subfamily tribe genus species species author date country valid available current valid name original combination fossil taxonomic history Dolichoderinae TRUE TRUE FALSE
Extant: 28 genera, 692 species, 130 subspecies
Fossil: 16 genera, 103 species
Dolichoderidae Forel, 1878: 364. Type-genus: Dolichoderus.
Taxonomic history
Dolichoderinae as family: Emery, 1894g: 378 [Dolichoderidae]; Ashmead, 1905b: 384 [Dolichoderidae]; Novák & Sadil, 1941: 94 [Dolichoderidae]; Bernard, 1951: 1071 [Dolichoderidae]; Bernard, 1953b: 255 [Dolichoderidae].
Dolichoderinae as tribe of Formicidae: André, 1882a: 127 [Dolichoderidae].
Dolichoderinae as subfamily of Formicidae: Forel, 1878: 364 [Dolichoderidae]; Emery & Forel, 1879a: 454 [Dolichoderidae]; Nasonov, 1889: 26 [Dolichoderidae]; Forel, 1892j: 220 [Dolichoderidae]; Forel, 1893a: 165; Dalla Torre, 1893: 156; Forel, 1895b: 107 [Dolichoderidae]; Emery, 1895j: 771 [subfamily spelled Dolichoderini]; Emery, 1896e: 186; Forel, 1899c: 98; Bingham, 1903: 288; Wheeler, W.M. 1910g: 142; Emery, 1913a: 2; Arnold, 1915: 143; Wheeler, W.M. 1915h: 71; Wheeler, 1915g: 811 [Dolichoderides]; Arnold, 1915: 143; Donisthorpe, 1915d: 178; Forel, 1917: 247; Escherich, 1917: 2 [Dolichoderini]; Bondroit, 1918: 86 [Dolichoderitae]; Wheeler, W.M. 1920: 53; Wheeler, W.M. 1922a: 199; Borgmeier, 1923: 80; Karavaiev, 1936: 163; Smith, M.R. 1951a: 833; Clark, 1951: 16; Brown, 1954e: 29; Kempf, 1972a: 266; Wheeler, G.C. & Wheeler, J. 1972a: 41; Brown, 1973b: 169; all subsequent authors.
Dolichoderinae as formicomorph subfamily of Formicidae: Bolton, 2003: 18, 80.
Dolichoderinae as formicoid subfamily of Formicidae: Brady, Schultz, et al. 2006: 18173; Moreau, Bell, et al. 2006: 102.
Dolichoderinae as formicoid dolichoderomorph subfamily of Formicidae: Ward, 2007a: 556.
Tribes (extant) of Dolichoderinae: Bothriomyrmecini, Dolichoderini, Leptomyrmecini, Tapinomini.
Tribes (extinct) incertae sedis in Dolichoderinae: †Miomyrmecini, †Zherichiniini.
Genera (extinct) incertae sedis in Dolichoderinae: †Alloiomma, †Asymphylomyrmex, †Elaeomyrmex, †Elaphrodites, †Eldermyrmex, †Emplastus, †Eotapinoma, †Eurymyrmex, †Kotshkorkia, †Leptomyrmula, †Miomyrmex, †Petraeomyrmex, †Proiridomyrmex, †Protazteca, †Zherichinius.
Subfamily Dolichoderinae and tribes references, world
Forel, 1878: 364, 380 (diagnosis, genera); Dalla Torre, 1893: 156 (catalogue); Emery, 1895j: 771 (synoptic classification); Emery, 1896e: 186 (genera key); Handlirsch, 1907: 869 (†fossil taxa catalogue); Wheeler, W.M. 1910g: 142 (diagnosis); Emery, 1913a: 2, 6 (diagnosis, tribe key, catalogue); Emery, 1913a: 17 (Tapinomini diagnosis, genera key, catalogue); Arnold, 1915: 144, (diagnosis); Gallardo, 1916a: 3 (diagnosis); Forel, 1917: 247 (synoptic classification); Forel, 1921c: 136 (diagnosis); Wheeler, W.M. 1922a: 199, 687, 688 (diagnosis, tribe key, Tapinomini genera key); Brown & Nutting, 1950: 127 (venation, phylogeny); Brown, 1954e: 29 (phylogeny); Pavan, 1955: 135 (gastral organs); Eisner, 1957: 453 (proventriculus morphology); Bernard, 1967: 246 (diagnosis); Gotwald, 1969: 118 (mouthparts morphology); Wheeler, G.C. & Wheeler, J. 1972a: 41 (diagnosis); Brown, 1973b: 169 (genera & distribution); Wheeler, G.C. & Wheeler, J. 1976b: 61 (larvae, review & synthesis); Snelling, R.R. 1981: 401 (synoptic classification); Dazzini Valcurone & Fanfani, 1985: 1 (gastral glands); Wheeler, G.C. & Wheeler, J. 1985: 258 (synoptic classification); Billen, 1986: 173 (Dufour's gland); Billen, 1987: 278 (abdominal glands); Baroni Urbani & Wilson, 1987: 1 (†fossil Leptomyrmecini); Dlussky & Fedoseeva, 1988: 77 (synoptic classification); Hölldobler & Wilson, 1990: 9 (synoptic classification, genera keys); Baroni Urbani, Bolton & Ward, 1992: 316 (phylogeny); Shattuck, 1992b: 199 (higher classification, phylogeny); Shattuck, 1992c: 20 (revision of subfamily, genera key); Shattuck, 1994: 3 (catalogue); Bolton, 1994: 22 (diagnosis, synoptic classification, genera keys); Shattuck, 1995: 217 (phylogeny, genera); Bolton, 1995a: 1038 (census); Bolton, 1995b: 10 (catalogue); Wenseleers, Schoeters, et al., 1998: 121 (cloacal gland); Brandão, Baroni Urbani, et al. 1999: 411 (phylogeny, genera); Chiotis, Jermiin & Crozier, 2000: 108 (phylogeny, genera); Dlussky & Rasnitsyn, 2002: 416 (diagnosis for impression fossils); Bolton, 2003: 18, 80 (diagnosis, synopsis); Dubovikov, 2005a: 91 (tribes of Dolichoderinae); Brady, Schultz, et al. 2006: 18173 (phylogeny); Moreau, Bell, et al. 2006: 102 (phylogeny); Ward, Brady, et al. 2010: 342 (phylogeny, tribe-rank classification).
Regional and national faunas with keys
André, 1882a: 127 (Europe & Algeria); Nasonov, 1889: 50 (Russia); Forel, 1891b: 9 (Madagascar genera); Forel, 1895e: 460 (India & Sri Lanka); Bingham, 1903: 288 (India, Sri Lanka & Burma); Ruzsky, 1905b: 102 (Russian Empire); Bondroit, 1910: 488 (Belgium); Wheeler, W.M. 1910g: 560 (North America genera); Stitz, 1914: 77 (Central Europe); Gallardo, 1915: 34 (Argentina genera); Forel, 1915d: 40 (Switzerland); Arnold, 1915: 145 (South Africa); Donisthorpe, 1915d: 178 (Britain); Gallardo, 1916a: 12 (Argentina); Emery, 1916b: 208 (Italy); Wheeler, W.M. 1916m: 589 (U.S.A., Connecticut); Bondroit, 1918: 86 (France & Belgium); Gallardo, 1919b: 253 (Argentina genera); Soudek, 1922: 58 (Czechoslovakia); Stärcke, 1926: 117 (Netherlands); Karavaiev, 1927c: 271 (Ukraine); Donisthorpe, 1927b: 199 (Britain); Menozzi & Russo, 1930: 172 (Dominican Republic); Arnol'di, 1933b: 600 (Russia); Menozzi, 1933b: 90 (Israel genera); Karavaiev, 1936: 164 (Ukraine); Smith, M.R. 1937: 861 (Puerto Rico); Stitz, 1939: 209 (Germany); Kratochvíl, 1941: 94 (Central Europe); Novák & Sadil, 1941: 94 (Central Europe); Cole, 1942: 370 (U.S.A., Utah); Smith, M.R. 1943f: 309 (U.S.A. males); Buren, 1944a: 290 (U.S.A., Iowa); Smith, M.R. 1947f: 592 (U.S.A. genera); Creighton, 1950a: 330 (North America); Kusnezov, 1956: 27 (Argentina); Brown, 1958h: 29 (New Zealand); Kusnezov, 1959: 40 (Neotropical genera); Gregg, 1963: 341 (U.S.A., Colorado); Wheeler, G.C. & Wheeler, J. 1963: 149 (U.S.A., North Dakota); Bernard, 1967: 246 (Western Europe); Wilson & Taylor, 1967: 17 (Polynesia); Bolton, 1973a: 329 (West Africa genera); Bolton & Collingwood, 1975: 3 (Britain); Boven, 1970b: 24 (Netherlands); Snelling, R.R. & Hunt, 1976: 90 (Chile); Tarbinsky, 1976: 118 (Kyrghyzstan); Boven, 1977: 123 (Belgium); Kutter, 1977c: 170 (Switzerland); Arnol'di & Dlussky, 1978: 546 (former European U.S.S.R.); Collingwood, 1978: 80 (Iberian Peninsula); Collingwood, 1979: 32 (Fennoscandia & Denmark); Greenslade, 1979: 28 (South Australia genera); Schembri & Collingwood, 1981: 420 (Malta); Allred, 1982: 443 (U.S.A., Utah); Baroni Urbani, 1984: 80 (Neotropical genera); Gösswald, 1985: 314 (Germany); Collingwood, 1985: 242 (Saudi Arabia); Wheeler, G.C. & Wheeler, J. 1986g: 55 (U.S.A., Nevada); Agosti & Collingwood, 1987b: 278 (Balkans); Dlussky, Soyunov & Zabelin, 1990: 168 (Turkmenistan); Kupyanskaya, 1990: 152 (Far Eastern Russia); Morisita, Kubota, Onoyama, et al., 1991: 2 (Japan); Atanasov & Dlussky, 1992: 49 (Bulgaria); Lattke, in Jaffe, 1993: 147 (Neotropical genera); Arakelian, 1994: 70 (Armenia); Wu, J. & Wang, 1995: 115 (China genera); Kupyanskaya, 1995: 331 (Far Eastern Russia); Collingwood & Agosti, 1996: 359 (Saudi Arabia); Seifert, 1996b: 161 (Central Europe); Skinner & Allen, 1996: 40 (Britain); Collingwood & Prince, 1998: 20 (Portugal); Kim & Kim, 1999: 18 (Korea); Shattuck, 1999: 26, 64 (Australia genera, synopsis); Andersen, 2000: 59 (northern Australia genera); Zhou, 2001b: 149 (China, Guangxi); Czechowski, Radchenko & Czechowska, 2002: 134 (Poland); Aktaç & Radchenko, 2002: 54 (Turkey genera); Yoshimura & Onoyama, 2002: 425 (Japan genera, males); Mackay & Mackay, 2002: 221 (U.S.A., New Mexico); Palacio & Fernández, in Fernández, 2003d: 239 (Neotropical genera); Radchenko, 2005b: 186 (North Korea); Coovert, 2005: 102 (U.S.A., Ohio); Clouse, 2007b: 190 (Micronesia); Seifert, 2007: 147 (North and Central Europe); Fisher, 2009: 51 (Malagasy genera); Terayama, 2009: 196 (Taiwan); Heterick, 2009: 28 (south-western Australia genera); Boer, 2010: 14 (Benelux); Yoshimura & Fisher, 2011: 7 (Malagasy males key).
Formicinae TRUE TRUE FALSEExtant: 49 genera, 2,831 species, 839 subspecies
Fossil: 29 genera, 193 species, 1 subspecies
Formicariae Latreille, 1809: 124. Type-genus: Formica.
Taxonomic history
Formicinae as group name: Latreille, 1809: 124 [Formicariae]; Lepeletier de Saint-Fargeau, 1835: 197 [Formicites]; Nylander, 1846a: 877 [Formicae].
Formicinae as family: Smith, F. 1858b: 1 [Formicidae]; Smith, F. 1861b: 36 [Formicidae]; Smith, 1871a: 302 [Formicidae]; André, 1882a: 125 [Formicidae]; Saunders, 1896: 18 [Formicidae]; Novák & Sadil, 1941: 97 [Formicidae]; Bernard, 1951: 1073 [Formicidae]; Bernard, 1953b: 256 [Formicidae].
Formicinae as tribe of Formicidae: André, 1874: 167 [Formicidae].
Formicinae as subfamily of Formicidae: Mayr, 1855: 286, 299 [Formicidae]; Mayr, 1861: 21 [Formicidae]; Mayr, 1862: 651 [Formicidae]; Mayr, 1865: 6 [Formicidae]; Mayr, 1868c: 24 [Formicidae]; Forel, 1870: 307 [Formicidae]; Forel, 1874: 21 [Formicidae]; Emery, 1877a: 70 [Formicidae]; André, 1881b: 54 [Formicidae]; Ashmead, 1905b: 384; Bondroit, 1918: 17 [Formicitae]; Wheeler, W.M. 1920: 53; Wheeler, W.M. 1922a: 210; Emery, 1925b: 2; Karavaiev, 1936: 172; Clark, 1951: 16; Brown, 1954e: 29; Kempf, 1972a: 266; Brown, 1973b: 169; all subsequent authors.
Formicinae as formicomorph subfamily of Formicidae: Bolton, 2003: 20, 93.
Formicinae as formicoid subfamily of Formicidae: Brady, Schultz, et al. 2006: 18173; Moreau, Bell, et al. 2006: 102.
Formicinae as formicoid formicomorph subfamily of Formicidae: Ward, 2007a: 556.
Tribes of Formicinae: Camponotini, Formicini, Gesomyrmecini, Gigantiopini, Lasiini, Melophorini, Myrmecorhynchini, Myrmoteratini, Notostigmatini, Oecophyllini, Plagiolepidini.
Genera (extinct) incertae sedis in Formicinae: †Camponotites, †Drymomyrmex, †Imhoffia, †Kyromyrma, †Leucotaphus, †Protrechina, †Zhangidris.
Hong (2002) genera (extinct) incertae sedis in Formicinae: †Curtipalpulus, †Eoleptocerites, †Eurytarsites, †Fushuniformica, †Huaxiaformica, †Leptogasteritus, †Liaoformica, †Longiformica, †Magnogasterites, †Orbicapitia, †Ovalicapito, †Ovaligastrula, †Sinoformica, †Sinotenuicapito, †Wilsonia (unresolved junior homonym).
Subfamily Formicinae references, world
Mayr, 1862: 651 (genera key); Mayr, 1865: 6 (diagnosis); Handlirsch, 1907: 859 (†fossil taxa catalogue); Dalla Torre, 1893: 171 (catalogue); Emery, 1895j: 772 (synoptic classification); Emery, 1896e: 187 (genera key); Wheeler, W.M. 1910g: 143 (diagnosis); Forel, 1912i: 88 (tribes key); Forel, 1917: 248 (synoptic classification); Arnold, 1920a: 551 (diagnosis); Forel, 1921c: 139 (diagnosis); Wheeler, W.M. 1922a: 210, 691 (diagnosis, tribes key); Emery, 1925b: 2 (diagnosis, tribe key, catalogue); Brown & Nutting, 1950: 127 (venation, phylogeny); Eisner, 1957: 465 (proventriculus morphology); Hung & Brown, 1966: 198 (gastric apex, structure); Bernard, 1967: 267 (diagnosis); Gotwald, 1969: 120 (mouthparts morphology); Wheeler, G.C. & Wheeler, J. 1972a: 41 (diagnosis); Brown, 1973b: 169 (genera, distribution); Wheeler, G.C. & Wheeler, J. 1976b: 62 (larvae, review & synthesis); Snelling, R.R. 1981: 402 (synoptic classification); Wheeler, G.C. & Wheeler, J. 1985: 258 (synoptic classification); Billen, 1986: 173 (Dufour's gland); Dlussky & Fedoseeva, 1988: 77 (synoptic classification); Hölldobler & Wilson, 1990: 9 (synoptic classification, genera keys); Agosti, 1991: 295 (genus group diagnoses); Shattuck, 1992b: 201 (phylogeny); Baroni Urbani, Bolton & Ward, 1992: 317 (phylogeny); Bolton, 1994: 42 (diagnosis, synoptic classification, genera keys); Bolton, 1995a: 1039 (census); Bolton, 1995b: 11 (catalogue); Wenseleers, Schoeters, et al. 1998: 121 (cloacal gland); Dlussky & Rasnitsyn, 2002: 417 (diagnosis for impression fossils); Bolton, 2003: 20, 93 (diagnosis, synopsis); Brady, Schultz, et al. 2006: 18173 (phylogeny); Moreau, Bell, et al. 2006: 102 (phylogeny).
Regional and national faunas with keys
Mayr, 1855: 299 (Austria); Mayr, 1861: 25 (Europe); Mayr, 1868c: 25 (†Baltic Amber); André, 1874: 167 (Europe); Forel, 1874: 22 (Switzerland); Saunders, E. 1880: 203 (Britain); André, 1882a: 126 (Europe & Algeria); Provancher, 1887: 225 (Canada); Cresson, 1887: 94 (U.S.A. genera); Nasonov, 1889: 50 (Russia); Forel, 1891b: 8 (Madagascar genera); Lameere, 1892: 62 (Belgium); Forel, 1892j: 220 (India & Sri Lanka); Bingham, 1903: 308 (India, Sri Lanka & Burma); Ruzsky, 1905b: 100 (Russian Empire); Wasmann, 1906: 7 (Luxemburg); Bondroit, 1910: 481 (Belgium); Wheeler, W.M. 1910g: 560 (North America genera); Stitz, 1914: 80 (Central Europe); Gallardo, 1915: 35 (Argentina genera); Forel, 1915d: 45 (Switzerland); Donisthorpe, 1915d: 184 (Britain); Emery, 1916b: 216 (Italy); Wheeler, W.M. 1916m: 590 (U.S.A., Connecticut); Bondroit, 1918: 17 (France & Belgium); Arnold, 1920a: 552 (South Africa); Kutter, 1920b: 134 (Switzerland); Soudek, 1922: 61 (Czechoslovakia); Lomnicki, 1925a: 160 (Poland); Stärcke, 1926: 118, 146 (Netherlands); Karavaiev, 1927c: 273 (Ukraine); Donisthorpe, 1927b: 205 (Britain); Menozzi & Russo, 1930: 172 (Dominican Republic); Arnol'di, 1933b: 601 (Russia); Menozzi, 1933b: 90 (Israel genera); Karavaiev, 1936: 173 (Ukraine); Smith, M.R. 1937: 865 (Puerto Rico); Stitz, 1939: 230 (Germany); Kratochvíl, 1941: 97 (Central Europe); Novák & Sadil, 1941: 97 (Central Europe); Cole, 1942: 373 (U.S.A., Utah); Smith, M.R. 1943f: 309 (U.S.A., males); Buren, 1944a: 292 (U.S.A., Iowa); Holgersen, 1943b: 173 (Norway); Holgersen, 1944: 199 (Norway); Smith, M.R. 1947f: 599 (U.S.A. genera); Boven, 1947: 181 (Belgium); Creighton, 1950a: 355 (North America); Kusnezov, 1956: 31 (Argentina); Brown, 1958h: 42 (New Zealand); Boven, 1959: 11 (Netherlands); Gregg, 1963: 447 (U.S.A., Colorado); Wheeler, G.C. & Wheeler, J. 1963: 160 (U.S.A., North Dakota); Collingwood, 1964: 104 (Britain); Bernard, 1967: 268 (Western Europe); Wilson & Taylor, 1967: 17 (Polynesia); Boven, 1970b: 26 (Netherlands); Kempf, 1972a: 266 (Neotropical, synoptic classification); Bolton, 1973a: 329 (West Africa genera); Bolton & Collingwood, 1975: 3 (Britain); Snelling & Hunt, 1976: 104 (Chile); Tarbinsky, 1976: 126 (Kyrghyzstan); Boven, 1977: 126 (Belgium); Kutter, 1977c: 183 (Switzerland); Arnol'di & Dlussky, 1978: 548 (former European U.S.S.R.); Collingwood, 1978: 88 (Iberian Peninsula); Collingwood, 1979: 85 (Fennoscandia & Denmark); Greenslade, 1979: 32 (South Australia genera); Schembri & Collingwood, 1981: 436 (Malta); Prins, 1983: 8 (Southern Africa genera); Allred, 1982: 444 (U.S.A., Utah); Verhaeghe, Deligne, et al., 1984: 106 (Belgium genera); Baroni Urbani, 1984: 81 (Neotropical genera); Gösswald, 1985: 263 (Germany); Collingwood, 1985: 273 (Saudi Arabia); Wheeler, G.C. & Wheeler, J. 1986g: 58 (U.S.A., Nevada); Nilsson & Douwes, 1987: 68 (Norway); Agosti & Collingwood, 1987b: 279 (Balkans); Dlussky, Soyunov & Zabelin, 1990: 124 (Turkmenistan); Kupyanskaya, 1990: 162 (Far Eastern Russia); Morisita, Kubota, Onoyama, et al., 1991: 10 (Japan); Atanasov & Dlussky, 1992: 49 (Bulgaria); Shattuck, 1992b: 199 (higher classification, phylogeny); Lattke, in Jaffe, 1993: 150 (Neotropical genera); Arakelian, 1994: 76 (Armenia); Wu, J. & Wang, 1995: 125 (China genera); Kupyanskaya, 1995: 332 (Far Eastern Russia); Collingwood & Agosti, 1996: 361 (Saudi Arabia); Seifert, 1996b: 166 (Central Europe); Skinner & Allen, 1996: 41 (Britain); Collingwood & Prince, 1998: 21 (Portugal); Shattuck, 1999: 25, 86 (Australia genera, synopsis); Andersen, 2000: 68 (northern Australia genera); Zhou, 2001b: 165 (China, Guangxi); Czechowski, Radchenko & Czechowska, 2002: 147 (Poland); Aktaç & Radchenko, 2002: 54 (Turkey genera); Yoshimura & Onoyama, 2002: 425 (Japan genera, males); Mackay & Mackay, 2002: 236 (U.S.A., New Mexico); Palacio & Fernández, in Fernández, 2003d: 242 (Neotropical genera); Radchenko, 2005b: 187 (North Korea); Coovert, 2005: 113 (U.S.A., Ohio); Clouse, 2007b: 190 (Micronesia); Seifert, 2007: 150 (North and Central Europe); Terayama, 2009: 202 (Taiwan); Heterick, 2009: 30 (south-western Australia genera); Boer, 2010: 17 (Benelux).
Myrmicinae TRUE TRUE FALSEExtant: 141 genera, 6,087 species, 864 subspecies
Fossil: 31 genera, 130 species
Myrmicites Lepeletier de Saint-Fargeau, 1835: 169. Type-genus: Myrmica.
Taxonomic history
Myrmicinae as group name: Lepeletier de Saint-Fargeau, 1835: 169 [Myrmicites]; Nylander, 1846a: 877 [Myrmicae].
Myrmicinae as family: Smith, F. 1851: 4 [Myrmicidae]; Smith, F. 1861b: 45 [Myrmicidae]; Smith, F. 1871a: 324 [Myrmicidae]; André, 1882a: 125 [Myrmicidae]; Cresson, 1887: 93 [Myrmicidae]; Emery, 1894g: 383 [Myrmicidae]; Saunders, 1896: 18 [Myrmicidae]; Ashmead, 1905b: 383 [Myrmicidae]; Novák & Sadil, 1941: 71 [Myrmicidae]; Bernard, 1951: 1058 [Myrmicidae]; Bernard, 1953b: 222 [Myrmicidae].
Myrmicinae as subfamily of Poneridae: Smith, F. 1858b: 114 [Myrmicidae].
Myrmicinae as tribe of Formicidae: André, 1874: 167 [Myrmicidae].
Myrmicinae as subfamily of Myrmicidae: Ashmead, 1905b: 383.
Myrmicinae as subfamily of Formicidae: Mayr, 1855: 290, 299 [Myrmicidae]; Smith, F. 1857a: 70 [Myrmicidae]; Mayr, 1861: 21 [Myrmicidae]; Smith, F. 1862b: 33 [Myrmicidae]; Mayr, 1862: 738 [Myrmicidae]; Mayr, 1865: 17 [Myrmicidae]; Mayr, 1868c: 24 [Myrmicidae]; Forel, 1870: 307 [Myrmicidae]; Forel, 1874: 22 [Myrmicidae]; Emery, 1877a: 70 [Myrmicidae]; Forel, 1878: 367 [Myrmicidae]; Emery & Forel, 1879a: 456 [Myrmicidae]; André, 1881b: 64 [Myrmicidae]; Nasonov, 1889: 28 [Myrmicidae]; Forel, 1891b: 11 [Myrmicidae]; Forel, 1892j: 220 [Myrmicidae]; Forel, 1893a: 163; Dalla Torre, 1893: 53; Emery, 1895j: 768 [subfamily spelled Myrmicini]; Emery, 1896e: 179; Forel, 1899c: 30; Forel, 1902f: 520; Bingham, 1903: 105; Wheeler, W.M. 1910g: 138; Emery, 1914a: 29; Wheeler, 1915g: 806 [Myrmicides]; Wheeler, W.M. 1915h: 40; Donisthorpe, 1915d: 74; Arnold, 1916: 166; Escherich, 1917: 2 [Myrmicini]; Forel, 1917: 240 [subfamily spelled Myrmicini]; Bondroit, 1918: 14 [Myrmicitae]; Wheeler, W.M. 1920: 53; Wheeler, W.M. 1922a: 124; Emery, 1921f: 3; Karavaiev, 1934: 59; Clark, 1951: 16; Brown, 1954e: 28; Wheeler, G.C. & Wheeler, J. 1972a: 40; Brown, 1973b: 166; all subsequent authors.
Myrmicinae as myrmicomorph subfamily of Formicidae: Bolton, 2003: 52, 182.
Myrmicinae as formicoid subfamily of Formicidae: Moreau, Bell, et al. 2006: 102; Brady, Schultz, et al. 2006: 18173.
Myrmicinae as formicoid myrmicomorph subfamily of Formicidae: Ward, 2007a: 556.
Tribes of Myrmicinae: Adelomyrmecini, Ankylomyrmini, Attini, Basicerotini, Blepharidattini, Cataulacini, Cephalotini, Crematogastrini, Dacetini, Formicoxenini, Lenomyrmecini, Liomyrmecini, Melissotarsini, Meranoplini, Metaponini, Myrmecinini, Myrmicariini, Myrmicini, Paratopulini, Phalacromyrmecini, Pheidolini, Solenopsidini, Stegomyrmecini, Stenammini, Tetramoriini.
Genera (extant) incertae sedis in Myrmicinae: Diaphoromyrma, Tyrannomyrmex.
Genera (extinct) incertae sedis in Myrmicinae: †Afromyrma, †Attopsis, †Cephalomyrmex, †Electromyrmex, †Eocenidris, †Eomyrmex, †Fallomyrma, †Miosolenopsis, †Myrmecites, †Solenopsites.
Hong (2002) genera (extinct) incertae sedis in Myrmicinae: †Brachytarsites, †Clavipetiola, †Fushunomyrmex, †Orbigastrula, †Quadrulicapito, †Quineangulicapito, †Sinomyrmex, †Sphaerogasterites, †Wumyrmex.
Collective group name in Myrmicinae: †Myrmicites.
Subfamily Myrmicinae references, world
Mayr, 1865: 17 (diagnosis); Mayr, 1867a: 91 (diagnosis); Forel, 1878: 367 (diagnosis); Handlirsch, 1907: 872 (†fossil taxa catalogue); Dalla Torre, 1893: 53 (catalogue); Emery, 1895j: 768 (diagnosis); Emery, 1896e: 179 (genera key); Wheeler, W.M. 1910g: 138 (diagnosis); Emery, 1912b: 101 (phylogeny); Emery, 1914a: 34 (phylogeny, tribe key); Arnold, 1916: 164 (diagnosis); Forel, 1917: 240 (synoptic classification); Forel, 1921c: 139 (diagnosis); Emery, 1921f: 3 (diagnosis, tribes & genera key, catalogue); Wheeler, W.M. 1922a: 124, 655 (diagnosis, tribes key); Brown & Nutting, 1950: 126 (venation, phylogeny); Brown, 1954e: 28 (phylogeny); Eisner, 1957: 477 (proventriculus morphology); Bernard, 1967: 93 (diagnosis); Gotwald, 1969: 99 (mouthparts morphology); Wheeler, G.C. & Wheeler, J. 1972a: 40 (diagnosis); Brown, 1973b: 166 (genera, distribution); Wheeler, G.C. & Wheeler, J. 1976b: 52 (larvae, review & synthesis); Kugler, C. 1978a: 413 (sting structure); Kugler, C. 1978b: 267 (pygidial glands); Kugler, C. 1979c: 117 (sting, evolution); Snelling, R.R. 1981: 393 (synoptic classification); Caetano, F.H. 1984: 257 (digestive tract, morphology); Wheeler, G.C. & Wheeler, J. 1985: 257 (synoptic classification); Billen, 1986: 167 (Dufour's gland); Dlussky & Fedoseeva, 1988: 79 (synoptic classification); Hölldobler & Wilson, 1990: 9 onward (synoptic classification, genera keys); Baroni Urbani, Bolton & Ward, 1992: 317 (phylogeny); Bolton, 1994: 75 (diagnosis, synoptic classification, genera keys); Bolton, 1995a: 1040 (census); Bolton, 1995b: 13 (catalogue); Hashimoto, 1996: 354 (phylogenetic position); Baroni Urbani, 2000: 480 (phylogeny); Dlussky & Rasnitsyn, 2002: 422 (diagnosis for wingless fossils); Bolton, 2003: 52, 182 (diagnosis, synopsis); Moreau, Bell, et al. 2006: 102 (phylogeny); Brady, Schultz, et al. 2006: 18173 (phylogeny); Ward, 2007a: 556 (classification).
Regional and national faunas with keys
Mayr, 1855: 391 (Austria); Mayr, 1861: 29 (Europe); Mayr, 1868c: 79 (†Baltic Amber); André, 1874: 171 (Europe); Forel, 1874: 29 (Switzerland); Saunders, E. 1880: 213 (Britain); André, 1882c: 256 (Europe & Algeria); Cresson, 1887: 98 (U.S.A. genera); Provancher, 1887: 243 (Canada); Nasonov, 1889: 54 (Russia); Forel, 1891b: 11 (Madagascar genera); Lameere, 1892: 66 (Belgium); Forel, 1902f: 520 (India & Sri Lanka genera); Bingham, 1903: 105 (India, Sri Lanka & Burma); Ruzsky, 1905b: 103 (Russian Empire); Wasmann, 1906: 13 (Luxemburg); Bondroit, 1910: 490 (Belgium); Wheeler, W.M. 1910g: 558 (North America genera); Stitz, 1914: 55 (Central Europe); Gallardo, 1915: 32 (Argentina genera); Forel, 1915d: 8 (Switzerland); Donisthorpe, 1915d: 74 (Britain); Arnold, 1916: 166, 170 (South Africa tribes, genera); Emery, 1916b: 112 (Italy); Wheeler, W.M. 1916m: 581 (U.S.A., Connecticut); Bondroit, 1918: 90 (France & Belgium); Kutter, 1920b: 144 (Switzerland); Soudek, 1922: 20 (Czechoslovakia); Stärcke, 1926: 84 (Netherlands); Karavaiev, 1927c: 256 (Ukraine); Donisthorpe, 1927b: 77 (Britain); Menozzi & Russo, 1930: 170 (Dominican Republic); Gallardo, 1932b: 91 (Argentina, tribes); Arnol'di, 1933b: 596 (Russia); Menozzi, 1933b: 88 (Israel genera); Karavaiev, 1934: 60 (Ukraine); Smith, M.R. 1937: 829 (Puerto Rico); Stitz, 1939: 63 (Germany); Kratochvíl, 1941: 71 (Central Europe); Novák & Sadil, 1941: 71 (Central Europe); Cole, 1942: 360 (U.S.A., Utah); Smith, M.R. 1943f: 291 (U.S.A., males); Holgersen, 1943b: 166 (Norway); Holgersen, 1944: 198 (Norway); Buren, 1944a: 281 (U.S.A., Iowa); Smith, M.R. 1947f: 543 (U.S.A. genera); Boven, 1947: 170 (Belgium); Creighton, 1950a: 83 (North America); Kusnezov, 1956: 15 (Argentina); Brown, 1958h: 25 (New Zealand); Boven, 1959: 7 (Netherlands); Gregg, 1963: 288 (U.S.A., Colorado); Wheeler, G.C. & Wheeler, J. 1963: 92 (U.S.A., North Dakota); Collingwood, 1964: 94 (Britain); Bernard, 1967: 95 (Western Europe); Wilson & Taylor, 1967: 13 (Polynesia); Boven, 1970b: 9 (Netherlands); Kempf, 1972a: 263 (Neotropical, synoptic classification); Bolton, 1973a: 325 (West Africa genera); Bolton & Collingwood, 1975: 3 (Britain); Snelling, R.R. & Hunt, 1976: 70 (Chile); Tarbinsky, 1976: 19 (Kyrghyzstan); Boven, 1977: 69 (Belgium); Kutter, 1977c: 31 (Switzerland); Arnol'di & Dlussky, 1978: 524 (former European U.S.S.R.); Collingwood, 1978: 75 (Iberian Peninsula); Collingwood, 1979: 36 (Fennoscandia & Denmark); Greenslade, 1979: 20 (South Australia genera); Schembri & Collingwood, 1981: 423 (Malta); Allred, 1982: 438 (U.S.A., Utah); Baroni Urbani, 1984: 76 (Neotropical genera); Verhaeghe, Deligne, et al. 1984: 112 (Belgium genera); Gösswald, 1985: 289 (Germany); Collingwood, 1985: 245 (Saudi Arabia); Wheeler, G.C. & Wheeler, J. 1986g: 20 (U.S.A., Nevada); Nilsson & Douwes, 1987: 57 (Norway); Agosti & Collingwood, 1987b: 265 (Balkans); Dlussky, Soyunov & Zabelin, 1990: 181 (Turkmenistan); Kupyanskaya, 1990: 89 (Far Eastern Russia); Ogata, 1991b: 61 (Japan genera); Morisita, Kubota, et al. 1992: 1 (Japan); Atanasov & Dlussky, 1992: 51 (Bulgaria); Lattke, in Jaffe, 1993: 153 (Neotropical genera); Arakelian, 1994: 15 (Armenia); Wu, J. & Wang, 1995: 57 (China genera); Kupyanskaya, 1995: 327 (Far Eastern Russia); Collingwood & Agosti, 1996: 308 (Saudi Arabia); Seifert, 1996b: 108 (Central Europe); Skinner & Allen, 1996: 43 (Britain); Collingwood & Prince, 1998: 10 (Portugal); Shattuck, 1999: 39, 122 (Australia genera, synopsis); Andersen, 2000: 37 (northern Australia genera); Zhou, 2001b: 69 (China, Guangxi); Czechowski, Radchenko & Czechowska, 2002: 135 (Poland); Aktaç & Radchenko, 2002: 55 (Turkey genera); Yoshimura & Onoyama, 2002: 424 (Japan genera, males); Mackay & Mackay, 2002: 58 (U.S.A., New Mexico); Palacio & Fernández, in Fernández, 2003d: 244 (Neotropical genera and synopsis); Coovert, 2005: 32 (U.S.A., Ohio); Radchenko, 2005b: 184 (North Korea); Clouse, 2007b: 190 (Micronesia); Seifert, 2007: 110 (North and Central Europe); Terayama, 2009: 131 (Taiwan); Heterick, 2009: 36 (south-western Australia genera); Boer, 2010: 46 (Benelux); Eguchi, Bui & Yamane, 2011: 8 (Vietnam genera).
