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DNA barcoding shows one clade with five subclusters, with less than 3% sequence divergence within the subclusters and about 10% divergence among them. So far no morphological correlates are known.
Types Mus. Comp. Zool. Harvard.
diagnosis Similar to bilimekiHNS, floridanaHNS, hazenaeHNS, rectilumaHNS, rectispinaHNS, rectitrudis, sospesHNS, and stomachosaHNS, differing as follows. Major: mostly blackish brown, with bicolored head; from margins of dorsal head surface to space between eyes and antennal fossae rugoreticulate, as well as lateral margins of frontal lobes; all of central third of head, including much of the vertex and occiput, solely foveolate; all of head and mesosoma overall foveolate; in dorsal-oblique view humerus subangulate; propodeal spines robust and nearly vertical to propodeal basal face in side view; postpetiole from above conulate.
Minor: eyes set well forward on head; propodeal spines robust, nearly vertical to propodeal basal face; almost all of head and all of mesosoma foveolate and opaque.
Measurements (mm) Holotype major: HW 0.90, HL 0.98, SL 0.50, EL 0.12, PW 0.48. Paratype minor: HW 0.50, HL 0.58, SL 0.46, EL 0.06, PW 0.34.
Color Major: head bicolored, with surface anterior to eye level medium reddish brown and rest blackish brown; also sides of pronotum and petiole medium reddish brown; and rest of body blackish brown; appendages light reddish brown. Minor: body blackish brown, appendages brownish yellow.
Range Known only from the type locality.
Biology The type colony was found in forest, nesting in a rotting twig in leaf litter in a tree crotch about one meter from the ground.
figure Upper: holotype, major. Lower: paratype, minor. NICARAGUA: Hotel Selva Negra, 139 km north of Matagalpa, 1200 m (Charles Kugler and J. Hahn). Scale bars = 1 mm.
Found most commonly in these habitats: 490 times found in cloud forest, 18 times found in disturbed cloud forest, 30 times found in mesophil forest, 15 times found in oak cloud forest, 16 times found in mixed hardwood forest, 21 times found in ridgetop cloud forest, 15 times found in cloud forest w/pine, 12 times found in 2º mesophil forest, 12 times found in disturbed pine-oak forest, 5 times found in mesophyl forest, ...
Found most commonly in these microhabitats: 346 times at bait, 334 times ex sifted leaf litter, 3 times nest, 4 times ex sifted litter, 1 times ex dead wood, 6 times beating vegetation, 1 times under epiphytes on ground, 2 times at baits, 1 times at bait on ground, 1 times rotten log, 2 times flight intercept trap, ...
Collected most commonly using these methods: 345 times Baiting, 257 times MiniWinkler, 70 times MaxiWinkler, 14 times search, 9 times Winkler, 5 times bait, 6 times Beating, 2 times Berlese, 2 times Flight Intercept Trap, 1 times Malaise.
Elevations: collected from 1080 - 2190 meters, 1636 meters average