznetsov-Ugamsky, 1928a: 7 (central Asia species key); Bernard, 1967: 282 (diagnosis, Western Europe species key); Dlussky, 1969a: 231 (Russia and contiguous countries, key); Tarbinsky, 1976: 165 (Kyrghyzstan species key); Collingwood, 1978: 92 (Iberian Peninsula species key); Agosti & Collingwood, 1987b: 287 (Balkans species key); Atanasov & Dlussky, 1992: 287 (Bulgaria species key); Agosti, 1994a: 112 (diagnosis, review of genus); Radchenko, 1994b: 115 (South Siberia species key); Arakelian, 1994: 109 (Armenia species key); Bolton, 1994: 50 (synoptic classification); Bolton, 1995a: 1052 (census); Bolton, 1995b: 368 (catalogue); Collingwood & Prince, 1998: 27 (Portugal species key); Wu, J. & Wang, 1995: 148 (China species key); Bolton, 2003: 130 (synopsis); Dubovikov, 2005b: 190 (Caucasian Isthmus species key).

Pseudomyrmecinae Pseudomyrmecini Pseudomyrmex TRUE TRUE FALSE

Extant: 131 species, 10 subspecies

Fossil: 12 species

Pseudomyrmex Lund, 1831b: 106. Type-species: Formica gracilis, by subsequent designation of Smith, M.R. 1952a: 98.

Taxonomic history

Pseudomyrmex in Pseudomyrmecinae: Smith, M.R. 1952a: 98; all subsequent authors.

Junior synonyms of PSEUDOMYRMEX

Leptalea Erichson, 1839: 309. Type-species: Formica gracilis, by subsequent designation of Wheeler, W.M. 1911f: 166.

Taxonomic history

Leptalea in Pseudomyrmecinae: Donisthorpe, 1943f: 656.

Leptalea as junior synonym of Pseudomyrma: Smith, F. 1858b: 153; Roger, 1863b: 24; Smith, F. 1877b: 58; Dalla Torre, 1893: 55; Emery, 1921f: 28.

[Leptalaea Spinola, 1851b: 52 and Spinola, 1853: 68, incorrect subsequent spellings.]

[Pseudomyrmex and Leptalea share the same type-species; synonymy is therefore absolute.]

Pseudomyrma Guérin-Méneville, 1844a: 427 [as subgenus of Formica]. Type-species: Pseudomyrma bicolor, by monotypy.

Taxonomic history

Pseudomyrma in Poneridae, Myrmicidae: Smith, F. 1858b: 153.

Pseudomyrma in Myrmicidae: Smith, F. 1871a: 328; Smith, F. 1877b: 58; Cresson, 1887: 262.

Pseudomyrma in Myrmicidae, Myrmeciidae: Emery, 1877a: 81.

Pseudomyrma in Myrmicinae: Mayr, 1865: 24 [Myrmicidae]; Dalla Torre, 1893: 55.

Pseudomyrma in Myrmicidae, Pseudomyrminae: Ashmead, 1905b: 383.

Pseudomyrma in Myrmicinae, Pseudomyrmini: Forel, 1893a: 164; Emery, 1895j: 768; Forel, 1899c: 86; Wheeler, W.M. 1910g: 139; Emery, 1914a: 40; Forel, 1917: 240; Emery, 1921f: 28.

Pseudomyrma in Pseudomyrminae: Wheeler, W.M. 1922a: 654; Donisthorpe, 1941d: 27; Donisthorpe, 1943g: 722.

Pseudomyrma as junior synonym of Pseudomyrmex: Smith, M.R. 1952a: 98; Kempf, 1972a: 214; Ward, 1990: 469; Bolton, 1994: 184.

Myrmex Guérin-Méneville, 1844a: 427. Type-species: Myrmex perboscii, by monotypy.

Taxonomic history

[Junior homonym of Myrmex Sturm, 1826: 32 (Coleoptera).]

Myrmex Guérin-Méneville junior synonym of Pseudomyrma: Smith, F. 1858b: 153; Roger, 1863b: 24; Smith, F. 1877b: 58; Dalla Torre, 1893: 55; Emery, 1921f: 28.

Clavanoda Enzmann, E.V. 1944: 61 [as "branch" of Pseudomyrma]. Type-species: Formica gracilis, by original designation.

Taxonomic history

Clavanoda as junior synonym of Pseudomyrmex: Ward, 1990: 469. [Pseudomyrmex and Clavanoda share the same type-species, synonymy is therefore absolute.]

Ornatinoda Enzmann, E.V. 1944: 61 [as "branch" of Pseudomyrma]. Type-species: Formica tenuis, by original designation.

Taxonomic history

Ornatinoda as junior synonym of Pseudomyrmex: Ward, 1990: 469.

Triangulinoda Enzmann, E.V. 1944: 61 [as "branch" of Pseudomyrma]. Type-species: Pseudomyrma spinicola, by original designation.

Taxonomic history

Triangulinoda as junior synonym of Pseudomyrmex: Ward, 1990: 469.

Apedunculata Enzmann, E.V. 1944: 62 [as "branch" of Pseudomyrma]. Type-species: Pseudomyrma sericea, by original designation.

Taxonomic history

Apedunculata as junior synonym of Pseudomyrmex: Ward, 1990: 469.

Latinoda Enzmann, E.V. 1944: 62 [as "branch" of Pseudomyrma]. Type-species: Pseudomyrma latinoda, by original designation.

Taxonomic history

Latinoda as junior synonym of Pseudomyrmex: Ward, 1990: 469.

Genus Pseudomyrmex references

Smith, F. 1858b: 153 (diagnosis); Roger, 1863b: 24 (catalogue); Mayr, 1863: 426, 451 (Leptalea, Pseudomyrma catalogues); Mayr, 1865: 24 (diagnosis); Mayr, 1870a: 407 (Colombia + Panama (= New Grenada) species key); Smith, 1877b: 58 (checklist); Cresson, 1887: 262 (U.S.A. catalogue); Dalla Torre, 1893: 55 (catalogue); Emery, 1921f: 28 (diagnosis, catalogue); Gallardo, 1932a: 44 (Argentina species key); Enzmann, E.V. 1944: 62 (all species keys); Creighton, 1950a: 79 (North America species key); Kempf, 1961a: 373 (P. gracilis group, partial key); Kempf, 1972a: 215 (Neotropical catalogue); Alayo, 1974: 9 (Cuba species key); Smith, D.R. 1979: 1345 (North America catalogue); Ward, 1985b: 215 (Nearctic species key); Billen, 1986: 173 (Dufour's gland); Ward, 1989: 407 (P. oculatus group, key); Ward, 1989: 430 (P. subtilissimus group, key); Ward, 1990: 469 (diagnosis, review of genus); Brandão, 1991: 373 (catalogue); Ward, 1992: 76 (†Dominican Amber species key); Ward, 1993: 130 (swollen-thorn Acacia-associated species key); Bolton, 1994: 184 (synoptic classification); Bolton, 1995a: 1052 (census); Bolton, 1995b: 370 (catalogue); Ward, 1999b: 473 (P. viduus group, and other Tachigali- and Triplaris-associated species key); Mackay & Mackay, 2002: 38 (U.S.A., New Mexico species key).

Cerapachyinae Cerapachyini Cerapachys TRUE TRUE FALSE

146 species

Cerapachys Smith, F. 1857a: 74. Type-species: Cerapachys antennatus, by subsequent designation of Bingham, 1903: 28.

Taxonomic history

Cerapachys in Myrmicinae: Smith, F. 1857a: 74 [Myrmicidae]; Smith, F. 1861b: 47 [Myrmicidae]; Smith, F. 1871a: 329 [Myrmicidae].

Cerapachys in Ponerinae: Dalla Torre, 1893: 17.

Cerapachys in Dorylinae, Cerapachyini: Emery, 1895j: 765; Emery, 1901a: 34.

Cerapachys in Ponerinae, Cerapachyini: Forel, 1893a: 162 (also misspelled Orapachys, same page); Forel, 1900d: 331; Ashmead, 1905b: 382; Wheeler, W.M. 1910g: 137; Emery, 1911d: 8; Arnold, 1915: 11; Forel, 1917: 239; Brown, 1975: 18; Snelling, R.R. 1981: 389; Dlussky & Fedoseeva, 1988: 79; Hölldobler & Wilson, 1990: 10.

Cerapachys in Cerapachyinae, Cerapachyini: Wheeler, W.M. 1902d: 185; Wheeler, W.M. 1922a: 639; Donisthorpe, 1943f: 632; Chapman & Capco, 1951: 17; Kempf, 1972a: 76; Wheeler, G.C. & Wheeler, J. 1985: 256; Bolton, 1990a: 67; Bolton, 1990c: 1357; Jaffe, 1993: 9; Bolton, 1994: 19; Bolton, 2003: 34.

Junior synonyms of CERAPACHYS

Syscia Roger, 1861a: 19. Type-species: Syscia typhla, by monotypy.

Taxonomic history

Syscia in Ponerinae: Mayr, 1862: 714 (in key) [Poneridae]; Mayr, 1865: 15 [Poneridae]; Dalla Torre, 1893: 17.

Syscia in Dorylinae, Cerapachyini: Emery, 1895j: 765.

Syscia in Ponerinae, Cerapachyini: Forel, 1893a: 162; Forel, 1900d: 329; Ashmead, 1905b: 382.

Syscia as genus: Dalla Torre, 1893: 17; Forel, 1900d: 329; Bingham, 1903: 31.

Syscia as subgenus of Cerapachys: Wheeler, W.M. 1902d: 185; Emery, 1902c: 24; Wheeler, W.M. 1910g: 137; Emery, 1911d: 10; Forel, 1917: 239; Wheeler, W.M. 1922a: 639; Donisthorpe, 1943g: 730.

Syscia as junior synonym of Cerapachys: Kempf, 1972a: 76; Brown, 1975: 18; all subsequent authors.

Ooceraea Roger, 1862a: 248. Type-species: Ooceraea fragosa, by monotypy.

Taxonomic history

Ooceraea in Myrmicinae: Mayr, 1865: 24 [Myrmicidae].

Ooceraea in Myrmicidae, Pheidolidae: Emery, 1877a: 81.

Ooceraea in Poneridae: Smith, F. 1871a: 324.

Ooceraea in Ponerinae: Dalla Torre, 1893: 17.

Ooceraea in Ponerinae, Cerapachyini: Forel, 1893a: 162.

Ooceraea in Dorylinae, Cerapachyini: Emery, 1895j: 765; Emery, 1901a: 34.

Ooceraea in Ecitoninae, Ecitonini: Ashmead, 1905b: 381; Ashmead, 1906: 25.

Ooceraea as genus: Roger, 1862a: 248; Smith, F. 1871a: 324; Dalla Torre, 1893: 17; Forel, 1900d: 329; Bingham, 1903: 31.

Ooceraea as subgenus of Cerapachys: Emery, 1902c: 24; Wheeler, W.M. 1902d: 185; Wheeler, W.M. 1910g: 137; Emery, 1911d: 10; Forel, 1917: 239; Wheeler, W.M. 1922a: 639; Donisthorpe, 1943f: 678.

Ooceraea as junior synonym of Cerapachys: Brown, 1973b: 183 [provisional]; Brown, 1975: 19; all subsequent authors.

Lioponera Mayr, 1879: 666. Type-species: Lioponera longitarsus, by monotypy.

Taxonomic history

Lioponera in Ponerinae: Dalla Torre, 1893: 17.

Lioponera in Pachycondylinae, Lioponerini: Ashmead, 1905b: 382.

Lioponera in Ponerinae, Cerapachyini: Forel, 1893a: 162; Forel, 1900d: 329; Wheeler, W.M. 1910g: 137; Emery, 1911d: 11; Forel, 1917: 239.

Lioponera in Dorylinae, Cerapachyini: Emery, 1895j: 765.

Lioponera in Cerapachyinae, Cerapachyini: Wheeler, W.M. 1902d: 185; Wheeler, W.M. 1922a: 639; Donisthorpe, 1943f: 657.

Lioponera as subgenus of Cerapachys: Forel, 1892l: 244.

Lioponera as genus: Mayr, 1879: 666; Dalla Torre, 1893: 17; Forel, 1900d: 329; Bingham, 1903: 26; Emery, 1911d: 11; Wheeler, W.M. 1922a: 639; Donisthorpe, 1943f: 657.

Lioponera as junior synonym of Cerapachys: Brown, 1975: 19; all subsequent authors.

Parasyscia Emery, in André, 1882c: 235. Type-species: Parasyscia piochardi, by monotypy.

Taxonomic history

Parasyscia in Ponerinae: Dalla Torre, 1893: 17.

Parasyscia in Dorylinae, Cerapachyini: Emery, 1895j: 765.

Parasyscia in Ponerinae, Cerapachyini: Forel, 1893a: 162 [Cerapachysii]; Ashmead, 1905b: 382.

Parasyscia as genus: Dalla Torre, 1893: 17.

Parasyscia as subgenus of Cerapachys: Forel, 1892l: 243; Wheeler, W.M. 1902d: 185; Wheeler, W.M. 1910g: 137; Emery, 1911d: 9; Forel, 1917: 239; Wheeler, W.M. 1922a: 639; Donisthorpe, 1943f: 681; Creighton, 1950a: 57.

Parasyscia as junior synonym of Cerapachys: Kempf, 1972a: 76; Brown, 1975: 18; all subsequent authors.

Phyracaces Emery, 1902c: 23. Type-species: Cerapachys mayri, by original designation.

Taxonomic history

Phyracaces in Ponerinae, Cerapachyini: Ashmead, 1905b: 382; Wheeler, W.M. 1910g: 137; Emery, 1911d: 10; Arnold, 1915: 17; Forel, 1917: 239.

Phyracaces in Cerapachyinae, Cerapachyini: Wheeler, W.M. 1902d: 185; Wheeler, W.M. 1922a: 639; Donisthorpe, 1943f: 683.

Phyracaces as junior synonym of Cerapachys: Brown, 1973b: 183 [provisional]; Brown, 1975: 19; all subsequent authors.

Cysias Emery, 1902c: 24 [as subgenus of Cerapachys]. Type-species: Ooceraea papuana, by original designation.

Taxonomic history

Cysias in Ponerinae, Cerapachyini: Ashmead, 1905b: 382.

Cysias as genus: Ashmead, 1905b: 382.

Cysias as subgenus of Cerapachys: Emery, 1902c: 24; Wheeler, W.M. 1910g: 137; Donisthorpe, 1943f: 637.

Cysias as junior synonym of Syscia: Emery, 1911d: 10; Brown, 1975: 19; all subsequent authors.

Ceratopachys Schulz, W.A. 1906: 155, unjustified emendation of Cerapachys.

Taxonomic history

Ceratopachys as junior synonym of Cerapachys: Wheeler, W.M. 1922a: 755.

Chrysapace Crawley, 1924: 380. Type-species: Chrysapace jacobsoni (junior secondary homonym in Cerapachys, replaced by Cerapachys crawleyi), by monotypy.

Taxonomic history

Chrysapace in Ponerinae: Crawley, 1924: 380.

Chrysapace in Cerapachyinae, Cerapachyini: Donisthorpe, 1943f: 633.

Chrysapace as junior synonym of Cerapachys: Brown, 1975: 19; all subsequent authors.

Neophyracaces Clark, 1941: 76. Type-species: Phyracaces clarus, by original designation.

Taxonomic history

Neophyracaces in Cerapachyinae: Clark, 1941: 76.

Neophyracaces as junior synonym of Cerapachys: Brown, 1973b: 183 [provisional]; Brown, 1975: 19; all subsequent authors.

Yunodorylus Xu, 2000b: 297. Type-species: Yunodorylus sexspinus, by original designation.

Taxonomic history

Yunodorylus in Dorylinae: Xu, 2000b: 297.

Yunodorylus in Cerapachyinae, Cerapachyini: Bolton, 2003: 141, 268.

Yunodorylus as junior synonym of Cerapachys: Bolton, 2003: 141, 268; Borowiec, 2009: 57.

Genus Cerapachys references

Roger, 1863b: 21, 25 (catalogue); Mayr, 1863: 403, 439, 455 (catalogue); Mayr, 1865: 15, 24 (Syscia, Ooceraea diagnoses); André, 1882c: 236 (Europe & Algeria species); Dalla Torre, 1893: 17 (Lioponera, Cerapachys, Parasyscia, Ooceraea, Syscia catalogues); Forel, 1900d: 331 (India & Sri Lanka species key); Bingham, 1903: 27, 28 (India, Sri Lanka & Burma Lioponera, Cerapachys species keys); Emery, 1911d: 8 (diagnosis, subgenera key, catalogue); Emery, 1911d: 9 (C. (Parasyscia) diagnosis, catalogue); Emery, 1911d: 10 (C. (Ooceraea) & C. (Syscia) diagnoses, catalogues); Emery, 1911d: 10 (Phyracaces diagnosis, catalogue); Emery, 1911d: 11 (Lioponera diagnosis, catalogue); Arnold, 1915: 11 (diagnosis, South Africa species key); Wheeler, W.M. 1918a: 239 (Australia Phyracaces species key); Mann, 1921: 408 (Fiji species key); Wheeler, W.M. 1922a: 52, 53, 639 (Cerapachys & Phyracaces diagnoses, subgenera key); Wheeler, W.M. 1924d: 224 (Chrysapace, comments); Arnold, 1926: 193 (South Africa species, revised key); Donisthorpe, 1939a: 253 (Lioponera diagnosis); Chapman & Capco, 1951: 17, 21, 22 (Asia Cerapachys, Lioponera, Phyracaces checklists); Wilson, 1959b: 44, 52, 55, 56 (Melanesia, Fiji Is, New Guinea & New Caledonia species keys); Kempf, 1972a: 76 (Neotropical catalogue); Brown, 1975: 18, 26 (diagnosis, keys); Smith, D.R. 1979: 1333 (North America catalogue); Taylor & Brown, D.R. 1985: 23 (Australia catalogue); Taylor, 1987a: 16 (Australia, New Caledonia checklist); Terayama, Kubota, Sakai & Kawazoe, 1988: 35 (Taiwan species key); Morisita, Kubota, Onoyama, et al. 1989: 30 (Japan species key); Bolton, 1995a: 1048 (census); Bolton, 1995b: 142 (catalogue); Wu, J. & Wang, 1995: 47 (China species key); Collingwood & Agosti, 1996: 312 (Saudi Arabia species key); Terayama, 1996: 18 (Japan species key); Shattuck, 1999: 60 (Australia synopsis); Mackay & Mackay, 2002: 36 (U.S.A., New Mexico species key); Borowiec, 2009: 46 (C. sexspinus group key); Terayama, 2009: 122 (Taiwan species key); Heterick, 2009: 126 (south-western Australia species key).

Ecitoninae Ecitonini Labidus TRUE TRUE FALSE

7 species, 1 subspecies

Labidus Jurine, 1807: 282. Type-species: Labidus latreillii (junior synonym of Formica coeca), by monotypy.

Taxonomic history

Labidus in Dorylidae: Leach, 1815: 147 [Dorylida]; Haliday, 1836: 331; Shuckard, 1840a: 196; Smith, F. 1859b: 1; Cresson, 1887: 259.

Labidus in Dorylinae: Mayr, 1865: 16 [Dorylidae]; Forel, 1878: 365 (footnote) [Dorylidae].

Labidus in Dorylinae, Ecitonini: Emery, 1910b: 21 [Ecitini]; Forel, 1917: 240; Donisthorpe, 1943f: 654; Borgmeier, 1955: 79; Kempf, 1972a: 126; Dlussky & Fedoseeva, 1988: 79.

Labidus in Ecitoninae: Brown, 1973b: 166; Snelling, R.R. 1981: 392.

Labidus in Ecitoninae, Ecitonini: Bolton, 1990c: 1357; Hölldobler & Wilson, 1990: 12; Jaffe, 1993: 13; Bolton, 1994: 39; all subsequent authors.

Labidus as junior synonym of Eciton: Dalla Torre, 1893: 1.

Labidus as subgenus of Eciton: Emery, 1910b: 21; Forel, 1917: 240; Gallardo, 1920: 312; Wheeler, W.M. 1922a: 635; Borgmeier, 1923: 41; Donisthorpe, 1943f: 654; Creighton, 1950a: 61; Smith, M.R. 1951a: 779; Kusnezov, 1956: 9.

Labidus as genus: Jurine, 1807: 282; Shuckard, 1840a: 196; Borgmeier, 1953: 4; Borgmeier, 1955: 80; Smith, M.R. 1958c: 108; all subsequent authors.

Junior synonyms of LABIDUS

Nycteresia Roger, 1861a: 21. Type-species: Formica coeca, by monotypy.

Taxonomic history

Nycteresia in Ponerinae: Mayr, 1862: 714 [Poneridae].

Nycteresia as junior synonym of Eciton: Mayr, 1865: 76; Dalla Torre, 1893: 1.

Nycteresia as junior synonym of Labidus: Emery, 1910b: 21; Donisthorpe, 1943f: 675; Borgmeier, 1955: 80; all subsequent authors.

Pseudodichthadia André, 1885: 838. Type-species: Pseudodichthadia incerta (junior synonym of Formica coeca), by monotypy.

Taxonomic history

Pseudodichthadia in Dorylidae: André, 1885: 838.

Pseudodichthadia in Dorylinae: Dalla Torre, 1893: 7.

Pseudodichthadia in Dorylinae, Ecitonini: Forel, 1893a: 163; Emery, 1895j: 765 [Ecitonii].

Pseudodichthadia as junior synonym of Eciton: André, in Forel, 1899c: 160 (footnote).

Pseudodichthadia as junior synonym of Labidus: Emery, 1910b: 21; Borgmeier, 1955: 80; all subsequent authors.

Genus Labidus references

Roger, 1863b: 21, 41 (catalogue); Mayr, 1863: 424, 436 (catalogue); Mayr, 1865: 16 (diagnosis); Cresson, 1887: 259 (U.S.A. catalogue); Emery, 1910b: 21 (diagnosis, catalogue); Gallardo, 1920: 312 (Argentina species key); Creighton, 1950a: 62 (North America species key); Borgmeier, 1955: 80, 84 (diagnosis, all species revision, key); Kempf, 1972a: 126 (Neotropical catalogue); Watkins, 1976: 8 (all species key); Smith, D.R. 1979: 1327 (North America catalogue); Watkins, 1982: 209 (Mexico species key); Bolton, 1995a: 1050 (census); Bolton, 1995b: 219 (catalogue); Palacio, 1999: 153 (Colombia species key).

Ecitoninae Ecitonini Neivamyrmex TRUE TRUE FALSE

Extant: 124 species, 1 subspecies

Fossil: 1 species

Neivamyrmex Borgmeier, 1940: 606 [as subgenus of Eciton].

Taxonomic history

[Replacement name for Acamatus Emery, 1894c: 181; junior homonym of Acamatus Schoenherr, 1833: 20 (Coleoptera.]

Neivamyrmex in Dorylinae, Ecitonini: Donisthorpe, 1943f: 673; Borgmeier, 1955: 79; Kempf, 1972a: 152; Dlussky & Fedoseeva, 1988: 79.

Neivamyrmex in Ecitoninae: Brown, 1973b: 166; Snelling, R.R. 1981: 392.

Neivamyrmex in Ecitoninae, Ecitonini: Bolton, 1990c: 1357; Hölldobler & Wilson, 1990: 12; Jaffe, 1993: 13; Bolton, 1994: 39; all subsequent authors.

Neivamyrmex as subgenus of Eciton: Creighton, 1950a: 64; Kusnezov, 1956: 8.

Neivamyrmex as genus: Borgmeier, 1950c: 624; Borgmeier, 1955: 277; Smith, M.R. 1958c: 108; all subsequent authors except the above entry.

Junior synonym of NEIVAMYRMEX

Woitkowskia Enzmann, E.V. 1952: 443. Type-species: Woitkowskia connectens (junior synonym of Labidus walkerii), by original designation.

Taxonomic history

Woitkowskia as junior synonym of Neivamyrmex: Borgmeier, 1955: 277; all subsequent authors.

Genus Neivamyrmex references

Emery, 1894c: 183 (species key); Emery, 1910b: 23 (diagnosis, catalogue); Gallardo, 1920: 348 (Argentina species key); Smith, M.R. 1942c: 542 (U.S.A. species key); Buren, 1944a: 280 (U.S.A., Iowa species key); Creighton, 1950a: 66 (North America species key); Borgmeier, 1955: 277, 290 (diagnosis, all species revision, key); Watkins, 1971: 93 (U.S.A. species key); Kempf, 1972a: 152 (Neotropical catalogue); Watkins, 1972: 348 (U.S.A. species key); Watkins, 1976: 11 (all species key); Smith, D.R. 1979: 1329 (North America catalogue); Watkins, 1982: 210 (Mexico species key); Watkins, 1985: 481 (U.S.A. species key); Wheeler, G.C. & Wheeler, J. 1986g: 19 (U.S.A., Nevada species key); Brandão, 1991: 359 (Neotropical catalogue); Bolton, 1995a: 1051 (census); Bolton, 1995b: 287 (catalogue); Ward, 1999a: 90 (modification of Watkins, 1985 key); Palacio, 1999: 154 (Colombia species key); Mackay & Mackay, 2002: 40 (U.S.A., New Mexico species key); Coovert, 2005: 30 (U.S.A., Ohio species key); Snelling, G.C & Snelling, R.R., 2007: 462 (U.S.A. species, keys).

Homonym replaced by NEIVAMYRMEX

Acamatus Emery, 1894c: 181 [as subgenus of Eciton]. Type-species: Eciton (Acamatus) schmitti (junior synonym of Labidus nigrescens), by subsequent designation of Ashmead, 1906: 24; Wheeler, W.M. 1911f: 157.

Taxonomic history

[Junior homonym of Acamatus Schoenherr, 1833: 20 (Coleoptera).]

Acamatus in Ecitoninae, Ecitonini: Ashmead, 1905b: 381; Ashmead, 1906: 24.

Acamatus in Dorylinae, Ecitonini: Emery, 1910b: 23 [Ecitini].

Acamatus as genus: Ashmead, 1905b: 381; Ashmead, 1906: 24.

Acamatus as subgenus of Eciton: Emery, 1894c: 181; Emery, 1895j: 765; Emery, 1910b: 23; Wheeler, W.M. 1910g: 138; Forel, 1917: 240; Gallardo, 1920: 345; Wheeler, W.M. 1922a: 635; Borgmeier, 1923: 44; Donisthorpe, 1943f: 618.

Heteroponerinae Heteroponerini Heteroponera TRUE TRUE FALSE

19 species

Heteroponera Mayr, 1887: 532. Type-species: Heteroponera carinifrons, by monotypy.

Taxonomic history

Heteroponera in Ponerinae: Dalla Torre, 1893: 43.

Heteroponera in Pachycondylinae, Pachycondylini: Ashmead, 1905b: 382.

Heteroponera in Ponerinae, Ectatommini: Emery, 1911d: 35; Forel, 1917: 236; all subsequent authors to the following.

Heteroponera in Heteroponerinae, Heteroponerini: Bolton, 2003: 47, 177.

Heteroponera as subgenus of Acanthoponera: Forel, 1917: 236.

Heteroponera as junior synonym of Acanthoponera: Emery, 1895j: 767; Emery, 1911d: 35; Wheeler, W.M. 1922a: 643; Wheeler, 1923e: 179.

Heteroponera as genus: Brown, 1952h: 70; Kusnezov, 1956: 13; Brown, 1958g: 194; all subsequent authors.

Junior synonyms of HETEROPONERA

Paranomopone Wheeler, W.M. 1915c: 117. Type-species: Paranomopone relicta, by monotypy.

Taxonomic history

Paranomopone in Ponerinae, Ectatommini: Forel, 1917: 236; Emery, 1919c: 106; Wheeler, W.M. 1922a: 642; Wheeler, W.M. 1930f: 13; Donisthorpe, 1943f: 681.

Paranomopone as junior synonym of Heteroponera: Brown, in Borgmeier, 1957: 112; Brown, 1958g: 194; all subsequent authors.

Anacanthoponera Wheeler, W.M. 1923e: 176 (in text) [as subgenus of Acanthoponera]. Type-species: Ponera dolo, by original designation.

Taxonomic history

Anacanthoponera as junior synonym of Heteroponera: Brown, 1952h: 70; Brown, 1958g: 194; all subsequent authors.

Genus Heteroponera references

Dalla Torre, 1893: 43 (catalogue); Emery, 1911d: 35 (diagnosis, catalogue as part of Acanthoponera); Wheeler, W.M. 1923e: 190 (all species key as part of Acanthoponera); Brown, 1958g: 194, 196 (review of genus, Australia-New Zealand species key, New World species key); Brown, 1958h: 17 (New Zealand species); Kempf, 1962a: 31 (Neotropical species key); Kempf, 1972a: 117 (Neotropical catalogue); Taylor & Brown, D.R. 1985: 30 (Australia catalogue); Taylor, 1987a: 28 (Australia, New Zealand checklist); Fernández, 1993: 250 (Colombia species key); Bolton, 1995a: 1050 (census); Bolton, 1995b: 212 (catalogue); Shattuck, 1999: 186 (Australia synopsis); Arias-Penna & Fernández, 2008: 113 (Neotropical species key); Heterick, 2009: 142 (south-western Australia species key); Taylor, R.W. 2011: 118 (Australia groups and leae group species key).

Myrmicinae Basicerotini Rhopalothrix TRUE TRUE FALSE

9 species

Rhopalothrix Mayr, 1870a: 415. Type-species: Rhopalothrix ciliata, by subsequent designation of Wheeler, W.M. 1911f: 172.

Taxonomic history

Rhopalothrix in Myrmicidae, Cryptoceridae: Emery, 1877a: 81.

Rhopalothrix in Myrmicinae: Dalla Torre, 1893: 145.

Rhopalothrix in Cryptoceridae, Dacetini: Ashmead, 1905b: 384.

Rhopalothrix in Myrmicinae, Dacetini: Forel, 1892d: 344; Forel, 1893a: 164; Forel, 1895b: 133; Emery, 1895j: 770; Forel, 1899c: 41; Wheeler, W.M. 1910g: 141; Emery, 1914a: 42; Forel, 1917: 246; Wheeler, W.M. 1922a: 666; Emery, 1924d: 328; Donisthorpe, 1943g: 724; Brown, 1948e: 102; Chapman & Capco, 1951: 105; Baroni Urbani & De Andrade, 1994: 32; Baroni Urbani & De Andrade, 2007: 88 (as junior synonym of Basiceros).

Rhopalothrix in Myrmicinae, Basicerotini: Brown, 1949f: 92; Brown & Kempf, 1960: 230; Taylor, 1990b: 397; Bolton, 1994: 105; Bolton, 1998: 67; Fernández, 2003d: 314; Bolton, 2003: 185.

Junior synonyms of RHOPALOTHRIX

Heptastruma Weber, 1934a: 54. Type-species: Heptastruma wheeleri (junior secondary homonym in Rhopalothrix, replaced by Rhopalothrix weberi), by original designation.

Taxonomic history

Heptastruma in Myrmicinae, Dacetini: Weber, 1934a: 54; Donisthorpe, 1943f: 650; Brown, 1948e: 102.

Heptastruma in Myrmicinae, Basicerotini: Brown, 1949f: 94.

Heptastruma as junior synonym of Rhopalothrix: Brown & Kempf, 1960: 230.

Acanthidris Weber, 1941a: 188. Type-species: Acanthidris isthmicus, by original designation.

Taxonomic history

Acanthidris in Myrmicinae, Dacetini: Weber, 1941a: 183 [Dacetonini]; Donisthorpe, 1943f: 618; Brown, 1948e: 102.

Acanthidris in Myrmicinae, Basicerotini: Brown, 1949f: 94.

Acanthidris as junior synonym of Rhopalothrix: Brown & Kempf, 1960: 230.

Genus Rhopalothrix references

Dalla Torre, 1893: 145 (catalogue); Emery, 1894c: 216 (Neotropical species key); Mann, 1919: 360 (Papuasian species key); Emery, 1924d: 328 (diagnosis, catalogue); Chapman & Capco, 1951: 105 (Asia checklist); Brown, 1949f: 93 (checklist); Brown & Kempf, 1960: 230, 247 (diagnosis, all species revision, key); Taylor, 1968b: 334 (Indo-Australian species key); Kempf, 1972a: 226 (Neotropical catalogue); Taylor & Brown, D.R. 1985: 86 (Australia catalogue); Taylor, 1987a: 66 (Australia checklist); Taylor, 1990b: 401 (Old World species key); Bolton, 1995a: 1052 (census); Bolton, 1995b: 377 (catalogue); Shattuck, 1999: 162 (Australia synopsis).

Myrmicinae Dacetini Pyramica TRUE TRUE FALSE

Extant: 344 species

Fossil: 4 species

Pyramica Roger, 1862a: 251. Type-species: Pyramica gundlachi, by monotypy.

Taxonomic history

Pyramica in Myrmicinae, Dacetini: Emery, 1924d: 320; all subsequent authors.

Pyramica as junior synonym of Strumigenys: Roger, 1863b: 40; Emery & Forel, 1879a: 464; Dalla Torre, 1893: 145; Bingham, 1903: 147; Emery, 1924d: 320; Donisthorpe, 1943g: 723; Brown & Wilson, 1959b: 281; Brown, 1960b: 37; Kempf, 1972a: 242; Bolton, 1983: 358; Bolton, 1994: 105; Baroni Urbani & De Andrade, 2007: 101.

Pyramica as subgenus of Strumigenys: Brown, 1948e: 110.

Pyramica as genus: Mayr, 1863: 453; Bolton, 1999: 1667; Bolton, 2000: 90; Eguchi, Bui & Yamane, 2011: 26.

Junior synonyms of PYRAMICA

Epitritus Emery, 1869a: 136. Type-species: Epitritus argiolus, by monotypy.

Taxonomic history

Epitritus in Myrmicidae, Cryptoceridae: Emery, 1877a: 81.

Epitritus in Myrmicinae: Emery & Forel, 1879a: 465 [Myrmicidae]; Dalla Torre, 1893: 148.

Epitritus in Cryptoceridae, Dacetonini: Ashmead, 1905b: 384.

Epitritus in Myrmicinae, Dacetini: Forel, 1892d: 344; Forel, 1893a: 164; Emery, 1895j: 770; Wheeler, W.M. 1910g: 141; Emery, 1914a: 42; Forel, 1917: 246; Arnold, 1917: 384; Wheeler, W.M. 1922a: 666; Emery, 1924d: 326; Donisthorpe, 1943f: 643; Brown, 1948e: 122; Brown, 1949b: 44; all subsequent authors.

Epitritus as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 12; Baroni Urbani & De Andrade, 2007: 101.

Epitritus as junior synonym of Pyramica: Bolton, 1999: 1667; Bolton, 2000: 90.

Trichoscapa Emery, 1869b: 24 [as subgenus of Strumigenys]. Type-species: Strumigenys (Trichoscapa) membranifera, by monotypy.

Taxonomic history

Trichoscapa in Myrmicinae: Emery & Forel, 1879a: 464 [Myrmicidae]; Dalla Torre, 1893: 145.

Trichoscapa in Myrmicinae, Dacetini: Forel, 1917: 246; Wheeler, W.M. 1922a: 668; Emery, 1924d: 323; Donisthorpe, 1943g: 734; Brown, 1948e: 112; all subsequent authors.

Trichoscapa as subgenus of Strumigenys: Emery & Forel, 1879a: 464; Forel, 1917: 246; Creighton, 1950a: 304.

Trichoscapa as junior synonym of Strumigenys: Dalla Torre, 1893: 145; Baroni Urbani & De Andrade, 1994: 13; Baroni Urbani & De Andrade, 2007: 101.

Trichoscapa as junior synonym of Cephaloxys: Wheeler, W.M. 1922a: 668; Emery, 1924d: 323; Donisthorpe, 1943g: 734.

Trichoscapa as genus: Brown, 1948e: 112; Kempf, 1972a: 255; Bolton, 1983: 319.

Trichoscapa as junior synonym of Pyramica: Bolton, 1999: 1667; Bolton, 2000: 90.

Pentastruma Forel, 1912a: 50. Type-species: Pentastruma sauteri, by monotypy.

Taxonomic history

Pentastruma in Myrmicinae, Dacetini: Emery, 1914a: 42; Forel, 1917: 246; Wheeler, W.M. 1922a: 666; Emery, 1924d: 326; Donisthorpe, 1943f: 682; Brown, 1948e: 102.

Pentastruma as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 15; Baroni Urbani & De Andrade, 2007: 101.

Pentastruma as junior synonym of Pyramica: Bolton, 1999: 1667; Bolton, 2000: 91.

Glamyromyrmex Wheeler, W.M. 1915e: 487. Type-species: Glamyromyrmex beebei, by monotypy.

Taxonomic history

Glamyromyrmex in Myrmicinae, Dacetini: Wheeler, W.M. 1915e: 491; Forel, 1917: 246; Wheeler, W.M. 1922a: 667; Emery, 1924d: 326; Donisthorpe, 1943f: 647; Brown, 1948e: 116; all subsequent authors.

Glamyromyrmex as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 14; Baroni Urbani & De Andrade, 2007: 101.

Glamyromyrmex as junior synonym of Pyramica: Bolton, 1999: 1667; Bolton, 2000: 91.

Codiomyrmex Wheeler, W.M. 1916d: 326. Type-species: Codiomyrmex thaxteri, by monotypy.

Taxonomic history

Codiomyrmex in Myrmicinae, Dacetini: Wheeler, W.M. 1922a: 667; Emery, 1924d: 325; Donisthorpe, 1943f: 633; Brown, 1948e: 114; all subsequent authors.

Codiomyrmex as subgenus of Strumigenys: Emery, 1924d: 325; Donisthorpe, 1943f: 633.

Codiomyrmex as genus: Wheeler, W.M. 1916d: 326; Brown, 1948e: 114.

Codiomyrmex as junior synonym of Glamyromyrmex: Brown, 1973b: 179 [provisional].

Codiomyrmex as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 14; Baroni Urbani & De Andrade, 2007: 101.

Codiomyrmex as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Tingimyrmex Mann, 1926: 104 [as subgenus of Strumigenys]. Type-species: Strumigenys (Tingimyrmex) mirabilis, by monotypy.

Taxonomic history

Tingimyrmex in Myrmicinae, Dacetini: Donisthorpe, 1943g: 732; Brown, 1948e: 111; all subsequent authors.

Tingimyrmex as subgenus of Strumigenys: Mann, 1926: 104; Donisthorpe, 1943g: 732.

Tingimyrmex as genus: Brown, 1948e: 111; Kempf, 1972a: 250.

Tingimyrmex as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 15; Baroni Urbani & De Andrade, 2007: 101.

Tingimyrmex as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Codioxenus Santschi, 1931c: 278 [as subgenus of Epitritus]. Type-species: Epitritus (Codioxenus) simulans, by monotypy.

Taxonomic history

Codioxenus in Myrmicinae, Dacetini: Donisthorpe, 1943f: 633; Brown, 1948e: 123; all subsequent authors.

Codioxenus as genus: Brown, 1948e: 123.

Codioxenus as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 14; Baroni Urbani & De Andrade, 2007: 101.

Codioxenus as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Smithistruma Brown, 1948e: 104.

Taxonomic history

[Replacement name for Cephaloxys Smith, F. 1865: 76; junior homonym of Cephaloxys Signoret, 1847: 294 (Hemiptera). Type-species not Strumigenys pulchella, unjustified subsequent designation by Brown, 1948e: 104, repeated in Kempf, 1972a: 230, Bolton, 1983: 274 and Bolton, 1995b: 46.]

Smithistruma in Myrmicinae, Dacetini: Brown, 1948e: 104.

Smithistruma as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 14; Baroni Urbani & De Andrade, 2007: 102.

Smithistruma as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Homonym replaced by Smithistruma

Cephaloxys Smith, F. 1865: 76. Type-species: Cephaloxys capitata, by monotypy.

Taxonomic history

[Cephaloxys Smith, F. junior homonym of Cephaloxys Signoret, 1847: 294 (Hemiptera).]

Cephaloxys in Cryptoceridae: Smith, F. 1871a: 335.

Cephaloxys in Myrmicinae, Dacetini: Wheeler, W.M. 1922a: 668.

Cephaloxys as junior synonym of Strumigenys: Dalla Torre, 1893: 15.

Cephaloxys as subgenus of Strumigenys: Wheeler, W.M. 1922a: 668; Emery, 1924d: 323; Bernard, 1953b: 254 (anachronism).

Weberistruma Brown, 1948e: 106 [as subgenus of Smithistruma]. Type-species: Strumigenys (Cephaloxys) leptothrix, by original designation.

Taxonomic history

Weberistruma in Myrmicinae, Dacetini: Brown, 1948e: 106.

Weberistruma as genus: Brown, 1949d: 7; Brown, 1953g: 24.

Weberistruma as junior synonym of Smithistruma: Brown, 1973a: 35.

Weberistruma as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Weberistruma as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 2007: 102.

Wessonistruma Brown, 1948e: 106 [as subgenus of Smithistruma]. Type-species: Strumigenys pergandei, by original designation.

Taxonomic history

Wessonistruma in Myrmicinae, Dacetini: Brown, 1948e: 106.

Wessonistruma junior synonym of Smithistruma: Brown, 1973a: 35.

Wessonistruma as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Wessonistruma as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 2007: 102.

Serrastruma Brown, 1948e: 107 [as subgenus of Smithistruma]. Type-species: Strumigenys simoni, by original designation.

Taxonomic history

Serrastruma in Myrmicinae, Dacetini: Brown, 1948e: 107, and all subsequent authors.

Serrastruma as genus: Brown, 1949d: 6; Brown, 1952e: 70; Bolton, 1983: 335; Bolton, 1994: 105.

Serrastruma as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 14; Baroni Urbani & De Andrade, 2007: 102.

Serrastruma as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Neostruma Brown, 1948e: 111. Type-species: Strumigenys crassicornis, by original designation.

Taxonomic history

Neostruma in Myrmicinae, Dacetini: Brown, 1948e: 111; all subsequent authors.

Neostruma as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 12; Baroni Urbani & De Andrade, 2007: 102.

Neostruma as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Dorisidris Brown, 1948e: 116. Type-species: Strumigenys (Codiomyrmex) nitens, by original designation.

Taxonomic history

Dorisidris in Myrmicinae, Dacetini: Brown, 1948e: 116; all subsequent authors.

Dorisidris as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 14; Baroni Urbani & De Andrade, 2007: 102.

Dorisidris as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Miccostruma Brown, 1948e: 123. Type-species: Epitritus mandibularis, by original designation.

Taxonomic history

Miccostruma in Myrmicinae, Dacetini: Brown, 1948e: 123.

Miccostruma as junior synonym of Smithistruma: Bolton, 1983: 274.

Miccostruma as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Miccostruma as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 2007: 102.

Kyidris Brown, 1949d: 3. Type-species: Kyidris mutica, by original designation.

Taxonomic history

Kyidris in Myrmicinae, Dacetini: Brown, 1949d: 3.

Kyidris as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 15; Baroni Urbani & De Andrade, 2007: 103.

Kyidris as junior synonymof Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Chelystruma Brown, 1950a: 33 [as subgenus of Glamyromyrmex]. Type-species: Glamyromyrmex (Chelystruma) lilloana, by monotypy.

Taxonomic history

Chelystruma in Myrmicinae, Dacetini: Brown, 1950a: 33; all subsequent authors.

Chelystruma as subgenus of Glamyromyrmex: Brown, 1953g: 16.

Chelystruma as genus: Kempf, 1959d: 338; Kempf, 1972a: 77.

Chelystruma as junior synonym of Glamyromyrmex: Brown, 1973b: 179 [provisional].

Chelystruma as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 14; Baroni Urbani & De Andrade, 2007: 103.

Chelystruma as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Polyhomoa Azuma, 1950: 36. Type-species: Polyhomoa itoi (junior synonym of Kyidris mutica), by monotypy.

Taxonomic history

Polyhomoa as junior synonym of Kyidris: Creighton, 1950b: 93; Brown & Yasumatsu, 1951: 93.

Polyhomoa as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Polyhomoa as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 2007: 103.

Borgmeierita Brown, 1953g: 23. Type-species: Codiomyrmex excisus, by original designation.

Taxonomic history

Borgmeierita in Myrmicinae, Dacetini: Brown, 1953g: 23.

Borgmeierita as junior synonym of Glamyromyrmex: Brown, 1973a: 35.

Borgmeierita as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Borgmeierita as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 2007: 103.

Platystruma Brown, 1953g: 112 [as subgenus of Smithistruma]. Type-species: Strumigenys (Cephaloxys) depressiceps, by original designation.

Taxonomic history

Platystruma in Myrmicinae, Dacetini: Brown, 1953g: 112; all subsequent authors.

Platystruma as junior synonym of Smithistruma: Brown, 1973a: 35.

Platystruma as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Platystruma as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 2007: 103.

Gymnomyrmex Borgmeier, 1954b: 279. Type-species: Gymnomyrmex splendens, by original designation.

Taxonomic history

Gymnomyrmex in Myrmicinae, Dacetini: Borgmeier, 1954b: 279; all subsequent authors.

Gymnomyrmex as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 14; Baroni Urbani & De Andrade, 2007: 103.

Gymnomyrmex as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Dysedrognathus Taylor, 1968d: 132. Type-species: Dysedrognathus extemenus, by original designation.

Taxonomic history

Dysedrognathus in Myrmicinae, Dacetini: Taylor, 1968d: 132.

Dysedrognathus as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 13; Baroni Urbani & De Andrade, 2007: 103.

Dysedrognathus as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Asketogenys Brown, 1972: 23. Type-species: Asketogenys acubecca, by original designation.

Taxonomic history

Asketogenys in Myrmicinae, Dacetini: Brown, 1972: 23.

Asketogenys as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 15; Baroni Urbani & De Andrade, 2007: 103.

Asketogenys as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Cladarogenys Brown, 1976b: 33. Type-species: Cladarogenys lasia, by original designation.

Taxonomic history

Cladarogenys in Myrmicinae, Dacetini: Brown, 1976b: 33.

Cladarogenys as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 14; Baroni Urbani & De Andrade, 2007: 103.

Cladarogenys as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Genus Pyramica references

Mayr, 1863: 453 (catalogue); André, 1883b: 398, 402 (Europe & Algeria Strumigenys (Trichoscapa), Epitritus species); Dalla Torre, 1893: 145, 148 (Strumigenys, Epitritus catalogues); Emery, 1895c: 325 (North America species key); Wheeler, W.M. 1908a: 146 (New World species key); Santschi, 1913b: 257 (Afrotropical species key); Emery, 1916b: 205 (Italy species key); Arnold, 1917: 373 (South Africa species key); Wheeler, W.M. 1922a: 918, 920 (Afrotropical Strumigenys (Cephaloxys), Epitritus catalogues); Emery, 1924d: 323, 325, 326 (Strumigenys (Cephaloxys), Strumigenys (Codiomyrmex), Glamyromyrmex, Pentastruma, Epitritus diagnoses, catalogues); Smith, M.R. 1931c: 691 (North America Strumigenys (Cephaloxys) species key); Buren, 1944a: 290 (U.S.A., Iowa species key); Brown, 1948e: 112 - 124 (minor genera reviews); Brown, 1949b: 43 (Epitritus diagnosis, review); Brown, 1949d: 8 (Weberistruma species, review); Brown, 1949d: 21 (Japan, China & Taiwan species key); Creighton, 1950a: 301 (North America Strumigenys (Trichoscapa) species key); Chapman & Capco, 1951: 104, 105, 106 (Asia Epitritus, Kyidris, Pentastruma, Smithistruma, Strumigenys (Cephaloxys) checklists); Brown, 1950a: 27 (Glamyromyrmex review); Brown, 1952e: 86 (Serrastruma diagnosis, species revision, key); Brown, 1953g: 20, 24 (Codiomyrmex, Weberistruma reviews); Brown, 1953g: 31, 46, 92, 114, 125, 130 (Smithistruma diagnosis, Nearctic, Neotropical, Malesian, Afrotropical, Palaearctic species revisions, keys); Wilson & Brown, 1956b: 439 (Kyidris species review); Brown, 1959b: 1 (Neostruma species review); Kempf, 1959d: 339 (Gymnomyrmex species key); Brown, 1960b: 37 (Strumigenys gundlachi group revision, key); Kempf, 1960f: 447 (Neotropical Glamyromyrmex species key); Kempf, 1960f: 451 (Gymnomyrmex species key); Brown, 1962b: 79 (Epitritus species key); Brown, 1964a: 183 (supplement to 1953e Smithistruma revision); Bernard, 1967: 243, 245 (Smithistruma, Epitritus diagnoses); Kempf, 1972a: 37, 77, 98, 110, 116, 163, 230, 250, 255 (Neotropical Borgmeierita, Chelystruma, Codiomyrmex, Codioxenus, Dorisidris, Glamyromyrmex, Gymnomyrmex, Neostruma, Smithistruma, Tingimyrmex, Trichoscapa catalogues); Bolton, 1972: 208 (Epitritus species key); Alayo, 1974: 19 (Cuba species key); Smith, D.R. 1979: 1405, 1408 (North America Smithistruma, Trichoscapa catalogues); Taylor, 1987a: 28 (Australia checklist); Brown & Boisvert, 1979: 201 (Pentastruma species review); Bolton, 1983: 274, 319, 320, 335, 353, 354 (Afrotropical Smithistruma, Trichoscapa, Glamyromyrmex, Serrastruma, Cladarogenys, Epitritus species revisions, keys); Taylor & Brown, D.R. 1985: 64 (Australia Glamyromyrmex catalogue); Perrault, 1986a: 3 (Gymnomyrmex species key); Ward, 1988: 122 (Western Nearctic Smithistruma species key); Terayama & Kubota, 1989: 787 (Taiwan Smithistruma species key); Hölldobler & Wilson, 1990: 15 (synoptic classification); Brandão, 1991: 344, 346, 360, 378 (Neotropical catalogue); Arakelian & Dlussky, 1991: 150 (former U.S.S.R. species key); Morisita, Kubota, et al. 1992: 68 (Japan Smithistruma species key); Morisita, Kubota, et al. 1992: 76 (Japan Epitritus species key); Bolton, 1994: 105 (synoptic classification); Bolton, 1995a: 1048 - 1053 (census); Bolton, 1995b: 75, 145, 146, 177, 183, 188, 207, 211, 219, 292, 316, 382, 383, 420, 421 (catalogue); Terayama, Lin & Wu, 1996: 328 (Taiwan Smithistruma species key); Ogata & Onoyama, 1998: 277 (Japan Smithistruma species key); Shattuck, 1999: 136, 170 (Australia Glamyromyrmex, Trichoscapa synopses); Bolton, 1999: 1657, 1667 (morphology, diagnosis, review of genus, phylogeny); Xu, 2000e: 297 (China Epitritus species key); Bolton, 2000: 95, 137, 272, 285, 342, 372, 472 (Nearctic, Neotropical, Afrotropical, West Palaearctic, Malagasy, Malesian-Oriental-East Palaearctic, Austral species revisions, keys); Xu & Zhou, 2004: 441 (China species, key); Longino, 2006b: 139 (Costa Rica species, additions to key); Terayama, 2009: 138 (Taiwan species key).

Myrmicinae Cephalotini Cephalotes TRUE TRUE FALSE

Extant: 115 species

Fossil: 16 species

Cephalotes Latreille, 1802a: 357. Type-species: Formica atrata, by monotypy.

Taxonomic history

[Type-species not Formica cephalotes, unjustified subsequent designation by Wheeler, W.M. 1911f: 160; corrected by Wheeler, W.M. 1913a: 78.]

Cephalotes in Myrmicinae, Cryptocerini: Emery, 1914a: 42; Forel, 1917: 246; Wheeler, W.M. 1922a: 665; Emery, 1924d: 303; all subsequent authors to 1949, and Dlussky & Fedoseeva, 1988: 79 (anachronism).

Cephalotes in Myrmicinae, Cephalotini: Smith, M.R. 1949c: 19; Kempf, 1951: 105; all subsequent authors except the above.

Cephalotes as junior synonym of Cryptocerus: Fabricius, 1804: 419.

Junior synonyms of CEPHALOTES

Cryptocerus Latreille, 1803: 311. Type-species: Formica atrata, by subsequent designation of Latreille, 1810: 437.

Taxonomic history

[Type-species not Cryptocerus umbraculatus, unjustified subsequent designation by Emery, 1914c: 38, repeated in Emery, 1924d: 305.]

Cryptocerus in Myrmicites: Lepeletier de Saint-Fargeau, 1835: 170.

Cryptocerus in Myrmicidae, Cryptoceridae: Smith, F. 1853: 214; Emery, 1877a: 81.

Cryptocerus in Poneridae, Cryptoceridae: Smith, F. 1858b: 187.

Cryptocerus in Cryptoceridae: Smith, F. 1876d: 605.

Cryptocerus in Cryptoceridae, Cryptocerinae: Ashmead, 1905b: 384.

Cryptocerus in Myrmicinae: Mayr, 1865: 25 [Myrmicidae]; Mayr, 1866b: 907 [Myrmicidae]; Dalla Torre, 1893: 140.

Cryptocerus in Myrmicinae, Cryptocerini: Forel, 1892d: 345; Forel, 1893a: 164; Forel, 1895b: 133; Emery, 1895j: 771; Forel, 1899c: 48; Wheeler, W.M. 1910g: 141; Emery, 1914a: 42; Forel, 1917: 246; Wheeler, W.M. 1922a: 665; Emery, 1924d: 305; Wheeler, G.C. & Wheeler, J. 1976b: 59.

Cryptocerus as junior synonym of Cephalotes: Wheeler, W.M. 1913a: 78; Donisthorpe, 1943f: 635; Smith, M.R. 1949c: 19; Kempf, 1951: 105; all subsequent authors except the above. [Cephalotes and Cryptocerus share the same type-species, synonymy is therefore absolute.]

[Cryptocephalus Lowne, 1865b: 336, incorrect subsequent spelling of Cryptocerus in the combination Cryptocephalus pubescens; current combination Meranoplus pubescens: Bolton, 1981a: 47.]

Zacryptocerus Wheeler, W.M. 1911f: 175 (see also footnote). Type-species: Cryptocerus clypeatus, by original designation.

Taxonomic history

[Zacryptocerus Ashmead, 1905b: 384, nomen nudum. (Based on a non-existent type-species: Cryptocerus multistrigus, nomen nudum, attributed to Smith, F.).]

Zacryptocerus in Myrmicinae, Cryptocerini: Forel, 1917: 246; Wheeler, W.M. 1922a: 665; Emery, 1924d: 304; all subsequent authors except the following; Dlussky & Fedoseeva, 1988: 79 (anachronism).

Zacryptocerus in Myrmicinae, Cephalotini: Kempf, 1951: 133; all subsequent authors.

Zacryptocerus as junior synonym of Cephalotes: De Andrade & Baroni Urbani, 1999: 59.

Paracryptocerus Emery, 1915i: 192 [as subgenus of Cryptocerus]. Type-species: Cryptocerus spinosus, by original designation.

Taxonomic history

Paracryptocerus in Myrmicinae, Cryptocerini: Forel, 1917: 246; Wheeler, W.M. 1922a: 665; Emery, 1924d: 306; Donisthorpe, 1943f: 680.

Paracryptocerus in Myrmicinae, Cephalotini: Smith, M.R. 1949c: 20; Kempf, 1951: 153; all subsequent authors.

Paracryptocerus as subgenus of Cryptocerus: Emery, 1915i: 192; Forel, 1917: 246; Wheeler, W.M. 1922a: 665; Emery, 1924d: 306.

Paracryptocerus as genus: Smith, M.R. 1949c: 20; Kempf, 1951: 153; Kempf, 1972a: 175.

Paracryptocerus as junior synonym of Zacryptocerus: Kempf, 1973c: 460; Smith, D.R. 1979: 1402.

Hypocryptocerus Wheeler, W.M. 1920: 53 [as subgenus of Cryptocerus]. Type-species: Formica haemorrhoidalis, by original designation.

Taxonomic history

Hypocryptocerus in Myrmicinae, Cryptocerini: Wheeler, W.M. 1922a: 665; Wheeler, W.M. 1936b: 200; Donisthorpe, 1943f: 652.

Hypocryptocerus in Myrmicinae, Cephalotini: Kempf, 1951: 143.

Hypocryptocerus as subgenus of Cryptocerus: Wheeler, W.M. 1920: 53; Wheeler, W.M. 1922a: 665.

Hypocryptocerus as genus: Wheeler, W.M. 1936b: 200; Kempf, 1951: 143; Kempf, 1972a: 120.

Hypocryptocerus as junior synonym of Zacryptocerus: Kempf, 1973c: 460; Smith, D.R. 1979: 1402.

Cyathomyrmex Creighton, 1933: 100 [as subgenus of Cryptocerus].

Taxonomic history

[Replacement name for Cyathocephalus Emery; junior homonym of Cyathocephalus Kessler, 1868: 135 (Cestoda).]

Cyathomyrmex in Myrmicinae, Cephalotini: Smith, M.R. 1949c: 21.

Cyathomyrmex as subgenus of Cryptocerus: Creighton, 1933: 100; Creighton, 1950a: 298.

Cyathomyrmex as subgenus of Paracryptocerus: Smith, M.R. 1949c: 21; Kempf, 1951: 156.

Cyathomyrmex as junior synonym of Paracryptocerus: Kempf, 1972a: 175.

Homonym replaced by Cyathomyrmex

Cyathocephalus Emery, 1915i: 192 [as subgenus of Cryptocerus]. Type-species: Cryptocerus pallens, by original designation.

Taxonomic history

[Cyathocephalus Emery junior homonym of Cyathocephalus Kessler, 1868: 135 (Cestoda).]

Cyathocephalus as subgenus of Cryptocerus: Emery, 1915i: 192; Forel, 1917: 246; Wheeler, W.M. 1922a: 665; Emery, 1924d: 311.

Harnedia Smith, M.R. 1949c: 20 [as subgenus of Paracryptocerus]. Type-species: Cryptocerus umbraculatus, by original designation.

Taxonomic history

Harnedia in Myrmicinae, Cephalotini: Smith, M.R. 1949c: 20; Kempf, 1951: 233.

Harnedia as subgenus of Paracryptocerus: Smith, M.R. 1949c: 20; Kempf, 1951: 233.

Harnedia as junior synonym of Paracryptocerus: Kempf, 1972a: 175.

Eucryptocerus Kempf, 1951: 127. Type-species: Cryptocerus oculatus, by original designation.

Taxonomic history

Eucryptocerus in Myrmicinae, Cephalotini: Kempf, 1951: 127; all subsequent authors except the following.

Eucryptocerus in Myrmicinae, Cryptocerini: Dlussky & Fedoseeva, 1988: 79 (anachronism).

Eucryptocerus as genus: Kempf, 1972a: 107; Kempf, 1973c: 460; Bolton, 1994: 105.

Eucryptocerus as junior synonym of Cephalotes: Brown, 1973b: 180 [provisional]; De Andrade & Baroni Urbani, 1999: 59.

†Exocryptocerus Vierbergen & Scheven, 1995: 159. Type-species: †Exocryptocerus serratus, by original designation.

Taxonomic history

†Exocryptocerus as junior synonym of Cephalotes: De Andrade & Baroni Urbani, 1999: 59.

Genus Cephalotes references

Smith, F. 1853: 215 (diagnosis); Smith, F. 1858b: 187 (diagnosis); Roger, 1863b: 38 (catalogue); Mayr, 1863: 405 (catalogue); Mayr, 1865: 25 (diagnosis); Dalla Torre, 1893: 140 (catalogue); Santschi, 1920f: 149 (species key); Wheeler, W.M. 1922a: 665 (Cryptocerus subgenera, key); Emery, 1924d: 303 (diagnosis, catalogue); Emery, 1924d: 304 (Zacryptocerus diagnosis, catalogue); Emery, 1924d: 305 (Cryptocerus diagnosis, subgenera key, catalogue); Emery, 1924d: 306 (C. (Paracryptocerus) diagnosis, catalogue); Emery, 1924d: 308 (C. (Cryptocerus) diagnosis, catalogue); Emery, 1924d: 311 (C. (Cyathocephalus) diagnosis, catalogue); Smith, M.R. 1947a: 29 (U.S.A. species key); Creighton, 1950a: 296 (North America species key); Kempf, 1951: 107 (diagnosis, revision, key); Kempf, 1951: 127 (Eucryptocerus species revision, key); Kempf, 1951: 134 (Zacryptocerus species revision, key); Kempf, 1951: 143 (Hypocryptocerus species revision, key); Kempf, 1951: 153 (Paracryptocerus species revision, key); Kempf, 1952: 5 (Paracryptocerus (Harnedia) pinellii complex revision, key); Kempf, 1958a: 10 (Argentina Paracryptocerus species key); Kempf, 1958a: 64 (Paracryptocerus subgenera, key); Kempf, 1958a: 66 (Paracryptocerus (Harnedia) species revision, key); Kempf, 1958a: 145 (Paracryptocerus (Cyathomyrmex) species revision, key); Kempf, 1960f: 443 (addition to 1958 key); Snelling, R.R. 1968a: 9 (C. wheeleri complex, key); Kempf, 1972a: 75, 107, 120, 175, 259 (Neotropical Cephalotes, Eucryptocerus, Hypocryptocerus, Paracryptocerus, Zacryptocerus catalogues); Kempf, 1974a: 73 (Paracryptocerus pinelii group, key); Smith, D.R. 1979: 1402 (North America catalogue); Hölldobler & Wilson, 1990: 13 (synoptic classification); Brandão, 1991: 337, 343, 383 (catalogue); Bolton, 1994: 105 (synoptic classification); Bolton, 1995a: 1048 (census); Bolton, 1995b: 140, 189, 424 (catalogue); De Andrade & Baroni Urbani, 1999: 736 (diagnosis, all species revision, key).

Myrmicinae Attini Atta TRUE TRUE FALSE

15 species

Atta Fabricius, 1804: 421. Type-species: Formica cephalotes, by subsequent designation of Wheeler, W.M. 1911f: 159.

Taxonomic history

Atta in Myrmicites: Lepeletier de Saint-Fargeau, 1835: 172.

Atta in Poneridae, Attidae: Smith, F. 1858b: 161.

Atta in Myrmicinae: Mayr, 1855: 459 [Myrmicidae]; Smith, F. 1857a: 77 [Myrmicidae]; Mayr, 1861: 65 [Myrmicidae]; Mayr, 1865: 18 [Myrmicidae]; Smith, F. 1871a: 333 [Myrmicidae]; Cresson, 1887: 259 [Myrmicidae]; Dalla Torre, 1893: 150.

Atta in Cryptoceridae, Attinae: Ashmead, 1905b: 384.

Atta in Myrmicinae, Attini: Emery, 1877a: 81 [Myrmicidae, Attidae]; Forel, 1893a: 163; Emery, 1895j: 770; Forel, 1899c: 30; Wheeler, W.M. 1910g: 141; Emery, 1914a: 42; Forel, 1917: 247; Wheeler, W.M. 1922a: 669; Emery, 1924d: 352; all subsequent authors.

[Myrmegis (nomen nudum) referable to Atta: Brown, 1973b: 182 (see under Myrmegis).]

Junior synonyms of ATTA

Oecodoma Latreille, 1818: 222. Type-species: Formica cephalotes, by subsequent designation of Shuckard, in Swainson & Shuckard, 1840: 174.

Taxonomic history

Oecodoma in Myrmicites: Lepeletier de Saint-Fargeau, 1835: 176.

Oecodoma in Poneridae, Attidae: Smith, F. 1858b: 180.

Oecodoma as junior synonym of Atta: Roger, 1863b: 35. [Atta and Oecodoma share the same type-species, synonymy is therefore absolute.]

Archeatta Gonçalves, 1942: 342 [as subgenus of Atta]. Type-species: Oecodoma mexicana, by original designation.

Taxonomic history

Archeatta as subgenus of Atta: Gonçalves, 1942: 342; Borgmeier, 1959b: 350; Kempf, 1972a: 27.

Archeatta as junior synonym of Atta: Smith, M.R. 1951a: 832; Weber, 1958a: 8; Brown, 1973b: 178 [provisional]; Bolton, 1994: 105.

Neoatta Gonçalves, 1942: 346 [as subgenus of Atta]. Type-species: Formica sexdens, by original designation.

Taxonomic history

Neoatta as subgenus of Atta: Gonçalves, 1942: 346; Borgmeier, 1959b: 358; Kempf, 1972a: 27.

Neoatta as junior synonym of Atta: Weber, 1958a: 8; Brown, 1973b: 183 [provisional]; Bolton, 1994: 105.

Palaeatta Borgmeier, 1950d: 244 [as subgenus of Atta]. Type-species: Atta bisphaerica, by original designation.

Taxonomic history

Palaeatta as subgenus of Atta: Borgmeier, 1950d: 244; Borgmeier, 1959b: 377; Kempf, 1972a: 28.

Palaeatta as junior synonym of Atta: Weber, 1958a: 8; Brown, 1973b: 183 [provisional]; Bolton, 1994: 105.

Epiatta Borgmeier, 1950d: 246 [as subgenus of Atta]. Type-species: Oecodoma laevigata, by original designation.

Taxonomic history

Epiatta as junior synonym of Atta: Weber, 1958a: 8.

Epiatta as junior synonym of Neoatta: Borgmeier, 1959b: 358.

Genus Atta references

Mayr, 1855: 459 (diagnosis); Smith, F. 1858b: 161, 180 (Atta, Oecodoma diagnoses); Mayr, 1861: 66 (Europe species key); Roger, 1863b: 35 (catalogue); Mayr, 1863: 395, 437 (catalogue); Mayr, 1865: 18 (diagnosis); Cresson, 1887: 259 (U.S.A. catalogue); Dalla Torre, 1893: 150 (catalogue); Gallardo, 1916d: 340 (Argentina species key); Emery, 1924d: 352 (diagnosis, catalogue); Gonçalves, 1942: 334 (all species key); Gonçalves, 1947a: 186 (south & central Brazil species key); Chapman & Capco, 1951: 82 (Asia checklist, anachronisms); Borgmeier, 1959b: 335 (subgenera key); Borgmeier, 1959b: 337 (diagnosis, all species revision, key); Kempf, 1972a: 26 (Neotropical catalogue); Smith, D.R. 1979: 1413 (North America catalogue); Mackay & Mackay, 1988: 25 (Colombia species key); Cherrett & Cherrett, 1989: 1 (bibliography); Brandão, 1991: 328 (catalogue); Bolton, 1994: 105 (synoptic classification); Bolton, 1995a: 1048 (census); Bolton, 1995b: 75 (catalogue).

Myrmicinae Attini Cyphomyrmex TRUE TRUE FALSE

Extant: 39 species

Fossil: 2 species

Cyphomyrmex Mayr, 1862: 651 (diagnosis in key), 690. Type-species: Cyphomyrmex minutus, by monotypy.

Taxonomic history

[Type-species not Cryptocerus rimosus, unjustified subsequent designation by Wheeler, W.M. 1911f: 161. Type-species not Meranoplus difformis, unjustified subsequent designation by Emery, 1924d: 340.]

Cyphomyrmex in Formicinae: Mayr, 1862: 651 (in key) [Formicidae].

Cyphomyrmex in Myrmicidae, Cryptoceridae: Emery, 1877a: 81.

Cyphomyrmex in Myrmicinae: Mayr, 1865: 25 [Myrmicidae]; Dalla Torre, 1893: 149.

Cyphomyrmex in Cryptoceridae, Dacetonini: Ashmead, 1905b: 384.

Cyphomyrmex in Myrmicinae, Attini: Forel, 1893a: 164; Emery, 1895j: 770; Forel, 1899c: 40; Wheeler, W.M. 1907c: 670; Wheeler, W.M. 1910g: 141; Emery, 1913b: 251; Emery, 1914a: 42; Forel, 1917: 247; Wheeler, W.M. 1922a: 669; Emery, 1924d; 339; all subsequent authors.

Cyphomyrmex as junior synonym of Cataulacus: Roger, 1863b: 39.

Cyphomyrmex as subgenus of Atta: Forel, 1912e: 188.

Cyphomyrmex as genus: Mayr, 1862: 690; Mayr, 1863: 406; Emery, 1913b: 251; Bruch, 1914: 217; Emery, 1914a: 42; Wheeler, W.M. 1922a: 669; Emery, 1924d; 339; all subsequent authors.

Junior synonym of CYPHOMYRMEX

Cyphomannia Weber, 1938b: 183 [as subgenus of Cyphomyrmex]. Type-species: Cyphomyrmex (Cyphomannia) laevigatus, by original designation.

Taxonomic history

Cyphomannia as subgenus of Cyphomyrmex: Weber, 1938b: 183; Weber, 1966: 166.

Cyphomannia as junior synonym of Cyphomyrmex: Kempf, 1962b: 29; Kempf, 1968a: 39; Kempf, 1972a: 92; Bolton, 1994: 105.

Genus Cyphomyrmex references

Mayr, 1863: 406 (catalogue); Mayr, 1865: 25 (diagnosis); Mayr, 1887: 556 (all species key); Dalla Torre, 1893: 149 (catalogue); Emery, 1924d; 340 (diagnosis, subgenera key, catalogue); Kusnezov, 1949d: 436 (Argentina species key); Weber, 1940a: 408 (all species key); Creighton, 1950a: 316 (North America species key); Kusnezov, 1957b: 9 (Neotropical species, partial key); Kempf, 1962b: 30 (species groups); Kempf, 1964d: 5 (C. strigatus group revision, key); Kempf, 1966: 161 (C. rimosus group revision, key); Kempf, 1972a: 92 (Neotropical catalogue); Smith, D.R. 1979: 1410 (North America catalogue); Brandão, 1991: 339 (catalogue); Snelling, R.R. & Longino, 1992: 480 (C. rimosus group revision, key); Bolton, 1994: 105 (synoptic classification); Bolton, 1995a: 1049 (census); Bolton, 1995b: 167 (catalogue); Mackay & Mackay, 2002: 38 (U.S.A., New Mexico species key).

Myrmicinae Attini Trachymyrmex TRUE TRUE FALSE

Extant: 47 species, 1 subspecies

Fossil: 1 species

Trachymyrmex Forel, 1893e: 600 [as subgenus of Atta]. Type-species: Atta septentrionalis, by subsequent designation of Wheeler, W.M. 1911f: 174.

Taxonomic history

Trachymyrmex in Cryptoceridae, Attinae: Ashmead, 1905b: 384.

Trachymyrmex in Myrmicinae, Attini: Emery, 1895j: 770; Emery, 1913b: 251; Forel, 1917: 247; Wheeler, W.M. 1922a: 669; Emery, 1924d: 344; all subsequent authors.

Trachymyrmex as subgenus of Atta: Forel, 1893e: 600 Emery, 1895j: 770; Forel, 1893e: 600; Forel, 1899c: 30; Wheeler, W.M. 1907c: 670; Wheeler, W.M. 1910g: 141.

Trachymyrmex as subgenus of Cyphomyrmex: Emery, 1913b: 251; Emery, 1924d: 344; Donisthorpe, 1943g: 733.

Trachymyrmex as subgenus of Acromyrmex: Forel, 1917: 247.

Trachymyrmex as genus: Gallardo, 1916b: 242; Gallardo, 1916d: 318; Wheeler, W.M. 1916c: 11; Wheeler, W.M. 1922a: 669; Borgmeier, 1927c: 128; Borgmeier, 1950b: 384; Kusnezov, 1964: 63; Kempf, 1972a: 252.

Genus Trachymyrmex references

Wheeler, W.M. 1911g: 248 (all species key); Emery, 1924d: 344 (diagnosis, catalogue); Creighton, 1950a: 320 (North America species key); Kempf, 1972a: 252 (Neotropical catalogue); Smith, D.R. 1979: 1411 (North America catalogue); Brandão, 1991: 382 (catalogue); Bolton, 1994: 105 (synoptic classification); Bolton, 1995a: 1053 (census); Bolton, 1995b: 420 (catalogue); Mackay & Mackay, 2002: 38 (U.S.A., New Mexico species key); Mayhé-Nunes & Brandão, 2002: 672 (diagnosis, T. opulentus group key); Mayhé-Nunes & Brandão, 2005: 274 (T. iheringi group, key); Brandão& Mayhé-Nunes, 2007: 72 (species group phylogeny). Mayhé-Nunes & Brandão, 2007: 4 (T. jamaicensis group, key); Rabeling, Cover, et al. 2007: 4 (North America species, key).