Ponerinae TRUE TRUE FALSEExtant: 27 genera, 1,033 species, 155 subspecies
Fossil: 9 genera, 68 species, 2 subspecies
Ponerites Lepeletier de Saint-Fargeau, 1835: 185. Type-genus: Ponera.
Taxonomic history
Ponerinae as group name: Lepeletier de Saint-Fargeau, 1835: 185 [Ponérites].
Ponerinae as family: Smith, F. 1851: 6 [Poneridae]; Smith, F. 1858b: 76 [Poneridae]; Smith, F. 1861b: 44 [Poneridae]; Smith, F. 1871a: 320 [Poneridae]; André, 1882a: 125 [Poneridae]; Cresson, 1887: 93 [Poneridae]; Emery, 1894g: 379 [Poneridae]; Saunders, 1896: 18 [Poneridae]; Ashmead, 1905b: 382 [Poneridae]; Novák & Sadil, 1941: 70 [Poneridae]; Bernard, 1951: 1042 [Poneridae]; Bernard, 1953b: 185 [Poneridae].
Ponerinae as tribe of Formicidae: André, 1874: 167 [Poneridae].
Ponerinae as subfamily of Poneridae: Ashmead, 1905b: 382.
Ponerinae as subfamily of Formicidae: Mayr, 1855: 289, 299 [Poneridae]; Mayr, 1861: 21 [Poneridae]; Mayr, 1862: 712 [Poneridae]; Smith, F. 1857a: 64 [Poneridae]; Smith, F. 1862b: 31 [Poneridae]; Mayr, 1865: 11 [Poneridae]; Mayr, 1868c: 24 [Poneridae]; Forel, 1870: 307 [Poneridae]; Forel, 1874: 21 [Poneridae]; Emery, 1877a: 70 [Poneridae]; Forel, 1878: 366 [Poneridae]; Emery & Forel, 1879a: 455 [Poneridae]; André, 1881b: 64 [Poneridae]; Forel, 1892j: 220 [Poneridae]; Dalla Torre, 1893: 13; Forel, 1893a: 162; Forel, 1895b: 111 [Poneridae]; Nasonov, 1889: 27 [Poneridae]; Emery, 1895j: 766 [subfamily spelled Ponerini]; Emery, 1896e: 176; Forel, 1899c: 2; Emery, 1901a: 36; Bingham, 1903: 23; Wheeler, W.M. 1910g: 134; Emery, 1911d: 2; Wheeler, 1915g: 805 [Ponerides]; Donisthorpe, 1915d: 65; Wheeler, W.M. 1915h: 25; Arnold, 1915: 9; Escherich, 1917: 2 [Ponerini]; Forel, 1917: 235; Bondroit, 1918: 78 [Poneritae]; Wheeler, W.M. 1920: 53; Wheeler, W.M. 1922a: 56, 632, 640; Borgmeier, 1923: 37; Karavaiev, 1934: 49; Clark, 1951: 15; Brown, 1954e: 24; Wheeler, G.C. & Wheeler, J. 1972a: 39; Brown, 1973b: 165; all subsequent authors.
Ponerinae as poneromorph subfamily of Formicidae: Bolton, 2003: 42, 156.
Ponerinae as poneroid subfamily of Formicidae: Ouellette, Fisher, et al. 2006: 365; Brady, Schultz, et al. 2006: 18173; Moreau, Bell et al. 2006: 102; Ward, 2007a: 555.
Tribes of Ponerinae: Platythyreini, Ponerini, Thaumatomyrmecini.
Genera (extinct) incertae sedis in Ponerinae: †Afropone, †Eogorgites, †Eoponerites, †Furcisutura, †Longicapitia.
Subfamily Ponerinae references, world
Smith, F. 1858b: 76 (diagnosis); Mayr, 1862: 712 (genera key); Mayr, 1865: 11 (Odontomachidae, Poneridae diagnoses), Mayr, 1867a: 79, 81 (Odontomachidae, Poneridae diagnoses); Forel, 1878: 366 (diagnosis); Dalla Torre, 1893: 13 (world catalogue); Emery, 1895j: 766 (diagnosis); Emery, 1896e: 176 (genera key); Handlirsch, 1907: 879 (†fossil taxa catalogue); Wheeler, W.M. 1910g: 134 (diagnosis); Emery, 1911d: 3 (diagnosis, tribes key); Forel, 1917: 235 (synoptic classification); Forel, 1921c: 133 (diagnosis); Wheeler, W.M. 1922a: 56, 636 (diagnosis, tribes key); Brown & Nutting, 1950: 124 (venation, phylogeny); Brown, 1954e: 24 (phylogeny); Eisner, 1957: 475 (proventriculus morphology); Bernard, 1967: 79 (diagnosis); Gotwald, 1969: 25 (mouthparts morphology); Wheeler, G.C. & Wheeler, J. 1972a: 39 (diagnosis); Brown, 1973b: 165 (genera, distribution); Wheeler, G.C. & Wheeler, J. 1976b: 48 (larvae, review & synthesis); Snelling, R.R. 1981: 387 (synoptic classification); Wheeler, G.C. & Wheeler, J. 1985: 256 (synoptic classification); Billen, 1986: 168 (Dufour's gland); Fanfani & Dazzini Valcurone, 1986: 115 (exocrine glands synopsis); Dlussky & Fedoseeva, 1988: 78 (synoptic classification); Hölldobler & Wilson, 1990: 9 onward (synoptic classification, genera keys); Baroni Urbani, Bolton & Ward, 1992: 317 (phylogeny); Bolton, 1994: 153 (diagnosis, synoptic classification, genera keys); Bolton, 1995a: 1042 (census); Bolton, 1995b: 14 (catalogue); Grimaldi, Agosti & Carpenter, 1997: 20 (†Cretaceous genera); Dlussky & Rasnitsyn, 2002: 422 (diagnosis for wingless fossils); Bolton, 2003: 42, 156 (diagnosis, synopsis); Ouellette, Fisher et al. 2006: 359 (phylogeny); Brady, Schultz, et al. 2006: 18173 (phylogeny); Moreau, Bell et al. 2006: 102 (phylogeny); Ward, 2007a: 555 (classification).
.
Regional and national faunas with keys
Mayr, 1861: 28 (Europe); Mayr, 1868c: 69 (†Baltic Amber); André, 1874: 170 (Europe); Forel, 1874: 29 (Switzerland); André, 1882b: 227 (Europe & Algeria); Cresson, 1887: 97 (U.S.A., genera); Provancher, 1887: 238 (Canada); Nasonov, 1889: 54 (Russia); Forel, 1891b: 9 (Madagascar genera); Forel, 1900c: 52 (India & Sri Lanka); Bingham, 1903: 23 (India, Sri Lanka & Burma); Ruzsky, 1905b: 108 (Russian Empire); Wheeler, W.M. 1910g: 557 (North America genera); Bondroit, 1910: 489 (Belgium); Stitz, 1914: 54 (Central Europe); Gallardo, 1915: 31 (Argentina genera); Forel, 1915d: 6 (Switzerland); Arnold, 1915: 10 (South Africa); Donisthorpe, 1915d: 65 (Britain); Emery, 1916b: 98 (Italy); Wheeler, W.M. 1916m: 580 (U.S.A., Connecticut); Bondroit, 1918: 79 (France & Belgium); Gallardo, 1918b: 4 (Argentina); Soudek, 1922: 18 (Czechoslovakia); Stärcke, 1926: 82 (Netherlands); Donisthorpe, 1927b: 68 (Britain); Menozzi & Russo, 1930: 169 (Dominican Republic); Arnol'di, 1933b: 596 (Russia); Karavaiev, 1934: 50 (Ukraine); Smith, M.R. 1937: 823 (Puerto Rico); Stitz, 1939: 58 (Germany); Smith, M.R. 1943f: 278 (U.S.A. males); Buren, 1944a: 279 (U.S.A., Iowa); Smith, M.R. 1947f: 530 (U.S.A. genera); Creighton, 1950a: 31 (North America); Kusnezov, 1956: 11 (Argentina); Brown, 1958h: 11 (New Zealand); Gregg, 1963: 280 (U.S.A., Colorado); Bernard, 1967: 80 (Western Europe); Wilson & Taylor, 1967: 10 (Polynesia); Kempf, 1972a: 262 (Neotropical synoptic classification); Bolton, 1973a: 323 (West Africa genera); Bolton & Collingwood, 1975: 3 (Britain); Boven, 1977: 66 (Belgium); Kutter, 1977c: 21 (Switzerland); Arnol'di & Dlussky, 1978: 522 (former European U.S.S.R.); Collingwood, 1978: 74 (Iberian Peninsula); Collingwood, 1979: 29 (Fennoscandia & Denmark); Greenslade, 1979: 14 (South Australia genera); Francoeur, 1979: 30 (Canada, Québec); Schembri & Collingwood, 1981: 417 (Malta); Allred, 1982: 438 (U.S.A., Utah); Baroni Urbani, 1984: 75 (Neotropical genera); Collingwood, 1985: 236 (Saudi Arabia); Wheeler, G.C. & Wheeler, J. 1986g: 17 (U.S.A., Nevada); Agosti & Collingwood, 1987b: 264 (Balkans); Ogata, 1987: 101 (Japan genera); Morisita, Kubota, et al. 1989: 8 (Japan); Dlussky, Soyunov & Zabelin, 1990: 173 (Turkmenistan); Kupyanskaya, 1990: 85 (Far Eastern Russia); Atanasov & Dlussky, 1992: 51 (Bulgaria); Lattke, in Jaffe, 1993: 166 (Neotropical genera); Arakelian, 1994: 10 (Armenia); Wu, J. & Wang, 1995: 31 (China genera); Collingwood & Agosti, 1996: 308 (Saudi Arabia); Seifert, 1996b: 107 (Central Europe); Collingwood & Prince, 1998: 10 (Portugal); Kim, Kim & Kim, 1998: 145 (Korea); Shattuck, 1999: 52, 179 (Australia genera synopsis); Andersen, 2000: 19 (northern Australia genera); Zhou, 2001b: 22 (China, Guangxi); Czechowski, Radchenko & Czechowska, 2002: 133 (Poland); Aktaç & Radchenko, 2002: 53 (Turkey genera); Yoshimura & Onoyama, 2002: 436 (Japan genera, males); Mackay & Mackay, 2002: 27 (U.S.A., New Mexico); Csösz, 2003: 147 (Carpathian Basin); Palacio & Fernández, in Fernández, 2003d: 256 (Neotropical genera); Radchenko, 2005b: 184 (North Korea); Coovert, 2005: 23 (U.S.A., Ohio); Yoshimura & Fisher, 2007: 27 (Malagasy genera, males); Clouse, 2007b: 190 (Micronesia); Seifert, 2007: 108 (North and Central Europe); Bolton & Fisher, 2008c: 32 (Afrotropical genera); Fernández & Arias-Penna, 2008: 32 (Neotropical genera); Dlussky, 2009: 1071 (Eocene amber); Terayama, 2009: 101 (Taiwan genera); Heterick, 2009: 34 (south-western Australia genera); Boer, 2010: 68 (Benelux).
Dolichoderinae Tapinomini Tapinoma TRUE TRUE FALSEExtant: 64 species, 26 subspecies
Fossil: 4 species
Tapinoma Foerster, 1850a: 43. Type-species: Tapinoma collina (junior synonym of Formica erratica), by monotypy.
Taxonomic history
Tapinoma in Formicidae: Smith, F. 1858b: 55.
Tapinoma in Formicinae: Mayr, 1855: 372 [Formicidae]; Mayr, 1861: 41 [Formicidae]; Mayr, 1862: 653 (in key) [Formicidae]; Mayr, 1865: 10 [Formicidae]; André, 1874: 168 [Formicidae]; André, 1881b: 63 [Formicidae].
Tapinoma in Dolichoderinae: Forel, 1878: 385 [Dolichoderidae]; Emery & Forel, 1879a: 454 [Dolichoderidae]; Forel, 1895b: 109 [Dolichoderidae]; Dalla Torre, 1893: 164; Emery, 1895j: 771; Forel, 1899c: 101; Wheeler, W.M. 1910g: 142.
Tapinoma in Dolichoderidae: Ashmead, 1905b: 384.
Tapinoma in Dolichoderinae, Tapinomini: Emery, 1913a: 38; Arnold, 1915: 152; Forel, 1917: 248; Bondroit, 1918: 89; Wheeler, W.M. 1922a: 690; subsequent authors except those below; then Jaffe, 1993: 9; Dubovikov, 2005a: 91 (implied); Ward, Brady, et al. 2010: 361.
Tapinoma in Dolichoderinae, Dolichoderini: Shattuck, 1992c: 146; Bolton, 1994: 26; Bolton, 2003: 91.
Junior synonyms of TAPINOMA
Micromyrma Dufour, 1857: 60. Type-species: Micromyrma pygmaea, by monotypy.
Taxonomic history
[Type-species not Formica erratica, unjustified subsequent designation by Wheeler, W.M. 1911f: 167.]
Micromyrma as subgenus of Tapinoma: Menozzi, 1925a: 19; Santschi, 1928e: 472; Chapman & Capco, 1951: 192.
Micromyrma as junior synonym of Tapinoma: Mayr, 1863: 455; Forel, 1878: 385; Emery & Forel, 1879a: 454; Dalla Torre, 1893: 164; Emery, 1913a: 38; Wheeler, W.M. 1922a: 923; Smith, M.R. 1951a: 837; Shattuck, 1992c: 146; Shattuck, 1994: 140; Bolton, 1994: 26; Bolton, 2003: 91; Fisher & Bolton, 2007: 65.
Semonius Forel, 1910f: 21. Type-species: Semonius schultzei, by monotypy.
Taxonomic history
Semonius in Dolichoderinae: Wheeler, W.M. 1910g: 142.
Semonius in Dolichoderinae, Tapinomini: Emery, 1913a: 44; Arnold, 1915: 157; Forel, 1917: 247; Wheeler, W.M. 1922a: 688; all subsequent authors.
Semonius as junior synonym of Tapinoma: Shattuck, 1992c: 146; Shattuck, 1994: 140; Bolton, 1994: 26; Bolton, 2003: 91; Fisher & Bolton, 2007: 66.
Pseudaphomomyrmex Wheeler, W.M. 1920: 53. Type-species: Aphomomyrmex emeryi, by original designation.
Taxonomic history
Pseudaphomomyrmex in Formicinae, Dimorphomyrmecini: Donisthorpe, 1943g: 721.
Pseudaphomomyrmex incertae sedis in Formicinae: Dlussky & Fedoseeva, 1988: 77.
Pseudaphomomyrmex in Formicinae, Myrmelachistini: Hölldobler & Wilson, 1990: 18.
Pseudaphomomyrmex in Formicinae, Brachymyrmecini: Bolton, 1994: 50.
Pseudaphomomyrmex in Formicinae, Plagiolepidini: Wheeler, W.M. 1922a: 695; Chapman & Capco, 1951: 214; Wheeler, G.C. & Wheeler, J. 1985: 258; Bolton, 2003: 106.
Pseudaphomomyrmex in Dolichoderinae: LaPolla & Longino, 2006: 305.
Pseudaphomomyrmex as junior synonym of Aphomomyrmex: Emery, 1925b: 44; Donisthorpe, 1943g: 721.
Pseudaphomomyrmex as genus: Wheeler, W.M. 1920: 53; Chapman & Capco, 1951: 214; Bolton, 1994: 50; Bolton, 1995b: 369; Bolton, 2003: 106.
Pseudaphomomyrmex as junior synonym of Tapinoma: Fisher & Bolton, 2007: 66.
Zatapinoma Wheeler, W.M. 1928a: 20. Type-species: Zatapinoma annandalei, by original designation.
Taxonomic history
Zatapinoma in Dolichoderinae, Tapinomini: Donisthorpe, 1943g: 737; all subsequent authors.
Zatapinoma as junior synonym of Tapinoma: Shattuck, 1992c: 146; Shattuck, 1994: 140; Bolton, 1994: 26; Bolton, 2003: 91; Fisher & Bolton, 2007: 66.
Neoclystopsenella Kurian, 1955: 133. Type-species: Neoclystopsenella luffae, by monotypy.
Taxonomic history
Neoclystopsenella in Bethylidae: Kurian, 1955: 133.
Neoclystopsenella in Dolichoderinae: Brown, 1988a: 337.
Neoclystopsenella as junior synonym of Tapinoma: Brown, 1988a: 337; Shattuck, 1992c: 146; Shattuck, 1994: 140; Bolton, 1994: 26; Bolton, 2003: 91; Fisher & Bolton, 2007: 66.
Genus Tapinoma references
Mayr, 1855: 372 (diagnosis); Smith, F. 1858b: 55 (diagnosis); Roger, 1863b: 13, 14 (Tapinoma, Micromyrma catalogues); Mayr, 1863: 455 (catalogue); Mayr, 1865: 10 (diagnosis); Mayr, 1867a: 78 (diagnosis); André, 1882b: 222 (Europe & Algeria species key); Cresson, 1887: 258 (U.S.A. catalogue); Dalla Torre, 1893: 164 (catalogue); Bingham, 1903: 304 (India, Sri Lanka & Burma species key); Emery, 1913a: 38 (diagnosis, catalogue); Emery, 1913a: 44 (Semonius diagnosis, catalogue); Arnold, 1915: 152, 157 (diagnosis, South Africa species key, Semonius diagnosis); Wheeler, W.M. 1922a: 695 (Pseudaphomomyrmex diagnosis, in key). Wheeler, W.M. 1922a: 923, 927 (Afrotropical Tapinoma, Semonius catalogues); Wheeler, W.M. 1922a: 1034 (Malagasy catalogue); Emery, 1925d: 63 (Palaearctic species key); Kuznetsov-Ugamsky, 1927d: 33 (Turkestan species key); Smith, M.R. 1928a: 311 (North America species key); Kratochvíl, Novák & Snoflák, 1944: 79 (Czechoslovakia species key); Creighton, 1950a: 352 (North America species key); Chapman & Capco, 1951: 192, 196 (Asia Semonius, Tapinoma, Zatapinoma checklists); Bernard, 1967: 254 (diagnosis, Western Europe species key); Boven, 1970b: 24 (Netherlands species key); Kempf, 1972a: 246 (Neotropical catalogue); Tarbinsky, 1976: 118 (Kyrghyzstan species key); Kutter, 1977c: 178 (Switzerland species key); Arnol'di & Dlussky, 1978: 547 (former European U.S.S.R. species key); Collingwood, 1978: 87 (Iberian Peninsula species key); Smith, D.R. 1979: 1421 (North America catalogue); Schembri & Collingwood, 1981: 421 (Malta species key); Gösswald, 1985: 317 (Germany species key); Taylor & Brown, D.R. 1985: 106 (Australia catalogue); Agosti & Collingwood, 1987b: 279 (Balkans species key); Taylor, 1987a: 77 (Australia, New Caledonia & New Zealand checklist); Dlussky, Soyunov & Zabelin, 1990: 168 (Turkmenistan species key); Brandão, 1991: 380 (Neotropical catalogue); Shattuck, 1992c: 146 (diagnosis, review of genus); Arakelian, 1994: 72 (Armenia species key); Shattuck, 1994: 140 (catalogue); Bolton, 1995a: 1053 (census); Bolton, 1995b: 399 (catalogue); Wu, J. & Wang, 1995: 116 (China species key); Collingwood & Agosti, 1996: 360 (Saudi Arabia species key); Seifert, 1996b: 165 (Central Europe species key); Skinner & Allen, 1996: 40 (Britain species key); Collingwood & Prince, 1998: 20 (Portugal species key); Shattuck, 1999: 81 (Australia synopsis); Zhou, 2001b: 151 (China, Guangxi species key); Czechowski, Radchenko & Czechowska, 2002: 135 (Poland species key); Seifert, 2007: 149 (North and Central Europe species key); Heterick, 2009: 58 (south-western Australia species key); Yoshimura & Fisher, 2011: 13 (Malagasy males).
Dolichoderinae Tapinomini Technomyrmex TRUE TRUE FALSEExtant: 93 species
Fossil: 4 species
Technomyrmex Mayr, 1872: 147. Type-species: Technomyrmex strenuus, by monotypy.
Taxonomic history
Technomyrmex in Dolichoderinae: Forel, 1878: 380 [Dolichoderidae]; Dalla Torre, 1893: 166; Emery, 1895j: 771; Wheeler, W.M. 1910g: 142.
Technomyrmex in Dolichoderinae, Tapinomini: Emery, 1913a: 42; Arnold, 1915: 147; Forel, 1917: 248; Wheeler, W.M. 1922a: 690; subsequent authors except those below; then Jaffe, 1993: 9; Ward, Brady, et al. 2010: 361.
Technomyrmex in Dolichoderinae, Dolichoderini: Shattuck, 1992c: 153; Bolton, 1994: 26; Bolton, 2003: 92.
Junior synonyms of TECHNOMYRMEX
Engramma Forel, 1905b: 180. Type-species: Engramma lujae, by monotypy.
Taxonomic history
Engramma in Dolichoderinae: Wheeler, W.M. 1910g: 142.
Engramma in Dolichoderinae, Tapinomini: Emery, 1913a: 38; Forel, 1917: 248; Wheeler, W.M. 1922a: 690; all subsequent authors.
Engramma as junior synonym of Technomyrmex: Shattuck, 1992c: 153; Shattuck, 1994: 156; Bolton, 1994: 26; Bolton, 2003: 92; Bolton, 2007a: 4.
Tapinoptera Santschi, 1925g: 348 [as subgenus of Tapinoma]. Type-species: Tapinoma vexatum, by monotypy.
Taxonomic history
Tapinoptera as junior synonym of Tapinoma: Shattuck, 1992c: 146; Shattuck, 1994: 140; Bolton, 1994: 26; Bolton, 2003: 91.
Tapinoptera as junior synonym of Technomyrmex: Fisher & Bolton, 2007: 66 (in text); Bolton, 2007a: 4.
Aphantolepis Wheeler, W.M. 1930d: 44. Type-species: Aphantolepis quadricolor, by monotypy.
Taxonomic history
Aphantolepis in Formicinae, Lasiini: Wheeler, W.M. 1930d: 44.
Aphantolepis in Formicinae, Prenolepidini: Donisthorpe, 1943f: 624.
Aphantolepis as junior synonym of Technomyrmex: Brown, 1953h: 5; Shattuck, 1992c: 153; Shattuck, 1994: 156; Bolton, 1994: 26; Bolton, 2003: 92; Bolton, 2007a: 4.
Genus Technomyrmex references
Forel, 1878: 380 (diagnosis); Dalla Torre, 1893: 166 (catalogue); Bingham, 1903: 301 (India, Sri Lanka & Burma species key); Emery, 1913a: 38 (Engramma diagnosis, catalogue); Emery, 1913a: 42 (diagnosis, catalogue); Arnold, 1915: 147 (diagnosis, South Africa species key); Wheeler, W.M. 1922a: 201, 208, 209 (Engramma, Tapinoma, Technomyrmex diagnoses); Wheeler, W.M. 1922a: 202 (Engramma species key); Wheeler, W.M. 1922a: 922, 925 (Afrotropical Engramma, Technomyrmex catalogues); Wheeler, W.M. 1922a: 1034 (Malagasy catalogue); Taylor & Brown, D.R. 1985: 106 (Australia catalogue); Taylor, 1987a: 77 (Australia, New Caledonia & New Zealand checklist); Shattuck, 1992c: 153 (diagnosis, review of genus); Shattuck, 1994: 156 (catalogue); Bolton, 1995a: 1053 (census); Bolton, 1995b: 402 (catalogue); Wu, J. & Wang, 1995: 118 (China species key); Collingwood & Agosti, 1996: 360 (Saudi Arabia species key); Shattuck, 1999: 83 (Australia synopsis); Zhou, 2001b: 158 (China, Guangxi species key); Bolton, 2007a: 1 (all species revision; Afrotropical & West Palaearctic species key: 13; Malagasy species key: 43; East Palaearctic, Oriental, Malesian & Polynesian species key: 59; Austral species key: 108; New World species key: 119); Fernández & Guerrero, 2008: 110 (New World synopsis, key); Terayama, 2009: 200 (Taiwan species key); Yoshimura & Fisher, 2011: 14 (Malagasy males); Sharaf, et al. 2011: 15 (Arabian Peninsula species key).
Tribes (extinct) incertae sedis in DOLICHODERINAE
Formicinae Lasiini Anoplolepis TRUE TRUE FALSE9 species, 5 subspecies
Anoplolepis Santschi, 1914b: 123 [as subgenus of Plagiolepis]. Type-species: Formica longipes (junior primary homonym in Formica; Formica gracilipes first available replacement name: Bolton, 1995b: 67), by monotypy.
Taxonomic history
Anoplolepis in Camponotinae, Plagiolepidini: Forel, 1917: 249; Arnold, 1922: 593.
Anoplolepis in Formicinae, Plagiolepidini: Wheeler, W.M. 1922a: 696; Emery, 1925b: 16; Santschi, 1926a: 13; all subsequent authors except the following.
Anoplolepis in Formicinae [part in Oecophylla genus group, part in Pseudolasius genus group]: Agosti, 1991: 295, 296.
Anoplolepis in Formicinae, Lasiini: Bolton, 2003: 22, 95.
Anoplolepis as subgenus of Plagiolepis: Santschi, 1914b: 123; Forel, 1917: 249; Arnold, 1922: 593; Wheeler, W.M. 1922a: 696.
Anoplolepis as genus: Emery, 1925b: 16; Santschi, 1926a: 13; all subsequent authors.
Junior synonym of ANOPLOLEPIS
Zealleyella Arnold, 1922: 579 (diagnosis in key) [as subgenus of Plagiolepis]. Type-species: Formica custodiens, by subsequent designation of Santschi, 1926a: 14.
Taxonomic history
Zealleyella as subgenus of Plagiolepis: Arnold, 1922: 579.
Zealleyella as subgenus of Anoplolepis: Santschi, 1926a: 14.
Zealleyella as junior synonym of Anoplolepis: Brown, 1973b: 185 [provisional]; Bolton, 2003: 96, 267.
Genus Anoplolepis references
Arnold, 1922: 593 (South Africa, species); Wheeler, W.M. 1922a: 931, 1036 (Afrotropical, Malagasy catalogues); Emery, 1925b: 16 (diagnosis, catalogue); Santschi, 1926a: 13 (subgenera diagnoses); Santschi, 1937g: 83 (subgenera key); Chapman & Capco, 1951: 213 (Asia checklist); Kempf, 1972a: 22 (Neotropical catalogue); Prins, 1982: 235 (South Africa A. (Zealleyella) partial revision, key); Taylor & Brown, D.R. 1985: 108 (Australia catalogue); Dlussky & Fedoseeva, 1988: 77 (synoptic classification); Bolton, 1994: 51 (synoptic classification); Bolton, 1995a: 1048 (census); Bolton, 1995b: 66 (catalogue); Collingwood & Agosti, 1996: 362 (Saudi Arabia species key); Shattuck, 1999: 88 (Australia synopsis); Bolton, 2003: 267 (genus diagnosis).
Formicinae Plagiolepidini Aphomomyrmex TRUE TRUE FALSE1 species
Aphomomyrmex Emery, 1899e: 493. Type-species: Aphomomyrmex afer, by subsequent designation of Wheeler, W.M. 1911f: 159.
Taxonomic history
Aphomomyrmex in Formicinae, Plagiolepidini: Ashmead, 1905b: 384.
Aphomomyrmex in Camponotinae, Plagiolepidini: Wheeler, W.M. 1910g: 143.
Aphomomyrmex in Camponotinae, Myrmelachistini: Forel, 1912i: 89; Forel, 1917: 249; Arnold, 1920a: 552.
Aphomomyrmex in Formicinae, Dimorphomyrmecini: Emery, 1925b: 44 [subtribe Brachymyrmecini]; Donisthorpe, 1943f: 624.
Aphomomyrmex incertae sedis in Formicinae: Dlussky & Fedoseeva, 1988: 77.
Aphomomyrmex in Formicinae, Myrmelachistini: Wheeler, W.M. 1922a: 697; Hölldobler & Wilson, 1990: 18.
Aphomomyrmex in Formicinae, Pseudolasius genus group: Agosti, 1991: 296.
Aphomomyrmex in Formicinae, Brachymyrmecini: Wheeler, W.M. 1929a: 12; Wheeler, G.C. & Wheeler, J. 1985: 258; Bolton, 1994: 50.
Aphomomyrmex in Formicinae, Plagiolepidini: Bolton, 2003: 23, 101.
[Aphomyrmex Ashmead, 1905a: 111, incorrect subsequent spelling.]
Genus Aphomomyrmex references
Arnold, 1920a: 552 (South Africa species); Wheeler, W.M. 1922a: 940 (Afrotropical catalogue); Emery, 1925b: 44 (diagnosis, catalogue); Snelling, R.R. 1979b: 3 (review of genus); Bolton, 1995a: 1048 (census); Bolton, 1995b: 74 (catalogue).
Formicinae Plagiolepidini Lepisiota TRUE TRUE FALSE81 species, 51 subspecies
Lepisiota Santschi, 1926a: 15 [as subgenus of Acantholepis]. Type-species: Plagiolepis rothneyi, by original designation.
Taxonomic history
[Lepisiota first available replacement name for Acantholepis Mayr, 1861 (junior homonym of Acantholepis Kroyer, 1846: 98 (Pisces)), hence valid name of genus: Bolton, 1995b: 33.]
Lepisiota in Formicinae, Plagiolepidini: Bolton, 1994: 51; Bolton, 2003: 23.
[Lepisiota incorrectly as junior synonym of Acanthomyrmex: Brown, 1973b: 181; Snelling, R.R. 1981: 397. In both publications Acanthomyrmex is an error for Acantholepis.]
Junior synonym of LEPISIOTA
Baroniurbania Pagliano & Scaramozzino, 1990: 4. Unnecessary replacement name for Acantholepis Mayr (junior homonym).
Taxonomic history
Baroniurbania as junior synonym of Lepisiota: Bolton, 1994: 51; Bolton, 2003: 102.
Baroniurbania as subgenus of Lepisiota: Dietrich, 2004: 324.
Genus Lepisiota references
[Note. Entries prior to Bolton, 1995b: 44, refer to genus as Acantholepis.]
Roger, 1863b: 11 (catalogue); Mayr, 1863: 394 (catalogue); Mayr, 1865: 9 (diagnosis); Forel, 1878: 378 (diagnosis); André, 1882b: 210 (Europe & Algeria species key); Forel, 1892a: 41 (all species key); Dalla Torre, 1893: 171 (catalogue); Forel, 1894c: 411 (India & Sri Lanka species key); Bingham, 1903: 315 (India, Sri Lanka & Burma species key); Arnold, 1920a: 554 (diagnosis, South Africa species key); Wheeler, W.M. 1922a: 214, 934, 1036 (diagnosis, Afrotropical, Malagasy catalogues); Emery, 1925b: 23 (diagnosis, catalogue); Kuznetsov-Ugamsky, 1929c: 480 (Turkestan species key); Finzi, 1936: 188 (Egypt species key); Chapman & Capco, 1951: 209 (Asia checklist); Collingwood, 1985: 292 (Saudi Arabia species key); Agosti & Collingwood, 1987b: 281 (Balkans species key); Atanasov & Dlussky, 1992: 205 (Bulgaria species key); Xu, 1994c: 232 (China species key); Arakelian, 1994: 80 (Armenia species key); Bolton, 1994: 51 (synoptic classification); Bolton, 1995a: 1050 (census); Bolton, 1995b: 226 (catalogue); Wu, J. & Wang, 1995: 128 (China species key); Collingwood & Agosti, 1996: 363 (Saudi Arabia species key); Zhou, 2001b: 167 (China, Guangxi species key).
Homonym replaced by LEPISIOTA
Acantholepis Mayr, 1861: 42. Type-species: Hypoclinea frauenfeldi, by monotypy.
Taxonomic history
[Junior homonym of Acantholepis Kroyer, 1846: 98 (Pisces)].
Acantholepis in Formicinae: Mayr, 1861: 42 [Formicinae]; Mayr, 1862: 652 [Formicidae]; Mayr, 1865: 9 [Formicidae].
Acantholepis in Camponotinae: Forel, 1878: 378 [Camponotidae]; Emery & Forel, 1879a: 453 [Camponotidae]; Dalla Torre, 1893: 171.
Acantholepis in Camponotinae, Plagiolepidini: Forel, 1886f: 212; Forel, 1893a: 165; Emery, 1895j: 771; Wheeler, W.M. 1910g: 143; Forel, 1912i: 88; Forel, 1917: 249; Arnold, 1920a: 554.
Acantholepis in Formicinae, Plagiolepidini: Ashmead, 1905b: 384; Wheeler, W.M. 1922a: 696; Emery, 1925b: 23; Santschi, 1926a: 15; all subsequent authors except the following.
Acantholepis in Formicinae, Pseudolasius genus group: Agosti, 1991: 296.
[Achantilepis Santschi, 1935b: 274, incorrect subsequent spelling.]
[Pseudacantholepis Bernard, 1953b: 256 (attributed to Santschi) [as subgenus of Acantholepis]. Unavailable name. Proposed without designation of type-species and therefore unavailable. Species included by Bernard (1953) are all referable to Lepisiota: Bolton, 1995b: 44.]
Extant: 57 species, 19 subspecies
Fossil: 10 species
Plagiolepis Mayr, 1861: 42. Type-species: Formica pygmaea, by monotypy.
Taxonomic history
Plagiolepis in Formicinae: Mayr, 1861: 42 [Formicinae]; Mayr, 1862: 652 [Formicidae]; Mayr, 1865: 7 [Formicidae]; Mayr, 1868c: 36 [Formicidae].
Plagiolepis in Camponotinae: Forel, 1878: 378 [Camponotidae]; Emery & Forel, 1879a: 453 [Camponotidae]; Dalla Torre, 1893: 172.
Plagiolepis in Camponotinae, Plagiolepidini: Forel, 1886f: 212; Forel, 1893a: 165; Emery, 1895j: 771; Wheeler, W.M. 1910g: 143; Forel, 1912i: 88; Wheeler, W.M. 1915h: 100; Forel, 1917: 249; Arnold, 1920a: 578.
Plagiolepis in Formicinae, Plagiolepidini: Ashmead, 1905b: 384; Bondroit, 1918: 18; Wheeler, W.M. 1922a: 696; Emery, 1925b: 19; Santschi, 1926a: 14; all subsequent authors except the one below; Bolton, 2003: 23, 105.
Plagiolepis in Formicinae, Pseudolasius genus group: Agosti, 1991: 296.
Junior synonyms of PLAGIOLEPIS
†Rhopalomyrmex Mayr, 1868c: 41. Type-species: †Rhopalomyrmex pygmaeus, by monotypy.
Taxonomic history
†Rhopalomyrmex in Formicinae: Mayr, 1868c: 41 [Formicidae].
†Rhopalomyrmex in Camponotinae: Forel, 1878: 376 [Camponotidae]; Dalla Torre, 1893: 175.
†Rhopalomyrmex in Camponotinae, Myrmelachistini: Forel, 1912i: 89.
†Rhopalomyrmex in Camponotinae, Plagiolepidini: Wheeler, W.M. 1910g: 143; Wheeler, W.M. 1915h: 103.
†Rhopalomyrmex in Formicinae, Plagiolepidini: Donisthorpe, 1943g: 724; Bolton, 1994: 51.
†Rhopalomyrmex as junior synonym of Plagiolepis: Dlussky, 1997: 624; Bolton, 2003: 105.
