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Species: Lasius (Chthonolasius) mixtus   (Nylander, 1846) 

Classification:
Download Data

Taxonomic History (provided by Barry Bolton, 2020)

Formica mixta Nylander, 1846b PDF: 1050 (w.q.m.) FINLAND. Palearctic. AntCat AntWiki HOL

Taxonomic history

Combination in Lasius: Mayr, 1861 PDF: 50 (in key).
Combination in Donisthorpea: Donisthorpe, 1915f: 236.
Combination in Lasius (Lasius): Forel, 1915d: 52.
Combination in Formicina: Emery, 1916b PDF: 63; Bondroit, 1918 PDF: 32.
Combination in Formicina (Formicina): Emery, 1916a PDF: 242.
Combination in Acanthomyops: Ruzsky, 1925b PDF: 45.
Junior synonym of Lasius umbratus: Wilson, 1955a PDF: 150; Bernard, 1967a PDF: 363; Smith, 1979: 1439; Coovert, 2005 PDF: 127 (error).
Subspecies of Lasius umbratus: Forel, 1874 PDF: 47; Emery & Forel, 1879 PDF: 453; Mayr, 1886d PDF: 430; Lameere, 1892: 65; Emery, in Dalla Torre, 1893 PDF: 191 (footnote); Emery, 1893k PDF: 640; Ruzsky, 1896 PDF: 71; Saunders, 1896 PDF: 24; Wheeler, 1904f PDF: 305; Ruzsky, 1905b: 288; Wheeler, 1905j PDF: 397; Wheeler, 1906g PDF: 13; Bondroit, 1910 PDF: 485; Wheeler, 1910h PDF: 239; Wheeler, 1910a PDF: 569; Karavaiev, 1912b PDF: 587; Stitz, 1914 PDF: 86; Emery, 1914c PDF: 159; Ruzsky, 1914a PDF: 61; Forel, 1915d: 52 (in key); Wheeler, 1916o PDF: 172; Wheeler, 1917a PDF: 528; Wheeler, 1917h PDF: 167; Wheeler, 1917k PDF: 463; Escherich, 1917: 333 (in key); Emery, 1922b PDF: 13; Kulmatycki, 1922 PDF: 79; Soudek, 1922b PDF: 72; Emery, 1925d PDF: 234; Ruzsky, 1925a PDF: 288; Ruzsky, 1925b PDF: 45; Schkaff, 1925b PDF: 276; Stärcke, 1926a PDF: 123 (in key); Karavaiev, 1927d: 279 (in key); Wheeler, 1928d PDF: 120; Smith, 1928c PDF: 277; Karavaiev, 1931e PDF: 215; Arnol'di, 1933a: 603 (in key); Cole, 1936a PDF: 37; Dennis, 1938 PDF: 296; Wesson & Wesson, 1940 PDF: 101; Teranishi, 1940: 22; Cole, 1942 PDF: 375; Smith, 1951c PDF: 852; Chapman & Capco, 1951 PDF: 202; Ceballos, 1956: 317.
Status as species: Foerster, 1850a: 41, 72; Schenck, 1852 PDF: 64; Mayr, 1855 PDF: 367 (redescription); Nylander, 1856b PDF: 70; Gredler, 1858 PDF: 15; Smith, 1858a PDF: 9; Mayr, 1861 PDF: 50 (in key); Meinert, 1861: 319; Roger, 1863b PDF: 12; Mayr, 1863a PDF: 426; Dours, 1873 PDF: 165; André, 1874b: 180 (in key); Emery, 1878: 47; André, 1882c PDF: 196 (in key); Nasonov, 1889: 24; Dalla Torre, 1893 PDF: 187; Ruzsky, 1902d PDF: 15; Ruzsky, 1903c PDF: 207; Forel, 1904g PDF: 424; Wasmann, 1906 PDF: 114 (in key); Bondroit, 1912 PDF: 352; Donisthorpe, 1915f: 236; Emery, 1916a PDF: 242; Ruzsky, 1916: 5; Bondroit, 1918 PDF: 32; Bondroit, 1920a PDF: 143; Müller, 1923b PDF: 123 (in key); Donisthorpe, 1927c: 276; Finzi, 1930d PDF: 317; Ruzsky, 1936 PDF: 91; Stärcke, 1937 PDF: 52; Stitz, 1939: 295; Holgersen, 1940 PDF: 185; Novák & Sadil, 1941 PDF: 103 (in key); Holgersen, 1943c PDF: 174 (in key); Holgersen, 1944a PDF: 182; Ruzsky, 1946 PDF: 70; Van Boven, 1947b PDF: 185 (in key); Arnol'di, 1948a PDF: 212 (in list); Azuma, 1953 PDF: 4; Collingwood, 1962c PDF: 219; Collingwood, 1963b PDF: 149; Collingwood & Yarrow, 1969 PDF: 80; Baroni Urbani, 1971c PDF: 211; Collingwood, 1971 PDF: 166; Banert & Pisarski, 1972 PDF: 354; Bourne, 1973 PDF: 26; Bolton & Collingwood, 1975: 7 (in key); Pisarski, 1975: 38; Van Boven, 1977 PDF: 151; Kutter, 1977c: 233; Arnol'di & Dlussky, 1978: 556 (in key); Collingwood, 1978 PDF: 90 (in key); Collingwood, 1979 PDF: 106; Agosti & Collingwood, 1987a PDF: 58; Agosti & Collingwood, 1987b PDF: 282 (in key); Nilsson & Douwes, 1987: 71; Seifert, 1988a PDF: 145 (redescription); Seifert, 1990 PDF: 11; Casevitz-Weulersse, 1990c PDF: 434; Atanassov & Dlussky, 1992: 249; Radchenko, 1994b: 115 (in key); Bolton, 1995b: 224; Douwes, 1995: 94; Poldi et al., 1995: 8; Espadaler, 1997g PDF: 28; Gallé et al., 1998: 217; Czechowski et al., 2002 PDF: 115; Radchenko, 2005b PDF: 170; Bračko, 2006 PDF: 149; Markó et al., 2006 PDF: 68; Petrov, 2006 PDF: 106 (in key); Bračko, 2007 PDF: 20; Radchenko, 2007 PDF: 34; Seifert, 2007: 281; Werner & Wiezik, 2007 PDF: 142; Zryanin & Zryanina, 2007 PDF: 235; Gratiashvili & Barjadze, 2008 PDF: 135; Casevitz-Weulersse & Galkowski, 2009 PDF: 484; Lapeva-Gjonova et al., 2010 PDF: 39; Boer, 2010: 39; Csosz et al., 2011 PDF: 58; Legakis, 2011 PDF: 27; Borowiec & Salata, 2012 PDF: 502; Czechowski et al., 2012: 283; Kiran & Karaman, 2012 PDF: 13; Borowiec, 2014 PDF: 87; Bračko et al., 2014 PDF: 20; Lebas et al., 2016: 206; Radchenko, 2016: 377; Salata & Borowiec, 2018c 10.5281/zenodo.2199191 PDF: 45; Seifert, 2018: 283.
// Distribution