Ichnotaxon attached to Attini

Ichnogenus †ATTAICHNUS

†Attaichnus Laza, 1982: 112. Included ichnospecies: †Attaichnus kuenzelii. [Ichnofossil, purportedly fossil traces of workings attributable to attine ants.]

Myrmicinae Stenammini Stenamma TRUE TRUE FALSE

Extant: 44 species

Fossil: 1 species

Stenamma Westwood, 1839: 219. Type-species: Stenamma westwoodii, by monotypy.

Taxonomic history

Stenamma in Myrmicidae, Myrmicidae: Emery, 1877a: 81.

Stenamma in Myrmicinae: Mayr, 1861: 55 [Myrmicidae]; Mayr, 1865: 23 [Myrmicidae]; Emery & Forel, 1879a: 456 [Myrmicidae]; Dalla Torre, 1893: 121.

Stenamma in Myrmicinae, Myrmicini: Emery, 1895j: 769; Wheeler, W.M. 1910g: 140; Wheeler, W.M. 1915h: 53; Kempf, 1972a: 242 (anachronism).

Stenamma in Myrmicinae, Pheidolini: Emery, 1914a: 40; Forel, 1917: 241; Emery, 1921f: 52 [subtribe Stenammini]; Wheeler, W.M. 1922a: 661; Karavaiev, 1934: 96; Kugler, C. 1994: 25.

Stenamma in Myrmicinae, Stenammini: Ashmead, 1905b: 383; Bolton, 1994: 106.

Junior synonyms of STENAMMA

Asemorhoptrum Mayr, 1861: 76. Type-species: Myrmica lippula, by monotypy.

Taxonomic history

Asemorhoptrum in Myrmicinae: Mayr, 1865: 20 [Myrmicidae]; Emery, 1877a: 81 [Myrmicidae]; Emery & Forel, 1879a: 460; [Myrmicidae].

Asemorhoptrum as junior synonym of Stenamma: André, 1883a: 310; all subsequent authors except the following.

[Asemorhoptrum as junior synonym of Proatta: Snelling, R.R. 1981: 395 (error).]

Theryella Santschi, 1921d: 68. Type-species: Theryella myops (provisional junior synonym of Stenamma punctiventre), by monotypy.

Taxonomic history

Theryella as junior synonym of Stenamma: Santschi, 1923a: 136; all subsequent authors except the following.

[Theryella as junior synonym of Proatta: Snelling, R.R. 1981: 395 (error).]

Genus Stenamma references

Roger, 1863b: 25, 27 (catalogue); Mayr, 1863: 395, 454 (catalogue); Mayr, 1865: 20, 23 (Asemorhoptrum, Stenamma diagnoses); André, 1883a: 311 (Europe & Algeria species key); Cresson, 1887: 261 (U.S.A. catalogue); Dalla Torre, 1893: 121 (catalogue); Forel, 1903a: 693 (India & Sri Lanka species key); Emery, 1908c: 306 (Palaearctic species key); Emery, 1916b: 127 (Italy species key); Emery, 1921f: 52 (diagnosis, catalogue); Arnol'di, 1928b: 215 (former U.S.S.R. species key); Arnol'di, 1933b: 599 (Russia species key); Buren, 1944a: 284 (U.S.A., Iowa species key); Creighton, 1950a: 135 (North America species key); Chapman & Capco, 1951: 153 (Asia checklist); Smith, M.R. 1957b: 141 (U.S.A. species key); Yasumatsu & Murakami, 1960: 28 (Japan species key); Smith, M.R. 1962a: 38 (Mexico and Central America species key); Gregg, 1963: 347 (U.S.A., Colorado species key); Wheeler, G.C. & Wheeler, J. 1963: 120 (U.S.A., North Dakota species key); Bernard, 1967: 126 (diagnosis, Western Europe species key); Kempf, 1972a: 242 (Neotropical catalogue); Snelling, R.R. 1973c: 5 (Western U.S.A. species key); Arnol'di, 1975: 1826 (former U.S.S.R. species key); Kutter, 1977c: 74 (Switzerland species key); Arnol'di & Dlussky, 1978: 535 (former European U.S.S.R. species key); Collingwood, 1978: 80 (Iberian Peninsula species key); Smith, D.R. 1979: 1357 (North America, catalogue); Allred, 1982: 443 (U.S.A., Utah species key); Gösswald, 1985: 297 (Germany species key); Wheeler, G.C. & Wheeler, J. 1986g: 33 (U.S.A., Nevada species key); Agosti & Collingwood, 1987b: 268 (Balkans species key); Kupyanskaya, 1990: 116 (Far Eastern Russia species key); Morisita, Kubota, et al. 1992: 14 (Japan species key); Arakelian, 1994: 29 (Armenia species key); Kupyanskaya, 1995: 344 (Far Eastern Russia species key); Bolton, 1995a: 1052 (census); Bolton, 1995b: 393 (catalogue); Seifert, 1996b: 155 (Central Europe species key); Skinner & Allen, 1996: 44 (Britain species key); Fernández, Baena & Palacio, 1996: 9 (Colombia species key); DuBois, 1998b: 215 (Palaearctic & Oriental species revision, keys); Mackay & Mackay, 2002: 209 (U.S.A., New Mexico species key); Lyu & Cho, 2004: 263 (Korea species key); Seifert, 2007: 143 (North and Central Europe species key); Branstetter, 2009: 43 (diagnosis).

Myrmicinae Solenopsidini Monomorium TRUE TRUE FALSE

Extant: 386 species, 30 subspecies

Fossil: 3 species

Monomorium Mayr, 1855: 452. Type-species: Monomorium monomorium (replacement name for Monomorium minutum, junior secondary homonym in Monomorium), by monotypy.

Taxonomic history

Monomorium in Myrmicidae, Myrmicidae: Emery, 1877a: 81.

Monomorium in Myrmicinae: Mayr, 1855: 452 [Myrmicidae]; Mayr, 1861: 71 [Myrmicidae]; Mayr, 1865: 22 [Myrmicidae]; Emery & Forel, 1879a: 456 [Myrmicidae]; Dalla Torre, 1893: 65.

Monomorium in Myrmicinae, Myrmicini: Forel, 1895b: 124; Emery, 1895j: 769; Forel, 1899c: 78; Wheeler, W.M. 1910g: 139.

Monomorium in Myrmicinae, Tetramoriini: Ashmead, 1905b: 383.

Monomorium in Myrmicinae, Monomoriini: Wheeler, W.M. 1915h: 45; Dlussky & Fedoseeva, 1988: 80.

Monomorium in Myrmicinae, Monomorium genus group: Ettershank, 1966: 81; in Solenopsis genus group: Bolton, 1987: 271.

Monomorium in Myrmicinae, Solenopsidini: Emery, 1914a: 41 [subtribe Monomoriini]; Arnold, 1916: 203; Forel, 1917: 242; Emery, 1922e: 166; Wheeler, W.M. 1922a: 662; Hölldobler & Wilson, 1990: 16; Jaffe, 1993: 10; Bolton, 1994: 106.

Junior synonyms of MONOMORIUM

Phacota Roger, 1862b: 260. Type-species: Phacota sichelii, by monotypy.

Taxonomic history

Phacota in Myrmicidae, Pheidolidae: Emery, 1877a: 81.

Phacota in Myrmicinae: Mayr, 1865: 22 [Myrmicidae]; Emery & Forel, 1879a: 464 [Myrmicidae]; Dalla Torre, 1893: 71.

Phacota in Myrmicinae, Myrmicini: Emery, 1895j: 769; Wheeler, W.M. 1910g: 139.

Phacota in Myrmicinae, Stenammini: Ashmead, 1905b: 383.

Phacota in Myrmicinae, Solenopsidini: Emery, 1914a: 41 [subtribe Monomoriini]; Forel, 1917: 243; Emery, 1922e: 187; Wheeler, W.M. 1922a: 662; all subsequent authors except the entry below; Hölldobler & Wilson, 1990: 16; Bolton, 1994: 106.

Phacota in Myrmicinae, Solenopsis genus group: Bolton, 1987: 271.

Phacota as genus: Bolton, 1987: 281.

Phacota as junior synonym of Monomorium: Ettershank, 1966: 82; Fernández, in Heterick, 2006: 79; Fernández, 2007b: 130.

Trichomyrmex Mayr, 1865: 19. Type-species: Trichomyrmex rogeri, by monotypy.

Taxonomic history

Trichomyrmex in Myrmicidae, Myrmicidae: Emery, 1877a: 81.

Trichomyrmex in Myrmicinae: Mayr, 1865: 19 [Myrmicidae]; Dalla Torre, 1893: 72.

Trichomyrmex in Myrmicinae, Myrmicini: Wheeler, W.M. 1910g: 140.

Trichomyrmex in Myrmicinae, Solenopsidini: Emery, 1914a: 41 [subtribe Monomoriini]; Forel, 1917: 243; Emery, 1922e: 185; Wheeler, W.M. 1922a: 662; Donisthorpe, 1943g: 733.

Trichomyrmex as junior synonym of Monomorium: Ettershank, 1966: 82; Smith, D.R. 1979: 1381; Bolton, 1987: 296.

†Lampromyrmex Mayr, 1868c: 93. Type-species: †Monomorium mayrianum (replacement name for †Lampromyrmex gracillimus, junior secondary homonym in Monomorium), by monotypy.

Taxonomic history

†Lampromyrmex in Myrmicidae, Pheidolidae: Emery, 1877a: 81.

†Lampromyrmex in Myrmicinae: Mayr, 1868c: 93 [Myrmicidae]; Dalla Torre, 1893: 78.

†Lampromyrmex in Myrmicinae, Solenopsidini: Emery, 1922e: 183 [subtribe Monomoriini]; Donisthorpe, 1943f: 655.

†Lampromyrmex as junior synonym of Monomorium: Wheeler, W.M. 1915h: 45.

†Lampromyrmex as subgenus of Monomorium: Emery, 1922e: 183; Wheeler, W.M. 1922a: 676.

†Lampromyrmex as junior synonym of Monomorium: Wheeler, W.M. 1915h: 45; Ettershank, 1966: 82; Smith, D.R. 1979: 1381; Bolton, 1987: 299.

Holcomyrmex Mayr, 1879: 671. Type-species: Holcomyrmex scabriceps, by subsequent designation of Bingham, 1903: 280.

Taxonomic history

[Holcomyrmex Smith, F. 1873: ix, nomen nudum, attributed to Mayr.]

Holcomyrmex in Myrmicinae, Myrmicini: Emery, 1895j: 769; Ashmead, 1905b: 383.

Holcomyrmex in Myrmicinae, Solenopsidini: Emery, 1922e: 181 [subtribe Monomoriini]; Donisthorpe, 1943f: 650.

Holcomyrmex as genus: Dalla Torre, 1893: 65; Forel, 1903a: 692; Bingham, 1903: 280.

Holcomyrmex as subgenus of Monomorium: Wheeler, W.M. 1910g: 139; Emery, 1915i: 190; Forel, 1917: 242; Emery, 1922e: 181; Wheeler, W.M. 1922a: 676.

Holcomyrmex as junior synonym of Monomorium: Emery, 1908h: 667; Ettershank, 1966: 82; Smith, D.R. 1979: 1381; Bolton, 1987: 295.

Epoecus Emery, 1893a: cclxxvi. Type-species: Epoecus pergandei, by monotypy.

Taxonomic history

Epoecus in Myrmicinae, Myrmicini: Emery, 1895j: 769; Wheeler, W.M. 1910g: 139.

Epoecus in Myrmicinae, Stenammini: Ashmead, 1905b: 383.

Epoecus in Myrmicinae, Solenopsidini: Emery, 1914a: 41 [subtribe Anergatini]; Forel, 1917: 243; Emery, 1922e: 204; Wheeler, W.M. 1922a: 663; Donisthorpe, 1943f: 643.

Epoecus as junior synonym of Monomorium: Ettershank, 1966: 82; Smith, D.R. 1979: 1381; Bolton, 1987: 287.

Wheeleriella Forel, 1907f: 145.

Taxonomic history

[Replacement name for Wheeleria Forel, 1905b: 171; junior homonym of Wheeleria Tutt, 1905: 37 (Lepidoptera).]

Wheeleriella in Myrmicinae, Myrmicini: Wheeler, W.M. 1910g: 139.

Wheeleriella in Myrmicinae, Solenopsidini: Emery, 1914a: 41 [subtribe Monomoriini]; Forel, 1917: 243; Emery, 1922e: 186; Wheeler, W.M. 1922a: 662; Donisthorpe, 1943g: 735.

Wheeleriella as junior synonym of Monomorium: Ettershank, 1966: 82; Smith, D.R. 1979: 1381; Bolton, 1987: 291.

Homonym replaced by Wheeleriella

Wheeleria Forel, 1905b: 171. Type-species: Wheeleria santschii, by monotypy.

Taxonomic history

[Junior homonym of Wheeleria Tutt, 1905: 37 (Lepidoptera).]

Epixenus Emery, 1908f: 556. Type-species: Epixenus andrei (junior secondary homonym in Monomorium, replaced by Monomorium advena), by subsequent designation of Wheeler, W.M. 1911f: 163.

Taxonomic history

Epixenus in Myrmicinae, Solenopsidini: Emery, 1914a: 41 [subtribe Monomoriini]; Forel, 1917: 243; Emery, 1922e: 184; Wheeler, W.M. 1922a: 662; Donisthorpe, 1943f: 643.

Epixenus as junior synonym of Monomorium: Brown & Wilson, 1957bb: 244; Ettershank, 1966: 82; Smith, D.R. 1979: 1381; Bolton, 1987: 287; Tinaut & Ortiz, 1988: 167.

Mitara Emery, 1913b: 261 [as subgenus of Monomorium]. Type-species: Monomorium laeve, by original designation.

Taxonomic history

Mitara in Myrmicinae, Solenopsidini: Arnold, 1916: 238; Forel, 1917: 242 [subtribe Monomoriini]; all subsequent authors.

Mitara as subgenus of Monomorium: Emery, 1913b: 261; Emery, 1915i: 190; Arnold, 1916: 238; Forel, 1917: 242.

Mitara as junior synonym of †Lampromyrmex: Emery, 1922e: 183; Wheeler, W.M. 1922a: 162, 676.

Mitara as genus: Donisthorpe, 1943f: 662.

Mitara as junior synoynm of Monomorium: Smith, D.R. 1979: 1381; Bolton, 1987: 299.

Chelaner Emery, 1914f: 410 [as subgenus of Monomorium]. Type-species: Monomorium (Chelaner) forcipatum, by subsequent designation of Emery, 1922e: 168.

Taxonomic history

Chelaner in Myrmicinae, Solenopsidini: Forel, 1917: 242 [subtribe Monomoriini]; Emery, 1922e: 168; all subsequent authors except the following.

Chelaner in Myrmicinae, Monomorium genus group: Ettershank, 1966: 81.

Chelaner in Myrmicinae, Monomoriini: Dlussky & Fedoseeva, 1988: 80.

Chelaner as subgenus of Monomorium: Emery, 1914f: 410; Forel, 1917: 242; Emery, 1922e: 168; Wheeler, W.M. 1922a: 676.

Chelaner as genus: Ettershank, 1966: 93; Taylor & Brown, D.R. 1985: 55; Wheeler, G.C. & Wheeler, J. 1985: 257; Dlussky & Fedoseeva, 1988: 80 (anachronism).

Chelaner (and its junior synonyms Notomyrmex, Protholcomyrmex, Schizopelta) as junior synonym of Monomorium: Bolton, 1987: 300; Heterick, 2001: 354.

Notomyrmex Emery, 1915i: 190 [as subgenus of Monomorium]. Type-species: Atta antarctica, by original designation.

Taxonomic history

Notomyrmex as subgenus of Monomorium: Emery, 1915i: 190; Forel, 1917: 242; Emery, 1922e: 168; Wheeler, W.M. 1922a: 676; Kusnezov, 1956: 20.

Notomyrmex as genus: Kusnezov, 1957a: 269 (in key); Kusnezov, 1960b: 343; Kusnezov, 1962b: 160; Kusnezov, 1964: 60.

Notomyrmex as junior synonym of Chelaner: Ettershank, 1966: 93.

Parholcomyrmex Emery, 1915i: 190 [as subgenus of Monomorium]. Type-species: Myrmica gracillima (junior synonym of Atta destructor), by original designation.

Taxonomic history

Parholcomyrmex as subgenus of Monomorium: Emery, 1915i: 190; Forel, 1917: 242; Emery, 1922e: 179; Wheeler, W.M. 1922a: 676; subsequent authors to the following.

Parholcomyrmex as junior synonym of Monomorium: Ettershank, 1966: 82; Smith, D.R. 1979: 1381; Bolton, 1987: 296.

[Paraholcomyrmex Emery, 1915i: 191, Forel, 1917: 242, incorrect subsequent spellings.]

Xeromyrmex Emery, 1915i: 190 [as subgenus of Monomorium]. Type-species: Formica salomonis, by original designation.

Taxonomic history

Xeromyrmex as subgenus of Monomorium: Forel, 1917: 242; Emery, 1922e: 175; Wheeler, W.M. 1922a: 677.

Xeromyrmex as junior synonym of Monomorium: Ettershank, 1966: 82; Smith, D.R. 1979: 1381; Bolton, 1987: 294.

Syllophopsis Santschi, 1915c: 259 [as subgenus of Monomorium.] Type-species: Monomorium modestum, by monotypy.

Taxonomic history

Syllophopsis in Myrmicinae, Solenopsidini: Forel, 1917: 242 [subtribe Monomoriini]; Emery, 1922e: 175; all subsequent authors to the following.

Syllophopsis in Myrmicinae, Monomorium genus group: Ettershank, 1966: 81.

Syllophopsis in Myrmicinae, Monomoriini: Dlussky & Fedoseeva, 1988: 80.

Syllophopsis as subgenus of Monomorium: Santschi, 1915c: 259; Forel, 1917: 242; Emery, 1922e: 175; Wheeler, W.M. 1922a: 676; Donisthorpe, 1943g: 729; Arnold, 1952a: 465.

Syllophopsis as genus: Santschi, 1921c: 119; Santschi, 1936a: 32; Ettershank, 1966: 100.

Syllophopsis as junior synonym of Monomorium: Bolton, 1987: 287.

[Syllopsis Santschi, 1921c: 120, incorrect subsequent spelling.]

Corynomyrmex Viehmeyer, 1916a: 133 [as subgenus of Monomorium]. Type-species: Monomorium (Corynomyrmex) hospitum, by monotypy.

Taxonomic history

Corynomyrmex as subgenus of Monomorium: Viehmeyer, 1916a: 133; Forel, 1917: 242; Emery, 1922e: 174.

Corynomyrmex as junior synonym of Monomorium: Ettershank, 1966: 82 [provisional]; Smith, D.R. 1979: 1381; Bolton, 1987: 287.

Isolcomyrmex Santschi, 1917b: 296 [as subgenus of Monomorium]. Type-species: Holcomyrmex santschii (junior secondary homonym in Monomorium, replaced by Monomorium santschianum), by monotypy.

Taxonomic history

Isolcomyrmex as subgenus of Monomorium: Santschi, 1917b: 296; Wheeler, W.M. 1922a: 676.

Isolcomyrmex as junior synonym of Monomorium: Ettershank, 1966: 82; Smith, D.R. 1979: 1381; Bolton, 1987: 296.

[Isholcomyrmex Santschi, 1936a: 32, incorrect subsequent spelling.]

Paraphacota Santschi, 1919d: 90. Type-species: Paraphacota surcoufi, by monotypy.

Taxonomic history

[Type-species not Phacota noualhieri, unjustified subsequent designation by Santschi, 1927d: 245; repeated in Ettershank, 1966: 82 and Bolton, 1973a: 352.]

Paraphacota in Myrmicinae, Solenopsidini: Wheeler, W.M. 1922a: 663; Donisthorpe, 1943f: 681.

Paraphacota as junior synonym of Monomorium: Santschi, 1927d: 243; Ettershank, 1966: 82; Smith, D.R. 1979: 1381; Bolton, 1987: 293.

Equestrimessor Santschi, 1919d: 92 [as subgenus of Monomorium]. Type-species: Holcomyrmex chobauti, by subsequent designation of Donisthorpe, 1943f: 644.

Taxonomic history

Equestrimessor as subgenus of Monomorium: Santschi, 1919d: 92;Wheeler, W.M. 1922a: 677; Santschi, 1936a: 32.

Equestrimessor as junior synonym of Monomorium: Ettershank, 1966: 82; Smith, D.R. 1979: 1381; Bolton, 1987: 287, 297.

[Equessimessor Santschi, 1936a: 32, incorrect subsequent spelling.]

Xenhyboma Santschi, 1919g: 405. Type-species: Xenhyboma mystes (junior synonym of Monomorium medinae), by monotypy.

Taxonomic history

Xenhyboma in Myrmicinae, Solenopsidini: Donisthorpe, 1943g: 736.

Xenhyboma as junior synonym of Monomorium: Ettershank, 1966: 82 [provisional]; Smith, D.R. 1979: 1381; Espadaler, 1982b: 112; Bolton, 1987: 293.

Protholcomyrmex Wheeler, W.M. 1922a: 162 [as subgenus of Monomorium]. Type-species: Monomorium rothsteini, by original designation.

Taxonomic history

Protholcomyrmex as junior synonym of Chelaner: Ettershank, 1966: 93.

Ireneidris Donisthorpe, 1943c: 81. Type-species: Ireneidris myops (junior synonym of Monomorium talpa), by original designation.

Taxonomic history

Ireneidris in Myrmicinae, Solenopsidini [subtribe Monomoriini]: Donisthorpe, 1943c: 82.

Ireneidris as junior synonym of Monomorium: Ettershank, 1966: 82; Smith, D.R. 1979: 1381; Bolton, 1987: 298.

Schizopelta McAreavey, 1949: 14. Type-species: Schizopelta falcata, by original designation.

Taxonomic history

Schizopelta as junior synonym of Chelaner: Ettershank, 1966: 93.

Nothidris Ettershank, 1966: 105. Type-species: Monomorium latastei, by original designation.

Taxonomic history

Nothidris in Myrmicinae, Megalomyrmex genus group: Ettershank, 1966: 81; in Solenopsis genus group: Bolton, 1987: 271.

Nothidris in Myrmicinae, Megalomyrmecini: Dlussky & Fedoseeva, 1988: 80.

Nothidris in Myrmicinae, Solenopsidini: Kempf, 1972a: 165; Hölldobler & Wilson, 1990: 16; Jaffe, 1993: 10; Bolton, 1994: 106.

Nothidris as junior synonym of Monomorium: Fernández, in Heterick, 2006: 79; Fernández, 2007b: 130.

Pharaophanes Bernard, 1967: 168 (attributed to Santschi) [as subgenus of Monomorium]. Type-species: Formica pharaonis, by monotypy.

Taxonomic history

[Pharaophanes Bernard, 1953b: 238 (attributed to Santschi) [as subgenus of Monomorium]. Unavailable name; proposed without designation of type-species. Species included by Bernard (1953) are all referable to Monomorium: Bolton, 1987: 288; Bolton, 1995b: 42 (see also note in Baroni Urbani, 1971c: 90).]

Pharaophanes as junior synonym of Monomorium: Bolton, 2003: 213.

Antichthonidris Snelling, R.R. 1975: 5. Type-species: Monomorium denticulatum, by original designation.

Taxonomic history

Antichthonidris in Myrmicinae, Solenopsis genus group: Bolton, 1987: 271.

Antichthonidris in Myrmicinae, Solenopsidini: Hölldobler & Wilson, 1990: 16; Jaffe, 1993: 10; Bolton, 1994: 106.

Antichthonidris as junior synonym of Monomorium: Heterick, 2001: 361.

Epelysidris Bolton, 1987: 279. Type-species: Epelysidris brocha, by original designation.

Taxonomic history

Epelysidris in Myrmicinae, Solenopsis genus group: Bolton, 1987: 271.

Epelysidris in Myrmicinae, Solenopsidini: Hölldobler & Wilson, 1990: 16; Bolton, 1994: 106; Bolton, 1995a: 1049; Bolton, 1995b: 188.

Epelysidris as junior synonym of Monomorium: Fernández, in Heterick, 2006: 79; Fernández, 2007b: 130.

Genus Monomorium references

Roger, 1863b: 30, 31 (Phacota, Monomorium catalogues); Mayr, 1863: 429, 440 (Monomorium, Phacota catalogues); Mayr, 1865: 19, 22 (Trichomyrmex, Monomorium, Phacota diagnoses); Mayr, 1867a: 95 (Monomorium diagnosis); Mayr, 1876: 100 (Australia Monomorium species key); Emery, 1881b: 530 (Mediterranean & Red Sea Monomorium species key); André, 1883a: 331, 343 (Europe & Algeria Monomorium, Holcomyrmex species keys); André, 1883b: 397 (Phacota); Cresson, 1887: 262 (U.S.A. Monomorium catalogue); Nasonov, 1889: 69 (Russia Monomorium species key); Dalla Torre, 1893: 65, 71, 72, 78 (Holcomyrmex, Monomorium, Phacota, Trichomyrmex, †Lampromyrmex catalogues); Forel, 1903a: 685, 692 (India & Sri Lanka Monomorium, Holcomyrmex species keys); Bingham, 1903: 200, 282 (India, Sri Lanka & Burma Monomorium, Holcomyrmex species keys); Ruzsky, 1905b: 633 (Russian Empire Monomorium species key); Emery, 1908h: 664 (Palaearctic Monomorium species key); Emery, 1915i: 190 (Monomorium subgenera key); Arnold, 1916: 203, 204 (Monomorium diagnosis, South Africa species key); Emery, 1916b: 161 (Italy Monomorium species key); Santschi, 1921c: 121 (Syllophopsis species key); Emery, 1922e: 166 (Monomorium diagnosis, subgenera key, catalogue); Emery, 1922e: 168 (M. (Chelaner) & M. (Notomyrmex) diagnoses, catalogues); Emery, 1922e: 170 (M. (Monomorium) diagnosis, catalogue); Emery, 1922e: 174 (M. (Corynomyrmex) diagnosis, catalogue); Emery, 1922e: 175 (M. (Syllophopsis) & M. (Xeromyrmex) diagnoses, catalogues); Emery, 1922e: 179 (M. (Parholcomyrmex) diagnosis, catalogue); Emery, 1922e: 181 (M. (Holcomyrmex) diagnosis, catalogue); Emery, 1922e: 183 (M. (Lampromyrmex) diagnosis, catalogue); Emery, 1922e: 184 (Epixenus diagnosis, catalogue); Emery, 1922e: 185 (Trichomyrmex diagnosis, catalogue); Emery, 1922e: 186 (Wheeleriella diagnosis, catalogue); Emery, 1922e: 187 (Phacota diagnosis, catalogue); Emery, 1922e: 204 (Epoecus diagnosis, catalogue); Wheeler, W.M. 1922a: 161, 675, 862, 1026 (Monomorium diagnosis, subgenera key, Afrotropical, Malagasy catalogues); Menozzi, 1933b: 64 (Israel Monomorium species key); Santschi, 1936a: 33 (Mediterranean & Africa M. (Xeromyrmex) key); Buren, 1944a: 289 (U.S.A., Iowa Monomorium species key); Kusnezov, 1949a: 431 (Argentina Monomorium species); Creighton, 1950a: 217 (North America Monomorium species key); Creighton, 1950a: 239 (Epoecus, review of genus); Chapman & Capco, 1951: 162, 169 (Asia Ireneidris, Monomorium, Trichomyrmex, Wheeleriella checklists); Bernard, 1955b: 282 (Epixenus species key); Brown, 1958h: 28 (New Zealand Monomorium species key); Gregg, 1963: 367 (U.S.A., Colorado Monomorium species key); Ettershank, 1966: 82, 93, 100 (Monomorium, Chelaner, Syllophopsis diagnoses, reviews of genera, checklists); Bernard, 1967: 165 (Monomorium diagnosis, Western Europe species key); Kempf, 1972a: 143 (Neotropical Monomorium catalogue); Kempf, 1972a: 165 (Nothidris catalogue); Alayo, 1974: 14 (Cuba Monomorium species key); Snelling, R.R. 1975: 5 (Nothidris all species key); Snelling, R.R. & Hunt, 1976: 79 (Nothidris all species key); Arnol'di & Dlussky, 1978: 538 (former European U.S.S.R. Monomorium species key); Collingwood, 1978: 82 (Iberian Peninsula Monomorium species key); Smith, D.R. 1979: 1381 (North America Monomorium catalogue); Tohmé & Tohmé, 1980a: 1087 (Lebanon & Syria Epixenus species key); Collingwood, 1985: 267 (Saudi Arabia Monomorium species key); Taylor & Brown, D.R. 1985: 55, 70 (Australia Chelaner, Monomorium catalogues); DuBois, 1986: 74 (Nearctic Monomorium species revision, key); Bolton, 1987: 281 (Phacota diagnosis, review of genus); Bolton, 1987: 283 (Antichthonidris, review of genus); Bolton, 1987: 284 (Nothidris review of genus); Bolton, 1987: 287 (Monomorium diagnosis, review of genus, Afrotropical species revision, key); Taylor, 1987a: 19, 40 and Taylor, 1987b: 2 (Australia, New Caledonia & New Zealand Monomorium checklist); Agosti & Collingwood, 1987b: 272 (Balkans Monomorium species key); Hölldobler & Wilson, 1990: 14 (synoptic classification); Dlussky, Soyunov & Zabelin, 1990: 231 (Turkmenistan Monomorium species key); Brandão, 1991: 326, 357 (Neotropical Monomorium catalogue); Brandão, 1991: 361 (Nothidris catalogue); Morisita, Kubota, et al. 1992: 38 (Japan Monomorium species key); Arakelian, 1994: 46 (Armenia Monomorium species key); Bolton, 1995a: 1048, 1050, 1051 (census); Bolton, 1995b: 67, 258, 292, 316 (Antichthonidris, Monomorium, Nothidris, Phacota catalogues); Wu, J. & Wang, 1995: 88 (China Monomorium species key); Collingwood & Agosti, 1996: 337 (Saudi Arabia Monomorium species key); Radchenko, 1997f: 211 (M. scabriceps group revision, key); Collingwood & Prince, 1998: 19 (Portugal Monomorium species key); Shattuck, 1999: 145 (Australia synopsis); Heterick, 2001: 367 (Australia Monomorium species revision, key); Zhou, 2001b: 112 (China, Guangxi Monomorium species key); Mackay & Mackay, 2002: 127 (U.S.A., New Mexico Monomorium species key); Heterick, 2006: 84 (Malagasy Monomorium species revision, key); Fernández, 2007b: 139 (Neotropical Monomorium species key); Terayama, 2009: 154 (Taiwan species key); Heterick, 2009: 158 (south-western Australia species key); Aldawood & Sharaf, 2011: 52 (Saudi Arabia M. monomorium group key).

Myrmicinae Solenopsidini Solenopsis TRUE TRUE FALSE

Extant: 183 species, 75 subspecies

Fossil: 8 species

Solenopsis Westwood, 1840b: 86. Type-species: Solenopsis mandibularis (junior synonym of Atta geminata), by monotypy.

Taxonomic history

Solenopsis in Poneridae, Attidae: Smith, F. 1858b: 177.

Solenopsis in Attidae: Smith, F. 1860a: 74; Smith, F. 1861b: 48.

Solenopsis in Myrmicidae: Smith, F. 1871a: 333; Cresson, 1887: 262.

Solenopsis in Myrmicidae, Pheidolidae: Emery, 1877a: 81.

Solenopsis in Myrmicinae: Mayr, 1865: 24 [Myrmicidae]; Emery & Forel, 1879a: 464 [Myrmicidae]; Dalla Torre, 1893: 75.

Solenopsis in Myrmicinae, Solenopsis genus group: Ettershank, 1966: 81; Bolton, 1987: 271.

Solenopsis in Myrmicinae, Solenopsidini: Forel, 1893a: 164; Forel, 1895b: 130; Emery, 1895j: 770; Forel, 1899c: 79; Ashmead, 1905b: 383; Wheeler, W.M. 1910g: 140; Emery, 1914a: 41; Arnold, 1916: 242; Forel, 1917: 243; Emery, 1922e: 195; Wheeler, W.M. 1922a: 663; Kempf, 1972a: 232; Smith, D.R. 1979: 1384; Dlussky & Fedoseeva, 1988: 80; Hölldobler & Wilson, 1990: 16; Jaffe, 1993: 10; Bolton, 1994: 106.

Junior synonyms of SOLENOPSIS

Diplorhoptrum Mayr, 1855: 449. Type-species: Formica fugax, by monotypy.

Taxonomic history

Diplorhoptrum in Myrmicinae: Mayr, 1855: 449 [Myrmicidae]; Mayr, 1861: 72 [Myrmicidae].

Diplorhoptrum in Myrmicinae, Solenopsidini: Emery, 1922e: 195; all subsequent authors.

Diplorhoptrum as genus: Mayr, 1855: 449; Baroni Urbani, 1968b: 68; Kutter, 1977c: 98; Bernard, 1978a: 543; Arnol'di & Dlussky, 1978: 539; Collingwood, 1979: 64; Dlussky & Fedoseeva, 1988: 80; Kupyanskaya, 1990: 132; Atanasov & Dlussky, 1992: 167; Dlussky & Radchenko, 1994: 102; Baroni Urbani, 1995: 20 (footnote).

Diplorhoptrum as subgenus of Solenopsis: Creighton, 1930b: 43; Santschi, 1934e: 566 (in text), 568; Creighton, 1950a: 233; Smith, M.R. 1951a: 813; Kusnezov, 1956: 27; Kusnezov, 1957a: 273; Kusnezov, 1962b: 160; Smith, M.R. 1967: 358.

Diplorhoptrum as junior synonym of Solenopsis: Mayr, 1862: 751; Roger, 1863b: 32; Mayr, 1863: 453; Emery & Forel, 1879a: 464; Dalla Torre, 1893: 75; Forel, 1915d: 9; Emery, 1922e: 195; Wheeler, W.M. 1922a: 877; Donisthorpe, 1943f: 639; Ettershank, 1966: 134; Kempf, 1972a: 232; Smith, D.R. 1979: 1384; Bolton, 1987: 285; Bolton, 1994: 106; Bolton, 1995b: 27.

Octella Forel, 1915b: 70 [as subgenus of Oligomyrmex]. Type-species: Oligomyrmex (Octella) pachycerus, by original designation.

Taxonomic history

Octella as subgenus of Oligomyrmex: Forel, 1917: 243; Wheeler, W.M. 1922a: 687; Emery, 1924d: 218.

Octella as junior synonym of Oligomyrmex: Ettershank, 1966: 119.

Octella as junior synonym of Solenopsis: Taylor, 1991db: 611.

Synsolenopsis Forel, 1918b: 155 [as subgenus of Solenopsis]. Type-species: Solenopsis (Synsolenopsis) bruchi (junior primary homonym in Solenopsis, replaced by Solenopsis bruchiella), by monotypy.

Taxonomic history

Synsolenopsis as subgenus of Solenopsis: Forel, 1918b: 155; Emery, 1922e: 196.

Synsolenopsis as genus: Creighton, 1930b: 41 (in text); Kusnezov, 1953c: 341; Kusnezov, 1957a: 272; Kusnezov, 1962b: 160; Kusnezov, 1964: 61.

Synsolenopsis as junior synonym of Solenopsis: Ettershank, 1966: 134.

Diagyne Santschi, 1923c: 268 [as subgenus of Solenopsis]. Type-species: Solenopsis succinea by monotypy.

Taxonomic history

Diagyne in Myrmicinae, Solenopsidini: Donisthorpe, 1943f: 638.

Diagyne as subgenus of Solenopsis: Santschi, 1923c: 268; Creighton, 1930b: 42; Kusnezov, 1962b: 160.

Diagyne junior synonym of Solenopsis: Ettershank, 1966: 134.

Labauchena Santschi, 1930d: 81. Type-species: Labauchena daguerrei, by monotypy.

Taxonomic history

Labauchena in Myrmicinae, Solenopsidini: Santschi, 1930d: 81; all subsequent authors.

Labauchena as junior synonym of Solenopsis: Ettershank, 1966: 134.

Euophthalma Creighton, 1930b: 43 [as subgenus of Solenopsis]. Type-species: Myrmica globularia, by original designation.

Taxonomic history

Euophthalma as junior synonym of Solenopsis: Ettershank, 1966: 134.

Oedaleocerus Creighton, 1930b: 43 [as subgenus of Solenopsis]. Type-species: Solenopsis angulata, by original designation.

Taxonomic history

Oedaleocerus in Myrmicinae, Solenopsidini: Donisthorpe, 1943f: 677.

Oedaleocerus as subgenus of Solenopsis: Creighton, 1930b: 43.

Oedaleocerus as genus: Kusnezov, 1957a: 272; Kusnezov, 1962b: 160; Kusnezov, 1964: 61.

Oedaleocerus as junior synonym of Solenopsis: Ettershank, 1966: 134.

Bisolenopsis Kusnezov, 1953d: 9. Type-species: Bisolenopsis sea, by monotypy.

Taxonomic history

Bisolenopsis in Myrmicinae, Solenopsidini: Kusnezov, 1953d: 9; Kusnezov, 1957a: 272; Kusnezov, 1962b: 159; Kusnezov, 1964: 61.

Bisolenopsis as junior synonym of Solenopsis: Ettershank, 1966: 134.

[Disolenopsis Snelling, R.R. 1981: 397, incorrect subsequent spelling.]

Paranamyrma Kusnezov, 1953d: 17. Type-species: Paranamyrma solenopsidis, by monotypy.

Taxonomic history

[Paranamyrma also described as new by Kusnezov, 1954a: 9.]

Paranamyrma in Myrmicinae, Solenopsidini: Kusnezov, 1957a: 267 (in key); Kusnezov, 1962b: 160; Kusnezov, 1964: 61.

Paranamyrma as junior synonym of Solenopsis: Ettershank, 1966: 134.

Granisolenopsis Kusnezov, 1956: 26 [as subgenus of Solenopsis] (diagnosis in key). Type-species: Solenopsis (Granisolenopsis) granivora, by original designation.

Taxonomic history

[Granisolenopsis also described as new by Kusnezov, 1957a: 277.]

Granisolenopsis as junior synonym of Solenopsis: Ettershank, 1966: 134.

Lilidris Kusnezov, 1956: 26 (diagnosis in key). Type-species: Lilidris metatarsalis, by original designation.

Taxonomic history

[Lilidris also described as new by Kusnezov, 1957a: 274 and Kusnezov, 1958b: 189.]

Lilidris in Myrmicinae, Solenopsidini: Kusnezov, 1957a: 274; Kusnezov, 1958b: 189; Kusnezov, 1962b: 159; Kusnezov, 1964: 61.

Lilidris as junior synonym of Solenopsis: Ettershank, 1966: 134.

Genus Solenopsis references

Smith, F. 1858b: 177 (diagnosis); Roger, 1863b: 32 (catalogue); Mayr, 1863: 453 (catalogue); Mayr, 1865: 24 (diagnosis); Mayr, 1867a: 109 (diagnosis); Mayr, 1870b: 996 (all species key); André, 1883b: 387 (Europe & Algeria species key); Cresson, 1887: 262 (U.S.A. catalogue); Dalla Torre, 1893: 75 (catalogue); Emery, 1896g: 82 (New World species key); Forel, 1903a: 689 (India & Sri Lanka species key); Bingham, 1903: 158 (India, Sri Lanka & Burma species key); Emery, 1909a: 29 (Palaearctic species key); Emery, 1916b: 165 (Italy species key); Arnold, 1916: 242 (diagnosis); Emery, 1922e: 195 (diagnosis, catalogue); Wheeler, W.M. 1922a: 163, 877, 1028 (diagnosis, Afrotropical, Malagasy catalogues); Emery, 1924d: 218 (Oligomyrmex (Octella) diagnosis, catalogue); Creighton, 1930b: 42 (subgenera key); Creighton, 1930b: 45 (S. (Solenopsis) species key); Creighton, 1930b: 106 (S. (Euophthalma) species key); Cole, 1942: 361 (U.S.A., Utah species key); Bernard, 1950a: 5 (France species key); Creighton, 1950a: 228 (North America species key); Chapman & Capco, 1951: 167 (Asia checklist); Wilson, 1952b: 50 (South America S. saevissima complex, key); Kusnezov, 1953c: 347 (Synsolenopsis species key); Gregg, 1963: 370 (U.S.A., Colorado species key); Snelling, R.R. 1963: 1 (U.S.A S. geminata group species); Ettershank, 1966: 134 (diagnosis, review of genus, checklist); Bernard, 1967: 170 (diagnosis, Western Europe species key); Kusnezov, 1969: 37 (Bisolenopsis species key); Buren, 1972: 1 (S. geminata group, partial revision); Kempf, 1972a: 232 (Neotropical catalogue); Snelling, R.R. & Hunt, 1976: 81 (Chile species key); Kutter, 1977c: 99 (Switzerland species key); Arnol'di & Dlussky, 1978: 539 (former European U.S.S.R. species key); Collingwood, 1978: 82 (Iberian Peninsula species key); Bernard, 1978a: 573 (France species key); Smith, D.R. 1979: 1385 (North America catalogue); Allred, 1982: 443 (U.S.A., Utah species key); Gösswald, 1985: 302 (Germany species key); Taylor & Brown, D.R. 1985: 87 (Australia catalogue); Wheeler, G.C. & Wheeler, J. 1986g: 50 (U.S.A., Nevada species key); Taylor, 1987a: 72 (Australia, New Caledonia checklist); Agosti & Collingwood, 1987b: 273 (Balkans species key); Bolton, 1987: 285 (synonymy notes); Thompson & Johnson, 1989: 697 (U.S.A., Florida species key); Dlussky, Soyunov & Zabelin, 1990: 240 (Turkmenistan species key); Brandão, 1991: 378 (Neotropical catalogue); Trager, 1991: 141 (S. geminata group revision, key); Morisita, Kubota, et al. 1992: 41 (Japan species key); Arakelian, 1994: 48 (Armenia species key); Dlussky & Radchenko, 1994: 102 (Central Palaearctic species key); Bolton, 1995a: 1052 (census); Bolton, 1995b: 386 (catalogue); Wu, J. & Wang, 1995: 70 (China species key); Collingwood & Agosti, 1996: 358 (Saudi Arabia species key); Collingwood & Prince, 1998: 18 (Portugal species key); Shattuck, 1999: 165 (Australia synopsis); Taber, 2000: 234 (U.S.A. S. geminata group, key); Snelling, R.R. 2001: 522 (Puerto Rico S. fugax group, key); Mackay & Mackay, 2002: 196 (U.S.A., New Mexico species key); Terayama, 2009: 157 (Taiwan species key); Heterick, 2009: 176 (south-western Australia species key).

Myrmicinae Myrmicini Manica TRUE TRUE FALSE

6 species

Manica Jurine, 1807: 276. Type-species: Formica rubida, by subsequent designation of Wheeler, W.M. 1911f: 166.

Taxonomic history

Manica in Myrmicinae, Myrmicini: Wheeler, W.M. 1922a: 660; all subsequent authors.

Manica as junior synonym of Myrmica: Roger, 1863b: 28; Mayr, 1863: 431; Emery & Forel, 1879a: 460; Dalla Torre, 1893: 108; Forel, 1915d: 9.

Manica as subgenus of Myrmica: Emery, 1921f: 42; Wheeler, W.M. 1922a: 660; Smith, M.R. 1951a: 791.

Manica as genus: Jurine, 1807: 276; Weber, 1947: 439; Creighton, 1950a: 105; Smith, M.R. 1958c: 113; all subsequent authors.

Junior synonyms of MANICA

Neomyrma Forel, 1914a: 275 [as subgenus of Aphaenogaster]. Type-species: Aphaenogaster (Neomyrma) calderoni (junior synonym of Myrmica bradleyi), by monotypy.

Taxonomic history

Neomyrma as subgenus of Aphaenogaster: Forel, 1914a: 275.

Neomyrma as subgenus of Myrmica: Forel, 1915c: 364 (in text); Emery, 1915d: 69; Emery, 1916b: 120; Forel, 1917: 241.

Neomyrma as genus: Bondroit, 1918: 97.

Neomyrma as junior synonym of Myrmica: Donisthorpe, 1916b: 242.

Neomyrma as junior synonym of Manica: Emery, 1921f: 42; Wheeler, W.M. 1922a: 660.

Oreomyrma Wheeler, W.M. 1914d: 118 (in text) [as subgenus of Myrmica]. Type-species: Formica rubida (quoted by Wheeler as Myrmica rubida), by original designation.

Taxonomic history

Oreomyrma as junior synonym of Neomyrma: Wheeler, W.M. 1915a: 50; Forel, 1915c: 364; Forel, 1915d: 9; Donisthorpe, 1916b: 242. [Manica and Oreomyrma share the same type-species, synonymy is therefore absolute.]

Genus Manica references

Wheeler, W.M. 1914d: 119 (U.S.A. species key); Emery, 1921f: 42 (diagnosis, catalogue); Creighton, 1950a: 108 (North America species key); Bernard, 1967: 125 (diagnosis); Smith, D.R. 1979: 1352 (North America catalogue); Allred, 1982: 440 (U.S.A., Utah species key); Wheeler, G.C. & Wheeler, J. 1986g: 25 (U.S.A., Nevada species key); Morisita, Kubota, et al. 1992: 13 (Japan species key); Bolton, 1994: 106 (synoptic classification); Bolton, 1995a: 1050 (census); Bolton, 1995b: 249 (catalogue).

Myrmicinae Myrmicini Myrmica TRUE TRUE FALSE

Extant: 171 species, 2 subspecies

Fossil: 5 species

Myrmica Latreille, 1804: 179. Type-species: Formica rubra, by subsequent designation of Latreille, 1810: 437; see also Yarrow, 1955b: 113.

Taxonomic history

Myrmica in Myrmicites: Lepeletier de Saint-Fargeau, 1835: 180.

Myrmica in Poneridae, Myrmicidae: Smith, F. 1858b: 114.

Myrmica in Myrmicidae: Smith, F. 1871a: 324; Cresson, 1887: 260.

Myrmica in Myrmicinae: Mayr, 1855: 396 [Myrmicidae]; Smith, F. 1857a: 70 [Myrmicidae]; Mayr, 1861: 62 [Myrmicidae]; Mayr, 1865: 19 [Myrmicidae]; Emery & Forel, 1879a: 460 [Myrmicidae]; Dalla Torre, 1893: 108.

Myrmica in Myrmicinae, Myrmicini: Forel, 1895b: 125; Emery, 1895j: 769; Forel, 1899c: 63; Ashmead, 1905b: 383; Wheeler, W.M. 1910g: 140; Emery, 1914a: 40; Wheeler, W.M. 1915h: 59; Forel, 1917: 241; Bondroit, 1918: 96; Emery, 1921f: 36; Wheeler, W.M. 1922a: 660; all subsequent authors.

Junior synonyms of MYRMICA

Sifolinia Emery, 1907: 49. Type-species: Sifolinia laurae, by monotypy.

Taxonomic history

Sifolinia in Myrmicinae, Pheidolini: Emery, 1914a: 40; Forel, 1917: 241; Emery, 1922e: 117; Wheeler, W.M. 1922a: 661.

Sifolinia in Myrmicinae, Myrmicini: Wheeler, W.M. 1910g: 139; Collingwood, 1979: 58; Dlussky & Fedoseeva, 1988: 79.

Sifolinia as junior synonym of Myrmica: Brown, 1973b: 184 [provisional]; Bolton, 1988a: 3; Radchenko & Elmes, 2003a: 219.

†Nothomyrmica Wheeler, W.M. 1915h: 60. Type-species: †Macromischa rudis, by original designation.

Taxonomic history

[†Nothomyrmica Wheeler, W.M. 1908g: 413, and Wheeler, W.M. 1910g: 167, nomina nuda.]

†Nothomyrmica in Myrmicinae, Myrmicini: Wheeler, W.M. 1915h: 60; all subsequent authors except the following.

†Nothomyrmica in Myrmicinae, Tetramoriini: Donisthorpe, 1943f: 675 (error).

†Nothomyrmica in Myrmicinae, Leptothoracini: Dlussky & Fedoseeva, 1988: 79; Bolton, 1994: 106; Bolton, 1995b: 292.

†Nothomyrmica incertae sedis in Myrmicinae, Myrmicini: Bolton, 2003: 62.

†Nothomyrmica as junior synonym of Myrmica: Radchenko, Dlussky & Elmes, 2007: 1495.

Sommimyrma Menozzi, 1925d: 25. Type-species: Sommimyrma symbiotica, by original designation.

Taxonomic history

Sommimyrma in Myrmicinae, Myrmicini: Donisthorpe, 1943g: 726.

Sommimyrma as junior synonym of Myrmica: Brown, 1973b: 184 [provisional]; Bolton, 1988a: 3; Radchenko & Elmes, 2003a: 224.

Symbiomyrma Arnol'di, 1930c: 267. Type-species: Symbiomyrma karavajevi, by monotypy.

Taxonomic history

[Symbiomyrma also described as new by Arnol'di, 1933a: 41.]

Symbiomyrma in Myrmicinae, Pheidolini: Karavaiev, 1934: 102.

Symbiomyrma in Myrmicinae, Myrmicini: Donisthorpe, 1943g: 729.

Symbiomyrma as genus: Karavaiev, 1934: 102; Seifert, 1994: 15; Seifert, 1996b: 236.

Symbiomyrma as junior synonym of Sifolinia: Samsinak, 1964: 156.

Symbiomyrma as junior synonym of Myrmica: Bolton, 1988a: 3; Bolton, 1994: 106; Radchenko & Elmes, 2003a: 219.

Paramyrmica Cole, 1957a: 37. Type-species: Paramyrmica colax, by original designation.

Taxonomic history

Paramyrmica in Myrmicinae, Myrmicini: Wheeler, G.C. & Wheeler, J. 1985: 257.

Paramyrmica as genus: Gregg, 1961: 215; Smith, M.R. 1967: 350; Smith, D.R. 1979: 1351.

Paramyrmica as junior synonym of Myrmica: Brown, 1973b: 183 [provisional]; Bolton, 1988a: 3.

Dodecamyrmica Arnol'di, 1968: 1803 [as subgenus of Myrmica]. Type-species: Myrmica arnoldii, by original designation.

Taxonomic history

Dodecamyrmica as junior synonym of Myrmica: Brown, 1973b: 180 [provisional]; Francoeur, 1981: 759; Bolton, 1988a: 4.

Genus Myrmica references

Mayr, 1855: 396, 397 (diagnosis, Austria species key); Smith, F. 1858b: 114 (diagnosis); Mayr, 1861: 62 (Europe species key); Roger, 1863b: 28 (catalogue); Mayr, 1863: 429 (catalogue); Mayr, 1865: 19 (diagnosis); André, 1874: 192 (Europe species key); Forel, 1874: 75 (Switzerland species key); André, 1883a: 315 (Europe & Algeria species key); Cresson, 1887: 260 (U.S.A. catalogue); Provancher, 1887: 246 (Canada species key); Nasonov, 1889: 71 (Russia species key); Lameere, 1892: 68 (Belgium species key); Dalla Torre, 1893: 108 (catalogue); Forel, 1903a: 696 (India species key); Bingham, 1903: 267 (India, Sri Lanka & Burma species key); Ruzsky, 1905b: 652 (Russian Empire species key); Wasmann, 1906: 15 (Luxemburg species key); Emery, 1908a: 165 (Palaearctic species key); Bondroit, 1910: 493 (Belgium species key); Stitz, 1914: 70 (Central Europe species key); Donisthorpe, 1915d: 110 (Britain species key); Emery, 1916b: 121 (Italy species key); Wheeler, W.M. 1916m: 581 (U.S.A., Connecticut species key); Bondroit, 1918: 98 (France & Belgium species key); Emery, 1921f: 36 (diagnosis, catalogue); Emery, 1922e: 117 (Sifolinia diagnosis, catalogue); Finzi, 1926: 77 (diagnosis, European species key); Kuznetsov-Ugamsky, 1927e: 190 (Turkestan species key); Karavaiev, 1927c: 258 (Ukraine species key); Donisthorpe, 1927b: 118 (Britain species key); Arnol'di, 1933b: 599 (Russia species key); Arnol'di, 1934: 151 (former European U.S.S.R. species, biometrics); Karavaiev, 1934: 63 (Ukraine species key); Menozzi, 1939a: 293 (Himalaya & Tibet species key); Stitz, 1939: 74 (Germany species key); Kratochvíl, 1941: 75 (Central Europe species key); Novák & Sadil, 1941: 75 (Central Europe species key); Cole, 1942: 367 (U.S.A., Utah species key); Holgersen, 1943b: 168 (Norway species key); Holgersen, 1944: 199 (Norway species key); Buren, 1944a: 281 (U.S.A., Iowa species key); Boven, 1947: 175 (Belgium species key); Weber, 1947: 444 (North America species key); Creighton, 1950a: 92 (North America species key); Chapman & Capco, 1951: 124 (Asia checklist); Sadil, 1952: 264 (Czechia & Slovakia species key); Collingwood, 1958b: 65 (Britain species key); Boven, 1959: 7 (Netherlands species key); Gregg, 1963: 290 (U.S.A., Colorado species key); Wheeler, G.C. & Wheeler, J. 1963: 96 (U.S.A., North Dakota species key); Collingwood, 1964: 101 (Britain species key); Bernard, 1967: 107 (diagnosis, Western Europe species key); Bernard, 1967: 156 (Sifolinia diagnosis); Boven, 1970b: 16 (Netherlands species key); Arnol'di, 1970b: 1832 (former European U.S.S.R. species key); Kutter, 1973c: 253 (morphology, satellite genera, Sifolinia, Sommimyrma, Symbiomyrma, key to first); Bolton & Collingwood, 1975: 5 (Britain species key); Arnol'di, 1976a: 554 (central Asia & Kazakhstan species key); Tarbinsky, 1976: 20 (Kyrghyzstan species key); Boven, 1977: 94 (Belgium species key); Kutter, 1977c: 43 (Switzerland species key); Arnol'di & Dlussky, 1978: 530 (former European U.S.S.R. species key); Collingwood, 1978: 78 (Iberian Peninsula species key); Collingwood, 1979: 41 (Fennoscandia & Denmark species key); Smith, D.R. 1979: 1347, 1351 (North America Myrmica, Paramyrmica catalogues); Francoeur, 1981: 755 (M. lampra group); Allred, 1982: 440 (U.S.A., Utah species key); Gösswald, 1985: 294 (Germany species key); Wheeler, G.C. & Wheeler, J. 1986g: 22 (U.S.A., Nevada species key); Kupyanskaya, 1986a: 83 (Far Eastern Russia M. lobicornis group, key); Nilsson & Douwes, 1987: 62 (Norway species key); Agosti & Collingwood, 1987b: 267 (Balkans species key); Seifert, 1988b: 43 (Europe, Asia Minor & Caucasus species key); Bolton, 1988a: 3 (diagnosis, review of genus, venation); Dlussky, Soyunov & Zabelin, 1990: 181 (Turkmenistan species key); Kupyanskaya, 1990: 92 (Far Eastern Russia species key); Morisita, Kubota, et al. 1992: 8 (Japan species key); Atanasov & Dlussky, 1992: 80 (Bulgaria species key); Arakelian, 1994: 20 (Armenia species key); Radchenko, 1994b: 105 (South Siberia species key); Radchenko, 1994c: 130 (Central & Eastern Palaearctic species key); Douwes, 1995: 86 (Sweden species key); Bolton, 1994: 106 (synoptic classification); Kupyanskaya, 1995: 337 (Far Eastern Russia species key); Bolton, 1995a: 1051 (census); Bolton, 1995b: 277 (catalogue); Wu, J. & Wang, 1995: 91 (China species key); Seifert, 1996b: 140 (Central Europe species key); Skinner & Allen, 1996: 45 (Britain species key); Radchenko, Czechowski & Czechowsa, 1997: 481 (Poland species key); Collingwood & Prince, 1998: 12 (Portugal species key); Radchenko & Elmes, 1998: 3 (M. ritae group, key); Elmes & Radchenko, 1998: 218 (Taiwan species key); Wei, Zhou, He & Liu, 2001: 561 (China species key); Radchenko & Elmes, 2001: 262 (Himalaya species key); Czechowski, Radchenko & Czechowska, 2002: 141 (Poland species key); Radchenko & Elmes, 2003a: 239 (Palaearctic socially parasitic species key); Mackay & Mackay, 2002: 133 (U.S.A., New Mexico species key); Radchenko, 2005b: 192 (North Korea species key); Radchenko, Elmes & Bui, 2006: 43 (Vietnam species key); Radchenko, Elmes & Alicata, 2006: 531 (Palaearctic schencki group key); Seifert, 2007: 131 (North and Central Europe species key); Francoeur, 2007: 156 (punctiventris group key); Radchenko, Dlussky & Elmes, 2007: 1497 (late Eocene amber species key); Terayama, 2009: 160 (Taiwan species key); Radchenko & Elmes, 2009a: 85 (pachei group key); Radchenko & Elmes, 2010: 713 (Palaearctic, Oriental, all species revision, keys).

Myrmicinae Myrmicini Pogonomyrmex TRUE TRUE FALSE

Extant: 67 species, 9 subspecies

Fossil: 1 species

Pogonomyrmex Mayr, 1868b: 169. Type-species: Formica badia, by subsequent designation of Wheeler, W.M. 1911f: 170.

Taxonomic history

Pogonomyrmex in Myrmicidae, Myrmicidae: Emery, 1877a: 81.

Pogonomyrmex in Myrmicidae: Cresson, 1887: 260.

Pogonomyrmex in Myrmicinae: Dalla Torre, 1893: 118.

Pogonomyrmex in Myrmicinae, Myrmicini: Forel, 1895b: 125; Emery, 1895j: 769; Forel, 1899c: 61; Ashmead, 1905b: 383; Wheeler, W.M. 1910g: 140; Emery, 1914a: 40; Forel, 1917: 240; Emery, 1921f: 44; Wheeler, W.M. 1922a: 660; all subsequent authors.

Junior synonyms of POGONOMYRMEX

Ephebomyrmex Wheeler, W.M. 1902c: 390 [as subgenus of Pogonomyrmex]. Type-species: Pogonomyrmex naegelii, by subsequent designation of Wheeler, W.M. 1911f: 163.

Taxonomic history

Ephebomyrmex in Myrmicinae, Myrmicini: Ashmead, 1905b: 383; Forel, 1917: 240; Emery, 1921f: 48; Wheeler, W.M. 1922a: 660; all subsequent authors.

Ephebomyrmex as subgenus of Pogonomyrmex: Wheeler, W.M. 1902c: 390; Wheeler, W.M. 1910g: 140; Forel, 1917: 240; Emery, 1921f: 48; Wheeler, W.M. 1922a: 660; Gallardo, 1932b: 94; Donisthorpe, 1943f: 643; Kusnezov, 1949c: 293; Creighton, 1950a: 132; Kusnezov, 1951a: 245; Smith, M.R. 1951a: 794; Smith, M.R. 1958c: 115; Kusnezov, 1964: 56; Kusnezov, 1956: 118; Smith, M.R. 1967: 351; Cole, 1968: 22; Smith, D.R. 1979: 1357; Snelling, R.R. 1982a: 108; Mackay, Mackay, Dominguez, et al. 1985: 42.

Ephebomyrmex as genus: Creighton, 1957a: 54; Kusnezov, 1960b: 350; Kempf, 1972a: 106; Brown, 1973b: 180; Snelling, R.R. 1981: 395; Wheeler, G.C. & Wheeler, J. 1985: 257; Dlussky & Fedoseeva, 1988: 79; Mackay & Vinson, 1989b: 14; Hölldobler & Wilson, 1990: 13; Taber, 1998: 146.

Ephebomyrmex as junior synonym of Pogonomyrmex: Lattke, 1991a: 305; Bolton, 1994: 106; Bolton, 1995b: 28; Fernández & Palacio, 1998: 1649; Bolton, 2003: 223.

Forelomyrmex Wheeler, W.M. 1913a: 80 [as subgenus of Pogonomyrmex].

Taxonomic history

[Replacement name for Janetia Forel, 1899c: 61 (footnote); junior homonym of Janetia Kieffer, 1896: 236 (Diptera).]

Forelomyrmex as subgenus of Pogonomyrmex: Wheeler, W.M. 1913a: 80; Forel, 1917: 240; Emery, 1921f: 48; Wheeler, W.M. 1922a: 660; Donisthorpe, 1943f: 646; Kusnezov, 1951a: 245; Kempf, 1972a: 209.

Forelomyrmex as junior synonym of Pogonomyrmex: Brown, 1973b: 180 [provisional]; Snelling, R.R. 1981: 395; Lattke, 1991a: 305; Bolton, 1994: 106; Bolton, 1995b: 29; Fernández & Palacio, 1998: 1649.

Homonym replaced by Forelomyrmex

Janetia Forel, 1899c: 61 (footnote) [as subgenus of Pogonomyrmex]. Type-species: Pogonomyrmex (Janetia) mayri, by monotypy.

Taxonomic history

[Junior homonym of Janetia Kieffer, 1896: 236 (Diptera).]

Genus Pogonomyrmex references

Mayr, 1870b: 970 (all species key); Mayr, 1887: 608 (all species key); Cresson, 1887: 260 (U.S.A. catalogue); Dalla Torre, 1893: 118 (catalogue); Wheeler, W.M. 1902a: 97 (North America species key); Emery, 1921f: 44 (diagnosis, subgenera key, catalogue); Emery, 1921f: 48 (P. (Ephebomyrmex) & P. (Forelomyrmex) diagnoses, catalogues); Gallardo, 1932b: 95 (Argentina species key); Cole, 1942: 364 (U.S.A., Utah species key); Kusnezov, 1949c: 301 (Argentina P. (Ephebomyrmex) species key); Creighton, 1950a: 113 (North America species key); Kusnezov, 1951a: 254 (Argentina species key); Kusnezov, 1954c: 85 (Bolivia species key); Creighton, 1957a: 54 (North America Ephebomyrmex); Gregg, 1963: 317 (U.S.A., Colorado species key); Cole, 1968: 21, 38, 155 (North America Pogonomyrmex & P. (Ephebomyrmex) diagnoses, species revision, keys); Kempf, 1972a: 106, 206 (Neotropical Ephebomyrmex, Pogonomyrmex catalogues); Snelling, R.R. & Hunt, 1976: 72 (Chile species key); Smith, D.R. 1979: 1353 (North America catalogue); Allred, 1982: 442 (U.S.A., Utah species key); Snelling, R.R. 1982a: 104 (supplement to Cole, 1968 key); Snelling, R.R. 1982a: 108 (North America P. (Ephebomyrmex) species key); Mackay, Mackay, Dominguez, et al. 1985: 42 (Mexico species key); Wheeler, G.C. & Wheeler, J. 1986g: 26 (U.S.A., Nevada species key); Wheeler, G.C. & Wheeler, J. 1986g: 33 (U.S.A., Nevada Ephebomyrmex species key); Shattuck, 1987: 175 (P. occidentalis complex, key); Taber, 1990: 307 (U.S.A. species complexes, phylogeny); Brandão, 1991: 342, 372 (Neotropical catalogue); Bolton, 1994: 106 (synoptic classification); Bolton, 1995a: 1052 (census); Bolton, 1995b: 339 (catalogue); Taber, 1998: 146, 147 (Ephebomyrmex, Pogonomyrmex species key); Fernández & Palacio, 1998: 1657 (northern South America species key); Mackay & Mackay, 2002: 180 (U.S.A., New Mexico species key); Lattke, 2006: 56 (northern South America species key); Johnson, R.A. & Overson, 2009: 313 (P. californicus group key); Cuezzo & Claver, 2009: 99 (Argentina species key).

Myrmicinae Tetramoriini Strongylognathus TRUE TRUE FALSE

24 species

Strongylognathus Mayr, 1853d: 389. Type-species: Eciton testaceum, by monotypy.

Taxonomic history

[Replacement name for Myrmus Schenck, 1853: 188; junior homonym of Myrmus Hahn, 1832: 81 (Hemiptera).]

Strongylognathus in Poneridae, Myrmicidae: Smith, F. 1858b: 134.

Strongylognathus in Myrmicidae, Myrmicidae: Emery, 1877a: 81.

Strongylognathus in Myrmicinae: Mayr, 1855: 430 [Myrmicidae]; Mayr, 1865: 21 [Myrmicidae]; Emery & Forel, 1879a: 457 [Myrmicidae]; Dalla Torre, 1893: 129.

Strongylognathus in Myrmicinae, Tetramoriini: Emery, 1895j: 770; Ashmead, 1905b: 383; Wheeler, W.M. 1910g: 141; Emery, 1914a: 42; Forel, 1917: 245; Wheeler, W.M. 1922a: 664; Emery, 1924d: 285; all subsequent authors.

Genus Strongylognathus references

Smith, F. 1858b: 134 (diagnosis); Mayr, 1863: 454 (catalogue); Mayr, 1865: 21 (diagnosis); Forel, 1874: 71 (Switzerland species key); André, 1883a: 281 (Europe & Algeria species key); Dalla Torre, 1893: 129 (catalogue); Forel, 1900e: 278 (all species key); Ruzsky, 1905b: 541 (Russian Empire species key); Emery, 1909d: 708 (Palaearctic species key); Emery, 1916b: 199 (Italy species key); Bondroit, 1918: 110 (France & Belgium species key); Emery, 1924d: 285 (diagnosis catalogue); Arnol'di, 1933b: 598 (Russia species key); Kratochvíl, 1941: 87 (Central Europe species key); Novák & Sadil, 1941: 87 (Central Europe species key); Bernard, 1967: 237 (Western Europe species key); Baroni Urbani, 1969c: 157 (Western Europe S. huberi group revision, key); Bolton, 1976: 304 (diagnosis, review of genus); Kutter, 1977c: 159 (Switzerland species key); Arnol'di & Dlussky, 1978: 545 (former European U.S.S.R. species key); Collingwood, 1978: 86 (Iberian Peninsula species key); Gösswald, 1985: 312 (Germany species key); Agosti & Collingwood, 1987b: 278 (Balkans species key); Dlussky, Soyunov & Zabelin, 1990: 209 (Turkmenistan species key); Radchenko, 1991: 89 (former U.S.S.R. species key); Atanasov & Dlussky, 1992: 156 (Bulgaria species key); Arakelian, 1994: 68 (Armenia species key); Bolton, 1994: 106 (synoptic classification); Bolton, 1995a: 1053 (census); Bolton, 1995b: 395 (catalogue); Wu, J. & Wang, 1995: 95 (China species key); Seifert, 1996b: 115 (Central Europe species key); Sanetra & Buschinger, 2000: 95 (phylogeny); Wei, Xu & He, 2001: 68 (China species key); Seifert, 2007: 115 (North and Central Europe species key).

Homonym replaced by STRONGYLOGNATHUS

Myrmus Schenck, 1853: 188. Type-species: Myrmus emarginatus (junior synonym of Eciton testaceum), by monotypy.

Taxonomic history

[Junior homonym of Myrmus Hahn, 1832: 81 (Hemiptera).]

Myrmicinae Tetramoriini Tetramorium TRUE TRUE FALSE

Extant: 461 species, 16 subspecies

Fossil:

Tetramorium Mayr, 1855: 423. Type-species: Formica caespitum, by subsequent designation of Girard, 1879: 1016.

Taxonomic history

Tetramorium in Myrmicidae, Myrmicidae: Emery, 1877a: 81.

Tetramorium in Myrmicinae: Mayr, 1855: 423 [Myrmicidae]; Mayr, 1861: 61 [Myrmicidae]; Mayr, 1865: 20 [Myrmicidae]; Emery & Forel, 1879a: 457 [Myrmicidae]; Dalla Torre, 1893: 130.

Tetramorium in Myrmicinae, Myrmicini: Forel, 1895b: 125; Forel, 1899c: 52; Kusnezov, 1964: 57 (anachronism).

Tetramorium in Myrmicinae, Tetramoriini: Emery, 1895j: 770; Ashmead, 1905b: 383; Wheeler, W.M. 1910g: 141; Emery, 1914a: 42; Arnold, 1917: 271; Forel, 1917: 245; Wheeler, W.M. 1922a: 664; Emery, 1924d: 275; all subsequent authors.

Junior synonyms of TETRAMORIUM

Tetrogmus Roger, 1857: 10. Type-species: Tetrogmus caldarius, by monotypy.

Taxonomic history

Tetrogmus as subgenus of Tetramorium: Wheeler, W.M. 1910g: 141.

Tetrogmus as junior synonym of Tetramorium: Roger, 1862c: 297; all subsequent authors except the above; Bolton, 1976: 359; Bolton, 1980: 195.

Xiphomyrmex Forel, 1887: 385 [as subgenus of Tetramorium]. Type-species: Tetramorium (Xiphomyrmex) kelleri, by subsequent designation of Wheeler, W.M. 1911f: 175.

Taxonomic history

Xiphomyrmex in Myrmicinae: Dalla Torre, 1893: 130.

Xiphomyrmex in Myrmicinae, Myrmicini: Ashmead, 1905b: 383.

Xiphomyrmex in Myrmicinae, Tetramoriini: Emery, 1895j: 770; Emery, 1914a: 42; Forel, 1917: 245; Wheeler, W.M. 1922a: 664; Emery, 1924d: 286; all subsequent authors.

Xiphomyrmex as subgenus of Tetramorium: Dalla Torre, 1893: 130; Forel, 1899c: 53; Forel, 1903a: 700; Wheeler, W.M. 1910g: 141; Arnold, 1917: 274; Arnold, 1926: 277.

Xiphomyrmex as genus: Emery, 1895j: 770; Emery, 1896e: 183; Emery, 1914a: 42; Forel, 1917: 245; Wheeler, W.M. 1922a: 193; Emery, 1924d: 286; Smith, M.R. 1938a: 126; Donisthorpe, 1943g: 736; Creighton, 1950a: 293; Smith, M.R. 1951a: 824; Chapman & Capco, 1951: 179; Bernard, 1953b: 250; Wheeler, G.C. & Wheeler, J. 1985: 257 (anachronism); Dlussky & Fedoseeva, 1988: 80 (anachronism).

Xiphomyrmex as junior synonym of Tetramorium: Bingham, 1903: 175; Bolton, 1976: 359; Bolton, 1980: 195; Bolton, 1994: 106.

Triglyphothrix Forel, 1890b: cvi. Type-species: Triglyphothrix walshi, by monotypy.

Taxonomic history

Triglyphothrix in Myrmicinae: Dalla Torre, 1893: 135.

Triglyphothrix in Myrmicinae, Tetramoriini: Emery, 1895j: 770; Ashmead, 1905b: 383; Wheeler, W.M. 1910g: 141; Emery, 1914a: 42; Forel, 1917: 245; Wheeler, W.M. 1922a: 664; Emery, 1924d: 272; all subsequent authors.

Triglyphothrix as subgenus of Tetramorium: Arnold, 1917: 274; Arnold, 1926: 272.

Triglyphothrix as genus: Forel, 1890b: cvi. Dalla Torre, 1893: 135; Emery, 1895j: 770; Forel, 1903a: 703; Bingham, 1903: 171; Wheeler, W.M. 1910g: 141; Emery, 1914a: 42; Forel, 1917: 245; Wheeler, W.M. 1922a: 196; Emery, 1924d: 272; Donisthorpe, 1943g: 734; Creighton, 1950a: 285; Smith, M.R. 1951a: 823; Chapman & Capco, 1951: 178; Bernard, 1953b: 248; Kempf, 1972a: 255; Bolton, 1976: 310; Smith, D.R. 1979: 1399; Wheeler, G.C. & Wheeler, J. 1985: 257; Taylor & Brown, D.R. 1985: 91.

Triglyphothrix as junior synonym of Tetramorium: Bolton, 1985: 247; Bolton, 1994: 106.

Atopula Emery, 1912b: 104. Type-species: Atopomyrmex nodifer, by original designation.

Taxonomic history

Atopula in Myrmicinae, Myrmecinini: Emery, 1912b: 105; Emery, 1914a: 41; Forel, 1917: 244; Wheeler, W.M. 1922a: 663; Emery, 1924d: 242 [subtribe Podomyrmini]; all subsequent authors to Bolton, 1976: 359.

Atopula in Myrmicinae, Podomyrmini: Dlussky & Fedoseeva, 1988: 79.