Anacantholepis Santschi, 1914e: 36 [as subgenus of Plagiolepis]. Type-species: Plagiolepis (Anacantholepis) decora, by original designation.
Taxonomic history
Anacantholepis as genus: Chapman & Capco, 1951: 212.
Anacantholepis as subgenus of Plagiolepis: Santschi, 1914e: 36 Forel, 1917: 249; Arnold, 1922: 581; Wheeler, W.M. 1922a: 696; Emery, 1925b: 22; Bolton, 1994: 51.
Anacantholepis as junior synonym of Plagiolepis: Brown, 1973b: 178 [provisional]; Bolton, 2003: 105, 268.
Aporomyrmex Faber, 1969: 52. Type-species: Aporomyrmex ampeloni, by original designation.
Taxonomic history
Aporomyrmex in Formicinae, Plagiolepidini: Faber, 1969: 52 (by implication).
Aporomyrmex as junior synonym of Plagiolepis: Brown, 1973b: 178 [provisional]; Bolton, 1994: 51; all subsequent authors.
Paraplagiolepis Faber, 1969: 65 [as subgenus of Plagiolepis]. Type-species: Plagiolepis xene, by monotypy.
Taxonomic history
Paraplagiolepis as junior synonym of Plagiolepis: Brown, 1973b: 183 [provisional]; Bolton, 1994: 51; all subsequent authors.
Genus Plagiolepis references
Roger, 1863b: 11 (catalogue); Mayr, 1863: 442 (catalogue); Mayr, 1865: 7 (diagnosis); Mayr, 1867a: 73 (diagnosis); Forel, 1878: 376, 378 (†Rhopalomyrmex, Plagiolepis diagnoses); André, 1882b: 208 (Europe & Algeria species key); Dalla Torre, 1893: 172, 175 (Plagiolepis, †Rhopalomyrmex, catalogue); Forel, 1894c: 414 (India & Sri Lanka species key); Bingham, 1903: 320 (India, Sri Lanka & Burma species key); Arnold, 1920a: 578 (diagnosis, subgenera key); Arnold, 1922: 579 (South Africa species key); Wheeler, W.M. 1922a: 211, 696, 928, 1035 (diagnosis, subgenera key, Afrotropical, Malagasy catalogues); Emery, 1925b: 19 (diagnosis, catalogue); Emery, 1925b: 22 (P. (Anacantholepis) diagnosis, catalogue); Stitz, 1939: 231 (Germany species key); Kratochvíl, 1941: 98 (Central Europe species key); Novák & Sadil, 1941: 98 (Central Europe species key); Chapman & Capco, 1951: 212, 213 (Asia Anacantholepis, Plagiolepis checklists); Bernard, 1967: 272 (diagnosis, Western Europe species key); Kempf, 1972a: 205 (Neotropical catalogue); Kutter, 1977c: 185 (Switzerland species key); Collingwood, 1978: 89 (Iberian Peninsula species key); Smith, D.R. 1979: 1422 (North America, catalogue); Gösswald, 1985: 264 (Germany species key); Collingwood, 1985: 297 (Saudi Arabia species key); Taylor & Brown, D.R. 1985: 130 (Australia catalogue); Taylor, 1987a: 55 (Australia, New Caledonia checklist); Agosti & Collingwood, 1987b: 280 (Balkans species key); Radchenko, 1989b: 153 (European former U.S.S.R. species key); Dlussky, Soyunov & Zabelin, 1990: 161 (Turkmenistan species key); Morisita, Kubota, et al. 1991: 16 (Japan species key); Atanasov & Dlussky, 1992: 201 (Bulgaria species key); Arakelian, 1994: 78 (Armenia species key); Bolton, 1994: 51 (synoptic classification); Bolton, 1995a: 1052 (census); Bolton, 1995b: 334 (catalogue); Wu, J. & Wang, 1995: 130 (China species key); Radchenko, 1996a: 178 (Central & Southern Palaearctic species key); Collingwood & Agosti, 1996: 362 (Saudi Arabia species key); Seifert, 1996b: 178 (Central Europe species key); Collingwood & Prince, 1998: 22 (Portugal species key); Shattuck, 1999: 106 (Australia synopsis); Seifert, 2007: 158 (North and Central Europe species key); Terayama, 2009: 213 (Taiwan species key); Heterick, 2009: 106 (south-western Australia species key); Dlussky, 2010: 72 (Eocene European amber species key).
Formicinae Camponotini Camponotus TRUE TRUE FALSEExtant: 1,058 species, 495 subspecies
Fossil: 31 species
Camponotus Mayr, 1861: 35. Type-species: Formica ligniperda, by subsequent designation of Bingham, 1903: 347.
Taxonomic history
[Type-species not Formica herculeana, unjustified subsequent designation by Forel, 1914a: 259, repeated in Arnold, 1922: 612.]
Camponotus in Formicinae: Mayr, 1861: 35 [Formicidae]; Mayr, 1862: 652 (in key) [Formicidae]; Mayr, 1865: 6 [Formicidae]; Mayr, 1868c: 26 [Formicidae].
Camponotus in Camponotinae: Forel, 1878: 367 [Camponotidae]; Emery & Forel, 1879a: 447 [Camponotidae]; Forel, 1892j: 223 [Camponotidae]; Dalla Torre, 1893: 171.
Camponotus in Camponotinae, Camponotini: Forel, 1886f: 141; Forel, 1893a: 165; Forel, 1895b: 101; Emery, 1895j: 772; Forel, 1899c: 130; Ashmead, 1905b: 384; Wheeler, W.M. 1910g: 144; Forel, 1912i: 89; Wheeler, W.M. 1915h: 138; Forel, 1917: 250; Arnold, 1922: 610.
Camponotus in Formicinae, Camponotini: Bondroit, 1918: 65; Wheeler, W.M. 1922a: 703; Emery, 1925b: 59; all subsequent authors except the following.
Camponotus in Formicinae, Formica genus group: Agosti, 1991: 295.
Camponotus as subgenus of Formica: Smith, F. 1871a: 306.
Camponotus as genus: all authors except the entry above.
Junior synonym of CAMPONOTUS
†Paleosminthurus Pierce & Gibron, 1962: 146. Type-species: †Paleosminthurus juliae, by monotypy.
Taxonomic history
†Paleosminthurus in Collembola, family †Paleosminthuridae: Pierce & Gibron, 1962: 146.
†Paleosminthurus incertae sedis in Formicidae: Najt, 1987: 152.
†Paleosminthurus incertae sedis in Formicinae: Hölldobler & Wilson, 1990: 18.
†Paleosminthurus in †Paleosmithurinae: Bolton, 1994: 187 [provisional subfamily status]; Bolton, 1995b: 14, 311; Bolton, 2003: 76, 263.
†Paleosminthurus as junior synonym of Camponotus: Snelling (pers. comm., 2004). [Synonymy remained unpublished at his death in 2008.]
†Shanwangella Zhang, 1989: 307. Type-species: †Shanwangella palaeoptera, by original designation.
Taxonomic history
†Shanwangella as junior synonym of Camponotus: Hong & Wu, 2000: 20.
Subgenera of CAMPONOTUS include the nominal plus the following.
All the subgenera were given as provisional junior synonyms of Camponotus by Brown, 1973b: 179 - 185. Brown's list was repeated in Hölldobler & Wilson, 1990: 18 but with all subgenera listed as junior synonyms; subgeneric status was reinstated in Bolton, 1994: 50. These consecutive entries are not repeated individually below.
Subgenus CAMPONOTUS (COLOBOPSIS)
Colobopsis Mayr, 1861: 38. Type-species: Formica truncata, by subsequent designation of Bingham, 1903: 342.
Taxonomic history
Colobopsis in Formicinae: Mayr, 1861: 38 [Formicidae]; Mayr, 1862: 652 (in key) [Formicidae]; Mayr, 1865: 7 [Formicidae].
Colobopsis in Camponotinae: Forel, 1878: 368 [Camponotidae]; Emery & Forel, 1897: 449 [Camponotidae].
Colobopsis in Camponotinae, Camponotini: Forel, 1886f: 193; Emery, 1895j: 772; Ashmead, 1905b: 384.
Colobopsis in Formicinae, Camponotini: Bondroit, 1918: 65; Emery, 1925b: 144; all subsequent authors.
Colobopsis as genus: Mayr, 1861: 38; Mayr, 1862: 652; Mayr, 1865: 7; Mayr, 1870b: 940; Forel, 1878: 368; Forel, 1879a: 125; Emery & Forel, 1879a: 449; André, 1882b: 159; Bingham, 1903: 342; Forel, 1886f: 193; Ashmead, 1905b: 384; Bondroit, 1918: 66; Jaffe, 1993: 14.
Colobopsis as subgenus of Camponotus: Emery, 1889b: 517; Dalla Torre, 1893: 219; Forel, 1893a: 165; Forel, 1893b: 435; Emery, 1895j: 772; Wheeler, W.M. 1904b: 139; Wheeler, W.M. 1910g: 144; Forel, 1912i: 90; Forel, 1914a: 263; Forel, 1917: 251; Emery, 1920b: 247; Arnold, 1922: 613; Wheeler, W.M. 1922a: 708; Emery, 1925b: 144; Donisthorpe, 1943f: 634; Creighton, 1950a: 390; Chapman & Capco, 1951: 222; Kempf, 1972a: 42; Smith, D.R., 1979: 1433; Bolton, 1994: 50.
Junior synonyms of CAMPONOTUS (COLOBOPSIS)
Condylomyrma Santschi, 1928c: 72 [as subgenus of Camponotus]. Type-species: Camponotus (Condylomyrma) bryani, by monotypy.
Taxonomic history
Condylomyrma as subgenus of Camponotus: Santschi, 1928c: 72.
Condylomyrma as junior synonym of Colobopsis: Wheeler, W.M. 1934e: 422 (in text).
[Campylomyrma Wheeler, W.M. 1934e: 421 (in text), incorrect subsequent spelling.]
Dolophra Wu, J. & Wang, 1994: 35. Type-species: Dolophra politae, by original designation.
Taxonomic history
Dolophra as genus: Wu, J. & Wang, 1994: 35; Wu, J. & Wang, 1995: 158.
Dolophra as junior synonym of Camponotus: Bolton, 1995b: 27.
Dolophra as junior synonym of Camponotus (Colobopsis): Bolton, 2003: 113, 268.
Subgenus CAMPONOTUS (MAYRIA)
Mayria Forel, 1878: 369. Type-species: Mayria madagascarensis (junior secondary homonym in Camponotus, replaced by Camponotus repens), by monotypy.
Taxonomic history
Mayria in Camponotinae: Forel, 1878: 369 [Camponotidae]; Dalla Torre, 1893: 219.
Mayria in Camponotinae, Camponotini: Forel, 1893a: 165; Emery, 1895j: 772; Ashmead, 1905b: 384; Wheeler, W.M. 1910g: 144.
Mayria as subgenus of Camponotus: Forel, 1894e: 227; Forel, 1914a: 262; Forel, 1917: 251; Wheeler, W.M. 1922a: 706; Emery, 1925b: 121; all subsequent authors.
Junior synonym of CAMPONOTUS (MAYRIA)
Myrmosaga Forel, 1912i: 92 [as subgenus of Camponotus]. Type-species: Camponotus kelleri, by subsequent designation of Wheeler, W.M. 1913a: 81.
Taxonomic history
[Type-species not Camponotus quadrimaculatus, unjustified subsequent designation by Forel, 1914a: 260; repeated in Arnold, 1922: 613.]
Myrmosaga as subgenus of Camponotus Forel, 1912i: 92; Forel, 1914a: 260; Forel, 1917: 250; Emery, 1920b: 241; Arnold, 1922: 613; Wheeler, W.M. 1922a: 706; all subsequent authors.
Myrmosaga as junior synonym of Mayria: Emery, 1925b: 121.
Subgenus CAMPONOTUS (RHINOMYRMEX)
Rhinomyrmex Forel, 1886f: 192. Type-species: Rhinomyrmex klaesii, by monotypy.
Taxonomic history
Rhinomyrmex in Camponotinae: Forel, 1886f: 192 [Camponotii]; Dalla Torre, 1893: 219.
Rhinomyrmex in Camponotinae, Camponotini: Forel, 1893a: 165; Emery, 1895j: 772; Ashmead, 1905b: 384; Wheeler, W.M. 1910g: 144.
Rhinomyrmex as genus: Forel, 1886f: 192; Forel, 1893a: 165; Dalla Torre, 1893: 219; Emery, 1895j: 772; Wheeler, W.M. 1910g: 144.
Rhinomyrmex as subgenus of Camponotus: Emery, 1896d: 374; Forel, 1914a: 264; Forel, 1917: 251; Emery, 1920b: 247; Wheeler, W.M. 1922a: 708; Emery, 1925b: 142; all subsequent authors.
Subgenus CAMPONOTUS (DENDROMYRMEX)
Dendromyrmex Emery, 1895j: 772. Type-species: Formica chartifex, by subsequent designation of Wheeler, W.M. 1911f: 161.
Taxonomic history
Dendromyrmex in Camponotinae, Camponotini: Emery, 1895j: 772; Ashmead, 1905b: 384; Forel, 1912i: 89; Wheeler, W.M. 1910g: 144; Forel, 1917: 250.
Dendromyrmex in Formicinae, Camponotini: Wheeler, W.M. 1922a: 700; Emery, 1925b: 172; all subsequent authors except the following.
Dendromyrmex in Formicinae, Formica genus group: Agosti, 1991: 295.
Dendromyrmex as subgenus of Camponotus: Fernández, 2002: 51.
Subgenus CAMPONOTUS (DINOMYRMEX)
Dinomyrmex Ashmead, 1905b: 384. Type-species: Formica gigas, by original designation.
Taxonomic history
Dinomyrmex in Camponotinae, Camponotini: Ashmead, 1905b: 384.
Dinomyrmex as genus: Ashmead, 1905b: 384.
Dinomyrmex as subgenus of Camponotus: Forel, 1913b: 350; Forel, 1914a: 259; Forel, 1917: 250; Emery, 1920b: 236; Arnold, 1922: 612; Wheeler, W.M. 1922a: 704; Emery, 1925b: 69; all subsequent authors.
Junior synonym of CAMPONOTUS (DINOMYRMEX)
Myrmogigas Forel, 1912i: 91 [as subgenus of Camponotus]. Type-species: Formica gigas, by subsequent designation of Wheeler, W.M. 1913a: 81.
Taxonomic history
Myrmogigas as subgenus of Camponotus: Forel, 1912i: 91.
Myrmogigas as junior synonym of Dinomyrmex: Forel, 1913b: 350 (in text); Forel, 1914a: 259; Wheeler, W.M. 1922a: 704. [Dinomyrmex and Myrmogigas share the same type-species, synonymy is therefore absolute.]
Subgenus CAMPONOTUS (TANAEMYRMEX)
Tanaemyrmex Ashmead, 1905b: 384. Type-species: Formica longipes (junior primary homonym in Formica, replaced by Camponotus etiolipes), by original designation.
Taxonomic history
Tanaemyrmex in Camponotinae, Camponotini: Ashmead, 1905b: 384.
Tanaemyrmex as genus: Ashmead, 1905b: 384.
Tanaemyrmex as subgenus of Camponotus: Emery, 1925b: 75; all subsequent authors.
Junior synonym of CAMPONOTUS (TANAEMYRMEX)
Myrmoturba Forel, 1912i: 91 [as subgenus of Camponotus]. Type-species: Formica maculata, by subsequent designation of Wheeler, W.M. 1913a: 82.
Taxonomic history
Myrmoturba as subgenus of Camponotus: Forel, 1912i: 91; Forel, 1914a: 259; Forel, 1917: 250; Emery, 1920b: 235; Arnold, 1922: 612; Wheeler, W.M. 1922a: 704.
Myrmoturba as junior synonym of Tanaemyrmex: Emery, 1925b: 75; all subsequent authors.
Subgenus CAMPONOTUS (ORTHONOTOMYRMEX)
Orthonotomyrmex Ashmead, 1906: 31.
Taxonomic history
[Replacement name for Orthonotus Ashmead, 1905b: 384; junior homonym of Orthonotus Westwood, in Stephens, 1829: 344 (Hemiptera). Type-species not Formica lateralis, unjustified subsequent designation by Forel, 1914a: 264; repeated in Arnold, 1922: 613.]
Orthonotomyrmex as subgenus of Camponotus: Forel, 1913i: 350 (in text); Forel, 1914a: 264; Forel, 1917: 251; Emery, 1920b: 244; Arnold, 1922: 613; Wheeler, W.M. 1922a: 707; Emery, 1925b: 124; all subsequent authors.
Homonym replaced by ORTHONOTOMYRMEX
Orthonotus Ashmead, 1905b: 384. Type-species: Formica sericea, by original designation.
Taxonomic history
[Junior homonym of Orthonotus Westwood, in Stephens, 1829: 344 (Hemiptera).]
Orthonotus in Camponotinae, Camponotini: Ashmead, 1905b: 384.
Subgenus CAMPONOTUS (MYRMAMBLYS)
Myrmamblys Forel, 1912i: 90 [as subgenus of Camponotus]. Type-species: Camponotus reticulatus, by subsequent designation of Wheeler, W.M. 1913a: 80.
Taxonomic history
[Type-species not Camponotus fastigatus, unjustified subsequent designation by Forel, 1914a: 263; repeated in Arnold, 1922: 613.]
Myrmamblys as subgenus of Camponotus: Forel, 1912i: 90; Forel, 1914a: 263; Forel, 1917: 251; Emery, 1920b: 248; Arnold, 1922: 613; Wheeler, W.M. 1922a: 708; Emery, 1925b: 136; Santschi, 1926c: 601; all subsequent authors.
Subgenus CAMPONOTUS (MYRMOTHRIX)
Myrmothrix Forel, 1912i: 91 [as subgenus of Camponotus]. Type-species: Formica abdominalis (junior primary homonym; Formica atriceps is first available replacement name), by subsequent designation of Wheeler, W.M. 1913a: 81.
Taxonomic history
[Type-species not Formica rufipes, unjustified subsequent designation by Forel, 1914a: 260.]
Myrmothrix as subgenus of Camponotus: Forel, 1912i: 91; Forel, 1914a: 260; Forel, 1917: 250; Emery, 1920b: 236; Emery, 1925b: 107; all subsequent authors.
Subgenus CAMPONOTUS (MYRMOSERICUS)
Myrmosericus Forel, 1912i: 91 [as subgenus of Camponotus]. Type-species: Formica rufoglauca, by subsequent designation of Wheeler, W.M. 1913a: 81.
Taxonomic history
Myrmosericus as subgenus of Camponotus: Forel, 1912i: 91; Forel, 1914a: 259; Forel, 1917: 250; Emery, 1920b: 236; Arnold, 1922: 613; Wheeler, W.M. 1922a: 704; Emery, 1925b: 104; all subsequent authors.
[See note under Myrmosaulus.]
Subgenus CAMPONOTUS (MYRMOPHYMA)
Myrmophyma Forel, 1912i: 91 [as subgenus of Camponotus]. Type-species: Camponotus capito, by subsequent designation of Wheeler, W.M. 1913a: 81.
Taxonomic history
[Type-species not Camponotus quadrisectus, unjustified subsequent designation by Forel, 1914a: 261; repeated in Arnold, 1922: 612.]
Myrmophyma as subgenus of Camponotus: Forel, 1912i: 91; Forel, 1914a: 261; Forel, 1917: 250; Emery, 1920b: 239; Arnold, 1922: 612; Wheeler, W.M. 1922a: 706; Emery, 1925b: 109; all subsequent authors.
Junior synonym of CAMPONOTUS (MYRMOPHYMA)
Myrmocamelus Forel, 1914a: 261 [as subgenus of Camponotus]. Type-species: Formica ephippium, by original designation.
Taxonomic history
[Type-species not Camponotus gambeyi, unjustified subsequent designation by Forel, 1922: 101. Myrmocamelus also described as new by Forel, 1915b: 102.]
Myrmocamelus as subgenus of Camponotus: Forel, 1914a: 261; Forel, 1915b: 102; Forel, 1917: 250; Forel, 1922: 101.
Myrmocamelus as junior synonym of Myrmophyma: Emery, 1920b: 257; Wheeler, W.M. 1921a: 18; Emery, 1925b: 109.
Subgenus CAMPONOTUS (MYRMOTREMA)
Myrmotrema Forel, 1912i: 91 [as subgenus of Camponotus]. Type-species: Camponotus foraminosus, by subsequent designation of Wheeler, W.M. 1913a: 81.
Taxonomic history
Myrmotrema as subgenus of Camponotus: Forel, 1912i: 91; Forel, 1914a: 262; Forel, 1917: 250; Emery, 1920b: 245; Arnold, 1922: 613; Wheeler, W.M. 1922a: 707; Emery, 1925b: 130; all subsequent authors.
Subgenus CAMPONOTUS (MYRMOBRACHYS)
Myrmobrachys Forel, 1912i: 91 [as subgenus of Camponotus]. Type-species: Formica senex, by subsequent designation of Wheeler, W.M. 1913a: 81.
Taxonomic history
Myrmobrachys as subgenus of Camponotus: Forel, 1912i: 91; Forel, 1914a: 262; Forel, 1917: 251; Emery, 1920b: 251; Wheeler, W.M. 1922a: 709; Emery, 1925b: 161; all subsequent authors.
Subgenus CAMPONOTUS (MYRMENTOMA)
Myrmentoma Forel, 1912i: 92 [as subgenus of Camponotus]. Type-species: Formica lateralis, by subsequent designation of Wheeler, W.M. 1913a: 80.
Taxonomic history
Myrmentoma as junior synonym of Orthonotomyrmex: Forel, 1913b: 350 (in text); Forel, 1914a: 264; Wheeler, W.M. 1922a: 970.
Myrmentoma as subgenus of Camponotus: Forel, 1912i: 92; Emery, 1920b: 243; Wheeler, W.M. 1922a: 707; Emery, 1925a: 62; Emery, 1925b: 116; all subsequent authors.
Subgenus CAMPONOTUS (MYRMOSPHINCTA)
Myrmosphincta Forel, 1912i: 92 [as subgenus of Camponotus]. Type-species: Formica sexguttata, by subsequent designation of Wheeler, W.M. 1913a: 81.
Taxonomic history
[Type-species not Formica cinerascens, unjustified subsequent designation by Forel, 1914a: 265; repeated in Arnold, 1922: 614.]
Myrmosphincta as subgenus of Camponotus: Forel, 1912i: 92; Forel, 1914a: 265; Forel, 1917: 251; Emery, 1920b: 238; Arnold, 1922: 614; Wheeler, W.M. 1922a: 708; Emery, 1925b: 151; all subsequent authors.
Subgenus CAMPONOTUS (MYRMOTARSUS)
Myrmotarsus Forel, 1912i: 92 [as subgenus of Camponotus]. Type-species: Formica mistura, by subsequent designation of Wheeler, W.M. 1913a: 81.
Taxonomic history
[Type-species not Formica irritabilis, unjustified subsequent designation by Forel, 1914a: 260.]
Myrmotarsus as subgenus of Camponotus: Forel, 1912i: 92; Forel, 1914a: 260; Forel, 1917: 250; Emery, 1920b: 238; Wheeler, W.M. 1922a: 705; Emery, 1925b: 134; all subsequent authors.
Subgenus CAMPONOTUS (MYRMEPOMIS)
Myrmepomis Forel, 1912i: 92 [as subgenus of Camponotus]. Type-species: Formica sericeiventris, by subsequent designation of Wheeler, W.M. 1913a: 81.
Taxonomic history
[Type-species not Formica fulvopilosa, unjustified subsequent designation by Forel, 1914a: 265; repeated in Arnold, 1922: 614.]
Myrmepomis as subgenus of Camponotus: Forel, 1912i: 92; Forel, 1914a: 265; Forel, 1917: 251; Emery, 1920b: 245; Arnold, 1922: 614; Wheeler, W.M. 1922a: 705; Emery, 1925b: 170; all subsequent authors.
Junior synonym of CAMPONOTUS (MYRMEPOMIS)
Myrmolophus Emery, 1920b: 237 [as subgenus of Camponotus]. Type-species: Formica sericeiventris, by original designation.
Taxonomic history
Myrmolophus as subgenus of Camponotus: Emery, 1920b: 237.
Myrmolophus as junior synonym of Myrmepomis: Wheeler, W.M. 1921a: 17; Wheeler, W.M. 1922a: 705; Emery, 1925b: 170. [Myrmepomis and Myrmolophus share the same type-species, synonymy is therefore absolute.]
Subgenus CAMPONOTUS (MYRMOGONIA)
Myrmogonia Forel, 1912i: 92 [as subgenus of Camponotus]. Type-species: Camponotus laminatus, by subsequent designation of Wheeler, W.M. 1913a: 81.
Taxonomic history
Myrmogonia as subgenus of Camponotus: Forel, 1912i: 92; Forel, 1914a: 261; Forel, 1917: 250; Emery, 1920b: 240; Wheeler, W.M. 1922a: 706; Emery, 1925b: 143; all subsequent authors.
Subgenus CAMPONOTUS (MYRMEURYNOTA)
Myrmeurynota Forel, 1912i: 92 [as subgenus of Camponotus]. Type-species: Camponotus eurynotus, by subsequent designation of Wheeler, W.M. 1913a: 81.
Taxonomic history
[Type-species not Camponotus gilviventris, unjustified subsequent designation by Forel, 1914a: 266.]
Myrmeurynota as subgenus of Camponotus: Forel, 1912i: 92; Forel, 1914a: 266; Forel, 1917: 251; Emery, 1920b: 251; Wheeler, W.M. 1922a: 710; Emery, 1925b: 167; all subsequent authors.
Subgenus CAMPONOTUS (MYRMOPSAMMA)
Myrmopsamma Forel, 1914a: 261 [as subgenus of Camponotus]. Type-species: Camponotus mystaceus, by original designation.
Taxonomic history
Myrmopsamma as subgenus of Camponotus: Forel, 1914a: 261; Forel, 1917: 250; Emery, 1920b: 246; Arnold, 1922: 612; Wheeler, W.M. 1922a: 708; Emery, 1925b: 70; all subsequent authors.
Subgenus CAMPONOTUS (MYRMOMALIS)
Myrmomalis Forel, 1914a: 263 [as subgenus of Camponotus]. Type-species: Camponotus depressus, by original designation.
Taxonomic history
Myrmomalis as subgenus of Camponotus: Forel, 1914a: 263; Forel, 1917: 251; Emery, 1920b: 251; Wheeler, W.M. 1922a: 710; Emery, 1925b: 169; all subsequent authors.
Subgenus CAMPONOTUS (MYRMOPLATYS)
Myrmoplatys Forel, 1916: 460 [as subgenus of Camponotus]. Type-species: Camponotus korthalsiae, by original designation.
Taxonomic history
Myrmoplatys as genus: Wheeler, W.M. 1936e: 219 (in text); Donisthorpe, 1943f: 670.
Myrmoplatys as subgenus of Camponotus: Forel, 1916: 460; Forel, 1917: 251; Emery, 1920b: 238; Wheeler, W.M. 1922a: 705; Emery, 1925b: 135; Bolton, 1994: 50.
Subgenus CAMPONOTUS (MYRMAPHAENUS)
Myrmaphaenus Emery, 1920b: 237 [as subgenus of Camponotus]. Type-species: Camponotus leydigi, by original designation.
Taxonomic history
Myrmaphaenus as subgenus of Camponotus: Emery, 1920b: 237; Wheeler, W.M. 1922a: 705; Emery, 1925b: 152; all subsequent authors.
Junior synonyms of CAMPONOTUS (MYRMAPHAENUS)
Paracolobopsis Emery, 1920b: 249 [as subgenus of Camponotus]. Type-species: Camponotus salvini, by original designation.
Taxonomic history
Paracolobopsis as subgenus of Camponotus: Emery, 1920b: 249; Wheeler, W.M. 1922a: 709.
Paracolobopsis as junior synonym of Myrmaphaenus: Emery, 1925b: 152; Kempf, 1972a: 44.
Neomyrmamblys Wheeler, W.M. 1921a: 19 [as subgenus of Camponotus]. Type-species: Camponotus fastigatus, by subsequent designation of Santschi, 1921f: 311.
Taxonomic history
Neomyrmamblys as subgenus of Camponotus: Wheeler, W.M. 1921a: 19; Wheeler, W.M. 1922a: 708.
Neomyrmamblys as junior synonym of Myrmaphaenus: Emery, 1925b: 152; Kempf, 1972a: 44.
Subgenus CAMPONOTUS (MYRMONESITES)
Myrmonesites Emery, 1920b: 242 [as subgenus of Camponotus]. Type-species: Camponotus putatus, by original designation.
Taxonomic history
Myrmonesites as subgenus of Camponotus: Emery, 1920b: 242; Wheeler, W.M. 1922a: 706; Emery, 1925b: 123.
[Myrmensites Donisthorpe, 1943f: 666, incorrect subsequent spelling.]
Subgenus CAMPONOTUS (MYRMOPYTIA)
Myrmopytia Emery, 1920b: 243 [as subgenus of Camponotus]. Type-species: Camponotus imitator, by original designation.
Taxonomic history
Myrmopytia as subgenus of Camponotus: Emery, 1920b: 243; Wheeler, W.M. 1922a: 707; Emery, 1925b: 114; all subsequent authors.
Subgenus CAMPONOTUS (MYRMOTEMNUS)
Myrmotemnus Emery, 1920b: 246 [as subgenus of Camponotus]. Type-species: Camponotus moeschi, by original designation.
Taxonomic history
Myrmotemnus as junior synonym of Myrmamblys: Wheeler, W.M. 1921a: 19; Wheeler, W.M. 1922a: 708; Emery, 1925b: 137; Donisthorpe, 1943f: 671 (anachronism).
Myrmotemnus as subgenus of Camponotus: Emery, 1920b: 246; Santschi, 1926c: 601.
Subgenus CAMPONOTUS (PSEUDOCOLOBOPSIS)
Pseudocolobopsis Emery, 1920b: 249 [as subgenus of Camponotus]. Type-species: Camponotus macrocephalus, by original designation.
Taxonomic history
Pseudocolobopsis as subgenus of Camponotus: Emery, 1920b: 249; Wheeler, W.M. 1922a: 709; Emery, 1925b: 157; all subsequent authors.
Subgenus CAMPONOTUS (MYRMOSTENUS)
Myrmostenus Emery, 1920b: 250 [as subgenus of Camponotus]. Type-species: Camponotus mirabilis, by original designation.
Taxonomic history
Myrmostenus as subgenus of Camponotus: Emery, 1920b: 250; Wheeler, W.M. 1922a: 709; Emery, 1925b: 161; all subsequent authors.
Subgenus CAMPONOTUS (HYPERCOLOBOPSIS)
Hypercolobopsis Emery, 1920b: 250 [as subgenus of Camponotus]. Type-species: Colobopsis paradoxa, by original designation.
Taxonomic history
Hypercolobopsis as subgenus of Camponotus: Emery, 1920b: 250; Wheeler, W.M. 1922a: 709; Emery, 1925b: 160; all subsequent authors.
Junior synonym of CAMPONOTUS (HYPERCOLOBOPSIS)
Neocolobopsis Borgmeier, 1928b: 65 [as subgenus of Camponotus]. Type-species: Camponotus (Neocolobopsis) scrobifer (junior synonym of Camponotus coriolanus), by original designation.
Taxonomic history
Neocolobopsis as subgenus of Camponotus Borgmeier, 1928b: 65.
Neocolobopsis as junior synonym of Hypercolobopsis: Kempf, 1968b: 411; Kempf, 1972a: 43.
Subgenus CAMPONOTUS (MYRMOPIROMIS)
Myrmopiromis Wheeler, W.M. 1921a: 17 [as subgenus of Camponotus]. Type-species: Formica fulvopilosa, by subsequent designation of Wheeler, W.M. 1922a: 707.
Taxonomic history
Myrmopiromis as subgenus of Camponotus: Wheeler, W.M. 1921a: 17; Wheeler, W.M. 1922a: 707; Emery, 1925b: 1127.
Subgenus CAMPONOTUS (MYRMOSAULUS)
Myrmosaulus Wheeler, W.M. 1921a: 18 [as subgenus of Camponotus]. Type-species: Formica cinerascens, by subsequent designation of Wheeler, W.M. 1922a: 705.
Taxonomic history
[Type-species not Formica singularis, unjustified subsequent designation by Emery, 1925b: 113, repeated in Donisthorpe, 1943f: 670. Note that Donisthorpe, 1932c: 445, states that C. cinerascens belongs in subgenus Myrmosericus. If so then Myrmosaulus would fall as a junior synonym of Myrmosericus.]
Myrmosaulus as subgenus of Camponotus: Wheeler, W.M. 1921a: 18; Wheeler, W.M. 1922a: 705; Emery, 1925b: 112; all subsequent authors.
Subgenus CAMPONOTUS (MYRMOCLADOECUS)
Myrmocladoecus Wheeler, W.M. 1921a: 19 [as subgenus of Camponotus]. Type-species: Camponotus latangulus, by subsequent designation of Wheeler, W.M. 1922a: 709.
Taxonomic history
Myrmocladoecus as subgenus of Camponotus: Wheeler, W.M. 1921a: 19; Wheeler, W.M. 1922a: 709; Emery, 1925b: 166; all subsequent authors.
Subgenus CAMPONOTUS (MANNIELLA)
Manniella Wheeler, W.M. 1921a: 19 [as subgenus of Camponotus]. Type-species: Camponotus sphaericus, by original designation.
Taxonomic history
Manniella subgenus of Camponotus: Wheeler, W.M. 1921a: 19; Wheeler, W.M. 1922a: 710; Emery, 1925b: 159; all subsequent authors.
Subgenus CAMPONOTUS (MYRMISOLEPIS)
Myrmisolepis Santschi, 1921f: 310 [as subgenus of Camponotus]. Type-species: Camponotus epinotalis, by original designation.
Taxonomic history
Myrmisolepis as subgenus of Camponotus: Santschi, 1921f: 310; Emery, 1925b: 133; all subsequent authors.
Subgenus CAMPONOTUS (MYRMOPELTA)
Myrmopelta Santschi, 1921f: 310 [as subgenus of Camponotus]. Type-species: Camponotus arminius, by original designation.
Taxonomic history
Myrmopelta as junior synonym of Myrmamblys: Emery, 1925b: 137.
Myrmopelta as subgenus of Camponotus: Santschi, 1921f: 310; Wheeler, W.M. 1922a: 704 (footnote); Santschi, 1926a: 16; Santschi, 1926c: 603; all subsequent authors.
Subgenus CAMPONOTUS (MYRMOPLATYPUS)
Myrmoplatypus Santschi, 1921f: 311 [as subgenus of Camponotus]. Type-species: Camponotus platytarsus, by original designation.
Taxonomic history
Myrmoplatypus as subgenus of Camponotus: Santschi, 1921f: 311; Emery, 1925b: 169; all subsequent authors.
Subgenus CAMPONOTUS (MYRMEPINOTUS)
Myrmepinotus Santschi, 1921f: 312 [as subgenus of Camponotus]. Type-species: Camponotus echinoploides, by original designation.
Taxonomic history
Myrmepinotus as subgenus of Camponotus: Santschi, 1921f: 312; Emery, 1925b: 126; all subsequent authors.
Subgenus CAMPONOTUS (MYRMOXYGENYS)
Myrmoxygenys Emery, 1925b: 70 [as subgenus of Camponotus]. Type-species: Camponotus caesar, by monotypy.
Taxonomic history
Myrmoxygenys as subgenus of Camponotus: Emery, 1925b: 70; all subsequent authors.
