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Genus: Paratopula   Wheeler, 1919 

Classification:

Taxonomic History (provided by Barry Bolton, 2018)

Extant: 12 valid species

Paratopula Wheeler, 1919f PDF: 144. Type-species: Atopomyrmex ceylonicus, by original designation. AntCat AntWiki HOL

Taxonomic history

Paratopula as junior synonym of Atopula: Brown, 1973b PDF: 183 [provisional].
Paratopula as genus: Bolton, 1988b PDF: 125.
Genus Paratopula references
Chapman & Capco, 1951 PDF: 115, 118 (Asia checklist); Bolton, 1988b PDF: 125 (diagnosis, review of genus, all species revision, worker key, male key); Bolton, 1995a PDF: 1051 (census); Bolton, 1995b: 312 (catalogue); Xu & Xu, 2012: 595 (all species worker key); Luo et al., 2016 10.20362/am.008016 PDF: 1 (all species worker key); Cantone, 2017 PDF: 225 (brief male diagnosis)

Distribution:

   (According to curated Geolocale/Taxon lists)   Genus is native to: Indomalaya bioregion (based on species list records).

Indomalaya Region: Borneo, Brunei, India, Indonesia, Malaysia, Myanmar, Philippines, Sri Lanka, Vietnam
Palearctic Region: China

Taxonomic Treatment (provided by Plazi)

Treatment Citation: Eguchi, K., Viet, B. T. & Yamane, S., 2011, Generic synopsis of the Formicidae of Vietnam (Insecta: Hymenoptera), Part I - Myrmicinae and Pseudomyrmicinae., Zootaxa 2878, pp. 1-61

Paratopula WheelerHNS, 1919

Taxonomy. The genus ParatopulaHNS is assigned to the tribe ParatopuliniHNS by Bolton (2003). The genus was revised by Bolton (1988b). The worker of the single Vietnamese species has the following features.

Worker monomorphic; head in full-face view subrectangular; frontal carina indistinct or absent; antennal scrobe absent; anterior clypeal margin feebly emarginated medially, lacking an isolated median seta; posteromedian portion of clypeus very broadly inserted between frontal lobes; mandible triangular; masticatory margin with 9 teeth decreasing in size from apex to base; antennae 12-segmented, with distinct 3-segmented club; eye large, in full-face view strongly convex laterad; mesosoma in lateral view elongate and low; promesonotum in lateral view not domed; promesonotal suture absent dorsally; metanotal groove distinctly impressed dorsally; propodeal spine long and spinose; propodeal lobe extending well posteriad; petiole consisting of elongate peduncle and cubic node, with small anteroventral process; postpetiole in lateral view compressed dorsoventrally, lower than long; gastral shoulder absent; sting simple and strong; head and mesosoma strongly sculptured with rugation or rugoreticulation; body bearing moderately dense hairs that are short and blunt apically.

The worker of ParatopulaHNS is somewhat similar to that of TetramoriumHNS. However, in the latter, the lateral portion of clypeus is modified into a distinct ridge or wall in front of antennal insertion, the apex of sting bears a small lamellate appendage, and the body is much smaller.

Vietnamese species. Only one species is known from Vietnam: sp. eg-1 (Nam Cat Tien).

Bionomics. ParatopulaHNS sp. eg-1 seems to be an arboreal nester and forager, and so it is rarely found on the ground and lower vegetation.



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