Atopula as subgenus of Vollenhovia: Emery, 1912d: 273.

Atopula as subgenus of Terataner: Arnold, 1952b: 129.

Atopula as genus: Emery, 1912b: 104; Emery, 1914a: 41; Wheeler, W.M. 1922a: 663; Emery, 1924d: 242; Bernard, 1948: 177; Wheeler, G.C. & Wheeler, J. 1985: 257 (anachronism); Dlussky & Fedoseeva, 1988: 79 (anachronism).

Atopula as junior synonym of Tetramorium: Bolton, 1976: 359; Bolton, 1980: 195; Bolton, 1994: 106.

Macromischoides Wheeler, W.M. 1920: 53. Type-species: Macromischa aculeata, by original designation.

Taxonomic history

Macromischoides in Myrmicinae, Leptothoracini: Wheeler, W.M. 1922a: 664; Wheeler, G.C. & Wheeler, J. 1985: 257 (anachronism).

Macromischoides in Myrmicinae, Tetramoriini: Santschi, 1924b: 207; all subsequent authors.

Macromischoides as genus: Wheeler, W.M. 1922a: 187; Santschi, 1924b: 206; Bernard, 1953b: 248; Dlussky & Fedoseeva, 1988: 80 (anachronism).

Macromischoides as junior synonym of Tetramorium: Bolton, 1976: 359; Bolton, 1980: 196.

[Macromichoides Santschi, 1924b: 206, incorrect subsequent spelling.]

Lobomyrmex Kratochvíl, 1941: 84 [as subgenus of Tetramorium]. Type-species: Tetramorium (Lobomyrmex) ferox silhavyi (junior synonym of Tetramorium ferox), by monotypy.

Taxonomic history

Lobomyrmex as junior synonym of Tetramorium: Brown, 1973b: 181 [provisional]; Bolton, 1976: 359; Bolton, 1980: 196.

Sulcomyrmex Kratochvíl, 1941: 84 [as subgenus of Tetramorium]. Unavailable name. Proposed without designation of type-species and therefore unavailable. Species included by Kratochvíl (1941) are all referable to Tetramorium: Bolton, 1976: 359.]

Apomyrmex Calilung, 2000: 66. Type-species: Apomyrmex manobo, by original designation.

Taxonomic history

Apomyrmex incertae sedis in Myrmicinae: Calilung, 2000: 66.

Apomyrmex as junior synonym of Tetramorium: Bolton, 2003: 227, 269.

Genus Tetramorium references

Roger, 1863b: 26, 27 (catalogue); Mayr, 1863: 456 (catalogue); Mayr, 1865: 20 (diagnosis); Mayr, 1870b: 972 (all species key); André, 1883a: 285 (Europe & Algeria species key); Cresson, 1887: 261 (U.S.A. catalogue); Emery, 1893e: 214 (Triglyphothrix species key); Dalla Torre, 1893: 130, 135 (Tetramorium, Triglyphothrix catalogues); Forel, 1903a: 700, 703 (India & Sri Lanka Tetramorium, Triglyphothrix species keys); Bingham, 1903: 172, 175 (India, Sri Lanka & Burma Triglyphothrix, Tetramorium species keys); Ruzsky, 1905b: 517 (Russian Empire species key); Emery, 1909d: 695 (Palaearctic species key); Emery, 1916b: 195 (Italy species key); Arnold, 1917: 271, 275, 334, 345 (diagnosis, South Africa Tetramorium (Tetramorium), T. (Triglyphothrix), T. (Xiphomyrmex) species keys); Bondroit, 1918: 106 (France & Belgium species key); Mann, 1919: 353 (Papuasia Triglyphothrix species key); Wheeler, W.M. 1922a: 187, 190, 193, 196 (Macromischoides, Tetramorium, Xiphomyrmex, Triglyphothrix diagnoses); Wheeler, W.M. 1922a: 886, 889, 893, 906, 909 (Afrotropical Atopula, Macromischoides, Tetramorium, Xiphomyrmex, Triglyphothrix catalogues); Wheeler, W.M. 1922a: 1030, 1031, 1032 (Malagasy Tetramorium, Xiphomyrmex, Triglyphothrix catalogues); Emery, 1924d: 242 (Atopula diagnosis, catalogue); Emery, 1924d: 272 (Triglyphothrix diagnosis, catalogue); Emery, 1924d: 275 (Tetramorium diagnosis, catalogue); Emery, 1924d: 286 (Xiphomyrmex diagnosis, catalogue); Santschi, 1924b: 210 (Macromischoides species key); Arnold, 1926: 241 (South Africa Tetramorium (Tetramorium) species key); Menozzi, 1933b: 74 (Israel species key); Finzi, 1936: 183 (Egypt species key); Smith, M.R. 1938a: 127 (North America Xiphomyrmex species key); Kratochvíl, 1941: 84 (Central Europe species key); Novák & Sadil, 1941: 84 (Central Europe species key); Smith, M.R. 1943b: 2 (North America species key); Kratochvíl, Novák & Snoflák, 1944: 76, 95 (Central Europe species key); Creighton, 1950a: 290, 293 (North America Tetramorium, Xiphomyrmex species key); Chapman & Capco, 1951: 174, 178, 179 (Asia Tetramorium, Triglyphothrix, Xiphomyrmex checklists); Brown, 1958h: 27 (New Zealand species); Bernard, 1967: 227 (diagnosis, Western Europe species key); Kempf, 1972a: 249, 255 (Neotropical Tetramorium, Triglyphothrix catalogues); Alayo, 1974: 18 (Cuba species key); Tarbinsky, 1976: 105 (Kyrghyzstan species key); Bolton, 1976: 310, 314, 341 (Triglyphothrix diagnosis, Afrotropical, Oriental & Malesian species revisions, keys); Bolton, 1976: 359 (Tetramorium diagnosis, review of genus, synonymy); Bolton, 1977: 72, 133 (Oriental & Malesian, Australia species revisions, keys); Kutter, 1977c: 150 (Switzerland species key); Arnol'di & Dlussky, 1978: 544 (former European U.S.S.R. species key); Collingwood, 1978: 86 (Iberian Peninsula species key); Collingwood, 1979: 82 (Fennoscandia & Denmark species key); Smith, D.R. 1979: 1399, 1400 (North America Triglyphothrix, Tetramorium catalogues); Bolton, 1979: 132, 159 (Malagasy, New World species revisions, keys); Bolton, 1980: 196, 205 (diagnosis, review of genus, Afrotropical species revision, key); Collingwood, 1985: 262 (Saudi Arabia species key); Gösswald, 1985: 311 (Germany species key); Taylor & Brown, D.R. 1985: 88 (Australia catalogue); Taylor, 1987a: 78 (Australia, New Caledonia & New Zealand checklist); Agosti & Collingwood, 1987b: 277 (Balkans species key); Wang, M., Xiao & Wu, 1988: 264 (China species key); Radchenko & Arakelian, 1990: 371 (Crimea & Caucasus T. ferox complex, key); Dlussky, Soyunov & Zabelin, 1990: 197 (Turkmenistan species key); Atanasov & Dlussky, 1992: 149 (Bulgaria species key); Radchenko, 1992a: 44 (former U.S.S.R. species key); Morisita, Kubota, et al. 1992: 33 (Japan species key); Arakelian, 1994: 60 (Armenia species key); Radchenko, 1994b: 109 (South Siberia species key); Bolton, 1994: 106 (synoptic classification); Bolton, 1995a: 1053 (census); Bolton, 1995b: 403 (catalogue); Wu, J. & Wang, 1995: 79 (China species key); Collingwood & Agosti, 1996: 332 (Saudi Arabia species key); Seifert, 1996b: 158 (Central Europe species key); Cagniant, 1997: 89 (Morocco species key); Radchenko, Czechowski & Czechowsa, 1998: 107 (Poland species key); Collingwood & Prince, 1998: 17 (Portugal species key); Sanetra, Güsten & Schulz, 1999: 317 (Italy, species); Shattuck, 1999: 168 (Australia synopsis); Chang & He, 2001a: 1 (northwest China species key); Zhou, 2001b: 100 (China, Guangxi species key); Czechowski, Radchenko & Czechowska, 2002: 145 (Poland species key); Mackay & Mackay, 2002: 214 (U.S.A., New Mexico species key); Seifert, 2007: 145 (North and Central Europe species key); Csösz, Radchenko & Schulz, 2007: 8 (T. chefketi complex key); Terayama, 2009: 162 (Taiwan species key); Heterick, 2009: 177 (south-western Australia species key); Csösz & Schulz, 2010: 12 (T. ferox complex key); Hita Garcia, Fischer & Peters, 2010b: 12 (Afrotropical T. weitzeckeri species group key).

Myrmicinae Pheidolini Aphaenogaster TRUE TRUE FALSE

Extant: 176 species, 39 subspecies

Fossil: 19 species

Aphaenogaster Mayr, 1853b: 107. Type-species: Aphaenogaster sardoa, by subsequent designation of Bingham, 1903: 270.

Taxonomic history

Aphaenogaster in Myrmicinae: Mayr, 1855: 466 [Myrmicidae]; Mayr, 1865: 19 [Myrmicidae]; Emery & Forel, 1879a: 461 [Myrmicidae]; Cresson, 1887: 260 [Myrmicidae]; Dalla Torre, 1893: 98.

Aphaenogaster in Myrmicinae, Myrmicini: Forel, 1891b: 166; Forel, 1895b: 129; Emery, 1895j: 769; Forel, 1899c: 58; Wheeler, W.M. 1910g: 140; Wheeler, W.M. 1915h: 53; Santschi, 1932b: 13; Kempf, 1972a: 22 (anachronism).

Aphaenogaster in Myrmicinae, Aphaenogastrini: Enzmann, J. 1947b: 147.

Aphaenogaster in Myrmicinae, Pheidolini: Emery, 1877a: 81 [Pheidolidae]; Emery, 1914a: 40; Forel, 1917: 241; Emery, 1921f: 55 [subtribe Stenammini]; Wheeler, W.M. 1922a: 661; Brown, 1949a: 48; Chapman & Capco, 1951: 131; all subsequent authors.

Aphaenogaster as junior synonym of Atta: Mayr, 1863: 395.

Aphaenogaster as subgenus of Stenamma: Emery, 1895c: 298; Emery, 1895j: 769; Forel, 1903a: 693.

Aphaenogaster as genus: Emery, 1908c: 309; all subsequent authors.

Junior synonyms of APHAENOGASTER

Deromyrma Forel, 1913b: 350 [as subgenus of Aphaenogaster]. Type-species: Aphaenogaster (Ischnomyrmex) swammerdami, by original designation.

Taxonomic history

[Deromyrma also described as new by Forel, 1913k: 49.]

Deromyrma as subgenus of Aphaenogaster: Forel, 1913b: 350; Forel, 1913k: 49; Forel, 1917: 241; Emery, 1921f: 64; Wheeler, W.M. 1922a: 680; all subsequent authors to the following.

Deromyrma as junior synonym of Aphaenogaster: Brown, 1973b: 180 [provisional]; Smith, D.R. 1979: 1359; Bolton, 1982: 364.

Planimyrma Viehmeyer, 1914d: 604 [as subgenus of Aphaenogaster]. Type-species: Stenamma (Ischnomyrmex) loriai, by original designation.

Taxonomic history

Planimyrma as subgenus of Aphaenogaster: Viehmeyer, 1914d: 604; Forel, 1917: 241; Emery, 1921f: 65; Wheeler, W.M. 1922a: 680; all subsequent authors to the following.

Planimyrma as junior synonym of Aphaenogaster: Brown, 1973b: 184 [provisional]; Smith, D.R. 1979: 1359; Bolton, 1982: 364.

Attomyrma Emery, 1915d: 70 [as subgenus of Aphaenogaster]. Type-species: Formica subterranea, by original designation.

Taxonomic history

Attomyrma as subgenus of Aphaenogaster: Emery, 1915d: 70; Emery, 1916b: 129; Emery, 1921f: 56; all subsequent authors to the following.

Attomyrma as junior synonym of Aphaenogaster: Brown, 1973b: 178 [provisional]; Smith, D.R. 1979: 1359; Bolton, 1982: 364.

Novomessor Emery, 1915d: 73. Type-species: Aphaenogaster (Ischnomyrmex) cockerelli, by original designation.

Taxonomic history

Novomessor in Myrmicinae, Pheidolini: Forel, 1917: 241; Emery, 1921f: 66 [subtribe Stenammini]; Wheeler, W.M. 1922a: 661; all subsequent authors except the following.

Novomessor in Myrmicinae, Myrmicini: Donisthorpe, 1943f: 675; Kempf, 1972a: 166.

Novomessor as genus: Wheeler, W.M. & Creighton, 1934: 343; Creighton, 1950a: 155; Hölldobler, Stanton & Engel, 1976: 32.

Novomessor as junior synonym of Aphaenogaster: Brown, 1974b: 47; Bolton, 1982: 364 (discussion pp. 339-341).

Nystalomyrma Wheeler, W.M. 1916j: 215 [as subgenus of Aphaenogaster]. Type-species: Myrmica longiceps, by original designation.

Taxonomic history

Nystalomyrma as subgenus of Aphaenogaster: Wheeler, W.M. 1916j: 215; Emery, 1921f: 61; Wheeler, W.M. 1922a: 680; Donisthorpe, 1943f: 676.

Nystalomyrma as junior synonym of Aphaenogaster: Brown, 1973b: 183 [provisional]; Smith, D.R. 1979: 1359; Bolton, 1982: 364.

Brunella Forel, 1917: 234. Type-species: Aphaenogaster belti, by monotypy.

Taxonomic history

Brunella in Myrmicinae, Leptothoracini: Forel, 1917: 245.

Brunella in Myrmicinae, Myrmecinini: Wheeler, W.M. 1922a: 663; Emery, 1924d: 242; Donisthorpe, 1943f: 629.

Brunella incertae sedis in Formicidae: Wheeler, G.C. & Wheeler, J. 1985: 259 (anachronism, incomprehensible entry).

Brunella incertae sedis in Myrmicinae: Dlussky & Fedoseeva, 1988: 80 (anachronism).

Brunella as junior synonym of Atopula: Emery, 1924d: 242; Donisthorpe, 1943f: 629.

Brunella as junior synonym of Aphaenogaster: Bolton, 1982: 364; Bolton, 1994: 106.

†Sinaphaenogaster Zhang, 1989: 266 [as subgenus of Aphaenogaster]. Type-species: †Paraphaenogaster shanwangensis, by original designation.

Taxonomic history

†Sinaphaenogaster as junior synonym of Aphaenogaster: Bolton, 2003: 230, 273.

Genus Aphaenogaster references

Mayr, 1855: 466 (diagnosis); Roger, 1863b: 29 (catalogue); Mayr, 1865: 19 (diagnosis); Mayr, 1867a: 92 (diagnosis); André, 1874: 195 (Europe species key); Forel, 1874: 74 (Switzerland species key); André, 1883b: 348 (Europe & Algeria species key); Mayr, 1886d: 443 (U.S.A. species key); Cresson, 1887: 260 (U.S.A. catalogue); Emery, 1888a: 531 (A. (Ischnomyrmex) species key); Nasonov, 1889: 74 (Russia species key); Dalla Torre, 1893: 98 (catalogue); Forel, 1903a: 693 (India & Sri Lanka species key); Bingham, 1903: 270 (India, Sri Lanka & Burma species key); Ruzsky, 1905b: 714 (Russian Empire species key); Emery, 1908c: 310 (Palaearctic species key); Bondroit, 1910: 495 (Belgium species key); Emery, 1916b: 132 (Italy species key); Wheeler, W.M. 1916m: 585 (U.S.A., Connecticut species key); Wheeler, W.M. 1916j: 213 (Australia species); Bondroit, 1918: 156 (France & Belgium species key); Emery, 1921f: 55 (diagnosis, subgenera key, catalogue); Emery, 1921f: 56, 61, 64, 65 (A. (Attomyrma, Nystalomyrma, Deromyrma, Planimyrma) diagnoses, catalogues); Emery, 1921f: 66 (Novomessor diagnosis, catalogue); Wheeler, W.M. 1922a: 680 (subgenera key); Wheeler, W.M. 1922a: 1016, 1029 (Aphaenogaster, Brunella catalogues); Wheeler, W.M. & Creighton, 1934: 348 (Novomessor species key); Menozzi, 1939a: 296 (Himalaya & Tibet species key); Stitz, 1939: 112 (Germany species key); Kratochvíl, 1941: 81 (Central Europe species key); Novák & Sadil, 1941: 81 (Central Europe species key); Cole, 1942: 363 (U.S.A., Utah species key); Buren, 1944a: 284 (U.S.A., Iowa species key); Creighton, 1950a: 140, 155 (North America Aphaenogaster, Novomessor species keys); Chapman & Capco, 1951: 131 (Asia checklist); Smith, M.R. 1961b: 218 (New Guinea species key); Gregg, 1963: 336 (U.S.A., Colorado species key); Bernard, 1967: 128 (diagnosis, Western Europe species key); Kempf, 1972a: 22, 166 (Neotropical Aphaenogaster (Deromyrma), Novomessor catalogues); Arnol'di, 1976b: 1023 (former U.S.S.R. species key); Kutter, 1977c: 77 (Switzerland species key); Arnol'di & Dlussky, 1978: 536 (former European U.S.S.R. species key); Collingwood, 1978: 79 (Iberian Peninsula species key); Smith, D.R. 1979: 1359 (North America catalogue); Allred, 1982: 439 (U.S.A., Utah species key); Taylor & Brown, D.R. 1985: 54 (Australia catalogue); Gösswald, 1985: 298 (Germany species key); Wheeler, G.C. & Wheeler, J. 1986g: 35 (U.S.A., Nevada species key); Taylor, 1987a: 8 (Australia checklist); Agosti & Collingwood, 1987b: 269 (Balkans species key); Dlussky & Fedoseeva, 1988: 79 (synoptic classification); Dlussky, Soyunov & Zabelin, 1990: 211 (Turkmenistan species key); Brandão, 1991: 326 (Neotropical catalogue); Morisita, Kubota, et al. 1992: 15 (Japan species key); Atanasov & Dlussky, 1992: 107 (Bulgaria species key); Schulz, A. 1994a: 425 (A. (Attomyrma), partial key); Arakelian, 1994: 29 (Armenia species key); Bolton, 1994: 106 (synoptic classification); Bolton, 1995a: 1048 (census); Bolton, 1995b: 68 (catalogue); Wu, J. & Wang, 1995: 110 (China species key); Umphrey, 1996: 557 (North America A. fulva complex key); Cagniant, 1996a: 67 (Morocco species key); Seifert, 1996b: 156 (Central Europe species key); Collingwood & Prince, 1998: 13 (Portugal species key); Shattuck, 1999: 126 (Australia synopsis); Park & Kim, 2000: 108 (Korea species key); Zhou, 2001b: 142 (China, Guangxi species key); Mackay & Mackay, 2002: 60 (U.S.A., New Mexico species key); Longino & Cover, 2004: 1 (A. phalangium complex revision); Radchenko, 2005b: 192 (North Korea species key); Seifert, 2007: 144 (North and Central Europe species key); Shattuck, 2008a: 27 (Australia species revision, key); Terayama, 2009: 164 (Taiwan species key); Heterick, 2009: 144 (south-western Australia species key).

Myrmicinae Pheidolini Messor TRUE TRUE FALSE

Extant: 112 species, 49 subspecies

Fossil:

Messor Forel, 1890a: lxviii [as subgenus of Aphaenogaster]. Type-species: Formica barbara, by subsequent designation of Bingham, 1903: 277.

Taxonomic history

Messor in Myrmicinae: Dalla Torre, 1893: 98.

Messor in Myrmicinae, Myrmicini: Emery, 1895j: 769; Forel, 1899c: 59; Ashmead, 1905b: 383; Wheeler, W.M. 1910g: 140.

Messor in Myrmicinae, Pheidolini: Emery, 1914a: 40; Forel, 1917: 241; Arnold, 1920a: 404; Emery, 1921f: 68 [subtribe Stenammini]; Wheeler, W.M. 1922a: 661; all subsequent authors.

Messor as subgenus of Aphaenogaster: Forel, 1890a: lxviii; Dalla Torre, 1893: 98; Forel, 1899c: 59.

Messor as subgenus of Stenamma: Emery, 1895c: 298; Emery, 1895j: 769; Forel, 1903a: 693.

Messor as genus: Bingham, 1903: 277; Ruzsky, 1905b: 726; Emery, 1908e: 437; Wheeler, W.M. 1910g: 140; Forel, 1917: 241; Bondroit, 1918: 149; Arnold, 1920a: 404; Emery, 1921f: 68; Wheeler, W.M. 1922a: 661; all subsequent authors.

Junior synonyms of MESSOR

Cratomyrmex Emery, 1892d: 572. Type-species: Cratomyrmex regalis, by monotypy.

Taxonomic history

Cratomyrmex in Myrmicinae, Myrmicini: Emery, 1895j: 769; Ashmead, 1905b: 383; Wheeler, W.M. 1910g: 140; Emery, 1914a: 40; Forel, 1917: 240; Emery, 1921f: 43; Wheeler, W.M. 1922a: 660; Donisthorpe, 1943f: 635.

Cratomyrmex as subgenus of Messor: Santschi, 1920d: 378.

Cratomyrmex as genus: Emery, 1892d: 572; Wheeler, W.M. 1910g: 140; Forel, 1917: 240; Emery, 1921f: 43; Wheeler, W.M. 1922a: 660, 802; Bernard, 1971: 6.

Cratomyrmex as junior synonym of Messor: Emery, 1924d: 357; Donisthorpe, 1943f: 635; Bolton, 1982: 338; Hölldobler & Wilson, 1990: 14; Bolton, 1994: 106.

Veromessor Forel, 1917: 235 [as subgenus of Novomessor]. Type-species: Aphaenogaster andrei, by subsequent designation of Emery, 1921f: 67.

Taxonomic history

Veromessor in Myrmicinae, Pheidolini: Wheeler, W.M. 1922a: 661; all subsequent authors.

Veromessor as subgenus of Novomessor: Forel, 1917: 235; Emery, 1921f: 67; Donisthorpe, 1943g: 735.

Veromessor as genus: Wheeler, W.M. 1922a: 661; Wheeler, W.M. & Creighton, 1934: 354; Creighton, 1950a: 157; Smith, M.R. 1951a: 799; Kusnezov, 1952b: 10 (in key); Kempf, 1972a: 257; Smith, D.R. 1979: 1364.

Veromessor as junior synonym of Messor: Bolton, 1982: 338.

Lobognathus Enzmann, J. 1947b: 152 [as subgenus of Veromessor]. Erroneous entry for Veromessor lobognathus and hence junior synonym of Veromessor: Brown, 1949a: 49.

[Sphaeromessor Bernard, 1985: 48. Unavailable name. Proposed without designation of type-species and therefore unavailable. Species included by Bernard (1985) are all referable to Messor: Bolton, 1995b: 46.]

Genus Messor references

Forel, 1903a: 693 (India & Sri Lanka species key); Bingham, 1903: 278 (India, Sri Lanka & Burma species key); Ruzsky, 1905b: 726 (Russian Empire species key); Emery, 1908e: 437 (Palaearctic species key); Emery, 1916b: 143 (Italy species key); Bondroit, 1918: 149 (France & Belgium species key); Arnold, 1920a: 404 (diagnosis); Emery, 1921f: 43 (Cratomyrmex diagnosis, catalogue); Emery, 1921f: 67 (Veromessor diagnosis, catalogue); Emery, 1921f: 68 (diagnosis, catalogue); Wheeler, W.M. 1922a: 802 (Afrotropical Cratomyrmex, Messor catalogues); Santschi, 1927c: 247 (M. instabilis group key); Kuznetsov-Ugamsky, 1927a: 89 (Central Asia species key); Kuznetsov-Ugamsky, 1929d: 4 (Central Asia species key); Finzi, 1929: 77 (Italy species key); Menozzi, 1933b: 56 (Israel species key); Wheeler, W.M. & Creighton, 1934: 360 (Veromessor species key); Finzi, 1936: 160 (Egypt species key); Novák & Sadil, 1941: 82 (Central Europe species key); Creighton, 1950a: 158 (North America Veromesor species key); Chapman & Capco, 1951: 136 (Asia checklist); Bernard, 1955a: 360 (Mediterranean M. structor group key); Smith, M.R. 1956a: 36 (U.S.A. Veromessor species key); Bernard, 1967: 137 (diagnosis, Western Europe species key); Kempf, 1972a: 257 (Neotropical Veromessor catalogue); Tarbinsky, 1976: 50 (Kyrghyzstan species key); Arnol'di, 1977b: 1640 (former U.S.S.R. species key); Arnol'di & Dlussky, 1978: 537 (former European U.S.S.R. species key); Collingwood, 1978: 80 (Iberian Peninsula species key); Smith, D.R. 1979: 1364 (North America Veromessor catalogue); Bolton, 1982: 338 (diagnosis, review of genus, Afrotropical species revision, key); Collingwood, 1985: 248 (Saudi Arabia species key); Wheeler, G.C. & Wheeler, J. 1986g: 38 (U.S.A., Nevada Veromessor species key); Agosti & Collingwood, 1987b: 270 (Balkans species key); Dlussky, Soyunov & Zabelin, 1990: 215 (Turkmenistan species key); Casevitz-Weulersse, 1990b: 156 (Corsica species key); Atanasov & Dlussky, 1992: 113 (Bulgaria species key); Arakelian, 1994: 33 (Armenia species key); Bolton, 1994: 106 (synoptic classification); Bolton, 1995a: 1050 (census); Bolton, 1995b: 252 (catalogue); Collingwood & Agosti, 1996: 314 (Saudi Arabia species key); Cagniant & Espadaler, 1998: 419 (Morocco species key); Collingwood & Prince, 1998: 14 (Portugal species key).

Myrmicinae Pheidolini Pheidole TRUE TRUE FALSE

Extant: 970 species, 142 subspecies

Fossil: 4 species

Pheidole Westwood, 1839: 219. Type-species: Atta providens, by monotypy.

Taxonomic history

Pheidole in Poneridae, Attidae: Smith, F. 1858b: 172.

Pheidole in Attidae: Smith, F. 1860a: 74; Smith, F. 1862a: 49.

Pheidole in Myrmicidae: Smith, F. 1871a: 331; Cresson, 1887: 262.

Pheidole in Myrmicidae, Pheidolidae: Emery, 1877a: 81.

Pheidole in Myrmicinae: Mayr, 1861: 69 [Myrmicidae]; Mayr, 1865: 22 [Myrmicidae]; Emery & Forel, 1879a: 463 [Myrmicidae]; Dalla Torre, 1893: 88.

Pheidole in Myrmicinae, Myrmicini: Forel, 1895b: 127; Emery, 1895j: 769; Forel, 1899c: 64; Ashmead, 1905b: 383; Wheeler, W.M. 1910g: 139; Kempf, 1972a: 183 (anachronism).

Pheidole in Myrmicinae, Pheidolini: Forel, 1893a: 165; Emery, 1914a: 40; Forel, 1917: 241; Arnold, 1920a: 414; Emery, 1921f: 77; Wheeler, W.M. 1922a: 661; all subsequent authors except Kempf, 1972a: 183, above.

Junior synonyms of PHEIDOLE

Oecophthora Heer, 1852: 15. Type-species: Oecophthora pusilla (junior synonym of Formica megacephala), by monotypy.

Taxonomic history

Oecophthora in Myrmicinae: Mayr, 1855: 453 [Myrmicidae].

Oecophthora as junior synonym of Pheidole: Smith, F. 1858a: 282; Smith, F. 1858b: 172; Roger, 1863b: 30.

Ischnomyrmex Mayr, 1862: 738. Type-species: Myrmica longipes, by monotypy.

Taxonomic history

Ischnomyrmex in Myrmicinae: Mayr, 1865: 18 [Myrmicidae]; Dalla Torre, 1893: 98.

Ischnomyrmex in Myrmicinae, Myrmicini: Emery, 1895j: 769; Ashmead, 1905b: 383; Wheeler, W.M. 1910g: 140.

Ischnomyrmex in Myrmicinae, Pheidolini: Emery, 1914a: 40; Forel, 1917: 241; Emery, 1922e: 113; Wheeler, W.M. 1922a: 661; all subsequent authors.

Ischnomyrmex as junior synonym of Aphaenogaster: Emery & Forel, 1879a: 461; Bingham, 1903: 270.

Ischnomyrmex as subgenus of Aphaenogaster: Forel, 1891b: 166; Dalla Torre, 1893: 98; Forel, 1899c: 59; Emery, 1908c: 313; Wheeler, W.M. 1910g: 140.

Ischnomyrmex as subgenus of Stenamma: Emery, 1895c: 298; Emery, 1895j: 769; Forel, 1902h: 439; Forel, 1903a: 693.

Ischnomyrmex as subgenus of Pheidole: Forel, 1913k: 49.

Ischnomyrmex as genus: Mayr, 1862: 738; Emery, 1914a: 40; Viehmeyer, 1914d: 601; Forel, 1917: 241; Wheeler, W.M. 1922a: 661; Emery, 1922e: 113. Stitz, 1911a: 367; Kusnezov, 1952b: 10 (in key).

Ischnomyrmex as junior synonym of Pheidole: Brown, 1973b: 181 [provisional]; Smith, D.R. 1979: 1365; Snelling, R.R. 1981: 395.

Leptomyrma Motschoulsky, 1863: 17. Type-species: Leptomyrma gracilipes, by monotypy.

Taxonomic history

Leptomyrma in Myrmicidae: Smith, F. 1871a: 329.

Leptomyrma as junior synonym of Pheidole: Emery, 1892b: 166; all subsequent authors.

Pheidolacanthinus Smith, F. 1865: 75. Type-species: Pheidolacanthinus armatus (junior synonym of Myrmica quadrispinosa), by monotypy.

Taxonomic history

Pheidolacanthinus in Myrmicidae: Smith, F. 1871a: 333.

Pheidolacanthinus in Myrmicinae: Dalla Torre, 1893: 98.

Pheidolacanthinus as subgenus of Pheidole: Emery, 1915i: 190; Forel, 1917: 241; Emery, 1921f: 81; Wheeler, W.M. 1922a: 673; all subsequent authors to the following.

Pheidolacanthinus as junior synonym of Pheidole: Brown, 1973b: 183 [provisional]; Smith, D.R. 1979: 1365; Snelling, R.R. 1981: 395.

Ceratopheidole Pergande, 1896: 889 [as subgenus of Pheidole]. Type-species: Pheidole (Ceratopheidole) granulata, by monotypy.

Taxonomic history

Ceratopheidole in Myrmicinae, Pheidolini: Emery, 1914a: 40; Forel, 1917: 241; Emery, 1922e: 112; Wheeler, W.M. 1922a: 661; all subsequent authors.

Ceratopheidole as genus: Emery, 1914a: 40; Forel, 1917: 241; Emery, 1922e: 112; Wheeler, W.M. 1922a: 661; Chapman & Capco, 1951: 135; Wu, J. & Wang, 1995: 104 (anachronism).

Ceratopheidole as subgenus of Pheidole: Pergande, 1896: 889; Kempf, 1972a: 183.

Ceratopheidole as junior synonym of Pheidole: Brown, 1973b: 179 [provisional]; Smith, D.R. 1979: 1365; Snelling, R.R. 1981: 395; Wilson, 2003: 6.

Epipheidole Wheeler, W.M. 1903f: 664. Type-species: Epipheidole inquilina, by monotypy.

Taxonomic history

[Epipheidole also described as new by Wheeler, W.M. 1904a: 14.]

Epipheidole in Myrmicinae, Stenammini: Ashmead, 1905b: 383.

Epipheidole in Myrmicinae, Myrmicini: Wheeler, W.M. 1910g: 140.

Epipheidole in Myrmicinae, Pheidolini: Emery, 1914a: 40; Forel, 1917: 241; Emery, 1922e: 114; Wheeler, W.M. 1922a: 661; Creighton, 1950a: 192.

Epipheidole as junior synonym of Pheidole: Cole, 1965: 174; Smith, D.R. 1979: 1366.

Phidole Bingham, 1903: 220, unjustified emendation of Pheidole.

Taxonomic history

Phidole as junior synonym of Pheidole: Wheeler, 1922a: 806.

Sympheidole Wheeler, W.M. 1904a: 7. Type-species: Sympheidole elecebra, by monotypy.

Taxonomic history

Sympheidole in Myrmicinae, Myrmicini: Wheeler, W.M. 1910g: 140.

Sympheidole in Myrmicinae, Stenammini: Ashmead, 1905b: 383.

Sympheidole in Myrmicinae, Pheidolini: Emery, 1914a: 40; Forel, 1917: 241; Emery, 1922e: 115; Wheeler, W.M. 1922a: 661; all subsequent authors to the following.

Sympheidole as junior synonym of Pheidole: Brown, 1973b: 185 [provisional]; Smith, D.R. 1979: 1365; Snelling, R.R. 1981: 395; Wilson, 2003: 6.

Phidola Schulz, W.A. 1906: 155, unjustified emendation of Pheidole.

Taxonomic history

Phidola as junior synonym of Pheidole: Smith, D.R. 1979: 1365.

Allopheidole Forel, 1912f: 237 [as subgenus of Pheidole]. Type-species: Pheidole kingi (junior synonym of Pheidole tepicana), by subsequent designation of Wheeler, W.M. 1913a: 79.

Taxonomic history

Allopheidole as subgenus of Pheidole: Forel, 1912f: 237; Forel, 1917: 241.

Allopheidole as junior synonym of Pheidole: Emery, 1915i: 190; Emery, 1921f: 84; Smith, D.R. 1979: 1366.

Decapheidole Forel, 1912f: 237 [as subgenus of Pheidole]. Type-species: Pheidole perpusilla, by subsequent designation of Wheeler, W.M. 1913a: 80.

Taxonomic history

Decapheidole in Myrmicinae, Pheidolini: Wheeler, W.M. 1922a: 661.

Decapheidole as genus: Wheeler, W.M. 1922a: 661; Borgmeier, 1927c: 78.

Decapheidole as subgenus of Pheidole: Forel, 1912f: 237, Forel, 1917: 241; Emery, 1922e: 112; Kempf, 1972a: 183.

Decapheidole as junior synonym of Pheidole: Brown, 1973b: 180 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395; Wilson, 2003: 6.

Isopheidole Forel, 1912m: 765 [as subgenus of Pheidole]. Type-species: Myrmica longipes, by monotypy.

Taxonomic history

Isopheidole as junior synonym of Ischnomyrmex: Forel, 1913b: 350 (in text); Forel, 1913k: 49; Emery, 1922e: 113; Wheeler, W.M. 1922a: 680 (in key). [Ischnomyrmex and Isopheidole share the same type-species, synonymy is therefore absolute.]

Elasmopheidole Forel, 1913k: 43 [as subgenus of Pheidole]. Type-species: Pheidole aberrans, by subsequent designation of Emery, 1915i: 190.

Taxonomic history

Elasmopheidole as subgenus of Pheidole: Forel, 1913k: 43; Emery, 1915i: 190; Forel, 1917: 241; Emery, 1922e: 110; Wheeler, W.M. 1922a: 673; all subsequent authors to the following.

Elasmopheidole as junior synonym of Pheidole: Brown, 1973b: 180 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395.

Cardiopheidole Wheeler, W.M. 1914b: 48 [as subgenus of Pheidole]. Type-species: Pheidole vaslitii, by original designation.

Taxonomic history

Cardiopheidole as subgenus of Pheidole: Wheeler, W.M. 1914b: 48; Forel, 1917: 241.

Cardiopheidole as junior synonym of Pheidole: Emery, 1915i: 190; Emery, 1921f: 84; Smith, D.R. 1979: 1366.

Anergatides Wasmann, 1915b: 281. Type-species: Anergatides kohli (junior secondary homonym in Pheidole, replaced by Pheidole neokohli), by monotypy.

Taxonomic history

Anergatides in Myrmicinae, Pheidolini: Forel, 1917: 243 [subtribe Anergatini]; Emery, 1922e: 116 [subtribe Anergatidini]; Donisthorpe, 1943f: 623.

Anergatides in Myrmicinae, Solenopsidini: Wheeler, W.M. 1922a: 663; Wheeler, G.C. & Wheeler, J. 1985: 257 (anachronism).

Anergatides as junior synonym of Pheidole: Brown, 1973b: 178 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395; Wilson, 1984: 328; Bolton, 1994: 106.

Parapheidole Emery, 1915d: 68. Type-species: Aphaenogaster oculata, by monotypy.

Taxonomic history

Parapheidole in Myrmicinae, Pheidolini: Forel, 1917: 241; Emery, 1922e: 115; Wheeler, W.M. 1922a: 661; all subsequent authors.

Parapheidole as junior synonym of Pheidole: Brown, 1973b: 183 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395.

Macropheidole Emery, 1915i: 190 [as subgenus of Pheidole]. Type-species: Pheidole fimbriata, by monotypy.

Taxonomic history

Macropheidole as subgenus of Pheidole: Emery, 1915i: 190; Forel, 1917: 241; Emery, 1921f: 81; Wheeler, W.M. 1922a: 673; all subsequent authors to the following.

Macropheidole as junior synonym of Pheidole: Smith, M.R. 1951a: 800; Brown, 1973b: 181 [provisional]; Smith, D.R. 1979: 1366.

Scrobopheidole Emery, 1915i: 190 [as subgenus of Pheidole]. Type-species: Pheidole scrobifera, by monotypy.

Taxonomic history

Scrobopheidole as subgenus of Pheidole: Forel, 1917: 241; Emery, 1922e: 112; Wheeler, W.M. 1922a: 673; all subsequent authors to the following.

Scrobopheidole as junior synonym of Pheidole: Brown, 1973b: 184 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395; Wilson, 2003: 6.

Stegopheidole Emery, 1915i: 190 [as subgenus of Pheidole]. Type-species: Pheidole (Elasmopheidole) upeneci, by monotypy.

Taxonomic history

Stegopheidole as subgenus of Pheidole: Emery, 1921f: 83; Wheeler, W.M. 1922a: 673; all subsequent authors to the following.

Stegopheidole as junior synonym of Pheidole: Brown, 1973b: 185 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395.

Trachypheidole Emery, 1915i: 190 [as subgenus of Pheidole]. Type-species: Pheidole bicornis, by original designation.

Taxonomic history

Trachypheidole as subgenus of Pheidole: Forel, 1917: 241; Emery, 1922e: 111; Wheeler, W.M. 1922a: 673; all subsequent authors to the following.

Trachypheidole as junior synonym of Pheidole: Brown, 1973b: 185 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395; Wilson, 2003: 6.

Electropheidole Mann, 1921: 438 [as subgenus of Pheidole]. Type-species: Pheidole (Electropheidole) roosevelti, by subsequent designation of Donisthorpe, 1943f: 642.

Taxonomic history

Electropheidole as subgenus of Pheidole: Mann, 1921: 438; Wheeler, W.M. 1922a: 672; all subsequent authors to the following.

Electropheidole as junior synonym of Pheidole: Brown, 1973b: 180 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395.

Bruchomyrma Santschi, 1922d: 248. Type-species: Bruchomyrma acutidens, by monotypy.

Taxonomic history

Bruchomyrma in Myrmicinae, Pheidolini: Donisthorpe, 1943f: 629.

Bruchomyrma in Myrmicinae, Myrmicini: Kempf, 1972a: 42.

Bruchomyrma as junior synonym of Pheidole: Brown, 1973b: 179 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395; Wilson, 1984: 327; Bolton, 1994: 106; Wilson, 2003: 6.

Bruchomyrma as genus: Dlussky & Fedoseeva, 1988: 80 (anachronism).

Cephalomorium Forel, 1922: 91 [as subgenus of Tetramorium]. Type-species: Tetramorium (Cephalomorium) bahai, by monotypy.

Taxonomic history

Cephalomorium as junior synonym of Hendecapheidole: Santschi, 1925d: 228.

Hendecapheidole Wheeler, W.M. 1922d: 3 [as subgenus of Pheidole]. Type-species: Pheidole tachigaliae, by original designation.

Taxonomic history

Hendecapheidole as junior synonym of Pheidole: Brown, 1973b: 181 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395; Wilson, 2003: 6.

Gallardomyrma Bruch, 1932: 271. Type-species: Gallardomyrma argentina, by original designation.

Taxonomic history

Gallardomyrma in Myrmicinae, Pheidolini: Donisthorpe, 1943f: 646.

Gallardomyrma in Myrmicinae, Myrmicini: Kempf, 1972a: 110.

Gallardomyrma as junior synonym of Pheidole: Brown, 1973b: 180 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395; Wilson, 1984: 327; Wilson, 2003: 6.

Gallardomyrma as genus: Kempf, 1972a: 110; Dlussky & Fedoseeva, 1988: 80 (anachronism).

Conothoracoides Strand, 1935: 176.

Taxonomic history

[Replacement name for Conothorax Karavaiev; junior homonym of Conothorax Jekel, 1854: 9 bis (foldout pages) (Coleoptera).]

Conothoracoides as junior synonym of Pheidole: Brown, 1973b: 179 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395.

Homonym replaced by Conothoracoides

Conothorax Karavaiev, 1935a: 75. Type-species: Conothorax bilobum, by monotypy.

Taxonomic history

[Conothorax Karavaiev junior homonym of Conothorax Jekel, 1854: 9 bis (foldout pages) (Coleoptera).]

Conothorax in Myrmicinae, Pheidolini: Karavaiev, 1935a: 75.

Eriopheidole Kusnezov, 1952a: 10. Type-species: Eriopheidole symbiotica (unresolved junior secondary homonym in Pheidole), by monotypy.

Taxonomic history

Eriopheidole in Myrmicinae, Pheidolini: Kusnezov, 1952a: 10.

Eriopheidole in Myrmicinae, Myrmicini: Kempf, 1972a: 107.

Eriopheidole junior synonym of Pheidole: Brown, 1973b: 180 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395; Bolton, 1994: 106.

Eriopheidole as genus: Kempf, 1972a: 107; Dlussky & Fedoseeva, 1988: 80 (anachronism).

Xenoaphaenogaster Baroni Urbani, 1964b: 50. Type-species: Xenoaphaenogaster inquilina (junior synonym of Myrmica pallidula), by original designation.

Taxonomic history

Xenoaphaenogaster as junior synonym of Monomorium: Brown, 1973b: 185 [provisional].

Xenoaphaenogaster as junior synonym of Pheidole: Bolton, 1987: 291.

Genus Pheidole references

Smith, F. 1858b: 172 (diagnosis); Mayr, 1861: 70 (Europe species key); Roger, 1863b: 27, 30 (catalogue); Mayr, 1863: 424, 440 (Ischnomyrmex, Pheidole catalogues); Mayr, 1865: 18, 22 (Ischnomyrmex, Pheidole diagnoses); Mayr, 1867a: 92, 96 (Ischnomyrmex, Pheidole diagnoses); Mayr, 1870b: 977, 979 (Australia & New World species keys); Mayr, 1876: 102 (Australia species key); André, 1883b: 382 (Europe & Algeria species key); Mayr, 1879: 674 (Asia species key); Mayr, 1887: 582 (New World species key); Cresson, 1887: 262 (U.S.A., catalogue); Nasonov, 1889: 75 (Russia species key); Dalla Torre, 1893: 88, 98 (Pheidole, Pheidolacanthinus catalogues); Emery, 1895c: 288 (North America species key); Mayr, 1896: 241 (Africa species, partial key); Emery, 1896g: 80 (P. flavens group, key); Forel, 1902c: 165 (India species key); Forel, 1902f: 523 (India & Sri Lanka species key); Bingham, 1903: 223 (India, Sri Lanka & Burma species key); Wheeler, W.M. 1908a: 133 (Puerto Rico species key); Arnold, 1920a: 414 (diagnosis, South Africa species key); Emery, 1921f: 78 (diagnosis, subgenera key, catalogue); Emery, 1921f: 81 (P. (Macropheidole) & P. (Pheidolacanthinus) diagnoses, catalogues); Emery, 1921f: 83 (P. (Stegopheidole) diagnosis, catalogue); Mann, 1921: 442 (Fiji Is species key); Emery, 1922e: 110 (P. (Elasmopheidole) diagnosis, catalogue); Emery, 1922e: 111 (P. (Trachypheidole) diagnosis, catalogue); Emery, 1922e: 112 (P. (Scrobopheidole), P. (Decapheidole) & P. (Ceratopheidole) diagnoses, catalogues); Emery, 1922e: 113 (Ischnomyrmex diagnosis, catalogue); Emery, 1922e: 114 (Epipheidole diagnosis, catalogue); Emery, 1922e: 115 (Sympheidole, Parapheidole diagnoses, catalogues); Emery, 1922e: 116 (Anergatides diagnosis, catalogue); Wheeler, W.M. 1922a: 126, 672 (diagnosis, subgenera key); Wheeler, W.M. 1922a: 806, 879 (Afrotropical Pheidole, Anergatides catalogues); Wheeler, W.M. 1922a: 1017, 1021 (Malagasy Pheidole, Parapheidole catalogues); Gallardo, 1932c: 179 (Argentina Elasmopheidole species key); Finzi, 1936: 165 (Egypt species key); Menozzi, 1939a: 298 (Himalaya species key); Cole, 1942: 362 (U.S.A., Utah species key); Buren, 1944a: 285 (U.S.A., Iowa species key); Creighton, 1950a: 163 (North America species key); Creighton, 1950a: 192, 194 (North America Epipheidole, Sympheidole species); Chapman & Capco, 1951: 135, 137 (Asia Ceratopheidole, Conothorax, Ischnomyrmex, Pheidole checklists); Kusnezov, 1952b: 61 (Argentina species key); Smith, M.R. 1955c: 302 (P. (Hendecapheidole) species key); Gregg, 1959: 9 (U.S.A. species key); Gregg, 1963: 407 (U.S.A., Colorado species key); Wheeler, G.C. & Wheeler, J. 1963: 126 (U.S.A., North Dakota species key); Bernard, 1967: 149 (diagnosis, Western Europe species key); Gregg, 1969: 95 (addition to Gregg, 1959 U.S.A. key); Kempf, 1972a: 183 (Neotropical catalogue); Alayo, 1974: 11 (Cuba species key); Smith, D.R. 1979: 1365 (North America catalogue); Allred, 1982: 441 (U.S.A., Utah species key); Ogata, 1982: 191 (Japan species key); Collingwood, 1985: 253 (Saudi Arabia species key); Naves, 1985: 54 (U.S.A., Florida species key); Taylor & Brown, D.R. 1985: 74 (Australia catalogue); Wheeler, G.C. & Wheeler, J. 1986g: 41 (U.S.A., Nevada species key); Taylor, 1987a: 53 (Australia, New Caledonia & New Zealand checklists); Agosti & Collingwood, 1987b: 271 (Balkans species key); Hölldobler & Wilson, 1990: 15 (synoptic classification); Kupyanskaya, 1990: 123 (Far Eastern Russia species key); Brandão, 1991: 368 (Neotropical catalogue); Morisita, Kubota, et al. 1992: 22 (Japan species key); Bolton, 1994: 106 (synoptic classification); Kupyanskaya, 1995: 345 (Far Eastern Russia species key); Bolton, 1995a: 1051 (census); Bolton, 1995b: 316 (catalogue); Wu, J. & Wang, 1995: 99 (China species key); Collingwood & Agosti, 1996: 322 (Saudi Arabia species key); Park & Kim, 2000: 109 (Korea species key); Shattuck, 1999: 152 (Australia synopsis); Eguchi, 1999: 103 (Borneo P. longipes group, key); Ward, 2000: 96 (U.S.A. P. hyatti complex, key); Zhou, 2001b: 121 (China, Guangxi species key); Eguchi, 2001b: 10 (Borneo species key); ; Mackay & Mackay, 2002: 149 (U.S.A., New Mexico species key)Wilson, 2003: 27 (New World, all species revision, keys); Sarnat, 2008: 8 (Fiji Is P. roosevelti group, key); Eguchi, 2008: 8 (North Vietnam species key); Terayama, 2009: 171 (Taiwan species key); Heterick, 2009: 169 (south-western Australia species key); Sarnat & Moreau, 2011: 114 (P. roosevelti group phylogeny and radiation).

Genera incertae sedis in Pheidolini

Myrmicinae Formicoxenini Cardiocondyla TRUE TRUE FALSE

67 species, 2 subspecies

Cardiocondyla Emery, 1869b: 20. Type-species: Cardiocondyla elegans, by monotypy.

Taxonomic history

Cardiocondyla in Myrmicinae: Emery, 1877a: 81 [Myrmicidae]; Emery & Forel, 1879a: 456 [Myrmicidae]; Dalla Torre, 1893: 70.

Cardiocondyla in Myrmicinae, Myrmicini: Emery, 1895j: 769; Wheeler, W.M. 1910g: 139; Kusnezov, 1964: 57 (anachronism).

Cardiocondyla in Myrmicinae, Stenammini: Ashmead, 1905b: 383.

Cardiocondyla in Myrmicinae, Cardiocondylini: Emery, 1914a: 40; Arnold, 1916: 200; Forel, 1917: 242; Emery, 1922e: 124; Wheeler, W.M. 1922a: 661; all subsequent authors; Dlussky & Fedoseeva, 1988: 80 (anachronism); Jaffe, 1993: 10 (anachronism).

Cardiocondyla in Myrmicinae, Leptothoracini: Bolton, 1982: 311 (in text).

Cardiocondyla in Myrmicinae, Formicoxenini: Bolton, 1994: 105.

Junior synonyms of CARDIOCONDYLA

Emeryia Forel, 1890b: cx. Type-species: Emeryia wroughtonii, by monotypy.

Taxonomic history

Emeryia as junior synonym of Cardiocondyla: Forel, 1892h: 461; Forel, 1892i: 313; Dalla Torre, 1893: 70.

Xenometra Emery, 1917a: 96. Type-species: Xenometra monilicornis (junior synonym of Cardiocondyla emeryi), by monotypy.

Taxonomic history

Xenometra in Myrmicinae, Cardiocondylini: Emery, 1922e: 126; Wheeler, W.M. 1922a: 661; subsequent authors.

Xenometra as junior synonym of Cardiocondyla: Baroni Urbani, 1973: 199; Marikovsky & Yakushin, 1974: 60.

Dyclona Santschi, 1930b: 70 (footnote) [as subgenus of Cardiocondyla]. Type-species: Monomorium cristatum, by original designation.

Taxonomic history

Dyclona as junior synonym of Cardiocondyla: Brown, 1973b: 180 [provisional]; Smith, D.R. 1979: 1375; Bolton, 1982: 309.

Loncyda Santschi, 1930b: 70 [as subgenus of Cardiocondyla]. Type-species: Cardiocondyla (Loncyda) monardi, by monotypy.

Taxonomic history

Loncyda as junior synonym of Cardiocondyla: Brown, 1973b: 181 [provisional]; Smith, D.R. 1979: 1375; Bolton, 1982: 309.

Prosopidris Wheeler, W.M. 1935b: 40 [as subgenus of Cardiocondyla]. Type-species: Cardiocondyla (Prosopidris) sima, by original designation.

Taxonomic history

Prosopidris in Myrmicinae, Cardiocondylini: Donisthorpe, 1943f: 688.

Prosopidris as genus: Reiskind, 1965: 80.

Prosopidris as junior synonym of Cardiocondyla: Brown, 1973b: 184 [provisional]; Smith, D.R. 1979: 1375; Bolton, 1982: 309.

Genus Cardiocondyla references

André, 1883a: 327 (Europe & Algeria species key); Dalla Torre, 1893: 70 (catalogue); Forel, 1903a: 688 (India & Sri Lanka species key); Bingham, 1903: 287 (India, Sri Lanka & Burma species key); Ruzsky, 1905b: 623 (Russian Empire species key); Emery, 1909a: 20 (Palaearctic species key); Arnold, 1916: 200 (diagnosis); Emery, 1922e: 124 (diagnosis, catalogue); Emery, 1922e: 126 (Xenometra diagnosis, catalogue); Wheeler, W.M. 1922a: 149, 827, 1021 (diagnosis, Afrotropical, Malagasy catalogues); Kuznetsov-Ugamsky, 1927d: 37 (Turkestan species key); Karavaiev, 1934: 114 (Ukraine species key); Finzi, 1936: 172 (Egypt species key); Smith, M.R. 1944a: 32 (U.S.A. species key); Creighton, 1950a: 197 (North America species key); Chapman & Capco, 1951: 83 (Asia checklist); Bernard, 1956c: 301 (Palaearctic species key); Bernard, 1967: 156 (diagnosis, Western Europe species key); Bernard, 1967: 158 (Xenometra diagnosis); Kempf, 1972a: 73, 259 (Neotropical Cardiocondyla, Xenometra catalogues); Alayo, 1974: 12 (Cuba species key); Tarbinsky, 1976: 71 (Kyrghyzstan species key); Arnol'di & Dlussky, 1978: 538 (former European U.S.S.R. species key); Collingwood, 1978: 86 (Iberian Peninsula species key); Smith, D.R. 1979: 1375 (North America catalogue); Bolton, 1982: 309 (diagnosis, Afrotropical species key); Kugler, J. 1984: 17 (males, key); Collingwood, 1985: 256 (Saudi Arabia species key); Agosti & Collingwood, 1987b: 276 (Balkans species key); Dlussky, Soyunov & Zabelin, 1990: 193 (Turkmenistan species key); Morisita, Kubota, et al. 1992: 31 (Japan species key); Atanasov & Dlussky, 1992: 172 (Bulgaria species key); Arakelian, 1994: 42 (Armenia species key); Bolton, 1995a: 1048 (census); Bolton, 1995b: 132 (catalogue); Radchenko, 1995b: 452 (Palaearctic species key); Mackay, 1995: 170 (New World species key); Wu, J. & Wang, 1995: 68 (China species key); Collingwood & Agosti, 1996: 326 (Saudi Arabia species key); Collingwood & Prince, 1998: 16 (Portugal species key); Terayama, 1999d: 99 (Japan species, review); Shattuck, 1999: 129 (Australia synopsis); Zhou, 2001b: 85 (China, Guangxi species key); Rigato, 2002: 172 (Afrotropical species key); Seifert, 2003a: 292 (C. elegans, C. bulgarica, C. batesii, C. nuda, C. shuckardi, C. stambuloffii, C. wroughtonii, C. emeryi, C. minutior groups revision, key); Terayama, 2009: 179 (Taiwan species key).

Myrmicinae Formicoxenini Formicoxenus TRUE TRUE FALSE

7 species

Formicoxenus Mayr, 1855: 413. Type-species: Myrmica nitidula, by monotypy.

Taxonomic history

Formicoxenus in Myrmicinae: Mayr, 1855: 413 [Myrmicidae]; Dalla Torre, 1893: 62.

Formicoxenus in Myrmicinae, Myrmicini: Emery, 1895j: 769; Wheeler, W.M. 1910g: 139.

Formicoxenus in Myrmicinae, Stenammini: Ashmead, 1905b: 383.

Formicoxenus in Myrmicinae, Leptothoracini: Emery, 1914a: 42; Forel, 1917: 245; Wheeler, W.M. 1922a: 664; Emery, 1924d: 264; all subsequent authors except the following.

Formicoxenus in Myrmicinae, Formicoxenini: Forel, 1893a: 165; Bolton, 1994: 105.

Formicoxenus as junior synonym of Stenamma: Mayr, 1863: 422; Roger, 1863b: 25; Emery & Forel, 1879a: 457.

Formicoxenus as junior synonym of Leptothorax: Brown, 1973b: 180 [provisional].

Formicoxenus as genus: André, 1882c: 273; all subsequent authors except the above; Francoeur, Loiselle & Buschinger, 1985: 343.

Junior synonym of FORMICOXENUS

Symmyrmica Wheeler, W.M. 1904a: 3. Type-species: Symmyrmica chamberlini, by monotypy.

Taxonomic history

Symmyrmica in Myrmicinae, Myrmicini: Wheeler, W.M. 1910g: 139.

Symmyrmica in Myrmicinae, Stenammini: Ashmead, 1905b: 383.

Symmyrmica in Myrmicinae, Leptothoracini: Emery, 1914a: 42; Forel, 1917: 245; Wheeler, W.M. 1922a: 664; Emery, 1924d: 263; all subsequent authors.

Symmyrmica as junior synonym of Leptothorax: Brown, 1973b: 185 [provisional].

Symmyrmica as junior synonym of Formicoxenus: Francoeur, Loiselle & Buschinger, 1985: 347.

Genus Formicoxenus references

André, 1882c: 273 (Europe & Algeria species); Dalla Torre, 1893: 62 (catalogue); Emery, 1924d: 263 (Symmyrmica diagnosis, catalogue); Emery, 1924d: 264 (diagnosis, catalogue); Bernard, 1967: 225 (diagnosis); Smith, D.R. 1979: 1398 (Symmyrmica catalogue); Francoeur, Loiselle & Buschinger, 1985: 374 (diagnosis, all species revision, key); Kupyanskaya, 1990: 148 (Far Eastern Russia species key); Radchenko, 1994b: 111 (South Siberia species key); Bolton, 1995a: 1049 (census); Bolton, 1995b: 206 (catalogue); Mackay & Mackay, 2002: 91 (U.S.A., New Mexico species key).

Myrmicinae Formicoxenini Leptothorax TRUE TRUE FALSE

Extant: 18 species, 2 subspecies

Fossil:

Leptothorax Mayr, 1855: 431. Type-species: Formica acervorum, by subsequent designation of Bingham, 1903: 214.

Taxonomic history

[Type-species not Myrmica clypeata, unjustified subsequent designation by Emery, 1912d: 271; repeated in Wheeler, W.M. 1913a: 79 and Emery, 1924d: 248.]

Leptothorax in Myrmicidae, Myrmicidae: Emery, 1877a: 81.

Leptothorax in Myrmicinae: Mayr, 1855: 431 [Myrmicidae]; Mayr, 1861: 57 [Myrmicidae]; Mayr, 1865: 20 [Myrmicidae]; Emery & Forel, 1879a: 458 [Myrmicidae]; Dalla Torre, 1893: 122.

Leptothorax in Myrmicinae, Myrmicini: Forel, 1895b: 125; Emery, 1895j: 769; Forel, 1899c: 54; Wheeler, W.M. 1910g: 139; Wheeler, W.M. 1915h: 63; Kusnezov, 1964: 57 (anachronism).

Leptothorax in Myrmicinae, Stenammini: Ashmead, 1905b: 383.

Leptothorax in Myrmicinae, Leptothoracini: Emery, 1914a: 42; Arnold, 1916: 257; Forel, 1917: 244; Wheeler, W.M. 1922a: 664; Emery, 1924d: 247; subsequent authors to the following.

Leptothorax in Myrmicinae, Formicoxenini: Bolton, 1994: 105; Mackay, 2000: 267.

Junior synonyms of LEPTOTHORAX

Mychothorax Ruzsky, 1904a: 288. Type-species: Formica acervorum, by original designation.

Taxonomic history

Mychothorax as subgenus of Leptothorax: Ruzsky, 1905b: 609; Emery, 1915g: 24; Emery, 1916b: 176; Forel, 1917: 245; Wheeler, W.M. 1922a: 679; Emery, 1924d: 260; Creighton, 1950a: 274.

Mychothorax as junior synonym of Leptothorax: Smith, M.R. 1950: 29; Brown, 1973b: 182 [provisional]; Bolton, 1982: 319. [Leptothorax and Mychothorax share the same type-species, synonymy is therefore absolute.]

Doronomyrmex Kutter, 1945: 485. Type-species: Doronomyrmex pacis, by monotypy.

Taxonomic history

Doronomyrmex in Myrmicinae, Leptothoracini:Dlussky & Fedoseeva, 1988: 79.

Doronomyrmex in Myrmicinae, Formicoxenini: Bolton, 1994: 105.

Doronomyrmex as junior synonym of Leptothorax: Brown, 1973b: 180 [provisional]; Heinze, 1998: 195; Bolton, 2003: 270.

Genus Leptothorax references

Mayr, 1855: 433 (Austria species key); Mayr, 1861: 58 (Europe species key); Roger, 1863b: 26, 28, 30 (catalogue); Mayr, 1863: 426, 428, 456 (Leptothorax, Macromischa, Temnothorax catalogues); Mayr, 1865: 19, 20, 21 (Macromischa, Leptothorax, Temnothorax diagnoses); Mayr, 1868c: 83 (†Baltic Amber species key); André, 1874: 188 (Europe species key); Forel, 1874: 84 (Switzerland species key); André, 1883a: 293 (Europe & Algeria species key); Mayr, 1886d: 451 (U.S.A. species key); Cresson, 1887: 261 (U.S.A. catalogue); Nasonov, 1889: 70 (Russia species key); Emery, 1891b: 5 (North Africa species key); Dalla Torre, 1893: 120, 122 (Macromischa, Leptothorax catalogues); Emery, 1895c: 317 (North America species key); Emery, 1896g: 58 (Neotropical species key); Bingham, 1903: 215 (India, Sri Lanka & Burma species key); Wheeler, W.M. 1903c: 223 (North America species key); Ruzsky, 1905b: 570 (Russian Empire species key); Wasmann, 1906: 16 (Luxemburg species key); Wheeler, W.M. 1908a: 141 (Macromischa species key); Santschi, 1909b: 460 (L. rottenbergi group key); Bondroit, 1910: 496 (Belgium species key); Stitz, 1914: 60 (Central Europe species key); Crawley, 1914: 91 (Britain species key); Donisthorpe, 1915d: 147 (Britain species key); Emery, 1916b: 176 (Italy species key); Wheeler, W.M. 1916m: 581 (U.S.A., Connecticut species key); Arnold, 1916: 257 (diagnosis, South Africa species key); Bondroit, 1918: 117 (France & Belgium species key); Mann, 1920: 408 (Macromischa species key); Wheeler, W.M. 1922a: 677 (Macromischa subgenera key); Wheeler, W.M. 1922a: 679 (Leptothorax subgenera key); Wheeler, W.M. 1922a: 890 (Afrotropical catalogue); Wheeler, W.M. 1922a: 1029 (Malagasy catalogue); Emery, 1924d: 245 (Macromischa catalogue); Emery, 1924d: 247 (diagnosis, subgenera key, catalogue); Emery, 1924d: 249 (L. (Goniothorax) diagnosis, catalogue); Emery, 1924d: 251 (L. (Leptothorax) diagnosis, catalogue); Emery, 1924d: 259 (L. (Temnothorax) diagnosis, catalogue); Emery, 1924d: 260 (L. (Dichothorax) & L. (Mychothorax) diagnoses, catalogues); Emery, 1924d: 264 (Symmyrmica review, catalogue); Karavaiev, 1927c: 266 (Ukraine species key); Donisthorpe, 1927b: 163 (Britain species key); Kuznetsov-Ugamsky, 1927d: 38 (Turkestan species key); Wheeler, W.M. 1931b: 31 (Macromischa, Croesomyrmex, Antillaemyrmex checklists); Arnol'di, 1933b: 598 (Russia species key); Menozzi, 1933b: 68 (Israel species key); Karavaiev, 1934: 134 (Ukraine species key); Wheeler, W.M. 1937b: 463 (Macromischa checklist); Menozzi, 1939a: 307 (Himalaya & Tibet species key); Stitz, 1939: 158 (Germany species key); Smith, M.R. 1939e: 503 (U.S.A. Macromischa species key); Kratochvíl, 1941: 89 (Central Europe species key); Novák & Sadil, 1941: 89 (Central Europe species key); Cole, 1942: 369 (U.S.A., Utah species key); Holgersen, 1943b: 172 (Norway species key); Holgersen, 1944: 198 (Norway species key); Kratochvíl, Novák & Snoflák, 1944: 123 (Czechoslovakia males, key); Buren, 1944a: 286 (U.S.A., Iowa species key); Boven, 1947: 178 (Belgium species key); Creighton, 1950a: 251, 256 (North America Macromischa, Leptothorax species keys); Chapman & Capco, 1951: 110 (Asia checklist); Smith, M.R. 1952b: 97 (U.S.A. L. tricarinatus complex, key); Bernard, 1956a: 151 (Western Europe species groups); Kusnezov, 1958c: 266 (subgenera key); Boven, 1959: 8 (Netherlands species key); Gregg, 1963: 380 (U.S.A., Colorado species key); Wheeler, G.C. & Wheeler, J. 1963: 139 (U.S.A., North Dakota species key); Collingwood, 1964: 99 (Britain species key); Bernard, 1967: 185 (diagnosis, Western Europe species key); Bernard, 1967: 187 (diagnosis, Western Europe species key); Bernard, 1967: 224 (Doronomyrmex diagnosis); Boven, 1970b: 20 (Netherlands species key); Arnol'di, 1971: 1824 (Kazakhstan species key); Kempf, 1972a: 132, 135 (Neotropical Leptothorax, Macromischa catalogues); Alayo, 1974: 15 (Macromischa Cuba species key); Bolton & Collingwood, 1975: 5 (Britain species key); Tarbinsky, 1976: 83 (Kyrghyzstan species key); Boven, 1977: 94 (Belgium species key); Kutter, 1977c: 103 (Switzerland species key); Arnol'di & Dlussky, 1978: 540 (former European U.S.S.R. species key); Collingwood, 1978: 83 (Iberian Peninsula species key); Collingwood, 1979: 68 (Fennoscandia & Denmark species key); Baroni Urbani, 1978b: 533 (L. (Macromischa) all species revision, key); Smith, D.R. 1979: 1391 (North America catalogue); Buschinger, 1981: 211 (Doronomyrmex review); Allred, 1982: 440 (U.S.A., Utah species key); Bolton, 1982: 319 (diagnosis, review of genus, Afrotropical species key); Gösswald, 1985: 303 (Germany species key); Wheeler, G.C. & Wheeler, J. 1986g: 51 (U.S.A., Nevada species key); Nilsson & Douwes, 1987: 60 (Norway species key); Agosti & Collingwood, 1987b: 273 (Balkans species key); Dlussky & Soyunov, 1988: 29 (former U.S.S.R. Temnothorax species); Hölldobler & Wilson, 1990: 13 (synoptic classification); Dlussky, Soyunov & Zabelin, 1990: 184, 188 (Turkmenistan species key); Kupyanskaya, 1990: 136 (Far Eastern Russia species key); Brandão, 1991: 349 (Neotropical catalogue); Morisita, Kubota, et al. 1992: 26 (Japan species key); Atanasov & Dlussky, 1992: 129 (Bulgaria species key); Arakelian, 1994: 52 (Armenia species key); Radchenko, 1994b: 111 (South Siberia species key); Radchenko, 1994d: 146 (Central & Eastern Palaearctic species key); Bolton, 1995a: 1050 (census); Bolton, 1995b: 235 (catalogue); Douwes, 1995: 89 (Sweden species key); Kupyanskaya, 1995: 347 (Far Eastern Russia species key); Wu, J. & Wang, 1995: 108 (China species key); Collingwood & Agosti, 1996: 325 (Saudi Arabia species key); Seifert, 1996b: 13 (Central Europe Doronomyrmex species key); Seifert, 1996b: 119 (Central Europe species key); Skinner & Allen, 1996: 45 (Britain species key); Cagniant & Espadaler, 1997: 260 (Morocco species key); Collingwood & Prince, 1998: 16 (Portugal species key); Terayama & Onoyama, 1999: 73 (Japan species key); Mackay, 2000: 265, 288, 297 (New World L. (Myrafant) species revision key, Mexico & Guatemala species key, species complexes key); Fontenla Rizo, 2001: 15 (Cuba Macromischa species key); Chang & He, 2001b: 1 (northwest China species key); Czechowski, Radchenko & Czechowska, 2002: 143 (Poland species key); Mackay & Mackay, 2002: 93 (U.S.A., New Mexico species key); Bolton, 2003: 270 (genus diagnosis); Lyu & Cho, 2003b: 269 (Korea species key); Radchenko, 2004: 131 (Eastern Palaearctic species, key); Deyrup & Cover, 2004a: 56 (southeast U.S.A. species key); Radchenko, 2005b: 192 (North Korea species key); Seifert, 2007: 118 (North and Central Europe species key).

Myrmicinae Formicoxenini Tricytarus FALSE FALSE FALSE

Tricytarus Donisthorpe, 1947d: 187. Type-species: Tricytarus parviumgulatus, by original designation.

Taxonomic history

Tricytarus incertae sedis in Formicidae: Wheeler, G.C. & Wheeler, J. 1985: 259 (incomprehensible entry).

Tricytarus incertae sedis in Myrmicinae: Hölldobler & Wilson, 1990: 16.

Tricytarus in Myrmicinae, Formicoxenini?: Bolton, 1994: 105; Bolton, 1995b: 422.

[Tricytarus an unrecognisable taxon: a male-based genus, description inadequate. Type-material not present in California Academy of Sciences, San Francisco, U.S.A., nor in The Natural History Museum, London, U.K.; presumed lost: Bolton, 2003: 254.]

Myrmicinae Myrmecinini Myrmecina TRUE TRUE FALSE

51 species, 1 subspecies

Myrmecina Curtis, 1829: 265. Type-species: Myrmecina latreillii (junior synonym of Formica graminicola), by monotypy.

Taxonomic history

Myrmecina in Poneridae, Myrmicidae: Smith, F. 1858b: 132.

Myrmecina in Myrmicidae: Smith, F. 1871a: 327; Cresson, 1887: 261.

Myrmecina in Myrmicidae, Myrmicidae: Emery, 1877a: 81.

Myrmecina in Myrmicinae: Mayr, 1855: 420 [Myrmicidae]; Mayr, 1861: 73 [Myrmicidae]; Mayr, 1865: 21 [Myrmicidae]; Emery & Forel, 1879a: 457 [Myrmicidae]; Dalla Torre, 1893: 61.

Myrmecina in Myrmicinae, Myrmicini: Emery, 1895j: 769; Wheeler, W.M. 1910g: 139.

Myrmecina in Myrmicinae, Myrmecinini: Ashmead, 1905b: 383; Emery, 1912b: 105; Emery, 1914a: 41; Forel, 1917: 244; Wheeler, W.M. 1922a: 663; Emery, 1924d: 230; all subsequent authors.

Junior synonym of MYRMECINA

Archaeomyrmex Mann, 1921: 448. Type-species: Archaeomyrmex cacabau, by original designation.

Taxonomic history

Archaeomyrmex in Myrmicinae, Archaeomyrmecini: Mann, 1921: 449; Wheeler, W.M. 1922a: 663.

Archaeomyrmex as junior synonym of Myrmecina: Brown, 1971a: 1.

Genus Myrmecina references

Smith, F. 1858b: 132 (diagnosis); Roger, 1863b: 33 (catalogue); Mayr, 1863: 429 (catalogue); Mayr, 1865: 21 (diagnosis); André, 1882c: 275 (Europe & Algeria species); Cresson, 1887: 261 (U.S.A. catalogue); Dalla Torre, 1893: 61 (catalogue); Bingham, 1903: 197 (diagnosis); Emery, 1916b: 171 (Italy species key); Mann, 1919: 337 (Papuasian species key); Emery, 1924d: 230 (diagnosis, catalogue); Smith, M.R. 1948: 239 (U.S.A. species key); Creighton, 1950a: 246 (North America review); Chapman & Capco, 1951: 117 (Asia checklist); Bernard, 1967: 226 (diagnosis); Smith, D.R. 1979: 1399 (North America catalogue); Taylor & Brown, D.R. 1985: 71 (Australia catalogue); Taylor, 1987a: 47 (Australia checklist); Morisita, Kubota, et al. 1992: 54 (Japan species key); Bolton, 1994: 105 (synoptic classification); Bolton, 1995a: 1050 (census); Bolton, 1995b: 274 (catalogue); Terayama, 1996: 31 (Japan species key); Lin & Wu, 1998: 89 (Taiwan species key); Rigato, 1999: 83 (West Palaearctic species key); Shattuck, 1999: 147 (Australia synopsis); Zhou, 2001b: 138 (China, Guangxi species key); Radchenko, 2005b: 192 (North Korea species key); Zhou, Huang & Ma, 2008: 285 (China species key); Shattuck, 2009: 3 (Australia species key); Terayama, 2009: 187 (Taiwan species key).

Amblyoponinae Amblyoponini Amblyopone TRUE TRUE FALSE

Extant: 72 species

Fossil: 2 species

Amblyopone Erichson, 1842: 260. Type-species: Amblyopone australis, by monotypy.

Taxonomic history

Amblyopone in Poneridae: Smith, F. 1858b: 108; Smith, F. 1871a: 324.