Subgenus CAMPONOTUS (KARAVAIEVIA)
Karavaievia Emery, 1925b: 115 [as subgenus of Camponotus]. Type-species: Camponotus exsectus, by original designation.
Taxonomic history
Karavaievia as subgenus of Camponotus: Emery, 1925b: 115; all subsequent authors.
Subgenus CAMPONOTUS (MYRMODIRACHIS)
Myrmodirachis Emery, 1925b: 168 [as subgenus of Camponotus]. Type-species: Camponotus heathi, by original designation.
Taxonomic history
Myrmodirachis as subgenus of Camponotus: Emery, 1925b: 168; all subsequent authors.
Subgenus CAMPONOTUS (MYRMESPERA)
Myrmespera Santschi, 1926b: 247 [as subgenus of Camponotus]. Type-species: Camponotus (Myrmespera) debellator, by subsequent designation of Santschi, 1926c: 604.
Taxonomic history
Myrmespera as subgenus of Camponotus: Santschi, 1926b: 247; Santschi, 1926c: 603; all subsequent authors.
Subgenus CAMPONOTUS (PARAMYRMAMBLYS)
Paramyrmamblys Santschi, 1926c: 604 [as subgenus of Camponotus].Type-species: Camponotus (Myrmamblys) ostiarius, by original designation.
Taxonomic history
Paramyrmamblys as subgenus of Camponotus: Santschi, 1926c: 604; all subsequent authors.
Subgenus CAMPONOTUS (MYRMACRHAPHE)
Myrmacrhaphe Santschi, 1926c: 607 [as subgenus of Camponotus]. Type-species: Camponotus conradti, by original designation.
Taxonomic history
Myrmacrhaphe as subgenus of Camponotus: Santschi, 1926c: 607; all subsequent authors.
Subgenus CAMPONOTUS (THLIPSEPINOTUS)
Thlipsepinotus Santschi, 1928e: 483 [as subgenus of Camponotus]. Type-species: Camponotus claripes, by original designation.
Taxonomic history
Thlipsepinotus as subgenus of Camponotus: Santschi, 1928e: 483; all subsequent authors.
Subgenus CAMPONOTUS (MYRMOPALPELLA)
Myrmopalpella Stärcke, 1934: 30 [as subgenus of Camponotus]. Type-species: Camponotus megalonyx, by monotypy.
Taxonomic history
Myrmopalpella as genus: Wheeler, W.M. 1936e: 219 (in text); Donisthorpe, 1943f: 669.
Myrmopalpella as subgenus of Camponotus: Bolton, 1995b: 37.
Genus Camponotus references
Mayr, 1861: 35 (Europe species key); Roger, 1863b: 1, 9 (Camponotus, Colobopsis catalogues); Mayr, 1863: 397, 403 (Camponotus, Colobopsis catalogues); Mayr, 1865: 6, 7 (Camponotus, Colobopsis diagnosis); Mayr, 1867a: 67 (Colobopsis diagnois); Mayr, 1868c: 27 (†Baltic Amber species key); Mayr, 1870a: 373 (Colombia + Panama (= New Grenada) species key); Mayr, 1870b: 941 (Colobopsis species key); André, 1874: 175 (Europe species key); Forel, 1874: 38 (Switzerland species key); Mayr, 1876: 58 (Australia species key); Mayr, 1878: 869 (all Dendromyrmex species key (as part of Camponotus)); Forel, 1878: 367, 368 (Camponotus, Colobopsis diagnoses); André, 1882a: 138 (Europe & Algeria species key); André, 1882b: 159 (Europe & Algeria Colobopsis species key); Cresson, 1887: 255 (U.S.A. Camponotus, Colobopsis catalogue); Provancher, 1887: 228 (Canada species key); Nasonov, 1889: 57 (Russia species key); Forel, 1891b: 71, 216 (Madagascar species key); Forel, 1892j: 223 (India & Sri Lanka species key); Dalla Torre, 1893: 219 (Rhinomyrmex, Mayria, Camponotus catalogues); Emery, 1893i: 667 (North America species key); Forel, 1893b: 435 (India & Sri Lanka C. (Colobopsis) species key); Emery, 1895j: 772 (synoptic classification); Emery, 1896d: 370, 378 (species groups, catalogue); Emery, 1898a: 225 (additions to 1896a catalogue); Emery, 1903 (South America species key); Bingham, 1903: 343, 348 (India, Sri Lanka & Burma Colobopsis, Camponotus species keys); Ruzsky, 1905b: 185 (Russian Empire species key); Emery, 1908a: 183 (Palaearctic C. herculeanus & C. maculatus groups species key); Bondroit, 1910: 486 (Belgium species key); Wheeler, W.M. 1910d: 297 (North America species key); Forel, 1914a: 259, 266 (subgenera diagnoses, species listed by subgenus); Stitz, 1914: 94 (Central Europe species key); Santschi, 1915c: 266 (C. (Myrmotrema) species key); Emery, 1916b: 227 (Italy species key); Wheeler, W.M. 1916m: 600 (U.S.A., Connecticut species key); Mann, 1916: 489 (Dendromyrmex, all species key); Bondroit, 1918 (France & Belgium species key); Emery, 1920b: 235 (subgenera, diagoses); Emery, 1920c: 8 (Old World C. maculatus group, checklist & key); Wheeler, W.M. 1921a: 16 (subgenera, notes); Santschi, 1921f: 310 (subgenera, notes); Mann, 1921: 489, 499 (Fiji Is C. (Myrmogonia) & C. (Colobopsis) species keys); Wheeler, W.M. 1922a: 704 (subgenera synopsis); Wheeler, W.M. 1922a: 232, 948, 1039 (diagnosis, Afrotropical, Malagasy catalogues); Arnold, 1922: 610, 612, 619, 662, 673 (diagnosis, South Africa subgenera synopsis, South Africa C. (Tanaemyrmex), C. (Myrmophyma), C. (Myrmopsamma) species keys); Arnold, 1924: 683, 700, 709, 724, 735 (South Africa C. (Myrmosericus), C. (Myrmamblys), C. (Orthonotomyrmex), C. (Myrmotrema), C. (Myrmepomis) species keys); Emery, 1925a: 62 (Palaearctic C. lateralis group, key); Emery, 1925b: 59 (diagnosis, subgenera key, catalogue); Emery, 1925b: 69 (C. (Dinomyrmex) diagnosis, catalogue); Emery, 1925b: 70 (C. (Myrmoxygenys) & C. (Myrmopsamma) diagnoses, catalogues); Emery, 1925b: 71 (C. (Camponotus) diagnosis, catalogue); Emery, 1925b: 75 (C. (Tanaemyrmex) diagnosis, catalogue); Emery, 1925b: 104 (C. (Myrmosericus) diagnosis, catalogue); Emery, 1925b: 107 (C. (Myrmothrix) diagnosis, catalogue); Emery, 1925b: 109 (C. (Myrmophyma) diagnosis, catalogue); Emery, 1925b: 112 (C. (Myrmosaulus) diagnosis, catalogue); Emery, 1925b: 114 (C. (Myrmopytia) diagnosis, catalogue); Emery, 1925b: 115 (C. (Karavaievia) diagnosis, catalogue); Emery, 1925b: 116 (C. (Myrmentoma) diagnosis, catalogue); Emery, 1925b: 121 (C. (Mayria) diagnosis, catalogue); Emery, 1925b: 123 (C. (Myrmonesites) diagnosis, catalogue); Emery, 1925b: 124 (C. (Orthonotomyrmex) diagnosis, catalogue); Emery, 1925b: 126 (C. (Myrmepinotus) diagnosis, catalogue); Emery, 1925b: 127 (C. (Myrmopiromis) diagnosis, catalogue); Emery, 1925b: 130 (C. (Myrmotrema) diagnosis, catalogue); Emery, 1925b: 133 (C. (Myrmisolepis) diagnosis, catalogue); Emery, 1925b: 134 (C. (Myrmotarsus) diagnosis, catalogue); Emery, 1925b: 135 (C. (Myrmoplatys) diagnosis, catalogue); Emery, 1925b: 136 (C. (Myrmamblys) diagnosis, catalogue); Emery, 1925b: 142 (C. (Rhinomyrmex) diagnosis, catalogue); Emery, 1925b: 143 (C. (Myrmogonia) diagnosis, catalogue); Emery, 1925b: 144 (C. (Colobopsis) diagnosis, catalogue); Emery, 1925b: 151 (C. (Myrmosphincta) diagnosis, catalogue); Emery, 1925b: 152 (C. (Myrmaphaenus) diagnosis, catalogue); Emery, 1925b: 157 (C. (Pseudocolobopsis) diagnosis, catalogue); Emery, 1925b: 159 (C. (Manniella) diagnosis, catalogue); Emery, 1925b: 160 (C. (Hypercolobopsis) diagnosis, catalogue); Emery, 1925b: 161 (C. (Myrmostenus) & C. (Myrmobrachys) diagnoses, catalogues); Emery, 1925b: 166 (C. (Myrmocladoecus) diagnosis, catalogue); Emery, 1925b: 167 (C. (Myrmeurynota) diagnosis, catalogue); Emery, 1925b: 168 (C. (Myrmodirachis) diagnosis, catalogue); Emery, 1925b: 169 (C. (Myrmomalis) & C. (Myrmoplatypus) diagnoses, catalogues); Emery, 1925b: 170 (C. (Myrmepomis) diagnosis, catalogue); Emery, 1925b: 172 (Dendromyrmex diagnosis, catalogue); Santschi, 1926a: 16, 21 (C. (Myrmopelta) diagnosis, species key); Karavaiev, 1927c: 275 (Ukraine species key); Arnol'di, 1933b: 601 (Russia species key); Menozzi, 1933b: 81 (Israel species key); Menozzi, 1935c: 330 (Chile species key); Karavaiev, 1936: 175 (Ukraine species key); Finzi, 1936: 190 (Egypt species key); Menozzi, 1939a: 316 (Himalaya & Tibet species key); Stitz, 1939: 236 (Germany species key); Kratochvíl, 1941: 109 (Central Europe species key); Novák & Sadil, 1941: 109 (Central Europe species key); Cole, 1942: 387 (U.S.A., Utah species key); Donisthorpe, 1932g: 248 (C. (Orthonotomyrmex) notes); Holgersen, 1943b: 174 (Norway species key); Holgersen, 1944: 199 (Norway species key); Buren, 1944a: 293 (U.S.A., Iowa species key); Creighton, 1950a: 362 (North America species keys); Chapman & Capco, 1951: 220 (Asia checklist); Yasumatsu & Brown, 1951: 29 (C. herculeanus complex); Kusnezov, 1952d: 192 (Argentina species key); Yasumatsu & Brown, 1957: 45 (C. herculeanus complex); Pisarski, 1961a: 175 (Poland species key); Gregg, 1963: 655 (U.S.A., Colorado species key); Wheeler, G.C. & Wheeler, J. 1963: 163 (U.S.A., North Dakota species key); Arnol'di, 1967: 1817 (former U.S.S.R. C. (Camponotus) species key); Bernard, 1967: 330 (diagnosis, Western Europe species key); Kempf, 1972a: 42, 95 (Neotropical catalogue); Hashmi, 1973: 1 (C. (Myrmothrix) all species revision, key); Alayo, 1974: 27 (Cuba species key); Snelling & Hunt, 1976: 117 (Chile species key); Tarbinsky, 1976: 144 (Kyrghyzstan species key); Boven, 1977: 130 (Belgium species key); Kutter, 1977c: 191 (Switzerland species key); Arnol'di & Dlussky, 1978: 551 (former European U.S.S.R. species key); Collingwood, 1978: 90 (Iberian Peninsula species key); Collingwood, 1979: 87 (Fennoscandia & Denmark species key); Smith, D.R. 1979: 1424 (North America catalogue); Allred, 1982: 445 (U.S.A., Utah species key); Gösswald, 1985: 264 (Germany species key); Collingwood, 1985: 275 (Saudi Arabia species key); Taylor & Brown, D.R. 1985: 109 (Australia catalogue); Dumpert, 1986: 571 (C. (Karavaievia) species key); Wheeler, G.C. & Wheeler, J. 1986g: 60 (U.S.A., Nevada species key); Agosti & Collingwood, 1987b: 282 (Balkans species key); Taylor, 1987a: 10, 16 (Australia, New Caledonia, New Zealand checklists); Snelling, R.R. 1988: 57 (Nearctic C. (Myrmentoma) species key); Wang, C., Xiao & Wu, 1989a: 222 (China species key); Dlussky, Soyunov & Zabelin, 1990: 125 (Turkmenistan species key); Kupyanskaya, 1990: 164 (Far Eastern Russia species key); Brandão, 1991: 332 (Neotropical catalogue); Morisita, Kubota, Onoyama, et al. 1991: 38 (Japan species key); Terayama, 1991: 165 (Japan C. (Paramyrmamblys) species); Atanasov & Dlussky, 1992: 207 (Bulgaria species key); Wang, C. & Wu, 1994: 28 (China species key); Radchenko, 1994b: 116 (South Siberia species key); Arakelian, 1994: 82 (Armenia species key); Bolton, 1994: 50 (synoptic classification); Bolton, 1995a: 1048 (census); Bolton, 1995b: 83, 169 (catalogue); Douwes, 1995: 92 (Sweden species key); Dumpert, Maschwitz, Weissflog et al. 1995: 104 (C. (Karavaievia) species key); Kupyanskaya, 1995: 352 (Far Eastern Russia species key); Wu, J. & Wang, 1995: 168 (China species key); Collingwood & Agosti, 1996: 370 (Saudi Arabia species key); Seifert, 1996b: 170 (Central Europe species key); Radchenko, 1996b: 1195 (Asiatic Palaearctic species key); Cagniant, 1996b: 87 (Morocco species key); McArthur & Adams, 1996: 18 (Australia C. nigriceps group, key); Mackay, 1997: 194 (C. (Myrmostenus) species key); Mackay & Mackay, 1997b: 319 and Snelling, R.R. 2000: 607 (C. montivagus complex, keys); Robertson & Zachariades, 1997: 2 (C. fulvopilosus group, key); Snelling & Torres, 1998: 9 (Puerto Rico & Virgin Is species key); Collingwood & Prince, 1998: 24 (Portugal species key); Terayama, 1999b: 27 (Japan species key); Shattuck, 1999: 91 (Australia synopsis); Zhou, 2001b: 200 (China, Guangxi species key); McArthur & Shattuck, 2001: 27 (Australia C. macrocephalus group, key); Czechowski, Radchenko & Czechowska, 2002: 153 (Poland species key); Fernández, 2002: 58 (C. (Dendromyrmex) diagnosis, all species revision, key); Mackay & Mackay, 2002: 248 (U.S.A., New Mexico species key); Shattuck & McArthur, 2002: 65 (Australia C. wiederkehri group key); Bolton, 2003: 112 (synopsis); Shattuck, 2005: 3 (C. aureopilus group key); Radchenko, 2005b: 188 (North Korea species key); Hansen & Klotz, 2005: 75 (North American “Carpenter Ants” key); Snelling, R.R. 2006: 96 (U.S.A. C. festinatus complex, key); Karmaly & Narendran, 2006: 29 (India species, key); Dumpert, Maschwitz & Weissflog, 2006: 77 (C. (Karavaievia) species, keys); Seifert, 2007: 152 (North and Central Europe species key); McArthur, 2007a: 290 (Australia species key); Shattuck & Janda, 2009: 252 (C. aureopilus group key); Mackay & Delsinne, 2009: 496 (New World C. maculatus complex, key); Terayama, 2009: 216 (Taiwan species key); Heterick, 2009: 60 (south-western Australia species key); Ionescu-Hirsch, 2009: 59 (Israel species key); McArthur, 2010: 18 (south Australia species key).
Formicinae Camponotini Polyrhachis TRUE TRUE FALSEExtant: 614 species, 86 subspecies
Fossil: 1 species
Polyrhachis Smith, F. 1857a: 58. Type-species: Formica bihamata, by original designation.
Taxonomic history
[Polyrhachis Shuckard, in Swainson & Shuckard, 1840: 172, nomen nudum.]
Polyrhachis in Formicidae: Smith, F. 1858b: 58.
Polyrhachis in Formicinae: Mayr, 1862: 652 [Formicidae]; Mayr, 1865: 6 [Formicidae].
Polyrhachis in Camponotinae: Forel, 1878: 367 [Camponotidae]; Dalla Torre, 1893: 257.
Polyrhachis in Camponotinae, Polyrhachidini: Ashmead, 1905b: 384.
Polyrhachis in Camponotinae, Camponotini: Forel, 1886f: 194; Forel, 1893a: 165; Emery, 1895j: 772; Wheeler, W.M. 1910g: 144; Forel, 1912i: 89; Forel, 1917: 251; Arnold, 1924: 741.
Polyrhachis in Formicinae, Camponotini: Wheeler, W.M. 1922a: 701; Emery, 1925b: 175; all subsequent authors except the following.
Polyrhachis in Formicinae, Formica genus group: Agosti, 1991: 295.
Polyrhachis as subgenus of Myrma: Wheeler, W.M. 1911c: 860; Wheeler, W.M. 1911f: 170. [Myrma as subgenus of Polyrhachis: Forel, 1915b: 106; Forel, 1917: 251; Wheeler, W.M. 1922a: 702, 993; Emery, 1925b: 198; all subsequent authors (see note under Myrma).]
[Polyrachis Arnold, 1924: 741, incorrect subsequent spelling.]
Subgenera of POLYRHACHIS include the nominal plus the following.
All the earlier subgenera were given as provisional junior synonyms of Polyrhachis by Brown, 1973b: 178 - 184. Brown's list was repeated in Hölldobler & Wilson, 1990: 19 but with all subgenera listed as junior synonyms; subgeneric status was reinstated in Bolton, 1994: 50. These consecutive entries are not repeated individually below.
Subgenus POLYRHACHIS (MYRMA)
Myrma Billberg, 1820: 104. Type-species: Formica militaris, by subsequent designation of Wheeler, W.M. 1911c: 859.
Taxonomic history
[Although Myrma antedates Polyrhachis, the former has been treated as a subgenus of the latter since Forel, 1917: 251 (see Hung, 1967b: 396 for history). Dorow, Kohout & Taylor, 1997: 236 proposed the precedence of Polyrhachis over Myrma and that precedence was established by the ICZN (Opinion 1919) 1999: 92.]
Polyrhachis as subgenus of Myrma: Wheeler, W.M. 1911c: 860; Wheeler, W.M. 1911f: 170.
Myrma as subgenus of Polyrhachis: Forel, 1915b: 106; Forel, 1917: 251; Wheeler, W.M. 1922a: 702, 993; Emery, 1925b: 198; all subsequent authors, see note above.
Junior synonyms of POLYRHACHIS (MYRMA)
Hoplomyrmus Gerstäcker, 1859: 262. Type-species: Hoplomyrmus schistaceus, by monotypy.
Taxonomic history
[Hoplomyrmus also described as new by Gerstäcker, 1862: 508.]
Hoplomyrmus as junior synonym of Polyrhachis: Roger, 1861b: 174; Roger, 1863b: 6; Mayr, 1863: 446; Dalla Torre, 1893: 257.
Hoplomyrmus as junior synonym of Myrma: Wheeler, W.M. 1911c: 860; Wheeler, W.M. 1922a: 993; Emery, 1925b: 198; Bolton, 1973b: 288.
Pseudocyrtomyrma Emery, 1921e: 18 [as subgenus of Polyrhachis]. Type-species: Polyrhachis revoili, by original designation.
Taxonomic history
Pseudocyrtomyrma as subgenus of Polyrhachis: Emery, 1921e: 18; Emery, 1925b: 206.
Pseudocyrtomyrma as junior synonym of Myrma: Bolton, 1973b: 288.
Anoplomyrma Chapman, 1963: 258 [as subgenus of Polyrhachis]. Type-species: Polyrhachis (Anoplomyrma) parabiotica, by monotypy.
Taxonomic history
Anoplomyrma as junior synonym of Myrma: Dorow, 1995: 30.
Subgenus POLYRHACHIS (HEMIOPTICA)
Hemioptica Roger, 1862a: 238. Type-species: Hemioptica scissa, by monotypy.
Taxonomic history
Hemioptica in Formicinae: Mayr, 1862: 769 [Formicidae]; Mayr, 1865: 6 [Formicidae].
Hemioptica in Camponotinae: Dalla Torre, 1893: 271.
Hemioptica in Camponotinae, Camponotini: Emery, 1895j: 772; Wheeler, W.M. 1910g: 144; all subsequent authors except the following.
Hemioptica in Camponotinae, Polyrhachidini: Ashmead, 1905b: 384.
Hemioptica as junior synonym of Polyrhachis: Forel, 1878: 368.
Hemioptica as subgenus of Myrma: Wheeler, W.M. 1911c: 860; Wheeler, W.M. 1911f: 164.
Hemioptica as genus: Roger, 1862a: 238; Mayr, 1862: 769; Mayr, 1865: 6; Dalla Torre, 1893: 271; Emery, 1895j: 772; Ashmead, 1905b: 384; Wheeler, W.M. 1910g: 144; Emery, 1925b: 209; Donisthorpe, 1943f: 649; Chapman & Capco, 1951: 255.
Hemioptica as subgenus of Polyrhachis: Mayr, 1872: 138; Forel, 1915b: 107; Forel, 1917: 251; Wheeler, W.M. 1922a: 701; Dorow & Kohout, 1995: 95; Bolton, 1995b: 31; Dorow, 1995: 29.
Subgenus POLYRHACHIS (HAGIOMYRMA)
Hagiomyrma Wheeler, W.M. 1911c: 860 [as subgenus of Polyrhachis]. Type-species: Formica ammon, by original designation.
Taxonomic history
Hagiomyrma as subgenus of Polyrhachis: Wheeler, W.M. 1911c: 860; Forel, 1917: 251; Wheeler, W.M. 1922a: 702; Emery, 1925b: 184; all subsequent authors.
Subgenus POLYRHACHIS (HIRTOMYRMA)
Hirtomyrma Kohout, 2010: 199 [as subgenus of Polyrhachis]. Type-species: Polyrhachis hirta, by original designation.
Taxonomic history
Hirtomyrma as subgenus of Polyrhachis: Kohout, 2010: 199.
Subgenus POLYRHACHIS (CAMPOMYRMA)
Campomyrma Wheeler, W.M. 1911g: 860 [as subgenus of Polyrhachis]. Type-species: Polyrhachis clypeata (junior synonym of Formica exercita), by original designation.
Taxonomic history
Campomyrma as subgenus of Polyrhachis: Forel, 1917: 251; Wheeler, W.M. 1922a: 702; Emery, 1925b: 178; all subsequent authors.
Subgenus POLYRHACHIS (MYRMOTHRINAX)
Myrmothrinax Forel, 1915b: 107 [as subgenus of Polyrhachis]. Type-species: Polyrhachis thrinax, by original designation.
Taxonomic history
Myrmothrinax as subgenus of Polyrhachis: Forel, 1915b: 107; Forel, 1917: 251; Wheeler, W.M. 1922a: 704; Emery, 1925b: 178; all subsequent authors.
Junior synonym of POLYRHACHIS (MYRMOTHRINAX)
Evelyna Donisthorpe, 1937c: 273 [as subgenus of Polyrhachis]. Type-species: Polyrhachis (Evelyna) cheesmanae, by original designation.
Taxonomic history
Evelyna as junior synonym of Myrmothrinax: Hung, 1967b: 402.
Subgenus POLYRHACHIS (CYRTOMYRMA)
Cyrtomyrma Forel, 1915b: 107 [as subgenus of Polyrhachis]. Type-species: Formica rastellata, by original designation.
Taxonomic history
Cyrtomyrma as subgenus of Polyrhachis: Forel, 1915b: 107; Forel, 1917: 251; Wheeler, W.M. 1922a: 701; Emery, 1925b: 207; all subsequent authors.
Subgenus POLYRHACHIS (MYRMHOPLA)
Myrmhopla Forel, 1915b: 107 [as subgenus of Polyrhachis]. Type-species: Formica armata, by original designation.
Taxonomic history
Myrmhopla as subgenus of Polyrhachis: Forel, 1915b: 107; Forel, 1917: 251; Wheeler, W.M. 1922a: 701; Emery, 1925b: 190; all subsequent authors.
Junior synonyms of POLYRHACHIS (MYRMHOPLA)
Cephalomyrma Karavaiev, 1935a: 115 [as subgenus of Polyrhachis]. Type-species: Polyrhachis (Cephalomyrma) stylifera, by monotypy.
Taxonomic history
Cephalomyrma as junior synonym of Myrmhopla: Hung, 1967b: 402.
Florencea Donisthorpe, 1937a: 624 [as subgenus of Polyrhachis]. Type-species: Polyrhachis (Florencea) kirkae (junior synonym of Polyrhachis nigriceps), by original designation.
Taxonomic history
Florencea as junior synonym of Myrmhopla: Hung, 1967b: 402.
Subgenus POLYRHACHIS (CHARIOMYRMA)
Chariomyrma Forel, 1915b: 107 [as subgenus of Polyrhachis]. Type-species: Polyrhachis guerini, by original designation.
Taxonomic history
Chariomyrma as subgenus of Polyrhachis: Forel, 1915b: 107; Forel, 1917: 251; Wheeler, W.M. 1922a: 702; Emery, 1925b: 185; all subsequent authors.
Subgenus POLYRHACHIS (HEDOMYRMA)
Hedomyrma Forel, 1915b: 107 [as subgenus of Polyrhachis]. Type-species: Polyrhachis ornata, by original designation.
Taxonomic history
Hedomyrma as subgenus of Polyrhachis: Forel, 1915b: 107; Forel, 1917: 251; Wheeler, W.M. 1922a: 702; Emery, 1925b: 189; all subsequent authors.
Junior synonyms of POLYRHACHIS (HEDOMYRMA)
Dolichorhachis Mann, 1919: 386 [as subgenus of Polyrhachis]. Type-species: Polyrhachis (Dolichorhachis) malaensis, by monotypy.
Taxonomic history
Dolichorhachis as subgenus of Polyrhachis: Mann, 1919: 386; Wheeler, W.M. 1922a: 702; Emery, 1925b: 188.
Dolichorhachis as junior synonym of Hedomyrma: Dorow, 1995: 26.
Morleyidris Donisthorpe, 1944c: 64 [as subgenus of Polyrhachis]. Type-species: Polyrhachis (Morleyidris) trina, by original designation.
Taxonomic history
Morleyidris as junior synonym of Hedomyrma: Hung, 1967b: 402.
Subgenus POLYRHACHIS (MYRMATOPA)
Myrmatopa Forel, 1915b: 107 [as subgenus of Polyrhachis]. Type-species: Polyrhachis schang, by original designation.
Taxonomic history
Myrmatopa as subgenus of Polyrhachis: Forel, 1917: 251; Wheeler, W.M. 1922a: 702; Emery, 1925b: 180; all subsequent authors.
Junior synonym of POLYRHACHIS (MYRMATOPA)
Irenea Donisthorpe, 1938e: 502 [as subgenus of Dolichoderus]. Type-species: Dolichoderus (Irenea) omyrmex, by original designation.
Taxonomic history
Irenea in Dolichoderinae, Dolichoderini: Donisthorpe, 1938e: 502; Donisthorpe, 1943f: 653; Chapman & Capco, 1951: 186.
Irenea as subgenus of Dolichoderus: Donisthorpe, 1938e: 502.
Irenea as genus: Donisthorpe, 1941h: 59; Chapman & Capco, 1951: 186.
Irenea as junior synonym of Polyrhachis (Myrmatopa): Bolton, 1994: 50; Dorow, 1995: 42.
Subgenus POLYRHACHIS (AULACOMYRMA)
Aulacomyrma Emery, 1921e: 17 [as subgenus of Polyrhachis]. Type-species: Polyrhachis porcata, by original designation.
Taxonomic history
Aulacomyrma as subgenus of Polyrhachis: Emery, 1921e: 17; Emery, 1925b: 197; all subsequent authors.
Junior synonym of POLYRHACHIS (AULACOMYRMA)
Johnia Karavaiev, 1927e: 43 [as subgenus of Polyrhachis]. Type-species: Polyrhachis (Johnia) schizospina, by monotypy.
Taxonomic history
Johnia as junior synonym of Aulacomyrma: Hung, 1967b: 402.
Genus Polyrhachis references
Smith, F. 1858b: 58 (diagnosis); Roger, 1863b: 6 (catalogue); Mayr, 1863: 422, 443 (Hemioptica, Polyrhachis catalogues); Mayr, 1865: 6 (Polyrhachis, Hemioptica diagnoses); Mayr, 1867a: 40, 41 (diagnosis, species groups (as "turmae")); Mayr, 1876: 68 (Australia species key); Forel, 1878: 368 (diagnosis); Mayr, 1879: 648 (species groups); André, 1887: 286 (Afrotropical species key); Dalla Torre, 1893: 257, 271 (Polyrhachis, Hemioptica catalogues); Forel, 1893c: 17 (India & Sri Lanka species key); Emery, 1896d: 378 (catalogue); Emery, 1897d: 579, 582 (P. relucens & P. guerini groups, keys); Emery, 1898a: 227 (additions to 1896a catalogue); Bingham, 1903: 383 (India, Sri Lanka & Burma species key); Santschi, 1910h: 284 (P. rastellata group, key); Viehmeyer, 1914b: 50, 54 (P. rastellata & P. hostilis groups, keys); Wheeler, W.M. 1922a: 256, 701, 992 (diagnosis, subgenera key, Afrotropical catalogue); Arnold, 1924: 741 (diagnosis, South Africa species key); Emery, 1925b: 175 (diagnosis, subgenera key, catalogue); Emery, 1925b: 178 (P. (Campomyrma) diagnosis, catalogue); Emery, 1925b: 180 (P. (Myrmatopa) diagnosis, catalogue); Emery, 1925b: 181 (P. (Polyrhachis) diagnosis, catalogue); Emery, 1925b: 182 (P. (Myrmothrinax) diagnosis, catalogue); Emery, 1925b: 184 (P. (Hagiomyrma) diagnosis, catalogue); Emery, 1925b: 185 (P. (Chariomyrma) diagnosis, catalogue); Emery, 1925b: 188 (P. (Dolichorhachis) diagnosis, catalogue); Emery, 1925b: 189 (P. (Hedomyrma) diagnosis, catalogue); Emery, 1925b: 190 (P. (Myrmhopla) diagnosis, catalogue); Emery, 1925b: 197 (P. (Aulacomyrma) diagnosis, catalogue); Emery, 1925b: 198 (P. (Myrma) diagnosis, catalogue); Emery, 1925b: 206 (P. (Pseudocyrtomyrma) diagnosis, catalogue); Emery, 1925b: 207 (P. (Cyrtomyrma) diagnosis, catalogue); Emery, 1925b: 209 (Hemioptica diagnosis, catalogue); Donisthorpe, 1938c: 246 (P. (Cyrtomyrma) review); Santschi, 1939c: 13 (P. (Pseudocyrtomyrma) species key); Chapman & Capco, 1951: 186, 255, 256 (Asia Irenea, Hemioptica, Polyrhachis checklists); Hung, 1962: 24 (Taiwan species key); Hung, 1967b: 395 (review of subgenera); Hung, 1970: 5 (P. (Polyrhachis) species revision, key); Bolton, 1973b: 289 (Afrotropical species revision, key); Bolton, 1975c: 1 (P. sexspinosa group, key); Taylor & Brown, D.R. 1985: 131 (Australia catalogue); Taylor, 1987a: 59 (Australia, New Caledonia checklist); Kohout, 1987: 169 (Philippines P. sexspinosa group, key); Kohout, 1989: 515 (Australia P. relucens group, key); Kohout, 1990: 500 (P. viehmeyeri group, key); Kohout & Taylor, 1990: 509 (Australia checklist); Morisita, Kubota, Onoyama, et al. 1991: 45 (Japan species key); Wang C. & Wu, 1991: 596 (China species key); Dorow & Kohout, 1995: 96 (P. (Hemioptica) species key); Bolton, 1994: 50 (synoptic classification); Bolton, 1995a: 1052 (census); Bolton, 1995b: 342 (catalogue); Dorow, 1995: 9 (subgenera revision, subgenera key, catalogue); Dorow, 1995: 10-11 (P. (Myrmhopla) species groups, key; P. arachne-, P. cryptoceroides- P. flavoflagellata-, P. furcata- and P. hector species groups, keys); Dorow, 1995: 12 (P. (Aulacomyrma) synopsis); Dorow, 1995: 13 (P. (Campomoyrma) synopsis); Dorow, 1995: 16 (P. (Chariomyrma) synopsis); Dorow, 1995: 21 (P. (Cyrtomyrma) synopsis); Dorow, 1995: 24 (P. (Hagiomyrma) synopsis); Dorow, 1995: 26 (P. (Hedomyrma) synopsis); Dorow, 1995: 28 (P. (Hemioptica) synopsis); Dorow, 1995: 30 (P. (Myrma) synopsis); Dorow, 1995: 42 (P. (Myrmatopa) synopsis); Dorow, 1995: 45 (P. (Myrmhopla) synopsis); Dorow, 1995: 61 (P. (Myrmothrinax) synopsis); Dorow, 1995: 63 (P. (Polyrhachis) synopsis); Wu, J. & Wang, 1995: 159 (China species key); Shattuck, 1999: 107 (Australia synopsis); Zhou, 2001b: 190 (China, Guangxi species key); Xu, 2002b: 523 (China P. (Cyrtomyrma) species key; Bolton, 2003: 122 (synopsis); Karmaly, 2004: 542 (India species key); Kohout, 2006a: 146 (P. cryptoceroides group, key); Kohout, 2006b: 87 (P. Cyrtomyrma) of Australia, Borneo, New Guinea, Solomon Is, keys); Kohout, 2007a: 189 (P. Aulacomyrma) diagnosis, all species revision, key); Kohout, 2007c: 8 (P. (Campomyrma) Borneo, key); Kohout, 2008a: 255 (Sulawesi species, keys); Kohout, 2008b: 298 (P. vestita group key); Kohout, 2008d: 12 (flavoflagellata group key); Kohout, 2009: 40, 44 (modifications to Kohout 2006b key); Terayama, 2009: 218 (Taiwan species key); Heterick, 2009: 106 (south-western Australia species key); Kohout, 2010: 169, 171, 200 (Australia subgenera key, Australia P. (Myrmhopla) species key, Australia P. (Hirtomyrma) species key).
Pseudomyrmecinae Pseudomyrmecini Tetraponera TRUE TRUE FALSEExtant: 89 species, 19 subspecies
Fossil: 7 species
Tetraponera Smith, F. 1852: 44. Type-species: Tetraponera atrata (junior synonym of Eciton nigrum), by subsequent designation of Wheeler, W.M. 1911f: 173.
Taxonomic history
Tetraponera in Formicidae, Poneridae: Smith, F. 1857a: 70.
Tetraponera in Myrmicidae: Smith, F. 1877b: 68.
Tetraponera in Myrmicinae, Pseudomyrmini: Forel, 1917: 240; Emery, 1921f: 24.
Tetraponera in Pseudomyrminae: Wheeler, W.M. 1922a: 654; subsequent authors to the following.
Tetraponera in Pseudomyrmecinae: Taylor & Brown, D.R. 1985: 17; all subsequent authors.
Tetraponera as junior synonym of Pseudomyrma: Roger, 1863b: 24; Mayr, 1863: 451.
Tetraponera as junior synonym of Sima: Dalla Torre, 1893: 53; Bingham, 1903: 107; Emery, 1917a: 95.
