This species is one of the most common Pachycondyla species in Costa Rica. Foragers are common on the ground, never arboreal, and relatively more abundant at night. They occur in most samples of sifted leaf litter (Winkler samples), and I collected them once at a tuna bait. In Corcovado, I once observed a mid-morning mating swarm inside an insectary.
I have never found a nest of this common species. The nest must be subterranean. If they nested in the leaf litter or in dead wood, nests would be more frequently encountered.
Garcia-P. et al. (1997) observed harpax preying on termites (Gnathamitermes tubiformans) in the wild.
When pursued with forceps, workers release a stream of clear viscous secretion from the top of the abdomen (Overal 1987). Overal suggests these are defensive secretions employed in tunnels, where room is lacking in which to wield the sting.
harpax. Formica harpax Fabricius, 1804: 401 (w.) SOUTH AMERICA. Wheeler, W.M. 1900b: 4, 17 (q.m.l.); Wheeler, G.C. & Wheeler, J. 1952c: 617 (l.). Combination in Pachycondyla: Roger, 1863b: 18. Senior synonym of concinna, dibullana, irina, montezumia (and its junior synonyms amplinoda, orizabana): Brown, 1950e: 247.
Brown, W. L., Jr. 1950. Morphological, taxonomic, and other notes on ants. Wasmann J. Biol. 8(2):241-250.
Fabricius, J. C. 1804. Systema Piezatorum secundum ordines, genera, species, adjectis synonymis, locis, observationibus, descriptionibus. Brunswick: C. Reichard, xiv + 15-439 + 30pp. Ants - p. 395-428.
Garcia P., J. A., A. Blanco P., R. Mercado H., and M. Badii 1997. The predatory behavior of Pachycondyla harpax Fabr. on Gnathamitermes tubiformans Buckley in captivity conditions. Southwestern Entomologist 22:345-353 [Spanish].
Overal, W. L. 1987. Defensive chemical weaponry in the ant Pachycondyla harpax (Formicidae, Ponerinae). Journal of Entomological Science 22:268-269.