Amblyopone in Ponerinae: Mayr, 1862: 714 (in key) [Poneridae]; Mayr, 1865: 16 [Poneridae]; Emery & Forel, 1879a: 455 [Poneridae]; Dalla Torre, 1893: 13.

Amblyopone in Amblyoponinae: Forel, 1893a: 162; Clark, 1951: 15 (in key).

Amblyopone in Pachycondylinae, Amblyoponini: Ashmead, 1905b: 382.

Amblyopone in Ponerinae, Amblyoponini: Emery, 1895c: 261; Emery, 1895j: 766; Emery, 1901a: 34; Wheeler, W.M. 1910g: 134; Emery, 1911d: 25; Forel, 1917: 235; Wheeler, W.M. 1922a: 641; Donisthorpe, 1943f: 621; Brown, 1953c: 11; all subsequent authors to the following.

Amblyopone in Amblyoponinae, Amblyoponini: Bolton, 2003: 42, 154.

Junior synonyms of AMBLYOPONE

Stigmatomma Roger, 1859: 250. Type-species: Stigmatomma denticulatum, by subsequent designation of Bingham, 1903: 36.

Taxonomic history

Stigmatomma in Ponerinae: Mayr, 1861: 53 [Poneridae]; Mayr, 1862: 714 [Poneridae]; Mayr, 1865: 16 [Poneridae]; Dalla Torre, 1893: 14.

Stigmatomma in Amblyoponinae: Forel, 1893a: 162; Forel, 1895b: 110 [Amblyoponeridae]

Stigmatomma in Pachycondylinae, Amblyoponini: Ashmead, 1905b: 283.

Stigmatomma in Ponerinae, Amblyoponini: Emery, 1895j: 766; Emery, 1901a: 34; Wheeler, W.M. 1910g: 134; Emery, 1911d: 23; Forel, 1917: 235; Wheeler, W.M. 1922a: 641.

Stigmatomma as subgenus of Amblyopone: Forel, 1900c: 55; Clark, 1934b: 27; Brown, 1949c: 87.

Stigmatomma as genus: Roger, 1859: 250; Dalla Torre, 1893: 14; Emery, 1895j: 766; Bingham, 1903: 36; Emery, 1911d: 23; Forel, 1917: 235; Wheeler, W.M. 1922a: 641; Borgmeier, 1923: 52; Donisthorpe, 1943g: 728; Creighton, 1950a: 31; Kusnezov, 1956: 12.

Stigmatomma as junior synonym of Amblyopone: Emery & Forel, 1879a: 455; Mayr, 1887: 546; Brown, 1960a: 155; all subsequent authors.

Arotropus Provancher, 1881a: 205. Type-species: Arotropus binodosus (junior synonym of Typhlopone pallipes), by monotypy.

Taxonomic history

Arotropus as junior synonym of Amblyopone: Provancher, 1887: 240; Brown, 1960a: 155.

Amblyopopone Dalla Torre, 1893: 13, unjustified emendation of Amblyopone.

Taxonomic history

Amblyopopone as junior synonym of Amblyopone: Forel, 1893a: 166; Emery, 1911d: 25.

Amblyopopona Schulz, W.A. 1906: 154, unjustified emendation of Amblyopone.

Taxonomic history

Amblyopopona as junior synonym of Amblyopone: Bolton, 1994: 164.

Xymmer Santschi, 1914d: 311 [as subgenus of Stigmatomma]. Type-species: Stigmatomma (Xymmer) muticum, by monotypy.

Taxonomic history

Xymmer as subgenus of Stigmatomma: Santschi, 1914d: 311; Forel, 1917: 235; Emery, 1919c: 106; Donisthorpe, 1943g: 737.

Xymmer as genus: Wheeler, W.M. 1922a: 641 (in key).

Xymmer as subgenus of Amblyopone: Clark, 1934b: 27.

Xymmer as junior synonym of Stigmatomma: Brown, 1949c: 87.

Xymmer as junior synonym of Amblyopone: Brown, 1960a: 155.

Fulakora Mann, 1919: 279 [as subgenus of Stigmatomma]. Type-species: Stigmatomma (Fulakora) celata, by original designation.

Taxonomic history

Fulakora as junior synonym of Stigmatomma: Brown, 1949c: 88.

Fulakora as junior synonym of Amblyopone: Brown, 1960a: 155.

Neoamblyopone Wheeler, W.M. 1927c: 1 [as subgenus of Amblyopone]. Type-species: Amblyopone (Neoamblyopone) clarki, by monotypy.

Taxonomic history

Neoamblyopone as junior synonym of Amblyopone: Brown, 1949c: 87; Brown, 1960a: 155.

Protamblyopone Wheeler, W.M. 1927c: 1 [as subgenus of Amblyopone]. Type-species: Amblyopone (Protamblyopone) aberrans, by monotypy.

Taxonomic history

Protamblyopone as junior synonym of Amblyopone: Brown, 1949c: 87; Brown, 1960a: 155.

Lithomyrmex Clark, 1928a: 30. Type-species: Lithomyrmex glauerti, by original designation.

Taxonomic history

Lithomyrmex in Ponerinae, Amblyoponini: Clark, 1928a: 30.

Lithomyrmex as junior synonym of Amblyopone: Brown, 1960a: 156.

Ericapelta Kusnezov, 1955: 273. Type-species: Ericapelta egregia, by monotypy.

Taxonomic history

Ericapelta as junior synonym of Amblyopone: Brown, 1960a: 156.

Genus Amblyopone references

Roger, 1863b: 20 (catalogue); Mayr, 1863: 394, 454 (catalogue); Mayr, 1865: 16 (Amblyopone, Stigmatomma diagnoses); André, 1882c: 233 (Europe & Algeria species key); Cresson, 1887: 259 (U.S.A. catalogue); Dalla Torre, 1893: 13, 14 (Amblyopone, Stigmatomma catalogues); Forel, 1900c: 55 (India & Sri Lanka species key); Bingham, 1903: 37 (India, Sri Lanka & Burma Stigmatomma species key); Emery, 1911d: 23 (Stigmatomma diagnosis, catalogue); Emery, 1911d: 25 (diagnosis, catalogue); Santschi, 1915a: 55 (Mediterranean species key); Emery, 1916b: 100 (Italy species key); Bondroit, 1918: 80 (France & Belgium species key); Wheeler, W.M. 1922a: 758 (Afrotropical catalogue); Wheeler, W.M. 1927c: 3 (Austral & New Guinea species key); Chapman & Capco, 1951: 22, 24 (Asia Amblyopone, Stigmatomma checklists); Brown, 1958h: 11 (New Zealand species); Brown, 1960a: 155, 191 (diagnosis & review of genus, New World species key); Kempf, 1972a: 19 (Neotropical catalogue); Baroni Urbani, 1978a: 41 (Mediterranean species key); Smith, D.R. 1979: 1334 (North America catalogue); Taylor, 1979: 824 (Melanesia species key); Taylor & Brown, D.R. 1985: 18 (Australia catalogue); Taylor, 1987a: 6 (Australia, New Caledonia & New Zealand checklist); Agosti & Collingwood, 1987b: 264 (Balkans species key); Dlussky & Fedoseeva, 1988: 78 (synoptic classification); Morisita, Kubota, et al. 1989: 8 (Japan species key); Terayama, 1989b: 345 (Taiwan species key); Tinaut, 1990d: 189 (Iberian Peninsula species); Dlussky, Soyunov & Zabelin, 1990: 173 (Turkmenistan species key); Lattke, 1991c: 6 (New World species key); Atanasov & Dlussky, 1992: 61 (Bulgaria species key); Bolton, 1995a: 1047 (census); Bolton, 1995b: 61 (catalogue); Onoyama, 1999: 190 (Japan species key); Shattuck, 1999: 179 (Australia synopsis); Xu, 2001d: 552 (East & South Asia species key); Lacau & Delabie, 2002: 40 (Neotropical species key); Xu, 2006: 152 (China species key); Arias-Penna, 2008a: 43 (Neotropical species key); Terayama, 2009: 93 (Taiwan species key); Heterick, 2009: 131 (south-western Australia species key).

Ponerinae Ponerini Leptogenys TRUE TRUE FALSE

Extant: 216 species, 31 subspecies

Fossil: 1 species

Leptogenys Roger, 1861a: 41. Type-species: Leptogenys falcigera, by subsequent designation of Bingham, 1903: 52.

Taxonomic history

Leptogenys in Ponerinae: Dalla Torre, 1893: 46; Mayr, 1862: 714 (in key) [Poneridae]; Mayr, 1865: 14 [Poneridae].

Leptogenys in Ponerinae, Leptogenyini: Forel, 1893a: 162; Forel, 1899c: 17; Ashmead, 1905b: 382; Emery, 1911d: 97; Arnold, 1915: 89; Forel, 1917: 238; Gallardo, 1918b: 84; Wheeler, W.M. 1922a: 652; Donisthorpe, 1943f: 656; Chapman & Capco, 1951: 31; Dlussky & Fedoseeva, 1988: 78.

Leptogenys in Ponerinae, Ponerini: Emery, 1895j: 767; Forel, 1900d: 304; Wheeler, W.M. 1910g: 135; Brown, 1963: 3; Kempf, 1972a: 129; Jaffe, 1993: 8; Bolton, 1994: 164; Bolton, 2003: 163.

Junior synonyms of LEPTOGENYS

Lobopelta Mayr, 1862: 714 (diagnosis in key), 733. Type-species: Ponera diminuta, by subsequent designation of Bingham, 1903: 54.

Taxonomic history

Lobopelta in Ponerinae: Mayr, 1862: 714 (in key) [Poneridae]; Mayr, 1865: 15 [Poneridae]; Dalla Torre, 1893: 43.

Lobopelta in Ponerinae, Leptogenyini: Forel, 1893a: 162; Ashmead, 1905b: 382; Emery, 1911d: 101; Wheeler, W.M. 1922a: 653; Donisthorpe, 1943f: 658.

Lobopelta as genus: Mayr, 1862: 733; Cresson, 1887: 258; Dalla Torre, 1893: 43; Bingham, 1903: 54.

Lobopelta as subgenus of Leptogenys: Forel, 1892k: 520; Forel, 1899c: 18; Forel, 1900d: 304; Wheeler, W.M. 1910g: 135; Emery, 1911d: 101; Arnold, 1915: 95; Forel, 1917: 238; Gallardo, 1918b: 85; Wheeler, W.M. 1922a: 653; Wheeler, W.M. 1923d: 1; Borgmeier, 1923: 75; Creighton, 1950a: 50; Kempf, 1972a: 131; Smith, D.R. 1979: 1343 (anachronism).

Lobopelta as junior synonym of Leptogenys: Emery, 1896e: 177 (footnote); Brown, 1973b: 181 [provisional]; Bolton, 1975a: 240; Bolton, 2003: 163.

Prionogenys Emery, 1895g: 348. Type-species: Prionogenys podenzanai, by monotypy.

Taxonomic history

Prionogenys in Ponerinae, Ponerini: Emery, 1895j: 767; Wheeler, W.M. 1910g: 135;

Prionogenys in Ponerinae, Leptogenyini: Ashmead, 1905b: 382; Emery, 1911d: 106; Forel, 1917: 238; Wheeler, W.M. 1922a: 652; Donisthorpe, 1943f: 685; Dlussky & Fedoseeva, 1988: 78.

Prionogenys as junior synonym of Leptogenys: Taylor, 1988: 33.

Machaerogenys Emery, 1911d: 100 [as subgenus of Leptogenys]. Type-species: Leptogenys truncatirostris, by original designation.

Taxonomic history

Machaerogenys as subgenus of Leptogenys: Emery, 1911d: 100; Forel, 1917: 238; Wheeler, W.M. 1922a: 653; subsequent authors to the following.

Machaerogenys as junior synonym of Leptogenys: Brown, 1973b: 181 [provisional]; Bolton, 1975a: 240.

Odontopelta Emery, 1911d: 101 [as subgenus of Leptogenys]. Type-species: Leptogenys turneri, by monotypy.

Taxonomic history

Odontopelta as subgenus of Leptogenys: Emery, 1911d: 101; Forel, 1917: 238; Wheeler, W.M. 1922a: 653; Donisthorpe, 1943f: 677.

Odontopelta as junior synonym of Leptogenys: Brown, 1973b: 183 [provisional]; Snelling, R.R. 1981: 390; Taylor & Brown, D.R. 1985: 32.

Dorylozelus Forel, 1915b: 24. Type-species: Dorylozelus mjobergi (junior secondary homonym in Leptogenys, replaced by Leptogenys tricosa), by monotypy.

Taxonomic history

Dorylozelus in Ponerinae, Ponerini: Forel, 1917: 238.

Dorylozelus in Ponerinae, Dorylozelini: Wheeler, W.M. 1922a: 646; Donisthorpe, 1943f: 640.

Dorylozelus incertae sedis in Ponerinae, Amblyoponini: Brown, 1960a: 181.

Dorylozelus as junior synonym of Leptogenys: Taylor, 1969: 132; Bolton, 1975a: 240.

Microbolbos Donisthorpe, 1948f: 170. Type-species: Microbolbos testaceus, by original designation.

Taxonomic history

Microbolbos as junior synonym of Leptogenys: Wilson, 1955c: 136; Bolton, 1975a: 240.

Genus Leptogenys references

Roger, 1863b: 19 (Lobopelta, Leptogenys catalogues); Mayr, 1863: 426, 427 (Leptogenys, Lobopelta catalogues); Mayr, 1865: 14, 15 (Leptogenys, Lobopelta diagnoses); Mayr, 1867a: 89 (diagnosis); Mayr, 1879: 664 (Lobopelta species key); Cresson, 1887: 258 (U.S.A. catalogue); Dalla Torre, 1893: 43, 46 (Lobopelta, Leptogenys catalogues); Forel, 1900d: 304 (India & Sri Lanka species key); Bingham, 1903: 52, 55 (India, Sri Lanka & Burma Leptogenys, Lobopelta species keys); Emery, 1911d: 97 (diagnosis, subgenera key, catalogue); Arnold, 1915: 89, 95 (diagnosis, South Africa L. (Leptogenys) & L. (Lobopelta) species keys); Mann, 1919: 298 (Papuasia L. (Leptogenys) species key); Mann, 1921: 426 (Fiji L. (Lobopelta) species key); Wheeler, W.M. 1922a: 93, 786, 1010 (diagnosis, Afrotropical, Malagasy catalogues); Wheeler, W.M. 1923d: 14 (New World L. (Lobopelta) species key); Creighton, 1950a: 51 (North America L. (Lobopelta) species key); Chapman & Capco, 1951: 31 (Asia checklist); Wilson, 1958a: 113 (Melanesia & Moluccas species revision, key); Wilson, 1958a: 136 (New Caledonia species key); Kempf, 1972a: 129 (Neotropical catalogue); Alayo, 1974: 8 (Cuba species key); Bolton, 1975a: 239 (diagnosis, review of genus); Bolton, 1975a: 244 (Afrotropical species revision, key); Bolton, 1975a: 293 (Malagasy species key); Smith, D.R. 1979: 1343 (North America catalogue); Taylor & Brown, D.R. 1985: 32, 39 (Australia catalogue); Taylor, 1987a: 34, 64 (Australia, New Caledonia checklists); Trager & Johnson, 1988: 30 (U.S.A. species key); Brandão, 1991: 349 (Neotropical catalogue); Bolton, 1995a: 1050 (census); Bolton, 1995b: 229 (catalogue); Wu, J. & Wang, 1995: 41 (China species key); Shattuck, 1999: 189 (Australia synopsis); Xu, 2000a: 118 (China species key); Zhou, 2001b: 39 (China, Guangxi species key); Bolton, 2003: 163 (synopsis); Lattke, in Fernández, 2008: 145 (Colombia species, review); Terayama, 2009: 115 (Taiwan species key); Heterick, 2009: 133 (south-western Australia species key).

Ponerinae Ponerini Odontomachus TRUE TRUE FALSE

Extant: 66 species

Fossil: 2 species

Odontomachus Latreille, 1804: 179. Type-species: Formica haematoda, by monotypy.

Taxonomic history

Odontomachus in Ponérites: Lepeletier de Saint-Fargeau, 1835: 185.

Odontomachus in Poneridae: Smith, F. 1857a: 64; Smith, F. 1858b: 76.

Odontomachus in Formicidae, Odontomachidae: Mayr, 1862: 711; Mayr, 1865: 11; Smith, F. 1871a: 319.

Odontomachus in Odontomachidae: Ashmead, 1905b: 382.

Odontomachus in Ponerinae: Dalla Torre, 1893: 49.

Odontomachus in Ponerinae, Odontomachini: Forel, 1893a: 163; Emery, 1895j: 768; Forel, 1895b: 118; Forel, 1899c: 19; Wheeler, W.M. 1910g: 136; Emery, 1911d: 111; Arnold, 1915: 108; Forel, 1917: 238; Gallardo, 1918b: 93; Wheeler, W.M. 1922a: 653; Donisthorpe, 1943f: 676; Chapman & Capco, 1951: 42; Kempf, 1972a: 170; Wheeler, G.C. & Wheeler, J. 1985: 256; Dlussky & Fedoseeva, 1988: 78; Jaffe, 1993: 8.

Odontomachus in Ponerinae, Ponerini: Brown, 1976a: 70 [subtribe Odontomachiti]; Bolton, 1994: 164.

Junior synonyms of ODONTOMACHUS

Pedetes Bernstein, 1861: 7. Type-species: Pedetes macrorhynchus, by monotypy.

Taxonomic history

Pedetes as junior synonym of Odontomachus: Dalla Torre, 1893: 51; all subsequent authors.

Champsomyrmex Emery, 1892d: 558 (footnote). Type-species: Odontomachus coquereli, by monotypy.

Taxonomic history

Champsomyrmex in Ponerinae: Dalla Torre, 1893: 49.

Champsomyrmex in Odontomachidae: Ashmead, 1905b: 382.

Champsomyrmex in Ponerinae, Odontomachini: Emery, 1895j: 768; Wheeler, W.M. 1910g: 136; Emery, 1911d: 111; Wheeler, W.M. 1922a: 653; Donisthorpe, 1943f: 632.

Champsomyrmex as subgenus of Odontomachus: Forel, 1893a: 163 (misspelled as Thempsomyrmex); Forel, 1917: 238.

Champsomyrmex as genus: Emery, 1892d: 558 (footnote); Dalla Torre, 1893: 49; Emery, 1911d: 111; Wheeler, W.M. 1922a: 653.

Champsomyrmex as junior synonym of Odontomachus: Brown, 1973b: 179 [provisional]; Brown, 1976a: 96.

Myrtoteras Matsumura, 1912: 191. Type-species: Myrtoteras kuroiwae, by monotypy.

Taxonomic history

Myrtoteras in Ponerinae, Odontomachini: Donisthorpe, 1943f: 672.

Myrtoteras as junior synonym of Odontomachus: Brown, 1973b: 182 [provisional]; Brown, 1976a: 96.

Genus Odontomachus references

Roger, 1863b: 21 (catalogue); Mayr, 1863: 436 (catalogue); Mayr, 1865: 11 (diagnosis); Cresson, 1887: 258 (U.S.A. catalogue); Emery, 1892d: 559 (all species key); Dalla Torre, 1893: 49 (catalogue); Forel, 1900c: 57 (India & Sri Lanka species key); Bingham, 1903: 47 (India, Sri Lanka & Burma species key); Emery, 1911d: 111 (diagnosis, catalogue); Gallardo, 1918b: 95 (Argentina species key); Wheeler, W.M. 1922a: 99, 792, 1013 (diagnosis, Afrotropical, Malagasy catalogues); Creighton, 1950a: 55 (North America species key); Chapman & Capco, 1951: 42 (Asia checklist); Wilson, 1959a: 485 (Melanesia & Moluccas species revision, key); Kempf, 1962b: 17 (Neotropical species key); Kempf, 1972a: 170 (Neotropical catalogue); Brown, 1976a: 111, 114, 117 (New World species, Malesian & Austral species, Afrotropical & Malagasy species keys); Smith, D.R. 1979: 1344 (North America catalogue); Deyrup, Trager & Carlin, 1985: 188 (Southeastern U.S.A. species, review); Brown, 1978b: 281 (supplement to Brown, 1976a); Taylor & Brown, D.R. 1985: 36 (Australia catalogue); Taylor, 1987a: 48 (Australia, New Caledonia checklist); Wang, M. 1993: 220 (China species key); De Andrade, 1994: 24 (†Dominican Amber species key); Bolton, 1995a: 1051 (census); Bolton, 1995b: 294 (catalogue); Gronenberg, 1995: 391 (mandible morphology); Shattuck, 1999: 194 (Australia synopsis); Zhou, 2001b: 25 (China, Guangxi species key); Bolton, 2003: 164 (synopsis); Deyrup & Cover, 2004b: 141 (U.S.A. species key); Yoshimura, Onoyama & Ogata, 2007: 89 (Japan species key); Fisher & Smith, 2008: 13 (Malagasy species revision, key); Rodríguez, in Fernández, 2008: 151 (Neotropical species key); Sorger & Zettel, 2011: 143 (Philippines species key).

Ponerinae Ponerini Ponera TRUE TRUE FALSE

Extant: 52 species

Fossil: 5 species

Ponera Latreille, 1804: 179. Type-species: Formica coarctata, by subsequent designation of Westwood, 1840a: 83 (see discussion in Taylor, 1967a: 6).

Taxonomic history

Ponera in Ponérites: Lepeletier de Saint-Fargeau, 1835: 188.

Ponera in Poneridae: Smith, F. 1858b: 83; Smith, F. 1871a: 320.

Ponera in Ponerinae: Mayr, 1855: 386 [Poneridae]; Smith, F. 1857a: 65 [Poneridae]; Mayr, 1861: 54 [Poneridae]; Mayr, 1862: 713 [Poneridae]; Mayr, 1865: 13 [Poneridae]; Emery & Forel, 1879a: 455 [Poneridae]; Dalla Torre, 1893: 37.

Ponera in Ponerinae, Ponerini: Emery, 1895j: 767; Forel, 1899c: 15; Ashmead, 1905b: 382; Wheeler, W.M. 1910g: 135; Emery, 1911d: 88 [subtribe Ponerini]; Wheeler, W.M. 1915h: 38; Arnold, 1915: 77; Forel, 1917: 238; Wheeler, W.M. 1922a: 650; all subsequent authors.

Junior synonyms of PONERA

Pseudocryptopone Wheeler, W.M. 1933g: 12. Type-species: Cryptopone tenuis, by original designation.

Taxonomic history

Pseudocryptopone in Ponerinae, Ponerini: Donisthorpe, 1943g: 722.

Pseudocryptopone as junior synonym of Ponera: Wilson, 1957b: 356.

Selenopone Wheeler, W.M. 1933g: 19. Type-species: Ponera selenophora, by original designation.

Taxonomic history

Selenopone in Ponerinae, Ponerini: Donisthorpe, 1943g: 725.

Selenopone as junior synonym of Ponera: Wilson, 1957b: 356.

Pteroponera Bernard, 1950a: 3. Type-species: Pteroponera sysphinctoides, by monotypy.

Taxonomic history

Pteroponera as junior synonym of Ponera: Brown, 1973b: 184 [provisional]; Brown, in Bolton, 1994: 164.

Genus Ponera references

[References before Taylor, 1967a refer to both Ponera and Hypoponera.]

Mayr, 1855: 386 (diagnosis); Smith, F. 1858b: 83 (diagnosis); Mayr, 1862: 721 (diagnosis); Roger, 1863b: 16 (catalogue); Mayr, 1863: 447 (catalogue); Mayr, 1865: 13 (diagnosis); Mayr, 1867a: 85 (diagnosis); Forel, 1874: 64 (Switzerland species key); Mayr, 1879: 661 (all species key); André, 1882c: 239 (Europe & Algeria species key); Cresson, 1887: 258 (U.S.A. catalogue); Dalla Torre, 1893: 37 (catalogue); Emery, 1895b: 60 (Mediterranean species key); Emery, 1896g: 53 (New World species key); Emery, 1900c: 316 (Australia, Papuasia & Oceania species key); Forel, 1900d: 322 (India & Sri Lanka species key); Bingham, 1903: 89 (India, Sri Lanka & Burma species key); Emery, 1909c: 367 (Palaearctic species key); Emery, 1911d: 88 (diagnosis, catalogue); Arnold, 1915: 76 (diagnosis, South Africa species key); Emery, 1916b: 107 (Italy species key); Bondroit, 1918: 81 (France & Belgium species key); Gallardo, 1918b: 73 (Argentina species key); Mann, 1921: 419 (Fiji species key); Wheeler, W.M. 1922a: 780, 1009 (Afrotropical, Malagasy catalogues); Menozzi, 1931b: 266 (Costa Rica species key); Wheeler, W.M. 1933g: 13 (Pseudocryptopone species key); Smith, M.R. 1936: 420 (North America species key); Stitz, 1939: 58 (Germany species key); Creighton, 1950a: 47 (North America species key); Chapman & Capco, 1951: 68, 73, 74 (Asia Ponera, Pseudocryptopone, Selenopone checklists); Bernard, 1953b: 197 (West Africa species key); Wilson, 1957b: 359, 376 (P. tenuis, P. selenophora groups revisions, keys); Wilson, 1958d: 323, 343 (Melanesia & Moluccas species, Fiji species revisions, keys); Brown, 1958h: 22 (New Zealand species); Taylor, 1960: 180 (P. leae complex, key); Taylor, 1964: 139 (†fossil species, review); Taylor, 1967a: 18 (diagnosis, all species revision, key); Bernard, 1967: 83 (diagnosis, Western Europe species key); Smith, D.R. 1979: 1341 (North America catalogue); Taylor & Brown, D.R. 1985: 38 (Australia catalogue); Taylor, 1987a: 63 (Australia, New Caledonia & New Zealand, checklist); Morisita, Kubota, et al. 1989: 22 (Japan species key); Perrault, 1993: 337 (addition to Taylor, 1967a key); Bolton, 1994: 164 (synoptic classification); Bolton, 1995a: 1052 (census); Bolton, 1995b: 360 (catalogue); Terayama, 1996: 14 (Japan species key); Kim, Kim & Kim, 1998: 149 (Korea species key); Shattuck, 1999: 199 (Australia synopsis); Xu, 2001a: 52 and Xu, 2001c: 218 (China species keys); Zhou, 2001b: 37 (China, Guangxi species key); Bolton, 2003: 170 (synopsis); Yoshimura, Hosoishi, et al. 2009: 200 (Japan species key); Terayama, 2009: 109 (Taiwan species key).

Proceratiinae Proceratiini Discothyrea TRUE TRUE FALSE

Extant: 32 species

Fossil: 2 species

Discothyrea Roger, 1863a: 176. Type-species: Discothyrea testacea, by monotypy.

Taxonomic history

Discothyrea in Ponerinae: Mayr, 1865: 13 [Poneridae]; Dalla Torre, 1893: 18.

Discothyrea in Dorylinae, Proceratiini: Emery, 1895j: 765.

Discothyrea in Ponerinae, Proceratiini: Ashmead, 1905b: 382; Wheeler, W.M. 1910g: 136; Emery, 1911d: 51; Forel, 1917: 236; Wheeler, W.M. 1922a: 645; Donisthorpe, 1943f: 640; Chapman & Capco, 1951: 76; Kusnezov, 1964: 51; Wheeler, G.C. & Wheeler, J. 1985: 256; Jaffe, 1993: 8; Lattke, 1994: 112.

Discothyrea in Ponerinae, Ectatommini: Brown, 1958g: 248; Kempf, 1972a: 97; Smith, D.R. 1979: 1339; Dlussky & Fedoseeva, 1988: 78; Hölldobler & Wilson, 1990: 10; Bolton, 1994: 164.

Discothyrea in Proceratiinae, Proceratiini: Bolton, 2003: 49, 179.

Junior synonyms of DISCOTHYREA

Prodiscothyrea Wheeler, W.M. 1916i: 33. Type-species: Prodiscothyrea velutina, by monotypy.

Taxonomic history

Prodiscothyrea in Ponerinae, Proceratiini: Wheeler, W.M. 1916i: 36; Wheeler, W.M. 1922a: 645; Donisthorpe, 1943f: 687; Chapman & Capco, 1951: 77.

Prodiscothyrea as junior synonym of Discothyrea: Brown, 1958g: 248.

Pseudosysphincta Arnold, 1916: 161. Type-species: Pseudosysphincta poweri, by original designation.

Taxonomic history

Pseudosysphincta in Ponerinae, Proceratiini: Forel, 1917: 236; Wheeler, W.M. 1922a: 645; Donisthorpe, 1943g: 723.

Pseudosysphincta as junior synonym of Discothyrea: Brown, 1958g: 248.

[Pseudosphincta Wheeler, W.M. 1922a: 645, incorrect subsequent spelling.]

Genus Discothyrea references

Roger, 1863b: 21 (catalogue); Mayr, 1865: 13 (diagnosis); Cresson, 1887: 259 (U.S.A., catalogue); Dalla Torre, 1893: 18 (catalogue); Emery, 1911d: 51 (diagnosis, catalogue); Arnold, 1916: 159 (diagnosis); Wheeler, W.M. 1922a: 761, 762 (Afrotropical Discothyrea, Pseudosysphincta catalogues); Weber, 1939a: 99 (Neotropical species key); Chapman & Capco, 1951: 76, 77 (Asia Discothyrea, Prodiscothyrea checklists); Brown, 1958g: 248 (review of genus); Brown, 1958h: 19 (New Zealand species); Kempf, 1972a: 97 (Neotropical catalogue); Smith, D.R. 1979: 1339 (North America catalogue); Taylor & Brown, D.R. 1985: 29 (Australia catalogue); Taylor, 1987a: 24 (Australia, New Caledonia & New Zealand checklists); Morisita, Kubota, et al. 1989: 16 (Japan species key); Lattke, 1994: 113 (Neotropical species); Bolton, 1995a: 1049 (census); Bolton, 1995b: 171 (catalogue); Shattuck, 1999: 184 (Australia synopsis); Bolton, 2003: 179 (synopsis); Sosa-Calvo & Longino, 2008: 223 (Colombia species key); Yoshimura & Fisher, 2009: 9 (Malagasy males diagnosis); Terayama, 2009: 98 (Taiwan species key); Heterick, 2009: 143 (south-western Australia species key).

Proceratiinae Proceratiini Proceratium TRUE TRUE FALSE

Extant: 78 species

Fossil: 5 species

Proceratium Roger, 1863a: 171. Type-species: Proceratium silaceum, by monotypy.

Taxonomic history

Proceratium in Ponerinae: Mayr, 1865: 12 [Poneridae]; Dalla Torre, 1893: 18.

Proceratium in Ponerinae, Ponerini: Forel, 1895b: 111.

Proceratium in Dorylinae, Proceratiini: Emery, 1895j: 765.

Proceratium in Ponerinae, Ectatommini: Brown, 1958g: 241; Kempf, 1972a: 211; Smith, D.R. 1979: 1338; Dlussky & Fedoseeva, 1988: 79; Hölldobler & Wilson, 1990: 10; Bolton, 1994: 164.

Proceratium in Ponerinae, Proceratiini: Ashmead, 1905b: 382; Wheeler, W.M. 1910g: 136; Emery, 1911d: 50; Forel, 1917: 236; Wheeler, W.M. 1922a: 645; Wheeler, G.C. & Wheeler, J. 1985: 256; Jaffe, 1993: 8; Lattke, 1994: 112.

Proceratium in Proceratiinae, Proceratiini: Bolton, 2003: 49, 179.

Junior synonym of PROCERATIUM

Sysphingta Roger, 1863a: 175. Type-species: Sysphingta micrommata, by monotypy.

Taxonomic history

Sysphingta in Ponerinae: Mayr, 1865: 12 [Poneridae]; Dalla Torre, 1893: 18.

Sysphingta in Dorylinae, Proceratiini: Emery, 1895j: 765.

Sysphingta in Ponerinae, Proceratiini: Ashmead, 1905b: 382; Wheeler, W.M. 1910g: 136; Emery, 1911d: 50; Arnold, 1915: 34; Forel, 1917: 236; Wheeler, W.M. 1922a: 645; Donisthorpe, 1943g: 730.

Sysphingta as genus: Emery, 1895c: 262; Wheeler, W.M. 1910g: 136; Forel, 1917: 236; Wheeler, W.M. 1922a: 645; Creighton, 1950a: 40.

Sysphingta as junior synonym of Proceratium: Mayr, 1886d: 437; Dalla Torre, 1893: 18; Brown, in Borgmeier, 1957: 118; Brown, 1958g: 241; all subsequent authors.

[Sysphincta Mayr, 1865: 12 (and most subsequent authors), incorrect subsequent spelling.]

Genus Proceratium references

Roger, 1863b: 16, 21 (catalogue); Mayr, 1863: 451, 455 (catalogue); Mayr, 1865: 12 (Sysphingta, Proceratium diagnoses); Cresson, 1887: 258 (U.S.A. catalogue); Dalla Torre, 1893: 18 (catalogue); Emery, 1911d: 50 (diagnosis, catalogue); Emery, 1911d: 50 (Sysphingta diagnosis, catalogue); Arnold, 1915: 34 (Sysphingta diagnosis); Emery, 1916b: 103 (Italy species key); Wheeler, W.M. 1922a: 761 (Afrotropical catalogue); Kratochvíl, Novák & Snoflák, 1944: 58, 88 (West Palaearctic species key); Creighton, 1950a: 39, 41 (North America Proceratium, Sysphingta species keys); Chapman & Capco, 1951: 77 (Asia Proceratium, Sysphingta checklists); Brown, 1958g: 241 (review of genus); Bernard, 1967: 81 (diagnosis); Snelling, R.R. 1967: 8 (New World species key); Kempf, 1972a: 211 (Neotropical catalogue); Brown, 1974a: 81 (species notes); Baroni Urbani, 1977d: 91 (European species, synopsis); Smith, D.R. 1979: 1338 (North America catalogue); Brown, 1980b: 342 (New World species key); Terron, 1981: 102 (Afrotropical species notes); Taylor & Brown, D.R. 1985: 39 (Australia catalogue); Taylor, 1987a: 65 (Australia checklist); Agosti & Collingwood, 1987b: 264 (Balkans species key); Ward, 1988: 116 (New World species key); Morisita, Kubota, et al. 1989: 14 (Japan species key); Brandão, 1991: 373 (Neotropical catalogue); Lattke, 1994: 113 (phylogeny); Bolton, 1995a: 1052 (census); Bolton, 1995b: 366 (catalogue); Shattuck, 1999: 203 (Australia synopsis); Xu, 2000c: 434 (China species key); Onoyama & Yoshimura, 2002: 31 (Japan species key); Bolton, 2003: 179 (synopsis); Baroni Urbani & De Andrade, 2003b: 40 (diagnosis, all species revision, keys; Palaearctic species key: 445; Indomalayan species key: 447; Oceanean species key: 449; Afrotropical species key: 451; Nearctic and Neotropical species key: 452; world species key: 456); Xu, 2006: 153 (China species key); Sosa-Calvo & Longino, 2008: 228 (Neotropical species key); Yoshimura & Fisher, 2009: 13 (Malagasy males diagnosis); Terayama, 2009: 97 (Taiwan species key).

Cerapachyinae Acanthostichini Acanthostichus texanus TRUE TRUE FALSE

texanus. Acanthostichus texanus Forel, 1904d: 168 (q.) U.S.A. Combination in Acanthostichus (Ctenopyga): Emery, 1911d: 14; in Ctenopyga: Brown, 1975: 42; in Acanthostichus: Mackay, 1996: 168. Senior synonym of townsendi: Smith, M.R. 1955a: 49.

Myrmicinae Attini Acromyrmex versicolor chisosensis TRUE TRUE FALSE

chisosensis. Atta (Moellerius) versicolor subsp. chisosensis Wheeler, W.M. 1907c: 705 (w.) U.S.A. Combination in Acromyrmex: Emery, 1924d: 351; in Acromyrmex (Moellerius): Fowler, 1988: 291.

Formicinae Lasiini Acropyga epedana TRUE TRUE FALSE

epedana. Acropyga epedana Snelling, R.R. 1973a: 7, fig. 1 (w.) U.S.A. LaPolla, Cover & Mueller, 2002: 369 (q.m.). See also: LaPolla, 2004a: 46.

Amblyoponinae Amblyoponini Amblyopone oregonensis TRUE TRUE FALSE

oregonensis. Stigmatomma pallipes subsp. oregonense Wheeler, W.M. 1915b: 389 (w.q.) U.S.A. Combination in Amblyopone: Brown, 1960a: 169. Raised to species: Brown, 1960a: 169. See also: Ward, 1988: 105.

Amblyoponinae Amblyoponini Amblyopone pallipes TRUE TRUE FALSE

pallipes. Typhlopone pallipes Haldeman, 1844: 54 (w.) U.S.A. Emery, 1895c: 261 (q.m.); Wheeler G.C. & Wheeler, J. 1952a: 114 (l.). Combination in Amblyopone: Haldeman, 1849a: 201 (footnote); in Stigmatomma: Roger, 1863b: 20; in Amblyopone: Brown, 1960a: 169. [Stigmatomma pallidipes: Dalla Torre, 1893: 14, unjustified emendation.] Senior synonym of serratum: Mayr, 1886d: 439; of binodosus: Emery, 1895c: 261; of arizonense, wheeleri: Creighton, 1940b: 3; of montigena: Brown, 1949c: 84; of subterranea: Brown, 1960a: 169. See also: Whelden, 1958: 1; Traniello, 1982: 65.

Amblyoponinae Amblyoponini Amblyopone trigonignatha TRUE TRUE FALSE

trigonignatha. Amblyopone (Stigmatomma) trigonignatha Brown, 1949c: 81, fig. 1 (w.) U.S.A.

Aneuretinae Aneuretini Aneuretellus deformis TRUE TRUE TRUE

†deformis. †Aneuretellus deformis Dlussky, 1988: 54, fig. 2 (w.q.) RUSSIA (Paleocene).

Myrmicinae Pheidolini Aphaenogaster albisetosa TRUE TRUE FALSE

albisetosa. Aphaenogaster albisetosa Mayr, 1886d: 446 (w.) U.S.A. Wheeler, W.M. & Creighton, 1934: 350 (q.m.); Wheeler, G.C. & Wheeler, J. 1953b: 70 (l.). Combination in Stenamma (Aphaenogaster): Emery, 1895c: 306; in Novomessor: Emery, 1915d: 73; in Aphaenogaster: Brown, 1974b: 47. Senior synonym of minor Enzmann: Brown, 1949a: 49. See also: Hölldobler, Stanton & Engel, 1976: 32; Hölldobler, Stanton & Markl, 1978: 163; Smith, D.R. 1979: 1360; Bolton, 1982: 340.

Myrmicinae Pheidolini Aphaenogaster aquia FALSE FALSE FALSE

aquia. Myrmica (Monomorium) aquia Buckley, 1867: 341 (w.q.) U.S.A. Combination in Aphaenogaster: Mayr, 1886c: 365. Junior synonym of fulva: Mayr, 1886c: 365. Revived from synonymy as subspecies of fulva: Emery, 1895c: 304. Unidentifiable taxon; incertae sedis in Aphaenogaster: Creighton, 1950a: 148; Smith, D.R. 1979: 1363.

Myrmicinae Pheidolini Aphaenogaster ashmeadi TRUE TRUE FALSE

ashmeadi. Stenamma (Aphaenogaster) treatae var. ashmeadi Emery, 1895c: 302 (w.) U.S.A. Combination in Aphaenogaster: Wheeler, W.M. 1913c: 114; in Aphaenogaster (Attomyrma): Emery, 1921f: 60. Raised to species and senior synonym of harnedi: Creighton, 1950a: 142.

Myrmicinae Pheidolini Aphaenogaster boulderensis TRUE TRUE FALSE

1 subspecies

boulderensis. Aphaenogaster (Attomyrma) boulderensis Smith, M.R. 1941: 120 (w.) U.S.A. Current subspecies: nominal plus smithi.

Myrmicinae Pheidolini Aphaenogaster carolinensis TRUE TRUE FALSE

carolinensis. Aphaenogaster texana var. carolinensis Wheeler, W.M. 1915b: 414 (w.q.) U.S.A. Combination in Aphaenogaster (Attomyrma): Emery, 1921f: 60. Subspecies of texana: Creighton, 1950a: 153. Raised to species: Umphrey, 1996: 558 (in key).

Myrmicinae Pheidolini Aphaenogaster donisthorpei TRUE TRUE TRUE

†donisthorpei. †Aphaenogaster donisthorpei Carpenter, 1930: 31, pl. 7, fig. 4 (q.) U.S.A. (Oligocene).

Myrmicinae Pheidolini Aphaenogaster flemingi TRUE TRUE FALSE

flemingi. Aphaenogaster texana subsp. flemingi Smith, M.R. 1928c: 275 (w.) U.S.A. Wheeler, G.C. & Wheeler, J. 1972b: 238 (l.). Combination in Aphaenogaster (Attomyrma): Creighton, 1950a: 143. Raised to species: Creighton, 1950a: 143. Senior synonym of macrospina: Smith, M.R. 1958a: 113.

Myrmicinae Pheidolini Aphaenogaster floridana TRUE TRUE FALSE

floridana. Aphaenogaster (Attomyrma) floridana Smith, M.R. 1941: 118 (w.) U.S.A. Wheeler, G.C. & Wheeler, J. 1960b: 7 (l.).

Myrmicinae Pheidolini Aphaenogaster fulva TRUE TRUE FALSE

1 subspecies

fulva. Aphaenogaster fulva Roger, 1863a: 190 (w.) U.S.A. Mayr, 1886d: 445 (q.m.). Combination in Stenamma (Aphaenogaster): Emery, 1895c: 303; in Aphaenogaster: Wheeler, W.M. 1913c: 114; in Aphaenogaster (Attomyrma): Emery, 1921f: 57. Senior synonym of rubida: Brown, 1949a: 49. Current subspecies: nominal plus azteca. See also: Umphrey, 1966: 557 (key). [See also entry under pusilla.]

Myrmicinae Pheidolini Aphaenogaster huachucana TRUE TRUE FALSE

1 subspecies

huachucana. Aphaenogaster (Attomyrma) huachucana Creighton, 1934: 189 (w.) U.S.A. Creighton, 1952a: 94 (q.m.). Current subspecies: nominal plus crinimera.

Myrmicinae Pheidolini Aphaenogaster januschevi TRUE TRUE FALSE

januschevi. Aphaenogaster januschevi Arnol'di, 1976b: 1020, figs. 1-4 (w.q.m.) RUSSIA.

Myrmicinae Pheidolini Aphaenogaster kurdica TRUE TRUE FALSE

kurdica. Aphaenogaster subterranea var. kurdica Ruzsky, 1905b: 717 (w.) RUSSIA. Combination in Aphaenogaster (Attomyrma): Emery, 1921f: 59. Subspecies of smythiesii: Emery, 1908c: 332. Raised to species: Arnol'di, 1948: 211.

Myrmicinae Pheidolini Aphaenogaster lamellidens TRUE TRUE FALSE

lamellidens. Aphaenogaster lamellidens Mayr, 1886d: 444 (w.q.m.) U.S.A. Taber & Cokendolpher, 1988: 95 (k.). Combination in Stenamma (Aphaenogaster): Emery, 1895c: 302; in Aphaenogaster: Wheeler, W.M. 1913c: 114; in Aphaenogaster (Attomyrma): Emery, 1921f: 58. Senior synonym of nigripes: Creighton, 1950a: 144.

Myrmicinae Pheidolini Aphaenogaster mariae TRUE TRUE FALSE

mariae. Aphaenogaster mariae Forel, 1886b: xli (w.) U.S.A. Combination in Stenamma (Aphaenogaster): Emery, 1895c: 301; in Aphaenogaster (Attomyrma): Emery, 1921f: 58.

Myrmicinae Pheidolini Aphaenogaster mayri TRUE TRUE TRUE

†mayri. †Aphaenogaster mayri Carpenter, 1930: 30, pl. 5, fig. 5; pl. 8, fig. 4; pl. 9, fig. 5; pl. 11, fig. 4 (w.q.m.) U.S.A. (Oligocene).

Myrmicinae Pheidolini Aphaenogaster megommata TRUE TRUE FALSE

megommata. Aphaenogaster (Attomyrma) megommatus Smith, M.R. 1963: 244, fig. 1 (w.) U.S.A. Cole, 1966: 9 (q.m.); Wheeler, G.C. & Wheeler, J. 1972b: 238 (l.).

Myrmicinae Pheidolini Aphaenogaster miamiana TRUE TRUE FALSE

miamiana. Aphaenogaster (Attomyrma) texana var. miamiana Wheeler, W.M. 1932a: 5 (w.q.m.) U.S.A. Raised to species: Creighton, 1950a: 145. See also: Umphrey, 1996: 558 (in key).

Myrmicinae Pheidolini Aphaenogaster nana FALSE FALSE FALSE

nana. Aphaenogaster (Attomyrma) texana subsp. nana Wheeler, W.M. 1932a: 6 (w.) U.S.A. Unidentifiable taxon incertae sedis in Aphaenogaster: Creighton, 1950a: 151; Smith, D.R. 1979: 1364.

Myrmicinae Pheidolini Aphaenogaster obsidiana TRUE TRUE FALSE

obsidiana. Atta obsidiana Mayr, 1861: 67 (diagnosis in key) (w.) RUSSIA. Ruzsky, 1905b: 725 (q.); Karavaiev, 1926b: 98 (m.). Combination in Aphaenogaster: Roger, 1863b: 29; in Stenamma (Aphaenogaster): Ruzsky, 1902d: 26; in Aphaenogaster: Ruzsky, 1905b: 723; in Aphaenogaster (Attomyrma): Emery, 1921f: 58.

Myrmicinae Pheidolini Aphaenogaster occidentalis TRUE TRUE FALSE

occidentalis. Stenamma (Aphaenogaster) subterraneum subsp. occidentale Emery, 1895c: 301 (w.) U.S.A. Wheeler, G.C. & Wheeler, J. 1953b: 61 (l.). Combination in Aphaenogaster (Attomyrma): Emery, 1921f: 60. Senior synonym of borealis and material of the unavailable name manni referred here: Creighton, 1950a: 150. Raised to species: Hunt & Snelling, 1975: 21.

Myrmicinae Pheidolini Aphaenogaster opposita FALSE FALSE FALSE

opposita. Myrmica opposita Say, 1836: 292 (w.m.) U.S.A. Combination in Aphaenogaster (Attomyrma): Emery, 1921f: 58. Unidentifiable taxon; incertae sedis in Aphaenogaster: Smith, M.R. 1951a: 798; unplaced taxon in Myrmicinae: Smith, D.R. 1979: 1414.

Myrmicinae Pheidolini Aphaenogaster picea TRUE TRUE FALSE

picea. Stenamma (Aphaenogaster) fulvum var. piceum Wheeler, W.M. 1908f: 621 (w.q.m.) U.S.A. [First available use of Stenamma (Aphaenogaster) fulvum subsp. aquia var. piceum Emery, 1895c: 305; unavailable name.] Combination in Aphaenogaster (Attomyrma): Emery, 1921f: 57. Subspecies of fulva: Buren, 1944a: 284; of rudis: Creighton, 1950a: 148 (incorrect as picea made available earlier than rudis and hence picea is senior name). Raised to species: Bolton, 1995b: 72;  Umphrey, 1996: 558 (in key).

Myrmicinae Pheidolini Aphaenogaster punctaticeps TRUE TRUE FALSE

punctaticeps. Aphaenogaster punctaticeps Mackay, 1989: 47, figs. 1, 4, 6, 7 (w.) U.S.A. See also: Umphrey, 1996: 557 (key).

Myrmicinae Pheidolini Aphaenogaster punctithorax FALSE FALSE FALSE

punctithorax. Aphaenogaster texana subsp. punctithorax Cole, 1938a: 239, fig. 2 (w.) U.S.A. Unidentifiable taxon: Creighton, 1950a: 151; junior synonym of picea: Smith, M.R. 1951a: 796; unidentifiable taxon; incertae sedis in Aphaenogaster: Smith, D.R. 1979: 1364.

Myrmicinae Pheidolini Aphaenogaster pusilla FALSE FALSE FALSE

pusilla. Aphaenogaster fulva var. pusilla Enzmann, J. 1947b: 150 (w.) U.S.A. [First available use of Stenamma (Aphaenogaster) fulvum subsp. aquia var. pusillum Emery, 1895c: 306; unavailable name.] Unidentifiable taxon: Creighton, 1950a: 151; unidentifiable taxon; incertae sedis in Aphaenogaster: Smith, D.R. 1979: 1364.

Myrmicinae Pheidolini Aphaenogaster rudis TRUE TRUE FALSE

rudis. Aphaenogaster fulva var. rudis Enzmann, J. 1947b: 150 (w.q.) U.S.A. [First available use of Stenamma (Aphaenogaster) fulvum subsp. aquia var. rude Emery, 1895c: 305; unavailable name.] Wheeler, G.C & Wheeler, J. 1953b: 56 (l.); Crozier, 1970: 125 (k.). Raised to species: Creighton, 1950a: 147. Subspecies of picea as this name made available first (picea entered as subspecies of rudis in Creighton, 1950a: 147), and hence is senior name: Bolton, 1995b: 72.] Raised to species: Umphrey, 1996: 558 (in key).

Myrmicinae Pheidolini Aphaenogaster tennesseensis TRUE TRUE FALSE

tennesseensis. Atta tennesseensis Mayr, 1862: 743 (w.) U.S.A. Wheeler, G.C. & Wheeler, J. 1953b: 61 (l.). Combination in Aphaenogaster: Roger, 1863b: 30; in Stenamma (Aphaenogaster): Emery, 1895c: 301; in Aphaenogaster (Attomyrma): Emery, 1921f: 60. Senior synonym of subrubra: Mayr, 1886c: 365; of laevis: Mayr, 1886d: 446; of ecalcaratum: Creighton, 1950a: 151.

Myrmicinae Pheidolini Aphaenogaster texana TRUE TRUE FALSE

texana. Aphaenogaster texana Wheeler, W.M. 1915b: 412 (w.q.m.) U.S.A. [First available use of Stenamma (Aphaenogaster) fulvum subsp. aquia var. texanum Emery, 1895c: 306;unavailable name.] Wheeler, G.C. & Wheeler, J. 1953b: 62 (l.). Senior synonym of furvescens, silvestrii: Creighton, 1950a: 152. [Unidentifiable taxa originally attached to texana: see entries under nana and punctithorax.] See also: Umphrey, 1996: 558 (key).

Myrmicinae Pheidolini Aphaenogaster treatae TRUE TRUE FALSE

1 subspecies

treatae. Aphaenogaster treatae Forel, 1886b: xl (w.q.m.) U.S.A. Combination in Stenamma (Aphaenogaster): Emery, 1895c: 302; in Aphaenogaster: Wheeler, W.M. 1913c: 114; in Aphaenogaster (Attomyrma): Emery, 1921f: 60. Senior synonym of wheeleri and material of the unavailable name alabamensis referred here: Creighton, 1950a: 153.

Myrmicinae Pheidolini Aphaenogaster uinta TRUE TRUE FALSE

uinta. Aphaenogaster uinta Wheeler, W.M. 1917a: 517 (w.q.m.) U.S.A. Combination in Aphaenogaster (Attomyrma): Emery, 1921f: 60.

Myrmicinae Pheidolini Aphaenogaster umphreyi TRUE TRUE FALSE

umphreyi. Aphaenogaster umphreyi Deyrup & Davis, 1998: 88, fig. 1 (w.) U.S.A.

Myrmicinae Pheidolini Aphaenogaster subterranea valida TRUE TRUE FALSE

valida. Aphaenogaster subterranea subsp. valida Wheeler, W.M. 1915b: 411 (w.q.m.) U.S.A. Combination in Aphaenogaster (Attomyrma): Emery, 1921f: 60.

Myrmicinae Pheidolini Aphaenogaster boulderensis smithi TRUE TRUE FALSE

smithi. Aphaenogaster boulderensis subsp. smithi Gregg, 1949b: 171 (w.) U.S.A.

Myrmicinae Pheidolini Aphaenogaster treatae pluteicornis TRUE TRUE FALSE

pluteicornis. Aphaenogaster treatae subsp. pluteicornis Wheeler, G.C. & Wheeler, E.W. 1934: 7 (w.q.m.) U.S.A. Wheeler, G.C. & Wheeler, J. 1953b: 62 (l.). Combination in Aphaenogaster (Attomyrma), and material of the unavailable name oklahomensis referred here: Creighton, 1950a: 154.

Myrmicinae Pheidolini Aphaenogaster huachucana crinimera TRUE TRUE FALSE

crinimera. Aphaenogaster (Attomyrma) huachucana subsp. crinimera Cole, 1953b: 82, fig. 1 (w.q.m.) U.S.A.

Armaniinae Armaniini Archaeopone kzylzharica TRUE TRUE TRUE

†kzylzharica. †Archaeopone kzylzharica Dlussky, 1975: 121, fig. 132 (m.) RUSSIA (Cretaceous).

Armaniinae Armaniini Archaeopone taylori TRUE TRUE TRUE

†taylori. †Archaeopone taylori Dlussky, 1983: 72, fig. 6 (m.) RUSSIA (Cretaceous).

Ponerinae Ponerini Archiponera wheeleri TRUE TRUE TRUE

†wheeleri. †Archiponera wheeleri Carpenter, 1930: 27, pl. 1, pl. 2, fig. 1 (w.) U.S.A. (Oligocene).

Armaniinae Armaniini Armania capitata TRUE TRUE TRUE

†capitata. †Armania capitata Dlussky, 1999a: 64, fig. 1 (q.) RUSSIA (Cretaceous).

Armaniinae Armaniini Armania curiosa TRUE TRUE TRUE

†curiosa. †Armaniella curiosa Dlussky, 1983: 71, fig. 5 (q.) RUSSIA (Cretaceous). Combination in †Armania: Dlussky, 1999a: 63.

Armaniinae Armaniini Armania pristina TRUE TRUE TRUE

†pristina. †Armania pristina Dlussky, 1999a: 63, fig. 1 (q.) RUSSIA (Cretaceous).

Armaniinae Armaniini Armania robusta TRUE TRUE TRUE

†robusta. †Armania robusta Dlussky, 1983: 68, figs. 1, 2 (q.) RUSSIA (Cretaceous).

Myrmicinae Attini Atta pilosa FALSE FALSE FALSE

pilosa. Oecodoma pilosa Buckley, 1867: 348 (w.) U.S.A. Unidentifiable taxon; incertae sedis in Atta: Bolton, 1995b: 76. See notes in Wheeler, W.M. 1902f: 29. Unplaced myrmicine taxon: Smith, D.R. 1979: 1414.

Myrmicinae Attini Atta tardigrada FALSE FALSE FALSE

tardigrada. Oecodoma tardigrada Buckley, 1867: 349 (w.q.m.) U.S.A. Unidentifiable taxon; incertae sedis in Atta: Bolton, 1995b: 77. Included in Acromyrmex: Forel, 1885a: 358; in Trachymyrmex: Mayr, 1886d: 442; Forel, 1893e: 601; Wheeler, W.M. 1902f: 30. Unplaced myrmicine taxon: Smith, D.R. 1979: 1414.

Myrmicinae Attini Atta texana TRUE TRUE FALSE

texana. Myrmica (Atta) texana Buckley, 1860a: 233 (w.q.m.) U.S.A. Wheeler, G.C. & Wheeler, J. 1974d: 79 (l.). Combination in Atta: Roger, 1863b: 35; in Atta (Archeatta): Gonçalves, 1942: 343. Junior synonym of fervens: Forel, 1885a: 362. Revived from synonymy: Wheeler, W.M. 1907c: 700. Subspecies of insularis: Emery, 1913bb: 259; Emery, 1924d: 354; of fervens: Borgmeier, 1939: 423. Revived status as species: Gonçalves, 1942: 335. See also: Borgmeier, 1959b: 356; Smith, D.R., 1979: 1413; Petralia & Vinson, 1980: 386.

Sphecomyrminae Sphecomyrmini Baikuris casei TRUE TRUE TRUE

†casei. †Baikuris casei Grimaldi, Agosti & Carpenter, 1997: 15, figs. 1c, 5a-d, 12b (m.) U.S.A. (Cretaceous).

Sphecomyrminae Sphecomyrmini Baikuris mandibularis TRUE TRUE TRUE

†mandibularis. †Baikuris mandibularis Dlussky, 1987: 134, fig. 2 (m.) RUSSIA (Cretaceous).

Sphecomyrminae Sphecomyrmini Baikuris mirabilis TRUE TRUE TRUE

†mirabilis. †Baikuris mirabilis Dlussky, 1987: 134, fig. 3 (m.) RUSSIA (Cretaceous).

Dolichoderinae Bothriomyrmecini Bothriomyrmex anastasiae TRUE TRUE FALSE

anastasiae. Bothriomyrmex anastasiae Dubovikov, 2002: 918, figs. 1-3 (w.q.) RUSSIA.

Formicinae Plagiolepidini Brachymyrmex depilis TRUE TRUE FALSE

depilis. Brachymyrmex heeri subsp. depilis Emery, 1893i: 635 (w.q.) U.S.A. Wheeler, G.C. & Wheeler, J. 1953c: 139 (l.). Raised to species: Santschi, 1923b: 663. Senior synonym of nanellus: Creighton, 1950a: 359; of flavescens: Cole, 1953g: 266.

Brownimeciinae Brownimeciini Brownimecia clavata TRUE TRUE TRUE

†clavata. †Brownimecia clavata Grimaldi, Agosti & Carpenter, 1997: 20, figs. 1e-f, 7-11 (w.) U.S.A. (Cretaceous). See also: Engel & Grimaldi, 2005: 12.

Formicinae Camponotini Camponotus absquatulator TRUE TRUE FALSE

absquatulator. Camponotus absquatulator Snelling, R.R. 2006: 88, figs. 12-17 (s.w.q.) U.S.A.

Formicinae Camponotini Camponotus acutirostris TRUE TRUE FALSE

acutirostris. Camponotus acutirostris Wheeler, W.M. 1910d: 317 (s.w.q.m.) U.S.A. Combination in C. (Myrmoturba): Forel, 1914a: 266; in C. (Camponotus): Emery, 1925b: 74; in C. (Tanaemyrmex): Creighton, 1950a: 375. Senior synonym of clarigaster: Creighton, 1950a: 375.

Formicinae Camponotini Camponotus americanus TRUE TRUE FALSE

americanus. Camponotus americanus Mayr, 1862: 661 (w.q.) U.S.A. Emery, 1893i: 674 (m.); Wheeler, G.C. & Wheeler, J. 1968: 216 (l.). Combination in C. (Camponotus): Emery, 1925b: 74. Junior synonym of castaneus: Mayr, 1886d: 420. Revived from synonymy as subspecies of castaneus: Emery, 1893i: 674; Wheeler, W.M. 1913c: 117; Wheeler, W.M. 1917i: 465; Wheeler, W.M. 1932a: 13. Revived status as species and senior synonym of rufinasis: Creighton, 1950a: 365. See also: Eisner & Wilson, 1952: 47.

Formicinae Camponotini Camponotus americana FALSE FALSE FALSE

americana. Formica americana Buckley, 1866: 154 (w.q.) U.S.A. Combination in Camponotus: Emery, 1895c: 337. [Unresolved junior secondary homonym of Camponotus americanus Mayr, above (Bolton, 1995b: 85).] Unidentifiable taxon; incertae sedis in Camponotus: Bolton, 1995b: 85. [Note: Emery, 1895c: 337 and Wheeler, W.M. 1902f: 19, suspected this name was cognate with marginatus. Emery, 1925b: 117 provisionally synonymised it with minutus. Smith, M.R. 1951a: 845 and Smith, D.R. 1979: 1432, dubiously place it near nearcticus.]

Formicinae Camponotini Camponotus anthrax TRUE TRUE FALSE

anthrax. Camponotus anthrax Wheeler, W.M. 1911d: 96 (s.w.q.m.) U.S.A. Wheeler, G.C. & Wheeler, J. 1970: 650 (l.). Combination in C. (Camponotus): Forel, 1914a: 266; in C. (Myrmentoma): Emery, 1920b: 257. See also: Snelling, R.R. 1988: 59.

Formicinae Camponotini Camponotus atra FALSE FALSE FALSE

atra. Formica atra Buckley, 1866: 160 (w.) U.S.A. [Unresolved junior primary homonym of atra Zetterstedt, above (Bolton, 1995b: 86).] Unidentifiable taxon; incertae sedis in Camponotus: Emery, 1895c: 337; Wheeler, W.M. 1902f: 20. [Note: Wheeler, W.M. 1917c: 28 suggests synonymy with caryae, but Smith, D.R. 1979: 1467 retains atra among the unplaced taxa.]

Formicinae Camponotini Camponotus bakeri TRUE TRUE FALSE

bakeri. Camponotus hyatti var. bakeri Wheeler, W.M. 1904d: 271 (w.q.) U.S.A. Combination in C. (Myrmentoma): Emery, 1925b: 119. Subspecies of hyatti: Creighton, 1950a: 387; Smith, M.R. 1951a: 844. Raised to species: Snelling, R.R. 1985: 24; Snelling, R.R. 1988: 60.

Formicinae Camponotini Camponotus caryae TRUE TRUE FALSE

caryae. Formica caryae Fitch, 1855: 855 (w.q.m.) U.S.A. Combination in Camponotus: Roger, 1863b: 6; in C. (Myrmentoma): Emery, 1920b: 257. Junior synonym of pennsylvanicus: Forel, 1879a: 56; Dalla Torre, 1893: 247. Revived from synonymy: Wheeler, W.M. 1917c: 28. Material of the unavailable name cnemidatus referred here: Smith, M.R. 1940b: 139. See also: Creighton, 1950a: 385; Snelling, R.R. 1988: 60. Current subspecies: nominal plus kamensis.

Formicinae Camponotini Camponotus castaneus TRUE TRUE FALSE

castaneus. Formica castanea Latreille, 1802c: 118, pl. 3, figs. 11, 12 (w.q.m.) U.S.A.Wheeler, W.M. 1910d: 321 (s.). Combination in Camponotus: Roger, 1863b: 6; in C. (Camponotus): Forel, 1914a: 266; in C. (Tanaemyrmex): Creighton, 1950a: 375. Senior synonym of mellea (and its junior synonym clarus): Mayr, 1886d: 420; Forel, 1886f: 141.

Formicinae Camponotini Camponotus clarithorax TRUE TRUE FALSE

clarithorax. Camponotus caryae subsp. clarithorax Creighton, 1950a: 386 (s.w.q.m.) U.S.A. [First available use of Camponotus marginatus subsp. discolor var. clarithorax Emery, 1893i: 678; unavailable name.] Raised to species: Smith, M.R. 1951a: 844. See also: Snelling, R.R. 1988: 61.

Formicinae Camponotini Camponotus decipiens TRUE TRUE FALSE

decipiens. Camponotus marginatus var. decipiens Emery, 1893i: 676 (w.q.) U.S.A. Wheeler, W.M. 1910f: 227 (s.w.q.m.). Combination in C. (Myrmentoma): Emery, 1925b: 117. Subspecies of fallax: Wheeler, W.M. 1910f: 227; Wheeler, W.M. 1917a: 558; of caryae: Wheeler, W.M. 1917c: 28; Emery, 1925b: 117. Junior synonym of nearcticus: Creighton, 1950a: 388. Revived from synonymy, raised to species and senior synonym of rasilis: Snelling, R.R. 1988: 66.

Formicinae Camponotini Camponotus discolor TRUE TRUE FALSE

discolor. Formica discolor Buckley, 1866: 166 (w.q.) U.S.A. Emery, 1893i: 678 (m.); Wheeler, W.M. 1910f: 230 (s.). Combination in Camponotus: Mayr, 1886c: 365; in C. (Myrmentoma): Emery, 1925b: 117. Junior synonym of marginatus: Mayr, 1886d: 423. Revived from synonymy as subspecies of marginatus: Emery, 1893i: 677. Subspecies of fallax: Wheeler, W.M. 1910f: 230; Wheeler, W.M. 1917a: 558; of caryae: Wheeler, W.M. 1917c: 28; Creighton, 1950a: 386. Revived status as species: Snelling, R.R. 1988: 68.

Formicinae Camponotini Camponotus dumetorum TRUE TRUE FALSE

dumetorum. Camponotus maculatus subsp. dumetorum Wheeler, W.M. 1910d: 354 (s.w.m.) U.S.A. Combination in C. (Myrmoturba): Wheeler, W.M. 1917a: 560; in C. (Camponotus): Emery, 1925b: 75; in C. (Tanaemyrmex): Creighton, 1950a: 377. Subspecies of sansabeanus: Emery, 1925b: 75. Junior synonym of maccooki: Creighton, 1950a: 377. Revived from synonymy and raised to species: Snelling, R.R. 1970: 396.

Formicinae Camponotini Camponotus essigi TRUE TRUE FALSE

essigi. Camponotus caryae var. essigi Smith, M.R. 1923b: 306 (s.w.) U.S.A. Combination in C. (Myrmentoma): Creighton, 1950a: 387. Raised to species: Creighton, 1950a: 387. Senior synonym of nevadensis: Wheeler, G.C. & Wheeler, J. 1986g: 61; of pertusus: Snelling, R.R. 2000: 601. See also: Smith, M.R. 1951a: 844; Snelling, R.R. 1988: 68.

Formicinae Camponotini Camponotus etiolatus TRUE TRUE FALSE

etiolatus. Camponotus (Colobopsis) abditus var. etiolatus Wheeler, W.M. 1904b: 150, fig. 5 (s.w.q.m.) U.S.A. Wheeler, G.C. & Wheeler, J. 1970: 650 (l.). Raised to species: Wheeler, W.M. 1927f: 31; Wheeler, W.M. 1934g: 216.

Formicinae Camponotini Camponotus festinatus TRUE TRUE FALSE

festinatus. Formica festinata Buckley, 1866: 164 (w.q.) U.S.A. Wheeler, W.M. 1910d: 312 (s.m.); Wheeler, G.C. & Wheeler, J. 1970: 651 (l.). Combination in Camponotus: Dalla Torre, 1893: 231; in C. (Myrmoturba): Wheeler, W.M. 1917a: 561; in C. (Tanaemyrmex): Emery, 1925b: 80. Subspecies of fumidus: Wheeler, W.M. 1902f: 22; Creighton, 1950a: 376. Revived status as species: Snelling, R.R. 1968b: 350. Senior synonym of pubicornis: Wheeler, W.M. 1910d: 312; of spurcus: Creighton, 1950a: 376; of †juliae: Snelling, R.R. (pers. comm., unpublished). See also: Snelling, R.R. 2006: 85.

Formicinae Camponotini Camponotus floridanus TRUE TRUE FALSE

floridanus. Formica floridana Buckley, 1866: 161 (w.) U.S.A. Mayr, 1886d: 423 (q.m.). Combination in Camponotus: Mayr, 1886d: 423; in C. (Myrmothrix): Emery, 1925b: 108. Subspecies of atriceps: Mayr, 1886d: 423; of abdominalis: Emery, 1893i: 670. Revived status as species: Wheeler, W.M. 1910d: 325. Subspecies of abdominalis: Wheeler, W.M. 1913c: 117. Senior synonym of yankee: Mayr, 1886d: 423. Junior synonym of atriceps: Hashmi, 1973: 82. Revived from synonymy as subspecies of abdominalis: Deyrup & Trager, 1986: 219. Revived status as species: Deyrup, Johnson, et al. 1989: 100.

Formicinae Camponotini Camponotus fuscipennis TRUE TRUE TRUE

†fuscipennis. †Camponotus fuscipennis Carpenter, 1930: 59, pl. 5, fig. 4; pl. 11, fig. 6 (q.) U.S.A. (Oligocene). [†Camponotus carpenteri Özdikmen, 2010a: 525. Unnecessary replacement name.]

Formicinae Camponotini Camponotus hunteri TRUE TRUE FALSE

hunteri. Camponotus (Colobopsis) pylartes var. hunteri Wheeler, W.M. 1910d: 353 (s.w.) U.S.A. Raised to species: Creighton, 1950a: 393.

Formicinae Camponotini Camponotus hyatti TRUE TRUE FALSE

hyatti. Camponotus hyatti Emery, 1893i: 680, pl. 22, figs. 25, 26 (s.w.) U.S.A. Combination in C. (Camponotus): Forel, 1914a: 266; in C. (Myrmentoma): Emery, 1920b: 257. Senior synonym of californica: Snelling, R.R. 1988: 69.

Formicinae Camponotini Camponotus impressus TRUE TRUE FALSE

impressus. Colobopsis impressa Roger, 1863a: 160 (w.) U.S.A. Mayr, 1886d: 423 (s.q.); Emery, 1893i: 681 (m.). Combination in Camponotus (Colobopsis): Emery, 1889b: 517.

Formicinae Camponotini Camponotus incensus TRUE TRUE FALSE

incensus. Camponotus (Tanaemyrmex) incensus Wheeler, W.M. 1932a: 14 (s.w.) U.S.A.

Formicinae Camponotini Camponotus kopetdaghensis TRUE TRUE FALSE

kopetdaghensis. Camponotus (Myrmentoma) kopetdaghensis Dlussky & Zabelin, 1985: 236, fig. 8 (s.w.) RUSSIA.

Formicinae Camponotini Camponotus koseritzi TRUE TRUE FALSE

koseritzi. Camponotus tenuiscapus r. koseritzi Emery, 1888c: 366 (s.w.) BRAZIL. Combination in C. (Myrmoturba): Bruch, 1914: 229; in C. (Tanaemyrmex): Emery, 1925b: 78. Subspecies of tenuiscapus: Forel, 1901h: 70. Status as species: Dalla Torre, 1893: 237; Emery, 1894g: 375; Forel, 1895b: 103; Emery, 1903: 68; Kempf, 1972a: 68.

Formicinae Camponotini Camponotus kurdistanicus TRUE TRUE FALSE

kurdistanicus. Camponotus gestroi subsp. kurdistanicus Emery, 1898c: 151 (w.q.) RUSSIA. Combination in C. (Myrmentoma): Emery, 1920b: 257. Raised to species: Emery, 1920b: 257; Emery, 1920c: 25. See also: Emery, 1925a: 70.

Formicinae Camponotini Camponotus laevigatus TRUE TRUE FALSE

laevigatus. Formica laevigata Smith, F. 1858b: 55 (w.q.) U.S.A. Wheeler, W.M. 1910d: 327 (s.m.); Wheeler, G.C. & Wheeler, J. 1968: 216 (l.). Combination in Camponotus: Roger, 1863b: 5; in C. (Camponotus): Forel, 1914a: 266.

Formicinae Camponotini Camponotus lameerei TRUE TRUE FALSE

lameerei. Camponotus marginatus var. lameerei Emery, 1898c: 150 (s.w.q.m.) RUSSIA. Combination in C. (Myrmentoma): Emery, 1925b: 118. Subspecies of marginatus: Ruzsky, 1905b: 249; Kuznetsov-Ugamsky, 1923: 242; of lateralis: Forel, 1904b: 380; of caryae: Wheeler, W.M. 1917c: 29. Raised to species: Tarbinsky, 1976: 146; Dlussky & Zabelin, 1985: 237. Senior synonym of barbiger: Radchenko, 1997b: 704.

Formicinae Camponotini Camponotus maguassa TRUE TRUE FALSE

maguassa. Camponotus (Myrmoturba) maguassa Wheeler, W.M. 1922a: 237 (s.w.) DEMOCRATIC REPUBLIC OF CONGO. Combination in C. (Tanaemyrmex): Emery, 1925b: 92.

Formicinae Camponotini Camponotus maritimus TRUE TRUE FALSE

maritimus. Camponotus maritimus Ward, 2005: 10, figs. 1-4 (s.w.q.m.) U.S.A.

Formicinae Camponotini Camponotus microcephalus TRUE TRUE TRUE

†microcephalus. †Camponotus microcephalus Carpenter, 1930: 60, pl. 6, fig. 4; pl. 11, fig. 8 (q.) U.S.A. (Oligocene).

Formicinae Camponotini Camponotus microps TRUE TRUE FALSE

microps. Camponotus microps Snelling, R.R. 2006: 91, figs. 23-27 (s.w.q.) U.S.A.