Tetraponera as subgenus of Sima: Emery, 1900d: 673; Emery, 1915b: 266; Arnold, 1916: 180; Forel, 1917: 240; Emery, 1921f: 24.
Tetraponera as genus: Smith, F. 1852: 44; Smith, F. 1857a: 70; Smith, F. 1877b: 68; Donisthorpe, 1916b: 244; Wheeler, W.M. 1922a: 654; Taylor & Brown, D.R. 1985: 17; Ward, 1990: 470; Bolton, 1994: 184.
Junior synonyms of TETRAPONERA
Sima Roger, 1863a: 178. Type-species: Sima compressa (junior synonym of Pseudomyrma allaborans), by monotypy.
Taxonomic history
[Type-species not Eciton rufonigrum, unjustified subsequent designation by Emery, 1915b: 266, repeated in Emery, 1917a: 95 and Emery, 1921f: 22.]
Sima in Myrmicidae: Smith, F. 1871a: 328.
Sima in Myrmicidae, Myrmeciidae: Emery, 1877a: 81.
Sima in Myrmicinae: Mayr, 1865: 25 [Myrmicidae]; Mayr, 1866b: 906 [Myrmicidae]; Dalla Torre, 1893: 53.
Sima in Myrmicidae, Pseudomyrminae: Ashmead, 1905b: 383.
Sima in Myrmicinae, Pseudomyrmini: Forel, 1893a: 164; Emery, 1895j: 768; Emery, 1914a: 40; Forel, 1917: 240; Wheeler, W.M. 1915h: 40; Arnold, 1916: 173; Emery, 1921f: 21.
Sima as genus: Roger, 1863a: 178; Mayr, 1863: 453; Mayr, 1868c: 100; Dalla Torre, 1893: 53; Emery, 1900d: 672; Bingham, 1903: 107; Emery, 1914a: 40; Emery, 1915b: 266; Arnold, 1916: 173; Emery, 1917a: 95; Forel, 1917: 240; Emery, 1921f: 21.
Sima as junior synonym of Tetraponera: Smith, F. 1877b: 68; Donisthorpe, 1916b: 244; Wheeler, W.M. 1920: 47; Wheeler, W.M. 1922a: 105; Donisthorpe, 1943g: 726; Ward, 1990: 470.
Pachysima Emery, 1912b: 97 [as subgenus of Sima]. Type-species: Tetraponera aethiops, by monotypy.
Taxonomic history
Pachysima in Myrmicinae, Pseudomyrmini: Forel, 1917: 240; Emery, 1921f: 22.
Pachysima in Pseudomyrminae: Wheeler, W.M. 1922a: 112, 654.
Pachysima as genus: Wheeler, W.M. 1918c: 308 (footnote); Wheeler, W.M. 1922a: 112, 654.
Pachysima as subgenus of Sima: Emery, 1912b: 97; Arnold, 1916: 174; Emery, 1921f: 22.
Pachysima as junior synonym of Tetraponera: Brown, 1973b: 183 [provisional]; Ward, 1990: 470.
Viticicola Wheeler, W.M. 1919i: 130 (in text). Type-species: Sima tessmanni, by original designation.
Taxonomic history
[Viticicola also as new in Wheeler, W.M. 1920: 53.]
Viticicola in Pseudomyrminae: Wheeler, W.M. 1922a: 107, 654.
Viticicola as junior synonym of Tetraponera: Brown, 1973b: 185 [provisional]; Ward, 1990: 470.
[Viticola Donisthorpe, 1943g: 735, incorrect subsequent spelling.]
Parasima Donisthorpe, 1948d: 592 [as subgenus of Tetraponera]. [Unnecessary replacement name for Sima in the sense of Emery, 1921f: 23.]
Taxonomic history
Parasima as junior synonym of Tetraponera: Ward, 1990: 470.
Genus Tetraponera references
Mayr, 1863: 453 (catalogue); Mayr, 1865: 25 (Sima diagnosis); Mayr, 1867a: 82 (Sima diagnosis); Mayr, 1868c: 101 (†Baltic Amber species key); Smith, F. 1877b: 68 (checklist); Dalla Torre, 1893: 53 (catalogue); Emery, 1900d: 17 (Asia & Australia species key); Forel, 1903a: 708 (India & Sri Lanka species key); Bingham, 1903: 107 (India, Sri Lanka & Burma species key); Arnold, 1916: 173 (Sima diagnosis, South Africa species key); Emery, 1921f: 21 (Sima diagnosis, subgenera key, catalogue); Emery, 1921f: 22 (S. (Pachysima) diagnosis, catalogue); Emery, 1921f: 24 (S. (Tetraponera) diagnosis, catalogue); Wheeler, W.M. 1922a: 112 (Pachysima species key); Wheeler, W.M. 1922a: 104, 107, 112 (Tetraponera, Viticicola, Pachysima diagnoses); Wheeler, W.M. 1922a: 795, 801 (Afrotropical Tetraponera, Viticicola, Pachysima catalogues); Wheeler, W.M. 1922a: 1014 (Malagasy catalogue); Taylor & Brown, D.R. 1985: 17 (Australia catalogue); Taylor, 1987a: 79 (Australia checklist); Wu, J. & Wang, 1990: 516 (China species key); Dlussky & Radchenko, 1990: 120 (Vietnam species key); Ward, 1990: 470 (diagnosis, review of genus, phylogeny); Bolton, 1994: 184 (synoptic classification); Bolton, 1995a: 1053 (census); Bolton, 1995b: 416 (catalogue); Wu, J. & Wang, 1995: 55 (China species key); Collingwood & Agosti, 1996: 312 (Saudi Arabia species key); Shattuck, 1999: 208 (Australia synopsis); Zhou, 2001b: 64 (China, Guangxi species key); Ward, 2001: 596 (Oriental & Australian, all species, revision, key); Xu & Chai, 2004: 64 (China species key); Ward, 2006: 120 (Afrotropical species groups key); Ward, 2006: 123 (T. ambigua group, key); Ward, 2009: 288 (T. grandidieri group, key); Dlussky, 2009: 1084 (Eocene amber species key); Terayama, 2009: 128 (Taiwan species key).
Cerapachyinae Cerapachyini Cerapachys TRUE TRUE FALSE146 species
Cerapachys Smith, F. 1857a: 74. Type-species: Cerapachys antennatus, by subsequent designation of Bingham, 1903: 28.
Taxonomic history
Cerapachys in Myrmicinae: Smith, F. 1857a: 74 [Myrmicidae]; Smith, F. 1861b: 47 [Myrmicidae]; Smith, F. 1871a: 329 [Myrmicidae].
Cerapachys in Ponerinae: Dalla Torre, 1893: 17.
Cerapachys in Dorylinae, Cerapachyini: Emery, 1895j: 765; Emery, 1901a: 34.
Cerapachys in Ponerinae, Cerapachyini: Forel, 1893a: 162 (also misspelled Orapachys, same page); Forel, 1900d: 331; Ashmead, 1905b: 382; Wheeler, W.M. 1910g: 137; Emery, 1911d: 8; Arnold, 1915: 11; Forel, 1917: 239; Brown, 1975: 18; Snelling, R.R. 1981: 389; Dlussky & Fedoseeva, 1988: 79; Hölldobler & Wilson, 1990: 10.
Cerapachys in Cerapachyinae, Cerapachyini: Wheeler, W.M. 1902d: 185; Wheeler, W.M. 1922a: 639; Donisthorpe, 1943f: 632; Chapman & Capco, 1951: 17; Kempf, 1972a: 76; Wheeler, G.C. & Wheeler, J. 1985: 256; Bolton, 1990a: 67; Bolton, 1990c: 1357; Jaffe, 1993: 9; Bolton, 1994: 19; Bolton, 2003: 34.
Junior synonyms of CERAPACHYS
Syscia Roger, 1861a: 19. Type-species: Syscia typhla, by monotypy.
Taxonomic history
Syscia in Ponerinae: Mayr, 1862: 714 (in key) [Poneridae]; Mayr, 1865: 15 [Poneridae]; Dalla Torre, 1893: 17.
Syscia in Dorylinae, Cerapachyini: Emery, 1895j: 765.
Syscia in Ponerinae, Cerapachyini: Forel, 1893a: 162; Forel, 1900d: 329; Ashmead, 1905b: 382.
Syscia as genus: Dalla Torre, 1893: 17; Forel, 1900d: 329; Bingham, 1903: 31.
Syscia as subgenus of Cerapachys: Wheeler, W.M. 1902d: 185; Emery, 1902c: 24; Wheeler, W.M. 1910g: 137; Emery, 1911d: 10; Forel, 1917: 239; Wheeler, W.M. 1922a: 639; Donisthorpe, 1943g: 730.
Syscia as junior synonym of Cerapachys: Kempf, 1972a: 76; Brown, 1975: 18; all subsequent authors.
Ooceraea Roger, 1862a: 248. Type-species: Ooceraea fragosa, by monotypy.
Taxonomic history
Ooceraea in Myrmicinae: Mayr, 1865: 24 [Myrmicidae].
Ooceraea in Myrmicidae, Pheidolidae: Emery, 1877a: 81.
Ooceraea in Poneridae: Smith, F. 1871a: 324.
Ooceraea in Ponerinae: Dalla Torre, 1893: 17.
Ooceraea in Ponerinae, Cerapachyini: Forel, 1893a: 162.
Ooceraea in Dorylinae, Cerapachyini: Emery, 1895j: 765; Emery, 1901a: 34.
Ooceraea in Ecitoninae, Ecitonini: Ashmead, 1905b: 381; Ashmead, 1906: 25.
Ooceraea as genus: Roger, 1862a: 248; Smith, F. 1871a: 324; Dalla Torre, 1893: 17; Forel, 1900d: 329; Bingham, 1903: 31.
Ooceraea as subgenus of Cerapachys: Emery, 1902c: 24; Wheeler, W.M. 1902d: 185; Wheeler, W.M. 1910g: 137; Emery, 1911d: 10; Forel, 1917: 239; Wheeler, W.M. 1922a: 639; Donisthorpe, 1943f: 678.
Ooceraea as junior synonym of Cerapachys: Brown, 1973b: 183 [provisional]; Brown, 1975: 19; all subsequent authors.
Lioponera Mayr, 1879: 666. Type-species: Lioponera longitarsus, by monotypy.
Taxonomic history
Lioponera in Ponerinae: Dalla Torre, 1893: 17.
Lioponera in Pachycondylinae, Lioponerini: Ashmead, 1905b: 382.
Lioponera in Ponerinae, Cerapachyini: Forel, 1893a: 162; Forel, 1900d: 329; Wheeler, W.M. 1910g: 137; Emery, 1911d: 11; Forel, 1917: 239.
Lioponera in Dorylinae, Cerapachyini: Emery, 1895j: 765.
Lioponera in Cerapachyinae, Cerapachyini: Wheeler, W.M. 1902d: 185; Wheeler, W.M. 1922a: 639; Donisthorpe, 1943f: 657.
Lioponera as subgenus of Cerapachys: Forel, 1892l: 244.
Lioponera as genus: Mayr, 1879: 666; Dalla Torre, 1893: 17; Forel, 1900d: 329; Bingham, 1903: 26; Emery, 1911d: 11; Wheeler, W.M. 1922a: 639; Donisthorpe, 1943f: 657.
Lioponera as junior synonym of Cerapachys: Brown, 1975: 19; all subsequent authors.
Parasyscia Emery, in André, 1882c: 235. Type-species: Parasyscia piochardi, by monotypy.
Taxonomic history
Parasyscia in Ponerinae: Dalla Torre, 1893: 17.
Parasyscia in Dorylinae, Cerapachyini: Emery, 1895j: 765.
Parasyscia in Ponerinae, Cerapachyini: Forel, 1893a: 162 [Cerapachysii]; Ashmead, 1905b: 382.
Parasyscia as genus: Dalla Torre, 1893: 17.
Parasyscia as subgenus of Cerapachys: Forel, 1892l: 243; Wheeler, W.M. 1902d: 185; Wheeler, W.M. 1910g: 137; Emery, 1911d: 9; Forel, 1917: 239; Wheeler, W.M. 1922a: 639; Donisthorpe, 1943f: 681; Creighton, 1950a: 57.
Parasyscia as junior synonym of Cerapachys: Kempf, 1972a: 76; Brown, 1975: 18; all subsequent authors.
Phyracaces Emery, 1902c: 23. Type-species: Cerapachys mayri, by original designation.
Taxonomic history
Phyracaces in Ponerinae, Cerapachyini: Ashmead, 1905b: 382; Wheeler, W.M. 1910g: 137; Emery, 1911d: 10; Arnold, 1915: 17; Forel, 1917: 239.
Phyracaces in Cerapachyinae, Cerapachyini: Wheeler, W.M. 1902d: 185; Wheeler, W.M. 1922a: 639; Donisthorpe, 1943f: 683.
Phyracaces as junior synonym of Cerapachys: Brown, 1973b: 183 [provisional]; Brown, 1975: 19; all subsequent authors.
Cysias Emery, 1902c: 24 [as subgenus of Cerapachys]. Type-species: Ooceraea papuana, by original designation.
Taxonomic history
Cysias in Ponerinae, Cerapachyini: Ashmead, 1905b: 382.
Cysias as genus: Ashmead, 1905b: 382.
Cysias as subgenus of Cerapachys: Emery, 1902c: 24; Wheeler, W.M. 1910g: 137; Donisthorpe, 1943f: 637.
Cysias as junior synonym of Syscia: Emery, 1911d: 10; Brown, 1975: 19; all subsequent authors.
Ceratopachys Schulz, W.A. 1906: 155, unjustified emendation of Cerapachys.
Taxonomic history
Ceratopachys as junior synonym of Cerapachys: Wheeler, W.M. 1922a: 755.
Chrysapace Crawley, 1924: 380. Type-species: Chrysapace jacobsoni (junior secondary homonym in Cerapachys, replaced by Cerapachys crawleyi), by monotypy.
Taxonomic history
Chrysapace in Ponerinae: Crawley, 1924: 380.
Chrysapace in Cerapachyinae, Cerapachyini: Donisthorpe, 1943f: 633.
Chrysapace as junior synonym of Cerapachys: Brown, 1975: 19; all subsequent authors.
Neophyracaces Clark, 1941: 76. Type-species: Phyracaces clarus, by original designation.
Taxonomic history
Neophyracaces in Cerapachyinae: Clark, 1941: 76.
Neophyracaces as junior synonym of Cerapachys: Brown, 1973b: 183 [provisional]; Brown, 1975: 19; all subsequent authors.
Yunodorylus Xu, 2000b: 297. Type-species: Yunodorylus sexspinus, by original designation.
Taxonomic history
Yunodorylus in Dorylinae: Xu, 2000b: 297.
Yunodorylus in Cerapachyinae, Cerapachyini: Bolton, 2003: 141, 268.
Yunodorylus as junior synonym of Cerapachys: Bolton, 2003: 141, 268; Borowiec, 2009: 57.
Genus Cerapachys references
Roger, 1863b: 21, 25 (catalogue); Mayr, 1863: 403, 439, 455 (catalogue); Mayr, 1865: 15, 24 (Syscia, Ooceraea diagnoses); André, 1882c: 236 (Europe & Algeria species); Dalla Torre, 1893: 17 (Lioponera, Cerapachys, Parasyscia, Ooceraea, Syscia catalogues); Forel, 1900d: 331 (India & Sri Lanka species key); Bingham, 1903: 27, 28 (India, Sri Lanka & Burma Lioponera, Cerapachys species keys); Emery, 1911d: 8 (diagnosis, subgenera key, catalogue); Emery, 1911d: 9 (C. (Parasyscia) diagnosis, catalogue); Emery, 1911d: 10 (C. (Ooceraea) & C. (Syscia) diagnoses, catalogues); Emery, 1911d: 10 (Phyracaces diagnosis, catalogue); Emery, 1911d: 11 (Lioponera diagnosis, catalogue); Arnold, 1915: 11 (diagnosis, South Africa species key); Wheeler, W.M. 1918a: 239 (Australia Phyracaces species key); Mann, 1921: 408 (Fiji species key); Wheeler, W.M. 1922a: 52, 53, 639 (Cerapachys & Phyracaces diagnoses, subgenera key); Wheeler, W.M. 1924d: 224 (Chrysapace, comments); Arnold, 1926: 193 (South Africa species, revised key); Donisthorpe, 1939a: 253 (Lioponera diagnosis); Chapman & Capco, 1951: 17, 21, 22 (Asia Cerapachys, Lioponera, Phyracaces checklists); Wilson, 1959b: 44, 52, 55, 56 (Melanesia, Fiji Is, New Guinea & New Caledonia species keys); Kempf, 1972a: 76 (Neotropical catalogue); Brown, 1975: 18, 26 (diagnosis, keys); Smith, D.R. 1979: 1333 (North America catalogue); Taylor & Brown, D.R. 1985: 23 (Australia catalogue); Taylor, 1987a: 16 (Australia, New Caledonia checklist); Terayama, Kubota, Sakai & Kawazoe, 1988: 35 (Taiwan species key); Morisita, Kubota, Onoyama, et al. 1989: 30 (Japan species key); Bolton, 1995a: 1048 (census); Bolton, 1995b: 142 (catalogue); Wu, J. & Wang, 1995: 47 (China species key); Collingwood & Agosti, 1996: 312 (Saudi Arabia species key); Terayama, 1996: 18 (Japan species key); Shattuck, 1999: 60 (Australia synopsis); Mackay & Mackay, 2002: 36 (U.S.A., New Mexico species key); Borowiec, 2009: 46 (C. sexspinus group key); Terayama, 2009: 122 (Taiwan species key); Heterick, 2009: 126 (south-western Australia species key).
Myrmicinae Dacetini Pyramica TRUE TRUE FALSEExtant: 344 species
Fossil: 4 species
Pyramica Roger, 1862a: 251. Type-species: Pyramica gundlachi, by monotypy.
Taxonomic history
Pyramica in Myrmicinae, Dacetini: Emery, 1924d: 320; all subsequent authors.
Pyramica as junior synonym of Strumigenys: Roger, 1863b: 40; Emery & Forel, 1879a: 464; Dalla Torre, 1893: 145; Bingham, 1903: 147; Emery, 1924d: 320; Donisthorpe, 1943g: 723; Brown & Wilson, 1959b: 281; Brown, 1960b: 37; Kempf, 1972a: 242; Bolton, 1983: 358; Bolton, 1994: 105; Baroni Urbani & De Andrade, 2007: 101.
Pyramica as subgenus of Strumigenys: Brown, 1948e: 110.
Pyramica as genus: Mayr, 1863: 453; Bolton, 1999: 1667; Bolton, 2000: 90; Eguchi, Bui & Yamane, 2011: 26.
Junior synonyms of PYRAMICA
Epitritus Emery, 1869a: 136. Type-species: Epitritus argiolus, by monotypy.
Taxonomic history
Epitritus in Myrmicidae, Cryptoceridae: Emery, 1877a: 81.
Epitritus in Myrmicinae: Emery & Forel, 1879a: 465 [Myrmicidae]; Dalla Torre, 1893: 148.
Epitritus in Cryptoceridae, Dacetonini: Ashmead, 1905b: 384.
Epitritus in Myrmicinae, Dacetini: Forel, 1892d: 344; Forel, 1893a: 164; Emery, 1895j: 770; Wheeler, W.M. 1910g: 141; Emery, 1914a: 42; Forel, 1917: 246; Arnold, 1917: 384; Wheeler, W.M. 1922a: 666; Emery, 1924d: 326; Donisthorpe, 1943f: 643; Brown, 1948e: 122; Brown, 1949b: 44; all subsequent authors.
Epitritus as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 12; Baroni Urbani & De Andrade, 2007: 101.
Epitritus as junior synonym of Pyramica: Bolton, 1999: 1667; Bolton, 2000: 90.
Trichoscapa Emery, 1869b: 24 [as subgenus of Strumigenys]. Type-species: Strumigenys (Trichoscapa) membranifera, by monotypy.
Taxonomic history
Trichoscapa in Myrmicinae: Emery & Forel, 1879a: 464 [Myrmicidae]; Dalla Torre, 1893: 145.
Trichoscapa in Myrmicinae, Dacetini: Forel, 1917: 246; Wheeler, W.M. 1922a: 668; Emery, 1924d: 323; Donisthorpe, 1943g: 734; Brown, 1948e: 112; all subsequent authors.
Trichoscapa as subgenus of Strumigenys: Emery & Forel, 1879a: 464; Forel, 1917: 246; Creighton, 1950a: 304.
Trichoscapa as junior synonym of Strumigenys: Dalla Torre, 1893: 145; Baroni Urbani & De Andrade, 1994: 13; Baroni Urbani & De Andrade, 2007: 101.
Trichoscapa as junior synonym of Cephaloxys: Wheeler, W.M. 1922a: 668; Emery, 1924d: 323; Donisthorpe, 1943g: 734.
Trichoscapa as genus: Brown, 1948e: 112; Kempf, 1972a: 255; Bolton, 1983: 319.
Trichoscapa as junior synonym of Pyramica: Bolton, 1999: 1667; Bolton, 2000: 90.
Pentastruma Forel, 1912a: 50. Type-species: Pentastruma sauteri, by monotypy.
Taxonomic history
Pentastruma in Myrmicinae, Dacetini: Emery, 1914a: 42; Forel, 1917: 246; Wheeler, W.M. 1922a: 666; Emery, 1924d: 326; Donisthorpe, 1943f: 682; Brown, 1948e: 102.
Pentastruma as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 15; Baroni Urbani & De Andrade, 2007: 101.
Pentastruma as junior synonym of Pyramica: Bolton, 1999: 1667; Bolton, 2000: 91.
Glamyromyrmex Wheeler, W.M. 1915e: 487. Type-species: Glamyromyrmex beebei, by monotypy.
Taxonomic history
Glamyromyrmex in Myrmicinae, Dacetini: Wheeler, W.M. 1915e: 491; Forel, 1917: 246; Wheeler, W.M. 1922a: 667; Emery, 1924d: 326; Donisthorpe, 1943f: 647; Brown, 1948e: 116; all subsequent authors.
Glamyromyrmex as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 14; Baroni Urbani & De Andrade, 2007: 101.
Glamyromyrmex as junior synonym of Pyramica: Bolton, 1999: 1667; Bolton, 2000: 91.
Codiomyrmex Wheeler, W.M. 1916d: 326. Type-species: Codiomyrmex thaxteri, by monotypy.
Taxonomic history
Codiomyrmex in Myrmicinae, Dacetini: Wheeler, W.M. 1922a: 667; Emery, 1924d: 325; Donisthorpe, 1943f: 633; Brown, 1948e: 114; all subsequent authors.
Codiomyrmex as subgenus of Strumigenys: Emery, 1924d: 325; Donisthorpe, 1943f: 633.
Codiomyrmex as genus: Wheeler, W.M. 1916d: 326; Brown, 1948e: 114.
Codiomyrmex as junior synonym of Glamyromyrmex: Brown, 1973b: 179 [provisional].
Codiomyrmex as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 14; Baroni Urbani & De Andrade, 2007: 101.
Codiomyrmex as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.
Tingimyrmex Mann, 1926: 104 [as subgenus of Strumigenys]. Type-species: Strumigenys (Tingimyrmex) mirabilis, by monotypy.
Taxonomic history
Tingimyrmex in Myrmicinae, Dacetini: Donisthorpe, 1943g: 732; Brown, 1948e: 111; all subsequent authors.
Tingimyrmex as subgenus of Strumigenys: Mann, 1926: 104; Donisthorpe, 1943g: 732.
Tingimyrmex as genus: Brown, 1948e: 111; Kempf, 1972a: 250.
Tingimyrmex as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 15; Baroni Urbani & De Andrade, 2007: 101.
Tingimyrmex as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.
Codioxenus Santschi, 1931c: 278 [as subgenus of Epitritus]. Type-species: Epitritus (Codioxenus) simulans, by monotypy.
Taxonomic history
Codioxenus in Myrmicinae, Dacetini: Donisthorpe, 1943f: 633; Brown, 1948e: 123; all subsequent authors.
Codioxenus as genus: Brown, 1948e: 123.
Codioxenus as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 14; Baroni Urbani & De Andrade, 2007: 101.
Codioxenus as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.
Smithistruma Brown, 1948e: 104.
Taxonomic history
[Replacement name for Cephaloxys Smith, F. 1865: 76; junior homonym of Cephaloxys Signoret, 1847: 294 (Hemiptera). Type-species not Strumigenys pulchella, unjustified subsequent designation by Brown, 1948e: 104, repeated in Kempf, 1972a: 230, Bolton, 1983: 274 and Bolton, 1995b: 46.]
Smithistruma in Myrmicinae, Dacetini: Brown, 1948e: 104.
Smithistruma as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 14; Baroni Urbani & De Andrade, 2007: 102.
Smithistruma as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.
Homonym replaced by Smithistruma
Cephaloxys Smith, F. 1865: 76. Type-species: Cephaloxys capitata, by monotypy.
Taxonomic history
[Cephaloxys Smith, F. junior homonym of Cephaloxys Signoret, 1847: 294 (Hemiptera).]
Cephaloxys in Cryptoceridae: Smith, F. 1871a: 335.
Cephaloxys in Myrmicinae, Dacetini: Wheeler, W.M. 1922a: 668.
Cephaloxys as junior synonym of Strumigenys: Dalla Torre, 1893: 15.
Cephaloxys as subgenus of Strumigenys: Wheeler, W.M. 1922a: 668; Emery, 1924d: 323; Bernard, 1953b: 254 (anachronism).
Weberistruma Brown, 1948e: 106 [as subgenus of Smithistruma]. Type-species: Strumigenys (Cephaloxys) leptothrix, by original designation.
Taxonomic history
Weberistruma in Myrmicinae, Dacetini: Brown, 1948e: 106.
Weberistruma as genus: Brown, 1949d: 7; Brown, 1953g: 24.
Weberistruma as junior synonym of Smithistruma: Brown, 1973a: 35.
Weberistruma as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.
Weberistruma as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 2007: 102.
Wessonistruma Brown, 1948e: 106 [as subgenus of Smithistruma]. Type-species: Strumigenys pergandei, by original designation.
Taxonomic history
Wessonistruma in Myrmicinae, Dacetini: Brown, 1948e: 106.
Wessonistruma junior synonym of Smithistruma: Brown, 1973a: 35.
Wessonistruma as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.
Wessonistruma as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 2007: 102.
Serrastruma Brown, 1948e: 107 [as subgenus of Smithistruma]. Type-species: Strumigenys simoni, by original designation.
Taxonomic history
Serrastruma in Myrmicinae, Dacetini: Brown, 1948e: 107, and all subsequent authors.
Serrastruma as genus: Brown, 1949d: 6; Brown, 1952e: 70; Bolton, 1983: 335; Bolton, 1994: 105.
Serrastruma as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 14; Baroni Urbani & De Andrade, 2007: 102.
Serrastruma as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.
Neostruma Brown, 1948e: 111. Type-species: Strumigenys crassicornis, by original designation.
Taxonomic history
Neostruma in Myrmicinae, Dacetini: Brown, 1948e: 111; all subsequent authors.
Neostruma as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 12; Baroni Urbani & De Andrade, 2007: 102.
Neostruma as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.
Dorisidris Brown, 1948e: 116. Type-species: Strumigenys (Codiomyrmex) nitens, by original designation.
Taxonomic history
Dorisidris in Myrmicinae, Dacetini: Brown, 1948e: 116; all subsequent authors.
Dorisidris as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 14; Baroni Urbani & De Andrade, 2007: 102.
Dorisidris as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.
Miccostruma Brown, 1948e: 123. Type-species: Epitritus mandibularis, by original designation.
Taxonomic history
Miccostruma in Myrmicinae, Dacetini: Brown, 1948e: 123.
Miccostruma as junior synonym of Smithistruma: Bolton, 1983: 274.
Miccostruma as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.
Miccostruma as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 2007: 102.
Kyidris Brown, 1949d: 3. Type-species: Kyidris mutica, by original designation.
Taxonomic history
Kyidris in Myrmicinae, Dacetini: Brown, 1949d: 3.
Kyidris as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 15; Baroni Urbani & De Andrade, 2007: 103.
Kyidris as junior synonymof Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.
Chelystruma Brown, 1950a: 33 [as subgenus of Glamyromyrmex]. Type-species: Glamyromyrmex (Chelystruma) lilloana, by monotypy.
Taxonomic history
Chelystruma in Myrmicinae, Dacetini: Brown, 1950a: 33; all subsequent authors.
Chelystruma as subgenus of Glamyromyrmex: Brown, 1953g: 16.
Chelystruma as genus: Kempf, 1959d: 338; Kempf, 1972a: 77.
Chelystruma as junior synonym of Glamyromyrmex: Brown, 1973b: 179 [provisional].
Chelystruma as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 14; Baroni Urbani & De Andrade, 2007: 103.
Chelystruma as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.
Polyhomoa Azuma, 1950: 36. Type-species: Polyhomoa itoi (junior synonym of Kyidris mutica), by monotypy.
Taxonomic history
Polyhomoa as junior synonym of Kyidris: Creighton, 1950b: 93; Brown & Yasumatsu, 1951: 93.
Polyhomoa as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.
Polyhomoa as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 2007: 103.
Borgmeierita Brown, 1953g: 23. Type-species: Codiomyrmex excisus, by original designation.
Taxonomic history
Borgmeierita in Myrmicinae, Dacetini: Brown, 1953g: 23.
Borgmeierita as junior synonym of Glamyromyrmex: Brown, 1973a: 35.
Borgmeierita as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.
Borgmeierita as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 2007: 103.
Platystruma Brown, 1953g: 112 [as subgenus of Smithistruma]. Type-species: Strumigenys (Cephaloxys) depressiceps, by original designation.
Taxonomic history
Platystruma in Myrmicinae, Dacetini: Brown, 1953g: 112; all subsequent authors.
Platystruma as junior synonym of Smithistruma: Brown, 1973a: 35.
Platystruma as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.
Platystruma as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 2007: 103.
Gymnomyrmex Borgmeier, 1954b: 279. Type-species: Gymnomyrmex splendens, by original designation.
Taxonomic history
Gymnomyrmex in Myrmicinae, Dacetini: Borgmeier, 1954b: 279; all subsequent authors.
Gymnomyrmex as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 14; Baroni Urbani & De Andrade, 2007: 103.
Gymnomyrmex as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.
Dysedrognathus Taylor, 1968d: 132. Type-species: Dysedrognathus extemenus, by original designation.
Taxonomic history
Dysedrognathus in Myrmicinae, Dacetini: Taylor, 1968d: 132.
Dysedrognathus as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 13; Baroni Urbani & De Andrade, 2007: 103.
Dysedrognathus as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.
Asketogenys Brown, 1972: 23. Type-species: Asketogenys acubecca, by original designation.
Taxonomic history
Asketogenys in Myrmicinae, Dacetini: Brown, 1972: 23.
Asketogenys as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 15; Baroni Urbani & De Andrade, 2007: 103.
Asketogenys as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.
Cladarogenys Brown, 1976b: 33. Type-species: Cladarogenys lasia, by original designation.
Taxonomic history
Cladarogenys in Myrmicinae, Dacetini: Brown, 1976b: 33.
Cladarogenys as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 14; Baroni Urbani & De Andrade, 2007: 103.
Cladarogenys as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.
Genus Pyramica references
Mayr, 1863: 453 (catalogue); André, 1883b: 398, 402 (Europe & Algeria Strumigenys (Trichoscapa), Epitritus species); Dalla Torre, 1893: 145, 148 (Strumigenys, Epitritus catalogues); Emery, 1895c: 325 (North America species key); Wheeler, W.M. 1908a: 146 (New World species key); Santschi, 1913b: 257 (Afrotropical species key); Emery, 1916b: 205 (Italy species key); Arnold, 1917: 373 (South Africa species key); Wheeler, W.M. 1922a: 918, 920 (Afrotropical Strumigenys (Cephaloxys), Epitritus catalogues); Emery, 1924d: 323, 325, 326 (Strumigenys (Cephaloxys), Strumigenys (Codiomyrmex), Glamyromyrmex, Pentastruma, Epitritus diagnoses, catalogues); Smith, M.R. 1931c: 691 (North America Strumigenys (Cephaloxys) species key); Buren, 1944a: 290 (U.S.A., Iowa species key); Brown, 1948e: 112 - 124 (minor genera reviews); Brown, 1949b: 43 (Epitritus diagnosis, review); Brown, 1949d: 8 (Weberistruma species, review); Brown, 1949d: 21 (Japan, China & Taiwan species key); Creighton, 1950a: 301 (North America Strumigenys (Trichoscapa) species key); Chapman & Capco, 1951: 104, 105, 106 (Asia Epitritus, Kyidris, Pentastruma, Smithistruma, Strumigenys (Cephaloxys) checklists); Brown, 1950a: 27 (Glamyromyrmex review); Brown, 1952e: 86 (Serrastruma diagnosis, species revision, key); Brown, 1953g: 20, 24 (Codiomyrmex, Weberistruma reviews); Brown, 1953g: 31, 46, 92, 114, 125, 130 (Smithistruma diagnosis, Nearctic, Neotropical, Malesian, Afrotropical, Palaearctic species revisions, keys); Wilson & Brown, 1956b: 439 (Kyidris species review); Brown, 1959b: 1 (Neostruma species review); Kempf, 1959d: 339 (Gymnomyrmex species key); Brown, 1960b: 37 (Strumigenys gundlachi group revision, key); Kempf, 1960f: 447 (Neotropical Glamyromyrmex species key); Kempf, 1960f: 451 (Gymnomyrmex species key); Brown, 1962b: 79 (Epitritus species key); Brown, 1964a: 183 (supplement to 1953e Smithistruma revision); Bernard, 1967: 243, 245 (Smithistruma, Epitritus diagnoses); Kempf, 1972a: 37, 77, 98, 110, 116, 163, 230, 250, 255 (Neotropical Borgmeierita, Chelystruma, Codiomyrmex, Codioxenus, Dorisidris, Glamyromyrmex, Gymnomyrmex, Neostruma, Smithistruma, Tingimyrmex, Trichoscapa catalogues); Bolton, 1972: 208 (Epitritus species key); Alayo, 1974: 19 (Cuba species key); Smith, D.R. 1979: 1405, 1408 (North America Smithistruma, Trichoscapa catalogues); Taylor, 1987a: 28 (Australia checklist); Brown & Boisvert, 1979: 201 (Pentastruma species review); Bolton, 1983: 274, 319, 320, 335, 353, 354 (Afrotropical Smithistruma, Trichoscapa, Glamyromyrmex, Serrastruma, Cladarogenys, Epitritus species revisions, keys); Taylor & Brown, D.R. 1985: 64 (Australia Glamyromyrmex catalogue); Perrault, 1986a: 3 (Gymnomyrmex species key); Ward, 1988: 122 (Western Nearctic Smithistruma species key); Terayama & Kubota, 1989: 787 (Taiwan Smithistruma species key); Hölldobler & Wilson, 1990: 15 (synoptic classification); Brandão, 1991: 344, 346, 360, 378 (Neotropical catalogue); Arakelian & Dlussky, 1991: 150 (former U.S.S.R. species key); Morisita, Kubota, et al. 1992: 68 (Japan Smithistruma species key); Morisita, Kubota, et al. 1992: 76 (Japan Epitritus species key); Bolton, 1994: 105 (synoptic classification); Bolton, 1995a: 1048 - 1053 (census); Bolton, 1995b: 75, 145, 146, 177, 183, 188, 207, 211, 219, 292, 316, 382, 383, 420, 421 (catalogue); Terayama, Lin & Wu, 1996: 328 (Taiwan Smithistruma species key); Ogata & Onoyama, 1998: 277 (Japan Smithistruma species key); Shattuck, 1999: 136, 170 (Australia Glamyromyrmex, Trichoscapa synopses); Bolton, 1999: 1657, 1667 (morphology, diagnosis, review of genus, phylogeny); Xu, 2000e: 297 (China Epitritus species key); Bolton, 2000: 95, 137, 272, 285, 342, 372, 472 (Nearctic, Neotropical, Afrotropical, West Palaearctic, Malagasy, Malesian-Oriental-East Palaearctic, Austral species revisions, keys); Xu & Zhou, 2004: 441 (China species, key); Longino, 2006b: 139 (Costa Rica species, additions to key); Terayama, 2009: 138 (Taiwan species key).
