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HOLOTYPE WORKER. TL 2.5, HL 0.65, HW 0.58, CI 90, ML 0.32, MI 49, SL 0.42, SI 72, PW 0.31, AL 0.62. Characters of sphera-complex. Each mandible with 1 preapical acute tooth which is situated in the apical third of the length. Upper scrobe margin with a projecting lamellate flange throughout its length, eyes not visible in full-face view. Eye small, convex, its maximum diameter less than maximum width of scape. Basal portion of scape swollen, apical third subcylindrical; hairs on leading edge fine, narrowly spatulate. Cephalic dorsum densely clothed with small appressed scale-like ground-pilosity; upper scrobe margin fringed with short curved narrowly spatulate to spoon-shaped hairs. Cephalic dorsum with 3 pairs of stout standing remiform to clavate hairs arranged in a transverse row close to the occipital margin, and 1 pair on vertex. Pronotal humeral hair absent, the humeral angles bluntly marginate. Mesonotum with 1 pair of stout standing clavate hairs, on anterior margin. Ground-pilosity on alitrunk dorsum sparse, consisting of short narrowly spatulate to broadly spatulate (almost scale-like) hairs. Dorsum of alitrunk in outline convex anteriorly, posterior portion more or less flat to gently sloping to declivity. Metanotal groove shallowly impressed. Propodeal spines narrowly triangular, diverging; lamella absent from declivity. Alitrunk dorsum densely reticulate-punctate. Pronotal dorsum with superimposed rugulose sculpture, most evident at lateral margins. Sides of pronotum striolate to costulate with punctate sculpture on apical margin, remainder more or less smooth. Pleurae smooth and shiny with punctures peripherally. Petiole node in dorsal view approximately as broad as long. Postpetiole disc reticulate-punctate, on some specimens faint longitudinally striolate sculpture present on lateral margins of disc. In profile ventral spongiform tissue of petiolar peduncle a narrow curtain along the base of the peduncle, depth of curtain approximately equal to maximum width of eye. Ventral spongiform lobe of postpetiole moderately developed. Basigastral costulae short but distinct. Dorsal surface of petiole with one pair of erect stout clavate hairs; postpetiole and gaster with stout standing clavate hairs. Colour light to medium brown.
PARATYPE WORKERS. TL 2.6 - 2.7, HL 0.67 - 0.71, HW 0.58 - 0.62, CI 84 - 91, ML 0.33 - 0.36, MI 49 - 50, SL 0.44 - 0.46, SI 71 - 78, PW 0.32 - 0.34, AL 0.63 - 0.68 (6 measured). As holotype.
Holotype worker, Madagascar: 5.3 km. SSE Ambanizana, Andranobe, 15 Ã‚Â° 40 ' S, 49 Ã‚Â° 58 ' E, 425 m., 21. xi. 1993, sifted litter (leaf mold, rotten wood), rainforest, # 926 (l) - 6 (B. L. Fisher) (MCZ).
Paratypes. workers with same data as holotype but coded (10) - 7, (27) - 9, (28) - ll, (35) - 10, (40) - 16. (BMNH, SAM).
NON-PARATYPIC MATERIAL EXAMINED. Madagascar: 5 km. SW Antalaha, 50 m. (CD. Alpert); 19 km. ESE Maroantsetra, 350 m. (P. S. Vlard); Fotodriana, Cap Masoala, 25 m. (O. D. Alpert); 6.3 km. S Ambanizana, Andranobe, 25 m. (B. L. Fisher); Nosy Mangabe, 150 m. (P. S. Ward); Nosy Mangabe (J. - M. Belsch); Baie d'Antongil, Ivontaka, 150 m. (J. - M. Belsch); 1 km. W Andampibe, Cap Masoala, 125 m. (Alpert el al.); 2 km. W Andampibe, Cap Masoala, 125 m. (Alpert el al.); 10 km. NNE Fenoarivo Atm., S. F. Tampolo, 10 m. (B. L. Fisher).
Measurements of this material extend the range shown by the type-series: HL 0.64 - 0.69, HW 0.55 - 0.61, CI 82 - 90, ML 0.31 - 0.34, MI 46 - 52, SL 0.40 - 0.44, SI 70 - 79 (7 measured).
1 Upper scrobe margin with a projecting lamellate flange throughout its length.
2 Basal half of scape distinctly and sharply swollen; in dorsal view maximum width of scape about twice its apical width.
Found most commonly in these habitats: 18 times found in rainforest, 5 times found in littoral rainforest, 22 times found in lowland rainforest, 3 times found in littoral forest, 2 times found in littoral forest on sandy soil, 1 times found in dense humid forest, 3 times found in Secondary Forest, 1 times found in forest, 1 times found in montane rainforest.
Found most commonly in these microhabitats: 20 times sifted litter (leaf mold, rotten wood), 3 times sifted litter, 17 times decayed log, 4 times ex rotten log, 1 times under stone.
Collected most commonly using these methods: 26 times Winkler app., 3 times 10 MaxiWinks, mixed samples, 2 times 8 maxi winks, 3 times MW, 10 sifted litter, 2 times 9 MaxiWinks, mixed samples, 5 times MW 50 sample transect, 5m, 1 times MW 25 sample transect, 5m, 1 times 7 maxi winks, 1 times 9 maxi winks, 1 times MW 75 sample transect, 5,10m, 1 times winkler app, ...
Elevations: collected from 3 - 1175 meters, 126 meters average