Formicinae Camponotini Camponotus mississippiensis TRUE TRUE FALSE

mississippiensis. Camponotus (Colobopsis) mississippiensis Smith, M.R. 1923a: 83 (s.w.) U.S.A. Wheeler, G.C. & Wheeler, J. 1953e: 188 (l.).

Formicinae Camponotini Camponotus modoc TRUE TRUE FALSE

modoc. Camponotus (Camponotus) herculeanus var. modoc Wheeler, W.M. 1910d: 333 (s.w.q.m.) U.S.A. Combination in C. (Camponotus): Emery, 1925b: 72. Subspecies of pennsylvanicus: Creighton, 1950a: 369; of herculeanus: Brown, 1950d: 158. Raised to species: Hunt & Snelling, R.R. 1975: 22; Smith, D.R. 1979: 1426; Mackay, Lowrie, et al. 1988: 104 (in key).

Formicinae Camponotini Camponotus nearcticus TRUE TRUE FALSE

nearcticus. Camponotus marginatus var. nearcticus Emery, 1893i: 675 (w.q.) U.S.A. Wheeler, W.M. 1910f: 223 (s.m.); Wheeler, G.C. & Wheeler, J. 1953e: 193 (l.). Combination in C. (Myrmentoma): Emery, 1925b: 117. Subspecies of fallax: Wheeler, W.M. 1908g: 406; Wheeler, W.M. 1917a: 558. Junior synonym of caryae: Wheeler, W.M. 1917c: 27. Revived from synonymy as subspecies of caryae: Buren, 1944a: 294. Raised to species: Creighton, 1950a: 388; Smith, M.R. 1951a: 845. Senior synonym of minutus, pardus, tanquaryi: Creighton, 1950a: 388. See also: Snelling, R.R. 1988: 70.

Formicinae Camponotini Camponotus novaeboracensis TRUE TRUE FALSE

novaeboracensis. Formica novaeboracensis Fitch, 1855: 766 (w.) U.S.A. Wheeler, W.M. 1910d: 340 (s.w.q.m.); Wheeler, G.C. & Wheeler, J. 1953e: 182 (l.). Combination in Camponotus: Roger, 1863b: 6; in C. (Camponotus): Emery, 1925b: 72. Junior synonym of pennsylvanicus: Dalla Torre, 1893: 247; Emery, 1896d: 372. Revived from synonymy as subspecies of ligniperda: Forel, 1899c: 130. Subspecies of herculeanus: Emery, 1925b: 72; Buren, 1944a: 293. Revived status as species: Creighton, 1950a: 369. Senior synonym of pictus: Forel, 1899c: 130; of rubens: Creighton, 1950a: 370.

Formicinae Camponotini Camponotus obliquus TRUE TRUE FALSE

obliquus. Camponotus (Colobopsis) obliquus Smith, M.R. 1930b: 567 (s.) U.S.A.

Formicinae Camponotini Camponotus ocreatus TRUE TRUE FALSE

ocreatus. Camponotus maculatus subsp. ocreatus Emery, 1893i: 673 (s.w.) U.S.A. Wheeler, W.M. 1910d: 310 (q.); Wheeler, G.C. & Wheeler, J. 1970: 651 (l.). Combination in C. (Myrmoturba): Wheeler, W.M. 1917a: 561; in C. (Camponotus): Emery, 1920b: 255; in C. (Tanaemyrmex): Creighton, 1950a: 378. Raised to species: Wheeler, W.M. 1917a: 561. Senior synonym of primipilaris: Creighton, 1950a: 378.

Formicinae Camponotini Camponotus papago TRUE TRUE FALSE

papago. Camponotus (Colobopsis) papago Creighton, 1953c: 153, figs. 1, 2; pl. 13, figs. 1-3 (s.w.q.m.) U.S.A.

Formicinae Camponotini Camponotus pennsylvanicus TRUE TRUE FALSE

pennsylvanicus. Formica pensylvanica De Geer, 1773: 603, pl. 31, figs. 9, 10 (s.w.q.m.) U.S.A. Wheeler, G.C. & Wheeler, J. 1953e: 187 (l.). Combination in Camponotus: Mayr, 1862: 666; in C. (Camponotus): Forel, 1914a: 266. Subspecies of herculeanus: Forel, 1879a: 57; Mayr, 1886d: 420; Wheeler, W.M. 1913c: 117; Wheeler, W.M. 1917i: 465; Wheeler, W.M. 1932a: 13; Buren, 1944a: 293. Status as species: Ruzsky, 1896: 67; Forel, 1901h: 70; Forel, 1907e: 10; Wheeler, W.M. 1910d: 335; Emery, 1920b: 255; Creighton, 1950a: 367; Smith, D.R. 1979: 1427. Senior synonym of semipunctata: Mayr, 1886d: 420; of herculeanopennsylvanicus (and its junior synonym mahican): Creighton, 1950a: 367.

Formicinae Camponotini Camponotus petrifactus TRUE TRUE TRUE

†petrifactus. †Camponotus petrifactus Carpenter, 1930: 60, pl. 6, fig. 5 (w.) U.S.A. (Oligocene).

Formicinae Camponotini Camponotus punctulatus TRUE TRUE FALSE

11 subspecies

punctulatus. Camponotus punctulatus Mayr, 1868b: 161 (w.q.) ARGENTINA. Goni et al., 1983: 366 (k.). Combination in C. (Myrmoturba): Forel, 1913l: 248; in C. (Neomyrmamblys): Wheeler, W.M. 1922a: 708; in C. (Tanaemyrmex): Emery, 1925b: 78. Subspecies of tenuiscapus: Emery, 1888c: 365. Status as species: Dalla Torre, 1893: 248; Emery, 1894g: 375; Pergande, 1896: 864; Emery, 1903: 65; Santschi, 1922c: 104. Current subspecies: nominal plus andigenus, brevibarbis, chubutensis, cruentus, hybridus, imberbis, lillii, lizeri, minutior, pergandei, tenuibarbis.

Formicinae Camponotini Camponotus pylartes TRUE TRUE FALSE

1 subspecies

pylartes. Camponotus (Colobopsis) pylartes Wheeler, W.M. 1904b: 147, fig. 4 (s.w.q.) U.S.A. Petralia & Vinson, 1980: 386 (l.). Current subspecies: nominal plus fraxinicola.

Formicinae Camponotini Camponotus quercicola TRUE TRUE FALSE

quercicola. Camponotus (Camponotus) quercicola Smith, M.R. 1954b: 211 (w.) U.S.A. See also: Gadau, Brady & Ward, 1999: 514.

Formicinae Camponotini Camponotus sachalinensis TRUE TRUE FALSE

sachalinensis. Camponotus herculeanus var. sachalinensis Forel, 1904b: 38 (q.) RUSSIA. Karavaiev, 1912b: 592 (m.). Combination in C. (Camponotus): Emery, 1925b: 72. Subspecies of herculeanus: Ruzsky, 1926: 108. Provisional junior synonym of saxatilis: Kuznetsov-Ugamsky, 1928b: 4; Kuznetsov-Ugamsky, 1929a: 18. Junior synonym of herculeanus: Yasumatsu & Brown, 1957: 49. Revived from synonymy as subspecies of herculeanus: Arnol'di, 1967: 1821. Subspecies of herculeanus: Collingwood, 1976: 306; Kupyanskaya, 1990: 166; Radchenko, 1996b: 1203. Raised to species: Collingwood, 1981: 29; Bolton, 1995b: 121. Senior synonym of altaica, jacuticus: Arnol'di, 1967: 1821.

Formicinae Camponotini Camponotus sansabeanus TRUE TRUE FALSE

sansabeanus. Formica sansabeana Buckley, 1866: 167 (s.w.q.m.) U.S.A. Wheeler, G.C. & Wheeler, J. 1953e: 196 (l.). Combination in Camponotus: Mayr, 1886c: 365; in C. (Myrmoturba): Forel, 1914a: 267; in C. (Camponotus): Emery, 1920b: 255; in C. (Tanaemyrmex): Creighton, 1950a: 378. Junior synonym of marginatus: Mayr, 1886c: 423. Revived from synonymy as subspecies of marginatus: Dalla Torre, 1893: 242. Subspecies of maculatus: Emery, 1896d: 371; Wheeler, W.M. 1910d: 307. Revived status as species: Emery, 1920b: 232 (footnote); Cole, 1942: 387; Creighton, 1950a: 378. Senior synonym of bulimosus, torrefactus: Wheeler, G.C. & Wheeler, J. 1986g: 14.

Formicinae Camponotini Camponotus saxatilis TRUE TRUE FALSE

saxatilis. Camponotus herculeanus subsp. saxatilis Ruzsky, 1895: 7 (w.q.m.) RUSSIA. Arnol'di, 1967: 1825 (s.). Combination in C. (Camponotus): Emery, 1925b: 73. Subspecies of pennsylvanicus: Ruzsky, 1905b: 228; of herculeanus: Ruzsky, 1925b: 42. Junior synonym of japonicus: Yasumatsu & Brown, 1951: 36. Revived from synonymy: Arnol'di, 1967: 1825; Kupyanskaya, 1990: 167.

Formicinae Camponotini Camponotus sayi TRUE TRUE FALSE

sayi. Camponotus sayi Emery, 1893i: 679, pl. 22, figs. 27, 28 (s.w.) U.S.A. Wheeler, W.M. 1910d: 343 (s.w.). Combination in C. (Camponotus): Forel, 1914a: 266; in C. (Myrmentoma): Emery, 1920b: 257. See also: Snelling, R.R. 1988: 72.

Formicinae Camponotini Camponotus schaefferi TRUE TRUE FALSE

schaefferi. Camponotus schaefferi Wheeler, W.M. 1909e: 88 (s.w.q.) U.S.A. Wheeler, W.M. 1910d: 344 (s.). Combination in C. (Myrmentoma): Emery, 1920b: 257; in C. (Camponotus): Forel, 1914a: 266; Creighton, 1950a: 371.

Formicinae Camponotini Camponotus semitestaceus TRUE TRUE FALSE

semitestaceus. Camponotus (Tanaemyrmex) semitestaceus Snelling, R.R. 1970: 396 (w.) U.S.A. [First available use of Camponotus maculatus subsp. vicinus var. semitestaceus Emery, 1893i: 672; unavailable name.]

Formicinae Camponotini Camponotus septentrionalis FALSE FALSE FALSE

septentrionalis. Formica septentrionale Buckley, 1866: 161 (w.q.) U.S.A. Combination in Camponotus: Emery, 1895c: 338; Wheeler, W.M. 1902f: 21. Unidentifiable taxon; incertae sedis in Camponotus: Bolton, 1995b: 123.

Formicinae Camponotini Camponotus subbarbatus TRUE TRUE FALSE

subbarbatus. Camponotus marginatus subsp. subbarbatus Emery, 1893i: 676 (w.q.m.) U.S.A. Wheeler, W.M. 1910f: 229 (s.). Combination in C. (Myrmentoma): Emery, 1925b: 117. Subspecies of fallax: Wheeler, W.M. 1910f: 229; of caryae: Wheeler, W.M. 1917c: 28. Raised to species and material of the unavailable name paucipilis referred here: Creighton, 1950a: 390. See also: Snelling, R.R. 1988: 73.

Formicinae Camponotini Camponotus tejonia FALSE FALSE FALSE

tejonia. Formica tejonia Buckley, 1866: 161 (m.) U.S.A. Combination in Camponotus: Emery, 1895c: 338; Wheeler, W.M. 1902f: 21. Unidentifiable taxon; incertae sedis in Camponotus: Bolton, 1995b: 126.

Formicinae Camponotini Camponotus tenuiscapus TRUE TRUE FALSE

tenuiscapus. Camponotus tenuiscapus Roger, 1863a: 143 (w.) URUGUAY. Emery, 1888c: 79 (s.). Combination in C. (Myrmamblys): Bruch, 1914: 230; in C. (Myrmoturba): Forel, 1914a: 267; Emery, 1920b: 255; in C. (Tanaemyrmex): Emery, 1925b: 79.

Formicinae Camponotini Camponotus texanus TRUE TRUE FALSE

texanus. Camponotus texanus Wheeler, W.M. 1903b: 108, fig. 10 (s.w.q.m.) U.S.A. Combination in C. (Myrmentoma): Emery, 1920b: 257; in C. (Camponotus): Forel, 1914a: 266; Creighton, 1950a: 371.

Formicinae Camponotini Camponotus tortuganus TRUE TRUE FALSE

tortuganus. Camponotus maculatus subsp. tortuganus Emery, 1895c: 336 (w.) U.S.A. Wheeler, W.M. 1910d: 310 (s.q.m.). Combination in C. (Tanaemyrmex): Emery, 1925b: 80. Subspecies of picipes: Emery, 1920b: 233. Raised to species: Wheeler, W.M. 1932a: 13; Creighton, 1950a: 380; Wilson, 1964b: 12.

Formicinae Camponotini Camponotus trepidulus TRUE TRUE FALSE

trepidulus. Camponotus (Myrmobrachys) trepidulus Creighton, 1965b: 7, figs. 5-8 (s.w.q.m.) U.S.A.

Formicinae Camponotini Camponotus ulcerosus TRUE TRUE FALSE

ulcerosus. Camponotus ulcerosus Wheeler, W.M. 1910d: 351 (s.) U.S.A. Creighton, 1951: 58 (w.q.). Combination in C. (Colobopsis): Forel, 1914a: 272; in C. (Myrmamblys): Emery, 1920b: 259; in C. (Manniella): Emery, 1925b: 160; in C. (Myrmaphaenus): Creighton, 1951: 57. Senior synonym of bruesi: Creighton, 1951: 53. See also: Creighton, 1953d: 82.

Formicinae Camponotini Camponotus vafer TRUE TRUE FALSE

vafer. Camponotus vafer Wheeler, W.M. 1910d: 315 (s.w.q.) U.S.A. Combination in C. (Myrmoturba): Forel, 1914a: 268; in C. (Camponotus): Emery, 1925b: 75; in C. (Tanaemyrmex): Creighton, 1950a: 381. See also: Snelling, R.R. 2006: 95.

Formicinae Camponotini Camponotus vetus TRUE TRUE TRUE

†vetus. †Camponotus vetus Scudder, 1877a: 742 (q.?) U.S.A. (Eocene).

Formicinae Camponotini Camponotus vicinus TRUE TRUE FALSE

vicinus. Camponotus vicinus Mayr, 1870b: 940 (w.) U.S.A. Emery, 1893i: 671 (s.q.); Wheeler, W.M. 1910d: 301 (m.); Wheeler, G.C. & Wheeler, J. 1968: 220 (l.). Combination in C. (Myrmoturba): Forel, 1914a: 268; in C. (Camponotus): Emery, 1925b: 75; in C. (Tanaemyrmex): Creighton, 1950a: 381. Subspecies of sylvaticus: Mayr, 1886d: 422; of maculatus: Emery, 1893i: 671; of sansabeanus: Emery, 1920b: 232 (footnote); Cole, 1942: 387. Revived status as species: Creighton, 1950a: 381; Snelling, R.R. 1970: 396. Material of the unavailable names infernalis, luteangulus, maritimus, nitidiventris, plorabilis, subrostrata referred here by Creighton, 1950a: 381; material of the unavailable name berkeleyensis referred here by Snelling, R.R. 1970: 396.

Formicinae Camponotini Camponotus yogi TRUE TRUE FALSE

yogi. Camponotus yogi Wheeler, W.M. 1915b: 420 (s.w.) U.S.A. Creighton & Snelling, 1967: 191 (q.m.); Wheeler, G.C. & Wheeler, J. 1970: 650 (l.). Combination in C. (Colobopsis): Wheeler, W.M. 1917a: 562; in C. (Myrmaphaenus): Emery, 1925b: 156.

Formicinae Camponotini Camponotus punctulatus hybridus TRUE TRUE FALSE

hybridus. Camponotus tenuiscapus r. hybridus Forel, 1901h: 69 (s.w.q.) BRAZIL. Combination in C. (Myrmoturba): Forel, 1915c: 363; Bruch, 1915: 535; in C. (Tanaemyrmex): Emery, 1925b: 78. Currently subspecies of punctulatus: Forel, 1915c: 363; Bruch, 1915: 535.

Formicinae Camponotini Camponotus pylartes fraxinicola TRUE TRUE FALSE

fraxinicola. Camponotus (Colobopsis) pylartes subsp. fraxinicola Smith, M.R. 1923a: 86 (s.w.) U.S.A.

Formicinae Camponotini Camponotus punctulatus minutior TRUE TRUE FALSE

minutior. Camponotus punctulatus r. minutior Forel, 1886f: 173 (s.w.) ARGENTINA. Combination in C. (Myrmoturba): Forel, 1913l: 249; in C. (Tanaemyrmex): Emery, 1925b: 79. Subspecies of tenuiscapus: Emery, 1888c: 366; of punctulatus: Emery, 1894g: 376; Santschi, 1922c: 104. Raised to species: Dalla Torre, 1893: 243; Emery, 1903: 71; Emery, 1906c: 190. Provisional synonym of punctulatus: Wheeler, W.M. 1929e: 104 (footnote). Currently subspecies of punctulatus: Kempf, 1972a: 71.

Myrmicinae Formicoxenini Cardiocondyla koshewnikovi TRUE TRUE FALSE

koshewnikovi. Cardiocondyla koshewnikovi Ruzsky, 1902b: 480 (w.q.) RUSSIA. [Also described as new by Ruzsky, 1902c: 16.] Subspecies of stambuloffii: Forel, 1902h: 440; Kuznetsov-Ugamsky, 1927d: 37. Junior synonym of stambuloffii: Pisarski, 1970: 308. Revived from synonymy as subspecies of stambuloffii: Tarbinsky, 1976: 72; Dlussky & Zabelin, 1985: 213. Revived status as species: Dlussky, Soyunov & Zabelin, 1990: 195. See also: Radchenko, 1995b: 450; Seifert, 2003a: 266.

Myrmicinae Formicoxenini Cardiocondyla ulianini TRUE TRUE FALSE

ulianini. Cardiocondyla elegans var. ulianini Emery, 1889a: 441 (w.) RUSSIA. Karavaiev, 1910b: 57 (q.); Wheeler, G.C. & Wheeler, J. 1953d: 188 (l.); Marikovsky & Yakushin, 1974: 57 (w.q.m., ergatoids). Subspecies of elegans: Kuznetsov-Ugamsky, 1927d: 38; Pisarski, 1967: 388; Dlussky & Zabelin, 1985: 213. Raised to species: Agosti & Collingwood, 1987a: 56. Junior synonym of elegans: Dlussky, Soyunov & Zabelin, 1990: 194; Radchenko, 1995b: 449. Revived from synonymy: Seifert, 2003a: 229.

Myrmicinae Solenopsidini Carebara longii TRUE TRUE FALSE

longii. Erebomyrma longii Wheeler, W.M. 1903a: 140, figs. 1-5 (w.q.m.) U.S.A. Combination in Oligomyrmex: Ettershank, 1966: 123; in Erebomyrma: Wilson, 1986b: 61; in Oligomyrmex: Bolton, 1995b: 299; in Carebara: Fernández, 2004a: 199.

Amblyoponinae Amblyoponini Casaleia inversa TRUE TRUE TRUE

†inversa. †Protamblyopone inversa Dlussky, 1981b: 65, fig. 50 (w.) RUSSIA (Miocene). Combination in Casaleia: Bolton, 1995b: 134. [Note. Pagliano & Scaramozzino, 1990: 5, did not indicate the combination of this species in Casaleia.]

Formicinae Formicini Cataglyphis aenescens TRUE TRUE FALSE

aenescens. Formica aenescens Nylander, 1849: 37 (w.q.m.) RUSSIA. Combination in Tapinoma: Smith, F. 1858b: 56; in Myrmecocystus: Emery & Forel, 1879: 449; in Cataglyphis: Mayr, 1861: 45; Müller, 1923: 149; in C. (Monocombus): Santschi, 1929b: 35. Junior synonym of cursor: Mayr, 1861: 45. Revived from synonymy: Emery & Forel, 1879: 449. Subspecies of cursor: Emery, 1906d: 47; Wheeler, W.M. & Mann, 1916: 173; Santschi, 1921a: 116; Karavaiev, 1924: 301; Santschi, 1929b: 35; Karavaiev, 1936: 263. Revived status as species: Pisarski, 1967: 416; Tarbinsky, 1976: 203; Arnol'di & Dlussky, 1978: 554; Dlussky & Zabelin, 1985: 239; Agosti, 1990b: 1480; Atanassov & Dlussky, 1992: 294. Senior synonym of caspius: Karavaiev, 1910b: 18; Lomnicki, 1925a: 160; of jakobsoni and material of the unavailable name flavigastra referred here: Dlussky, Soyunov & Zabelin, 1990: 149; of aterrima, chatkalensis, georgica, rockingeri, tancrei and material of the unavailable names maeotica, kuenlunensis referred here: Radchenko, 1997c: 436. See also: Radchenko, 2007: 36.

Formicinae Formicini Cataglyphis cinnamomea TRUE TRUE FALSE

cinnamomea. Myrmecocystus albicans subsp. cinnamomeus Karavaiev, 1910a: 269 (w.) RUSSIA. [Also described as new, as Myrmecocystus albicans subsp. viaticoides var. cinnamomea Karavaiev, 1910b: 37; unavailable name.] Combination in Cataglyphis: Emery, 1925b: 263. Raised to species: Pisarski, 1967: 418; Arnol'di, 1968: 1815.

Formicinae Formicini Cataglyphis foreli TRUE TRUE FALSE

foreli. Myrmecocystus foreli Ruzsky, 1903a: 36 (w.) RUSSIA. Arnol'di, 1964: 1803 (q.m.). Combination in Cataglyphis: Emery, 1912f: 104; in C. (Monocombus): Santschi, 1929b: 29. Subspecies of altisquamis: Forel, 1904b: 382. Revived status as species: Emery, 1912f: 104; Arnol'di, 1964: 1812; Pisarski, 1967: 418.

Formicinae Formicini Cataglyphis gracilens TRUE TRUE FALSE

gracilens. Cataglyphis gracilens Santschi, 1929b: 65 (w.) RUSSIA. [First available use of Myrmecocystus altisquamis subsp. foreli var. gracilens Ruzsky, 1905b: 439; unavailable name.]

Myrmicinae Cephalotini Cephalotes rohweri TRUE TRUE FALSE

rohweri. Cryptocerus (Cyathocephalus) rohweri Wheeler, W.M. 1916f: 32, fig. 2 (s.w.) U.S.A. Smith, M.R. 1947a: 35 (q.). Combination in Cryptocerus (Cryptocerus): Smith, M.R. 1947a: 34; in Paracryptocerus (Harnedia): Smith, M.R. 1951a: 825; in Zacryptocerus: Smith, D.R. 1979: 1403; in Cephalotes: De Andrade & Baroni Urbani, 1999: 568. See also: Kempf, 1958a: 129; Creighton & Nutting, 1965: 59.

Myrmicinae Cephalotini Cephalotes texanus TRUE TRUE FALSE

texanus. Cryptocerus texanus Santschi, 1915b: 208, fig. 2 (s.w.) U.S.A. Smith, M.R. 1947a: 38 (q.m.); Creighton & Gregg, 1954: 51 (q.). Combination in Paracryptocerus (Harnedia): Smith, M.R. 1951a: 825; in Zacryptocerus: Smith, D.R. 1979: 1403; in Cephalotes: De Andrade & Baroni Urbani, 1999: 591. See also: Kempf, 1958a: 123.

Cerapachyinae Cerapachyini Cerapachys augustae TRUE TRUE FALSE

augustae. Cerapachys (Parasyscia) augustae Wheeler, W.M. 1902d: 182, figs. 1, 2 (w.q.) U.S.A. Wheeler, W.M. 1903g: 206 (l.); Wheeler, G.C. 1950: 106 (l.); Smith, M.R. 1942b: 63 (m.).

Cerapachyinae Cerapachyini Cerapachys davisi TRUE TRUE FALSE

davisi. Cerapachys (Parasyscia) davisi Smith, M.R. 1942b: 64 (m.) U.S.A.

Formicinae Camponotini Chimaeromyrma brachycephala TRUE TRUE TRUE

†brachycephala. †Chimaeromyrma brachycephala Dlussky, 1988: 60, fig. 5 (w.) RUSSIA (Paleocene).

Myrmicinae Crematogastrini Crematogaster ashmeadi TRUE TRUE FALSE

ashmeadi. Crematogaster ashmeadi Mayr, 1886d: 463 (w.m.) U.S.A. Combination in C. (Acrocoelia): Emery, 1922e: 141; in C. (Crematogaster): Buren, 1968b: 92. Senior synonym of matura: Creighton, 1950a: 206. See also: Johnson, C. 1988: 315.

Myrmicinae Crematogastrini Crematogaster atkinsoni TRUE TRUE FALSE

atkinsoni. Crematogaster atkinsoni Wheeler, W.M. 1919g: 108, fig. 16 (w.) U.S.A. Wheeler, W.M. 1932a: 9 (q.m.). Combination in C. (Acrocoelia): Creighton, 1950a: 207; in C. (Crematogaster): Buren, 1968b: 92. Senior synonym of helveola: Creighton, 1950a: 207. See also: Johnson, C. 1988: 316.

Myrmicinae Crematogastrini Crematogaster browni TRUE TRUE FALSE

browni. Crematogaster (Crematogaster) browni Buren, 1968b: 100 (w.q.) U.S.A.

Myrmicinae Crematogastrini Crematogaster californica TRUE TRUE FALSE

californica. Crematogaster laeviuscula var. californica Wheeler, W.M. 1919g: 111 (w.) U.S.A. [First available use of Crematogaster lineolata subsp. laeviuscula var. californica Emery, 1895c: 285; unavailable name.] Combination in C. (Crematogaster): Buren, 1968b: 94. Subspecies of lineolata: Wheeler, W.M. 1934f: 135. Junior synonym of coarctata: Creighton, 1950a: 207. Revived from synonymy and raised to species: Smith, M.R. 1951a: 808; Buren, 1968b: 94 (in key).

Myrmicinae Crematogastrini Crematogaster cerasi TRUE TRUE FALSE

cerasi. Myrmica cerasi Fitch, 1855: 835 (w.) U.S.A. Wheeler, G.C. & Wheeler, J. 1973a: 28 (l.). Combination in Crematogaster: Roger, 1863b: 37; in C. (Acrocoelia): Emery, 1922e: 141; in C. (Crematogaster): Buren, 1968b: 92. Junior synonym of lineolata: Dalla Torre, 1893: 83; Creighton, 1950a: 213. Revived from synonymy as subspecies of lineolata: Emery, 1895c: 282; Smith, M.R. 1951a: 809. Revived status as species, senior synonym of kennedyi and material of the unavailable name punctinodis referred here: Buren, in Smith, M.R. 1958c: 125. See also: Buren, 1968b: 92; Johnson, C. 1988: 318.

Myrmicinae Crematogastrini Crematogaster coarctata TRUE TRUE FALSE

coarctata. Crematogaster coarctata Mayr, 1870b: 992 (w.) U.S.A. Combination in C. (Acrocoelia): Emery, 1922e: 141; in C. (Crematogaster): Buren, 1968b: 94. Junior synonym of lineolata: Mayr, 1886d: 462. Revived from synonymy as subspecies of lineolata: Emery, 1895c: 283; Wheeler, W.M. 1904d: 270. Revived status as species: Wheeler, W.M. 1919g: 111. Subspecies of lineolata: Emery, 1922e: 141; of opaca: Enzmann, J. 1946c: 94. Revived status as species: Creighton, 1950a: 207; Buren, 1968b: 94 (in key).

Myrmicinae Crematogastrini Crematogaster colei TRUE TRUE FALSE

colei. Crematogaster (Crematogaster) colei Buren, 1968b: 108 (w.q.) U.S.A.

Myrmicinae Crematogastrini Crematogaster depilis TRUE TRUE FALSE

depilis. Crematogaster opaca var. depilis Wheeler, W.M. 1919g: 111 (w.) U.S.A. [First available use of Crematogaster lineolata subsp. opaca var. depilis Wheeler, W.M. 1908e: 478; unavailable name.] Wheeler, G.C. & Wheeler, J. 1973a: 28 (l.). Combination in C. (Acrocoelia): Emery, 1922e: 141; in C. (Crematogaster): Buren, 1968b: 93. Subspecies of punctulata: Enzmann, J. 1946c: 93. Raised to species: Creighton, 1950a: 209; Buren, in Smith, M.R. 1958c: 126.

Myrmicinae Crematogastrini Crematogaster emeryana TRUE TRUE FALSE

emeryana. Crematogaster (Acrocoelia) lineolata subsp. emeryana Creighton, 1950a: 213 (w.) U.S.A. Combination in C. (Crematogaster): Buren, 1968b: 94 (in key). Raised to species: Buren, 1968b: 94 (in key).

Myrmicinae Crematogastrini Crematogaster hespera TRUE TRUE FALSE

hespera. Crematogaster (Crematogaster) hespera Buren, 1968b: 98 (w.q.) U.S.A.

Myrmicinae Crematogastrini Crematogaster isolata TRUE TRUE FALSE

isolata. Crematogaster (Crematogaster) isolata Buren, 1968b: 106 (w.) U.S.A.

Myrmicinae Crematogastrini Crematogaster laeviuscula TRUE TRUE FALSE

laeviuscula. Crematogaster laeviuscula Mayr, 1870b: 993 (w.) U.S.A. Wheeler, W.M. 1908e: 480 (q.); Wheeler, G.C. & Wheeler, J. 1952b: 255 (l.); Petralia & Vinson, 1980: 383 (l.). Combination in C. (Acrocoelia): Emery, 1922e: 141; in C. (Crematogaster): Buren, 1968b: 92. Subspecies of lineolata: Emery, 1895c: 284; Wheeler, W.M. 1904e: 301; Emery, 1922e: 141. Status as species: Wheeler, W.M. 1919g: 111; Creighton, 1950a: 210. Senior synonym of clara: Creighton, 1950a: 210; Johnson, C. 1988: 322; of arborea: Smith, M.R. 1951a: 809 (provisional); Buren, in Smith, M.R. 1958c: 126.

Myrmicinae Crematogastrini Crematogaster larreae TRUE TRUE FALSE

larreae. Crematogaster (Crematogaster) larreae Buren, 1968b: 117 (w.q.m.) U.S.A.

Myrmicinae Crematogastrini Crematogaster lineolata TRUE TRUE FALSE

lineolata. Myrmica lineolata Say, 1836: 290 (w.q.m.) U.S.A. Wheeler, G.C. & Wheeler, J. 1952b: 250 (l.). Combination in Crematogaster: Roger, 1863b: 37; in C. (Acrocoelia): Emery, 1922e: 141; in C. (Crematogaster): Buren, 1968b: 93. Senior synonym of columbiana, marylandica, novaeboracensis: Mayr, 1886d: 462; of punctulata, subopaca: Johnson, C. 1988: 319, 320. Material of the unavailable name lutescens referred here by Creighton, 1950a: 213; of the unavailable name wheldeni referred here by Brown, 1949a: 48.

Myrmicinae Crematogastrini Crematogaster marioni TRUE TRUE FALSE

marioni. Crematogaster (Crematogaster) marioni Buren, 1968b: 105 (w.) U.S.A.

Myrmicinae Crematogastrini Crematogaster minutissima TRUE TRUE FALSE

2 subspecies

minutissima. Crematogaster minutissima Mayr, 1870b: 995 (w.q.) U.S.A. Wheeler, W.M. 1908e: 484 (m.); Wheeler, G.C. & Wheeler, J. 1960b: 14 (l.). Combination in C. (Orthocrema): Emery, 1922e: 135. See also: Longino, 2003a: 84. Current subspecies: nominal plus missuriensis, smithi.

Myrmicinae Crematogastrini Crematogaster mormonum TRUE TRUE FALSE

mormonum. Crematogaster coarctata var. mormonum Wheeler, W.M. 1919g: 111 (w.) U.S.A. [First available use of Crematogaster lineolata subsp. coarctata var. mormonum Emery, 1895c: 284; unavailable name.] Combination in C. (Acrocoelia): Emery, 1922e: 141; in C. (Crematogaster): Buren, 1968b: 94. Raised to species: Creighton, 1950a: 215.

Myrmicinae Crematogastrini Crematogaster mutans TRUE TRUE FALSE

mutans. Crematogaster (Crematogaster) mutans Buren, 1968b: 115 (w.q.) U.S.A.

Myrmicinae Crematogastrini Crematogaster navajoa TRUE TRUE FALSE

navajoa. Crematogaster (Crematogaster) navajoa Buren, 1968b: 102 (w.q.) U.S.A.

Myrmicinae Crematogastrini Crematogaster nocturna TRUE TRUE FALSE

nocturna. Crematogaster (Crematogaster) nocturna Buren, 1968b: 112 (w.m.) U.S.A.

Myrmicinae Crematogastrini Crematogaster opuntiae TRUE TRUE FALSE

opuntiae. Crematogaster (Crematogaster) opuntiae Buren, 1968b: 120 (w.) U.S.A.

Myrmicinae Crematogastrini Crematogaster pilosa TRUE TRUE FALSE

pilosa. Crematogaster lineolata subsp. pilosa Emery, 1895c: 285, attributed to Pergande, (w.) U.S.A. Combination in C. (Acrocoelia): Emery, 1922e: 141; in C. (Crematogaster): Buren, 1968b: 92. Raised to species: Wheeler, W.M. 1919g: 111; Creighton, 1950a: 216; Johnson, C. 1988: 321. Senior synonym of creightoni and material of the nomen nudum subpilosa referred here: Buren, in Smith, M.R. 1958c: 127.

Myrmicinae Crematogastrini Crematogaster pinicola TRUE TRUE FALSE

pinicola. Crematogaster pinicola Deyrup & Cover, 2007: 101, figs. 1-3 (w.q.m.) U.S.A.

Myrmicinae Crematogastrini Crematogaster rifelna TRUE TRUE FALSE

rifelna. Crematogaster (Crematogaster) rifelna Buren, 1968b: 96 (w.q.m.) U.S.A.

Myrmicinae Crematogastrini Crematogaster vermiculata TRUE TRUE FALSE

vermiculata. Crematogaster vermiculata Emery, 1895c: 286 (w.) U.S.A. Wheeler, G.C. & Wheeler, J. 1960b: 13 (l.). Combination in C. (Acrocoelia): Emery, 1922e: 141; in C. (Crematogaster): Buren, 1968b: 91. Subspecies of coarctata: Creighton, 1950a: 209. Revived status as species: Buren, 1968b: 91 (in key); Johnson, C. 1988: 323. Material of the nomen nudum coachellai referred here by Brown, 1949a: 48.

Myrmicinae Crematogastrini Crematogaster minutissima missuriensis TRUE TRUE FALSE

missuriensis. Crematogaster victima subsp. missuriensis Emery, 1895c: 287, attributed to Pergande, (w.) U.S.A. Wheeler, G.C. & Wheeler, J. 1952b: 260 (l.). Combination in C. (Orthocrema): Emery, 1922e: 136. Currently subspecies of minutissima: Creighton, 1950a: 205.

Sphecomyrminae Sphecomyrmini Cretomyrma arnoldii TRUE TRUE TRUE

†arnoldii. †Cretomyrma arnoldii Dlussky, 1975: 116, fig. 128 (w.) RUSSIA (Cretaceous).

Sphecomyrminae Sphecomyrmini Cretomyrma unicornis TRUE TRUE TRUE

†unicornis. †Cretomyrma unicornis Dlussky, 1975: 116, fig. 129 (w.) RUSSIA (Cretaceous).

Myrmicinae Attini Cyphomyrmex wheeleri TRUE TRUE FALSE

wheeleri. Cyphomyrmex wheeleri Forel, 1900e: 282 (w.q.) U.S.A. Wheeler, W.M. 1907c: 726 (m.). See also: Kempf, 1966: 167.

Proceratiinae Proceratiini Discothyrea testacea TRUE TRUE FALSE

testacea. Discothyrea testacea Roger, 1863a: 177 (w.q.) U.S.A. See also: Smith, M.R. & Wing, 1955: 105.

Sphecomyrminae Sphecomyrmini Dlusskyidris zherichini TRUE TRUE TRUE

†zherichini. †Palaeomyrmex zherichini Dlussky, 1975: 118, fig. 130, pl. 7, fig. 29 (m.) RUSSIA (Cretaceous). Combination in †Dlusskyidris: Bolton, 1995b: 172.

Dolichoderinae Dolichoderini Dolichoderus antiquus TRUE TRUE TRUE

†antiquus. †Dolichoderus (Hypoclinea) antiquus Carpenter, 1930: 39, pl. 4, fig. 6; pl. 9, fig. 1; pl. 10, fig. 2 (w.q.) U.S.A. (Oligocene).

Dolichoderinae Dolichoderini Dolichoderus kohlsi TRUE TRUE TRUE

†kohlsi. †Dolichoderus kohlsi Dlussky & Rasnitsyn, 2002: 417, figs. 4-8 (q.) U.S.A. (Eocene).

Dolichoderinae Dolichoderini Dolichoderus mariae TRUE TRUE FALSE

mariae. Dolichoderus mariae Forel, 1885a: 349 (w.) U.S.A. Mayr, 1886d: 436 (q.); Wheeler, W.M. 1905d: 306 (m.); Wheeler, G.C. & Wheeler, J. 1966: 726 (l.). Combination in D. (Hypoclinea): Emery, 1894c: 229. Senior synonym of blatchleyi, davisi: Creighton, 1950a: 334. See also: Smith, D.R. 1979: 1415; Johnson, C. 1989a: 2; Mackay, 1993b: 75; Shattuck, 1994: 58.

Dolichoderinae Dolichoderini Dolichoderus plagiatus TRUE TRUE FALSE

plagiatus. Hypoclinea plagiata Mayr, 1870b: 960 (w.) U.S.A. Wheeler, W.M. 1905d: 311 (q.m.); Wheeler, G.C. & Wheeler, J. 1951: 175 (l.). Combination in Dolichoderus: Mayr, 1886d: 436; in D. (Hypoclinea): Emery, 1894c: 229. Senior synonym of borealis: Emery, 1895c: 330; of inornatus: Creighton, 1950a: 335. See also: Wheeler, G.C. & Wheeler, J. 1966: 727; Smith, D.R. 1979: 1416; Johnson, C. 1989a: 3; Mackay, 1993b: 81; Shattuck, 1994: 61.

Dolichoderinae Dolichoderini Dolichoderus pustulatus TRUE TRUE FALSE

pustulatus. Dolichoderus pustulatus Mayr, 1886d: 436 (w.q.) U.S.A. Wheeler, G.C. & Wheeler, J. 1966: 727 (l.); Mackay, 1993b: 85 (m.). Combination in D. (Hypoclinea): Emery, 1894c: 229. Subspecies of plagiatus: Wheeler, W.M. 1905d: 313. Revived status as species and senior synonym of beutenmuelleri: Creighton, 1950a: 335. See also: Smith, D.R. 1979: 1416; Johnson, C. 1989a: 3; Mackay, 1993b: 84.

Dolichoderinae Dolichoderini Dolichoderus rohweri TRUE TRUE TRUE

†rohweri. †Dolichoderus (Hypoclinea) rohweri Carpenter, 1930: 40, pl. 4, fig. 1; pl. 9, fig. 7 (q.) U.S.A. (Oligocene). See also: Mackay, 1993b: 88.

Dolichoderinae Dolichoderini Dolichoderus sibiricus TRUE TRUE FALSE

sibiricus. Dolichoderus quadripunctatus subsp. sibiricus Emery, 1889a: 442 (w.) RUSSIA. Kupyanskaya, 1990: 155 (q.m.). Combination in D. (Hypoclinea): Emery, 1894c: 229. Raised to species: Yasumatsu, 1962: 96. Senior synonym of abietis, japonicus, yoshiokae: Yasumatsu, 1941: 182; of atriceps, sinensis: Yasumatsu, 1962: 96.

Dolichoderinae Dolichoderini Dolichoderus taschenbergi TRUE TRUE FALSE

taschenbergi. Hypoclinea taschenbergi Mayr, 1866a: 498 (w.) U.S.A. Wheeler, G.C. & Wheeler, J. 1951: 173 (l.); Mackay, 1993b: 100 (q.m.). Combination in Dolichoderus: Mayr, 1886d: 437; in D. (Hypoclinea): Emery, 1894c: 229. Senior synonym of aterrimus: Creighton, 1950a: 336; of the unnecessary replacement name wheeleriella: Smith, M.R. 1951a: 834. See also: Wheeler, G.C & Wheeler, J. 1966: 728; Smith, D.R. 1979: 1416; Francoeur & Elias, 1985: 303; Johnson, C. 1989a: 4; Mackay, 1993b: 99; Shattuck, 1994: 69.

Dolichoderinae Dolichoderini Dolichoderus tauricus TRUE TRUE TRUE

†tauricus. †Dolichoderus tauricus Dlussky, 1981b: 70, fig. 52 (q.) RUSSIA (Miocene).

Armaniinae Armaniini Dolichomyrma latipes TRUE TRUE TRUE

†latipes. †Dolichomyrma latipes Dlussky, 1975: 122, fig. 134 (w.) RUSSIA (Cretaceous).

Armaniinae Armaniini Dolichomyrma longiceps TRUE TRUE TRUE

†longiceps. †Dolichomyrma longiceps Dlussky, 1975: 121, fig. 133 (w.) RUSSIA (Cretaceous).

Myrmicinae Solenopsidini Dolopomyrmex pilatus TRUE TRUE FALSE

pilatus. Dolopomyrmex pilatus Cover & Deyrup, 2007: 92, figs. 1 – 3 (w.q.m.) U.S.A. [D. pilatus Fisher & Cover, 2007: 83. Unavailable name.]

Dolichoderinae Leptomyrmecini Dorymyrmex bicolor TRUE TRUE FALSE

bicolor. Dorymyrmex pyramicus var. bicolor Wheeler, W.M. 1906d: 342 (w.q.) U.S.A. Gallardo, 1916a: 63 (q.); Crozier, 1970: 114 (k.). Combination in D. (Conomyrma): Gallardo, 1916a: 63; in D. (Biconomyrma): Kusnezov, 1952g: 430; in Conomyrma: Snelling, R.R. 1973b: 4; in Dorymyrmex: Shattuck, 1992c: 85. Subspecies of pyramicus: Creighton, 1950a: 349. Raised to species: Kusnezov, 1952g: 430; Cole, 1958b: 130. See also: Wilson, 1958b: 76; Johnson, C. 1989b: 192; Shattuck, 1994: 75.

Dolichoderinae Leptomyrmecini Dorymyrmex bossutus TRUE TRUE FALSE

bossutus. Conomyrma bossuta Trager, 1988: 15, figs. 1, 8 (w.q.) U.S.A. Combination in Dorymyrmex: Shattuck, 1992c: 85. See also: Johnson, C. 1989b: 192.

Dolichoderinae Leptomyrmecini Dorymyrmex bureni TRUE TRUE FALSE

bureni. Conomyrma bureni Trager, 1988: 19, figs. 2, 9, 19, 20 (w.q.) U.S.A. Combination in Dorymyrmex: Snelling, R.R. 1995: 7. Material of the nomen nudum edeni referable here: Trager, 1988: 19. Junior synonym of flavus: Johnson, C. 1989b: 187. Revived from synonymy: Snelling, R.R. 1995: 7.

Dolichoderinae Leptomyrmecini Dorymyrmex elegans TRUE TRUE FALSE

elegans. Conomyrma elegans Trager, 1988: 21, figs. 3, 10 (w.q.) U.S.A. Combination in Dorymyrmex: Shattuck, 1992c: 85. See also: Johnson, C. 1989b: 191.

Dolichoderinae Leptomyrmecini Dorymyrmex flavopectus TRUE TRUE FALSE

flavopectus. Dorymyrmex pyramicus subsp. flavopectus Smith, M.R. 1944d: 15 (w.) U.S.A. Combination in Conomyrma: Kusnezov, 1952g: 430; in Dorymyrmex: Shattuck, 1992c: 85. Raised to species: Kusnezov, 1952g: 430. See also: Snelling, R.R. 1973b: 4; Buren, Nickerson & Thompson, 1976: 306; Trager, 1988: 22; Johnson, C. 1989b: 192 (in key).

Dolichoderinae Leptomyrmecini Dorymyrmex flavus TRUE TRUE FALSE

flavus. Dorymyrmex flavus McCook, 1879: 188, figs. a-c (w.) U.S.A. Johnson, C. 1989b: 190 (q.m.); Cokendolpher & Francke, 1985: 349 (k.). Combination in Conomyrma: Kempf, 1972a: 80; in Dorymyrmex: Shattuck, 1992c: 85. Subspecies of pyramicus: Mayr, 1886d: 433; Forel, 1911e: 285. Junior synonym of pyramicus: Creighton, 1950a: 348; of insanus: Snelling, R.R. 1973b: 5. Revived from synonymy, revived status as species and senior synonym of nigra: Johnson, C. 1989b: 187. See also: Snelling, R.R. 1995: 5.

Dolichoderinae Leptomyrmecini Dorymyrmex grandulus TRUE TRUE FALSE

grandulus. Prenolepis (Nylanderia) parvula var. grandula Forel, 1922: 98 (w.) U.S.A. Trager, 1988: 24 (q.). Combination in Paratrechina (Nylanderia): Emery, 1925b: 222; Creighton, 1950a: 409; in Conomyrma: Trager, 1984b: 64; in Dorymyrmex: Shattuck, 1992c: 85. Junior synonym of parvula: Creighton, 1950a: 409. Revived from synonymy and raised to species: Trager, 1984: 64. See also: Johnson, C. 1989b: 192 (in key).

Dolichoderinae Leptomyrmecini Dorymyrmex insanus TRUE TRUE FALSE

insanus. Formica insana Buckley, 1866: 165 (w.q.) U.S.A. Combination in Dorymyrmex: McCook, 1879: 185; in D. (Conomyrma): Smith, M.R. 1951a: 837; in Conomyrma: Snelling, R.R. 1973b: 4; in Dorymyrmex: Shattuck, 1992c: 85. Junior synonym of pyramicus: Mayr, 1886d: 433. Revived from synonymy as subspecies of pyramicus: Santschi, 1920d: 381. Junior synonym of pyramicus: Creighton, 1950a: 348. Revived from synonymy as species: Snelling, R.R. 1973b: 5. Material of the unavailable name antillana referred here: Snelling, R.R. 1973b: 5. See also: Buren, Nickerson & Thompson, 1976: 306; Shattuck, 1994: 84; Snelling, R.R. 1995: 3.

Dolichoderinae Leptomyrmecini Dorymyrmex lipan TRUE TRUE FALSE

lipan. Dorymyrmex lipan Snelling, R.R. 1995: 8, figs. 16, 17 (w.) U.S.A.

Dolichoderinae Leptomyrmecini Dorymyrmex paiute TRUE TRUE FALSE

paiute. Dorymyrmex paiute Snelling, R.R. 1995: 9, figs. 14, 23 (w.) U.S.A.

Dolichoderinae Leptomyrmecini Dorymyrmex reginicula TRUE TRUE FALSE

reginicula. Conomyrma reginicula Trager, 1988: 27, figs. 7, 14, 17, 18 (w.q.) U.S.A. Combination in Dorymyrmex: Snelling, R.R. 1995: 7. Junior synonym of insanus: Johnson, C. 1989b: 185. Revived from synonymy: Snelling, R.R. 1995: 7.

Dolichoderinae Leptomyrmecini Dorymyrmex smithi TRUE TRUE FALSE

smithi. Dorymyrmex pyramicus var. smithi Cole, 1936b: 120 (w.) U.S.A. Combination in Conomyrma: Snelling, R.R. 1973b: 4; in Dorymyrmex: Snelling, R.R. 1995: 7. Junior synonym of pyramicus: Creighton, 1950a: 349; of insanus: Snelling, R.R. 1973b: 5. Revived from synonymy, raised to species and senior synonym of medeis: Snelling, R.R. 1995: 7.

Dolichoderinae Leptomyrmecini Dorymyrmex wheeleri TRUE TRUE FALSE

wheeleri. Conomyrma (Biconomyrma) wheeleri Kusnezov, 1952g: 438 (w.) U.S.A. Combination in Dorymyrmex: Snelling, R.R. 1995: 8. Junior synonym of insanus: Snelling, R.R. 1973b: 5. Revived from synonymy: Snelling, R.R. 1995: 8.

Dolichoderinae Leptomyrmecini Dorymyrmex exsanguis anaemicus TRUE TRUE FALSE

anaemicus. Dorymyrmex (Ammomyrma) exsanguis var. anaemica Santschi, 1922b: 368 (w.q.) ARGENTINA. Combination in Araucomyrmex: Brandão, 1991: 327.

Ecitoninae Ecitonini Eciton lucanoides TRUE TRUE FALSE

1 subspecies

lucanoides. Eciton lucanoides Emery, 1894c: 176, pl.2, fig. 1 (s.) PERU. Current subspecies: nominal plus conquistador. See also: Borgmeier, 1955: 262.

Ecitoninae Ecitonini Eciton lucanoides conquistador TRUE TRUE FALSE

conquistador. Eciton (Eciton) conquistador Weber, 1949c: 3, figs. 1-7 (s.w.l.) PANAMA. Reichensperger, 1934: 240 (q.). Currently subspecies of lucanoides: Borgmeier, 1955: 264.

Agroecomyrmecinae Agroecomyrmecini Eulithomyrmex rugosus TRUE TRUE TRUE

†rugosus. †Lithomyrmex rugosus Carpenter, 1930: 35, pl. 5, figs. 1, 3; pl. 8, fig. 2; pl. 11, fig. 3 (q.m.) U.S.A. (Oligocene). Combination in †Eulithomyrmex: Carpenter, 1935: 91.

Agroecomyrmecinae Agroecomyrmecini Eulithomyrmex striatus TRUE TRUE TRUE

†striatus. †Lithomyrmex striatus Carpenter, 1930: 36, pl. 6, fig. 1 (w.q.) U.S.A. (Oligocene). Combination in †Eulithomyrmex: Carpenter, 1935: 91.

Myrmicinae Basicerotini Eurhopalothrix floridana TRUE TRUE FALSE

floridana. Eurhopalothrix floridana Brown & Kempf, 1960: 207, fig. 34 (w.) U.S.A. Deyrup, Johnson & Davis, 1997: 183 (m.).

Dolichoderinae Leptomyrmecini Forelius mccooki TRUE TRUE FALSE

mccooki. Iridomyrmex mccooki McCook, 1880: 187 (w.) U.S.A. Junior synonym of foetida Buckley: Wheeler, W.M. 1901b: 520 and hence first available replacement name for Formica foetida Buckley, 1866: 167. [Junior primary homonym of Formica foetida Linnaeus, 1758: 582.] [Iridomyrmex maccooki Forel, 1878: 382. Nomen nudum.] Mayr, 1886d: 432 (q.m.); Forel, 1886b: xxxix (w.). Combination in Forelius: Emery, 1888d: 389. Senior synonym of andrei: Smith, D.R. 1979: 1419. See also: Cuezzo, 2000: 249; Ward, 2005: 9.

Formicinae Formicini Formica adamsi TRUE TRUE FALSE

2 subspecies

adamsi. Formica adamsi Wheeler, W.M. 1909e: 84 (w.) U.S.A. [Formica adamsi Wheeler, W.M. 1908g: 408. Nomen nudum.] Subspecies of whymperi: Wheeler, W.M. 1917a: 544; Creighton, 1950a: 509. [First available name is adamsi as this has priority over whymperi (see there): Bolton, 1995b: 191.] Current subspecies: nominal plus alpina, whymperi.

Formicinae Formicini Formica aerata TRUE TRUE FALSE

aerata. Formica aerata Francoeur, 1973: 116, figs. 183-189 (w.q.) U.S.A.

Formicinae Formicini Formica altipetens TRUE TRUE FALSE

altipetens. Formica cinerea var. altipetens Wheeler, W.M. 1913f: 399 (diagnosis in key) (w.) U.S.A. [Formica cinerea subsp. cinerea var. altipetens Wheeler, W.M. 1913f: 523 (w.q.m.); unavailable name.] Wheeler, G.C. & Wheeler, J. 1953c: 160 (l.). Subspecies of cinerea: Wheeler, W.M. 1917a: 550; Wheeler, G.C. & Wheeler, E.W. 1944: 259. Raised to species: Creighton, 1950a: 531. See also: Francoeur, 1973: 52.

Formicinae Formicini Formica archboldi TRUE TRUE FALSE

archboldi. Formica pallidefulva subsp. archboldi Smith, M.R. 1944d: 16 (w.) U.S.A. Trager, MacGown & Trager, 2007: 615 (q.m.). Combination in F. (Neoformica): Creighton, 1950a: 549. Raised to species: Creighton, 1950a: 549.

Formicinae Formicini Formica arenicola FALSE FALSE FALSE

arenicola. Formica arenicola Buckley, 1866; 160 (w.) U.S.A. Unidentifiable to genus; incertae sedis in Formica: Bolton, 1995b: 191.

Formicinae Formicini Formica aterrima TRUE TRUE FALSE

aterrima. Formica aterrima Cresson, 1865: 426 (m.) U.S.A. See also: Brown, 1947: 8.

Formicinae Formicini Formica biophilica TRUE TRUE FALSE

biophilica. Formica biophilica Trager, J.C., in Trager, MacGown & Trager, 2007: 617, figs. 4b, 5b, 6b (w.q.m.) U.S.A.

Formicinae Formicini Formica bradleyi TRUE TRUE FALSE

bradleyi. Formica bradleyi Wheeler, W.M. 1913f: 423 (w.m.) U.S.A. Wheeler, W.M. 1917a: 535 (q.); Wheeler, G.C. & Wheeler, J. 1953c: 168 (l.). Combination in F. (Raptiformica): Emery, 1925b: 259. Senior synonym of morbida: Wilson & Brown, 1955: 126. See also: Smith, D.R. 1979: 1449.

Formicinae Formicini Formica calviceps TRUE TRUE FALSE

calviceps. Formica calviceps Cole, 1954c: 164 (w.) U.S.A.

Formicinae Formicini Formica ciliata TRUE TRUE FALSE

ciliata. Formica ciliata Mayr, 1886d: 428 (q.) U.S.A. Wheeler, W.M. 1903e: 640 (w.m.). See also: Wheeler, W.M. 1913f: 452.

Formicinae Formicini Formica clara TRUE TRUE FALSE

clara. Formica rufibarbis var. clara Forel, 1886f: 206 (w.) RUSSIA. Ruzsky, 1902b: 471 (q.m.). Combination in F. (Serviformica): Emery, 1925b: 250. Subspecies of rufibarbis: Forel, 1901h: 65. Raised to species: Collingwood, 1961a: 68; Dlussky, 1967a: 77. Material of the unavailable names transversa, clara referred here by Dlussky, 1967a: 77. Senior synonym of rizadaghi: Dlussky, Soyunov & Zabelin, 1990: 139; of caucasica, lusatica: Seifert & Schultz, 2009: 263. Current subspecies: nominal plus sinae.

Formicinae Formicini Formica clarissima TRUE TRUE FALSE

clarissima. Formica (Serviformica) rufibarbis var. clarissima Emery, 1925b: 250 (w.) RUSSIA. [First available name for material formerly referred to clarior Ruzsky, 1915a: 421, above; unavailable name.] Senior synonym of ruzskyi (unnecessary replacement name): Bolton, 1995b: 193. Subspecies of subpilosa: Dlussky, 1965a: 30 (under the name ruzskyi). Raised to species: Seifert & Schultz, 2008: 79.

Formicinae Formicini Formica cockerelli TRUE TRUE TRUE

†cockerelli. †Formica cockerelli Carpenter, 1930: 57, pl. 4, fig. 3 (q.m.) U.S.A. (Oligocene).

Formicinae Formicini Formica coloradensis TRUE TRUE FALSE

coloradensis. Formica rufa subsp. coloradensis Creighton, 1940a: 1, fig. 1 (w.q.) U.S.A. [First available use of Formica truncicola subsp. integroides var. coloradensis Wheeler, W.M. 1913f: 440; unavailable name.] Subspecies of integroides: Creighton, 1950a: 489. Raised to species: Wheeler, G.C. & Wheeler, J. 1986g: 81; Mackay, Lowrie, et al. 1988: 111 (in key).

Formicinae Formicini Formica comata TRUE TRUE FALSE

comata. Formica comata Wheeler, W.M. 1909e: 85 (w.q.m.) U.S.A. See also: Wheeler, W.M. 1913f: 454.

Formicinae Formicini Formica connecticutensis FALSE FALSE FALSE

connecticutensis. Formica connecticutensis Buckley, 1866: 154 (w.q.) U.S.A. Unidentifiable but possibly a Formica; see Wheeler, W.M. 1902f: 19. Incertae sedis in Formica: Bolton, 1995b: 193.

Formicinae Formicini Formica creightoni TRUE TRUE FALSE

creightoni. Formica (Raptiformica) creightoni Buren, 1968a: 35, fig. 3 (w.) U.S.A.

Formicinae Formicini Formica curiosa TRUE TRUE FALSE

curiosa. Formica curiosa Creighton, 1935: 5, fig. 2 (w.q.) U.S.A. Senior synonym of parcipappa: Snelling, R.R. 1969b: 194.

Formicinae Formicini Formica dakotensis TRUE TRUE FALSE

dakotensis. Formica dakotensis Emery, 1893i: 652, pl. 22, fig. 5 (w.) U.S.A. Wasmann, 1902a: 4 (q.m.); Wheeler, W.M. 1913f: 462 (q.). Senior synonym of saturata, wasmanni (and its junior synonym specularis): Creighton, 1950a: 484; of montigena: Brown, 1957f: 167.

Formicinae Formicini Formica densiventris TRUE TRUE FALSE

densiventris. Formica fusca var. densiventris Viereck, 1903: 74 (w.) U.S.A. Cole, 1954a: 90 (m.). Junior synonym of subaenescens: Wheeler, W.M. 1913f: 504. Revived from synonymy as subspecies of fusca: Brown, 1947: 7. Raised to species: Creighton, 1950a: 499. Senior synonym of spicata: Cole, 1954a: 89; Cole, 1955b: 50; of rasilis: Mackay, Lowrie, et al. 1988: 114.

Formicinae Formicini Formica difficilis TRUE TRUE FALSE

difficilis. Formica rufa subsp. difficilis Emery, 1893i: 651, pl. 22, figs. 9, 14 (w.q.m.) U.S.A. Wheeler, G.C. & Wheeler, J. 1953c: 164 (l.). Raised to species: Wheeler, W.M. 1904e: 305; Wheeler, W.M. 1904f: 348. Senior synonym of consocians, habrogyna: Cole, 1943a: 389.

Formicinae Formicini Formica dirksi TRUE TRUE FALSE

dirksi. Formica dirksi Wing, 1949: 13 (q.) U.S.A.

Formicinae Formicini Formica dislocata FALSE FALSE FALSE

dislocata. Formica dislocata Say, 1836: 288 (q.m.) U.S.A. Unidentifiable to genus; incertae sedis in Formica: Bolton, 1995b: 194.

Formicinae Formicini Formica dolosa TRUE TRUE FALSE

dolosa. Formica (Neoformica) pallidefulva subsp. dolosa Buren, 1944a: 309 (w.) U.S.A. [First available use of Formica pallidefulva subsp. schaufussi var. dolosa Wheeler, W.M. 1912c: 90; unavailable name and unnecessary replacement name for meridionalis Wheeler, W.M. 1904f: 370, unavailable name; see there.] Wheeler, W.M. 1913f: 554 (q.). Subspecies of schaufussi: Creighton, 1950a: 551. Raised to species: Trager, MacGown & Trager, 2007: 619.

Formicinae Formicini Formica emeryi TRUE TRUE FALSE

emeryi. Formica emeryi Wheeler, 1913f: 419 (w.q.) U.S.A. Combination in F. (Raptiformica): Emery, 1925b: 259. Junior synonym of subnuda: Wilson & Brown, 1955: 118. Revived from synonymy: Buren, 1968a: 34.

Formicinae Formicini Formica eoptera TRUE TRUE TRUE

†eoptera. †Formica eoptera Cockerell, 1923b: 399, fig. 1 (wing) U.S.A. (Eocene).

Formicinae Formicini Formica exsectoides TRUE TRUE FALSE

exsectoides. Formica exsectoides Forel, 1886b: xxxviii (w.q.) U.S.A. Emery, 1893i: 653 (m.). Senior synonym of davisi, hesperia: Creighton, 1950a: 513. See also: Wheeler, W.M. 1913f: 481; Smith, D.R. 1979: 1456.

Formicinae Formicini Formica ferocula TRUE TRUE FALSE

ferocula. Formica ferocula Wheeler, W.M. 1913f: 461 (w.) U.S.A.

Formicinae Formicini Formica foreliana TRUE TRUE FALSE

foreliana. Formica foreliana Wheeler, W.M. 1913f: 451 (w.) U.S.A. Francoeur, 1973: 235 (q.m.). Junior synonym of gnava: Creighton, 1950a: 539. Revived from synonymy: Francoeur, 1973: 235.

Formicinae Formicini Formica fossaceps TRUE TRUE FALSE

fossaceps. Formica fossaceps Buren, 1942: 402 (w.q.m.) U.S.A.

Formicinae Formicini Formica gagatoides TRUE TRUE FALSE

gagatoides. Formica fusca var. gagatoides Ruzsky, 1904a: 289 (w.q.) RUSSIA. Ruzsky, 1915a: 423 (m.); Stärcke, 1935: 267 (footnote) (m.). Combination in F. (Serviformica): Emery, 1925b: 249. Subspecies of picea Nylander: Emery, 1925b: 249; Stitz, 1939: 368. Raised to species: Holgersen, 1942: 15; Holgersen, 1943a: 9. Junior synonym of picea Nylander: Bernard, 1967: 299. Revived from synonymy and status as species: Kutter, 1977c: 255; Collingwood, 1979: 121; Kupyanskaya, 1990: 188.

Formicinae Formicini Formica glacialis TRUE TRUE FALSE

glacialis. Formica fusca var. glacialis Wheeler, W.M. 1908f: 624 (w.q.m.) U.S.A. Combination in F. (Serviformica): Emery, 1925b: 247. Junior synonym of fusca: Wheeler, W.M. 1913f: 494; Creighton, 1950a: 532. Revived from synonymy and raised to species: Francoeur, 1973: 152. Senior synonym of fairchildi: Francoeur, 1977a: 11. Junior synonym of subsericea: Wheeler, G.C. & Wheeler, J. 1986g: 92. Revived from synonymy: Bolton, 1995b: 196.

Formicinae Formicini Formica glauca FALSE FALSE FALSE

glauca. Formica rufibarbis var. glauca Ruzsky, 1896: 70 (w.) RUSSIA. [First available use of Formica fusca subsp. rufibarbis var. glauca Ruzsky, 1895: 20; unavailable name.] Emery, 1909b: 198 (q.m.). Combination in F. (Serviformica): Emery, 1925b: 250. Subspecies of rufibarbis: Emery, 1925b: 250; of cunicularia: Pisarski, 1967: 416; Dlussky, 1967a: 74; Tarbinsky, 1976: 184. Raised to species: Agosti & Collingwood, 1987a: 59. Senior synonym of katuniensis, montaniformis, montivaga, volgensis and material of the unavailable name minor referred here: Dlussky, 1967a: 74 (note that the name montaniformis is misspelled as montanoides here). Junior synonym of cunicularia: Atanassov & Dlussky, 1992: 267. Unidentifiable taxon, incertae sedis in Formica: Seifert, 2007: 301 [Note that Seifert’s decision renders earlier taxonomy untenable].

Formicinae Formicini Formica gnava TRUE TRUE FALSE

gnava. Formica gnava Buckley, 1866: 156 (w.q.m.) U.S.A. Combination in F. (Serviformica): Emery, 1925b: 250. Subspecies of fusca: Wheeler, W.M. 1902f: 20; of rufibarbis: Wheeler, W.M. 1913f: 518; Creighton, 1950a: 539. Revived status as species: Francoeur, 1973: 238. Material of the nomen nudum subsericeoneorufibarbis (misspelled as subsericeorufibarbis by Emery, 1925b: 250) referred here by Wheeler, W.M. 1913f: 518.

Formicinae Formicini Formica grandis TRUE TRUE TRUE

†grandis. †Formica grandis Carpenter, 1930: 57, pl. 6, fig. 3 (q.) U.S.A. (Oligocene).

Formicinae Formicini Formica gynocrates TRUE TRUE FALSE

gynocrates. Formica gynocrates Snelling, R.R. & Buren, 1985: 72, figs. 7-15 (w.q.m.) U.S.A.

Formicinae Formicini Formica impexa TRUE TRUE FALSE

impexa. Formica impexa Wheeler, W.M. 1905c: 273 (w.) U.S.A. Wheeler, W.M. 1906a: 40 (q.). See also: Wheeler, W.M. 1913f: 472.

Formicinae Formicini Formica incerta TRUE TRUE FALSE

incerta. Formica (Neoformica) pallidefulva subsp. incerta Buren, 1944a: 309 (w.q.m.) U.S.A. [First available use of Formica pallidefulva subsp. schaufussi var. incerta Emery, 1893i: 655; unavailable name.] Junior synonym of nitidiventris: Creighton, 1950a: 551. Revived from synonymy and raised to species: Trager, MacGown & Trager, 2007: 621.

Formicinae Formicini Formica indianensis TRUE TRUE FALSE

indianensis. Formica indianensis Cole, 1940: 224 (w.m.) U.S.A.

Formicinae Formicini Formica integra TRUE TRUE FALSE

integra. Formica integra Nylander, 1856b: 62 (footnote) (w.) U.S.A. Mayr, 1886d: 425 (q.m.). Subspecies of rufa: Emery, 1893i: 652; Wheeler, W.M. 1908g: 406; Creighton, 1940a: 1; of truncicola: Wheeler, W.M. 1913c: 116; Wheeler, W.M. 1913f: 444; Wheeler, W.M. 1917a: 464. Revived status as species: Creighton, 1950a: 487. Senior synonym of similis: Emery, 1893i: 652; Wheeler, W.M. 1906b: 16.

Formicinae Formicini Formica integroides TRUE TRUE FALSE

integroides. Formica truncicola subsp. integroides Wheeler, W.M. 1913f: 438 (w.q.) U.S.A. [First available use of Formica rufa subsp. obscuriventris var. integroides Emery, 1893i: 649; unavailable name.] Subspecies of rufa: Creighton, 1940a: 3. Raised to species and senior synonym of subcaviceps: Creighton, 1950a: 489. Senior synonym of subfasciata: Brown, 1965d: 185.

Formicinae Formicini Formica knighti TRUE TRUE FALSE

knighti. Formica (Formica) knighti Buren, 1944a: 303 (w.) U.S.A.

Formicinae Formicini Formica laeviceps TRUE TRUE FALSE

laeviceps. Formica rufa subsp. laeviceps Creighton, 1940a: 9, fig. 1 (w.q.) U.S.A. Raised to species: Creighton, 1950a: 491.

Formicinae Formicini Formica lasioides TRUE TRUE FALSE

lasioides. Formica lasioides Emery, 1893i: 664 (w.) U.S.A. Combination in Proformica: Emery, 1912f: 100; in Formica (Proformica): Wheeler, W.M. 1913f: 539; in Formica: Francoeur, 1973: 114. Subspecies of fusca: Wheeler, W.M. 1908g: 409; of neogagates: Wheeler, W.M. 1913f: 539. Revived status as species and senior synonym of vetula: Creighton, 1950a: 457.

Formicinae Formicini Formica lauta FALSE FALSE FALSE

lauta. Formica lauta Say, 1836: 286 (q.m.) U.S.A. Combination in Camponotus: Dalla Torre, 1893: 238. Unidentifiable taxon, incertae sedis in Formica: Smith, D.R. 1979: 1467.

Formicinae Formicini Formica lepida TRUE TRUE FALSE

lepida. Formica cinerea var. lepida Wheeler, W.M. 1913f: 399 (diagnosis in key) (w.) U.S.A. [Formica cinerea subsp. cinerea var. lepida Wheeler, W.M. 1913f: 526; unavailable name.] Wheeler, W.M. 1917a: 551 (q.). Subspecies of cinerea: Wheeler, W.M. 1917a: 551; Creighton, 1950a: 531. Raised to species: Francoeur, 1973: 122.

Formicinae Formicini Formica limata TRUE TRUE FALSE

limata. Formica (Proformica) limata Wheeler, W.M. 1913f: 541 (w.) U.S.A.

Formicinae Formicini Formica lincecumii FALSE FALSE FALSE

lincecumii. Formica lincecumii Buckley, 1866: 163 (w.q.m.) U.S.A. Unidentifiable to genus; incertae sedis in Formica: Bolton, 1995b: 197. See also: Wheeler, W.M. 1902f: 22.

Formicinae Formicini Formica longipilosa TRUE TRUE FALSE

longipilosa. Formica longipilosa Francoeur, 1973: 125, figs. 208-214 (w.) U.S.A. Francoeur & Snelling, R.R. 1979: 4 (m.).

Formicinae Formicini Formica manni TRUE TRUE FALSE

manni. Formica manni Wheeler, W.M. 1913f: 420 (w.q.) U.S.A. Combination in F. (Raptiformica): Emery, 1925b: 259.

Formicinae Formicini Formica martynovi TRUE TRUE TRUE

†martynovi. †Formica martynovi Popov, 1933: 19, figs. 2, 3 (q.) RUSSIA. See also: Dlussky, 1981b: 75.

Formicinae Formicini Formica mesasiatica TRUE TRUE FALSE

mesasiatica. Formica (Coptoformica) mesasiatica Dlussky, 1964: 1030, figs. (w.q.m.) RUSSIA. Junior synonym of fukaii: Sonobe & Dlussky, 1977: 23. Revived status as species: Seifert, 2000a: 532. See also: Tarbinsky, 1976: 179.

Formicinae Formicini Formica microgyna TRUE TRUE FALSE

microgyna. Formica microgyna Wheeler, W.M. 1903e: 645, fig. 3 (w.q.m.) U.S.A. Senior synonym of recidiva: Creighton, 1950a: 504. See also: Wheeler, W.M. 1903f: 656; Wheeler, W.M. 1913f: 465; Letendre & Huot, 1972: 117.

Formicinae Formicini Formica microphthalma TRUE TRUE FALSE

microphthalma. Formica microphthalma Francoeur, 1973: 201, figs. 344-359 (w.m.) U.S.A.

Formicinae Formicini Formica moki TRUE TRUE FALSE

moki. Formica moki Wheeler, W.M. 1906d: 343 (w.) U.S.A. Cole, 1966: 25 (m.). Combination in F. (Neoformica): Wheeler, W.M. 1913f: 558. Junior synonym of occidua: Francoeur, 1973: 254. [Note that moki has seniority over occidua and hence is the valid name; see Smith, D.R. 1979: 1453.] See also: Smith, M.R. 1939f: 582.

Formicinae Formicini Formica montana TRUE TRUE FALSE

montana. Formica subpolita var. montana Wheeler, W.M. 1910g: 571 (w.) U.S.A. [First available use of Formica fusca subsp. subpolita var. montana Emery, 1893i: 663; unavailable name.] Francoeur, 1973: 70 (q.m.); Hung, 1969: 455 (k.). Raised to species: Wheeler, W.M. 1913f: 529. Senior synonym of neocinerea, rutilans: Creighton, 1950a: 534; Francoeur, 1973: 67.

Formicinae Formicini Formica morsei TRUE TRUE FALSE

morsei. Formica morsei Wheeler, W.M. 1906a: 39, pl. 4, fig. 1 (w.) U.S.A. See also: Wheeler, W.M. 1913f: 480.

Formicinae Formicini Formica mucescens TRUE TRUE FALSE

mucescens. Formica truncicola subsp. mucescens Wheeler, W.M. 1913f: 442 (w.q.m.) U.S.A. Subspecies of rufa: Creighton, 1940a: 3. Raised to species: Creighton, 1950a: 491.

Formicinae Formicini Formica neoclara TRUE TRUE FALSE

neoclara. Formica fusca var. neoclara Emery, 1893i: 661 (w.) U.S.A. Wheeler, W.M. 1913f: 509 (q.m.); Wheeler, G.C. & Wheeler, J. 1953c: 165 (l.). Combination in F. (Serviformica): Emery, 1925b: 248. Subspecies of fusca: Buren, 1944a: 301. Raised to species and material of the unavailable name lutescens referred here: Creighton, 1950a: 535. Senior synonym of pruinosa: Francoeur, 1973: 84.