Myrmicinae Dacetini Strumigenys TRUE TRUE FALSEExtant: 487 species
Fossil:
Strumigenys Smith, F. 1860c: 72. Type-species: Strumigenys mandibularis, by monotypy.
Taxonomic history
Strumigenys in Myrmicidae: Smith, F. 1871a: 334; Cresson, 1887: 262.
Strumigenys in Myrmicidae, Cryptoceridae: Emery, 1877a: 81.
Strumigenys in Myrmicinae: Mayr, 1865: 26 [Myrmicidae]; Emery & Forel, 1879a: 464 [Myrmicidae]; Dalla Torre, 1893: 145.
Strumigenys in Cryptoceridae, Dacetini: Ashmead, 1905b: 384.
Strumigenys in Myrmicinae, Dacetini: Forel, 1892d: 344; Forel, 1893a: 164; Forel, 1895b: 133; Emery, 1895j: 770; Forel, 1899c: 42; Wheeler, W.M. 1910g: 141; Emery, 1914a: 42; Forel, 1917: 246; Arnold, 1917: 372; Wheeler, W.M. 1922a: 668; Emery, 1924d: 319; Brown, 1948e: 108; all subsequent authors.
Junior synonyms of STRUMIGENYS
Labidogenys Roger, 1862a: 249. Type-species: Labidogenys lyroessa, by monotypy.
Taxonomic history
Labidogenys in Myrmicinae, Dacetini: Donisthorpe, 1943f: 654; Brown, 1948e: 102.
Labidogenys as genus: Roger, 1862a: 249; Brown, 1948e: 102.
Labidogenys as junior synonym of Strumigenys: Roger, 1863b: 40; Emery & Forel, 1879a: 464; Dalla Torre, 1893: 145; Bingham, 1903: 147; Emery, 1924d: 320; Brown & Wilson, 1959b: 281; Brown, 1960b: 38; Bolton, 1983: 358; Bolton, 1999: 1673; Bolton, 2000: 492.
Proscopomyrmex Patrizi, 1946: 294. Type-species: Proscopomyrmex londianensis, by monotypy.
Taxonomic history
Proscopomyrmex in Myrmicinae, Dacetini: Patrizi, 1946: 294 [Dacetonini].
Proscopomyrmex as junior synonym of Strumigenys: Brown, 1949d: 15; Bolton, 1999: 1673; Bolton, 2000: 492.
Eneria Donisthorpe, 1948d: 598. Type-species: Eneria excisa (junior synonym of Strumigenys loriae), by original designation.
Taxonomic history
Eneria as genus: Chapman & Capco, 1951: 104 (anachronism).
Eneria as junior synonym of Strumigenys: Brown, 1949d: 15; Bolton, 1999: 1673; Bolton, 2000: 492.
Quadristruma Brown, 1949b: 47. Type-species: Epitritus emmae, by original designation.
Taxonomic history
Quadristruma in Myrmicinae, Dacetini: Brown, 1949b: 47; all subsequent authors.
Quadristruma as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 12; Bolton, 1999: 1673; Bolton, 2000: 492.
Genus Strumigenys references
Roger, 1863b: 40 (catalogue); Mayr, 1863: 424, 455 (catalogue); Mayr, 1865: 26 (diagnosis); Forel, 1886a: 216 (all species key); Mayr, 1887: 568 (all species key); Dalla Torre, 1893: 145 (catalogue); Emery, 1895c: 325 (North America species key); Emery, 1897c: 574 (New Guinea species key); Forel, 1903a: 707 (India & Sri Lanka species key); Bingham, 1903: 147 (India, Sri Lanka & Burma species key); Wheeler, W.M. 1908a: 146 (New World species key); Santschi, 1913b: 257 (Afrotropical species key); Arnold, 1917: 373 (diagnosis, South Africa species key); Mann, 1921: 467 (Fiji Is species key); Wheeler, W.M. 1922a: 917, 1034 (Afrotropical, Malagasy catalogues); Emery, 1924d: 319 (diagnosis, catalogue); Brown, 1948e: 108 (diagnosis, review of genus); Brown, 1949d: 21 (Japan, China & Taiwan species key); Creighton, 1950a: 301 (North America species key); Chapman & Capco, 1951: 104, 107 (Asia Eneria, Strumigenys (Strumigenys) checklists); Brown, 1954k: 10 (Afrotropical species revision, key); Brown, 1958h: 38 (New Zealand species); Brown, 1962c: 238 (Neotropical species synopsis, key); Kempf, 1972a: 226, 243 (Neotropical Quadristruma, Strumigenys catalogues); Alayo, 1974: 19 (Cuba species key); Kempf, 1976b: 43 (additions to Brown, 1962c key); Smith, D.R. 1979: 1403, 1409 (North America Strumigenys, Quadristruma catalogues); Bolton, 1983: 358, 400 (Afrotropical Strumigenys, Quadristruma species revisions, keys); Taylor & Brown, D.R. 1985: 85, 87 (Australia Quadristruma, Strumigenys catalogues); Taylor, 1987a: 66, 76 (Australia, New Caledonia & New Zealand checklists); Terayama & Kubota, 1989: 780 (Taiwan species key); Brandão, 1991: 380 (Neotropical catalogue); Arakelian & Dlussky, 1991: 150 (former U.S.S.R. species key); Morisita, Kubota, et al. 1992: 63 (Japan species key); Dlussky, 1993: 53 (Fiji Is, Tonga and Samoa species key); Bolton, 1994: 105 (synoptic classification); Bolton, 1995a: 1053 (census); Bolton, 1995b: 377, 395 (catalogue); Gronenberg, 1996: 2021 (mandible morphology); Lin & Wu, 1996: 139 (Taiwan species key); Lattke & Goitía, 1997: 370 (Venezuela species key); Shattuck, 1999: 160, 167 (Australia synopsis); Bolton, 1999: 1657, 1673 (morphology, diagnosis, review of genus, phylogeny); Bolton, 2000: 496, 583, 716, 942 (Neotropical-Nearctic, Afrotropical, Malesian-Oriental-East Palaearctic, Austral species revisions, keys); Fisher, 2000: 614 (Malagasy species revision, key); Zhou, 2001b: 78 (China, Guangxi species key); Lin & Wu, 2001: 166 (Taiwan species key); Zhou & Xu, 2003: 737 (China species key); Yoshimura & Onoyama, 2007: 676 (Japan species key); Terayama, 2009: 142 (Taiwan species key); Heterick, 2009: 176 (south-western Australia species key).
Myrmicinae Solenopsidini Carebara TRUE TRUE FALSEExtant: 160 species, 13 subspecies
Fossil: 6 species
Carebara Westwood, 1840b: 86. Type-species: Carebara lignata, by monotypy.
Taxonomic history
Carebara in Poneridae, Attidae: Smith, F. 1858b: 178.
Carebara in Myrmicinae: Mayr, 1865: 23 [Myrmicidae]; Smith, F. 1871a: 334 [Myrmicidae]; Dalla Torre, 1893: 74.
Carebara in Myrmicinae, Pheidolini: Emery, 1877a: 81 [Myrmicidae, Pheidolidae].
Carebara in Myrmicinae, Myrmicariini: Ashmead, 1905b: 383.
Carebara in Myrmicinae, Solenopsidini: Forel, 1893a: 164; Emery, 1895j: 770; Wheeler, W.M. 1910g: 140; Wheeler, W.M. 1922a: 663; Kusnezov, 1964: 61; Kempf, 1972a: 74; Jaffe, 1993: 10; Bolton, 2003: 61.
Carebara in Myrmicinae, Pheidologeton genus group: Ettershank, 1966: 81; Bolton, 1987: 265.
Carebara in Myrmicinae, Pheidologetini: Emery, 1914a: 41; Arnold, 1916: 248; Forel, 1917: 244; Emery, 1924d: 219; Dlussky & Fedoseeva, 1988: 80; Hölldobler & Wilson, 1990: 16; Bolton, 1994: 106 [Pheidologetonini].
Junior synonyms of CAREBARA
Oligomyrmex Mayr, 1867a: 110. Type-species: Oligomyrmex concinnus, by monotypy.
Taxonomic history
Oligomyrmex in Myrmicinae: Dalla Torre, 1893: 74.
Oligomyrmex in Myrmicinae, Stenammini: Ashmead, 1905b: 383.
Oligomyrmex in Myrmicinae, Solenopsidini: Forel, 1893a: 164; Emery, 1895j: 770; Wheeler, W.M. 1910g: 140; Wheeler, W.M. 1922a: 663; Kempf, 1972a: 172; Smith, D.R. 1979: 1389; Jaffe, 1993: 10.
Oligomyrmex in Myrmicinae, Pheidologeton genus group: Ettershank, 1966: 81; Bolton, 1987: 265.
Oligomyrmex in Myrmicinae, Pheidologetini: Emery, 1914a: 41; Forel, 1917: 243; Emery, 1924d: 215; Dlussky & Fedoseeva, 1988: 80; Hölldobler & Wilson, 1990: 16; Bolton, 1994: 106 [Pheidologetonini].
Oligomyrmex in Myrmicinae, Solenopsidini: Bolton, 2003: 61.
Oligomyrmex as junior synonym of Carebara: Fernández, 2004a: 194.
Aeromyrma Forel, 1891b: 198. Type-species: Aeromyrma nosindambo, by monotypy.
Taxonomic history
[Aeromyrma also described as new by Forel, 1891a: cccvii; no species-rank taxa named.]
Aeromyrma in Myrmicinae: Dalla Torre, 1893: 78.
Aeromyrma in Myrmicinae, Solenopsidini: Forel, 1893a: 164; Emery, 1895j: 770; Ashmead, 1905b: 383; Wheeler, W.M. 1910g: 140; Wheeler, W.M. 1922a: 663.
Aeromyrma in Myrmicinae, Pheidologetini: Emery, 1914a: 41; Arnold, 1916: 256; Forel, 1917: 243; Emery, 1924d: 215; all subsequent authors.
Aeromyrma as genus: Forel, 1891b: 198; Arnold, 1916: 256; Forel, 1917: 243; Wheeler, W.M. 1922a: 663, 882.
Aeromyrma as subgenus of Oligomyrmex: Emery, 1915c: 59 (footnote); Emery, 1924d: 215; Donisthorpe, 1943f: 620.
Aeromyrma as junior synonym of Oligomyrmex: Ettershank, 1966: 119; all subsequent authors to the following.
Aeromyrma as junior synonym of Carebara: Fernández, 2004a: 194.
Aneleus Emery, 1900c: 327 [as subgenus of Pheidologeton]. Type-species: Solenopsis similis, by subsequent designation of Wheeler, W.M. 1911f: 158.
Taxonomic history
[Type-species not Pheidologeton pygmaeus, unjustified subsequent designation by Wheeler, W.M. 1913a: 77; repeated in Emery, 1924d: 214.]
Aneleus in Myrmicinae, Solenopsidini: Wheeler, W.M. 1910g: 140; Wheeler, W.M. 1922a: 663.
Aneleus in Myrmicinae, Pheidologetini: Emery, 1914a: 41; Arnold, 1916: 254; Forel, 1917: 243; Emery, 1924d: 213; all subsequent authors.
Aneleus as genus: Emery, 1914a: 41; Arnold, 1916: 254; Forel, 1917: 243; Wheeler, W.M. 1922a: 663; Emery, 1924d: 213.
Aneleus as junior synonym of Oligomyrmex: Ettershank, 1966: 119; all subsequent authors to the following.
Aneleus as junior synonym of Carebara: Fernández, 2004a: 194.
Erebomyrma Wheeler, W.M. 1903a: 138. Type-species: Erebomyrma longii, by monotypy.
Taxonomic history
Erebomyrma in Myrmicinae, Solenopsidini: Wheeler, W.M. 1903a: 145; Wheeler, W.M. 1910g: 140; Wheeler, W.M. 1915h: 46; Wheeler, W.M. 1922a: 663.
Erebomyrma in Myrmicinae, Stenammini: Ashmead, 1905b: 383.
Erebomyrma in Myrmicinae, Pheidologetini: Emery, 1914a: 41; Forel, 1917: 244; Emery, 1924d: 218; all subsequent authors.
Erebomyrma as genus: Wheeler, W.M. 1922a: 663; Emery, 1924d: 218; Creighton, 1950a: 245; Kusnezov, 1964: 62; Wilson, 1986b: 61; Hölldobler & Wilson, 1990: 13.
Erebomyrma as junior synonym of Oligomyrmex: Ettershank, 1966: 119; Kempf, 1972a: 172; Smith, D.R. 1979: 1389; Bolton, 1994: 106; Bolton, 1995b: 298.
Erebomyrma as junior synonym of Carebara: Fernández, 2004a: 194.
Paedalgus Forel, 1911i: 217. Type-species: Paedalgus escherichi, by monotypy.
Taxonomic history
Paedalgus in Myrmicinae, Solenopsidini: Forel, 1911i: 218; Wheeler, W.M. 1922a: 663.
Paedalgus in Myrmicinae, Pheidologeton genus group: Ettershank, 1966: 81; Bolton, 1987: 265.
Paedalgus in Myrmicinae, Pheidologetini: Emery, 1914a: 41; Forel, 1917: 244; Emery, 1924d: 221; Dlussky & Fedoseeva, 1988: 80; Bolton, 1994: 106 [Pheidologetonini].
Paedalgus in Myrmicinae, Solenopsidini: Bolton, 2003: 61.
Paedalgus as junior synonym of Carebara: Fernández, 2004a: 194.
Lecanomyrma Forel, 1913k: 56 [as subgenus of Pheidologeton]. Type-species: Pheidologeton (Lecanomyrma) butteli, by monotypy.
Taxonomic history
Lecanomyrma as subgenus of Pheidologeton: Forel, 1913k: 56; Forel, 1917: 243.
Lecanomyrma as subgenus of Aneleus: Emery, 1924d: 215.
Lecanomyrma as junior synonym of Oligomyrmex: Ettershank, 1966: 119; all subsequent authors to the following.
Lecanomyrma as junior synonym of Carebara: Fernández, 2004a: 194.
Spelaeomyrmex Wheeler, W.M. 1922c: 9. Type-species: Spelaeomyrmex urichi, by original designation.
Taxonomic history
Spelaeomyrmex in Myrmicinae, Pheidologitonini: Wheeler, W.M. 1922c: 11; Donisthorpe, 1944b: 59.
Spelaeomyrmex in Myrmicinae, Solenopsidini: Donisthorpe, 1943g: 727; Kusnezov, 1964: 61.
Spelaeomyrmex as junior synonym of Erebomyrma: Wilson, 1962a: 63.
Spelaeomyrmex as junior synonym of Carebara: Fernández, 2004a: 194.
Hendecatella Wheeler, W.M. 1927h: 93 [as subgenus of Oligomyrmex]. Type-species: Oligomyrmex (Hendecatella) capreolus, by monotypy.
Taxonomic history
Hendecatella as junior synonym of Oligomyrmex: Ettershank, 1966: 119; all subsequent authors.
Hendecatella as junior synonym of Carebara: Fernández, 2004a: 194.
Solenops Karavaiev, 1930a: 207 [as subgenus of Solenopsis]. Type-species: Solenopsis (Solenops) weyeri, by monotypy.
Taxonomic history
Solenops as junior synonym of Oligomyrmex: Ettershank, 1966: 119; all subsequent authors.
Solenops as junior synonym of Carebara: Fernández, 2004a: 194.
Crateropsis Patrizi, 1948: 174 [as subgenus of Solenopsis]. Type-species: Solenopsis (Crateropsis) elmenteitae, by original designation.
Taxonomic history
Crateropsis as junior synonym of Oligomyrmex: Ettershank, 1966: 120; all subsequent authors.
Crateropsis as junior synonym of Carebara: Fernández, 2004a: 194.
Sporocleptes Arnold, 1948: 219. Type-species: Sporocleptes nicotianae, by original designation.
Taxonomic history
Sporocleptes as junior synonym of Aneleus: Consani, 1951: 169; Arnold, 1952a: 460.
Sporocleptes as junior synonym of Carebara: Fernández, 2004a: 194.
Nimbamyrma Bernard, 1953b: 240. Type-species: Nimbamyrma villiersi, by monotypy.
Taxonomic history
Nimbamyrma in Myrmicinae, Pheidologetini: Bernard, 1953b: 240.
Nimbamyrma as junior synonym of Oligomyrmex: Ettershank, 1966: 120; all subsequent authors.
Nimbamyrma as junior synonym of Carebara: Fernández, 2004a: 194; Fernández, 2010: 201.
Afroxyidris Belshaw & Bolton, 1994: 631. Type-species: Afroxyidris crigensis, by original designation.
Taxonomic history
Afroxyidris in Myrmicinae, Pheidologetonini: Belshaw & Bolton, 1994: 632; Bolton, 1994: 106.
Afroxyidris in Myrmicinae, Solenopsidini: Bolton, 2003: 61.
Afroxyidris as junior synonym of Carebara: Fernández, 2004a: 194.
Neoblepharidatta Sheela & Narendran, 1997: 88. Type-species: Neoblepharidatta nayana, by original designation.
Taxonomic history
Neoblepharidatta incertae sedis in Myrmicinae: Sheela & Narendran, 1997: 89.
Neoblepharidatta as junior synonym of Oligomyrmex: Bolton, 2003: 216, 273.
Neoblepharidatta as junior synonym of Carebara: Fernández, 2004a: 194.
Parvimyrma Eguchi & Bui, 2007: 40. Type-species: Parvimyrma sangi, by original designation.
Taxonomic history
Parvimyrma in Myrmicinae, Solenopsis genus group: Eguchi & Bui, 2007: 40.
Parvimyrma as junior synonym of Carebara: Fernández, 2010: 195.
Genus Carebara references
Smith, F. 1858b: 178 (Carebara diagnosis); Roger, 1863b: 32 (Carebara catalogue); Mayr, 1863: 402 (Carebara catalogue); Mayr, 1865: 23 (Carebara diagnosis); Mayr, 1867a: 106 (Carebara diagnosis); Dalla Torre, 1893: 74, 78 (Carebara, Oligomyrmex, Aeromyrma catalogues); Bingham, 1903: 150 (Carebara diagnosis); Bingham, 1903: 154 (Oligomyrmex India, Sri Lanka & Burma species key); Santschi, 1913e: 459 (African Oligomyrmex, key); Santschi, 1914d: 363 (Afrotropical Carebara species key); Arnold, 1916: 248, 254, 256 (Carebara, Aneleus, Aeromyrma diagnoses); Wheeler, W.M. 1922a: 165, 687 (Aeromyrma diagnosis, Oligomyrmex subgenera key); Wheeler, W.M. 1922a: 168, 172, 883 (Carebara diagnosis, Afrotropical species key, Afrotropical catalogue); Wheeler, W.M. 1922a: 177, 884 (Paedalgus diagnosis, Afrotropical catalogue); Wheeler, W.M. 1922a: 880, 881, 882 (Afrotropical Aneleus, Oligomyrmex, Aeromyrma catalogues); Wheeler, W.M. 1922a: 1028 (Malagasy Oligomyrmex, Aeromyrma catalogues); Emery, 1924d: 213 (Aneleus diagnosis, catalogue); Emery, 1924d: 215 (Aneleus (Lecanomyrma) diagnosis, catalogue); Emery, 1924d: 215 (Oligomyrmex (Aeromyrma) diagnosis, catalogue); Emery, 1924d: 217 (O. (Oligomyrex) diagnosis, catalogue); Emery, 1924d: 218 (Erebomyrma diagnosis, catalogue); Emery, 1924d: 219 (Carebara diagnosis, catalogue); Emery, 1924d: 221 (Paedalgus diagnosis, catalogue); Weber, 1950a: 1 (Africa Oligomyrmex species, synopsis); Chapman & Capco, 1951: 151, 154, 156, 158 (Asia Carebara, Aeromyrma, Aneleus, Oligomyrmex, Paedalgus checklists); Weber, 1952: 11 (Africa O. (Aeromyrma) species key); Ettershank, 1966: 119 (Oligomyrmex diagnosis, review of genus, checklist); Ettershank, 1966: 125 (Carebara diagnosis, review of genus, checklist); Ettershank, 1966: 128 (Paedalgus diagnosis, review of genus, checklist); Kempf, 1972a: 74 (Carebara Neotropical catalogue); Kempf, 1972a: 172 (Oligomyrmex Neotropical catalogue); Smith, D.R. 1979: 1389 (Oligomyrmex North America catalogue); Taylor & Brown, D.R. 1985: 71 (Oligomyrmex Australia catalogue); Taylor, 1987a: 50 (Oligomyrmex Australia, New Caledonia checklist); Taylor, 1991db: 605 (Oligomyrmex Australia species, partial key); Brandão, 1991: 342 (Oligomyrmex Neotropical catalogue); Morisita, Kubota, et al. 1992: 45 (Oligomyrmex Japan species key); Bolton & Belshaw, 1993: 181 (Paedalgus diagnosis, all species revision, key); Bolton, 1995a: 1048, 1051 (Carebara, Oligomyrmex, Paedalgus census); Bolton, 1995b: 133, 298, 311 (Carebara, Oligomyrmex, Paedalgus catalogues); Wu, J. & Wang, 1995: 74 (Oligomyrmex China species key); Terayama, 1996: 24 (Oligomyrmex Japan species key); Shattuck, 1999: 148 (Oligomyrmex Australia synopsis); Xu, 2003: 311 (Oligomyrmex China species key); Terayama, 2009: 151 (Taiwan species key).
Myrmicinae Solenopsidini Monomorium TRUE TRUE FALSEExtant: 386 species, 30 subspecies
Fossil: 3 species
Monomorium Mayr, 1855: 452. Type-species: Monomorium monomorium (replacement name for Monomorium minutum, junior secondary homonym in Monomorium), by monotypy.
Taxonomic history
Monomorium in Myrmicidae, Myrmicidae: Emery, 1877a: 81.
Monomorium in Myrmicinae: Mayr, 1855: 452 [Myrmicidae]; Mayr, 1861: 71 [Myrmicidae]; Mayr, 1865: 22 [Myrmicidae]; Emery & Forel, 1879a: 456 [Myrmicidae]; Dalla Torre, 1893: 65.
Monomorium in Myrmicinae, Myrmicini: Forel, 1895b: 124; Emery, 1895j: 769; Forel, 1899c: 78; Wheeler, W.M. 1910g: 139.
Monomorium in Myrmicinae, Tetramoriini: Ashmead, 1905b: 383.
Monomorium in Myrmicinae, Monomoriini: Wheeler, W.M. 1915h: 45; Dlussky & Fedoseeva, 1988: 80.
Monomorium in Myrmicinae, Monomorium genus group: Ettershank, 1966: 81; in Solenopsis genus group: Bolton, 1987: 271.
Monomorium in Myrmicinae, Solenopsidini: Emery, 1914a: 41 [subtribe Monomoriini]; Arnold, 1916: 203; Forel, 1917: 242; Emery, 1922e: 166; Wheeler, W.M. 1922a: 662; Hölldobler & Wilson, 1990: 16; Jaffe, 1993: 10; Bolton, 1994: 106.
Junior synonyms of MONOMORIUM
Phacota Roger, 1862b: 260. Type-species: Phacota sichelii, by monotypy.
Taxonomic history
Phacota in Myrmicidae, Pheidolidae: Emery, 1877a: 81.
Phacota in Myrmicinae: Mayr, 1865: 22 [Myrmicidae]; Emery & Forel, 1879a: 464 [Myrmicidae]; Dalla Torre, 1893: 71.
Phacota in Myrmicinae, Myrmicini: Emery, 1895j: 769; Wheeler, W.M. 1910g: 139.
Phacota in Myrmicinae, Stenammini: Ashmead, 1905b: 383.
Phacota in Myrmicinae, Solenopsidini: Emery, 1914a: 41 [subtribe Monomoriini]; Forel, 1917: 243; Emery, 1922e: 187; Wheeler, W.M. 1922a: 662; all subsequent authors except the entry below; Hölldobler & Wilson, 1990: 16; Bolton, 1994: 106.
Phacota in Myrmicinae, Solenopsis genus group: Bolton, 1987: 271.
Phacota as genus: Bolton, 1987: 281.
Phacota as junior synonym of Monomorium: Ettershank, 1966: 82; Fernández, in Heterick, 2006: 79; Fernández, 2007b: 130.
Trichomyrmex Mayr, 1865: 19. Type-species: Trichomyrmex rogeri, by monotypy.
Taxonomic history
Trichomyrmex in Myrmicidae, Myrmicidae: Emery, 1877a: 81.
Trichomyrmex in Myrmicinae: Mayr, 1865: 19 [Myrmicidae]; Dalla Torre, 1893: 72.
Trichomyrmex in Myrmicinae, Myrmicini: Wheeler, W.M. 1910g: 140.
Trichomyrmex in Myrmicinae, Solenopsidini: Emery, 1914a: 41 [subtribe Monomoriini]; Forel, 1917: 243; Emery, 1922e: 185; Wheeler, W.M. 1922a: 662; Donisthorpe, 1943g: 733.
Trichomyrmex as junior synonym of Monomorium: Ettershank, 1966: 82; Smith, D.R. 1979: 1381; Bolton, 1987: 296.
†Lampromyrmex Mayr, 1868c: 93. Type-species: †Monomorium mayrianum (replacement name for †Lampromyrmex gracillimus, junior secondary homonym in Monomorium), by monotypy.
Taxonomic history
†Lampromyrmex in Myrmicidae, Pheidolidae: Emery, 1877a: 81.
†Lampromyrmex in Myrmicinae: Mayr, 1868c: 93 [Myrmicidae]; Dalla Torre, 1893: 78.
†Lampromyrmex in Myrmicinae, Solenopsidini: Emery, 1922e: 183 [subtribe Monomoriini]; Donisthorpe, 1943f: 655.
†Lampromyrmex as junior synonym of Monomorium: Wheeler, W.M. 1915h: 45.
†Lampromyrmex as subgenus of Monomorium: Emery, 1922e: 183; Wheeler, W.M. 1922a: 676.
†Lampromyrmex as junior synonym of Monomorium: Wheeler, W.M. 1915h: 45; Ettershank, 1966: 82; Smith, D.R. 1979: 1381; Bolton, 1987: 299.
Holcomyrmex Mayr, 1879: 671. Type-species: Holcomyrmex scabriceps, by subsequent designation of Bingham, 1903: 280.
Taxonomic history
[Holcomyrmex Smith, F. 1873: ix, nomen nudum, attributed to Mayr.]
Holcomyrmex in Myrmicinae, Myrmicini: Emery, 1895j: 769; Ashmead, 1905b: 383.
Holcomyrmex in Myrmicinae, Solenopsidini: Emery, 1922e: 181 [subtribe Monomoriini]; Donisthorpe, 1943f: 650.
Holcomyrmex as genus: Dalla Torre, 1893: 65; Forel, 1903a: 692; Bingham, 1903: 280.
Holcomyrmex as subgenus of Monomorium: Wheeler, W.M. 1910g: 139; Emery, 1915i: 190; Forel, 1917: 242; Emery, 1922e: 181; Wheeler, W.M. 1922a: 676.
Holcomyrmex as junior synonym of Monomorium: Emery, 1908h: 667; Ettershank, 1966: 82; Smith, D.R. 1979: 1381; Bolton, 1987: 295.
Epoecus Emery, 1893a: cclxxvi. Type-species: Epoecus pergandei, by monotypy.
Taxonomic history
Epoecus in Myrmicinae, Myrmicini: Emery, 1895j: 769; Wheeler, W.M. 1910g: 139.
Epoecus in Myrmicinae, Stenammini: Ashmead, 1905b: 383.
Epoecus in Myrmicinae, Solenopsidini: Emery, 1914a: 41 [subtribe Anergatini]; Forel, 1917: 243; Emery, 1922e: 204; Wheeler, W.M. 1922a: 663; Donisthorpe, 1943f: 643.
Epoecus as junior synonym of Monomorium: Ettershank, 1966: 82; Smith, D.R. 1979: 1381; Bolton, 1987: 287.
Wheeleriella Forel, 1907f: 145.
Taxonomic history
[Replacement name for Wheeleria Forel, 1905b: 171; junior homonym of Wheeleria Tutt, 1905: 37 (Lepidoptera).]
Wheeleriella in Myrmicinae, Myrmicini: Wheeler, W.M. 1910g: 139.
Wheeleriella in Myrmicinae, Solenopsidini: Emery, 1914a: 41 [subtribe Monomoriini]; Forel, 1917: 243; Emery, 1922e: 186; Wheeler, W.M. 1922a: 662; Donisthorpe, 1943g: 735.
Wheeleriella as junior synonym of Monomorium: Ettershank, 1966: 82; Smith, D.R. 1979: 1381; Bolton, 1987: 291.
Homonym replaced by Wheeleriella
Wheeleria Forel, 1905b: 171. Type-species: Wheeleria santschii, by monotypy.
Taxonomic history
[Junior homonym of Wheeleria Tutt, 1905: 37 (Lepidoptera).]
Epixenus Emery, 1908f: 556. Type-species: Epixenus andrei (junior secondary homonym in Monomorium, replaced by Monomorium advena), by subsequent designation of Wheeler, W.M. 1911f: 163.
Taxonomic history
Epixenus in Myrmicinae, Solenopsidini: Emery, 1914a: 41 [subtribe Monomoriini]; Forel, 1917: 243; Emery, 1922e: 184; Wheeler, W.M. 1922a: 662; Donisthorpe, 1943f: 643.
Epixenus as junior synonym of Monomorium: Brown & Wilson, 1957bb: 244; Ettershank, 1966: 82; Smith, D.R. 1979: 1381; Bolton, 1987: 287; Tinaut & Ortiz, 1988: 167.
Mitara Emery, 1913b: 261 [as subgenus of Monomorium]. Type-species: Monomorium laeve, by original designation.
Taxonomic history
Mitara in Myrmicinae, Solenopsidini: Arnold, 1916: 238; Forel, 1917: 242 [subtribe Monomoriini]; all subsequent authors.
Mitara as subgenus of Monomorium: Emery, 1913b: 261; Emery, 1915i: 190; Arnold, 1916: 238; Forel, 1917: 242.
Mitara as junior synonym of †Lampromyrmex: Emery, 1922e: 183; Wheeler, W.M. 1922a: 162, 676.
Mitara as genus: Donisthorpe, 1943f: 662.
Mitara as junior synoynm of Monomorium: Smith, D.R. 1979: 1381; Bolton, 1987: 299.
Chelaner Emery, 1914f: 410 [as subgenus of Monomorium]. Type-species: Monomorium (Chelaner) forcipatum, by subsequent designation of Emery, 1922e: 168.
Taxonomic history
Chelaner in Myrmicinae, Solenopsidini: Forel, 1917: 242 [subtribe Monomoriini]; Emery, 1922e: 168; all subsequent authors except the following.
Chelaner in Myrmicinae, Monomorium genus group: Ettershank, 1966: 81.
Chelaner in Myrmicinae, Monomoriini: Dlussky & Fedoseeva, 1988: 80.
Chelaner as subgenus of Monomorium: Emery, 1914f: 410; Forel, 1917: 242; Emery, 1922e: 168; Wheeler, W.M. 1922a: 676.
Chelaner as genus: Ettershank, 1966: 93; Taylor & Brown, D.R. 1985: 55; Wheeler, G.C. & Wheeler, J. 1985: 257; Dlussky & Fedoseeva, 1988: 80 (anachronism).
Chelaner (and its junior synonyms Notomyrmex, Protholcomyrmex, Schizopelta) as junior synonym of Monomorium: Bolton, 1987: 300; Heterick, 2001: 354.
Notomyrmex Emery, 1915i: 190 [as subgenus of Monomorium]. Type-species: Atta antarctica, by original designation.
Taxonomic history
Notomyrmex as subgenus of Monomorium: Emery, 1915i: 190; Forel, 1917: 242; Emery, 1922e: 168; Wheeler, W.M. 1922a: 676; Kusnezov, 1956: 20.
Notomyrmex as genus: Kusnezov, 1957a: 269 (in key); Kusnezov, 1960b: 343; Kusnezov, 1962b: 160; Kusnezov, 1964: 60.
Notomyrmex as junior synonym of Chelaner: Ettershank, 1966: 93.
Parholcomyrmex Emery, 1915i: 190 [as subgenus of Monomorium]. Type-species: Myrmica gracillima (junior synonym of Atta destructor), by original designation.
Taxonomic history
Parholcomyrmex as subgenus of Monomorium: Emery, 1915i: 190; Forel, 1917: 242; Emery, 1922e: 179; Wheeler, W.M. 1922a: 676; subsequent authors to the following.
Parholcomyrmex as junior synonym of Monomorium: Ettershank, 1966: 82; Smith, D.R. 1979: 1381; Bolton, 1987: 296.
[Paraholcomyrmex Emery, 1915i: 191, Forel, 1917: 242, incorrect subsequent spellings.]
Xeromyrmex Emery, 1915i: 190 [as subgenus of Monomorium]. Type-species: Formica salomonis, by original designation.
Taxonomic history
Xeromyrmex as subgenus of Monomorium: Forel, 1917: 242; Emery, 1922e: 175; Wheeler, W.M. 1922a: 677.
Xeromyrmex as junior synonym of Monomorium: Ettershank, 1966: 82; Smith, D.R. 1979: 1381; Bolton, 1987: 294.
Syllophopsis Santschi, 1915c: 259 [as subgenus of Monomorium.] Type-species: Monomorium modestum, by monotypy.
Taxonomic history
Syllophopsis in Myrmicinae, Solenopsidini: Forel, 1917: 242 [subtribe Monomoriini]; Emery, 1922e: 175; all subsequent authors to the following.
Syllophopsis in Myrmicinae, Monomorium genus group: Ettershank, 1966: 81.
Syllophopsis in Myrmicinae, Monomoriini: Dlussky & Fedoseeva, 1988: 80.
Syllophopsis as subgenus of Monomorium: Santschi, 1915c: 259; Forel, 1917: 242; Emery, 1922e: 175; Wheeler, W.M. 1922a: 676; Donisthorpe, 1943g: 729; Arnold, 1952a: 465.
Syllophopsis as genus: Santschi, 1921c: 119; Santschi, 1936a: 32; Ettershank, 1966: 100.
Syllophopsis as junior synonym of Monomorium: Bolton, 1987: 287.
[Syllopsis Santschi, 1921c: 120, incorrect subsequent spelling.]
Corynomyrmex Viehmeyer, 1916a: 133 [as subgenus of Monomorium]. Type-species: Monomorium (Corynomyrmex) hospitum, by monotypy.
Taxonomic history
Corynomyrmex as subgenus of Monomorium: Viehmeyer, 1916a: 133; Forel, 1917: 242; Emery, 1922e: 174.
Corynomyrmex as junior synonym of Monomorium: Ettershank, 1966: 82 [provisional]; Smith, D.R. 1979: 1381; Bolton, 1987: 287.