Distribution:

  Geographic regions (According to curated Geolocale/Taxon lists):
    Europe: Belgium, Czech Republic, Finland, Germany, Greece, Ireland, Netherlands, Spain, Sweden, Switzerland
  Biogeographic regions (According to curated Bioregion/Taxon lists):
    Palearctic

Taxonomic Treatment (provided by Plazi)

Treatment Citation: Collingwood, C. A., 1979, The Formicidae (Hymenoptera) of Fennoscandia and Denmark., Fauna Entomologica Scandinavica 8, pp. 1-174

41. Lasius mixtusHNS (Nylander, 1846) Figs. 149-152.

Formica mixta NylanderHNS, 1846b: 1050.

Worker. Yellow to brownish yellow; funiculus segments 2 to 5 not longer than broad; scape broadly oval in cross section. Petiole low, narrowing to emarginate dorsal border. Body pubescence coarse but sparse. Body hairs short, longest gaster hairs less than one third maximum hind tibial width. Front tibiae and scape without erect hairs, occasionally present on hind tibiae. Length: 3.5-4.5 mm.

Queen. Brownish black; head about as broad as maximum width of alitrunk. Funiculus segments 2 to 6 as broad as long. Scape broadly oval in cross section. Sides of petiole convex with emarginate dorsal border. Body pubescence sparse; microsculpture shallow but coarse. Body hairs short, rather sparse. Front tibiae and scapes without suberect hairs, occasionally one or two present on hind tibiae. Eyes with short hairs between facets. Head width: 1.42-1.68 mm. Length: 6.0-7.5 mm.

Figs. 149-152. Lasius mixtus (Nyl.)HNS. - 149: worker in profile; 150: petiole scale of queen in anterior view; 151: head of queen in dorsal view; 152: head of male in dorsal view. Scale: 1 mm.

Male. Dull brownish black; mandibular dentition weak and indistinct. Pubescence sparse, microsculpture coarse with clypeus and frons weakly rugose. Body hairs short, scape and tibiae without suberect hairs. Eyes indistinctly haired. Length: 4.2-4.8 mm.

Distribution. Throughout Denmark and South Fennoscandia to 62°. - Throughout the British Isles. - Range: throughout north Eurasia and subalpine regions of Central Europe.

Biology. This species nests deep in the ground often among shrub roots and under deep boulders, but occasionally also constructs mound nests of fine loose soil. Alatae fly in August and September and fertilised queens are thought to found fresh colonies through adoption by Lasius alienusHNS and L. nigerHNS but actual recorded instances are very few or dubious. Dealate queens are often found wandering singly above ground in spring.

Note. L. mixtusHNS was synonymised by Wilson (1955) as one extreme of a variable species L. umbratus (Nyl.)HNS. However in North Europe the characteristic L. umbratusHNS with setose appendages is consistently distinct and justifies species separation (CoUingwood, 1963b).

Specimen Habitat Summary

Found most commonly in these habitats: 90 times found in Unknown, 20 times found in Rocks (rocky-calcareous grasslands), 12 times found in dry grassland, 11 times found in Wet grassland, 10 times found in Forest, 6 times found in heathlands, 4 times found in Anthropogenic, 2 times found in moorland, 2 times found in dunes & inland dunes, 1 times found in garden, ...

Found most commonly in these microhabitats: 4 times under rock, 2 times under stone, 2 times pasture, under rock, 1 times under ground, 1 times in soil, 1 times ground nest, 1 times grassy mound, pasture, 1 times grassy mound, at hedge, 1 times Forrajeando al sol, 1 times forest margin, under boulder, 1 times foraging with L. fuliginosus workers, ...

Collected most commonly using these methods: 62 times Pitfall trap, 18 times Manual catch, 7 times search, 3 times Hand, 3 times Malaise trap.

Elevations: collected from 250 - 1750 meters, 668 meters average

Collect Date Range: collected between 1881-11-09 and 2019-10-13

Type specimens:



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