Formicinae Formicini Formica neogagates TRUE TRUE FALSE

neogagates. Formica fusca var. neogagates Viereck, 1903: 74 (w.q.m.) U.S.A. [First available use of Formica fusca subsp. subpolita var. neogagates Emery, 1893i: 661; unavailable name.] Wheeler, G.C. & Wheeler, J. 1953c: 158 (l.). Combination in F. (Proformica): Wheeler, W.M. 1913f: 536. Subspecies of subpolita: Wheeler, W.M. 1908f: 625. Raised to species: Wheeler, W.M. 1913f: 536.

Formicinae Formicini Formica neorufibarbis TRUE TRUE FALSE

neorufibarbis. Formica fusca var. neorufibarbis Emery, 1893i: 660 (w.) U.S.A. Forel, 1902i: 699 (q.); Wheeler, W.M. 1917a: 547 (m.); Wheeler, G.C. & Wheeler, J. 1953c: 165 (l.). Combination in F. (Serviformica): Emery, 1925b: 248. Subspecies of fusca: Wheeler, G.C. & Wheeler, E.W. 1944: 261. Raised to species: Creighton, 1950a: 537. Senior synonym of algida, gelida: Francoeur, 1973: 215.

Formicinae Formicini Formica nepticula TRUE TRUE FALSE

nepticula. Formica nepticula Wheeler, W.M. 1905c: 270 (w.q.m.) U.S.A. See also: Wheeler, W.M. 1913f: 475.

Formicinae Formicini Formica nevadensis TRUE TRUE FALSE

nevadensis. Formica microgyna var. nevadensis Wheeler, W.M. 1904f: 373 (q.) U.S.A. Cole, 1956f: 256 (w.). Raised to species: Wheeler, W.M. 1905c: 272; Cole, 1956f: 257.

Formicinae Formicini Formica nortonii FALSE FALSE FALSE

nortonii. Formica nortonii Buckley, 1866: 153 (w.q.) U.S.A. Unidentifiable to genus; incertae sedis in Formica: Bolton, 1995b: 198. See also: Wheeler, W.M. 1902f: 19.

Formicinae Formicini Formica novaeanglae FALSE FALSE FALSE

novaeanglae. Formica novaeanglae Buckley, 1866: 153 (w.q.) U.S.A. Unidentifiable to genus; incertae sedis in Formica: Bolton, 1995b: 199.

Formicinae Formicini Formica obscuripes TRUE TRUE FALSE

obscuripes. Formica rufa r. obscuripes Forel, 1886b: xxxix (w.) U.S.A. Emery, 1893i: 650 (q.m.); Wheeler, G.C. & Wheeler, J. 1953c: 165 (l.); Hung, 1969: 456 (k.). Raised to species: Creighton, 1950a: 492. Senior synonym of aggerans: Forel, 1914c: 619; Creighton, 1940a: 1; of melanotica: Creighton, 1950a: 492. Material of the unavailable name rubiginosa referred here by Creighton, 1940a: 1. See also: Weber, 1935: 165.

Formicinae Formicini Formica obscuriventris TRUE TRUE FALSE

1 subspecies

obscuriventris. Formica truncicola var. obscuriventris Mayr, 1870b: 951 (w.) U.S.A. Wheeler, W.M. 1913f: 446 (q.m.). Subspecies of rufa: Mayr, 1886d: 426; Emery, 1893i: 649; Wheeler, W.M. 1908g: 408; Creighton, 1940a: 1; of truncicola: Wheeler, W.M. 1913c: 116; Wheeler, W.M. 1913f: 445; Wheeler, W.M. 1917i: 464. Raised to species: Creighton, 1950a: 493. Senior synonym of dryas: Wheeler, W.M. 1913f: 446; of gymnomma: Creighton, 1950a: 493. Current subspecies: nominal plus clivia.

Formicinae Formicini Formica obtusopilosa TRUE TRUE FALSE

obtusopilosa. Formica obtusopilosa Emery, 1893i: 648 (w.) U.S.A. Combination in F. (Raptiformica): Emery, 1925b: 259. Subspecies of sanguinea: Wheeler, W.M. 1901c: 713; Wheeler, W.M. 1913f: 414. Revived status as species: Emery, 1925b: 259; Cole, 1942: 377. Senior synonym of munda: Emery, 1925b: 259; Creighton, 1950a: 465; of alticola: Wilson & Brown, 1955: 128.

Formicinae Formicini Formica occidentalis FALSE FALSE FALSE

occidentalis. Formica occidentalis Buckley, 1866: 157 (w.q.m.) U.S.A. Unidentifiable to genus; incertae sedis in Formica: Bolton, 1995b: 200. See also: Emery, 1895c: 337; Wheeler, W.M. 1902f: 20.

Formicinae Formicini Formica occulta TRUE TRUE FALSE

occulta. Formica occulta Francoeur, 1973: 94, figs. 132-147 (w.q.m.) U.S.A.

Formicinae Formicini Formica omnivora FALSE FALSE FALSE

omnivora. Formica omnivora Linnaeus, 1758: 581 (w.) U.S.A. Unidentifiable to genus; incertae sedis in Formica: Bolton, 1995b: 200. [Wheeler, W.M. 1908b: 160, gives a combination in Monomorium but there is no evidence that this is correct. See also note under pusilla.]

Formicinae Formicini Formica opaciventris TRUE TRUE FALSE

opaciventris. Formica exsectoides var. opaciventris Emery, 1893i: 653 (w.m.) U.S.A. Gregg, 1952b: 14 (q.). Subspecies of exsectoides: Wheeler, W.M. 1913f: 484. Raised to species: Creighton, 1950a: 514.

Formicinae Formicini Formica oreas TRUE TRUE FALSE

1 subspecies

oreas. Formica oreas Wheeler, W.M. 1903e: 643, fig. 2 (w.q.m.) U.S.A. Current subspecies: nominal plus comptula. See also: Wheeler, W.M. 1913f: 458.

Formicinae Formicini Formica oregonensis TRUE TRUE FALSE

oregonensis. Formica oregonensis Cole, 1938c: 368 (w.) U.S.A. Combination in F. (Raptiformica): Creighton, 1950a: 465. Junior synonym of manni: Wilson & Brown, 1955: 127. Revived from synonymy: Cole, 1956d: 212.

Formicinae Formicini Formica pacifica TRUE TRUE FALSE

pacifica. Formica pacifica Francoeur, 1973: 131, figs. 190-197 (w.q.) U.S.A.

Formicinae Formicini Formica pallidefulva TRUE TRUE FALSE

pallidefulva. Formica pallidefulva Latreille, 1802c: 174 (w.) U.S.A. Mayr, 1866b: 889 (q.); Emery, 1893i: 657 (m.). Combination in F. (Neoformica): Wheeler, W.M. 1913a: 82. Senior synonym of succinea: Creighton, 1950a: 550; of nitidiventris, schaufussi: Trager, MacGown & Trager, 2007: 624. See also: Wheeler, W.M. 1913f: 548.

Formicinae Formicini Formica pergandei TRUE TRUE FALSE

pergandei. Formica pergandei Emery, 1893i: 646, pl. 22, fig. 1 (w.) U.S.A. Combination in F. (Raptiformica): Emery, 1925b: 259. Junior synonym of subintegra: Wilson & Brown, 1955: 120. Revived from synonymy and senior synonym of sublucida: Buren, 1968a: 28. See also: Snelling, R.R. & Buren, 1985: 72.

Formicinae Formicini Formica perpilosa TRUE TRUE FALSE

perpilosa. Formica perpilosa Wheeler, W.M. 1913f: 421 (w.q.m.) U.S.A. [First available use of Formica fusca subsp. subpolita var. perpilosa Wheeler, W.M. 1902b: 141; unavailable name.] Combination in F. (Raptiformica): Emery, 1925b: 259.

Formicinae Formicini Formica planipilis TRUE TRUE FALSE

planipilis. Formica rufa subsp. planipilis Creighton, 1940a: 9, fig. 1 (w.) U.S.A. Cole, 1956f: 259 (q.). Subspecies of integroides: Creighton, 1950a: 490. Raised to species: Cole, 1956f: 259.

Formicinae Formicini Formica politurata FALSE FALSE FALSE

politurata. Formica politurata Buckley, 1866: 160 (w.) U.S.A. Unidentifiable to genus; incertae sedis in Formica: Bolton, 1995b: 200.

Formicinae Formicini Formica postoculata TRUE TRUE FALSE

postoculata. Formica postoculata Kennedy & Dennis, 1937: 540, figs. 16-18 (w.) U.S.A.

Formicinae Formicini Formica prociliata TRUE TRUE FALSE

prociliata. Formica prociliata Kennedy & Dennis, 1937: 531, figs. 1-9 (w.q.m.) U.S.A.

Formicinae Formicini Formica propinqua TRUE TRUE FALSE

propinqua. Formica rufa subsp. propinqua Creighton, 1940a: 1, fig. 1 (w.) U.S.A. [First available use of Formica truncicola subsp. integroides var. propinqua Wheeler, W.M. 1917a: 538; unavailable name.] Cole, 1956f: 257 (q.m.). Subspecies of integroides: Creighton, 1950a: 490. Raised to species: Wheeler, G.C. & Wheeler, J. 1986g: 85; Mackay, Lowrie, et al. 1988: 112 (in key).

Formicinae Formicini Formica puberula TRUE TRUE FALSE

puberula. Formica sanguinea subsp. puberula Emery, 1893i: 648 (w.) U.S.A. Wheeler, W.M. 1913f: 413 (q.m.). Combination in F. (Raptiformica): Emery, 1925b: 260. Raised to species: Creighton, 1950a: 468. Junior synonym of subintegra: Wilson & Brown, 1955: 120. Revived from synonymy: Buren, 1968a: 30.

Formicinae Formicini Formica querquetulana TRUE TRUE FALSE

querquetulana. Formica querquetulana Kennedy & Dennis, 1937: 536, figs. 10-15 (w.q.) U.S.A.

Formicinae Formicini Formica ravida TRUE TRUE FALSE

ravida. Formica rufa subsp. ravida Creighton, 1940a: 1, fig. 1 (w.q.m.) U.S.A. [First available use of Formica truncicola subsp. integroides var. ravida Wheeler, W.M. 1913f: 560; unavailable name.] Subspecies of obscuripes: Creighton, 1950a: 493. Junior synonym of haemorrhoidalis Creighton: Brown, 1965d: 181. [F. haemorrhoidalis Creighton, 1940a: 1, junior primary homonym of F. haemorrhoidalis Latreille, 1802c: 276.] F. ravida designated as first available replacement name and senior synonym of tahoensis: Bolton, 1995b: 202.

Formicinae Formicini Formica reflexa TRUE TRUE FALSE

reflexa. Formica reflexa Buren, 1942: 399 (w.q.m.) U.S.A.

Formicinae Formicini Formica robusta TRUE TRUE TRUE

†robusta. †Formica robusta Carpenter, 1930: 56, pl. 6, fig. 6; pl. 7, fig. 6 (q.m.) U.S.A. (Oligocene).

Formicinae Formicini Formica rubicunda TRUE TRUE FALSE

rubicunda. Formica sanguinea subsp. rubicunda Emery, 1893i: 647, pl. 22, fig. 2 (w.q.) U.S.A. Wheeler, W.M. 1913f: 407 (m.). Combination in F. (Raptiformica): Emery, 1925b: 260. Raised to species: Creighton, 1950a: 468. Junior synonym of subintegra: Wilson & Brown, 1955: 120. Revived from synonymy: Buren, 1968a: 30.

Formicinae Formicini Formica saccharivora FALSE FALSE FALSE

saccharivora. Formica saccharivora Linnaeus, 1758: 580 (w.) U.S.A. Unidentifiable to genus; incertae sedis in Formica: Bolton, 1995b: 203.

Formicinae Formicini Formica saxicola FALSE FALSE FALSE

saxicola. Formica saxicola Buckley, 1866: 166 (w.q.) U.S.A. Unidentifiable to genus; incertae sedis in Formica: Bolton, 1995b: 203. See also: Emery, 1895c: 338; Wheeler, W.M. 1902f: 23.

Formicinae Formicini Formica scitula TRUE TRUE FALSE

scitula. Formica microgyna subsp. scitula Wheeler, W.M. 1913f: 470 (q.) U.S.A. Raised to species: Creighton, 1950a: 507.

Formicinae Formicini Formica sibylla TRUE TRUE FALSE

sibylla. Formica sibylla Wheeler, W.M. 1913f: 530 (w.m.) U.S.A. See also: Wheeler, W.M. 1917a: 551; Francoeur, 1973: 263.

Formicinae Formicini Formica spatulata TRUE TRUE FALSE

spatulata. Formica microgyna subsp. spatulata Buren, 1944a: 305 (w.q.m.) U.S.A. Raised to species: Creighton, 1950a: 508.

Formicinae Formicini Formica subaenescens TRUE TRUE FALSE

subaenescens. Formica fusca var. subaenescens Emery, 1893i: 659 (w.) U.S.A. Wheeler, W.M. 1913f: 504 (q.m.). Subspecies of fusca: Buren, 1944a: 300. Junior synonym of fusca: Creighton, 1950a: 532; Francoeur, 1973: 189; Wheeler, G.C. & Wheeler, J. 1986g: 16. Revived from synonymy and raised to species: Francoeur, 1977b: 208; Bolton, 1995b: 204.

Formicinae Formicini Formica subelongata TRUE TRUE FALSE

subelongata. Formica subelongata Francoeur, 1973: 268, figs. 482-485 (w.) U.S.A.

Formicinae Formicini Formica subintegra TRUE TRUE FALSE

subintegra. Formica sanguinea subsp. subintegra Wheeler, W.M. 1908f: 627 (w.q.) U.S.A. [First available use of Formica sanguinea subsp. rubicunda var. subintegra Emery, 1893i: 648, pl. 22, fig. 3; unavailable name.] Wheeler, W.M. 1913f: 411 (m.); Hung, 1969: 456 (k.). Combination in F. (Raptiformica): Emery, 1925b: 260. Subspecies of sanguinea: Wheeler, W.M. 1913f: 410. Raised to species and material of the unavailable name gilvescens referred here: Creighton, 1950a: 470. See also: Talbot & Kennedy, 1940: 560.

Formicinae Formicini Formica subnitens TRUE TRUE FALSE

subnitens. Formica rufa subsp. subnitens Creighton, 1940a: 10, fig. 1 (w.) U.S.A. Miller, 1957: 255 (q.m.). Subspecies of integroides: Creighton, 1950a: 490. Raised to species: Cole, 1955b: 50; Miller, 1957: 253; Mackay, Lowrie, et al. 1988: 111 (in key).

Formicinae Formicini Formica subpilosa TRUE TRUE FALSE

subpilosa. Formica rufibarbis subsp. subpilosa Ruzsky, 1902b: 472 (w.) RUSSIA. Karavaiev, 1910b: 17 (q.m.). Combination in F. (Serviformica): Emery, 1925b: 250. Raised to species: Bondroit, 1918: 51; Collingwood, 1961a: 70. Junior synonym of rufibarbis: Bernard, 1967: 297. Revived from synonymy, with status as species: Dlussky, 1965a: 28; Dlussky, 1967a: 70; Tarbinsky, 1976: 189. Senior synonym of bipilosa and material of the unavailable names clara, fallax referred here: Dlussky, 1965a: 28. Material of the unavailable name subpilosoclara referred here by Dlussky, 1967a: 70. See also: Seifert & Schultz, 2008: 79.

Formicinae Formicini Formica subpolita TRUE TRUE FALSE

subpolita. Formica fusca var. subpolita Mayr, 1886d: 426 (w.q.) U.S.A. Wheeler, W.M. 1913f: 534 (m.). Combination in F. (Serviformica): Emery, 1925b: 251. Subspecies of gagates: Dalla Torre, 1893: 199; of fusca: Emery, 1893i: 661. Raised to species: Wheeler, W.M. 1908g: 625; Wheeler, W.M. 1913f: 532. Senior synonym of flammiventris: Wheeler, W.M. 1913f: 532; Creighton, 1950a: 541; of camponoticeps, ficticia: Francoeur, 1973: 107.

Formicinae Formicini Formica subsericea TRUE TRUE FALSE

subsericea. Formica subsericea Say, 1836: 289 (w.m.) U.S.A. Junior synonym of fusca: Mayr, 1886d: 426; Creighton, 1950a: 532. Revived from synonymy as subspecies of fusca: Emery, 1893i: 659; Buren, 1944a: 300. Revived from synonymy, raised to species and senior synonym of lecontei: Francoeur, 1973: 172.

Formicinae Formicini Formica subspinosa FALSE FALSE FALSE

subspinosa. Formica (Hypochira) subspinosa Buckley, 1866: 169 (w.) U.S.A.; Unidentifiable to genus [unrecognisable and unplaceable taxon] incertae sedis in Formica: Bolton, 1995b: 204. [May possibly be a dolichoderine, see: Emery, 1895c: 338; Emery, 1925b: 271; Smith, D.R. 1979: 1422.]

Formicinae Formicini Formica talbotae TRUE TRUE FALSE

talbotae. Formica talbotae Wilson, 1977: 277, fig. 1 (q.m.) U.S.A. See also: Talbot, 1977: 282.

Formicinae Formicini Formica tenuissima FALSE FALSE FALSE

tenuissima. Formica tenuissima Buckley, 1866: 162 (w.) U.S.A. Unidentifiable to genus; incertae sedis in Formica: Bolton, 1995b: 205. See also: Mayr, 1886d: 432; Emery, 1895c: 338; Wheeler, W.M. 1902f: 21.

Formicinae Formicini Formica transkaucasica FALSE FALSE FALSE

transkaucasica. Formica transkaucasica Nasonov, 1889: 21 (w.) RUSSIA. Subspecies of gagates: Ruzsky, 1902d: 13; of fusca: Ruzsky, 1905b: 384. Junior synonym of picea Nylander: Emery, 1909b: 195; Donisthorpe, 1918a: 9; Emery, 1925b: 249; Dlussky, 1965a: 31; Dlussky, 1967a: 61. First available replacement name for picea Nylander: Donisthorpe, 1918a: 9; Yarrow, 1954a: 233; Atanassov & Dlussky, 1992: 263. Senior synonym of piceogagates: Arakelian, 1994: 102. Junior synonym of candida: Bolton, 1995b: 205. Unidentifiable taxon, incertae sedis in Formica: Seifert, 2004: 32.

Formicinae Formicini Formica triangularis FALSE FALSE FALSE

triangularis. Formica triangularis Say, 1836: 288 (q.m.) U.S.A. Unidentifiable to genus; incertae sedis in Formica: Bolton, 1995b: 205.

Formicinae Formicini Formica ulkei TRUE TRUE FALSE

ulkei. Formica ulkei Emery, 1893i: 653, pl. 22, fig. 7 (w.) U.S.A. Wheeler, W.M. 1913f: 486 (q.m.); Wheeler, G.C. & Wheeler, J. 1953c: 168 (l.); Hung, 1969: 456 (k.). Senior synonym of hebescens: Creighton, 1950a: 515. See also: Smith, D.R. 1979: 1456.

Formicinae Formicini Formica uralensis TRUE TRUE FALSE

uralensis. Formica uralensis Ruzsky, 1895: 13 (w.q.m.) RUSSIA. [Also described as new by Ruzsky, 1896: 69.] Senior synonym of superba: Wu, 1990: 4. See also: Emery, 1909b: 189; Wheeler, W.M. 1913f: 448; Karavaiev, 1936: 249; Stitz, 1939: 325; Dlussky, 1967a: 79; Kutter, 1977c: 274; Collingwood, 1979: 135; Kupyanskaya, 1990: 189.

Formicinae Formicini Formica vinculans TRUE TRUE FALSE

vinculans. Formica (Proformica) neogagates var. vinculans Wheeler, W.M. 1913f: 400 (in key) (w.q.) U.S.A. [Formica (Proformica) neogagates subsp. neogagates var. vinculans Wheeler, W.M. 1913f: 539; unavailable name.] Subspecies of neogagates: Buren, 1944a: 309. Junior synonym of neogagates: Creighton, 1950a: 459. Revived from synonymy and raised to species: Snelling, R.R. & Buren, 1985: 76.

Formicinae Formicini Formica virginiana FALSE FALSE FALSE

virginiana. Formica virginiana Buckley, 1866: 159 (w.) U.S.A. Unidentifiable to genus; incertae sedis in Formica: Bolton, 1995b: 206.  See also: Emery, 1895c: 337; Wheeler, W.M. 1902f: 20.

Formicinae Formicini Formica volgensis FALSE FALSE FALSE

volgensis. Formica rufibarbis subsp. volgensis Ruzsky, 1914c: 323, fig. (w.) RUSSIA. Junior synonym of glauca: Dlussky, 1967a: 74. Unidentifiable taxon: Seifert & Schultz, 2009: 271.

Formicinae Formicini Formica wheeleri TRUE TRUE FALSE

wheeleri. Formica wheeleri Creighton, 1935: 1, fig. 1 (w.q.) U.S.A. Combination in F. (Raptiformica): Creighton, 1950a: 472.

Formicinae Formicini Formica xerophila TRUE TRUE FALSE

xerophila. Formica (Neoformica) moki subsp. xerophila Smith, M.R. 1939f: 583 (w.) U.S.A. Raised to species and senior synonym of grundmanni: Francoeur, 1973: 259.

Formicinae Formicini Formica rufa tshugunovi TRUE TRUE FALSE

tshugunovi. Formica rufa var. tshugunovi Ruzsky, 1914b: 102 (w.) RUSSIA. Currently subspecies of rufa: Ruzsky, 1926: 110.

Formicinae Formicini Formica rufibarbis subpilosorufibarbis TRUE TRUE FALSE

subpilosorufibarbis. Formica rufibarbis var. subpilosorufibarbis Ruzsky, 1905b: 398 (w.) RUSSIA.

Formicinae Formicini Formica rufa rufobrevisetosa TRUE TRUE FALSE

rufobrevisetosa. Formica rufa var. rufobrevisetosa Ruzsky, 1926: 110 (w.) RUSSIA.

Formicinae Formicini Formica pratensis nuda TRUE TRUE FALSE

nuda. Formica pratensis subsp. nuda Ruzsky, 1926: 110 (w.) RUSSIA.

Formicinae Formicini Formica fusca hyrcana TRUE TRUE FALSE

hyrcana. Formica (Serviformica) fusca subsp. hyrcana Arnol'di, 1968: 1821 (w.q.) RUSSIA. [Formica fusca subsp. hyrcana Arnol'di, 1948: 212. Nomen nudum.]

Formicinae Formicini Formica rufa constricta TRUE TRUE FALSE

constricta. Formica (Formica) rufa var. constricta Karavaiev, 1929b: 215, fig. 6 (w.) RUSSIA.

Formicinae Formicini Formica oreas comptula TRUE TRUE FALSE

comptula. Formica oreas var. comptula Wheeler, W.M. 1913f: 460 (w.q.) U.S.A. Currently subspecies of oreas: Creighton, 1950a: 494.

Formicinae Formicini Formica obscuriventris clivia TRUE TRUE FALSE

clivia. Formica rufa subsp. clivia Creighton, 1940a: 8, fig. 1 (w.q.m.) U.S.A. Currently subspecies of obscuriventris: Creighton, 1950a: 494.

Formicinae Formicini Formica rufibarbis clarorufibarbis TRUE TRUE FALSE

clarorufibarbis. Formica rufibarbis var. clarorufibarbis Wheeler, W.M. & Mann, 1916: 173 (w.) RUSSIA. [First available use of Formica rufibarbis subsp. clara var. clarorufibarbis Ruzsky, 1905b: 400; unavailable name.]

Formicinae Formicini Formica cinerea cinereoimitans TRUE TRUE FALSE

cinereoimitans. Formica cinerea var. cinereoimitans Ruzsky, 1905b: 405 (w.) RUSSIA. See also comment in Seifert, 2002b: 267.

Formicinae Formicini Formica rufa brevisetosa TRUE TRUE FALSE

brevisetosa. Formica rufa subsp. brevisetosa Ruzsky, 1926: 110 (w.) RUSSIA.

Formicinae Formicini Formica adamsi alpina TRUE TRUE FALSE

alpina. Formica adamsi var. alpina Wheeler, W.M. 1909e: 85 (w.) U.S.A. Subspecies of whymperi: Wheeler, W.M. 1917a: 544; Creighton, 1950a: 509; Smith, D.R. 1979: 1463. Relationship with densiventris: Cole, 1954a: 89. Subspecies of adamsi: Bolton, 1995b: 191.

Myrmicinae Formicoxenini Formicoxenus chamberlini TRUE TRUE FALSE

chamberlini. Symmyrmica chamberlini Wheeler, W.M. 1904a: 5, pl. 1, figs. 1-7 (w.q.m.) U.S.A. Combination in Formicoxenus: Francoeur, Loiselle & Buschinger, 1985: 379. See also: Buschinger & Francoeur, 1983: 297.

Myrmicinae Formicoxenini Formicoxenus diversipilosus TRUE TRUE FALSE

diversipilosus. Leptothorax (Mychothorax) diversipilosus Smith, M.R. 1939c: 179 (w.q.) U.S.A. Smith, M.R. 1956c: 274 (q.m.); Wheeler, G.C. & Wheeler, J. 1989a: 321 (l.). Combination in Formicoxenus: Buschinger, 1979a: 61. See also: Francoeur, Loiselle & Buschinger, 1985: 381.

Myrmicinae Formicoxenini Formicoxenus hirticornis TRUE TRUE FALSE

hirticornis. Leptothorax (Leptothorax) hirticornis Emery, 1895c: 319 (w.) U.S.A. Smith, M.R. 1939c: 176 (q. ergatoid); Snelling, R.R. 1965a: 18 (q.m. ergatoid). Combination in L. (Mychothorax): Emery, 1924d: 262; in Mychothorax: Ruzsky, 1936: 96; in Formicoxenus: Buschinger, 1979a: 61. Senior synonym of formidolosus: Smith, M.R. 1939c: 177. See also: Wheeler, W.M. 1903c: 224; Francoeur, Loiselle & Buschinger, 1985: 382.

Myrmicinae Formicoxenini Formicoxenus sibiricus TRUE TRUE FALSE

sibiricus. Leptothorax sibiricus Forel, 1899b: 306 (w.) RUSSIA. Combination in L. (Mychothorax): Emery, 1924d: 262; in Formicoxenus: Francoeur, Loiselle & Buschinger, 1985: 382. Junior synonym of hirticornis: Ruzsky, 1905b: 622. Revived from synonymy as subspecies of hirticornis: Emery, 1924d: 262. Revived status as species and senior synonym of orientalis: Francoeur, Loiselle & Buschinger, 1985: 382.

Ectatomminae Ectatommini Gnamptogenys hartmani TRUE TRUE FALSE

hartmani. Ectatomma (Parectatomma) hartmani Wheeler, W.M. 1915b: 390 (w.) U.S.A. Brown, 1961b: 69 (q.); Wheeler, G.C. & Wheeler, J. 1964b: 450 (l.). Combination in Gnamptogenys: Brown, 1958g: 228. Senior synonym of nigrifrons, turmalis: Lattke, 1995: 169.

Ponerinae Ponerini Hypoponera inexorata TRUE TRUE FALSE

1 subspecies

inexorata. Ponera inexorata Wheeler, W.M. 1903b: 94, fig. 2 (w.q.) U.S.A. Combination in Hypoponera: Taylor, 1968a: 65. Current subspecies: nominal plus fallax.

Dolichoderinae Leptomyrmecini Iridomyrmex florissantius FALSE FALSE TRUE

†florissantius. †Iridomyrmex florissantius Carpenter, 1930: 50, pl. 2, fig. 4; pl. 10, fig. 7 (q.) U.S.A. (Oligocene). Provisionally retained in Iridomyrmex: Shattuck, 1992a: 15. Unidentifiable taxon, incertae sedis in Iridomyrmex: Heterick & Shattuck, 2011: 168.

Dolichoderinae Leptomyrmecini Iridomyrmex mapesi FALSE FALSE TRUE

†mapesi. †Iridomyrmex mapesi Wilson, 1985d: 211, figs. 4, 5 (w.) U.S.A. Provisionally retained in Iridomyrmex: Shattuck, 1992a: 16. Unidentifiable taxon, incertae sedis in Iridomyrmex: Heterick & Shattuck, 2011: 168.

Dolichoderinae Leptomyrmecini Iridomyrmex obscurans FALSE FALSE TRUE

†obscurans. †Iridomyrmex obscurans Carpenter, 1930: 51 (q.) U.S.A. (Oligocene). Provisionally retained in Iridomyrmex: Shattuck, 1992a: 16. Unidentifiable taxon, incertae sedis in Iridomyrmex: Heterick & Shattuck, 2011: 168.

Armaniinae Armaniini Khetania mandibulata TRUE TRUE TRUE

†mandibulata. †Khetania mandibulata Dlussky, 1999a: 65, fig. 1 (q.) RUSSIA (Cretaceous).

Formicinae Lasiini Lasius arizonicus TRUE TRUE FALSE

arizonicus. Lasius (Acanthomyops) interjectus subsp. arizonicus Wheeler, W.M. 1917a: 532 (w.) U.S.A. Wing, 1968: 90 (q.m.).Combination in Acanthomyops: Creighton, 1950a: 431; in Lasius: Ward, 2005: 13. Raised to species: Buren, 1950: 184. See also: Wing, 1968: 90.

Formicinae Lasiini Lasius atopus TRUE TRUE FALSE

atopus. Lasius (Chthonolasius) atopus Cole, 1958a: 75, fig. 1 (w.) U.S.A.

Formicinae Lasiini Lasius bureni TRUE TRUE FALSE

bureni. Acanthomyops bureni Wing, 1968: 135, figs. 159-167 (w.q.m.) U.S.A. Combination in Lasius: Ward, 2005: 13.

Formicinae Lasiini Lasius californicus TRUE TRUE FALSE

californicus. Lasius (Acanthomyops) interjectus subsp. californicus Wheeler, W.M. 1917a: 531 (w.q.) U.S.A. Wing, 1968: 85 (m.). Combination in Acanthomyops: Creighton, 1950a: 430; in Lasius: Ward, 2005: 13. Subspecies of claviger: Creighton, 1950a: 430. Raised to species: Buren, 1950: 185. See also: Wing, 1968: 85.

Formicinae Lasiini Lasius capitatus TRUE TRUE FALSE

capitatus. Acanthomyops fuliginosus subsp. capitatus Kuznetsov-Ugamsky, 1927e: 187 (w.) RUSSIA. [Also described as new by Kuznetsov-Ugamsky, 1928b: 18.] Combination in Lasius: Wilson, 1955a: 138. Junior synonym of fuliginosus: Wilson, 1955a: 138. Revived from synonymy and raised to species: Kupyanskaya, 1989: 785. See also: Radchenko, 2005a: 85.

Formicinae Lasiini Lasius claviger TRUE TRUE FALSE

claviger. Formica clavigera Roger, 1862a: 241, pl. 1, fig. 13 (q.) U.S.A. Mayr, 1870b: 950 (w.m.); Wheeler G.C. & Wheeler, J. 1953c: 155 (l.). Combination in Acanthomyops: Mayr, 1862: 700; in Lasius: Mayr, 1870b: 950; in Lasius (Acanthomyops): Emery, 1893i: 642; in Acanthomyops: Donisthorpe, 1916c: 276; in Lasius: Ward, 2005: 13. Senior synonym of parvula: Wing, 1968: 67. See also: Creighton, 1950a: 429; Regnier & Wilson, 1968: 955; Smith, D.R. 1979: 1440.

Formicinae Lasiini Lasius colei TRUE TRUE FALSE

colei. Acanthomyops colei Wing, 1968: 88, figs. 64-67 (w.m.) U.S.A. Combination in Lasius: Ward, 2005: 13.

Formicinae Lasiini Lasius coloradensis TRUE TRUE FALSE

coloradensis. Lasius (Acanthomyops) interjectus subsp. coloradensis Wheeler, W.M. 1917a: 532 (w.q.m.) U.S.A. Combination in Acanthomyops: Creighton, 1950a: 429; in Lasius: Ward, 2005: 13. Subspecies of claviger: Creighton, 1950a: 429; Raised to species: Buren, 1950: 185. See also: Wing, 1968: 78.

Formicinae Lasiini Lasius creightoni TRUE TRUE FALSE

creightoni. Acanthomyops creightoni Wing, 1968: 141, figs. 177-185 (w.q.m.) U.S.A. Combination in Lasius: Ward, 2005: 13.

Formicinae Lasiini Lasius crypticus TRUE TRUE FALSE

crypticus. Lasius (Lasius) crypticus Wilson, 1955a: 104 (w.q.m.) U.S.A.

Formicinae Lasiini Lasius fallax TRUE TRUE FALSE

fallax. Lasius (Cautolasius) fallax Wilson, 1955a: 130 (w.q.m.) U.S.A.

Formicinae Lasiini Lasius humilis TRUE TRUE FALSE

humilis. Lasius (Formicina) humilis Wheeler, W.M. 1917a: 528 (w.q.) U.S.A. Combination in L. (Chthonolasius): Emery, 1925b: 233. See also: Wilson, 1955a: 185.

Formicinae Lasiini Lasius interjectus TRUE TRUE FALSE

interjectus. Lasius interjectus Mayr, 1866b: 888, pl. 20, fig. 3 (q.) U.S.A. Emery, 1893i: 642 (w.); Wing, 1968: 93 (m.); Wheeler, G.C. & Wheeler, J. 1953c: 156 (l.). Combination in Lasius (Acanthomyops): Emery, 1893i: 642; Wheeler, W.M. 1917a: 531; in Acanthomyops: Creighton, 1950a: 430; in Lasius: Ward, 2005: 13. See also: Wing, 1968: 92; Smith, D.R. 1979: 1441.

Formicinae Lasiini Lasius latipes TRUE TRUE FALSE

latipes. Formica latipes Walsh, 1863: 311 (w.q.m.) U.S.A. Combination in Lasius: Mayr, 1866b: 889; in Lasius (Acanthomyops): Emery, 1893i: 642; in Acanthomyops: Creighton, 1950a: 431; in Lasius: Ward, 2005: 13. See also: Wheeler, W.M. & McClendon, 1903: 149; Wing, 1968: 98; Smith, D.R. 1979: 1441.

Formicinae Lasiini Lasius minutus TRUE TRUE FALSE

minutus. Lasius umbratus subsp. minutus Emery, 1893i: 641 (w.q.m.) U.S.A. Combination in L. (Chthonolasius): Ruzsky, 1914a: 61. Subspecies of bicornis: Emery, 1925b: 233. Raised to species: Wilson, 1955a: 180. See also: Wheeler, W.M. 1910e: 241; Smith, D.R. 1979: 1483.

Formicinae Lasiini Lasius monticola FALSE FALSE FALSE

monticola. Formica monticola Buckley, 1866: 157 (w.q.m.) U.S.A. Combination in Lasius: Emery, 1895c: 337; Wheeler, W.M. 1902f: 20; Emery, 1925b: 237. Unrecognisable taxon; incertae sedis in Lasius: Bolton, 1995b: 224.

Formicinae Lasiini Lasius murphyi TRUE TRUE FALSE

murphyi. Lasius (Acanthomyops) murphii Forel, 1901e: 367 (w.q.m.) U.S.A. [Justified emendation of spelling to murphyi: Wheeler, W.M. 1905f: 398.] Combination in Acanthomyops: Creighton, 1950a: 431; in Lasius: Ward, 2005: 13. See also: Wing, 1968: 113.

Formicinae Lasiini Lasius nearcticus TRUE TRUE FALSE

nearcticus. Lasius flavus subsp. nearcticus Wheeler, W.M. 1906a: 38 (w.q.m.) U.S.A. Wheeler, G.C. & Wheeler, J. 1953c: 151 (l.); Crozier, 1970: 114 (k.). Combination in L. (Chthonolasius): Creighton, 1950a: 422; in L. (Cautolasius): Wilson, 1955a: 133. Raised to species: Wilson, 1955a: 133. Senior synonym of mellea: Francoeur & Béique, 1966: 144.

Formicinae Lasiini Lasius neoniger TRUE TRUE FALSE

neoniger. Lasius niger var. neoniger Emery, 1893i: 639 (w.q.m.) U.S.A. Subspecies of niger: Wheeler, W.M. 1917a: 463; Buren, 1944a: 296; Creighton, 1950a: 420. Status as species: Gregg, E.V. 1945: 530; Wilson, 1955a: 97; Smith, D.R. 1979: 1436.

Formicinae Lasiini Lasius nevadensis TRUE TRUE FALSE

nevadensis. Lasius (Chthonolasius) nevadensis Cole, 1956a: 26 (w.q.m.) U.S.A.

Formicinae Lasiini Lasius nigrescens TRUE TRUE FALSE

nigrescens. Lasius emarginatus var. nigrescens Stitz, 1930: 240 (q.) RUSSIA. Junior synonym of niger: Wilson, 1955a: 60. Revived from synonymy and raised to species: Seifert, 1992b: 38.

Formicinae Lasiini Lasius occidentalis TRUE TRUE FALSE

occidentalis. Lasius (Acanthomyops) occidentalis Wheeler, W.M. 1909e: 83 (w.q.m.) U.S.A. Combination in Acanthomyops: Creighton, 1950a: 432; in Lasius: Ward, 2005: 13. See also: Wing, 1968: 143.

Formicinae Lasiini Lasius peritulus TRUE TRUE TRUE

†peritulus. †Tetramorium peritulum Cockerell, 1927: 165 (m.) U.S.A. (Oligocene). Combination in Lasius: Carpenter, 1930: 58. See also: Wilson, 1955a: 58.

Formicinae Lasiini Lasius plumopilosus TRUE TRUE FALSE

plumopilosus. Lasius (Acanthomyops) plumopilosus Buren, 1941: 231, fig. 1 (w.q.m.) U.S.A. Combination in Acanthomyops: Creighton, 1950a: 433; in Lasius: Ward, 2005: 13. See also: Wing, 1968: 132.

Formicinae Lasiini Lasius pogonogynus TRUE TRUE FALSE

pogonogynus. Lasius (Acanthomyops) pogonogynus Buren, 1950: 186 (w.q.) U.S.A. Combination in Acanthomyops: Wing, 1968: 117; in Lasius: Ward, 2005: 13. [According to Wing (1968: 117) pogonogynus is a murphyi × latipes hybrid; the name is not, however, formally synonymised.]

Formicinae Lasiini Lasius pubescens TRUE TRUE FALSE

pubescens. Lasius (Acanthomyops) pubescens Buren, 1942: 405 (w.q.) U.S.A. Combination in Acanthomyops: Creighton, 1950a: 433; in Lasius: Ward, 2005: 13. See also: Wing, 1968: 138.

Formicinae Lasiini Lasius sitiens TRUE TRUE FALSE

sitiens. Lasius (Lasius) sitiens Wilson, 1955a: 108 (w.q.m.) U.S.A.

Formicinae Lasiini Lasius speculiventris TRUE TRUE FALSE

speculiventris. Lasius speculiventris Emery, 1893i: 641 (w.m.) U.S.A. Wilson, 1955a: 172 (q.). Combination in L. (Chthonolasius): Emery, 1925b: 235. Subspecies of umbratus: Wheeler, W.M. 1910e: 237; Wheeler, W.M. 1917h: 167. Revived status as species: Creighton, 1950a: 423.

Formicinae Lasiini Lasius subglaber TRUE TRUE FALSE

subglaber. Lasius (Acanthomyops) claviger var. subglaber Emery, 1893i: 642 (w.q.m.) U.S.A. Wheeler, G.C. & Wheeler, J. 1953c: 155 (l.). Combination in Acanthomyops: Creighton, 1950a: 433; in Lasius: Ward, 2005: 13. Subspecies of claviger: Wheeler, W.M. 1917a: 533. Raised to species and senior synonym of clavigeroides: Creighton, 1950a: 433. See also: Buren, 1950: 185; Wing, 1968: 121.

Formicinae Lasiini Lasius subumbratus TRUE TRUE FALSE

subumbratus. Lasius umbratus subsp. subumbratus Viereck, 1903: 73 (q.) U.S.A. Wheeler, W.M. 1910e: 238 (w.m.). Combination in L. (Chthonolasius): Emery, 1925b: 235. Raised to species: Creighton, 1950a: 424. See also: Wheeler, W.M. 1917h: 167; Wilson, 1955a: 175.

Formicinae Lasiini Lasius tertiarius TRUE TRUE TRUE

†tertiarius. †Lasius tertiarius Zalessky, 1949: 51 (caste?) RUSSIA.

Formicinae Lasiini Lasius vestitus TRUE TRUE FALSE

vestitus. Lasius umbratus subsp. vestitus Wheeler, W.M. 1910e: 242 (q.) U.S.A. Wilson, 1955a: 174 (w.). Combination in L. (Chthonolasius): Emery, 1925b: 235. Raised to species: Creighton, 1950a: 425. Senior synonym of pilosus: Wilson, 1955a: 173.

Formicinae Lasiini Lasius vetulus TRUE TRUE TRUE

†vetulus. †Lasius vetulus Dlussky, 1981b: 72, figs. 53, 54 (q.) RUSSIA (Miocene).

Formicinae Lasiini Lasius xerophilus TRUE TRUE FALSE

xerophilus. Lasius xerophilus Mackay, W.P. & Mackay, E.E., 1994: 37, figs. 1-7, 9 (w.q.m.) U.S.A.

Formicinae Lasiini Lasius niger pinetorum TRUE TRUE FALSE

pinetorum. Lasius niger var. pinetorum Ruzsky, 1907: 68 (footnote) (w.) RUSSIA.

Formicinae Lasiini Lasius jensi longiceps TRUE TRUE FALSE

longiceps. Lasius (Chthonolasius) jensi subsp. longiceps Seifert, 1988: 158, figs. 12, 22, 31 (w.q.) RUSSIA.

Formicinae Plagiolepidini Lepisiota semenovi TRUE TRUE FALSE

semenovi. Acantholepis frauenfeldi var. semenovi Ruzsky, 1905b: 461 (w.) RUSSIA. Combination in Lepisiota: Bolton, 1995b: 228. Subspecies of frauenfeldi: Kuznetsov-Ugamsky, 1929c: 485. Raised to species: Pisarski, 1967: 410; Dlussky & Zabelin, 1985: 235. Senior synonym of arnoldovi, coriacea, deserticola, halinae, integrisquama, latisquama, litoralis, mediorubra, transcaspiensis, turkmena and material of the unavailable names infumata, minuta referred here: Dlussky, Soyunov & Zabelin, 1990: 163-165.

Ponerinae Ponerini Leptogenys elongata TRUE TRUE FALSE

elongata. Ponera elongata Buckley, 1866: 172 (w.) U.S.A. Wheeler, W.M. 1900b: 7, 16 (q.m.l.); Wheeler, W.M. 1904c: 258 (w.q.m.); Wheeler, G.C. & Wheeler, J. 1952c: 641 (l.); Petralia & Vinson, 1980: 378 (l.). Combination in Leptogenys: Emery, 1895c: 338; Wheeler, W.M. 1900b: 2. Senior synonym of septentrionalis, texana: Wheeler, W.M. 1902f: 25; Wheeler, W.M. 1904c: 257.

Ponerinae Ponerini Leptogenys manni TRUE TRUE FALSE

manni. Leptogenys (Lobopelta) elongata subsp. manni Wheeler, W.M. 1923d: 14 (w.) U.S.A. Trager & Johnson, 1988: 31 (q.m.). Raised to species: Trager & Johnson, 1988: 31.

Myrmicinae Formicoxenini Leptothorax calderoni TRUE TRUE FALSE

calderoni. Leptothorax (Mychothorax) canadensis ssp. calderoni Creighton, 1950a: 276 U.S.A. (as junior synonym of septentrionalis). [First available use of Leptothorax (Mychothorax) acervorum r. canadensis var. calderoni Forel, 1914c: 617; unavailable name.] Revived from synonymy and raised to species: Ward, 2005: 13.

Myrmicinae Formicoxenini Leptothorax crassipilis TRUE TRUE FALSE

crassipilis. Leptothorax (Mychothorax) acervorum subsp. crassipilis Wheeler, W.M. 1917a: 513 (w.q.m.) U.S.A. Raised to species: Creighton, 1950a: 278. See also: Loiselle, Francoeur, Fischer & Buschinger, 1990: 329.

Myrmicinae Formicoxenini Leptothorax oceanicus TRUE TRUE FALSE

oceanicus. Mychothorax muscorum subsp. oceanicum Kuznetsov-Ugamsky, 1928b: 29 (w.) RUSSIA. Kupyanskaya, 1990: 140 (q.). Combination in Leptothorax: Kupyanskaya, 1990: 140. Raised to species: Heinze, Schulz & Radchenko, 1993: 178; Radchenko, 1994d: 147 (in key). See also: Radchenko & Heinze, 1997: 79.

Myrmicinae Formicoxenini Leptothorax wilsoni TRUE TRUE FALSE

wilsoni. Leptothorax wilsoni Heinze, 1989: 50, figs. 1-5 (q.m.) U.S.A.

Dolichoderinae Tapinomini Liometopum luctuosum TRUE TRUE FALSE

luctuosum. Liometopum apiculatum subsp. luctuosum Wheeler, W.M. 1905e: 325 (w.) U.S.A. Forel, 1914c: 619 (m.); Del Toro, et al. 2009: 321 (q.). Subspecies of occidentale: Creighton, 1950a: 339. Raised to species: Wheeler, G.C. & Wheeler, J. 1986g: 55; Mackay, Lowrie, et al. 1988: 102.

Dolichoderinae Tapinomini Liometopum miocenicum TRUE TRUE TRUE

†miocenicum. †Liometopum miocenicum Carpenter, 1930: 46, pl. 3, fig. 6; pl. 8, fig. 3; pl. 9, fig. 8; pl. 10, fig. 8; pl. 11, fig. 7 (w.q.m.) U.S.A. (Oligocene).

Dolichoderinae Tapinomini Liometopum occidentale TRUE TRUE FALSE

occidentale. Liometopum microcephalum var. occidentale Emery, 1895c: 330 (w.m.) U.S.A. Wheeler, W.M. 1917a: 522 (q.). Subspecies of apiculatum: Wheeler, W.M. 1905e: 324. Raised to species: Wheeler, W.M. 1917a: 521. See also: Del Toro, et al. 2009: 330.

Dolichoderinae Tapinomini Liometopum orientale TRUE TRUE FALSE

orientale. Liometopum microcephalum var. orientalis Karavaiev, 1927d: 339 (w.) RUSSIA. Kupyanskaya, 1988: 31 (q.m.). Raised to species: Kupyanskaya, 1988: 29. See also: Kupyanskaya, 1990: 157; Del Toro, et al. 2009: 337.

Dolichoderinae Tapinomini Liometopum scudderi TRUE TRUE TRUE

†scudderi. †Liometopum scudderi Carpenter, 1930: 47, pl. 4, fig. 4; pl. 9, fig. 4 (w.q.) U.S.A. (Oligocene).

Myrmicinae Myrmicini Manica bradleyi TRUE TRUE FALSE

bradleyi. Myrmica bradleyi Wheeler, W.M. 1909e: 77 (w.) U.S.A. Cole, 1957c: 210 (q.m.); Wheeler, G.C. & Wheeler, J. 1960b: 6 (l.). Combination in Myrmica (Oreomyrma): Wheeler, W.M. 1914d: 120; in M. (Neomyrma): Emery, 1915d: 69 (footnote); in M. (Manica): Emery, 1921f: 43; in Manica: Weber, 1947: 440; Creighton, 1950a: 108. Senior synonym of calderoni: Wheeler, W.M. 1915a: 50.

Myrmicinae Myrmicini Manica hunteri TRUE TRUE FALSE

hunteri. Myrmica (Oreomyrma) hunteri Wheeler, W.M. 1914d: 121, fig. 1 (w.) U.S.A. Cole, 1957c: 212 (m.); Wheeler, G.C. & Wheeler, J. 1972b: 235 (l.). Combination in Myrmica (Neomyrma): Wheeler, W.M. 1917a: 507; in M. (Manica): Emery, 1921f: 43; in Manica: Weber, 1947: 440; Creighton, 1950a: 109. Senior synonym of aldrichi: Cole, 1956g: 262.

Myrmicinae Myrmicini Manica parasitica TRUE TRUE FALSE

parasitica. Myrmica (Manica) parasitica Creighton, 1934: 185 (w.) U.S.A. Combination in Manica: Weber, 1947: 440.

Myrmicinae Solenopsidini Megalomyrmex pacova TRUE TRUE FALSE

pacova. Megalomyrmex pacova Brandão, 1990: 437, figs. 46, 55 (w.) U.S.A. [Intercepted in quarantine, from Neotropics.]

Myrmicinae Pheidolini Messor andrei TRUE TRUE FALSE

andrei. Aphaenogaster andrei Mayr, 1886d: 448 (w.) U.S.A. Emery, 1895c: 306 (q.); Wheeler, G.C. & Wheeler, J. 1972b: 240 (l.); Taber & Cokendolpher, 1988: 95 (k.). Combination in Stenamma (Messor): Emery, 1895c: 306; in Novomessor: Emery, 1915d: 73; in N. (Veromessor): Forel, 1917: 235; in Veromessor: Wheeler, W.M. & Creighton, 1934: 362; in Messor: Bolton, 1982: 341. Senior synonym of castaneus (and its junior synonym flavus): Creighton, 1953a: 3. See also: Smith, D.R. 1979: 1364.

Myrmicinae Pheidolini Messor aralocaspius TRUE TRUE FALSE

1 subspecies

aralocaspius. Aphaenogaster barbara var. aralocaspia Ruzsky, 1902c: 20 (w.) RUSSIA. Karavaiev, 1910b: 63 (q.m.); Stitz, 1930: 240 (q.). Combination in Messor: Ruzsky, 1905b: 745. Subspecies of minor: Santschi, 1927c: 239. Status as species: Pisarski, 1967: 384; Pisarski, 1969b: 306; Tarbinsky, 1976: 65; Arnol'di, 1977b: 1642. Material of the unavailable name tamerlani referred here by Dlussky, Soyunov & Zabelin, 1990: 218. Current subspecies: nominal plus infumatus.

Myrmicinae Pheidolini Messor caducus TRUE TRUE FALSE

1 subspecies

caducus. Formica caduca Victor, 1839: 47, pl. 2, fig. d (w.) RUSSIA. Combination in Aphaenogaster: Roger, 1863b: 29; in Stenamma (Messor): Emery, 1898c: 125; in Messor: Ruzsky, 1905b: 754. Junior synonym of barbarus: Roger, 1863b: 29; Emery & Forel, 1879: 461; Nasonov, 1889: 37. Revived from synonymy as subspecies of barbarus: Emery, 1891b: 12. Revived status as species: Ruzsky, 1905b: 754. Material of the unavailable name semirubra referred here by Arnol'di, 1977b: 1638. Senior synonym of meridionalis: Atanassov & Dlussky, 1992: 121; of caucasicola: Arakelian, 1994: 34. Current subspecies: nominal plus glabrescens. See also: Agosti & Collingwood, 1987a: 54; Baroni Urbani, Aktaç & Camiltepe, 1989: 291.

Myrmicinae Pheidolini Messor chamberlini TRUE TRUE FALSE

chamberlini. Messor chamberlini Wheeler, W.M. 1915b: 410 (w.) U.S.A. Wheeler, G.C. & Wheeler, J. 1972b: 240 (l.). Combination in Veromessor: Wheeler, W.M. & Creighton, 1934: 366; in Messor: Bolton, 1982: 341. Subspecies of andrei: Enzmann, J. 1947b: 152. Revived status as species: Creighton, 1950a: 159.

Myrmicinae Pheidolini Messor chicoensis TRUE TRUE FALSE

chicoensis. Veromessor stoddardi subsp. chicoensis Smith, M.R. 1956a: 37 (w.) U.S.A. Combination in Messor: Bolton, 1995b: 253. Raised to species: Johnson, R.A. 2000: 96 (in list); Ward, 2005: 66 (in list).

Myrmicinae Pheidolini Messor fraternus TRUE TRUE FALSE

fraternus. Messor lobuliferus var. fraternus Ruzsky, 1905b: 740 (w.) RUSSIA. Raised to species: Ruzsky, 1915a: 444.

Myrmicinae Pheidolini Messor lariversi TRUE TRUE FALSE

lariversi. Veromessor lariversi Smith, M.R. 1951c: 94 (w.) U.S.A. Cole, 1955c: 52 (q.); Cole, 1963: 680 (m.). Combination in Messor: Bolton, 1982: 341.

Myrmicinae Pheidolini Messor lobognathus TRUE TRUE FALSE

lobognathus. Messor lobognathus Andrews, 1916: 82, fig. 1 (w.) U.S.A. Wheeler, G.C. & Wheeler, J. 1957c: 142 (l.); Cole, 1963: 680 (q.m.). Combination in Veromessor: Wheeler, W.M. & Creighton, 1934: 371; in Messor: Bolton, 1982: 341. See also: Gregg, 1955b: 45.

Myrmicinae Pheidolini Messor pergandei TRUE TRUE FALSE

pergandei. Aphaenogaster pergandei Mayr, 1886d: 448 (w.) U.S.A. Wheeler, W.M. & Creighton, 1934: 376 (s.q.m.); Wheeler, G.C. & Wheeler, J. 1972b: 240 (l.). Combination in Stenamma (Messor): Emery, 1895c: 307; in Novomessor: Emery, 1915d: 73; in N. (Veromessor): Forel, 1917: 234; in Veromessor: Wheeler, W.M. & Creighton, 1934: 374; in Messor: Bolton, 1982: 341. See also: Smith, D.R. 1979: 1365.

Myrmicinae Pheidolini Messor rufus TRUE TRUE FALSE

1 subspecies

rufus. Messor rufus Santschi, 1923f: 326 (w.) RUSSIA. [First available use of Messor barbarus subsp. meridionalis var. rufa Karavaiev, 1910a: 272; unavailable name.] Kuznetsov-Ugamsky, 1929d: 7 (q.m.). Subspecies of minor: Santschi, 1927c: 240; Santschi, 1927d: 247. Revived status as species: Kuznetsov-Ugamsky, 1929b: 45; Arnol'di, 1977b: 1642. Senior synonym of similis: Santschi, 1927d: 248; of karawajevi: Arnol'di, 1977b: 1642. Current subspecies: nominal plus striolatus.

Myrmicinae Pheidolini Messor smithi TRUE TRUE FALSE

smithi. Veromessor smithi Cole, 1963: 678, figs. 1-4 (w.q.m.) U.S.A. Wheeler, G.C. & Wheeler, J. 1972b: 240 (l.). Combination in Messor: Bolton, 1982: 341.

Myrmicinae Pheidolini Messor stoddardi TRUE TRUE FALSE

stoddardi. Stenamma (Messor) stoddardi Emery, 1895c: 307 (w.) U.S.A. Combination in Novomessor: Emery, 1915d: 73; in N. (Veromessor): Forel, 1917: 235; in Veromessor: Wheeler, W.M. & Creighton, 1934: 385; in Messor: Bolton, 1982: 341.

Myrmicinae Pheidolini Messor subgracilinodis TRUE TRUE FALSE

subgracilinodis. Messor subgracilinoclis Arnol'di, 1970a: 73, fig. 1 (w.q.) RUSSIA. Spelling emended to subgracilinodis: Arnol'di, 1970a: 87 (caption of fig. 1); Arnol'di, 1977b: 1642.

Myrmicinae Pheidolini Messor julianus manni TRUE TRUE FALSE

manni. Veromessor julianus subsp. manni Wheeler, W.M. & Creighton, 1934: 371 (w.) U.S.A. Combination in Messor: Bolton, 1995b: 255.

Aneuretinae Aneuretini Mianeuretus eocenicus TRUE TRUE TRUE

†eocenicus. †Mianeuretus eocenicus Dlussky & Rasnitsyn, 2002: 414, figs. 1, 2 (q.) U.S.A. (Eocene).

Aneuretinae Aneuretini Mianeuretus mirabilis TRUE TRUE TRUE

†mirabilis. †Mianeuretus mirabilis Carpenter, 1930: 38, pl. 3, fig. 5; pl. 10, fig. 1 (q.) U.S.A. (Oligocene).

Dolichoderinae Miomyrmecini Miomyrmex impactus TRUE TRUE TRUE

†impactus. †Formica impacta Cockerell, 1927: 165 (q.) U.S.A. (Oligocene). Combination in †Miomyrmex: Carpenter, 1930: 52.

Dolichoderinae Miomyrmecini Miomyrmex striatus TRUE TRUE TRUE

†striatus. †Miomyrmex striatus Carpenter, 1930: 54, pl. 8, fig. 6; pl. 10, fig. 9 (w.m.) U.S.A. (Oligocene).

Myrmicinae Solenopsidini Monomorium emarginatum TRUE TRUE FALSE

emarginatum. Monomorium emarginatum DuBois, 1986: 101, figs. 111-118 (w.q.) U.S.A.

Myrmicinae Solenopsidini Monomorium ergatogyna TRUE TRUE FALSE

ergatogyna. Monomorium minutum subsp. ergatogyna Wheeler, W.M. 1904d: 269 (w.q.) U.S.A. DuBois, 1986: 88 (m.). Subspecies of minimum: Wheeler, W.M. 1914b: 42. Raised to species: Essig, 1926: 857. Junior synonym of minimum: Creighton, 1950a: 219. Revived from synonymy, status as species: DuBois, 1986: 86. Senior synonym of wheelerorum: Ward, 2005: 14.

Myrmicinae Solenopsidini Monomorium minimum TRUE TRUE FALSE

minimum. Myrmica (Monomorium) minima Buckley, 1867: 338 (w.q.) U.S.A. Emery, 1895c: 274 (m.); Wheeler, G.C. & Wheeler, J. 1955c: 122 (l.); Crozier, 1970: 116 (k.). Combination in Monomorium: Emery, 1895c: 274. Subspecies of minutum: Emery, 1895c: 274; Wheeler, W.M. 1904e: 301; Wheeler, W.M. 1906b: 2. Revived status as species: Wheeler, W.M. 1910g: 562; Wheeler, W.M. 1913c: 115; Wheeler, W.M. 1914b: 42; Dennis, 1938: 279. Senior synonym of atra: Emery, 1895c: 274; of metoecus: Brown, in Ettershank, 1966: 90. See also: Wheeler, G.C. & Wheeler, J. 1973a: 30; DuBois, 1986: 76.

Myrmicinae Solenopsidini Monomorium pergandei TRUE TRUE FALSE

pergandei. Epoecus pergandei Emery, 1893a: cclxxvi (q.m.) U.S.A. [Also described as new by Emery, 1895c: 273.] Combination in Monomorium: Ettershank, 1966: 91. See also: DuBois, 1986: 113; Bolton, 1987: 299.

Myrmicinae Solenopsidini Monomorium talbotae TRUE TRUE FALSE

talbotae. Monomorium talbotae DuBois, 1981: 31, fig. 1 (q.m.) U.S.A. See also: DuBois, 1986: 112.

Myrmicinae Solenopsidini Monomorium trageri TRUE TRUE FALSE

trageri. Monomorium trageri DuBois, 1986: 109, figs. 7, 20, 133-148 (w.q.m.) U.S.A.

Myrmicinae Solenopsidini Monomorium viridis TRUE TRUE FALSE

viridis. Monomorium viridis Brown, 1943: 243 (w.q.) U.S.A. Crozier, 1970: 116 (k.); DuBois, 1986: 98 (m.). Senior synonym of peninsulatum: DuBois, 1986: 96.

Myrmicinae Attini Mycetosoritis hartmanni TRUE TRUE FALSE

hartmanni. Atta (Mycetosoritis) hartmanni Wheeler, W.M. 1907c: 714, pl. 49, figs. 6, 7 (w.q.m.) U.S.A. Wheeler, G.C. & Wheeler, J. 1974d: 77 (l.). Combination in Cyphomyrmex (Mycetosoritis): Emery, 1913b: 251; in Mycetosoritis: Creighton, 1950a: 318.

Myrmicinae Myrmecinini Myrmecina americana TRUE TRUE FALSE

americana. Myrmecina latreillei subsp. americana Emery, 1895c: 271 (w.) U.S.A. Wheeler, G.C. & Wheeler, J. 1954c: 130 (l.). Subspecies of graminicola: Wheeler, W.M. 1904e: 300; Wheeler, W.M. 1906b: 1. Raised to species: Brown, 1949a: 44; Creighton, 1950a: 248. Senior synonym of brevispinosa: Smith, D.R. 1979: 1399; of quadrispina: Brown, 1949a: 44; of texana: Brown, 1951: 103; Brown, 1967c: 236; of californica: Brown, 1967c: 236. See also: Snelling, R.R. 1965b: 101.

Formicinae Lasiini Myrmecocystus christineae TRUE TRUE FALSE

christineae. Myrmecocystus (Myrmecocystus) christineae Snelling, R.R. 1982b: 80, figs. 7-12, 14, 16, 20, 22, 24 (w.q.m.) U.S.A.

Formicinae Lasiini Myrmecocystus colei TRUE TRUE FALSE

colei. Myrmecocystus (Eremnocystus) colei Snelling, R.R. 1976: 94, figs. 203-211, 255-258 (w.q.m.) U.S.A.

Formicinae Lasiini Myrmecocystus creightoni TRUE TRUE FALSE

creightoni. Myrmecocystus creightoni Snelling, R.R. 1971a: 6, fig. 2 (w.q.m.) U.S.A. See also: Snelling, R.R. 1976: 98.

Formicinae Lasiini Myrmecocystus ewarti TRUE TRUE FALSE

ewarti. Myrmecocystus ewarti Snelling, R.R. 1971a: 2, fig. 1 (w.q.m.) U.S.A. See also: Snelling, R.R. 1976: 129.

Formicinae Lasiini Myrmecocystus flaviceps TRUE TRUE FALSE

flaviceps. Myrmecocystus yuma var. flaviceps Wheeler, W.M. 1912d: 177 (w.) U.S.A. Snelling, R.R. 1976: 84 (q.m.). Raised to species: Creighton, 1950a: 443.

Formicinae Lasiini Myrmecocystus hammettensis TRUE TRUE FALSE

hammettensis. Myrmecocystus hammettensis Cole, 1938b: 678 (w.q.m.) U.S.A. See also: Snelling, R.R. 1976: 101.

Formicinae Lasiini Myrmecocystus kathjuli TRUE TRUE FALSE

kathjuli. Myrmecocystus (Endiodioctes) kathjuli Snelling, R.R. 1976: 59, figs. 92-99, 160, 172, 185, 193 (w.q.m.) U.S.A.

Formicinae Lasiini Myrmecocystus kennedyi TRUE TRUE FALSE

kennedyi. Myrmecocystus kennedyi Snelling, R.R. 1969a: 6 (w.q.m.) U.S.A. [First available use of Myrmecocystus melliger subsp. semirufus var. kennedyi Cole, 1936b: 119; unavailable name.] See also: Snelling, R.R. 1976: 62.

Formicinae Lasiini Myrmecocystus koso TRUE TRUE FALSE

koso. Myrmecocystus (Endiodioctes) koso Snelling, R.R. 1976: 74, figs. 127-135, 164, 176, 199, 200 (w.q.m.) U.S.A.

Formicinae Lasiini Myrmecocystus lugubris TRUE TRUE FALSE

lugubris. Myrmecocystus lugubris Wheeler, W.M. 1909f: 98 (w.) U.S.A. Wheeler, G.C. & Wheeler, J. 1968: 213 (l.). See also: Creighton, 1956: 1; Snelling, R.R. 1976: 104.

Formicinae Lasiini Myrmecocystus mendax TRUE TRUE FALSE

mendax. Myrmecocystus melliger subsp. mendax Wheeler, W.M. 1908d: 351, fig. 4 (w.q.m.) U.S.A. Raised to species: Creighton, 1950a: 445. Senior synonym of orbiceps: Snelling, R.R. 1976: 35.

Formicinae Lasiini Myrmecocystus mimicus TRUE TRUE FALSE

mimicus. Myrmecocystus melliger subsp. mimicus Wheeler, W.M. 1908d: 353 (w.q.m.) U.S.A. Wheeler, G.C. & Wheeler, J. 1968: 213 (l.). Raised to species: Creighton, 1950a: 446. Senior synonym of lomaensis and material of the unavailable name jesuita referred here: Creighton, 1950a: 447; Snelling, R.R. 1976: 54.

Formicinae Lasiini Myrmecocystus navajo TRUE TRUE FALSE

navajo. Myrmecocystus mexicanus subsp. navajo Wheeler, W.M. 1908d: 360, fig. 11 (w.q.) U.S.A. Snelling, R.R. 1976: 126 (m.). Raised to species: Creighton, 1950a: 449.

Formicinae Lasiini Myrmecocystus pyramicus TRUE TRUE FALSE

pyramicus. Myrmecocystus pyramicus Smith, M.R. 1951c: 91, figs. 1-3 (w.) U.S.A. Cole, 1958b: 129 (q.m.). See also: Snelling, R.R. 1976: 132.

Formicinae Lasiini Myrmecocystus romainei TRUE TRUE FALSE

romainei. Myrmecocystus romainei Snelling, in Hunt & Snelling, 1975: 22 (w.) U.S.A. [First available use of Myrmecocystus melliger subsp. semirufus var. romainei Cole, 1936b: 120; unavailable name.] Snelling, R.R. 1976: 78 (q.m.).

Formicinae Lasiini Myrmecocystus semirufus TRUE TRUE FALSE

semirufus. Myrmecocystus melliger var. semirufus Emery, 1893i: 667 (w.m.) U.S.A. Wheeler, G.C. & Wheeler, J. 1968: 213 (l.); Snelling, R.R. 1976: 46 (q.). Subspecies of melliger: Wheeler, W.M. 1908d: 355. Raised to species: Creighton, 1950a: 449.

Formicinae Lasiini Myrmecocystus tenuinodis TRUE TRUE FALSE

tenuinodis. Myrmecocystus (Eremnocystus) tenuinodis Snelling, R.R. 1976: 107, figs. 241-247 (w.q.) U.S.A.

Formicinae Lasiini Myrmecocystus testaceus TRUE TRUE FALSE

testaceus. Myrmecocystus melliger var. testaceus Emery, 1893i: 667 (w.) U.S.A. Snelling, R.R. 1976: 136 (q.m.). Junior synonym of semirufus: Creighton, 1950a: 449. Revived from synonymy, raised to species and senior synonym of mojave: Snelling, R.R. 1969a: 6. Senior synonym of idahoensis: Snelling, R.R. 1976: 135.

Formicinae Lasiini Myrmecocystus wheeleri TRUE TRUE FALSE

wheeleri. Myrmecocystus wheeleri Snelling, R.R. 1971a: 11, fig. 3 (w.q.m.) U.S.A. See also: Snelling, R.R. 1976: 70.

Formicinae Lasiini Myrmecocystus yuma TRUE TRUE FALSE

yuma. Myrmecocystus yuma Wheeler, W.M. 1912d: 176 (w.) U.S.A. Snelling, R.R. 1976: 110 (q.). Junior synonym of lugubris: Creighton, 1956: 1. Revived from synonymy: Snelling, in Hunt & Snelling, 1975: 23.

Myrmicinae Myrmicini Myrmica alaskensis TRUE TRUE FALSE

alaskensis. Myrmica brevinodis var. alaskensis Wheeler, W.M. 1917a: 503 (w.) U.S.A. Junior synonym of kuschei: Creighton, 1950a: 97; of brevinodis: Weber, 1950b: 189; Smith, M.R. 1951a: 789. Revived from synonymy, raised to species and senior synonym of kuschei: Francoeur, 1986b: 57.

Myrmicinae Myrmicini Myrmica americana TRUE TRUE FALSE

americana. Myrmica sabuleti subsp. americana Weber, 1939b: 144 (w.q.m.) U.S.A. Wheeler, G.C. & Wheeler, J. 1953a: 116 (l.). Raised to species: Creighton, 1950a: 94.

Myrmicinae Myrmicini Myrmica angulinodis TRUE TRUE FALSE

angulinodis. Myrmica scabrinodis subsp. angulinodis Ruzsky, 1905b: 689 (w.q.) RUSSIA. Radchenko, 1994g: 84 (m.). Raised to species: Collingwood, 1962: 217; Collingwood, 1976: 302. Senior synonym of baikalensis: Radchenko, 1994g: 84; of incurvata: Radchenko, 2005b: 138. See also: Radchenko & Elmes, 2010: 89.

Myrmicinae Myrmicini Myrmica arnoldii TRUE TRUE FALSE

arnoldii. Myrmica arnoldii Dlussky, 1963: 191, figs. 1, 4, 5 (w.) RUSSIA. Arnol'di, 1968: 1803 (q.m.). Combination in M. (Dodecamyrmica): Arnol'di, 1968: 1803. See also: Francoeur, 1981: 759; Radchenko & Elmes, 2003a: 222; Radchenko & Elmes, 2010: 94.

Myrmicinae Myrmicini Myrmica brevispinosa TRUE TRUE FALSE

brevispinosa. Myrmica brevinodis var. brevispinosa Wheeler, W.M. 1917a: 502 (w.q.m.) U.S.A. [First available use of Myrmica rubra subsp. brevinodis var. brevispinosa Wheeler, W.M. 1907b: 74; unavailable name.] Wheeler, G.C. & Wheeler, J. 1953a: 117 (l.). Subspecies of brevinodis: Wheeler, G.C. & Wheeler, E.W. 1944: 241. Raised to species: Creighton, 1950a: 97. Senior synonym of decedens: Weber, 1950b: 198.

Myrmicinae Myrmicini Myrmica colax TRUE TRUE FALSE

colax. Paramyrmica colax Cole, 1957a: 37, fig. 1 (w.q.) U.S.A. Wheeler, G.C. & Wheeler, J. 1959: 219 (l.). Combination in Myrmica: Bolton, 1988a: 4.

Myrmicinae Myrmicini Myrmica commarginata TRUE TRUE FALSE

commarginata. Myrmica commarginata Ruzsky, 1905b: 708 (w.) RUSSIA. See also: Radchenko, 1994c: 138; Radchenko & Elmes, 2010: 112.

Myrmicinae Myrmicini Myrmica corrugata FALSE FALSE FALSE

corrugata. Myrmica corrugata Say, 1836: 291 (q.m.) U.S.A. Unidentifiable to genus; incertae sedis in Myrmica: Bolton, 1995b: 278.

Myrmicinae Myrmicini Myrmica crassirugis TRUE TRUE FALSE

crassirugis. Myrmica crassirugis Francoeur, 2007: 164, figs. 28 – 31 (w.q.m.) U.S.A.

Myrmicinae Myrmicini Myrmica deplanata TRUE TRUE FALSE

deplanata. Myrmica lobicornis var. deplanata Emery, 1921f: 38 (w.) UKRAINE, RUSSIA, GEORGIA. [First available use of Myrmica scabrinodis subsp. lobicornis var. deplanata Ruzsky, 1905b: 700; unavailable name.] Subspecies of lobicornis: Karavaiev, 1927a: 283; of schencki: Finzi, 1926: 111. Raised to species: Arnol'di, 1934: 169; Karavaiev, 1934: 83; Stitz, 1939: 105; Tarbinsky, 1976: 31. Senior synonym of moravica, plana: Karavaiev, 1934: 83; Arnol'di, 1970b: 1843; Seifert, 1988b: 36; Radchenko & Elmes, 2009c: 511; Radchenko & Elmes, 2010: 116; of dagmarae: Radchenko & Elmes, 2010: 116. Junior synonym of lacustris: Radchenko, 1994f: 77; Seifert, 2003b: 150; Radchenko, Elmes & Alicata, 2006: 516. Revived from synonymy: Radchenko & Elmes, 2009c: 511. See also: Radchenko & Elmes, 2010: 116.

Myrmicinae Myrmicini Myrmica discontinua TRUE TRUE FALSE

discontinua. Myrmica brevinodis subsp. discontinua Weber, 1939b: 150 (w.) U.S.A. Subspecies of brevispinosa: Creighton, 1950a: 98. Raised to species: Talbot, 1976: 245; Francoeur, 1977b: 206.

Myrmicinae Myrmicini Myrmica divergens TRUE TRUE FALSE

divergens. Myrmica bergi var. divergens Karavaiev, 1931c: 105, fig. 1 (w.) RUSSIA. Radchenko, Elmes & Woyciechowski, 2002: 411 (q.m.). Subspecies of bergi: Pisarski, 1969a: 227; Dlussky & Pisarski, 1970: 85. Junior synonym of bergi: Radchenko, 1994e: 76. Revived from synonymy and raised to species: Radchenko, Elmes & Woyciechowski, 2002: 411. See also: Radchenko & Elmes, 2010: 121.