Isolcomyrmex Santschi, 1917b: 296 [as subgenus of Monomorium]. Type-species: Holcomyrmex santschii (junior secondary homonym in Monomorium, replaced by Monomorium santschianum), by monotypy.
Taxonomic history
Isolcomyrmex as subgenus of Monomorium: Santschi, 1917b: 296; Wheeler, W.M. 1922a: 676.
Isolcomyrmex as junior synonym of Monomorium: Ettershank, 1966: 82; Smith, D.R. 1979: 1381; Bolton, 1987: 296.
[Isholcomyrmex Santschi, 1936a: 32, incorrect subsequent spelling.]
Paraphacota Santschi, 1919d: 90. Type-species: Paraphacota surcoufi, by monotypy.
Taxonomic history
[Type-species not Phacota noualhieri, unjustified subsequent designation by Santschi, 1927d: 245; repeated in Ettershank, 1966: 82 and Bolton, 1973a: 352.]
Paraphacota in Myrmicinae, Solenopsidini: Wheeler, W.M. 1922a: 663; Donisthorpe, 1943f: 681.
Paraphacota as junior synonym of Monomorium: Santschi, 1927d: 243; Ettershank, 1966: 82; Smith, D.R. 1979: 1381; Bolton, 1987: 293.
Equestrimessor Santschi, 1919d: 92 [as subgenus of Monomorium]. Type-species: Holcomyrmex chobauti, by subsequent designation of Donisthorpe, 1943f: 644.
Taxonomic history
Equestrimessor as subgenus of Monomorium: Santschi, 1919d: 92;Wheeler, W.M. 1922a: 677; Santschi, 1936a: 32.
Equestrimessor as junior synonym of Monomorium: Ettershank, 1966: 82; Smith, D.R. 1979: 1381; Bolton, 1987: 287, 297.
[Equessimessor Santschi, 1936a: 32, incorrect subsequent spelling.]
Xenhyboma Santschi, 1919g: 405. Type-species: Xenhyboma mystes (junior synonym of Monomorium medinae), by monotypy.
Taxonomic history
Xenhyboma in Myrmicinae, Solenopsidini: Donisthorpe, 1943g: 736.
Xenhyboma as junior synonym of Monomorium: Ettershank, 1966: 82 [provisional]; Smith, D.R. 1979: 1381; Espadaler, 1982b: 112; Bolton, 1987: 293.
Protholcomyrmex Wheeler, W.M. 1922a: 162 [as subgenus of Monomorium]. Type-species: Monomorium rothsteini, by original designation.
Taxonomic history
Protholcomyrmex as junior synonym of Chelaner: Ettershank, 1966: 93.
Ireneidris Donisthorpe, 1943c: 81. Type-species: Ireneidris myops (junior synonym of Monomorium talpa), by original designation.
Taxonomic history
Ireneidris in Myrmicinae, Solenopsidini [subtribe Monomoriini]: Donisthorpe, 1943c: 82.
Ireneidris as junior synonym of Monomorium: Ettershank, 1966: 82; Smith, D.R. 1979: 1381; Bolton, 1987: 298.
Schizopelta McAreavey, 1949: 14. Type-species: Schizopelta falcata, by original designation.
Taxonomic history
Schizopelta as junior synonym of Chelaner: Ettershank, 1966: 93.
Nothidris Ettershank, 1966: 105. Type-species: Monomorium latastei, by original designation.
Taxonomic history
Nothidris in Myrmicinae, Megalomyrmex genus group: Ettershank, 1966: 81; in Solenopsis genus group: Bolton, 1987: 271.
Nothidris in Myrmicinae, Megalomyrmecini: Dlussky & Fedoseeva, 1988: 80.
Nothidris in Myrmicinae, Solenopsidini: Kempf, 1972a: 165; Hölldobler & Wilson, 1990: 16; Jaffe, 1993: 10; Bolton, 1994: 106.
Nothidris as junior synonym of Monomorium: Fernández, in Heterick, 2006: 79; Fernández, 2007b: 130.
Pharaophanes Bernard, 1967: 168 (attributed to Santschi) [as subgenus of Monomorium]. Type-species: Formica pharaonis, by monotypy.
Taxonomic history
[Pharaophanes Bernard, 1953b: 238 (attributed to Santschi) [as subgenus of Monomorium]. Unavailable name; proposed without designation of type-species. Species included by Bernard (1953) are all referable to Monomorium: Bolton, 1987: 288; Bolton, 1995b: 42 (see also note in Baroni Urbani, 1971c: 90).]
Pharaophanes as junior synonym of Monomorium: Bolton, 2003: 213.
Antichthonidris Snelling, R.R. 1975: 5. Type-species: Monomorium denticulatum, by original designation.
Taxonomic history
Antichthonidris in Myrmicinae, Solenopsis genus group: Bolton, 1987: 271.
Antichthonidris in Myrmicinae, Solenopsidini: Hölldobler & Wilson, 1990: 16; Jaffe, 1993: 10; Bolton, 1994: 106.
Antichthonidris as junior synonym of Monomorium: Heterick, 2001: 361.
Epelysidris Bolton, 1987: 279. Type-species: Epelysidris brocha, by original designation.
Taxonomic history
Epelysidris in Myrmicinae, Solenopsis genus group: Bolton, 1987: 271.
Epelysidris in Myrmicinae, Solenopsidini: Hölldobler & Wilson, 1990: 16; Bolton, 1994: 106; Bolton, 1995a: 1049; Bolton, 1995b: 188.
Epelysidris as junior synonym of Monomorium: Fernández, in Heterick, 2006: 79; Fernández, 2007b: 130.
Genus Monomorium references
Roger, 1863b: 30, 31 (Phacota, Monomorium catalogues); Mayr, 1863: 429, 440 (Monomorium, Phacota catalogues); Mayr, 1865: 19, 22 (Trichomyrmex, Monomorium, Phacota diagnoses); Mayr, 1867a: 95 (Monomorium diagnosis); Mayr, 1876: 100 (Australia Monomorium species key); Emery, 1881b: 530 (Mediterranean & Red Sea Monomorium species key); André, 1883a: 331, 343 (Europe & Algeria Monomorium, Holcomyrmex species keys); André, 1883b: 397 (Phacota); Cresson, 1887: 262 (U.S.A. Monomorium catalogue); Nasonov, 1889: 69 (Russia Monomorium species key); Dalla Torre, 1893: 65, 71, 72, 78 (Holcomyrmex, Monomorium, Phacota, Trichomyrmex, †Lampromyrmex catalogues); Forel, 1903a: 685, 692 (India & Sri Lanka Monomorium, Holcomyrmex species keys); Bingham, 1903: 200, 282 (India, Sri Lanka & Burma Monomorium, Holcomyrmex species keys); Ruzsky, 1905b: 633 (Russian Empire Monomorium species key); Emery, 1908h: 664 (Palaearctic Monomorium species key); Emery, 1915i: 190 (Monomorium subgenera key); Arnold, 1916: 203, 204 (Monomorium diagnosis, South Africa species key); Emery, 1916b: 161 (Italy Monomorium species key); Santschi, 1921c: 121 (Syllophopsis species key); Emery, 1922e: 166 (Monomorium diagnosis, subgenera key, catalogue); Emery, 1922e: 168 (M. (Chelaner) & M. (Notomyrmex) diagnoses, catalogues); Emery, 1922e: 170 (M. (Monomorium) diagnosis, catalogue); Emery, 1922e: 174 (M. (Corynomyrmex) diagnosis, catalogue); Emery, 1922e: 175 (M. (Syllophopsis) & M. (Xeromyrmex) diagnoses, catalogues); Emery, 1922e: 179 (M. (Parholcomyrmex) diagnosis, catalogue); Emery, 1922e: 181 (M. (Holcomyrmex) diagnosis, catalogue); Emery, 1922e: 183 (M. (Lampromyrmex) diagnosis, catalogue); Emery, 1922e: 184 (Epixenus diagnosis, catalogue); Emery, 1922e: 185 (Trichomyrmex diagnosis, catalogue); Emery, 1922e: 186 (Wheeleriella diagnosis, catalogue); Emery, 1922e: 187 (Phacota diagnosis, catalogue); Emery, 1922e: 204 (Epoecus diagnosis, catalogue); Wheeler, W.M. 1922a: 161, 675, 862, 1026 (Monomorium diagnosis, subgenera key, Afrotropical, Malagasy catalogues); Menozzi, 1933b: 64 (Israel Monomorium species key); Santschi, 1936a: 33 (Mediterranean & Africa M. (Xeromyrmex) key); Buren, 1944a: 289 (U.S.A., Iowa Monomorium species key); Kusnezov, 1949a: 431 (Argentina Monomorium species); Creighton, 1950a: 217 (North America Monomorium species key); Creighton, 1950a: 239 (Epoecus, review of genus); Chapman & Capco, 1951: 162, 169 (Asia Ireneidris, Monomorium, Trichomyrmex, Wheeleriella checklists); Bernard, 1955b: 282 (Epixenus species key); Brown, 1958h: 28 (New Zealand Monomorium species key); Gregg, 1963: 367 (U.S.A., Colorado Monomorium species key); Ettershank, 1966: 82, 93, 100 (Monomorium, Chelaner, Syllophopsis diagnoses, reviews of genera, checklists); Bernard, 1967: 165 (Monomorium diagnosis, Western Europe species key); Kempf, 1972a: 143 (Neotropical Monomorium catalogue); Kempf, 1972a: 165 (Nothidris catalogue); Alayo, 1974: 14 (Cuba Monomorium species key); Snelling, R.R. 1975: 5 (Nothidris all species key); Snelling, R.R. & Hunt, 1976: 79 (Nothidris all species key); Arnol'di & Dlussky, 1978: 538 (former European U.S.S.R. Monomorium species key); Collingwood, 1978: 82 (Iberian Peninsula Monomorium species key); Smith, D.R. 1979: 1381 (North America Monomorium catalogue); Tohmé & Tohmé, 1980a: 1087 (Lebanon & Syria Epixenus species key); Collingwood, 1985: 267 (Saudi Arabia Monomorium species key); Taylor & Brown, D.R. 1985: 55, 70 (Australia Chelaner, Monomorium catalogues); DuBois, 1986: 74 (Nearctic Monomorium species revision, key); Bolton, 1987: 281 (Phacota diagnosis, review of genus); Bolton, 1987: 283 (Antichthonidris, review of genus); Bolton, 1987: 284 (Nothidris review of genus); Bolton, 1987: 287 (Monomorium diagnosis, review of genus, Afrotropical species revision, key); Taylor, 1987a: 19, 40 and Taylor, 1987b: 2 (Australia, New Caledonia & New Zealand Monomorium checklist); Agosti & Collingwood, 1987b: 272 (Balkans Monomorium species key); Hölldobler & Wilson, 1990: 14 (synoptic classification); Dlussky, Soyunov & Zabelin, 1990: 231 (Turkmenistan Monomorium species key); Brandão, 1991: 326, 357 (Neotropical Monomorium catalogue); Brandão, 1991: 361 (Nothidris catalogue); Morisita, Kubota, et al. 1992: 38 (Japan Monomorium species key); Arakelian, 1994: 46 (Armenia Monomorium species key); Bolton, 1995a: 1048, 1050, 1051 (census); Bolton, 1995b: 67, 258, 292, 316 (Antichthonidris, Monomorium, Nothidris, Phacota catalogues); Wu, J. & Wang, 1995: 88 (China Monomorium species key); Collingwood & Agosti, 1996: 337 (Saudi Arabia Monomorium species key); Radchenko, 1997f: 211 (M. scabriceps group revision, key); Collingwood & Prince, 1998: 19 (Portugal Monomorium species key); Shattuck, 1999: 145 (Australia synopsis); Heterick, 2001: 367 (Australia Monomorium species revision, key); Zhou, 2001b: 112 (China, Guangxi Monomorium species key); Mackay & Mackay, 2002: 127 (U.S.A., New Mexico Monomorium species key); Heterick, 2006: 84 (Malagasy Monomorium species revision, key); Fernández, 2007b: 139 (Neotropical Monomorium species key); Terayama, 2009: 154 (Taiwan species key); Heterick, 2009: 158 (south-western Australia species key); Aldawood & Sharaf, 2011: 52 (Saudi Arabia M. monomorium group key).
Myrmicinae Tetramoriini Rhoptromyrmex TRUE TRUE FALSE10 species
Rhoptromyrmex Mayr, 1901b: 18. Type-species: Rhoptromyrmex globinodis, by subsequent designation of Wheeler, W.M. 1911f: 172.
Taxonomic history
Rhoptromyrmex in Myrmicinae, Stenammini: Ashmead, 1905b: 383.
Rhoptromyrmex in Myrmicinae, Tetramoriini: Wheeler, W.M. 1910g: 141; Emery, 1914a: 42; Forel, 1917: 245; Arnold, 1917: 351; Wheeler, W.M. 1922a: 664; Emery, 1924d: 289; all subsequent authors.
Junior synonyms of RHOPTROMYRMEX
Hagioxenus Forel, 1910a: 8. Type-species: Hagioxenus schmitzi, by monotypy.
Taxonomic history
Hagioxenus in Myrmicinae, Solenopsidini: Emery, 1914a: 41 [subtribe Monomoriini]; Forel, 1917: 243; Emery, 1922e: 186; Wheeler, W.M. 1922a: 662; subsequent authors to the following.
Hagioxenus in Myrmicinae, Monomorium genus group: Ettershank, 1966: 81.
Hagioxenus as junior synonym of Rhoptromyrmex: Bolton, 1986a: 2.
Acidomyrmex Emery, 1915i: 191 [as subgenus of Rhoptromyrmex]. Type-species: Rhoptromyrmex wroughtonii, by original designation.
Taxonomic history
Acidomyrmex in Myrmicinae, Tetramoriini: Forel, 1917: 245; Wheeler, W.M. 1922a: 664; Emery, 1924d: 290; all subsequent authors.
Acidomyrmex as genus: Wheeler, W.M. 1922a: 664, 672; Chapman & Capco, 1951: 173.
Acidomyrmex as subgenus of Rhoptromyrmex: Emery, 1915i: 191; Forel, 1917: 245; Emery, 1924d: 290; Donisthorpe, 1943f: 619.
Acidomyrmex as junior synonym of Rhoptromyrmex: Brown, 1964c: 11; Bolton, 1976: 298; Bolton, 1986a: 2.
Ireneella Donisthorpe, 1941i: 175. Type-species: Ireneella papuensis (junior synonym of Tetramorium melleum), by original designation.
Taxonomic history
Ireneella in Myrmicinae, Tetramoriini: Donisthorpe, 1941i: 175. Donisthorpe, 1943f: 653.
Ireneella incertae sedis in Formicidae: Wheeler, G.C. & Wheeler, J. 1985: 259 (incomprehensible entry).
Ireneella as junior synonym of Rhoptromyrmex: Bolton, 1976: 298; Bolton, 1986a: 2.
Genus Rhoptromyrmex references
Arnold, 1917: 351 (diagnosis, South Africa species key); Wheeler, W.M. 1922a: 194, 908 (diagnosis, Afrotropical catalogue); Emery, 1922e: 186 (Hagioxenus diagnosis, catalogue); Emery, 1924d: 289 (Rhoptromyrmex diagnosis, catalogue); Emery, 1924d: 290 (R. (Acidomyrmex) diagnosis, catalogue); Chapman & Capco, 1951: 173, 174 (Asia Acidomyrmex, Ireneela, Rhoptromyrmex checklists); Brown, 1964c: 11 (diagnosis, all species revision, key); Bolton, 1976: 298 (diagnosis, all species key); Taylor & Brown, D.R. 1985: 86 (Australia catalogue); Bolton, 1986a: 5 (diagnosis, review of genus, all species key); Taylor, 1987a: 67 (Australia checklist); Bolton, 1994: 106 (synoptic classification); Bolton, 1995a: 1052 (census); Bolton, 1995b: 377 (catalogue); Shattuck, 1999: 163 (Australia synopsis).
Myrmicinae Tetramoriini Tetramorium TRUE TRUE FALSEExtant: 461 species, 16 subspecies
Fossil:
Tetramorium Mayr, 1855: 423. Type-species: Formica caespitum, by subsequent designation of Girard, 1879: 1016.
Taxonomic history
Tetramorium in Myrmicidae, Myrmicidae: Emery, 1877a: 81.
Tetramorium in Myrmicinae: Mayr, 1855: 423 [Myrmicidae]; Mayr, 1861: 61 [Myrmicidae]; Mayr, 1865: 20 [Myrmicidae]; Emery & Forel, 1879a: 457 [Myrmicidae]; Dalla Torre, 1893: 130.
Tetramorium in Myrmicinae, Myrmicini: Forel, 1895b: 125; Forel, 1899c: 52; Kusnezov, 1964: 57 (anachronism).
Tetramorium in Myrmicinae, Tetramoriini: Emery, 1895j: 770; Ashmead, 1905b: 383; Wheeler, W.M. 1910g: 141; Emery, 1914a: 42; Arnold, 1917: 271; Forel, 1917: 245; Wheeler, W.M. 1922a: 664; Emery, 1924d: 275; all subsequent authors.
Junior synonyms of TETRAMORIUM
Tetrogmus Roger, 1857: 10. Type-species: Tetrogmus caldarius, by monotypy.
Taxonomic history
Tetrogmus as subgenus of Tetramorium: Wheeler, W.M. 1910g: 141.
Tetrogmus as junior synonym of Tetramorium: Roger, 1862c: 297; all subsequent authors except the above; Bolton, 1976: 359; Bolton, 1980: 195.
Xiphomyrmex Forel, 1887: 385 [as subgenus of Tetramorium]. Type-species: Tetramorium (Xiphomyrmex) kelleri, by subsequent designation of Wheeler, W.M. 1911f: 175.
Taxonomic history
Xiphomyrmex in Myrmicinae: Dalla Torre, 1893: 130.
Xiphomyrmex in Myrmicinae, Myrmicini: Ashmead, 1905b: 383.
Xiphomyrmex in Myrmicinae, Tetramoriini: Emery, 1895j: 770; Emery, 1914a: 42; Forel, 1917: 245; Wheeler, W.M. 1922a: 664; Emery, 1924d: 286; all subsequent authors.
Xiphomyrmex as subgenus of Tetramorium: Dalla Torre, 1893: 130; Forel, 1899c: 53; Forel, 1903a: 700; Wheeler, W.M. 1910g: 141; Arnold, 1917: 274; Arnold, 1926: 277.
Xiphomyrmex as genus: Emery, 1895j: 770; Emery, 1896e: 183; Emery, 1914a: 42; Forel, 1917: 245; Wheeler, W.M. 1922a: 193; Emery, 1924d: 286; Smith, M.R. 1938a: 126; Donisthorpe, 1943g: 736; Creighton, 1950a: 293; Smith, M.R. 1951a: 824; Chapman & Capco, 1951: 179; Bernard, 1953b: 250; Wheeler, G.C. & Wheeler, J. 1985: 257 (anachronism); Dlussky & Fedoseeva, 1988: 80 (anachronism).
Xiphomyrmex as junior synonym of Tetramorium: Bingham, 1903: 175; Bolton, 1976: 359; Bolton, 1980: 195; Bolton, 1994: 106.
Triglyphothrix Forel, 1890b: cvi. Type-species: Triglyphothrix walshi, by monotypy.
Taxonomic history
Triglyphothrix in Myrmicinae: Dalla Torre, 1893: 135.
Triglyphothrix in Myrmicinae, Tetramoriini: Emery, 1895j: 770; Ashmead, 1905b: 383; Wheeler, W.M. 1910g: 141; Emery, 1914a: 42; Forel, 1917: 245; Wheeler, W.M. 1922a: 664; Emery, 1924d: 272; all subsequent authors.
Triglyphothrix as subgenus of Tetramorium: Arnold, 1917: 274; Arnold, 1926: 272.
Triglyphothrix as genus: Forel, 1890b: cvi. Dalla Torre, 1893: 135; Emery, 1895j: 770; Forel, 1903a: 703; Bingham, 1903: 171; Wheeler, W.M. 1910g: 141; Emery, 1914a: 42; Forel, 1917: 245; Wheeler, W.M. 1922a: 196; Emery, 1924d: 272; Donisthorpe, 1943g: 734; Creighton, 1950a: 285; Smith, M.R. 1951a: 823; Chapman & Capco, 1951: 178; Bernard, 1953b: 248; Kempf, 1972a: 255; Bolton, 1976: 310; Smith, D.R. 1979: 1399; Wheeler, G.C. & Wheeler, J. 1985: 257; Taylor & Brown, D.R. 1985: 91.
Triglyphothrix as junior synonym of Tetramorium: Bolton, 1985: 247; Bolton, 1994: 106.
Atopula Emery, 1912b: 104. Type-species: Atopomyrmex nodifer, by original designation.
Taxonomic history
Atopula in Myrmicinae, Myrmecinini: Emery, 1912b: 105; Emery, 1914a: 41; Forel, 1917: 244; Wheeler, W.M. 1922a: 663; Emery, 1924d: 242 [subtribe Podomyrmini]; all subsequent authors to Bolton, 1976: 359.
Atopula in Myrmicinae, Podomyrmini: Dlussky & Fedoseeva, 1988: 79.
Atopula as subgenus of Vollenhovia: Emery, 1912d: 273.
Atopula as subgenus of Terataner: Arnold, 1952b: 129.
Atopula as genus: Emery, 1912b: 104; Emery, 1914a: 41; Wheeler, W.M. 1922a: 663; Emery, 1924d: 242; Bernard, 1948: 177; Wheeler, G.C. & Wheeler, J. 1985: 257 (anachronism); Dlussky & Fedoseeva, 1988: 79 (anachronism).
Atopula as junior synonym of Tetramorium: Bolton, 1976: 359; Bolton, 1980: 195; Bolton, 1994: 106.
Macromischoides Wheeler, W.M. 1920: 53. Type-species: Macromischa aculeata, by original designation.
Taxonomic history
Macromischoides in Myrmicinae, Leptothoracini: Wheeler, W.M. 1922a: 664; Wheeler, G.C. & Wheeler, J. 1985: 257 (anachronism).
Macromischoides in Myrmicinae, Tetramoriini: Santschi, 1924b: 207; all subsequent authors.
Macromischoides as genus: Wheeler, W.M. 1922a: 187; Santschi, 1924b: 206; Bernard, 1953b: 248; Dlussky & Fedoseeva, 1988: 80 (anachronism).
Macromischoides as junior synonym of Tetramorium: Bolton, 1976: 359; Bolton, 1980: 196.
[Macromichoides Santschi, 1924b: 206, incorrect subsequent spelling.]
Lobomyrmex Kratochvíl, 1941: 84 [as subgenus of Tetramorium]. Type-species: Tetramorium (Lobomyrmex) ferox silhavyi (junior synonym of Tetramorium ferox), by monotypy.
Taxonomic history
Lobomyrmex as junior synonym of Tetramorium: Brown, 1973b: 181 [provisional]; Bolton, 1976: 359; Bolton, 1980: 196.
Sulcomyrmex Kratochvíl, 1941: 84 [as subgenus of Tetramorium]. Unavailable name. Proposed without designation of type-species and therefore unavailable. Species included by Kratochvíl (1941) are all referable to Tetramorium: Bolton, 1976: 359.]
Apomyrmex Calilung, 2000: 66. Type-species: Apomyrmex manobo, by original designation.
Taxonomic history
Apomyrmex incertae sedis in Myrmicinae: Calilung, 2000: 66.
Apomyrmex as junior synonym of Tetramorium: Bolton, 2003: 227, 269.
Genus Tetramorium references
Roger, 1863b: 26, 27 (catalogue); Mayr, 1863: 456 (catalogue); Mayr, 1865: 20 (diagnosis); Mayr, 1870b: 972 (all species key); André, 1883a: 285 (Europe & Algeria species key); Cresson, 1887: 261 (U.S.A. catalogue); Emery, 1893e: 214 (Triglyphothrix species key); Dalla Torre, 1893: 130, 135 (Tetramorium, Triglyphothrix catalogues); Forel, 1903a: 700, 703 (India & Sri Lanka Tetramorium, Triglyphothrix species keys); Bingham, 1903: 172, 175 (India, Sri Lanka & Burma Triglyphothrix, Tetramorium species keys); Ruzsky, 1905b: 517 (Russian Empire species key); Emery, 1909d: 695 (Palaearctic species key); Emery, 1916b: 195 (Italy species key); Arnold, 1917: 271, 275, 334, 345 (diagnosis, South Africa Tetramorium (Tetramorium), T. (Triglyphothrix), T. (Xiphomyrmex) species keys); Bondroit, 1918: 106 (France & Belgium species key); Mann, 1919: 353 (Papuasia Triglyphothrix species key); Wheeler, W.M. 1922a: 187, 190, 193, 196 (Macromischoides, Tetramorium, Xiphomyrmex, Triglyphothrix diagnoses); Wheeler, W.M. 1922a: 886, 889, 893, 906, 909 (Afrotropical Atopula, Macromischoides, Tetramorium, Xiphomyrmex, Triglyphothrix catalogues); Wheeler, W.M. 1922a: 1030, 1031, 1032 (Malagasy Tetramorium, Xiphomyrmex, Triglyphothrix catalogues); Emery, 1924d: 242 (Atopula diagnosis, catalogue); Emery, 1924d: 272 (Triglyphothrix diagnosis, catalogue); Emery, 1924d: 275 (Tetramorium diagnosis, catalogue); Emery, 1924d: 286 (Xiphomyrmex diagnosis, catalogue); Santschi, 1924b: 210 (Macromischoides species key); Arnold, 1926: 241 (South Africa Tetramorium (Tetramorium) species key); Menozzi, 1933b: 74 (Israel species key); Finzi, 1936: 183 (Egypt species key); Smith, M.R. 1938a: 127 (North America Xiphomyrmex species key); Kratochvíl, 1941: 84 (Central Europe species key); Novák & Sadil, 1941: 84 (Central Europe species key); Smith, M.R. 1943b: 2 (North America species key); Kratochvíl, Novák & Snoflák, 1944: 76, 95 (Central Europe species key); Creighton, 1950a: 290, 293 (North America Tetramorium, Xiphomyrmex species key); Chapman & Capco, 1951: 174, 178, 179 (Asia Tetramorium, Triglyphothrix, Xiphomyrmex checklists); Brown, 1958h: 27 (New Zealand species); Bernard, 1967: 227 (diagnosis, Western Europe species key); Kempf, 1972a: 249, 255 (Neotropical Tetramorium, Triglyphothrix catalogues); Alayo, 1974: 18 (Cuba species key); Tarbinsky, 1976: 105 (Kyrghyzstan species key); Bolton, 1976: 310, 314, 341 (Triglyphothrix diagnosis, Afrotropical, Oriental & Malesian species revisions, keys); Bolton, 1976: 359 (Tetramorium diagnosis, review of genus, synonymy); Bolton, 1977: 72, 133 (Oriental & Malesian, Australia species revisions, keys); Kutter, 1977c: 150 (Switzerland species key); Arnol'di & Dlussky, 1978: 544 (former European U.S.S.R. species key); Collingwood, 1978: 86 (Iberian Peninsula species key); Collingwood, 1979: 82 (Fennoscandia & Denmark species key); Smith, D.R. 1979: 1399, 1400 (North America Triglyphothrix, Tetramorium catalogues); Bolton, 1979: 132, 159 (Malagasy, New World species revisions, keys); Bolton, 1980: 196, 205 (diagnosis, review of genus, Afrotropical species revision, key); Collingwood, 1985: 262 (Saudi Arabia species key); Gösswald, 1985: 311 (Germany species key); Taylor & Brown, D.R. 1985: 88 (Australia catalogue); Taylor, 1987a: 78 (Australia, New Caledonia & New Zealand checklist); Agosti & Collingwood, 1987b: 277 (Balkans species key); Wang, M., Xiao & Wu, 1988: 264 (China species key); Radchenko & Arakelian, 1990: 371 (Crimea & Caucasus T. ferox complex, key); Dlussky, Soyunov & Zabelin, 1990: 197 (Turkmenistan species key); Atanasov & Dlussky, 1992: 149 (Bulgaria species key); Radchenko, 1992a: 44 (former U.S.S.R. species key); Morisita, Kubota, et al. 1992: 33 (Japan species key); Arakelian, 1994: 60 (Armenia species key); Radchenko, 1994b: 109 (South Siberia species key); Bolton, 1994: 106 (synoptic classification); Bolton, 1995a: 1053 (census); Bolton, 1995b: 403 (catalogue); Wu, J. & Wang, 1995: 79 (China species key); Collingwood & Agosti, 1996: 332 (Saudi Arabia species key); Seifert, 1996b: 158 (Central Europe species key); Cagniant, 1997: 89 (Morocco species key); Radchenko, Czechowski & Czechowsa, 1998: 107 (Poland species key); Collingwood & Prince, 1998: 17 (Portugal species key); Sanetra, Güsten & Schulz, 1999: 317 (Italy, species); Shattuck, 1999: 168 (Australia synopsis); Chang & He, 2001a: 1 (northwest China species key); Zhou, 2001b: 100 (China, Guangxi species key); Czechowski, Radchenko & Czechowska, 2002: 145 (Poland species key); Mackay & Mackay, 2002: 214 (U.S.A., New Mexico species key); Seifert, 2007: 145 (North and Central Europe species key); Csösz, Radchenko & Schulz, 2007: 8 (T. chefketi complex key); Terayama, 2009: 162 (Taiwan species key); Heterick, 2009: 177 (south-western Australia species key); Csösz & Schulz, 2010: 12 (T. ferox complex key); Hita Garcia, Fischer & Peters, 2010b: 12 (Afrotropical T. weitzeckeri species group key).
Myrmicinae Pheidolini Ocymyrmex TRUE TRUE FALSE37 species
Ocymyrmex Emery, 1886: 364. Type-species: Ocymyrmex barbiger, by monotypy.
Taxonomic history
Ocymyrmex in Myrmicinae: Dalla Torre, 1893: 72.
Ocymyrmex in Myrmicinae, Myrmicini: Emery, 1895j: 769; Wheeler, W.M. 1910g: 139.
Ocymyrmex in Myrmicinae, Ocymyrmecini: Emery, 1914a: 41; Arnold, 1916: 194; Forel, 1917: 245; Wheeler, W.M. 1922a: 664; Emery, 1924d: 270; all subsequent authors to the following.
Ocymyrmex in Myrmicinae, Pheidolini: Bolton & Marsh, 1989: 1281; Hölldobler & Wilson, 1990: 16; Bolton, 1994: 106
Genus Ocymyrmex references
Emery, 1892a: 116 (all species key); Arnold, 1916: 194 (diagnosis, South Africa species key); Wheeler, W.M. 1922a: 891 (catalogue); Emery, 1924d: 270 (diagnosis, catalogue); Bolton, 1981b: 260 (diagnosis, all species revision, key); Bolton & Marsh, 1989: 1267 (diagnosis, revised key); Bolton, 1995a: 1051 (census); Bolton, 1995b: 294 (catalogue).
Myrmicinae Pheidolini Pheidole TRUE TRUE FALSEExtant: 970 species, 142 subspecies
Fossil: 4 species
Pheidole Westwood, 1839: 219. Type-species: Atta providens, by monotypy.
Taxonomic history
Pheidole in Poneridae, Attidae: Smith, F. 1858b: 172.
Pheidole in Attidae: Smith, F. 1860a: 74; Smith, F. 1862a: 49.
Pheidole in Myrmicidae: Smith, F. 1871a: 331; Cresson, 1887: 262.
Pheidole in Myrmicidae, Pheidolidae: Emery, 1877a: 81.
Pheidole in Myrmicinae: Mayr, 1861: 69 [Myrmicidae]; Mayr, 1865: 22 [Myrmicidae]; Emery & Forel, 1879a: 463 [Myrmicidae]; Dalla Torre, 1893: 88.
Pheidole in Myrmicinae, Myrmicini: Forel, 1895b: 127; Emery, 1895j: 769; Forel, 1899c: 64; Ashmead, 1905b: 383; Wheeler, W.M. 1910g: 139; Kempf, 1972a: 183 (anachronism).
Pheidole in Myrmicinae, Pheidolini: Forel, 1893a: 165; Emery, 1914a: 40; Forel, 1917: 241; Arnold, 1920a: 414; Emery, 1921f: 77; Wheeler, W.M. 1922a: 661; all subsequent authors except Kempf, 1972a: 183, above.
Junior synonyms of PHEIDOLE
Oecophthora Heer, 1852: 15. Type-species: Oecophthora pusilla (junior synonym of Formica megacephala), by monotypy.
Taxonomic history
Oecophthora in Myrmicinae: Mayr, 1855: 453 [Myrmicidae].
Oecophthora as junior synonym of Pheidole: Smith, F. 1858a: 282; Smith, F. 1858b: 172; Roger, 1863b: 30.
Ischnomyrmex Mayr, 1862: 738. Type-species: Myrmica longipes, by monotypy.
Taxonomic history
Ischnomyrmex in Myrmicinae: Mayr, 1865: 18 [Myrmicidae]; Dalla Torre, 1893: 98.
Ischnomyrmex in Myrmicinae, Myrmicini: Emery, 1895j: 769; Ashmead, 1905b: 383; Wheeler, W.M. 1910g: 140.
Ischnomyrmex in Myrmicinae, Pheidolini: Emery, 1914a: 40; Forel, 1917: 241; Emery, 1922e: 113; Wheeler, W.M. 1922a: 661; all subsequent authors.
Ischnomyrmex as junior synonym of Aphaenogaster: Emery & Forel, 1879a: 461; Bingham, 1903: 270.
Ischnomyrmex as subgenus of Aphaenogaster: Forel, 1891b: 166; Dalla Torre, 1893: 98; Forel, 1899c: 59; Emery, 1908c: 313; Wheeler, W.M. 1910g: 140.
Ischnomyrmex as subgenus of Stenamma: Emery, 1895c: 298; Emery, 1895j: 769; Forel, 1902h: 439; Forel, 1903a: 693.
Ischnomyrmex as subgenus of Pheidole: Forel, 1913k: 49.
Ischnomyrmex as genus: Mayr, 1862: 738; Emery, 1914a: 40; Viehmeyer, 1914d: 601; Forel, 1917: 241; Wheeler, W.M. 1922a: 661; Emery, 1922e: 113. Stitz, 1911a: 367; Kusnezov, 1952b: 10 (in key).
Ischnomyrmex as junior synonym of Pheidole: Brown, 1973b: 181 [provisional]; Smith, D.R. 1979: 1365; Snelling, R.R. 1981: 395.
Leptomyrma Motschoulsky, 1863: 17. Type-species: Leptomyrma gracilipes, by monotypy.
Taxonomic history
Leptomyrma in Myrmicidae: Smith, F. 1871a: 329.
Leptomyrma as junior synonym of Pheidole: Emery, 1892b: 166; all subsequent authors.
Pheidolacanthinus Smith, F. 1865: 75. Type-species: Pheidolacanthinus armatus (junior synonym of Myrmica quadrispinosa), by monotypy.
Taxonomic history
Pheidolacanthinus in Myrmicidae: Smith, F. 1871a: 333.
Pheidolacanthinus in Myrmicinae: Dalla Torre, 1893: 98.
Pheidolacanthinus as subgenus of Pheidole: Emery, 1915i: 190; Forel, 1917: 241; Emery, 1921f: 81; Wheeler, W.M. 1922a: 673; all subsequent authors to the following.
Pheidolacanthinus as junior synonym of Pheidole: Brown, 1973b: 183 [provisional]; Smith, D.R. 1979: 1365; Snelling, R.R. 1981: 395.
Ceratopheidole Pergande, 1896: 889 [as subgenus of Pheidole]. Type-species: Pheidole (Ceratopheidole) granulata, by monotypy.