Myrmicinae Myrmicini Myrmica diversa FALSE FALSE FALSE

diversa. Myrmica (Monomorium) diversa Buckley, 1867: 337 (w.q.) U.S.A. Unidentifiable to genus; incertae sedis in Myrmica: Bolton, 1995b: 278. See also: Emery, 1895c: 338; Wheeler, W.M. 1902f: 26.

Myrmicinae Myrmicini Myrmica eidmanni TRUE TRUE FALSE

eidmanni. Myrmica eidmanni Menozzi, 1930d: 331, fig. 3 (w.) RUSSIA. Subspecies of scabrinodis: Weber, 1950b: 204. Revived status as species and senior synonym of littoralis: Radchenko, 2005b: 139. See also: Radchenko & Elmes, 2010: 127.

Myrmicinae Myrmicini Myrmica excelsa TRUE TRUE FALSE

excelsa. Myrmica excelsa Kupyanskaya, 1990: 115, figs. 16, 19, 20 (w.q.m.) RUSSIA. Senior synonym of cadusa: Radchenko, 2005b: 140; of sinica: Radchenko, Zhou, et al. 2008: 783. See also: Yamane, 2008: 29; Radchenko & Elmes, 2010: 131.

Myrmicinae Myrmicini Myrmica forcipata TRUE TRUE FALSE

forcipata. Myrmica (Myrmica) forcipata Karavaiev, 1931c: 105, fig. 2 (w.) RUSSIA. See also: Radchenko & Elmes, 2010: 134.

Myrmicinae Myrmicini Myrmica fracticornis TRUE TRUE FALSE

fracticornis. Myrmica lobicornis var. fracticornis Forel, 1901e: 371 (in text) (w.) U.S.A. [First available use of Myrmica rubra subsp. scabrinodis var. fracticornis Emery, 1895c: 313; unavailable name.] Weber, 1948a: 280 (q.m.); Wheeler, G.C. & Wheeler, J. 1953a: 117 (l.). Subspecies of scabrinodis: Wheeler, W.M. 1916m: 587; of lobicornis: Cole, 1942: 368; Wheeler, G.C. & Wheeler, E.W. 1944: 241; Creighton, 1950a: 100. Raised to species: Talbot, 1976: 245; Francoeur, 1977b: 206; Wheeler, G.C. & Wheeler, J. 1986g: 12. Senior synonym of detritinodis: Weber, 1948a: 276; Smith, M.R. 1951a: 790.

Myrmicinae Myrmicini Myrmica granulinodis FALSE FALSE FALSE

granulinodis. Myrmica granulinodis Nylander, 1846b: 1060 (q.m.) RUSSIA. Subspecies of scabrinodis: André, 1883a: 319; Weber, 1950b: 205. Junior synonym of sabuleti: Emery, 1908a: 177 (footnote); Sadil, 1952: 246. Unidentifiable taxon, incertae sedis in Myrmica: Radchenko, 1994g: 90. See also: Radchenko & Elmes, 2010: 659.

Myrmicinae Myrmicini Myrmica hamulata TRUE TRUE FALSE

hamulata. Myrmica sabuleti subsp. hamulata Weber, 1939b: 146 (w.q.m.) U.S.A. Raised to species: Creighton, 1950a: 99. Current subspecies: nominal plus trullicornis.

Myrmicinae Myrmicini Myrmica inflecta FALSE FALSE FALSE

inflecta. Myrmica inflecta Say, 1836: 292 (m.) U.S.A. Unidentifiable to genus; incertae sedis in Myrmica: Bolton, 1995b: 279.

Myrmicinae Myrmicini Myrmica kamtschatica TRUE TRUE FALSE

kamtschatica. Myrmica kamtschatica Kupyanskaya, 1986a: 88, figs. 3, 6, 9, 10, 14 (w.q.m.) RUSSIA. Senior synonym of aborigenica: Radchenko, 1994g: 86. See also: Kupyanskaya, 1990: 112; Radchenko & Elmes, 2010: 161.

Myrmicinae Myrmicini Myrmica kasczenkoi TRUE TRUE FALSE

kasczenkoi. Myrmica scabrinodis subsp. kasczenkoi Ruzsky, 1905b: 702 (w.m.) RUSSIA. Kupyanskaya, 1990: 113 (q.); Radchenko, 1994g: 82 (w.q.m.). Raised to species: Pisarski, 1969b: 296; Dlussky & Pisarski, 1970: 85; Collingwood, 1976: 302. Material of the unavailable name subaltaica referred here by Radchenko, 1994g: 82. See also: Radchenko & Elmes, 2010: 166.

Myrmicinae Myrmicini Myrmica latifrons TRUE TRUE FALSE

latifrons. Myrmica schencki subsp. latifrons Stärcke, 1927: 84 (w.) U.S.A. Raised to species: Bolton, 1995b: 280. Senior synonym of emeryana: Weber, 1948a: 298; Smith, M.R. 1954d: 138; Wheeler, G.C. & Wheeler, J. 1986g: 12. [All give latifrons as the junior synonym, but it has priority over emeryana and is the first available name for this taxon: Bolton, 1995b: 280.]

Myrmicinae Myrmicini Myrmica lineolata FALSE FALSE FALSE

lineolata. Myrmica (Monomorium) lineolata Buckley, 1867: 340 (w.q.) U.S.A. [Unresolved junior primary homonym of lineolta Say, 1836: 290 (Bolton, 1995b: 280).] Unidentifiable to genus; incertae sedis in Myrmica: Bolton, 1995b: 280.

Myrmicinae Myrmicini Myrmica lobifrons TRUE TRUE FALSE

lobifrons. Myrmica sabuleti var. lobifrons Pergande, 1900: 521 (w.) U.S.A. Subspecies of scabrinodis: Emery, 1921f: 40; of lobicornis: Creighton, 1950a: 100. Junior synonym of fracticornis: Smith, M.R. 1951a: 790; Cole, 1953d: 242. Revived from synonymy as subspecies of lobicornis: Gregg, 1961: 212. Raised to species: Yensen, Clark & Francoeur, 1977: 182.

Myrmicinae Myrmicini Myrmica luteola TRUE TRUE FALSE

luteola. Myrmica luteola Kupyanskaya, 1990: 103, Figs. 16, 17(w.q.) RUSSIA. Radchenko, 1994f: 75 (m.). See also: Masuko & Terayama, 2002: 228; Radchenko & Elmes, 2003a: 239; Radchenko & Elmes, 2010: 197.

Myrmicinae Myrmicini Myrmica montana FALSE FALSE FALSE

montana. Myrmica (Monomorium) montana Buckley, 1867: 339 (w.) U.S.A. [Unresolved junior primary homonym of montana Labram & Imhoff, 1838, pl. 36, figs. 1-3, above (Bolton, 1995b: 281).] Unidentifiable to genus; incertae sedis in Myrmica: Bolton, 1995b: 281. See also: Emery, 1895c: 338; Wheeler, W.M. 1902f: 27.

Myrmicinae Myrmicini Myrmica monticola TRUE TRUE FALSE

monticola. Myrmica monticola Creighton, 1950a: 101 (w.m.) U.S.A. [First available use of Myrmica scabrinodis subsp. schencki var. monticola Wheeler, W.M. 1917a: 505; unavailable name.] Wheeler, G.C. & Wheeler, J. 1953a: 117 (l.).

Myrmicinae Myrmicini Myrmica nearctica TRUE TRUE FALSE

nearctica. Myrmica sabuleti subsp. nearctica Weber, 1939b: 148 (w.q.m.) U.S.A. Junior synonym of monticola: Creighton, 1950a: 101. Revived from synonymy and raised to species: Francoeur, 1977b: 206.

Myrmicinae Myrmicini Myrmica pinetorum TRUE TRUE FALSE

pinetorum. Myrmica punctiventris subsp. pinetorum Wheeler, W.M. 1905f: 384 (w.q.) U.S.A. Weber, 1950b: 217 (m.). Raised to species: Creighton, 1950a: 102. See also: Francoeur, 2007: 156.

Myrmicinae Myrmicini Myrmica pisarskii TRUE TRUE FALSE

pisarskii. Myrmica pisarskii Radchenko, 1994h: 208, figs. 1-12 (w.q.m.) RUSSIA, MONGOLIA. See also: Radchenko & Elmes, 2010: 216.

Myrmicinae Myrmicini Myrmica punctinops TRUE TRUE FALSE

punctinops. Myrmica punctinops Francoeur, 2007: 163, figs. 20 – 27 (w.) U.S.A.

Myrmicinae Myrmicini Myrmica punctiventris TRUE TRUE FALSE

punctiventris. Myrmica punctiventris Roger, 1863a: 190 (w.) U.S.A. Mayr, 1886d: 450 (q.); Emery, 1895c: 312 (m.). Senior synonym of isfahani: Weber, 1950b: 512. See also: Francoeur, 2007: 158.

Myrmicinae Myrmicini Myrmica rugiventris TRUE TRUE FALSE

rugiventris. Tetramorium rugiventris Smith, M.R. 1943b: 4 (w.) U.S.A. Gregg, 1961: 218 (q.m.). Combination in Myrmica: Brown, 1957d: 6; in Paramyrmica: Gregg, 1961: 215; in Myrmica: Bolton, 1988a: 6.

Myrmicinae Myrmicini Myrmica salina TRUE TRUE FALSE

salina. Myrmica scabrinodis var. salina Ruzsky, 1905b: 687 (w.q.m.) RUSSIA & KAZAKHSTAN. Subspecies of schencki: Weber, 1948a: 302. Junior synonym of lonae: Sadil, 1952: 249. Revived from synonymy and raised to species: Arnol'di, 1970b: 1842; Arnol'di & Dlussky, 1978: 534; Seifert, 1988: 25. Junior synonym of lacustris: Radchenko, 1994f: 77. Revived from synonymy: Seifert, 1994: 13; Seifert, 2002a: 96, 99. See also: Radchenko & Elmes, 2004: 228; Radchenko & Elmes, 2009c: 520; Radchenko & Elmes, 2010: 254.

Myrmicinae Myrmicini Myrmica scabrata FALSE FALSE FALSE

scabrata. Myrmica scabrata Buckley, 1867: 343 (w.) U.S.A. Unidentifiable to genus; incertae sedis in Myrmica: Bolton, 1995b: 283.

Myrmicinae Myrmicini Myrmica semiparasitica TRUE TRUE FALSE

semiparasitica. Myrmica semiparasitica Francoeur, 2007: 161, figs. 14 – 19 (w.q.m.) U.S.A.

Myrmicinae Myrmicini Myrmica spatulata TRUE TRUE FALSE

spatulata. Myrmica schencki var. spatulata Smith, M.R. 1930b: 566, figs. 1-4 (w.q.) U.S.A. Raised to species: Creighton, 1950a: 105.

Myrmicinae Myrmicini Myrmica stangeana TRUE TRUE FALSE

stangeana. Myrmica stangeana Ruzsky, 1902a: 234 (w.) RUSSIA, KAZAKHSTAN. Seifert, 1988b: 23 (q.m.). [Also described as new by Ruzsky, 1902b: 474.] Subspecies of bergi: Ruzsky, 1903b: 314; Ruzsky, 1905b: 678; of scabrinodis: Emery, 1908a: 178. Revived status as species: Arnol'di, 1970b: 1841; Arnol'di & Dlussky, 1978: 533. Senior synonym of medvedevi: Seifert, 1988b: 23; Radchenko, 1994e: 78; of lacustris: Radchenko & Elmes, 2009c: 509. See also: Radchenko & Elmes, 2010: 291.

Myrmicinae Myrmicini Myrmica striolagaster TRUE TRUE FALSE

striolagaster. Myrmica striolagaster Cole, 1953a: 81 (w.) U.S.A. Cole, 1957c: 208 (q.m.); Wheeler, G.C. & Wheeler, J. 1972b: 234 (l.).

Myrmicinae Myrmicini Myrmica sublanuginosa FALSE FALSE FALSE

sublanuginosa. Myrmica (Atta) sublanuginosa Buckley, 1867: 343 (w.) U.S.A. Unidentifiable to genus; incertae sedis in Myrmica: Bolton, 1995b: 284.

Myrmicinae Myrmicini Myrmica tahoensis TRUE TRUE FALSE

tahoensis. Myrmica schencki var. tahoensis Weber, 1948a: 304 (w.q.m.) U.S.A. [First available use of Myrmica scabrinodis subsp. schencki var. tahoensis Wheeler, W.M. 1917a: 504; unavailable name.] Subspecies of emeryana: Creighton, 1950a: 99. Raised to species: Yensen, Clark & Francoeur, 1977: 182.

Myrmicinae Myrmicini Myrmica transsibirica TRUE TRUE FALSE

transsibirica. Myrmica transsibirica Radchenko, 1994h: 212, figs. 23-32 (w.q.m.) RUSSIA. Senior synonym of taediosa: Radchenko & Elmes, 2010: 308.

Myrmicinae Myrmicini Myrmica tschekanovskii TRUE TRUE FALSE

tschekanovskii. Myrmica tschekanovskii Radchenko, 1994h: 215, figs. 33-39 (w.m.) RUSSIA. See also: Radchenko & Elmes, 2010: 310.

Myrmicinae Myrmicini Myrmica ussuriensis FALSE FALSE FALSE

ussuriensis. Myrmica scabrinodis subsp. ussuriensis Kuznetsov-Ugamsky, 1928b: 36 (w.) RUSSIA. Unidentifiable taxon, incertae sedis in Myrmica: Radchenko & Elmes, 2010: 662.

Myrmicinae Myrmicini Myrmica wheeleri TRUE TRUE FALSE

wheeleri. Myrmica wheeleri Weber, 1939b: 150 (w.q.m.) U.S.A.

Myrmicinae Myrmicini Myrmica wheelerorum TRUE TRUE FALSE

wheelerorum. Myrmica wheelerorum Francoeur, 2007: 166, figs. 32 – 35 (w.q.) U.S.A.

Myrmicinae Myrmicini Myrmica whymperi TRUE TRUE FALSE

whymperi. Myrmica brevinodis var. whymperi Forel, 1913l: 215 (w.) U.S.A. [First available use of Myrmica rubra r. brevinodis var. whymperi Forel, 1904a: 154; unavailable name.] Junior synonym of sulcinodoides: Wheeler, W.M. 1917a: 502. [Wheeler gives sulcinodoides as the senior synonym, but whymperi has priority and hence is senior synonym of sulcinodoides and its junior synonym frigida: Bolton, 1995b: 284.]

Myrmicinae Myrmicini Myrmica zojae TRUE TRUE FALSE

zojae. Myrmica zojae Radchenko, 1994h: 210, figs. 13-22 (w.q.m.) RUSSIA. See also: Radchenko & Elmes, 2010: 330.

Myrmicinae Myrmicini Myrmica sabuleti trullicornis TRUE TRUE FALSE

trullicornis. Myrmica sabuleti subsp. trullicornis Buren, 1944a: 281 (w.q.) U.S.A. Junior synonym of americana: Weber, 1948a: 288. Revived from synonymy as subspecies of hamulata: Creighton, 1950a: 99.

Myrmicinae Myrmicini Myrmica rubra neolaevinodis TRUE TRUE FALSE

neolaevinodis. Myrmica rubra r. neolaevinodis Forel, 1901h: 80 (w.) U.S.A. Subspecies of laevinodis: Weber, 1947: 455. Currently subspecies of rubra: Smith, D.R. 1979: 1351.

Myrmicinae Formicoxenini Myrmoxenus gordiagini TRUE TRUE FALSE

gordiagini. Myrmoxenus gordiagini Ruzsky, 1902b: 475, figs. A-H (1-8) (w.q.m.) RUSSIA. Combination in Epimyrma: Bolton, 1995b: 188; in Myrmoxenus: Schulz & Sanetra, 2002: 159. Senior synonym of menozzii and provisional senior synonym of microcellatus: Buschinger, Winter & Faber, 1984: 336. See also: Kupyanskaya, 1990: 145.

Ecitoninae Ecitonini Neivamyrmex baylori TRUE TRUE FALSE

baylori. Neivamyrmex baylori Watkins, 1973: 430, figs. 1-8 (m.) U.S.A.

Ecitoninae Ecitonini Neivamyrmex californicus TRUE TRUE FALSE

californicus. Eciton californicum Mayr, 1870b: 969 (w.) U.S.A. Watkins, 1972: 364 (q.). Combination in E. (Acamatus): Emery, 1894c: 182 (in text); in E. (Neivamyrmex): Smith, M.R. 1942c: 560; in Neivamyrmex: Borgmeier, 1953: 8. Senior synonym of obscura: Borgmeier, 1955: 517. See also: Ward, 1999a: 76; Snelling, G.C. & Snelling, 2007: 467.

Ecitoninae Ecitonini Neivamyrmex carolinensis TRUE TRUE FALSE

carolinensis. Eciton (Acamatus) carolinense Emery, 1894c: 184 (diagnosis in key) (w.) U.S.A. Wheeler, W.M. 1921d: 314 (q.m.). Combination in E. (Neivamyrmex): Smith, M.R. 1942c: 564; in Neivamyrmex: Borgmeier, 1953: 6. See also: Borgmeier, 1955: 507; Smith, D.R. 1979: 1330; Snelling, G.C. & Snelling, 2007: 468.

Ecitoninae Ecitonini Neivamyrmex coeca FALSE FALSE FALSE

coeca. Myrmica (Monomorium) coeca Buckley, 1867: 339 (w.) U.S.A. [Unresolved junior primary homonym of Myrmica caeca Jerdon, 1851: 116 (Bolton, 1995b: 288).] Combination in Eciton: Emery, 1895c: 260; possibly synonymous with opacithorax or schmitti: Wheeler, W.M. 1902f: 27; unplaced taxon of Ecitoninae: Smith, D.R. 1979: 1332. Unidentifiable taxon; incertae sedis in Neivamyrmex: Bolton, 1995b: 288.

Ecitoninae Ecitonini Neivamyrmex fuscipennis TRUE TRUE FALSE

fuscipennis. Eciton (Neivamyrmex) fuscipennis Smith, M.R. 1942c: 578, pl. 4, fig. 15 (m.) U.S.A. [Acamatus fuscipennis Wheeler, W.M. 1908e: 417. Nomen nudum, attributed to Cresson.] Combination in Neivamyrmex: Borgmeier, 1953: 18. Senior synonym of macropterus: Snelling, G.C. & Snelling, 2007: 469. See also: Borgmeier, 1955: 642; Watkins, 1975d: 85; Smith, D.R. 1979: 1330.

Ecitoninae Ecitonini Neivamyrmex harrisii TRUE TRUE FALSE

harrisii. Labidus harrissii Haldeman, 1852: 367, pl. 9, figs. 4-6 (m.) U.S.A. Wheeler, G.C. & Wheeler, J. 1984: 273 (l.). Combination in Eciton (Labidus): Mayr, 1886d: 441; in E. (Acamatus): Emery, 1900a: 187; in E. (Neivamyrmex): Smith, M.R. 1942c: 572; in Neivamyrmex: Borgmeier, 1955: 522. Senior synonym of wheeleri (and its junior synonym dubia): Watkins, 1968: 275. See also: Forbes & Do-Van-Quy, 1965: 95.

Ecitoninae Ecitonini Neivamyrmex kiowapache TRUE TRUE FALSE

kiowapache. Neivamyrmex kiowapache Snelling, G.C. & Snelling, 2007: 474, figs. 7, 21, 37, 52, 67, 91, 93, 97-99, 126, 138 (w.q.m.) U.S.A.

Ecitoninae Ecitonini Neivamyrmex leonardi TRUE TRUE FALSE

leonardi. Eciton (Acamatus) leonardi Wheeler, W.M. 1915b: 392 (w.) U.S.A. Combination in E. (Neivamyrmex): Smith, M.R. 1942c: 570; in Neivamyrmex: Borgmeier, 1953: 10. Senior synonym of peninsulare: Borgmeier, 1955: 431. See also: Snelling, G.C. & Snelling, 2007: 477.

Ecitoninae Ecitonini Neivamyrmex mandibularis TRUE TRUE FALSE

mandibularis. Eciton (Neivamyrmex) pilosum subsp. mandibulare Smith, M.R. 1942c: 548, pl. 3, fig. 14 (m.) U.S.A. Combination in Neivamyrmex: Borgmeier, 1955: 375. Raised to species: Snelling, G.C. & Snelling, 2007: 478.

Ecitoninae Ecitonini Neivamyrmex melshaemeri TRUE TRUE FALSE

melshaemeri. Labidus melshaemeri Haldeman, 1852: 368, pl. 9, figs. 7-9 (m.) U.S.A. Combination in Eciton (Labidus): Mayr, 1886d: 442; in E. (Acamatus): Emery, 1900a: 187; in E. (Neivamyrmex): Smith, M.R. 1942c: 576; in Neivamyrmex: Borgmeier, 1953: 8. See also: Borgmeier, 1955: 647; Snelling, G.C. & Snelling, 2007: 479.

Ecitoninae Ecitonini Neivamyrmex microps TRUE TRUE FALSE

microps. Neivamyrmex microps Borgmeier, 1955: 635, pl. 56, figs. 6, 10; pl. 80, fig. 4 (m.) U.S.A. Snelling, G.C. & Snelling, 2007: 480 (w.).

Ecitoninae Ecitonini Neivamyrmex minor TRUE TRUE FALSE

minor. Labidus minor Cresson, 1872: 195 (m.) U.S.A. Combination in Eciton (Labidus): Mayr, 1886d: 441; in E. (Acamatus): Emery, 1900a: 187; in E. (Neivamyrmex): Smith, M.R. 1942c: 574; in Neivamyrmex: Borgmeier, 1953: 8. [Misspelled as minus by Dalla Torre, 1893: 4.] See also: Borgmeier, 1955: 630; Snelling, G.C. & Snelling, 2007: 481.

Ecitoninae Ecitonini Neivamyrmex mojave TRUE TRUE FALSE

mojave. Eciton (Neivamyrmex) mojave Smith, M.R. 1943e: 196, fig. 1 (m.) U.S.A. See also: Borgmeier, 1955: 633; Snelling, G.C. & Snelling, 2007: 482.

Ecitoninae Ecitonini Neivamyrmex moseri TRUE TRUE FALSE

moseri. Neivamyrmex moseri Watkins, 1969: 529, figs. 1-5 (w.) U.S.A. Watkins, 1971: 96 (q.). Senior synonym of isodentatus: Snelling, G.C. & Snelling, 2007: 483.

Ecitoninae Ecitonini Neivamyrmex ndeh TRUE TRUE FALSE

ndeh. Neivamyrmex ndeh Snelling, G.C. & Snelling, 2007: 483, figs. 118, 132, 143 (m.) U.S.A. Senior synonym of goyahkla: Snelling, G.C. & Cover, 2009: 282.

Ecitoninae Ecitonini Neivamyrmex nigrescens TRUE TRUE FALSE

nigrescens. Labidus nigrescens Cresson, 1872: 194 (m.) U.S.A. Wheeler, W.M. 1900a: 563 (q.); Smith, M.R. 1942c: 551 (w.); Wheeler, G.C. & Wheeler, J. 1984: 273 (l.). Combination in Eciton: Dalla Torre, 1893: 5; in E. (Labidus): Emery, 1895c: 261; in E. (Acamatus): Emery, 1900a: 187; in E. (Neivamyrmex): Smith, M.R. 1942c: 550; Gregg, 1963: 286; in Neivamyrmex: Borgmeier, 1955: 494; Watkins, 1972: 349. Senior synonym of schmitti: Smith, M.R. 1938b: 160; Borgmeier, 1955: 494. See also: Watkins, 1972: 358; Smith, D.R. 1979: 1331; Petralia & Vinson, 1980: 378; Ward, 1999a: 84; Snelling, G.C. & Snelling, 2007: 484.

Ecitoninae Ecitonini Neivamyrmex nyensis TRUE TRUE FALSE

nyensis. Neivamyrmex nyensis Watkins, 1977b: 421, figs. 1-4 (w.) U.S.A. See also: Snelling, G.C. & Snelling, 2007: 486.

Ecitoninae Ecitonini Neivamyrmex opacithorax TRUE TRUE FALSE

opacithorax. Eciton (Acamatus) californicum subsp. opacithorax Emery, 1894c: 184 (diagnosis in key) (w.) U.S.A. Wheeler, W.M. & Long, 1901: 163, 173 (m.q., respectively); Wheeler, G.C. & Wheeler, J. 1984: 273 (l.). Combination in E. (Neivamyrmex): Smith, M.R. 1942c: 555; in Neivamyrmex: Borgmeier, 1953: 5. Raised to species: Emery, 1900a: 186. Senior synonym of castaneum: Borgmeier, 1955: 502. See also: Smith, D.R. 1979: 1332; Wheeler, G.C. & Wheeler, J. 1990a: 461; Snelling, G.C. & Snelling, 2007: 487.

Ecitoninae Ecitonini Neivamyrmex pauxillus TRUE TRUE FALSE

pauxillus. Eciton (Acamatus) pauxillum Wheeler, W.M. 1903b: 93, fig. 1 (w.) U.S.A. Watkins, 1971: 99 (q.). Combination in E. (Neivamyrmex): Smith, M.R. 1942c: 569; in Neivamyrmex: Borgmeier, 1953: 19. See also: Borgmeier, 1955: 570.

Ecitoninae Ecitonini Neivamyrmex texanus TRUE TRUE FALSE

texanus. Neivamyrmex texanus Watkins, 1972: 353, figs. 2-4, 11, 14, 17-19, 33, 34, 49, 61 (w.q.m.) U.S.A. Wheeler, G.C. & Wheeler, J. 1984: 273 (l.); Taber & Cokendolpher, 1988: 95 (k.). See also: Ward, 1999a: 88; Snelling, G.C. & Snelling, 2007: 491.

Ecitoninae Ecitonini Neivamyrmex wilsoni TRUE TRUE FALSE

wilsoni. Neivamyrmex wilsoni Snelling, G.C. & Snelling, 2007: 491, figs. 16, 26, 48, 61, 74, 87 (w.) U.S.A.

Myrmicinae Formicoxenini Nesomyrmex wilda TRUE TRUE FALSE

wilda. Leptothorax (Goniothorax) wilda Smith, M.R. 1943d: 155 (w.q.) U.S.A. Francoeur & Loiselle, 1988b: 44 (m.). Combination in L. (Nesomyrmex): Smith, M.R. 1951a: 817; in Nesomyrmex: Bolton, 2003: 272. See also: Kempf, 1959c: 421.

Ecitoninae Ecitonini Nomamyrmex esenbeckii wilsoni TRUE TRUE FALSE

wilsoni. Eciton (Labidus) esenbecki st. wilsoni Santschi, 1920d: 366 (m.) U.S.A. Combination in E. (Nomamyrmex): Smith, M.R. 1951a: 779; in Nomamyrmex: Borgmeier, 1953: 13. Currently subspecies of esenbeckii: Borgmeier, 1955: 139; Watkins, 1977a: 205.

Formicinae Plagiolepidini Nylanderia arenivaga TRUE TRUE FALSE

arenivaga. Prenolepis arenivaga Wheeler, W.M. 1905f: 391, fig. 3 (w.m.) U.S.A. Combination in Pr. (Nylanderia): Forel, 1922: 98; in Paratrechina (Nylanderia): Emery, 1925b: 221; in Nylanderia: LaPolla, Brady & Shattuck, 2010a: 127. Subspecies of melanderi: Creighton, 1950a: 408. Revived status as species: Trager, 1984b: 119.

Formicinae Plagiolepidini Nylanderia austroccidua TRUE TRUE FALSE

austroccidua. Paratrechina austroccidua Trager, 1984b: 113, fig. 17, 20 (w.q.m.) U.S.A. Combination in Nylanderia: LaPolla, Brady & Shattuck, 2010a: 127.

Formicinae Plagiolepidini Nylanderia bruesii TRUE TRUE FALSE

bruesii. Prenolepis bruesii Wheeler, W.M. 1903b: 106, fig. 9 (w.q.m.) U.S.A. Wheeler, G.C. & Wheeler, J. 1968: 211 (l.). Combination in Pr. (Nylanderia): Emery, 1914f: 422; in Paratrechina (Nylanderia): Emery, 1925b: 222; in Nylanderia: LaPolla, Brady & Shattuck, 2010a: 127. See also: Trager, 1984b: 129.

Formicinae Plagiolepidini Nylanderia concinna TRUE TRUE FALSE

concinna. Paratrechina concinna Trager, 1984b: 88, figs. 11, 14, 41 (w.q.m.) U.S.A. Combination in Nylanderia: LaPolla, Brady & Shattuck, 2010a: 127.

Formicinae Plagiolepidini Nylanderia faisonensis TRUE TRUE FALSE

faisonensis. Prenolepis (Nylanderia) arenivaga var. faisonensis Forel, 1922: 98 (w.) U.S.A. Combination in Paratrechina (Nylanderia): Emery, 1925b: 221; in Nylanderia: LaPolla, Brady & Shattuck, 2010a: 127. Junior synonym of arenivaga: Creighton, 1950a: 408. Revived from synonymy and raised to species: Trager, 1984b: 93.

Formicinae Plagiolepidini Nylanderia hystrix TRUE TRUE FALSE

hystrix. Paratrechina hystrix Trager, 1984b: 134, figs. 28, 30, 40 (w.q.m.) U.S.A. Combination in Nylanderia: LaPolla, Brady & Shattuck, 2010a: 127.

Formicinae Plagiolepidini Nylanderia parvula TRUE TRUE FALSE

parvula. Prenolepis parvula Mayr, 1870b: 948 (w.q.m.) U.S.A. Wheeler, G.C. & Wheeler, J. 1968: 211 (l.). Combination in Pr. (Nylanderia): Forel, 1922: 98; in Paratrechina (Nylanderia): Emery, 1925b: 222; in Nylanderia: LaPolla, Brady & Shattuck, 2010a: 127. Subspecies of vividula: Forel, 1885a: 348; Mayr, 1886d: 431. Revived status as species: Emery, 1893i: 636. See also: Trager, 1984b: 104.

Formicinae Plagiolepidini Nylanderia perminuta FALSE FALSE FALSE

perminuta. Formica perminuta Buckley, 1866: 162 (w.) U.S.A. Combination in Brachymyrmex: Emery, 1895c: 338; in Prenolepis: Mayr, 1876: 78; Wheeler, W.M. 1902f: 21. Unidentifiable taxon; incertae sedis in Paratrechina: Bolton, 1995b: 315. [Trager, 1984b: 119 suggests possible synonymy with arenivaga, a junior name, but the original description of perminuta does not contain sufficient data to justify this.]

Formicinae Plagiolepidini Nylanderia phantasma TRUE TRUE FALSE

phantasma. Paratrechina phantasma Trager, 1984b: 124, figs. 24, 26, 48 (w.q.m.) U.S.A. Combination in Nylanderia: LaPolla, Brady & Shattuck, 2010a: 127.

Formicinae Plagiolepidini Nylanderia terricola TRUE TRUE FALSE

terricola. Formica terricola Buckley, 1866: 168 (w.q.m.) U.S.A. Combination in Prenolepis: Mayr, 1876: 78; in Paratrechina: Emery, 1925b: 223; in Nylanderia: LaPolla, Brady & Shattuck, 2010a: 127. Junior synonym of vividula: Mayr, 1876: 78; Smith, M.R. 1951a: 849. Revived from synonymy and senior synonym of melanderi: Trager, 1984b: 81.

Formicinae Plagiolepidini Nylanderia wojciki TRUE TRUE FALSE

wojciki. Paratrechina wojciki Trager, 1984b: 108, figs. 18, 21 (w.q.m.) U.S.A. Wheeler, G.C. & Wheeler, J. 1986d: 337 (l.). Combination in Nylanderia: LaPolla, Brady & Shattuck, 2010a: 127.

Ponerinae Ponerini Odontomachus brunneus TRUE TRUE FALSE

brunneus. Atta brunnea Patton, 1894: 618 (w.) U.S.A. Deyrup, Trager & Carlin, 1985: 191 (m.). Combination in Odontomachus: Emery, 1895c: 268. Junior synonym of insularis: Emery, 1895c: 268. Revived from synonymy and senior synonym of fuscus Stitz, 1925: Brown, 1976a: 103. See also: Brown, 1976a: 139.

Ponerinae Ponerini Odontomachus clarus TRUE TRUE FALSE

clarus. Odontomachus clarus Roger, 1861a: 26 (w.) U.S.A. Patton, 1894: 618 (m.); Wheeler, W.M. 1908e: 407 (q.); Petralia & Vinson, 1980: 381 (l.). Subspecies of haematodus: Emery, 1911d: 115; Wheeler, W.M. 1914b: 40; Smith, M.R. 1939d: 129. Revived status as species: Taylor & Wilson, 1962: 142; Brown, 1976a: 136. Senior synonym of clarionensis: Taylor & Wilson, 1962: 142; of coninodis, desertorum: Brown, 1976a: 103; of texana: Wheeler, W.M. 1902f: 26. See also: Deyrup, Trager & Carlin, 1985: 192.

Ponerinae Ponerini Odontomachus relictus TRUE TRUE FALSE

relictus. Odontomachus relictus Deyrup & Cover, 2004b: 137, figs. 1, 2A (w.q.m.) U.S.A.

Ponerinae Ponerini Pachycondyla agilis TRUE TRUE FALSE

agilis. Neoponera (Eumecopone) agilis Forel, 1901e: 336 (w.) U.S.A. (locality in error; type-locality possibly Colombia: Mackay & Mackay, 2010: 199). Combination in Pachycondyla: Brown, in Bolton, 1995b: 302. Senior synonym of golbachi, goyana: Mackay & Mackay, 2010: 199.

Ponerinae Ponerini Pachycondyla gilva TRUE TRUE FALSE

gilva. Ponera gilva Roger, 1863a: 170 (w.) U.S.A. Creighton & Tulloch, 1930: 74 (q.m.); Wheeler, G.C. & Wheeler, J. 1952c: 625 (l.). Combination in Pachycondyla (Pseudoponera): Emery, 1901a: 46; in Euponera (Trachymesopus): Emery, 1911d: 86; in Trachymesopus: Kempf, 1960f: 424; in Cryptopone: Brown, 1963: 3; in Pachycondyla: Mackay & Mackay, 2010: 352. Senior synonym of harnedi: Creighton & Tulloch, 1930: 74; of obsoleta: Longino, 2006b: 135; of guatemalensis: Mackay & Mackay, 2010: 352.

Ponerinae Ponerini Pachycondyla labandeirai TRUE TRUE TRUE

†labandeirai. †Pachycondyla labandeirai Dlussky & Rasnitsyn, 2002: 424, fig. 25 (q.) U.S.A. (Eocene).

Myrmicinae Pheidolini Paraphaenogaster microphthalma TRUE TRUE TRUE

†microphthalma. †Paraphaenogaster microphthalmus Dlussky, 1981b: 69, fig. 51 (q.) RUSSIA (Miocene).

Myrmicinae Pheidolini Pheidole adrianoi TRUE TRUE FALSE

adrianoi. Pheidole adrianoi Naves, 1985: 56, figs. 25, 26, 33, 53, 54 (s.w.q.) U.S.A. See also: Wilson, 2003: 555.

Myrmicinae Pheidolini Pheidole aurea TRUE TRUE FALSE

aurea. Pheidole aurea Wilson, 2003: 557, figs. (s.w.) U.S.A.

Myrmicinae Pheidolini Pheidole bahai FALSE FALSE FALSE

bahai. Tetramorium (Cephalomorium) bahai Forel, 1922: 91 (w.) U.S.A. Combination in Pheidole (Hendecapheidole): Santschi, 1925d: 228. Unidentifiable taxon, incertae sedis in Pheidole: Wilson, 2003: 779.

Myrmicinae Pheidolini Pheidole barbata TRUE TRUE FALSE

barbata. Pheidole barbata Wheeler, W.M. 1908e: 448 (s.w.) U.S.A. See also: Wilson, 2003: 560.

Myrmicinae Pheidolini Pheidole bicarinata TRUE TRUE FALSE

bicarinata. Pheidole bicarinata Mayr, 1870b: 989 (s.) U.S.A. Senior synonym of hayesi: Creighton, 1950a: 171; of longula, vinelandica (and its junior synonyms buccalis, laeviuscula, nebrascensis): Wilson, 2003: 561.

Myrmicinae Pheidolini Pheidole bureni TRUE TRUE FALSE

bureni. Pheidole bureni Wilson, 2003: 150, figs. (s.w.) U.S.A.

Myrmicinae Pheidolini Pheidole californica TRUE TRUE FALSE

californica. Pheidole californica Mayr, 1870b: 987 (s.w.) U.S.A. Wheeler, W.M. 1915b: 406 (q.); Wheeler, G.C. & Wheeler, J. 1972b: 243 (l.). Senior synonym of incenata, satura: Creighton, 1950a: 173; of pyramidensis: Wheeler, G.C. & Wheeler, J. 1986g: 13; of oregonica (and its junior synonyms hagermani, shoshoni): Wilson, 2003: 564.

Myrmicinae Pheidolini Pheidole carrolli TRUE TRUE FALSE

carrolli. Pheidole carrolli Naves, 1985: 58, figs. 14, 43 (s.w.) U.S.A. See also: Wilson, 2003: 566.

Myrmicinae Pheidolini Pheidole casta TRUE TRUE FALSE

casta. Pheidole casta Wheeler, W.M. 1908e: 454, pl. 26, figs, 22, 23 (s.w.) U.S.A. See also: Wilson, 2003: 392.

Myrmicinae Pheidolini Pheidole cavigenis TRUE TRUE FALSE

cavigenis. Pheidole tepicana subsp. cavigenis Wheeler, W.M. 1915b: 403 (s.) U.S.A. Wilson, 2003: 567 (w.). Junior synonym of soritis: Gregg, 1959: 30. Revived from synonymy and raised to species: Wilson, 2003: 567.

Myrmicinae Pheidolini Pheidole cerebrosior TRUE TRUE FALSE

cerebrosior. Pheidole vinelandica subsp. cerebrosior Wheeler, W.M. 1915b: 405 (s.w.) U.S.A. Raised to species: Creighton, 1950a: 175. See also: Wilson, 2003: 569.

Myrmicinae Pheidolini Pheidole ceres TRUE TRUE FALSE

ceres. Pheidole ceres Wheeler, W.M. 1904a: 10 (s.w.q.m.) U.S.A. Senior synonym of tepaneca: Wilson, 2003: 570.

Myrmicinae Pheidolini Pheidole clementensis TRUE TRUE FALSE

clementensis. Pheidole clementensis Gregg, 1969: 93, fig. 1 (s.w.) U.S.A. See also: Wilson, 2003: 572.

Myrmicinae Pheidolini Pheidole clydei TRUE TRUE FALSE

clydei. Pheidole (Ceratopheidole) clydei Gregg, 1950: 89, fig. 1 (w.) U.S.A. Gregg, 1953b: 1 (s.). See also: Creighton, 1965a: 169; Wilson, 2003: 539.

Myrmicinae Pheidolini Pheidole cockerelli TRUE TRUE FALSE

cockerelli. Pheidole cockerelli Wheeler, W.M. 1908e: 464 (s.w.) U.S.A. See also: Wilson, 2003: 277.

Myrmicinae Pheidolini Pheidole constipata TRUE TRUE FALSE

constipata. Pheidole constipata Wheeler, W.M. 1908e: 468 (s.w.q.m.) U.S.A. Subspecies of floridana: Smith, D.R. 1979: 1370. Revived status as species: Wilson, 2003: 402.

Myrmicinae Pheidolini Pheidole crassicornis TRUE TRUE FALSE

crassicornis. Pheidole crassicornis Emery, 1895c: 296 (s.) U.S.A. Forel, 1901e: 350 (w.m.). See also: Wilson, 2003: 152.

Myrmicinae Pheidolini Pheidole creightoni TRUE TRUE FALSE

creightoni. Pheidole creightoni Gregg, 1955a: 19, pl. 2, figs. a-c; pl. 3, figs. a-c (s.w.q.m.) U.S.A. See also: Wilson, 2003: 573.

Myrmicinae Pheidolini Pheidole davisi TRUE TRUE FALSE

davisi. Pheidole davisi Wheeler, W.M. 1905f: 380 (s.w.) U.S.A. See also: Wilson, 2003: 574.

Myrmicinae Pheidolini Pheidole dentata TRUE TRUE FALSE

dentata. Pheidole morrisi var. dentata Mayr, 1886d: 457 (s.w.m.) U.S.A. Forel, 1901e: 351 (q.); Wheeler, G.C. & Wheeler, J. 1953b: 71 (l.); Crozier, 1970: 117 (k.). Raised to species: Forel, 1901e: 351. Senior synonym of faisonsica, commutata (and its junior synonym tennesseensis): Creighton, 1950a: 178. See also: Wilson, 1976b: 63; Wilson, 2003: 282.

Myrmicinae Pheidolini Pheidole dentigula TRUE TRUE FALSE

dentigula. Pheidole dentigula Smith, M.R. 1927: 310 (s.w.) U.S.A. Smith, M.R. 1928b: 245 (q.); Wheeler, G.C. & Wheeler, J. 1960b: 12 (l.); Crozier, 1970: 117 (k.). See also: Wilson, 2003: 407.

Myrmicinae Pheidolini Pheidole desertorum TRUE TRUE FALSE

desertorum. Pheidole desertorum Wheeler, W.M. 1906d: 337 (s.w.q.m.) U.S.A. Taber & Cokendolpher, 1988: 95 (k.). Senior synonym of comanche, maricopa: Creighton, 1950a: 178. See also: Wilson, 2003: 284.

Myrmicinae Pheidolini Pheidole diversipilosa TRUE TRUE FALSE

diversipilosa. Pheidole crassicornis var. diversipilosa Wheeler, W.M. 1908e: 467 (s.w.q.) U.S.A. Junior synonym of crassicornis: Creighton, 1950a: 175; Gregg, 1959: 20. Revived from synonymy and raised to species: Naves, 1985: 61. See also: Wilson, 2003: 153.

Myrmicinae Pheidolini Pheidole elecebra TRUE TRUE FALSE

elecebra. Sympheidole elecebra Wheeler, W.M. 1904a: 8, pl. 2, figs. 8-11 (q.m.) U.S.A. Combination in Pheidole: Smith, D.R. 1979: 1370. See also: Wilson, 1984: 328; Wilson, 2003: 576.

Myrmicinae Pheidolini Pheidole floridana TRUE TRUE FALSE

floridana. Pheidole flavens subsp. floridana Emery, 1895c: 293 (s.w.q.) U.S.A. Raised to species: Pergande, 1896: 883; Emery, 1896g: 77. Senior synonym of lauta: Gregg, 1959: 21. See also: Wilson, 2003: 424.

Myrmicinae Pheidolini Pheidole furtiva TRUE TRUE FALSE

furtiva. Pheidole furtiva Wilson, 2003: 425, figs. (s.w.) U.S.A.

Myrmicinae Pheidolini Pheidole gilvescens TRUE TRUE FALSE

gilvescens. Pheidole gilvescens Creighton & Gregg, 1955: 5 (s.w.) U.S.A. [First available use of Pheidole xerophila subsp. tucsonica var. gilvescens Wheeler, W.M. 1908e: 448; unavailable name.] See also: Wilson, 2003: 577.

Myrmicinae Pheidolini Pheidole grundmanni TRUE TRUE FALSE

grundmanni. Pheidole (Ceratopheidole) grundmanni Smith, M.R. 1953e: 144, fig. 1 (w.) U.S.A. See also: Wilson, 2003: 543.

Myrmicinae Pheidolini Pheidole hoplitica TRUE TRUE FALSE

hoplitica. Pheidole hoplitica Wilson, 2003: 579, figs. (s.w.) U.S.A.

Myrmicinae Pheidolini Pheidole humeralis TRUE TRUE FALSE

humeralis. Pheidole humeralis Wheeler, W.M. 1908e: 456, pl. 27, fig. 39 (s.w.) U.S.A. See also: Wilson, 2003: 438.

Myrmicinae Pheidolini Pheidole hyatti TRUE TRUE FALSE

hyatti. Pheidole hyatti Emery, 1895c: 295 (s.w.) U.S.A. Wheeler, W.M. 1908e: 463 (q.); Wheeler, G.C. & Wheeler, J. 1953b: 74 (l.); Wheeler, G.C. & Wheeler, J. 1972b: 244 (l.); Taber & Cokendolpher, 1988: 95 (k.). Senior synonym of ecitonodora: Creighton, 1950a: 180; of vaslitii, solitanea: Ward, 2000: 94. See also: Wilson, 2003: 302.

Myrmicinae Pheidolini Pheidole inquilina TRUE TRUE FALSE

inquilina. Epipheidole inquilina Wheeler, W.M. 1903f: 664, fig. 5 (gynandromorph) U.S.A. Wheeler, W.M. 1904a: 15 (q.m.); Smith, M.R. 1940a: 106 (w.); Cole, 1965: 174 (s.). [Also described as new by Wheeler, W.M. 1904a: 15.] Combination in Pheidole: Cole, 1965: 174. See also: Wilson, 2003: 580.

Myrmicinae Pheidolini Pheidole juniperae TRUE TRUE FALSE

juniperae. Pheidole juniperae Wilson, 2003: 440, figs. (s.w.) U.S.A.

Myrmicinae Pheidolini Pheidole lamia TRUE TRUE FALSE

lamia. Pheidole lamia Wheeler, W.M. 1901b: 534, fig. 11 (s.w.) U.S.A. See also: Feener, 1981: 269; Wilson, 2003: 546.

Myrmicinae Pheidolini Pheidole littoralis TRUE TRUE FALSE

littoralis. Pheidole sitarches subsp. littoralis Cole, 1952b: 443 (s.w.) U.S.A. Raised to species: Naves, 1985: 64. See also: Wilson, 2003: 581.

Myrmicinae Pheidolini Pheidole macclendoni TRUE TRUE FALSE

macclendoni. Pheidole macclendoni Wheeler, W.M. 1908e: 450, pl. 27, fig. 36 (s.w.) U.S.A. See also: Wilson, 2003: 582.

Myrmicinae Pheidolini Pheidole marcidula TRUE TRUE FALSE

marcidula. Pheidole marcidula Wheeler, W.M. 1908e: 457 (s.w.) U.S.A. See also: Wilson, 2003: 584.

Myrmicinae Pheidolini Pheidole meihuashanensis TRUE TRUE FALSE

meihuashanensis. Pheidole meihuashanensis Li & Chen, 1992: 57 (s.w.) CHINA. See also: Tang, Li, et al. 1995: 57.

Myrmicinae Pheidolini Pheidole mera TRUE TRUE FALSE

mera. Pheidole mera Wilson, 2003: 452, figs. (s.w.) U.S.A.

Myrmicinae Pheidolini Pheidole metallescens TRUE TRUE FALSE

metallescens. Pheidole metallescens Emery, 1895c: 294 (w.) U.S.A. Wheeler, W.M. 1908e: 476 (s.); Wheeler, G.C. & Wheeler, J. 1960b: 12 (l.). Senior synonym of splendidula: Wilson, 2003: 453.

Myrmicinae Pheidolini Pheidole micula TRUE TRUE FALSE

micula. Pheidole californica subsp. micula Wheeler, W.M. 1915b: 408 (s.w.) U.S.A. Wheeler, G.C. & Wheeler, J. 1972b: 244 (l.). Raised to species: Gregg, 1959: 24. See also: Wilson, 2003: 585.

Myrmicinae Pheidolini Pheidole militicida TRUE TRUE FALSE

militicida. Pheidole militicida Wheeler, W.M. 1915b: 398 (s.w.) U.S.A. Creighton & Gregg, 1955: 11 (q.m.). See also: Wilson, 2003: 586.

Myrmicinae Pheidolini Pheidole morrisii TRUE TRUE FALSE

morrisii. Pheidole morrisii Forel, 1886b: xlvi (s.w.) U.S.A. Forel, 1901e: 350 (q.m.); Wheeler, G.C. & Wheeler, J. 1960b: 12 (l.). Senior synonym of vanceae: Creighton, 1950a: 183; of impexa: Wilson, 2003: 325. See also: Feener, 1987: 569.

Myrmicinae Pheidolini Pheidole nuculiceps TRUE TRUE FALSE

nuculiceps. Pheidole nuculiceps Wheeler, W.M. 1908e: 473 (s.w.) U.S.A. See also: Wilson, 2003: 475.

Myrmicinae Pheidolini Pheidole paiute TRUE TRUE FALSE

paiute. Pheidole bicarinata subsp. paiute Gregg, 1959: 17 (s.w.) U.S.A. Subspecies of bicarinata: Smith, D.R. 1979: 1367. Raised to species: Wheeler, G.C. & Wheeler, J. 1973e: 75; Wilson, 2003: 588.

Myrmicinae Pheidolini Pheidole pelor TRUE TRUE FALSE

pelor. Pheidole pelor Wilson, 2003: 547, figs. (s.w.) U.S.A.

Myrmicinae Pheidolini Pheidole perpilosa TRUE TRUE FALSE

perpilosa. Pheidole perpilosa Wilson, 2003: 336, figs. (s.w.) U.S.A.

Myrmicinae Pheidolini Pheidole picea FALSE FALSE FALSE

picea. Atta picea Buckley, 1867: 344 (s.w.) U.S.A. Combination in Pheidole: Emery, 1895c: 338. Unidentifiable taxon; incertae sedis in Pheidole: Smith, M.R. 1951a: 806.

Myrmicinae Pheidolini Pheidole pilifera TRUE TRUE FALSE

pilifera. Leptothorax pilifer Roger, 1863a: 180 (w.) U.S.A. Emery, 1895c: 290 (s.); Wheeler, G.C. & Wheeler, J. 1953b: 79 (l.). Combination in Pheidole: Emery, 1895c: 290. Senior synonym of pennsylvanica: Emery, 1895c: 290; of septentrionalis and material of the unavailable name simulans referred here: Creighton, 1950a: 186; of artemisia, coloradensis, pacifica: Wilson, 2003: 589.

Myrmicinae Pheidolini Pheidole pilosior TRUE TRUE FALSE

pilosior. Pheidole pilosior Wilson, 2003: 157, figs. (s.w.) U.S.A.

Myrmicinae Pheidolini Pheidole pinealis TRUE TRUE FALSE

pinealis. Pheidole pinealis Wheeler, W.M. 1908e: 459, pl. 27, fig. 38 (s.w.) U.S.A. See also: Wilson, 2003: 591.

Myrmicinae Pheidolini Pheidole porcula TRUE TRUE FALSE

porcula. Pheidole crassicornis subsp. porcula Wheeler, W.M. 1908e: 466, pl. 27, fig. 35 (s.w.) U.S.A. Taber & Cokendolpher, 1988: 95 (k.). Raised to species: Creighton, 1950a: 187. See also: Wilson, 2003: 159.

Myrmicinae Pheidolini Pheidole portalensis TRUE TRUE FALSE

portalensis. Pheidole portalensis Wilson, 2003: 338, figs. (s.w.) U.S.A.

Myrmicinae Pheidolini Pheidole psammophila TRUE TRUE FALSE

psammophila. Pheidole psammophila Creighton & Gregg, 1955: 15, figs. 4, 5 (s.w.) U.S.A. See also: Wilson, 2003: 593.

Myrmicinae Pheidolini Pheidole rhea TRUE TRUE FALSE

rhea. Pheidole rhea Wheeler, W.M. 1908e: 452 (q.) U.S.A. Smith, M.R. 1943c: 7 (s.w.). Combination in P. (Macropheidole): Smith, M.R. 1943c: 5. Junior synonym of fimbriata: Wheeler, W.M. 1915b: 403; Emery, 1921f: 81. Revived from synonymy: Smith, M.R. 1943c: 7. See also: Gregg, 1949a: 70; Creighton, 1950a: 168; Wilson, 2003: 594.

Myrmicinae Pheidolini Pheidole rugulosa TRUE TRUE FALSE

rugulosa. Pheidole rugulosa Gregg, 1959: 26 (s.w.q.m.) U.S.A. See also: Wilson, 2003: 595.

Myrmicinae Pheidolini Pheidole sciara TRUE TRUE FALSE

sciara. Pheidole sciara Cole, 1955a: 47 (s.w.) U.S.A. See also: Wilson, 2003: 346.

Myrmicinae Pheidolini Pheidole sciophila TRUE TRUE FALSE

sciophila. Pheidole sciophila Wheeler, W.M. 1908e: 443, pl. 26, figs. 18, 19 (s.w.q.m.) U.S.A. Senior synonym of proserpina, semilaevicephala: Creighton & Gregg, 1955: 19. See also: Wilson, 2003: 347.

Myrmicinae Pheidolini Pheidole senex TRUE TRUE FALSE

senex. Pheidole senex Gregg, 1952a: 1 (s.w.) U.S.A. Senior synonym of anfracta: Cole, 1953e: 298; Gregg, 1959: 30. See also: Wilson, 2003: 596.

Myrmicinae Pheidolini Pheidole sitiens TRUE TRUE FALSE

sitiens. Pheidole sitiens Wilson, 2003: 348, figs. (s.w.) U.S.A.

Myrmicinae Pheidolini Pheidole soritis TRUE TRUE FALSE

soritis. Pheidole soritis Wheeler, W.M. 1908e: 439 (s.w.) U.S.A. Subspecies of sitarches: Creighton, 1950a: 190. Revived status as species: Cole, 1953e: 298. Junior synonym of sitarches: Cole, 1956c: 115. Revived from synonymy as subspecies of sitarches: Gregg, 1959: 30; Smith, D.R. 1979: 1373. Revived status as species and senior synonym of campestris, rufescens, sitarches, transvarians: Wilson, 2003: 598.

Myrmicinae Pheidolini Pheidole spadonia TRUE TRUE FALSE

spadonia. Pheidole spadonia Wheeler, W.M. 1915b: 400 (s.w.) U.S.A. Creighton & Gregg, 1955: 23 (q.). See also: Wilson, 2003: 600.

Myrmicinae Pheidolini Pheidole tertiaria TRUE TRUE TRUE

†tertiaria. †Pheidole tertiaria Carpenter, 1930: 33, pl. 5, fig. 2; pl. 11, fig. 2 (q.) U.S.A. (Oligocene).

Myrmicinae Pheidolini Pheidole tetra TRUE TRUE FALSE

tetra. Pheidole crassicornis subsp. tetra Creighton, 1950a: 176 (s.w.) U.S.A. [First available use of Pheidole crassicornis subsp. porcula var. tetra Wheeler, W.M. 1908e: 467; unavailable name.] Raised to species: Wilson, 2003: 161.

Myrmicinae Pheidolini Pheidole texana TRUE TRUE FALSE

texana. Pheidole texana Wheeler, W.M. 1903b: 97, fig. 4 (s.w.) U.S.A. See also: Wilson, 2003: 353.

Myrmicinae Pheidolini Pheidole titanis TRUE TRUE FALSE

titanis. Pheidole titanis Wheeler, W.M. 1903b: 95, fig. 3 (s.w.) U.S.A. See also: Wilson, 2003: 602.

Myrmicinae Pheidolini Pheidole tysoni TRUE TRUE FALSE

tysoni. Pheidole tysoni Forel, 1901e: 348 (s.w.m.) U.S.A. See also: Wilson, 2003: 528.

Myrmicinae Pheidolini Pheidole vallicola TRUE TRUE FALSE

vallicola. Pheidole crassicornis subsp. vallicola Wheeler, W.M. 1915b: 409 (s.w.) U.S.A. Raised to species: Creighton, 1950a: 191. See also: Wilson, 2003: 162.

Myrmicinae Pheidolini Pheidole virago TRUE TRUE FALSE

virago. Pheidole virago Wheeler, W.M. 1915b: 401 (s.w.) U.S.A. See also: Wilson, 2003: 604.

Myrmicinae Pheidolini Pheidole vistana TRUE TRUE FALSE

vistana. Pheidole longipes var. vistana Forel, 1914d: 272 (w.) U.S.A. Raised to species: Smith, D.R. 1979: 1374. Senior synonym of grallipes: Gregg, 1959: 22. See also: Wilson, 2003: 360.

Myrmicinae Pheidolini Pheidole wheelerorum TRUE TRUE FALSE

wheelerorum. Pheidole wheelerorum Mackay, in Mackay, Lowrie, Fisher, et al. 1988: 96, figs. 29, 30 (s.w.) U.S.A.

Myrmicinae Pheidolini Pheidole xerophila TRUE TRUE FALSE

xerophila. Pheidole xerophila Wheeler, W.M. 1908e: 446, pl. 27, fig. 37 (s.w.m.) U.S.A. Senior synonym of tucsonica: Wilson, 2003: 605.

Myrmicinae Pheidolini Pheidole yaqui TRUE TRUE FALSE

yaqui. Pheidole yaqui Creighton & Gregg, 1955: 43, fig. 10 (s.w.) U.S.A. See also: Wilson, 2003: 607.

Myrmicinae Pheidolini Pheidole pallidula koshewnikovi TRUE TRUE FALSE

koshewnikovi. Pheidole pallidula subsp. koshewnikovi Ruzsky, 1905b: 648 (s.w.) RUSSIA.

Myrmicinae Myrmicini Pogonomyrmex anergismus TRUE TRUE FALSE

anergismus. Pogonomyrmex (Pogonomyrmex) anergismus Cole, 1954b: 115 (q.m.) U.S.A. See also: Cole, 1968: 46; Johnson, R.A. 1995: 257.

Myrmicinae Myrmicini Pogonomyrmex anzensis TRUE TRUE FALSE

anzensis. Pogonomyrmex (Pogonomyrmex) anzensis Cole, 1968: 87, pl. 3, fig. 13; pl. 4, fig. 11; pl. 6, fig. 12; pl. 7, fig. 15 (w.) U.S.A. Snelling, R.R. et al. 2009: 316 (q.m.).

Myrmicinae Myrmicini Pogonomyrmex apache TRUE TRUE FALSE

apache. Pogonomyrmex apache Wheeler, W.M. 1902c: 392 (w.) U.S.A. Cole, 1954e: 269 (q.m.); Taber, Cokendolpher & Francke, 1988: 51 (k.). Senior synonym of sanctihyacinthi: Cole, 1954e: 267. See also: Cole, 1968: 49.

Myrmicinae Myrmicini Pogonomyrmex badius TRUE TRUE FALSE

badius. Formica badia Latreille, 1802c: 238, pl. 11, fig. 71 (w.q.) U.S.A. Lepeletier, 1835: 175 (m.); Wheeler, G.C. & Wheeler, J. 1960b: 2 (l.); Taber, Cokendolpher & Francke, 1988: 51 (k.). Combination in Aphaenogaster: Roger, 1863b: 30; in Pogonomyrmex: Mayr, 1870b: 971. Senior synonym of transversa (and its junior synonyms brevipennis, crudelis): Emery, 1895c: 310. See also: Cole, 1968: 148; Gordon, 1984: 413; Hölldobler & Wilson, 1971: 385; Smith, D.R. 1979: 1356.

Myrmicinae Myrmicini Pogonomyrmex bicolor TRUE TRUE FALSE

bicolor. Pogonomyrmex (Pogonomyrmex) bicolor Cole, 1968: 59, pl. 3, fig. 15; pl. 4, fig. 17; pl. 5, fig. 4; pl. 7, fig. 21; pl. 8, fig. 14; pl. 10, fig. 14; pl. 11, fig. 15 (w.q.m.) U.S.A.

Myrmicinae Myrmicini Pogonomyrmex bigbendensis TRUE TRUE FALSE

bigbendensis. Pogonomyrmex (Pogonomyrmex) bigbendensis Francke & Merickel, 1982: 371, figs. 1, 2, 9, 12, 15 (w.) U.S.A.

Myrmicinae Myrmicini Pogonomyrmex brevispinosus TRUE TRUE FALSE

brevispinosus. Pogonomyrmex (Pogonomyrmex) brevispinosus Cole, 1968: 89, pl.3, fig. 8; pl. 4, fig. 6; pl. 6, fig. 6; pl. 7, fig. 6; pl. 8, fig. 6; pl. 10, fig. 5; pl. 11, fig. 5 (w.q.m.) U.S.A. Taber, Cokendolpher & Francke, 1988: 51 (k.).

Myrmicinae Myrmicini Pogonomyrmex californicus TRUE TRUE FALSE

californicus. Myrmica californica Buckley, 1867: 336 (w.) U.S.A. Wheeler, W.M. 1914e: 153 (q.m.); Taber, Cokendolpher & Francke, 1988: 51 (k.). Combination in Pogonomyrmex: Emery, 1895c: 311. Senior synonym of estebanius, longinodis, nitratus: Cole, 1968: 120; of hindleyi: Snelling, R.R. 1982a: 98. See also: Tulloch, 1930: 61.

Myrmicinae Myrmicini Pogonomyrmex colei TRUE TRUE FALSE

colei. Pogonomyrmex (Pogonomyrmex) colei Snelling, R.R. 1982a: 99, figs. 1-11 (q.m.) U.S.A. See also: Rissing, 1983: 321; Johnson, R.A., Parker & Rissing, 1996: 69.

Myrmicinae Myrmicini Pogonomyrmex comanche TRUE TRUE FALSE

comanche. Pogonomyrmex occidentalis subsp. comanche Wheeler, W.M. 1902c: 392 (w.) U.S.A. Cole, 1968: 126 (q.m.); Taber, Cokendolpher & Francke, 1988: 51 (k.). Subspecies of occidentalis: Creighton, 1950a: 128; Gregg, 1963: 333. Status as species: Wheeler, W.M. 1914e: 156; Olsen, 1934: 498; Smith, M.R. 1951a: 793; Cole, 1968: 126.

Myrmicinae Myrmicini Pogonomyrmex desertorum TRUE TRUE FALSE

desertorum. Pogonomyrmex desertorum Wheeler, W.M. 1902c: 387 (w.) U.S.A. Cole, 1968: 67 (q.m.); Taber, Cokendolpher & Francke, 1988: 51 (k.). Senior synonym of ferrugineus: Cole, 1968: 65.

Myrmicinae Myrmicini Pogonomyrmex fossilis TRUE TRUE TRUE

†fossilis. †Pogonomyrmex fossilis Carpenter, 1930: 33, pl. 9, fig. 6 (w.) U.S.A. (Oligocene).

Myrmicinae Myrmicini Pogonomyrmex huachucanus TRUE TRUE FALSE

huachucanus. Pogonomyrmex huachucanus Wheeler, W.M. 1914e: 151 (w.) U.S.A. Creighton, 1952b: 79 (q.m.); Wheeler, G.C. & Wheeler, J. 1972b: 237 (l.); Taber, Cokendolpher & Francke, 1988: 51 (k.). Combination in P. (Ephebomyrmex): Cole, 1968: 157.

Myrmicinae Myrmicini Pogonomyrmex imberbiculus TRUE TRUE FALSE

imberbiculus. Pogonomyrmex imberbiculus Wheeler, W.M. 1902a: 87, figs. 1, 2 (w.) U.S.A. Creighton, 1957a: 63 (q.); Cole, 1968: 171 (m.); Wheeler, G.C. & Wheeler, J. 1972b: 237 (l.); Taber, Cokendolpher & Francke, 1988: 51 (k.); Heinze, Hölldobler & Cover, 1992: 267 (ergatoid q.). Combination in P. (Ephebomyrmex): Wheeler, W.M. 1902c: 390. Senior synonym of townsendi: Creighton, 1957a: 56.

Myrmicinae Myrmicini Pogonomyrmex magnacanthus TRUE TRUE FALSE

magnacanthus. Pogonomyrmex (Pogonomyrmex) magnacanthus Cole, 1968: 133, pl. 2, fig. 5; pl. 3, fig. 12; pl. 4, fig. 10; pl. 6, fig. 9; pl. 7, fig. 16; pl. 8, fig. 10; pl. 11, fig. 10 (w.q.m.) U.S.A. Taber, Cokendolpher & Francke, 1988: 51 (k.).

Myrmicinae Myrmicini Pogonomyrmex maricopa TRUE TRUE FALSE

maricopa. Pogonomyrmex californicus subsp. maricopa Wheeler, W.M. 1914e: 155 (w.q.) U.S.A. Cole, 1968: 140 (m.); Taber, Cokendolpher & Francke, 1988: 51 (k.). Raised to species: Creighton, 1950a: 126. Senior synonym of barnesi: Cole, 1968: 138; of sinaloanus: Brown, in Cole, 1954b: 120.

Myrmicinae Myrmicini Pogonomyrmex mohavensis TRUE TRUE FALSE

mohavensis. Pogonomyrmex mohavensis Johnson, in Johnson, R.A. & Overson, 2009: 308, figs. 1A-E (w.) U.S.A.

Myrmicinae Myrmicini Pogonomyrmex montanus TRUE TRUE FALSE

montanus. Pogonomyrmex montanus Mackay, 1980: 151, figs. 1-7 (w.q.m.) U.S.A. Taber, Cokendolpher & Francke, 1988: 51 (k.). See also: Mackay, 1981: 25; Shattuck, 1987: 172.

Myrmicinae Myrmicini Pogonomyrmex occidentalis TRUE TRUE FALSE

occidentalis. Myrmica occidentalis Cresson, 1865: 426 (w.q.) U.S.A. Cole, 1968: 94 (m.); Wheeler, G.C. & Wheeler, J. 1953a: 110 (l.); Taber, Cokendolpher & Francke, 1988: 51 (k.); Taber & Francke, 1986: 274 (gynandromorph). Combination in Pogonomyrmex: Cresson, 1887: 22. Senior synonym of opaciceps, seminigra: Mayr, 1886d: 449; Forel, 1886b: xlii; of ruthveni: Olsen, 1934: 507; of utahensis: Cole, 1968: 95. See also: Smith, D.R. 1979: 1355; Taber & Francke, 1986: 274; Shattuck, 1987: 172.

Myrmicinae Myrmicini Pogonomyrmex pima TRUE TRUE FALSE

pima. Pogonomyrmex (Ephebomyrmex) pima Wheeler, W.M. 1909e: 79 (w.) U.S.A. Cole, 1968: 175 (q.m.); Johnson, R.A. et al. 2007: 79 (ergatoid q.).

Myrmicinae Myrmicini Pogonomyrmex rugosus TRUE TRUE FALSE

rugosus. Pogonomyrmex barbatus subsp. rugosus Emery, 1895c: 309 (w.m.) U.S.A. Cole, 1968: 72 (q.); Taber, Cokendolpher & Francke, 1988: 51 (k.). Raised to species: Cole, 1968: 70. Senior synonym of similis: Creighton, 1950a: 120; of fuscatus (and its junior synonyms curvispinosus, marfensis), spadix: Cole, 1968: 70.

Myrmicinae Myrmicini Pogonomyrmex salinus TRUE TRUE FALSE

salinus. Pogonomyrmex salinus Olsen, 1934: 510, pl. 5, fig. 3 (w.) U.S.A. Cole, 1968: 108 (q.m.); Wheeler, G.C. & Wheeler, J. 1972b: 237 (l.). Junior synonym of occidentalis: Smith, M.R. 1953g: 132. Revived from synonymy: Cole, 1968: 106. Senior synonym of owyheei: Shattuck, 1987: 173.

Myrmicinae Myrmicini Pogonomyrmex subdentatus TRUE TRUE FALSE

subdentatus. Pogonomyrmex subdentatus Mayr, 1870b: 971 (diagnosis in key) (w.) U.S.A. Cole, 1968: 111 (q.m.). See also: Shattuck, 1987: 175.

Myrmicinae Myrmicini Pogonomyrmex subnitidus TRUE TRUE FALSE

subnitidus. Pogonomyrmex occidentalis var. subnitidus Emery, 1895c: 310 (w.) U.S.A. Cole, 1968: 115 (q.m.); Taber, Cokendolpher & Francke, 1988: 51 (k.). Raised to species: Wheeler, W.M. 1914e: 156. See also: Mackay, 1981: 25.

Myrmicinae Myrmicini Pogonomyrmex tenuispinus TRUE TRUE FALSE

tenuispinus. Pogonomyrmex desertorum var. tenuispina Forel, 1914d: 269 (w.) U.S.A. Subspecies of desertorum: Creighton, 1950a: 125. Raised to species and senior synonym of dentatus: Cole, 1968: 77.

Myrmicinae Myrmicini Pogonomyrmex texanus TRUE TRUE FALSE

texanus. Pogonomyrmex (Pogonomyrmex) texanus Francke & Merickel, 1982: 375, figs. 4, 7, 8, 11, 13, 14, 16 (w.m.) U.S.A. Taber, 1988: 244 (q.).

Formicinae Formicini Polyergus breviceps TRUE TRUE FALSE

breviceps. Polyergus rufescens subsp. breviceps Emery, 1893i: 666 (w.) U.S.A. Raised to species: Kannowski, 1956: 185; Wheeler, J. 1968: 163. Senior synonym of bicolor, umbratus: Wheeler, J. 1968: 163. Material of the unavailable names fusciventris, montezuma, silvestrii referred here by Creighton, 1950a: 559. See also: Hölldobler, 1985: 225.

Formicinae Formicini Polyergus lucidus TRUE TRUE FALSE

1 subspecies

lucidus. Polyergus lucidus Mayr, 1870b: 952 (w.q.m.) U.S.A. Wheeler, G.C. & Wheeler, J. 1968: 214 (l.); Wheeler, W.M. 1903f: 659 (gynandromorph). Subspecies of rufescens: Forel, 1886f: 200. Revived status as species: Dalla Torre, 1893: 214; Wheeler, W.M. 1917i: 465; Smith, M.R. 1947g: 152; Creighton, 1950a: 557. Senior synonym of montivagus: Creighton, 1950a: 557. Current subspecies: nominal plus longicornis. See also: Talbot, 1968: 299.

Formicinae Formicini Polyergus nigerrimus TRUE TRUE FALSE

nigerrimus. Polyergus nigerrimus Marikovsky, 1963: 110, figs. 1-7 (w.q.m.) RUSSIA. See also: Kupyanskaya, 1990: 208.

Formicinae Formicini Polyergus texana FALSE FALSE FALSE

texana. Polyergus texana Buckley, 1866: 170 (q.) U.S.A. Unidentifiable to genus; incertae sedis in Polyergus: Bolton, 1995b: 342. See also: Wheeler, W.M. 1902f: 24.

Formicinae Formicini Polyergus lucidus longicornis TRUE TRUE FALSE

longicornis. Polyergus lucidus subsp. longicornis Smith, M.R. 1947g: 155 (w.) U.S.A.

Formicinae Formicini Polyergus rufescens laeviceps TRUE TRUE FALSE

laeviceps. Polyergus rufescens subsp. laeviceps Wheeler, W.M. 1915b: 420 (w.) U.S.A. See also: Creighton, 1950a: 559.

Ponerinae Ponerini Ponera exotica TRUE TRUE FALSE

exotica. Ponera exotica Smith, M.R. 1962b: 378, fig. 1 (w.q.) U.S.A. See also: Taylor, 1967a: 96; Johnson, C. 1987: 358.

Ponerinae Ponerini Ponera pennsylvanica TRUE TRUE FALSE

pennsylvanica. Ponera pennsylvanica Buckley, 1866: 171 (w.) U.S.A. Emery, 1895c: 267 (q.m.); Wheeler, G.C. & Wheeler, J. 1952c: 631 (l.). Subspecies of coarctata: Emery, 1895c: 267; Dennis, 1938: 277; Creighton, 1950a: 48. Revived status as species: Taylor, 1967a: 29.

Ponerinae Ponerini Ponerites coloradensis TRUE TRUE TRUE

†coloradensis. †Ponerites coloradensis Dlussky & Rasnitsyn, 2002: 423, figs. 22, 23 (q.) U.S.A. (Eocene).

Ponerinae Ponerini Ponerites eocenicus TRUE TRUE TRUE

†eocenicus. †Ponerites eocenicus Dlussky & Rasnitsyn, 2002: 423, fig. 21 (q.) U.S.A. (Eocene).

Ponerinae Ponerini Ponerites hypoponeroides TRUE TRUE TRUE

†hypoponeroides. †Ponerites hypoponeroides Dlussky & Rasnitsyn, 2002: 424, fig. 24 (q.) U.S.A. (Eocene).

Ponerinae Ponerini Ponerites karaganensis TRUE TRUE TRUE

†karaganensis. †Ponerites karaganensis Dlussky, 1981b: 68, fig. 50 (wing) RUSSIA (Miocene).

Ponerinae Ponerini Ponerites stauropolitanus TRUE TRUE TRUE

†stauropolitanus. †Ponerites stauropolitanus Dlussky, 1981b: 67, fig. 50 (wing) RUSSIA (Miocene).

Armaniinae Armaniini Poneropterus sphecoides TRUE TRUE TRUE

†sphecoides. †Poneropterus sphecoides Dlussky, 1983: 73, fig. 7 (m.) RUSSIA (Cretaceous).

Formicinae Plagiolepidini Prenolepis imparis TRUE TRUE FALSE

4 subspecies

imparis. Formica imparis Say, 1836: 287 (q.m.) U.S.A. Wheeler, W.M. 1930b: 16 (w.); Wheeler, G.C. & Wheeler, J. 1953c: 142 (l.); Hauschteck, 1962: 219 (k.). Combination in Prenolepis: Mayr, 1886d: 431. Senior synonym of americana: Emery, 1893i: 635; Dalla Torre, 1893: 178; of wichita: Dalla Torre, 1893: 178; Wheeler, W.M. 1930b: 15; of minuta, pumila, testacea: Creighton, 1950a: 414; of californica: Wheeler, G.C. & Wheeler, J. 1986g: 14. Current subspecies: nominal plus arizonica, colimana, coloradensis, veracruzensis. See also: Wheeler, W.M. 1930b: 16.

Formicinae Plagiolepidini Prenolepis imparis coloradensis TRUE TRUE FALSE

coloradensis. Prenolepis imparis var. coloradensis Wheeler, W.M. 1930b: 22, fig. 4 (w.) U.S.A. Currently subspecies of imparis: Creighton, 1950a: 415.

Formicinae Plagiolepidini Prenolepis imparis arizonica TRUE TRUE FALSE

arizonica. Prenolepis imparis var. arizonica Wheeler, W.M. 1930b: 22 (w.q.m.) U.S.A. Currently subspecies of imparis: Creighton, 1950a: 414.

Proceratiinae Proceratiini Proceratium californicum TRUE TRUE FALSE

californicum. Proceratium californicum Cook, 1953: 45, fig. (m.) U.S.A. Snelling, R.R. 1967: 4 (q.); Ward, 1988: 109 (w.). See also: Brown, 1958g: 332; Baroni Urbani & De Andrade, 2003b: 243.

Proceratiinae Proceratiini Proceratium chickasaw TRUE TRUE FALSE

chickasaw. Proceratium chickasaw De Andrade, in Baroni Urbani & De Andrade, 2003b: 235, figs. 98, 99 (w.) U.S.A.

Proceratiinae Proceratiini Proceratium compitale TRUE TRUE FALSE

compitale. Proceratium compitale Ward, 1988: 113, fig. 6 (w.q.) U.S.A. See also: Baroni Urbani & De Andrade, 2003b: 203.

Proceratiinae Proceratiini Proceratium crassicorne TRUE TRUE FALSE

crassicorne. Proceratium crassicorne Emery, 1895c: 265, pl. 8, fig. 9 (w.) U.S.A. Smith, M.R. 1928b: 244 (q.); Wheeler, G.C. & Wheeler, J. 1952a: 137 (l.); Baroni Urbani & De Andrade, 2003b: 426 (m.). Junior synonym of silaceum: Creighton, 1950a: 40; Brown, 1958g: 248. Revived from synonymy: Baroni Urbani & De Andrade, 2003b: 422.

Proceratiinae Proceratiini Proceratium creek TRUE TRUE FALSE

creek. Proceratium creek De Andrade, in Baroni Urbani & De Andrade, 2003b: 207, figs. 86-89 (w.q.m.) U.S.A.

Proceratiinae Proceratiini Proceratium croceum TRUE TRUE FALSE

croceum. Ponera crocea Roger, 1860: 288 (q.) U.S.A. Emery, 1895c: 265 (w.); Smith, M.R. 1930a: 390 (m.); Wheeler, G.C. & Wheeler, J. 1952a: 135 (l.). Combination in Sysphingta: Mayr, 1866a: 501; in Proceratium: Mayr, 1886d: 437. See also: Baroni Urbani & De Andrade, 2003b: 397.

Proceratiinae Proceratiini Proceratium melinum TRUE TRUE FALSE

melinum. Ponera melina Roger, 1860: 291 (w.q.m.) EUROPE (not U.S.A.; see Baroni Urbani, 1977d: 91). Combination in Proceratium: Mayr, 1886d: 438; in Sysphincta: Emery, 1895c: 263; in Proceratium: Brown, 1958g: 248. Senior synonym of europaea, fialai, rossica: Brown, 1958g: 248. See also: Atanassov & Dlussky, 1992: 64; Baroni Urbani & De Andrade, 2003b: 251.

Proceratiinae Proceratiini Proceratium pergandei TRUE TRUE FALSE

pergandei. Sysphincta pergandei Emery, 1895c: 264, pl. 8, fig. 4 (w.) U.S.A. Smith, M.R. 1928b: 242 (m.). Baroni Urbani & De Andrade, 2003b: 227 (q.). Combination in Proceratium: Brown, 1958g: 248.

Proceratiinae Proceratiini Proceratium silaceum TRUE TRUE FALSE

silaceum. Proceratium silaceum Roger, 1863a: 172 (w.) U.S.A. Emery, 1895c: 265 (q.); Kennedy & Talbot, 1939: 202 (m.); Crozier, 1970: 116 (k.). Senior synonym of rugulosum, vestitum: Creighton, 1950a: 40. See also: Brown, 1958g: 330; Brown, 1974a: 82; Baroni Urbani & De Andrade, 2003b: 408.

Myrmicinae Formicoxenini Protomognathus americanus TRUE TRUE FALSE

americanus. Tomognathus americanus Emery, 1895c: 272 (w.) U.S.A. Creighton, 1927: 27 (q.m.); Wheeler, G.C. & Wheeler, J. 1955b: 26 (l.). Combination in T. (Protomognathus): Wheeler, W.M. 1905a: 3; in Harpagoxenus: Wheeler, W.M. 1910g: 494; in Protomognathus: Cover, in Hölldobler & Wilson, 1990: 65. See also: Smith, M.R. 1939b: 166.

Ponerinae Ponerini Protopone primigena TRUE TRUE TRUE

†primigena. †Protopone primigena Dlussky, 1988: 53, fig. 2 (w.) RUSSIA (Palaeocene).

Armaniinae Armaniini Pseudarmania aberrans TRUE TRUE TRUE

†aberrans. †Pseudarmania aberrans Dlussky, 1983: 70, fig. 4 (q.) RUSSIA (Cretaceous).

Armaniinae Armaniini Pseudarmania rasnitsyni TRUE TRUE TRUE

†rasnitsyni. †Pseudarmania rasnitsyni Dlussky, 1983: 69, fig. 3 (q.) RUSSIA (Cretaceous).

Formicinae Camponotini Pseudocamponotus elkoanus TRUE TRUE TRUE

†elkoanus. †Pseudocamponotus elkoanus Carpenter, 1930: 22, pl. 2, fig. 2 (q.) U.S.A. (Miocene).

Pseudomyrmecinae Pseudomyrmecini Pseudomyrmex apache TRUE TRUE FALSE

apache. Pseudomyrmex apache Creighton, 1953b: 134, pl. 12, figs. 1-7 (w.q.m.) U.S.A. Wheeler, G.C. & Wheeler, J. 1956: 380 (l.). See also: Creighton, 1954: 9; Ward, 1985b: 229.

Pseudomyrmecinae Pseudomyrmecini Pseudomyrmex extinctus TRUE TRUE TRUE

†extinctus. †Pseudomyrma extincta Carpenter, 1930: 29, pl. 3, fig. 4 (q.) U.S.A. (Oligocene). Combination in Pseudomyrmex: Ward, 1990: 487.

Pseudomyrmecinae Pseudomyrmecini Pseudomyrmex leptosus TRUE TRUE FALSE

leptosus. Pseudomyrmex leptosus Ward, 1985b: 233, figs. 19, 26, 34, 35, 38 (q.m.) U.S.A.

Pseudomyrmecinae Pseudomyrmecini Pseudomyrmex pallidus TRUE TRUE FALSE

pallidus. Pseudomyrma pallidus Smith, F. 1855c: 160 (w.) U.S.A. Wheeler, G.C. & Wheeler, J. 1956: 386 (l.); Petralia & Vinson, 1980: 381 (l.). Combination in Leptalea: Smith, M.R. 1951a: 788; in Pseudomyrmex: Wheeler, G.C. & Wheeler, J. 1956: 386. Subspecies of flavidulus: Enzmann, E.V. 1944: 66. Revived status as species: Smith, M.R. 1947f: 544; Creighton, 1950a: 80. See also: Blum & Callahan, 1963: 69; Ward, 1985b: 235.

Pseudomyrmecinae Pseudomyrmecini Pseudomyrmex seminole TRUE TRUE FALSE

seminole. Pseudomyrmex seminole Ward, 1985b: 237, figs. 18, 21, 28, 44, 45 (w.q.m.) U.S.A. Wheeler, G.C. & Wheeler, J. 1991b: 90 (l.).

Myrmicinae Dacetini Pyramica abdita TRUE TRUE FALSE

abdita. Strumigenys (Cephaloxys) abdita Wesson L.G. & Wesson, R.G. 1939: 106, pl. 3, fig. 6 (w.) U.S.A. Brown, 1953g: 89 (q.); Brown, 1964a: 189 (m.). Combination in S. (Trichoscapa): Smith, M.R., 1947f: 587; Creighton, 1950a: 304; in Smithistruma: Smith, M.R., 1951a: 827; Brown, 1953g: 89; in Pyramica: Bolton, 1999: 1672. See also: Bolton, 2000: 118.

Myrmicinae Dacetini Pyramica angulata TRUE TRUE FALSE

angulata. Strumigenys (Cephaloxys) angulata Smith, M.R. 1931c: 697, pl. 1, fig. 3 (w.) U.S.A. Brown, 1953g: 54 (q.). Combination in S. (Trichoscapa): Smith, M.R., 1947f: 587; Creighton, 1950a: 304; in Smithistruma (Wessonistruma): Brown, 1948e: 106; Brown, 1953g: 54; in Pyramica: Bolton, 1999: 1673. See also: Bolton, 2000: 115.

Myrmicinae Dacetini Pyramica apalachicolensis TRUE TRUE FALSE

apalachicolensis. Pyramica apalachicolensis Deyrup & Lubertazzi, 2001: 15, figs. 1-3 (w.) U.S.A.

Myrmicinae Dacetini Pyramica archboldi TRUE TRUE FALSE

archboldi. Smithistruma archboldi Deyrup & Cover, 1998: 217, fig. 2 (w.q.) U.S.A. Combination in Pyramica: Bolton, 1999: 1673. See also: Bolton, 2000: 131.

Myrmicinae Dacetini Pyramica arizonica TRUE TRUE FALSE

arizonica. Smithistruma arizonica Ward, 1988: 121 (w.m.) U.S.A. Combination in Pyramica: Bolton, 1999: 1673. See also: Bolton, 2000: 125.

Myrmicinae Dacetini Pyramica bimarginata TRUE TRUE FALSE

bimarginata. Strumigenys (Cephaloxys) bimarginata Wesson L.G. & Wesson, R.G. 1939: 95, pl. 3, fig. 2 (w.q.) U.S.A. Combination in S. (Trichoscapa): Smith, M.R., 1947f: 587; Creighton, 1950a: 304; in Smithistruma: Smith, M.R., 1951a: 827; Brown, 1953g: 62; in Pyramica: Bolton, 1999: 1673. See also: Bolton, 2000: 103.

Myrmicinae Dacetini Pyramica boltoni TRUE TRUE FALSE

boltoni. Pyramica boltoni Deyrup, 2006: 1, fig. 1 (w.q.) U.S.A.

Myrmicinae Dacetini Pyramica bunki TRUE TRUE FALSE

bunki. Smithistruma (Smithistruma) bunki Brown, 1950b: 41, pl. 3, figs. 2, 3 (w.) U.S.A. Brown, 1953g: 83 (q.). Combination in Pyramica: Bolton, 1999: 1673. See also: Bolton, 2000: 125.

Myrmicinae Dacetini Pyramica californica TRUE TRUE FALSE

californica. Smithistruma (Smithistruma) californica Brown, 1950b: 40 (q.) U.S.A. Ward, 1988: 120 (w.). Combination in Pyramica: Bolton, 1999: 1673. See also: Brown, 1953g: 85; Bolton, 2000: 126.

Myrmicinae Dacetini Pyramica carolinensis TRUE TRUE FALSE

carolinensis. Smithistruma carolinensis Brown, 1964a: 185, pl. 16, fig. 2 (w.) U.S.A. Combination in Pyramica: Bolton, 1999: 1673. See also: Bolton, 2000: 127.

Myrmicinae Dacetini Pyramica chiricahua TRUE TRUE FALSE

chiricahua. Smithistruma chiricahua Ward, 1988: 119, fig. 8 (w.) U.S.A. Combination in Pyramica: Bolton, 1999: 1673. See also: Bolton, 2000: 127.

Myrmicinae Dacetini Pyramica cloydi TRUE TRUE FALSE

cloydi. Smithistruma cloydi Pfitzer, 1951: 198, figs. (w.) U.S.A. Combination in Pyramica: Bolton, 1999: 1673. See also: Bolton, 2000: 111.

Myrmicinae Dacetini Pyramica clypeata TRUE TRUE FALSE

clypeata. Strumigenys clypeata Roger, 1863a: 213 (w.) U.S.A. [Misspelled as clipeata in original description; justified emendation to clypeata by Mayr, 1887: 571.] Emery, 1895c: 328 (q.m.). Combination in S. (Cephaloxys): Emery, 1924d: 325; in S. (Trichoscapa): Smith, M.R., 1947f: 587; Creighton, 1950a: 304; in Smithistruma: Smith, M.R., 1951a: 827; Brown, 1953g: 58; in Pyramica: Bolton, 1999: 1673. See also: Wilson, 1954: 486; Smith, D.R. 1979: 1406; Bolton, 2000: 103.

Myrmicinae Dacetini Pyramica creightoni TRUE TRUE FALSE

creightoni. Strumigenys (Cephaloxys) creightoni Smith, M.R. 1931c: 705, pl. 4, fig. 16 (w.) U.S.A.Brown, 1953g: 80 (q.); Brown, 1964a: 191 (m.). Combination in S. (Trichoscapa): Smith, M.R., 1947f: 587; Creighton, 1950a: 305; in Smithistruma: Smith, M.R., 1951a: 827; Brown, 1953g: 80; in Pyramica: Bolton, 1999: 1673. See also: Bolton, 2000: 118.

Myrmicinae Dacetini Pyramica deyrupi TRUE TRUE FALSE

deyrupi. Pyramica deyrupi Bolton, 2000: 119 (w.) U.S.A.

Myrmicinae Dacetini Pyramica dietrichi TRUE TRUE FALSE

dietrichi. Strumigenys (Cephaloxys) dietrichi Smith, M.R. 1931c: 696, pl. 2, fig. 6 (w.) U.S.A. Brown, 1953g: 67 (q.). Combination in S. (Trichoscapa): Smith, M.R., 1947f: 587; Creighton, 1950a: 305; in Smithistruma: Smith, M.R., 1951a: 827; Brown, 1953g: 67; in Pyramica: Bolton, 1999: 1673. See also: Kennedy & Schramm, 1933: 100; Wilson, 1954: 487; Bolton, 2000: 112.

Myrmicinae Dacetini Pyramica filirrhina TRUE TRUE FALSE

filirrhina. Smithistruma (Smithistruma) filirrhina Brown, 1950b: 37 (w.) U.S.A. Combination in Pyramica: Bolton, 1999: 1673. See also: Brown, 1953g: 63; Bolton, 2000: 104.

Myrmicinae Dacetini Pyramica filitalpa TRUE TRUE FALSE

filitalpa. Smithistruma (Smithistruma) filitalpa Brown, 1950b: 39, pl. 3, fig. 4 (w.) U.S.A. Combination in Pyramica: Bolton, 1999: 1673. See also: Brown, 1953g: 79; Bolton, 2000: 132.

Myrmicinae Dacetini Pyramica hyalina TRUE TRUE FALSE

hyalina. Pyramica hyalina Bolton, 2000: 128 (w.q.) U.S.A.

Myrmicinae Dacetini Pyramica inopina TRUE TRUE FALSE

inopina. Smithistruma inopina Deyrup & Cover, 1998: 215, fig. 1 (q.) U.S.A. Combination in Pyramica: Bolton, 1999: 1673. See also: Bolton, 2000: 129.

Myrmicinae Dacetini Pyramica laevinasis TRUE TRUE FALSE

laevinasis. Strumigenys (Cephaloxys) clypeata var. laevinasis Smith, M.R. 1931c: 701, pl. 3, fig. 11 (w.) U.S.A. Brown, 1953g: 62 (q.). Combination in S. (Trichoscapa): Smith, M.R., 1947f: 587; Creighton, 1950a: 308; in Smithistruma: Smith, M.R., 1951a: 827; Brown, 1953g: 61; in Pyramica: Bolton, 1999: 1673. Subspecies of pilinasis: Creighton, 1950a: 308. Raised to species: Brown, 1953g: 61. See also: Bolton, 2000: 105.

Myrmicinae Dacetini Pyramica memorialis TRUE TRUE FALSE

memorialis. Smithistruma memorialis Deyrup, 1998: 81, fig. 1 (w.q.) U.S.A. Combination in Pyramica: Bolton, 1999: 1673. See also: Bolton, 2000: 120.

Myrmicinae Dacetini Pyramica metazytes TRUE TRUE FALSE

metazytes. Pyramica metazytes Bolton, 2000: 121, fig. 108 (w.) U.S.A.

Myrmicinae Dacetini Pyramica missouriensis TRUE TRUE FALSE

missouriensis. Strumigenys (Cephaloxys) missouriensis Smith, M.R. 1931c: 701, pl. 4, fig. 14 (w.) U.S.A. Brown, 1953g: 72 (q.). Combination in S. (Trichoscapa): Smith, M.R., 1947f: 587; Creighton, 1950a: 307; in Smithistruma: Smith, M.R., 1951a: 828; Brown, 1953g: 72; in Pyramica: Bolton, 1999: 1673. Senior synonym of sculpturata: Smith, M.R. 1951a: 828; Brown, 1953g: 72. See also: Wilson, 1954: 485; Bolton, 2000: 121.

Myrmicinae Dacetini Pyramica ohioensis TRUE TRUE FALSE

ohioensis. Strumigenys ohioensis Kennedy & Schramm, 1933: 98, figs. 1, 2 (w.) U.S.A. Brown, 1953g: 87 (q.m.). Combination in S. (Cephaloxys): Wesson, L.G. & Wesson, R.G., 1939: 108; in S. (Trichoscapa): Smith, M.R., 1947f: 587; Creighton, 1950a: 308; in Smithistruma: Smith, M.R.1951a: 828; Brown, 1953g: 87; in Pyramica: Bolton, 1999: 1673. Senior synonym of manni: Smith, M.R. 1951a: 828; Brown, 1953g: 87. See also: Bolton, 2000: 109.

Myrmicinae Dacetini Pyramica ornata TRUE TRUE FALSE

ornata. Strumigenys ornata Mayr, 1887: 571 (footnote) (w.) U.S.A. Brown, 1953g: 66 (q.). Combination in S. (Cephaloxys): Emery, 1924d: 325; in S. (Trichoscapa): Smith, M.R., 1947f: 587; Creighton, 1950a: 308; in Smithistruma: Smith, M.R., 1951a: 828; Brown, 1953g: 65; in Pyramica: Bolton, 1999: 1673. See also: Bolton, 2000: 113.

Myrmicinae Dacetini Pyramica pergandei TRUE TRUE FALSE

pergandei. Strumigenys pergandei Emery, 1895c: 326, pl. 8, figs. 17, 18 (w.q.m.) U.S.A. Wheeler, G.C. & Wheeler, J. 1955a: 144 (l.). Combination in S. (Cephaloxys): Emery, 1924d: 325; in S. (Trichoscapa): Smith, M.R., 1943f: 307; Creighton, 1950a: 308; in Smithistruma (Wessonistruma): Brown, 1948e: 106; Brown, 1953g: 51; in Pyramica: Bolton, 1999: 1673. See also: Bolton, 2000: 116.

Myrmicinae Dacetini Pyramica pilinasis TRUE TRUE FALSE

pilinasis. Strumigenys clypeata var. pilinasis Forel, 1901e: 339 (w.) U.S.A. Combination in S. (Cephaloxys): Emery, 1924d: 325; in S. (Trichoscapa): Smith, M.R., 1947f: 587; in Smithistruma: Smith, M.R., 1951a: 827; Brown, 1953g: 60; in Pyramica: Bolton, 1999: 1673. Raised to species: Wesson L.G. & Wesson, R.G. 1939: 109. Senior synonym of medialis: Smith, M.R. 1951a: 828; Brown, 1953g: 60; of brevisetosa: Brown, 1964a: 197. See also: Bolton, 2000: 106.

Myrmicinae Dacetini Pyramica pulchella TRUE TRUE FALSE

pulchella. Strumigenys pulchella Emery, 1895c: 327, pl. 8, fig. 19 (w.) U.S.A. Brown, 1953g: 71 (q.m.). Combination in S. (Cephaloxys): Emery, 1924d: 325; in S. (Trichoscapa): Smith, M.R., 1943f: 307; Creighton, 1950a: 309; in Smithistruma: Smith, M.R. 1951a: 828; Brown, 1953g: 70; in Pyramica: Bolton, 1999: 1673. See also: Bolton, 2000: 122.

Myrmicinae Dacetini Pyramica reflexa TRUE TRUE FALSE

reflexa. Strumigenys (Cephaloxys) reflexa Wesson L.G. & Wesson, R.G. 1939: 102, pl. 3, fig. 4 (w.) U.S.A. Brown, 1953g: 74 (q.m.). Combination in S. (Trichoscapa): Smith, M.R., 1943f: 307; Creighton, 1950a: 309; in Smithistruma: Smith, M.R., 1951a: 828; Brown, 1953g: 73; in Pyramica: Bolton, 1999: 1673. See also: Bolton, 2000: 123.

Myrmicinae Dacetini Pyramica reliquia TRUE TRUE FALSE

reliquia. Smithistruma reliquia Ward, 1988: 117, fig. 7 (w.) U.S.A. Combination in Pyramica: Bolton, 1999: 1673. See also: Bolton, 2000: 110.

Myrmicinae Dacetini Pyramica rohweri TRUE TRUE FALSE

rohweri. Strumigenys (Cephaloxys) rohweri Smith, M.R. 1935: 214 (w.) U.S.A. Combination in S. (Trichoscapa): Smith, M.R., 1947f: 587; Creighton, 1950a: 309; in Smithistruma: Smith, M.R., 1951a: 828; Brown, 1953g: 57; in Pyramica: Bolton, 1999: 1673. See also: Bolton, 2000: 107.

Myrmicinae Dacetini Pyramica rostrata TRUE TRUE FALSE

rostrata. Strumigenys rostrata Emery, 1895c: 329, pl. 8, figs. 23, 24 (w.q.m.) U.S.A. Wheeler, G.C. & Wheeler, J. 1955a: 144 (l.). Combination in S. (Cephaloxys): Emery, 1924d: 325; in S. (Trichoscapa): Smith, M.R., 1943f: 307; Creighton, 1950a: 309; in Smithistruma: Smith, M.R., 1951a: 828; Brown, 1953g: 84; in Pyramica: Bolton, 1999: 1673. See also: Wilson, 1954: 485; Bolton, 2000: 129.

Myrmicinae Dacetini Pyramica subnuda TRUE TRUE FALSE

subnuda. Pyramica subnuda MacGown & Hill, 2010: 572, fig. 1 (q.) U.S.A.

Myrmicinae Dacetini Pyramica talpa TRUE TRUE FALSE

talpa. Strumigenys (Cephaloxys) talpa Weber, 1934b: 63, fig. 1 (w.) U.S.A. Brown, 1953g: 77 (q.m.); Wheeler, G.C. & Wheeler, J. 1955a: 141 (l.). Combination in S. (Trichoscapa): Smith, M.R., 1947f: 587; Creighton, 1950a: 310; in Smithistruma: Smith, M.R., 1951a: 828; Brown, 1953g: 76; in Pyramica: Bolton, 1999: 1673. Senior synonym of venatrix: Wesson, 1949: 21. See also: Wilson, 1954: 486; Bolton, 2000: 132.

Myrmicinae Dacetini Pyramica wrayi TRUE TRUE FALSE

wrayi. Smithistruma (Smithistruma) wrayi Brown, 1950b: 38, pl. 3, fig. 5 (w.) U.S.A. Combination in Pyramica: Bolton, 1999: 1673. See also: Brown, 1953g: 78; Bolton, 2000: 133.

Ectatomminae Ectatommini Rhytidoponera kirghizorum TRUE TRUE TRUE

†kirghizorum. †Rhytidoponera kirghizorum Dlussky, 1981b: 66, fig. 50 (w.) RUSSIA (Miocene).

Myrmicinae Stenammini Rogeria creightoni TRUE TRUE FALSE

creightoni. Rogeria creightoni Snelling, R.R. 1973a: 2, fig. 1 (w.) U.S.A. See also: Kugler, C. 1994: 50.

Formicinae Formicini Rossomyrmex proformicarum TRUE TRUE FALSE

proformicarum. Rossomyrmex proformicarum Arnol'di, 1928c: 300, figs. 1-6 (w.) RUSSIA. See also: Arnol'di, 1932a: 319.

Myrmicinae Solenopsidini Solenopsis abdita TRUE TRUE FALSE

abdita. Solenopsis abdita Thompson, 1989: 275, figs. 1-4 (w.q.m.) U.S.A. See also: Thompson & Johnson, 1989: 698.

Myrmicinae Solenopsidini Solenopsis amblychila TRUE TRUE FALSE

amblychila. Solenopsis aurea subsp. amblychila Wheeler, W.M. 1915b: 394 (w.q.m.) U.S.A. Subspecies of xyloni: Creighton, 1930b: 104; of aurea: Creighton, 1950a: 230. Junior synonym of aurea: Snelling, R.R. 1963: 7; Wheeler, G.C. & Wheeler, J. 1986g: 13. Revived from synonymy and raised to species: Trager, 1991: 168.

Myrmicinae Solenopsidini Solenopsis aurea TRUE TRUE FALSE

aurea. Solenopsis geminata var. aurea Wheeler, W.M. 1906d: 336 (w.q.m.) U.S.A. Crozier, 1970: 116 (k.). Raised to species: Forel, 1909a: 269. Subspecies of xyloni: Creighton, 1930b: 103. Revived status as species: Creighton, 1950a: 230; Snelling, R.R. 1963: 6. Senior synonym of huachucana: Trager, 1991: 170.

Myrmicinae Solenopsidini Solenopsis brazoensis FALSE FALSE FALSE

brazoensis. Atta brazoensis Buckley, 1867: 345 (s.w.) U.S.A. Junior synonym of geminata: Emery, 1895c: 276. Unidentifiable taxon, incertae sedis in Solenopsis: Bolton, 1995b: 386.

Myrmicinae Solenopsidini Solenopsis carolinensis TRUE TRUE FALSE

carolinensis. Solenopsis texana r. carolinensis Forel, 1901e: 345 (w.q.m.) U.S.A. Combination in S. (Diplorhoptrum): Creighton, 1950a: 236. Raised to species: Creighton, 1950a: 236. See also: Thompson & Johnson, 1989: 698.

Myrmicinae Solenopsidini Solenopsis krockowi TRUE TRUE FALSE

krockowi. Solenopsis krockowi Wheeler, W.M. 1908e: 428, pl. 26, figs. 28, 29 (w.q.) U.S.A.

Myrmicinae Solenopsidini Solenopsis madara FALSE FALSE FALSE

madara. Solenopsis madara Roger, 1863a: 200 (w.) U.S.A. Unidentifiable taxon, incertae sedis in Solenopsis: Smith, M.R. 1951a: 814.

Myrmicinae Solenopsidini Solenopsis molesta TRUE TRUE FALSE

molesta. Myrmica molesta Say, 1836: 293 (q.) U.S.A. Wheeler, G.C. & Wheeler, J. 1955c: 134 (l.); Petralia & Vinson, 1980: 383 (l.); Crozier, 1970: 116 (k.). Combination in Solenopsis: Mayr, 1863: 407; in S. (Diplorhoptrum): Creighton, 1950a: 237. Senior synonym of debilis, exigua, minuta: Emery, 1895c: 277; Emery, 1896g: 85; of validiuscula: Smith, D.R. 1979: 1387.

Myrmicinae Solenopsidini Solenopsis nickersoni TRUE TRUE FALSE

nickersoni. Solenopsis (Diplorhoptrum) nickersoni Thompson, 1982: 486, fig. 1 (w.) U.S.A. Thompson, 1989: 281 (q.).

Myrmicinae Solenopsidini Solenopsis pergandei TRUE TRUE FALSE

pergandei. Solenopsis pergandei Forel, 1901e: 343 (w.q.m.) U.S.A. Wheeler, G.C. & Wheeler, J. 1960b: 21 (l.). Combination in S. (Diplorhoptrum): Smith, M.R. 1947f: 568. See also: Thompson & Johnson, 1989: 698.

Myrmicinae Solenopsidini Solenopsis phoretica TRUE TRUE FALSE

phoretica. Solenopsis phoretica Davis & Deyrup, 2006: 141, fig. 1 (q.) U.S.A.

Myrmicinae Solenopsidini Solenopsis picta TRUE TRUE FALSE

picta. Solenopsis picta Emery, 1895c: 278 (w.) U.S.A. Wheeler, G.C. & Wheeler, J. 1960b: 22 (l.). Senior synonym of moerens: Creighton, 1950a: 237. See also: Thompson & Johnson, 1989: 697.

Myrmicinae Solenopsidini Solenopsis pilosula TRUE TRUE FALSE

pilosula. Solenopsis pilosula Wheeler, W.M. 1908e: 426, pl. 26, figs. 26, 27 (w.m.) U.S.A.

Myrmicinae Solenopsidini Solenopsis puncticeps TRUE TRUE FALSE

puncticeps. Solenopsis (Diplorhoptrum) puncticeps Mackay & Vinson, 1989a: 177, figs. 5-8 (w.) U.S.A.

Myrmicinae Solenopsidini Solenopsis sabeana FALSE FALSE FALSE

sabeana. Myrmica (Atta) sabeana Buckley, 1867: 343 (w.) U.S.A. Junior synonym of geminata: Emery, 1895c: 276. Synonymy with xyloni postulated but cannot be proved: Wheeler, W.M. 1915b: 395; Creighton, 1930b: 98; Smith, D.R. 1979: 1389; Trager, 1991: 167. Unidentifiable taxon, incertae sedis in Solenopsis: Bolton, 1995b: 390. 

Myrmicinae Solenopsidini Solenopsis salina TRUE TRUE FALSE

salina. Solenopsis salina Wheeler, W.M. 1908e: 427, pl. 26, figs. 24, 25 (w.) U.S.A.

Myrmicinae Solenopsidini Solenopsis subterranea TRUE TRUE FALSE

subterranea. Solenopsis (Diplorhoptrum) subterranea Mackay & Vinson, 1989a: 175, figs. 1-4 (w.) U.S.A.

Myrmicinae Solenopsidini Solenopsis texana TRUE TRUE FALSE

1 subspecies

texana. Solenopsis pollux var. texana Emery, 1895c: 278 (w.) U.S.A. Forel, 1901e: 345 (q.m.); Wheeler, G.C. & Wheeler, J. 1955c: 136 (l.). Combination in S. (Diplorhoptrum): Creighton, 1950a: 238. Raised to species: Forel, 1901e: 345. Senior synonym of rosella: Creighton, 1950a: 238. Current subspecies: nominal plus catalinae.

Myrmicinae Solenopsidini Solenopsis tonsa TRUE TRUE FALSE

tonsa. Solenopsis tonsa Thompson, 1989: 272, figs. 29-32 (w.q.m.) U.S.A.

Myrmicinae Solenopsidini Solenopsis truncorum TRUE TRUE FALSE

truncorum. Solenopsis texana r. truncorum Forel, 1901e: 346 (w.q.) U.S.A. Raised to species and senior synonym of castanea: Creighton, 1950a: 239. See also: Thompson & Johnson, 1989: 697.

Myrmicinae Solenopsidini Solenopsis vorax TRUE TRUE FALSE

vorax. Solenopsis (Diplorhoptrum) vorax Santschi, 1934e: 580, figs. 23, 29, 40 (q.m.) RUSSIA.

Myrmicinae Solenopsidini Solenopsis xyloni TRUE TRUE FALSE

xyloni. Solenopsis xyloni McCook, 1879: 188, figs. 37, 38 (s.w.q.) U.S.A. Wheeler, W.M. 1915b: 396 (m.); Wheeler, G.C. & Wheeler, J. 1955c: 133 (l.); Taber & Cokendolpher, 1988: 95 (k.). Junior synonym of geminata: Dalla Torre, 1893: 76. Revived from synonymy as subspecies of geminata: Wheeler, W.M. 1910g: 563; Wheeler, W.M. 1915b: 395. Revived status as species: Creighton, 1930b: 98; Creighton, 1950a: 232; Snelling, R.R. 1963: 9. Senior synonym of maniosa, pylades: Trager, 1991: 166.

Myrmicinae Solenopsidini Solenopsis texana catalinae TRUE TRUE FALSE

catalinae. Solenopsis texana subsp. catalinae Wheeler, W.M. 1904d: 269 (w.q) U.S.A.

Myrmicinae Solenopsidini Solenopsis globularia littoralis TRUE TRUE FALSE

littoralis. Solenopsis (Euophthalma) globularia subsp. littoralis Creighton, 1930b: 113, pl. 6, fig. 3 (w.q.m.) U.S.A. Wheeler, G.C. & Wheeler, J. 1960b: 22 (l.). Senior synonym of mobilensis: Smith, M.R. 1951a: 813.

Sphecomyrminae Sphecomyrmini Sphecomyrma freyi TRUE TRUE TRUE

†freyi. †Sphecomyrma freyi Wilson & Brown, in Wilson, Carpenter & Brown, 1967: 8, pls. 1-4, figs. 1, 2 (w.) U.S.A. (Cretaceous). Designation of neotype: Grimaldi, Agosti & Carpenter, 1997: 12. See also: Engel & Grimaldi, 2005: 9.

Sphecomyrminae Sphecomyrmini Sphecomyrma mesaki TRUE TRUE TRUE

†mesaki. †Sphecomyrma mesaki Engel & Grimaldi, 2005: 7, fig. 5 (w.) U.S.A. (Cretaceous).

Myrmicinae Stenammini Stenamma asiaticum FALSE FALSE FALSE

asiaticum. Stenamma westwoodi subsp. asiaticum Ruzsky, 1905b: 711 (w.) RUSSIA. Unidentifiable taxon, incertae sedis in Stenamma: DuBois, 1998b: 310.

Myrmicinae Stenammini Stenamma brevicorne TRUE TRUE FALSE

brevicorne. Aphaenogaster brevicornis Mayr, 1886d: 447 (w.q.) U.S.A. Emery, 1895c: 299 (m.); Smith, M.R. 1957b: 167 (m.); Hauschteck, 1962: 219 (k.). Combination in Stenamma: Emery, 1895c: 299. Senior synonym of neoarcticum: Smith, M.R. 1957b: 165.

Myrmicinae Stenammini Stenamma californicum TRUE TRUE FALSE

californicum. Stenamma californicum Snelling, R.R. 1973c: 16, figs. 16, 17, 24, 25, 27 (w.q.) U.S.A.

Myrmicinae Stenammini Stenamma chiricahua TRUE TRUE FALSE

chiricahua. Stenamma chiricahua Snelling, R.R. 1973c: 7, figs. 6-11 (w.q.m.) U.S.A.

Myrmicinae Stenammini Stenamma dyscheres TRUE TRUE FALSE

dyscheres. Stenamma dyscheres Snelling, R.R. 1973c: 21, figs. 28, 29, 34, 35 (w.q.m.) U.S.A.

Myrmicinae Stenammini Stenamma exasperatum TRUE TRUE FALSE

exasperatum. Stenamma exasperatum Snelling, R.R. 1973c: 28, figs. 40, 41, 46, 47, 63 (w.) U.S.A.

Myrmicinae Stenammini Stenamma foveolocephalum TRUE TRUE FALSE

foveolocephalum. Stenamma fovolocephala Smith, M.R. 1930b: 564 (w.) U.S.A. Spelling emended to foveolocephalum and senior synonym of carolinense: DuBois & Davis, 1998: 132.

Myrmicinae Stenammini Stenamma georgii TRUE TRUE FALSE

georgii. Stenamma georgii Arnol'di, 1975: 1823 (page number misprinted as 1283 in text), fig. 3 (w.q.) RUSSIA. See also: DuBois, 1998b: 269.

Myrmicinae Stenammini Stenamma heathi TRUE TRUE FALSE

heathi. Stenamma brevicorne subsp. heathi Wheeler, W.M. 1915b: 410 (w.) U.S.A. Snelling, R.R. 1973c: 31 (q.). Raised to species: Creighton, 1950a: 137. See also: Smith, M.R. 1957b: 154.

Myrmicinae Stenammini Stenamma hissarianum TRUE TRUE FALSE

hissarianum. Stenamma hissarianum Arnol'di, 1975: 1824, fig. 5 (w.q.m.) RUSSIA. See also: DuBois, 1998b: 273.

Myrmicinae Stenammini Stenamma huachucanum TRUE TRUE FALSE

huachucanum. Stenamma huachucanum Smith, M.R. 1957b: 153, pl. 2, fig. 8 (w.) U.S.A.

Myrmicinae Stenammini Stenamma impar TRUE TRUE FALSE

impar. Stenamma brevicorne r. impar Forel, 1901e: 347 (w.q.) U.S.A. Smith, M.R. 1957b: 152 (m.). Raised to species: Creighton, 1950a: 137.

Myrmicinae Stenammini Stenamma kurilense TRUE TRUE FALSE

kurilense. Stenamma kurilense Arnol'di, 1975: 1825, fig. 2 (w.) RUSSIA. See also: DuBois, 1998b: 296.

Myrmicinae Stenammini Stenamma lippulum TRUE TRUE FALSE

lippulum. Myrmica lippula Nylander, 1849: 41 (w.) RUSSIA. Smith, F. 1858a: 280 (q.); DuBois, 1993: 322 (w.q.m.). Combination in Tetramorium: Roger, 1859: 258; in Asemorhoptrum: Mayr, 1861: 76; in Stenamma: Forel, 1889: 258. Junior synonym of westwoodii: Forel, 1889: 258. Revived from synonymy and senior synonym of caucasicum, hirtula: DuBois, 1993: 321. See also: DuBois, 1998b: 265. See Radchenko, 2007: 28 for doubts about type-material.

Myrmicinae Stenammini Stenamma meridionale TRUE TRUE FALSE

meridionale. Stenamma meridionale Smith, M.R. 1957b: 169, pl. 4, fig. 15 (w.q.) U.S.A.

Myrmicinae Stenammini Stenamma punctatoventre TRUE TRUE FALSE

punctatoventre. Stenamma punctatoventre Snelling, R.R. 1973c: 10, figs. 12, 13, 20, 21, 55 (w.q.m.) U.S.A.

Myrmicinae Stenammini Stenamma schmittii TRUE TRUE FALSE

schmittii. Stenamma brevicorne subsp. schmittii Wheeler, W.M. 1903d: 167 (diagnosis in key) (w.) U.S.A. Smith, M.R. 1957b: 144 (q.m.). Raised to species: Creighton, 1950a: 138.

Myrmicinae Stenammini Stenamma sequoiarum TRUE TRUE FALSE

sequoiarum. Stenamma brevicorne subsp. sequoiarum Wheeler, W.M. 1917a: 520 (w.q.) U.S.A. Subspecies of diecki: Creighton, 1950a: 137. Raised to species: Smith, M.R. 1957b: 156.

Myrmicinae Stenammini Stenamma smithi TRUE TRUE FALSE

smithi. Stenamma smithi Cole, 1966: 7, fig. (w.) U.S.A. Senior synonym of knowltoni: Snelling, R.R. 1973c: 12.

Myrmicinae Stenammini Stenamma ussuriense TRUE TRUE FALSE

ussuriense. Stenamma ussuriense Arnol'di, 1975: 1825, figs. 2, 6 (w.q.m.) RUSSIA. See also: Kupyanskaya, 1990: 117; DuBois, 1998b: 299.

Myrmicinae Stenammini Stenamma wheelerorum TRUE TRUE FALSE

wheelerorum. Stenamma wheelerorum Snelling, R.R. 1973c: 34, figs. 50-54 (w.q.) U.S.A.

Myrmicinae Tetramoriini Strongylognathus arnoldii TRUE TRUE FALSE

arnoldii. Strongylognathus arnoldii Radchenko, 1985a: 1514, fig. 1 (w.q.m.) RUSSIA.

Myrmicinae Tetramoriini Strongylognathus chelifer TRUE TRUE FALSE

chelifer. Strongylognathus cheliferus Radchenko, 1985a: 1516, fig. 2 (q.m.) RUSSIA.

Myrmicinae Tetramoriini Strongylognathus christophi TRUE TRUE FALSE

christophi. Strongylognathus christophi Emery, 1889a: 439 (q.) RUSSIA. Forel, 1900e: 280 (w.); Radchenko, 1985a: 1519 (m.). Subspecies of huberi: Emery, 1898c: 145; Emery, 1909d: 709; Wheeler, W.M. 1909g: 186; Emery, 1924d: 285. Status as species: Forel, 1904c: 1; Ruzsky, 1905b: 545; Pisarski, 1966: 510; Bolton, 1976: 305; Tarbinsky, 1976: 116; Radchenko, 1985a: 1519. Senior synonym of ruzskyi: Radchenko, 1991: 88.

Myrmicinae Dacetini Strumigenys infidelis TRUE TRUE FALSE

infidelis. Strumigenys infidelis Santschi, 1919f: 48 (w.) ARGENTINA. Junior synonym of louisianae: Brown, 1953g: 28; revived from synonymy: Bolton, 2000: 523.

Myrmicinae Dacetini Strumigenys louisianae TRUE TRUE FALSE

louisianae. Strumigenys louisianae Roger, 1863a: 211 (w.) U.S.A. Emery, 1895c: 326 (q.); Haug, 1932: 170 (m.); Wheeler, G.C. & Wheeler, J. 1955a: 136 (l.). Senior synonym of unispinulosa: Emery, 1895c: 326; Brown, 1953g: 28; of laticephala: Smith, M.R. 1951a: 826; Brown, 1953g: 28; of bruchi, costaricensis, cubaensis, guatemalensis, longicornis, obscuriventris, soledadensis: Brown, 1953g: 28; of clasmospongia, fusca, unidentata: Brown, 1961a: 64; of producta: Bolton, 2000: 524; of wani: Bolton, Sosa-Calvo, et al. 2008: 62. See also: Wilson, 1954: 481; Bolton, 2000: 525.

Dolichoderinae Tapinomini Tapinoma schreiberi TRUE TRUE FALSE

schreiberi. Tapinoma schreiberi Hamm, 2010: 25, figs. 5-8 (w.q.m.) U.S.A.

Dolichoderinae Tapinomini Tapinoma sessile TRUE TRUE FALSE

sessile. Formica sessilis Say, 1836: 287 (w.q.) U.S.A. Emery, 1895c: 333 (q.m.); Wheeler, G.C. & Wheeler, J. 1951: 196 (l.); Crozier, 1970: 119 (k.); Taber & Cokendolpher, 1988: 95 (k.). Combination in Tapinoma: Smith, F. 1858b: 57. Senior synonym of parva, boreale Roger (and its junior synonym boreale Provancher): Mayr, 1886d: 434; Creighton, 1950a: 353; of gracilis: Emery, 1895c: 337; Wheeler, W.M. 1902f: 20; Creighton, 1950a: 353; of dimmocki: Shattuck, 1992c: 153. See also: Smith, M.R. 1928a: 307; Smith, D.R. 1979: 1422; Kupyanskaya, 1990: 156; Shattuck, 1994: 153; Hamm, 2010: 24.

Myrmicinae Formicoxenini Temnothorax alinae TRUE TRUE FALSE

alinae. Leptothorax alinae Radchenko, 1994i: 31, fig. 6 (w.) RUSSIA. Combination in Temnothorax: Bolton, 2003: 271. See also: Radchenko, 1994d: 153.

Myrmicinae Formicoxenini Temnothorax alpinus TRUE TRUE FALSE

alpinus. Leptothorax alpinus Ruzsky, 1902d: 22 (w.) RUSSIA. Combination in Temnothorax: Bolton, 2003: 271. Subspecies of tuberum: Ruzsky, 1905b: 598. Revived status as species: Radchenko, 1994d: 150 (in key); Radchenko, 1996c: 17.

Myrmicinae Formicoxenini Temnothorax ambiguus TRUE TRUE FALSE

2 subspecies

ambiguus. Leptothorax (Leptothorax) curvispinosus subsp. ambiguus Emery, 1895c: 320 (w.) U.S.A. Wesson, L.G. & Wesson, R.G. 1940: 97 (q.m.); Wheeler, G.C. & Wheeler, J. 1955b: 22 (l.). Combination in L. (Myrafant): Smith, D.R. 1979: 1392; in Temnothorax: Bolton, 2003: 271. Raised to species: Wesson, L.G. & Wesson, R.G. 1940: 97. See also: Mackay, 2000: 312. Current subspecies: nominal plus foveatus, pinetorum.

Myrmicinae Formicoxenini Temnothorax andersoni TRUE TRUE FALSE

andersoni. Leptothorax (Myrafant) andersoni Mackay, 2000: 314, figs. 14, 83 (w.) U.S.A. Combination in Temnothorax: Bolton, 2003: 271.

Myrmicinae Formicoxenini Temnothorax andrei TRUE TRUE FALSE

andrei. Leptothorax (Leptothorax) andrei Emery, 1895c: 322, pl. 8, fig. 15 (w.) U.S.A. Cole, 1958c: 537 (q.m.). Combination in L. (Myrafant): Smith, D.R. 1979: 1392; in Temnothorax: Bolton, 2003: 271. Senior synonym of heathii, occidentalis, ocellatus: Ward, 2005: 15. See also: Wheeler, W.M. 1903c: 256; Mackay, 2000: 316.

Myrmicinae Formicoxenini Temnothorax bestelmeyeri TRUE TRUE FALSE

bestelmeyeri. Leptothorax (Myrafant) bestelmeyeri Mackay, 2000: 318, figs. 26, 41, 42, 87-89 (w.) U.S.A. Combination in Temnothorax: Bolton, 2003: 271.

Myrmicinae Formicoxenini Temnothorax bradleyi TRUE TRUE FALSE

bradleyi. Leptothorax bradleyi Wheeler, W.M. 1913c: 113 (w.) U.S.A. Wilson, 1952a: 69 (q.m.). Combination in L. (Myrafant): Smith, D.R. 1979: 1392; in Temnothorax: Bolton, 2003: 271. See also: Mackay, 2000: 321.

Myrmicinae Formicoxenini Temnothorax bristoli TRUE TRUE FALSE

bristoli. Leptothorax (Myrafant) bristoli Mackay, 2000: 326, figs. 28, 96 (w.) U.S.A. Combination in Temnothorax: Bolton, 2003: 271.

Myrmicinae Formicoxenini Temnothorax carinatus TRUE TRUE FALSE

carinatus. Leptothorax (Leptothorax) carinatus Cole, 1957d: 213, fig. 1 (w.q.) U.S.A. Wheeler, G.C. & Wheeler, J. 1973b: 70 (l.). Combination in L. (Myrafant): Smith, D.R. 1979: 1392; in Temnothorax: Bolton, 2003: 271. See also: Mackay, 2000: 328.

Myrmicinae Formicoxenini Temnothorax chandleri TRUE TRUE FALSE

chandleri. Leptothorax (Myrafant) chandleri Mackay, 2000: 330, figs. 35, 99 (w.) U.S.A. Combination in Temnothorax: Bolton, 2003: 271.

Myrmicinae Formicoxenini Temnothorax cokendolpheri TRUE TRUE FALSE

cokendolpheri. Leptothorax (Myrafant) cokendolpheri Mackay, 2000: 332, figs. 30, 100, 101 (w.q.) U.S.A. Combination in Temnothorax: Bolton, 2003: 271.

Myrmicinae Formicoxenini Temnothorax coleenae TRUE TRUE FALSE

coleenae. Leptothorax (Myrafant) coleenae Mackay, 2000: 335, figs. 37, 102 (w.) U.S.A. Combination in Temnothorax: Bolton, 2003: 271.

Myrmicinae Formicoxenini Temnothorax curvispinosus TRUE TRUE FALSE

curvispinosus. Leptothorax curvispinosus Mayr, 1866a: 508, pl., fig. 13 (w.) U.S.A. Mayr, 1886d: 451 (q.); Mackay, 2000: 338 (m.). Combination in L. (Myrafant): Smith, M.R. 1950: 30; in Temnothorax: Bolton, 2003: 271. Senior synonym of gallarum: Mayr, 1886d: 453. See also: Wheeler, W.M. 1903c: 239; Mackay, 2000: 337.

Myrmicinae Formicoxenini Temnothorax desertorum TRUE TRUE FALSE

desertorum. Temnothorax desertorum Dlussky & Soyunov, 1988: 35, figs. 1, 2 (w.q.) RUSSIA. Combination in Leptothorax: Bolton, 1995b: 238; in Temnothorax: Bolton, 2003: 271.

Myrmicinae Formicoxenini Temnothorax discoloratus TRUE TRUE FALSE

discoloratus. Leptothorax discoloratus Arnol'di, 1977a: 200 (w.) RUSSIA. Combination in Temnothorax: Bolton, 2003: 271.

Myrmicinae Formicoxenini Temnothorax duloticus TRUE TRUE FALSE

duloticus. Leptothorax (Mychothorax) duloticus Wesson, L.G. 1937: 125, fig. 1 (w.q.) U.S.A. Wesson, L.G. 1940: 81 (m.). Combination in Temnothorax: Bolton, 2003: 271. See also: Wilson, 1975: 108.

Myrmicinae Formicoxenini Temnothorax emmae TRUE TRUE FALSE

emmae. Leptothorax (Myrafant) emmae Mackay, 2000: 342, figs. 67, 107, 108 (w.q.) U.S.A. Combination in Temnothorax: Bolton, 2003: 271.

Myrmicinae Formicoxenini Temnothorax furunculus TRUE TRUE FALSE

furunculus. Leptothorax furunculus Wheeler, W.M. 1909e: 82 (w.) U.S.A. Combination in L. (Myrafant): Smith, D.R. 1979: 1393; in Temnothorax: Bolton, 2003: 271. See also: Mackay, 2000: 344.

Myrmicinae Formicoxenini Temnothorax gallae TRUE TRUE FALSE

gallae. Leptothorax (Leptothorax) gallae Smith, M.R. 1949e: 112 (w.) U.S.A. Combination in L. (Myrafant): Smith, D.R. 1979: 1393; in Temnothorax: Bolton, 2003: 271.See also: Mackay, 2000: 346.

Myrmicinae Formicoxenini Temnothorax hispidus TRUE TRUE FALSE

hispidus. Leptothorax (Leptothorax) hispidus Cole, 1957b: 42, fig. 1 (w.q.) U.S.A. Wheeler, G.C. & Wheeler, J. 1973b: 71 (l.). Combination in L. (Myrafant): Smith, D.R. 1979: 1393; in Temnothorax: Bolton, 2003: 271.See also: Mackay, 2000: 347.

Myrmicinae Formicoxenini Temnothorax josephi TRUE TRUE FALSE

josephi. Leptothorax (Myrafant) josephi Mackay, 2000: 349, figs. 78, 115-120 (w.q.m.) U.S.A. Combination in Temnothorax: Bolton, 2003: 271.

Myrmicinae Formicoxenini Temnothorax knipovitshi TRUE TRUE FALSE

knipovitshi. Leptothorax tuberum subsp. knipovitshi Karavaiev, 1916: 499 (w.q.) RUSSIA. Combination in Temnothorax: Bolton, 2003: 271. Raised to species and senior synonym of junipereti, nikitae: Radchenko, 1994d: 156 (in key); Radchenko, 1995c: 17.

Myrmicinae Formicoxenini Temnothorax kurilensis TRUE TRUE FALSE

kurilensis. Leptothorax kurilensis Radchenko, 1994i: 32, fig. 7 (w.q.) RUSSIA (Kunashir I.). Combination in Temnothorax: Bolton, 2003: 271. See also: Radchenko, 1994d: 148.

Myrmicinae Formicoxenini Temnothorax liebi TRUE TRUE FALSE

liebi. Leptothorax (Myrafant) liebi Mackay, 2000: 353, figs. 16, 36, 42, 121, 122 (w.) U.S.A. Combination in Temnothorax: Bolton, 2003: 271.

Myrmicinae Formicoxenini Temnothorax longispinosus TRUE TRUE FALSE

longispinosus. Leptothorax longispinosus Roger, 1863a: 180 (w.) U.S.A. Emery, 1895c: 321 (q.); Wheeler, W.M. 1903c: 238 (m.); Crozier, 1970: 117 (k.); Wheeler, G.C. & Wheeler, J. 1989a: 323 (l.). Combination in L. (Myrafant): Smith, M.R. 1950: 30; in Temnothorax: Bolton, 2003: 271.Senior synonym of iowensis: Creighton, 1950a: 264. See also: Mackay, 2000: 358.

Myrmicinae Formicoxenini Temnothorax minutissimus TRUE TRUE FALSE

minutissimus. Leptothorax minutissimus Smith, M.R. 1942a: 59, pl. 6 (q.) U.S.A. Buschinger & Linksvayer, 2004: 71 (m.). Combination in L. (Myrafant): Smith, D.R. 1979: 1393; in Temnothorax: Bolton, 2003: 271.  See also: Mackay, 2000: 373.

Myrmicinae Formicoxenini Temnothorax nassanowi TRUE TRUE FALSE

nassanowi. Leptothorax nassanowi Ruzsky, 1895: 26 (w.) RUSSIA. [Also described as new by Ruzsky, 1896: 72.] Kupyanskaya, 1990: 143 (q.m.). Combination in Temnothorax: Bolton, 2003: 271. Senior synonym of firssovi: Kupyanskaya, 1990: 143; Radchenko, 1995d: 9; of subnudus: Radchenko, 1995d: 9; Radchenko, 2005b: 151; of galeatus: Radchenko, 2004: 126. See also: Tarbinsky, 1976: 94. [This name is frequently misspelled as nassonovi, for example Ruzsky, 1905b: 582; Emery, 1924d: 255; Radchenko, 1994d: 155.]

Myrmicinae Formicoxenini Temnothorax neomexicanus TRUE TRUE FALSE

neomexicanus. Leptothorax neomexicanus Wheeler, W.M. 1903c: 248, pl. 12, fig. 18 (w.) U.S.A. Combination in Temnothorax: Bolton, 2003: 271. Subspecies of tricarinatus: Creighton, 1950a: 273; Smith, M.R. 1952b: 100. Revived status as species: Mackay, 2000: 374.

Myrmicinae Formicoxenini Temnothorax nevadensis TRUE TRUE FALSE

nevadensis. Leptothorax nevadensis Wheeler, W.M. 1903c: 252, pl. 12, fig. 20 (w.q.m.) U.S.A. Wheeler, G.C. & Wheeler, J. 1973b: 71 (l.). Combination in L. (Myrafant): Smith, D.R. 1979: 1393; in Temnothorax: Bolton, 2003: 271. Senior synonym of eldoradensis, melanderi: Mackay, 2000: 376; of lindae, maryanae: Ward, 2005: 16.

Myrmicinae Formicoxenini Temnothorax nitens TRUE TRUE FALSE

nitens. Leptothorax (Leptothorax) nitens Emery, 1895c: 322, pl. 8, fig. 16 (w.) U.S.A. Cole, 1958c: 536 (q.m.); Wheeler, G.C. & Wheeler, J. 1973b: 71 (l.). Combination in L. (Myrafant): Smith, D.R. 1979: 1394; in Temnothorax: Bolton, 2003: 271. Senior synonym of mariposa, melinus: Ward, 2005: 17. See also: Wheeler, W.M. 1903c: 244; Mackay, 2000: 379.

Myrmicinae Formicoxenini Temnothorax obliquicanthus TRUE TRUE FALSE

obliquicanthus. Leptothorax (Myrafant) obliquicanthus Cole, 1953c: 28, figs. 1, 2 (w.) U.S.A. See also: Mackay, 2000: 380.

Myrmicinae Formicoxenini Temnothorax obturator TRUE TRUE FALSE

obturator. Leptothorax obturator Wheeler, W.M. 1903c: 249, pl. 12, fig. 19 (w.q.m.) U.S.A. Wheeler, W.M. 1903f: 662 (gynandromorph); Wheeler, G.C. & Wheeler, J. 1955b: 24 (l.). Combination in L. (Myrafant): Smith, D.R. 1979: 1394; in Temnothorax: Bolton, 2003: 271.See also: Mackay, 2000: 382.

Myrmicinae Formicoxenini Temnothorax oxynodis TRUE TRUE FALSE

oxynodis. Leptothorax (Myrafant) oxynodis Mackay, 2000: 385, figs. 63, 146 (w.) U.S.A. Combination in Temnothorax: Bolton, 2003: 272.

Myrmicinae Formicoxenini Temnothorax palustris TRUE TRUE FALSE

palustris. Leptothorax palustris Deyrup & Cover, 2004a: 52, figs. 1, 2 (w.q.m.) U.S.A. Combination in Temnothorax: new combination (unpublished).

Myrmicinae Formicoxenini Temnothorax pamiricus TRUE TRUE FALSE

pamiricus. Leptothorax pamiricus Ruzsky, 1902b: 478 (w.) RUSSIA. Combination in Temnothorax: Bolton, 2003: 271. Menozzi, 1939a: 303 (q.m.). Subspecies of bulgaricus: Forel, 1904b: 375; Kuznetsov-Ugamsky, 1926a: 75; Menozzi, 1939a: 303. Revived status as species: Radchenko, 1994d: 152 (in key); Radchenko, 1995a: 26.

Myrmicinae Formicoxenini Temnothorax pergandei TRUE TRUE FALSE

pergandei. Leptothorax (Dichothorax) pergandei Emery, 1895c: 323, pl. 8, fig. 13 (w.) U.S.A. Wheeler, W.M. 1903c: 257 (q.m.); Wheeler, G.C. & Wheeler, J. 1960b: 23 (l.). Combination in Temnothorax: Bolton, 2003: 272. Senior synonym of manni: Wesson, L.G. 1939: 180; of floridanus (and its junior synonym flavus): Mackay, 1993a: 289.

Myrmicinae Formicoxenini Temnothorax politus TRUE TRUE FALSE

politus. Macromischa polita Smith, M.R. 1939e: 506, pl. 1, fig. B (w.) U.S.A. Combination in Leptothorax: Baroni Urbani, 1978b: 478; in Temnothorax: Bolton, 2003: 272.

Myrmicinae Formicoxenini Temnothorax rudis TRUE TRUE FALSE

rudis. Leptothorax nevadensis subsp. rudis Wheeler, W.M. 1917a: 508 (w.q.) U.S.A. Combination in Temnothorax: Ward, 2005: 19. Junior synonym of nevadensis: Mackay, 2000: 376. Revived from synomymy and raised to species: Ward, 2005: 19.

Myrmicinae Formicoxenini Temnothorax rugatulus TRUE TRUE FALSE

rugatulus. Leptothorax (Leptothorax) rugatulus Emery, 1895c: 321 (w.) U.S.A. Taber & Cokendolpher, 1988: 95 (k.). Combination in L. (Myrafant): Smith, M.R. 1950: 30; in Temnothorax: Bolton, 2003: 272.  Subspecies of curvispinosus: Wheeler, W.M. 1903c: 241. Revived status as species: Wheeler, W.M. 1917a: 509. Senior synonym of annectens, cockerelli, mediorufus: Creighton, 1950a: 269; of brunnescens: Mackay, 2000: 394.

Myrmicinae Formicoxenini Temnothorax schaumii TRUE TRUE FALSE

schaumii. Leptothorax schaumii Roger, 1863a: 180 (w.) U.S.A. Mayr, 1886d: 451 (m.); Wesson L.G. & Wesson, R.G. 1940: 95 (q.); Wheeler, G.C. & Wheeler, J. 1960b: 23 (l.). Combination in L. (Myrafant): Smith, D.R. 1979: 1394; in Temnothorax: Bolton, 2003: 272. Senior synonym of fortinodis, gilvus, melanoticus: Creighton, 1950a: 271. See also: Wheeler, W.M. 1903c: 232; Mackay, 2000: 403.

Myrmicinae Formicoxenini Temnothorax schmittii TRUE TRUE FALSE

schmittii. Leptothorax schmittii Wheeler, W.M. 1903c: 242, pl. 12, fig. 14 (w.) U.S.A. Combination in L. (Myrafant): Smith, D.R. 1979: 1395; Combination in Temnothorax: Bolton, 2003: 272. See also: Mackay, 2000: 405.

Myrmicinae Formicoxenini Temnothorax serviculus TRUE TRUE FALSE

serviculus. Leptothorax serviculus Ruzsky, 1902b: 476 (w.) RUSSIA. Ruzsky, 1905b: 768 (q.); Radchenko, 1996c: 16 (m.). Combination in Temnothorax: Bolton, 2003: 271. See also: Kupyanskaya, 1990: 140.

Myrmicinae Formicoxenini Temnothorax silvestrii TRUE TRUE FALSE

silvestrii. Tetramorium silvestrii Santschi, 1911d: 6 (w.) U.S.A. Creighton, 1953f: 5 (q.m.). Combination in Leptothorax: Santschi, 1922a: 68; in L. (Myrafant): Smith, D.R. 1979: 1395; in Temnothorax: Bolton, 2003: 272.See also: Mackay, 2000: 407.

Myrmicinae Formicoxenini Temnothorax subditivus TRUE TRUE FALSE

subditivus. Macromischa subditiva Wheeler, W.M. 1903b: 99, fig. 5 (w.) U.S.A. Wheeler, W.M. 1931b: 15 (q.); Smith, M.R. 1939e: 504 (m.). Combination in Leptothorax: Baroni Urbani, 1978b: 512; in Temnothorax: Bolton, 2003: 272. Senior synonym of flavitarsis, laevissima, luciliae: Baroni Urbani, 1978b: 512. See also: Creighton, 1966a: 282.

Myrmicinae Formicoxenini Temnothorax terrigena TRUE TRUE FALSE

terrigena. Leptothorax terrigena Wheeler, W.M. 1903c: 254, pl. 12, fig. 21 (w.q.) U.S.A. Combination in L. (Myrafant): Smith, D.R. 1979: 1395; in Temnothorax: Bolton, 2003: 272. See also: Mackay, 2000: 414.

Myrmicinae Formicoxenini Temnothorax texanus TRUE TRUE FALSE

texanus. Leptothorax texanus Wheeler, W.M. 1903c: 245, pl. 12, fig. 16 (w.q.m.) U.S.A. Wheeler, G.C. & Wheeler, J. 1960b: 24 (l.). Combination in L. (Myrafant): Smith, M.R. 1950a: 30; in Temnothorax: Bolton, 2003: 272. Senior synonym of davisi: Deyrup & Cover, 2004a: 56. See also: Smith, M.R. 1952b: 102; Mackay, 2000: 415.

Myrmicinae Formicoxenini Temnothorax tricarinatus TRUE TRUE FALSE

tricarinatus. Leptothorax (Leptothorax) tricarinatus Emery, 1895c: 321, pl. 8, fig. 14 (w.) U.S.A. Smith, M.R. 1952b: 99 (m.). Combination in L. (Myrafant): Smith, M.R. 1950: 30; in Temnothorax: Bolton, 2003: 272. See also: Wheeler, W.M. 1903c: 247; Mackay, 2000: 417.

Myrmicinae Formicoxenini Temnothorax tuscaloosae TRUE TRUE FALSE

tuscaloosae. Leptothorax tuscaloosae Wilson, 1951: 128 (w.q.) U.S.A. Combination in Temnothorax: Bolton, 2003: 272. See also: Mackay, 2000: 419.

Myrmicinae Formicoxenini Temnothorax usunkul TRUE TRUE FALSE

usunkul. Leptothorax usunkul Ruzsky, 1924: 152 (w.) RUSSIA. Combination in Temnothorax: Bolton, 2003: 271.

Myrmicinae Formicoxenini Temnothorax volgensis TRUE TRUE FALSE

volgensis. Leptothorax nassonovi subsp. volgensis Ruzsky, 1905b: 582 (w.) RUSSIA. Kupyanskaya, 1990: 142 (q.). Combination in Temnothorax: Bolton, 2003: 271. Raised to species: Arnol'di, 1977a: 203.

Myrmicinae Formicoxenini Temnothorax whitfordi TRUE TRUE FALSE

whitfordi. Leptothorax (Myrafant) whitfordi Mackay, 2000: 421, figs. 65, 67, 188-190 (w.q.m.) U.S.A. Combination in Temnothorax: Bolton, 2003: 272.

Myrmicinae Formicoxenini Temnothorax ambiguus pinetorum TRUE TRUE FALSE

pinetorum. Leptothorax ambiguus var. pinetorum Wesson L.G. & Wesson, R.G. 1940: 97 (w.q.m.) U.S.A. Combination in L. (Myrafant): Smith, D.R. 1979: 1392. Currently subspecies of ambiguus: Creighton, 1950a: 263.

Myrmicinae Formicoxenini Temnothorax ambiguus foveatus TRUE TRUE FALSE

foveatus. Leptothorax foveata Smith, M.R. 1934b: 211 (w.) U.S.A. Combination in L. (Myrafant): Smith, D.R. 1979: 1392. Currently subspecies of ambiguus: Creighton, 1950a: 263.

Myrmicinae Tetramoriini Tetramorium bicarinatum TRUE TRUE FALSE

bicarinatum. Myrmica bicarinata Nylander, 1846b: 1061 (w.q.) U.S.A. Forel, 1891b: 151 (m., misidentified as guineense); Wheeler, G.C. & Wheeler, J. 1954d: 449 (l., misidentified as guineense); Wheeler, W.M. 1924c: 136 (gynandromorph, misidentified as guineense). Combination in Tetramorium: Mayr, 1862: 740; Bolton, 1977: 94. Junior synonym of Formica guineensis Fabricius, 1793: 357 (now in Pheidole): Mayr, 1862: 740. Revived from synonymy and senior synonym of cariniceps (and its junior synonym kollari), modesta Smith, reticulata: Bolton, 1977: 94. [Note. The names cariniceps, kollari and reticulata had previously been incorrectly synonymised with guineensis Fabricius by Roger, 1862c: 293; modesta Smith was wrongly synonymised with guineensis by Donisthorpe, 1932c: 463.] See also: Bolton, 1979: 164.

Myrmicinae Tetramoriini Tetramorium ferox TRUE TRUE FALSE

ferox. Tetramorium caespitum var. ferox Ruzsky, 1903b: 309 (w.q.m.) RUSSIA. Subspecies of caespitum: Emery, 1909d: 703; Emery, 1916b: 194. Raised to species: Cori & Finzi, 1931: 239; Soudek, 1931: 10; Menozzi, 1933b: 75 (in key); Kratochvil, 1944: 72; Tarbinsky, 1976: 112; Atanassov & Dlussky, 1992: 155. Senior synonym of silhavyi: Bernard, 1967: 235; Dlussky, Soyunov & Zabelin, 1990: 201; Radchenko, 1992b: 55; of confinis, perspicax: Csösz & Schulz, 2010: 20; material of the unavailable name striaticeps referred here by Radchenko, 1992b: 55.

Myrmicinae Tetramoriini Tetramorium hispidum TRUE TRUE FALSE

hispidum. Xiphomyrmex spinosus subsp. hispidus Wheeler, W.M. 1915b: 415 (w.) U.S.A. Combination in Tetramorium: Bolton, 1979: 161. Raised to species: Bolton, 1979: 161.

Myrmicinae Tetramoriini Tetramorium pacificum TRUE TRUE FALSE

pacificum. Tetramorium pacificum Mayr, 1870b: 976 (w.q.) TONGA. Imai, Kubota, et al. 1985: 47 (k.). Senior synonym of subscabrum: Bolton, 1977: 102. See also: Schlick-Steiner, Steiner & Zettel, 2006: 182.

Myrmicinae Tetramoriini Tetramorium caespitum barabense TRUE TRUE FALSE

barabense. Tetramorium caespitum var. barabensis Ruzsky, 1925a: 288 (w.) RUSSIA.

Formiciinae Formiciini Titanomyrma lubei TRUE TRUE TRUE

†lubei. †Titanomyrma lubei Archibald, et al. 2011: 2, figs. 2, 3a,b (q.) U.S.A (Eocene).

Myrmicinae Attini Trachymyrmex arizonensis TRUE TRUE FALSE

arizonensis. Atta (Trachymyrmex) arizonensis Wheeler, W.M. 1907c: 710, pl. 49, figs. 9, 10 (q.m.) U.S.A. Wheeler, W.M. 1911e: 93 (w.). Combination in Trachymyrmex: Gallardo, 1916b: 242. See also: Rabeling, Cover, et al. 2007: 7.

Myrmicinae Attini Trachymyrmex desertorum TRUE TRUE FALSE

desertorum. Atta (Trachymyrmex) desertorum Wheeler, W.M. 1911e: 98, fig. 2 (w.) U.S.A. Rabeling, Cover, et al. 2007: 10 (q.m.). Combination in Trachymyrmex: Creighton, 1950a: 321.

Myrmicinae Attini Trachymyrmex nogalensis TRUE TRUE FALSE

nogalensis. Trachymyrmex nogalensis Byars, 1951: 109, figs. 1, 2 (w.) U.S.A. Rabeling, Cover, et al. 2007: 13 (q.).

Myrmicinae Attini Trachymyrmex pomonae TRUE TRUE FALSE

pomonae. Trachymyrmex pomonae Rabeling & Cover, in Rabeling, Cover, et al. 2007: 14, figs. 12-14 (w.q.m.) U.S.A.

Myrmicinae Attini Trachymyrmex septentrionalis TRUE TRUE FALSE

septentrionalis. Atta septentrionalis McCook, 1881a: 362 (w.) U.S.A. Wheeler, W.M. 1907c: 707 (q.m.); Wheeler, G.C. 1949: 673 (l.). Combination in Atta (Trachymyrmex): Forel, 1912e: 182; in Trachymyrmex: Gallardo, 1916b: 242. Senior synonym of virginiana: Wheeler, W.M. 1902f: 29; of vertebrata: Creighton, 1950a: 323; of obscurior, seminole: Weber, 1958b: 53. See also: Wheeler, W.M. 1911g: 246; Smith, D.R. 1979: 1411; Petralia & Vinson, 1980: 383; Rabeling, Cover, et al. 2007: 17.

Myrmicinae Attini Trachymyrmex turrifex TRUE TRUE FALSE

turrifex. Atta (Trachymyrmex) turrifex Wheeler, W.M. 1903b: 100, fig. 6 (w.q.) U.S.A. Combination in Trachymyrmex: Gallardo, 1916b: 242. Senior synonym of caroli: Rabeling, Cover, et al. 2007: 23.

Myrmicinae Formicoxenini Xenomyrmex floridanus TRUE TRUE FALSE

1 subspecies

floridanus. Xenomyrmex stolli subsp. floridanus Emery, 1895c: 275 (w.m.) U.S.A. Wheeler, W.M. 1931a: 136 (q.). Raised to species: Creighton, 1957c: 6. Senior synonym of rufescens: Creighton, 1950a: 226; of cubanus, lucayanus: Creighton, 1957c: 6. Current subspecies: nominal plus skwarrae.

Dolichoderinae Zherichiniini Zherichinius horribilis TRUE TRUE TRUE

†horribilis. †Zherichinius horribilis Dlussky, 1988: 57, figs. 4, 5 (w.) RUSSIA (Paleocene).

Dolichoderinae Zherichiniini Zherichinius rapax TRUE TRUE TRUE

†rapax. †Zherichinius rapax Dlussky, 1988: 57, figs. 4, 5 (w.) RUSSIA (Paleocene).