Taxonomic history
Ceratopheidole in Myrmicinae, Pheidolini: Emery, 1914a: 40; Forel, 1917: 241; Emery, 1922e: 112; Wheeler, W.M. 1922a: 661; all subsequent authors.
Ceratopheidole as genus: Emery, 1914a: 40; Forel, 1917: 241; Emery, 1922e: 112; Wheeler, W.M. 1922a: 661; Chapman & Capco, 1951: 135; Wu, J. & Wang, 1995: 104 (anachronism).
Ceratopheidole as subgenus of Pheidole: Pergande, 1896: 889; Kempf, 1972a: 183.
Ceratopheidole as junior synonym of Pheidole: Brown, 1973b: 179 [provisional]; Smith, D.R. 1979: 1365; Snelling, R.R. 1981: 395; Wilson, 2003: 6.
Epipheidole Wheeler, W.M. 1903f: 664. Type-species: Epipheidole inquilina, by monotypy.
Taxonomic history
[Epipheidole also described as new by Wheeler, W.M. 1904a: 14.]
Epipheidole in Myrmicinae, Stenammini: Ashmead, 1905b: 383.
Epipheidole in Myrmicinae, Myrmicini: Wheeler, W.M. 1910g: 140.
Epipheidole in Myrmicinae, Pheidolini: Emery, 1914a: 40; Forel, 1917: 241; Emery, 1922e: 114; Wheeler, W.M. 1922a: 661; Creighton, 1950a: 192.
Epipheidole as junior synonym of Pheidole: Cole, 1965: 174; Smith, D.R. 1979: 1366.
Phidole Bingham, 1903: 220, unjustified emendation of Pheidole.
Taxonomic history
Phidole as junior synonym of Pheidole: Wheeler, 1922a: 806.
Sympheidole Wheeler, W.M. 1904a: 7. Type-species: Sympheidole elecebra, by monotypy.
Taxonomic history
Sympheidole in Myrmicinae, Myrmicini: Wheeler, W.M. 1910g: 140.
Sympheidole in Myrmicinae, Stenammini: Ashmead, 1905b: 383.
Sympheidole in Myrmicinae, Pheidolini: Emery, 1914a: 40; Forel, 1917: 241; Emery, 1922e: 115; Wheeler, W.M. 1922a: 661; all subsequent authors to the following.
Sympheidole as junior synonym of Pheidole: Brown, 1973b: 185 [provisional]; Smith, D.R. 1979: 1365; Snelling, R.R. 1981: 395; Wilson, 2003: 6.
Phidola Schulz, W.A. 1906: 155, unjustified emendation of Pheidole.
Taxonomic history
Phidola as junior synonym of Pheidole: Smith, D.R. 1979: 1365.
Allopheidole Forel, 1912f: 237 [as subgenus of Pheidole]. Type-species: Pheidole kingi (junior synonym of Pheidole tepicana), by subsequent designation of Wheeler, W.M. 1913a: 79.
Taxonomic history
Allopheidole as subgenus of Pheidole: Forel, 1912f: 237; Forel, 1917: 241.
Allopheidole as junior synonym of Pheidole: Emery, 1915i: 190; Emery, 1921f: 84; Smith, D.R. 1979: 1366.
Decapheidole Forel, 1912f: 237 [as subgenus of Pheidole]. Type-species: Pheidole perpusilla, by subsequent designation of Wheeler, W.M. 1913a: 80.
Taxonomic history
Decapheidole in Myrmicinae, Pheidolini: Wheeler, W.M. 1922a: 661.
Decapheidole as genus: Wheeler, W.M. 1922a: 661; Borgmeier, 1927c: 78.
Decapheidole as subgenus of Pheidole: Forel, 1912f: 237, Forel, 1917: 241; Emery, 1922e: 112; Kempf, 1972a: 183.
Decapheidole as junior synonym of Pheidole: Brown, 1973b: 180 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395; Wilson, 2003: 6.
Isopheidole Forel, 1912m: 765 [as subgenus of Pheidole]. Type-species: Myrmica longipes, by monotypy.
Taxonomic history
Isopheidole as junior synonym of Ischnomyrmex: Forel, 1913b: 350 (in text); Forel, 1913k: 49; Emery, 1922e: 113; Wheeler, W.M. 1922a: 680 (in key). [Ischnomyrmex and Isopheidole share the same type-species, synonymy is therefore absolute.]
Elasmopheidole Forel, 1913k: 43 [as subgenus of Pheidole]. Type-species: Pheidole aberrans, by subsequent designation of Emery, 1915i: 190.
Taxonomic history
Elasmopheidole as subgenus of Pheidole: Forel, 1913k: 43; Emery, 1915i: 190; Forel, 1917: 241; Emery, 1922e: 110; Wheeler, W.M. 1922a: 673; all subsequent authors to the following.
Elasmopheidole as junior synonym of Pheidole: Brown, 1973b: 180 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395.
Cardiopheidole Wheeler, W.M. 1914b: 48 [as subgenus of Pheidole]. Type-species: Pheidole vaslitii, by original designation.
Taxonomic history
Cardiopheidole as subgenus of Pheidole: Wheeler, W.M. 1914b: 48; Forel, 1917: 241.
Cardiopheidole as junior synonym of Pheidole: Emery, 1915i: 190; Emery, 1921f: 84; Smith, D.R. 1979: 1366.
Anergatides Wasmann, 1915b: 281. Type-species: Anergatides kohli (junior secondary homonym in Pheidole, replaced by Pheidole neokohli), by monotypy.
Taxonomic history
Anergatides in Myrmicinae, Pheidolini: Forel, 1917: 243 [subtribe Anergatini]; Emery, 1922e: 116 [subtribe Anergatidini]; Donisthorpe, 1943f: 623.
Anergatides in Myrmicinae, Solenopsidini: Wheeler, W.M. 1922a: 663; Wheeler, G.C. & Wheeler, J. 1985: 257 (anachronism).
Anergatides as junior synonym of Pheidole: Brown, 1973b: 178 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395; Wilson, 1984: 328; Bolton, 1994: 106.
Parapheidole Emery, 1915d: 68. Type-species: Aphaenogaster oculata, by monotypy.
Taxonomic history
Parapheidole in Myrmicinae, Pheidolini: Forel, 1917: 241; Emery, 1922e: 115; Wheeler, W.M. 1922a: 661; all subsequent authors.
Parapheidole as junior synonym of Pheidole: Brown, 1973b: 183 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395.
Macropheidole Emery, 1915i: 190 [as subgenus of Pheidole]. Type-species: Pheidole fimbriata, by monotypy.
Taxonomic history
Macropheidole as subgenus of Pheidole: Emery, 1915i: 190; Forel, 1917: 241; Emery, 1921f: 81; Wheeler, W.M. 1922a: 673; all subsequent authors to the following.
Macropheidole as junior synonym of Pheidole: Smith, M.R. 1951a: 800; Brown, 1973b: 181 [provisional]; Smith, D.R. 1979: 1366.
Scrobopheidole Emery, 1915i: 190 [as subgenus of Pheidole]. Type-species: Pheidole scrobifera, by monotypy.
Taxonomic history
Scrobopheidole as subgenus of Pheidole: Forel, 1917: 241; Emery, 1922e: 112; Wheeler, W.M. 1922a: 673; all subsequent authors to the following.
Scrobopheidole as junior synonym of Pheidole: Brown, 1973b: 184 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395; Wilson, 2003: 6.
Stegopheidole Emery, 1915i: 190 [as subgenus of Pheidole]. Type-species: Pheidole (Elasmopheidole) upeneci, by monotypy.
Taxonomic history
Stegopheidole as subgenus of Pheidole: Emery, 1921f: 83; Wheeler, W.M. 1922a: 673; all subsequent authors to the following.
Stegopheidole as junior synonym of Pheidole: Brown, 1973b: 185 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395.
Trachypheidole Emery, 1915i: 190 [as subgenus of Pheidole]. Type-species: Pheidole bicornis, by original designation.
Taxonomic history
Trachypheidole as subgenus of Pheidole: Forel, 1917: 241; Emery, 1922e: 111; Wheeler, W.M. 1922a: 673; all subsequent authors to the following.
Trachypheidole as junior synonym of Pheidole: Brown, 1973b: 185 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395; Wilson, 2003: 6.
Electropheidole Mann, 1921: 438 [as subgenus of Pheidole]. Type-species: Pheidole (Electropheidole) roosevelti, by subsequent designation of Donisthorpe, 1943f: 642.
Taxonomic history
Electropheidole as subgenus of Pheidole: Mann, 1921: 438; Wheeler, W.M. 1922a: 672; all subsequent authors to the following.
Electropheidole as junior synonym of Pheidole: Brown, 1973b: 180 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395.
Bruchomyrma Santschi, 1922d: 248. Type-species: Bruchomyrma acutidens, by monotypy.
Taxonomic history
Bruchomyrma in Myrmicinae, Pheidolini: Donisthorpe, 1943f: 629.
Bruchomyrma in Myrmicinae, Myrmicini: Kempf, 1972a: 42.
Bruchomyrma as junior synonym of Pheidole: Brown, 1973b: 179 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395; Wilson, 1984: 327; Bolton, 1994: 106; Wilson, 2003: 6.
Bruchomyrma as genus: Dlussky & Fedoseeva, 1988: 80 (anachronism).
Cephalomorium Forel, 1922: 91 [as subgenus of Tetramorium]. Type-species: Tetramorium (Cephalomorium) bahai, by monotypy.
Taxonomic history
Cephalomorium as junior synonym of Hendecapheidole: Santschi, 1925d: 228.
Hendecapheidole Wheeler, W.M. 1922d: 3 [as subgenus of Pheidole]. Type-species: Pheidole tachigaliae, by original designation.
Taxonomic history
Hendecapheidole as junior synonym of Pheidole: Brown, 1973b: 181 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395; Wilson, 2003: 6.
Gallardomyrma Bruch, 1932: 271. Type-species: Gallardomyrma argentina, by original designation.
Taxonomic history
Gallardomyrma in Myrmicinae, Pheidolini: Donisthorpe, 1943f: 646.
Gallardomyrma in Myrmicinae, Myrmicini: Kempf, 1972a: 110.
Gallardomyrma as junior synonym of Pheidole: Brown, 1973b: 180 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395; Wilson, 1984: 327; Wilson, 2003: 6.
Gallardomyrma as genus: Kempf, 1972a: 110; Dlussky & Fedoseeva, 1988: 80 (anachronism).
Conothoracoides Strand, 1935: 176.
Taxonomic history
[Replacement name for Conothorax Karavaiev; junior homonym of Conothorax Jekel, 1854: 9 bis (foldout pages) (Coleoptera).]
Conothoracoides as junior synonym of Pheidole: Brown, 1973b: 179 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395.
Homonym replaced by Conothoracoides
Conothorax Karavaiev, 1935a: 75. Type-species: Conothorax bilobum, by monotypy.
Taxonomic history
[Conothorax Karavaiev junior homonym of Conothorax Jekel, 1854: 9 bis (foldout pages) (Coleoptera).]
Conothorax in Myrmicinae, Pheidolini: Karavaiev, 1935a: 75.
Eriopheidole Kusnezov, 1952a: 10. Type-species: Eriopheidole symbiotica (unresolved junior secondary homonym in Pheidole), by monotypy.
Taxonomic history
Eriopheidole in Myrmicinae, Pheidolini: Kusnezov, 1952a: 10.
Eriopheidole in Myrmicinae, Myrmicini: Kempf, 1972a: 107.
Eriopheidole junior synonym of Pheidole: Brown, 1973b: 180 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395; Bolton, 1994: 106.
Eriopheidole as genus: Kempf, 1972a: 107; Dlussky & Fedoseeva, 1988: 80 (anachronism).
Xenoaphaenogaster Baroni Urbani, 1964b: 50. Type-species: Xenoaphaenogaster inquilina (junior synonym of Myrmica pallidula), by original designation.
Taxonomic history
Xenoaphaenogaster as junior synonym of Monomorium: Brown, 1973b: 185 [provisional].
Xenoaphaenogaster as junior synonym of Pheidole: Bolton, 1987: 291.
Genus Pheidole references
Smith, F. 1858b: 172 (diagnosis); Mayr, 1861: 70 (Europe species key); Roger, 1863b: 27, 30 (catalogue); Mayr, 1863: 424, 440 (Ischnomyrmex, Pheidole catalogues); Mayr, 1865: 18, 22 (Ischnomyrmex, Pheidole diagnoses); Mayr, 1867a: 92, 96 (Ischnomyrmex, Pheidole diagnoses); Mayr, 1870b: 977, 979 (Australia & New World species keys); Mayr, 1876: 102 (Australia species key); André, 1883b: 382 (Europe & Algeria species key); Mayr, 1879: 674 (Asia species key); Mayr, 1887: 582 (New World species key); Cresson, 1887: 262 (U.S.A., catalogue); Nasonov, 1889: 75 (Russia species key); Dalla Torre, 1893: 88, 98 (Pheidole, Pheidolacanthinus catalogues); Emery, 1895c: 288 (North America species key); Mayr, 1896: 241 (Africa species, partial key); Emery, 1896g: 80 (P. flavens group, key); Forel, 1902c: 165 (India species key); Forel, 1902f: 523 (India & Sri Lanka species key); Bingham, 1903: 223 (India, Sri Lanka & Burma species key); Wheeler, W.M. 1908a: 133 (Puerto Rico species key); Arnold, 1920a: 414 (diagnosis, South Africa species key); Emery, 1921f: 78 (diagnosis, subgenera key, catalogue); Emery, 1921f: 81 (P. (Macropheidole) & P. (Pheidolacanthinus) diagnoses, catalogues); Emery, 1921f: 83 (P. (Stegopheidole) diagnosis, catalogue); Mann, 1921: 442 (Fiji Is species key); Emery, 1922e: 110 (P. (Elasmopheidole) diagnosis, catalogue); Emery, 1922e: 111 (P. (Trachypheidole) diagnosis, catalogue); Emery, 1922e: 112 (P. (Scrobopheidole), P. (Decapheidole) & P. (Ceratopheidole) diagnoses, catalogues); Emery, 1922e: 113 (Ischnomyrmex diagnosis, catalogue); Emery, 1922e: 114 (Epipheidole diagnosis, catalogue); Emery, 1922e: 115 (Sympheidole, Parapheidole diagnoses, catalogues); Emery, 1922e: 116 (Anergatides diagnosis, catalogue); Wheeler, W.M. 1922a: 126, 672 (diagnosis, subgenera key); Wheeler, W.M. 1922a: 806, 879 (Afrotropical Pheidole, Anergatides catalogues); Wheeler, W.M. 1922a: 1017, 1021 (Malagasy Pheidole, Parapheidole catalogues); Gallardo, 1932c: 179 (Argentina Elasmopheidole species key); Finzi, 1936: 165 (Egypt species key); Menozzi, 1939a: 298 (Himalaya species key); Cole, 1942: 362 (U.S.A., Utah species key); Buren, 1944a: 285 (U.S.A., Iowa species key); Creighton, 1950a: 163 (North America species key); Creighton, 1950a: 192, 194 (North America Epipheidole, Sympheidole species); Chapman & Capco, 1951: 135, 137 (Asia Ceratopheidole, Conothorax, Ischnomyrmex, Pheidole checklists); Kusnezov, 1952b: 61 (Argentina species key); Smith, M.R. 1955c: 302 (P. (Hendecapheidole) species key); Gregg, 1959: 9 (U.S.A. species key); Gregg, 1963: 407 (U.S.A., Colorado species key); Wheeler, G.C. & Wheeler, J. 1963: 126 (U.S.A., North Dakota species key); Bernard, 1967: 149 (diagnosis, Western Europe species key); Gregg, 1969: 95 (addition to Gregg, 1959 U.S.A. key); Kempf, 1972a: 183 (Neotropical catalogue); Alayo, 1974: 11 (Cuba species key); Smith, D.R. 1979: 1365 (North America catalogue); Allred, 1982: 441 (U.S.A., Utah species key); Ogata, 1982: 191 (Japan species key); Collingwood, 1985: 253 (Saudi Arabia species key); Naves, 1985: 54 (U.S.A., Florida species key); Taylor & Brown, D.R. 1985: 74 (Australia catalogue); Wheeler, G.C. & Wheeler, J. 1986g: 41 (U.S.A., Nevada species key); Taylor, 1987a: 53 (Australia, New Caledonia & New Zealand checklists); Agosti & Collingwood, 1987b: 271 (Balkans species key); Hölldobler & Wilson, 1990: 15 (synoptic classification); Kupyanskaya, 1990: 123 (Far Eastern Russia species key); Brandão, 1991: 368 (Neotropical catalogue); Morisita, Kubota, et al. 1992: 22 (Japan species key); Bolton, 1994: 106 (synoptic classification); Kupyanskaya, 1995: 345 (Far Eastern Russia species key); Bolton, 1995a: 1051 (census); Bolton, 1995b: 316 (catalogue); Wu, J. & Wang, 1995: 99 (China species key); Collingwood & Agosti, 1996: 322 (Saudi Arabia species key); Park & Kim, 2000: 109 (Korea species key); Shattuck, 1999: 152 (Australia synopsis); Eguchi, 1999: 103 (Borneo P. longipes group, key); Ward, 2000: 96 (U.S.A. P. hyatti complex, key); Zhou, 2001b: 121 (China, Guangxi species key); Eguchi, 2001b: 10 (Borneo species key); ; Mackay & Mackay, 2002: 149 (U.S.A., New Mexico species key)Wilson, 2003: 27 (New World, all species revision, keys); Sarnat, 2008: 8 (Fiji Is P. roosevelti group, key); Eguchi, 2008: 8 (North Vietnam species key); Terayama, 2009: 171 (Taiwan species key); Heterick, 2009: 169 (south-western Australia species key); Sarnat & Moreau, 2011: 114 (P. roosevelti group phylogeny and radiation).
Genera incertae sedis in Pheidolini
Myrmicinae Formicoxenini Leptothorax TRUE TRUE FALSEExtant: 18 species, 2 subspecies
Fossil:
Leptothorax Mayr, 1855: 431. Type-species: Formica acervorum, by subsequent designation of Bingham, 1903: 214.
Taxonomic history
[Type-species not Myrmica clypeata, unjustified subsequent designation by Emery, 1912d: 271; repeated in Wheeler, W.M. 1913a: 79 and Emery, 1924d: 248.]
Leptothorax in Myrmicidae, Myrmicidae: Emery, 1877a: 81.
Leptothorax in Myrmicinae: Mayr, 1855: 431 [Myrmicidae]; Mayr, 1861: 57 [Myrmicidae]; Mayr, 1865: 20 [Myrmicidae]; Emery & Forel, 1879a: 458 [Myrmicidae]; Dalla Torre, 1893: 122.
Leptothorax in Myrmicinae, Myrmicini: Forel, 1895b: 125; Emery, 1895j: 769; Forel, 1899c: 54; Wheeler, W.M. 1910g: 139; Wheeler, W.M. 1915h: 63; Kusnezov, 1964: 57 (anachronism).
Leptothorax in Myrmicinae, Stenammini: Ashmead, 1905b: 383.
Leptothorax in Myrmicinae, Leptothoracini: Emery, 1914a: 42; Arnold, 1916: 257; Forel, 1917: 244; Wheeler, W.M. 1922a: 664; Emery, 1924d: 247; subsequent authors to the following.
Leptothorax in Myrmicinae, Formicoxenini: Bolton, 1994: 105; Mackay, 2000: 267.
Junior synonyms of LEPTOTHORAX
Mychothorax Ruzsky, 1904a: 288. Type-species: Formica acervorum, by original designation.
Taxonomic history
Mychothorax as subgenus of Leptothorax: Ruzsky, 1905b: 609; Emery, 1915g: 24; Emery, 1916b: 176; Forel, 1917: 245; Wheeler, W.M. 1922a: 679; Emery, 1924d: 260; Creighton, 1950a: 274.
Mychothorax as junior synonym of Leptothorax: Smith, M.R. 1950: 29; Brown, 1973b: 182 [provisional]; Bolton, 1982: 319. [Leptothorax and Mychothorax share the same type-species, synonymy is therefore absolute.]
Doronomyrmex Kutter, 1945: 485. Type-species: Doronomyrmex pacis, by monotypy.
Taxonomic history
Doronomyrmex in Myrmicinae, Leptothoracini:Dlussky & Fedoseeva, 1988: 79.
Doronomyrmex in Myrmicinae, Formicoxenini: Bolton, 1994: 105.
Doronomyrmex as junior synonym of Leptothorax: Brown, 1973b: 180 [provisional]; Heinze, 1998: 195; Bolton, 2003: 270.
Genus Leptothorax references
Mayr, 1855: 433 (Austria species key); Mayr, 1861: 58 (Europe species key); Roger, 1863b: 26, 28, 30 (catalogue); Mayr, 1863: 426, 428, 456 (Leptothorax, Macromischa, Temnothorax catalogues); Mayr, 1865: 19, 20, 21 (Macromischa, Leptothorax, Temnothorax diagnoses); Mayr, 1868c: 83 (†Baltic Amber species key); André, 1874: 188 (Europe species key); Forel, 1874: 84 (Switzerland species key); André, 1883a: 293 (Europe & Algeria species key); Mayr, 1886d: 451 (U.S.A. species key); Cresson, 1887: 261 (U.S.A. catalogue); Nasonov, 1889: 70 (Russia species key); Emery, 1891b: 5 (North Africa species key); Dalla Torre, 1893: 120, 122 (Macromischa, Leptothorax catalogues); Emery, 1895c: 317 (North America species key); Emery, 1896g: 58 (Neotropical species key); Bingham, 1903: 215 (India, Sri Lanka & Burma species key); Wheeler, W.M. 1903c: 223 (North America species key); Ruzsky, 1905b: 570 (Russian Empire species key); Wasmann, 1906: 16 (Luxemburg species key); Wheeler, W.M. 1908a: 141 (Macromischa species key); Santschi, 1909b: 460 (L. rottenbergi group key); Bondroit, 1910: 496 (Belgium species key); Stitz, 1914: 60 (Central Europe species key); Crawley, 1914: 91 (Britain species key); Donisthorpe, 1915d: 147 (Britain species key); Emery, 1916b: 176 (Italy species key); Wheeler, W.M. 1916m: 581 (U.S.A., Connecticut species key); Arnold, 1916: 257 (diagnosis, South Africa species key); Bondroit, 1918: 117 (France & Belgium species key); Mann, 1920: 408 (Macromischa species key); Wheeler, W.M. 1922a: 677 (Macromischa subgenera key); Wheeler, W.M. 1922a: 679 (Leptothorax subgenera key); Wheeler, W.M. 1922a: 890 (Afrotropical catalogue); Wheeler, W.M. 1922a: 1029 (Malagasy catalogue); Emery, 1924d: 245 (Macromischa catalogue); Emery, 1924d: 247 (diagnosis, subgenera key, catalogue); Emery, 1924d: 249 (L. (Goniothorax) diagnosis, catalogue); Emery, 1924d: 251 (L. (Leptothorax) diagnosis, catalogue); Emery, 1924d: 259 (L. (Temnothorax) diagnosis, catalogue); Emery, 1924d: 260 (L. (Dichothorax) & L. (Mychothorax) diagnoses, catalogues); Emery, 1924d: 264 (Symmyrmica review, catalogue); Karavaiev, 1927c: 266 (Ukraine species key); Donisthorpe, 1927b: 163 (Britain species key); Kuznetsov-Ugamsky, 1927d: 38 (Turkestan species key); Wheeler, W.M. 1931b: 31 (Macromischa, Croesomyrmex, Antillaemyrmex checklists); Arnol'di, 1933b: 598 (Russia species key); Menozzi, 1933b: 68 (Israel species key); Karavaiev, 1934: 134 (Ukraine species key); Wheeler, W.M. 1937b: 463 (Macromischa checklist); Menozzi, 1939a: 307 (Himalaya & Tibet species key); Stitz, 1939: 158 (Germany species key); Smith, M.R. 1939e: 503 (U.S.A. Macromischa species key); Kratochvíl, 1941: 89 (Central Europe species key); Novák & Sadil, 1941: 89 (Central Europe species key); Cole, 1942: 369 (U.S.A., Utah species key); Holgersen, 1943b: 172 (Norway species key); Holgersen, 1944: 198 (Norway species key); Kratochvíl, Novák & Snoflák, 1944: 123 (Czechoslovakia males, key); Buren, 1944a: 286 (U.S.A., Iowa species key); Boven, 1947: 178 (Belgium species key); Creighton, 1950a: 251, 256 (North America Macromischa, Leptothorax species keys); Chapman & Capco, 1951: 110 (Asia checklist); Smith, M.R. 1952b: 97 (U.S.A. L. tricarinatus complex, key); Bernard, 1956a: 151 (Western Europe species groups); Kusnezov, 1958c: 266 (subgenera key); Boven, 1959: 8 (Netherlands species key); Gregg, 1963: 380 (U.S.A., Colorado species key); Wheeler, G.C. & Wheeler, J. 1963: 139 (U.S.A., North Dakota species key); Collingwood, 1964: 99 (Britain species key); Bernard, 1967: 185 (diagnosis, Western Europe species key); Bernard, 1967: 187 (diagnosis, Western Europe species key); Bernard, 1967: 224 (Doronomyrmex diagnosis); Boven, 1970b: 20 (Netherlands species key); Arnol'di, 1971: 1824 (Kazakhstan species key); Kempf, 1972a: 132, 135 (Neotropical Leptothorax, Macromischa catalogues); Alayo, 1974: 15 (Macromischa Cuba species key); Bolton & Collingwood, 1975: 5 (Britain species key); Tarbinsky, 1976: 83 (Kyrghyzstan species key); Boven, 1977: 94 (Belgium species key); Kutter, 1977c: 103 (Switzerland species key); Arnol'di & Dlussky, 1978: 540 (former European U.S.S.R. species key); Collingwood, 1978: 83 (Iberian Peninsula species key); Collingwood, 1979: 68 (Fennoscandia & Denmark species key); Baroni Urbani, 1978b: 533 (L. (Macromischa) all species revision, key); Smith, D.R. 1979: 1391 (North America catalogue); Buschinger, 1981: 211 (Doronomyrmex review); Allred, 1982: 440 (U.S.A., Utah species key); Bolton, 1982: 319 (diagnosis, review of genus, Afrotropical species key); Gösswald, 1985: 303 (Germany species key); Wheeler, G.C. & Wheeler, J. 1986g: 51 (U.S.A., Nevada species key); Nilsson & Douwes, 1987: 60 (Norway species key); Agosti & Collingwood, 1987b: 273 (Balkans species key); Dlussky & Soyunov, 1988: 29 (former U.S.S.R. Temnothorax species); Hölldobler & Wilson, 1990: 13 (synoptic classification); Dlussky, Soyunov & Zabelin, 1990: 184, 188 (Turkmenistan species key); Kupyanskaya, 1990: 136 (Far Eastern Russia species key); Brandão, 1991: 349 (Neotropical catalogue); Morisita, Kubota, et al. 1992: 26 (Japan species key); Atanasov & Dlussky, 1992: 129 (Bulgaria species key); Arakelian, 1994: 52 (Armenia species key); Radchenko, 1994b: 111 (South Siberia species key); Radchenko, 1994d: 146 (Central & Eastern Palaearctic species key); Bolton, 1995a: 1050 (census); Bolton, 1995b: 235 (catalogue); Douwes, 1995: 89 (Sweden species key); Kupyanskaya, 1995: 347 (Far Eastern Russia species key); Wu, J. & Wang, 1995: 108 (China species key); Collingwood & Agosti, 1996: 325 (Saudi Arabia species key); Seifert, 1996b: 13 (Central Europe Doronomyrmex species key); Seifert, 1996b: 119 (Central Europe species key); Skinner & Allen, 1996: 45 (Britain species key); Cagniant & Espadaler, 1997: 260 (Morocco species key); Collingwood & Prince, 1998: 16 (Portugal species key); Terayama & Onoyama, 1999: 73 (Japan species key); Mackay, 2000: 265, 288, 297 (New World L. (Myrafant) species revision key, Mexico & Guatemala species key, species complexes key); Fontenla Rizo, 2001: 15 (Cuba Macromischa species key); Chang & He, 2001b: 1 (northwest China species key); Czechowski, Radchenko & Czechowska, 2002: 143 (Poland species key); Mackay & Mackay, 2002: 93 (U.S.A., New Mexico species key); Bolton, 2003: 270 (genus diagnosis); Lyu & Cho, 2003b: 269 (Korea species key); Radchenko, 2004: 131 (Eastern Palaearctic species, key); Deyrup & Cover, 2004a: 56 (southeast U.S.A. species key); Radchenko, 2005b: 192 (North Korea species key); Seifert, 2007: 118 (North and Central Europe species key).
Myrmicinae Formicoxenini Myrmoxenus TRUE TRUE FALSE12 species
Myrmoxenus Ruzsky, 1902b: 474. Type-species: Myrmoxenus gordiagini, by monotypy.
Taxonomic history
Myrmoxenus in Myrmicinae, Myrmicini: Wheeler, W.M. 1910g: 139.
Myrmoxenus in Myrmicinae, Leptothoracini: Emery, 1914a: 42; Forel, 1917: 245; Wheeler, W.M. 1922a: 664; Emery, 1924d: 262; all subsequent authors to the following.
Myrmoxenus in Myrmicinae, Formicoxenini: Bolton, 1994: 105.
Myrmoxenus as junior synonym of Epimyrma: Buschinger, 1990: 245 (provisional synoynym); Bolton, 1994: 105 [seniority incorrect, Myrmoxenus has priority; see below.]
Junior synonyms of MYRMOXENUS
Epimyrma Emery, 1915a: 262. Type-species: Epimyrma kraussei, by original designation.
Taxonomic history
Epimyrma in Myrmicinae, Leptothoracini: Forel, 1917: 245; Wheeler, W.M. 1922a: 664; Emery, 1924d: 263; all subsequent authors to the following.
Epimyrma in Myrmicinae, Formicoxenini: Bolton, 1994: 105.
Epimyrma as junior synonym of Leptothorax: Brown, 1973b: 180 [provisional].
Epimyrma as junior synonym of Myrmoxenus: Schulz & Sanetra, 2002: 162.
Myrmetaerus Soudek, 1925a: 33. Type-species: Myrmetaerus microcellatus (junior synonym of Myrmoxenus gordiagini), by monotypy.
Taxonomic history
[Myrmetaerus also described as new by Soudek, 1925b: 14.]
Myrmetaerus in Myrmicinae, Leptothoracini: Donisthorpe, 1943f: 667.
Myrmetaerus as junior synonym of Leptothorax: Brown, 1973b: 182 [provisional].
Myrmetaerus as junior synonym of Camponotus: Snelling, R.R. 1981: 404 (error).
Myrmetaerus as junior synonym of Myrmoxenus: Buschinger, Winter & Faber, 1984: 336.
Subgenera of MYRMOXENUS include the nominal plus the following.
Subgenus MYRMOXENUS (GONEPIMYRMA)
Gonepimyrma Bernard, 1948: 146 [as subgenus of Epimyrma]. Type-species: Epimyrma (Gonepimyrma) africana, by monotypy.
Taxonomic history
Gonepimyrma junior synonym of Epimyrma: Brown, 1973b: 181 [provisional].
Genus Myrmoxenus references
Emery, 1916b: 189 (Italy species key); Bondroit, 1918: 139 (France & Belgium species key); Emery, 1924d: 262 (Myrmoxenus diagnosis, catalogue); Emery, 1924d: 263 (Epimyma diagnosis, catalogue); Menozzi, 1931c: 36 (diagnosis, all species key); Bernard, 1967: 216 (diagnosis, Western Europe species key); Kutter, 1973e: 281 (species, notes); Kutter, 1977c: 138 (Switzerland species key); Buschinger, Fischer, et al. 1987: 253 (partial revision); Agosti & Collingwood, 1987b: 276 (Balkans species key); Buschinger, 1989: 265 (evolution, speciation); Bolton, 1995a: 1049 (census); Bolton, 1995b: 188 (catalogue); Seifert, 1996b: 118 (Central Europe species key); Cagniant & Espadaler, 1997: 268 (Morocco species key); Seifert, 2007: 117 (North and Central Europe species key).
Myrmicinae Formicoxenini Nesomyrmex TRUE TRUE FALSEExtant: 51 species, 1 subspecies
Fossil: 2 species
Nesomyrmex Wheeler, W.M. 1910a: 259. Type-species: Nesomyrmex clavipilis, by monotypy.
Taxonomic history
[Nesomyrmex is the oldest synonym of Goniothorax Emery (a junior homonym, see below) and hence is the first available replacement name for Goniothorax Emery: Smith, M.R. 1950: 30.]
Nesomyrmex in Myrmicinae, Leptothoracini: Forel, 1917: 244; subsequent authors to the following.
Nesomyrmex in Myrmicinae, Formicoxenini: Bolton, 1994: 105.
Nesomyrmex as subgenus of Leptothorax: Emery, 1915g: 24; Forel, 1917: 244; Wheeler, W.M. 1922a: 679; Smith, M.R. 1950: 30; Smith, M.R. 1951a: 816; Kempf, 1959c: 393; Brown, 1971a: 4; Kempf, 1972a: 133; Smith, D.R. 1979: 1391; Mackay, 2000: 272 (in key).
Nesomyrmex as junior synonym of Goniothorax: Emery, 1924d: 249; Donisthorpe, 1943f: 674. [Goniothorax is a junior homonym and hence Nesomyrmex takes priority as first available replacement name.]
Nesomyrmex as junior synonym of Leptothorax: Brown, 1973b: 183 [provisional]; Bolton, 1982: 319; Bolton, 1994: 105; Bolton, 1995b: 39.
Nesomyrmex as genus: Wheeler, W.M. 1910a: 259; Francoeur & Loiselle, 1988b: 43 (possibly valid); Hölldobler & Wilson, 1990: 14 [junior synonyms of Nesomyrmex left behind under Leptothorax]; Jaffe, 1993: 11; Bolton, 2003: 70.
Junior synonyms of NESOMYRMEX
Tetramyrma Forel, 1912m: 766 [as subgenus of Dilobocondyla]. Type-species: Dilobocondyla (Tetramyrma) braunsi, by monotypy.
Taxonomic history
Tetramyrma in Myrmicinae, Tetramoriini: Emery, 1914a: 42; Forel, 1917: 245; Arnold, 1917: 358; Wheeler, W.M. 1922a: 664; Emery, 1924d: 291; Donisthorpe, 1943g: 732; Wheeler, G.C. & Wheeler, J. 1985: 257 (anachronism); Dlussky & Fedoseeva, 1988: 80 (anachronism).
Tetramyrma incertae sedis in Myrmicinae: Bolton, 1976: 291.
Tetramyrma as subgenus of Dilobocondyla: Forel, 1912m: 766.
Tetramyrma as genus: Forel, 1913a: 122; Arnold, 1917: 358; Wheeler, W.M. 1922a: 664; Emery, 1924d: 291; Donisthorpe, 1943g: 732; Bolton, 1976: 291.
Tetramyrma as junior synonym of Leptothorax: Bolton, 1982: 319; Bolton, 1994: 105.
Tetramyrma as junior synonym of Nesomyrmex: Bolton, 2003: 249, 272.
Caulomyrma Forel, 1914d: 233 [as subgenus of Leptothorax]. Type-species: Leptothorax echinatinodis, by original designation.
Taxonomic history
Caulomyrma as junior synonym of Nesomyrmex: Forel, 1915c: 364 [provisional]; Emery, 1915g: 24; Wheeler, W.M. 1922a: 679.
Caulomyrma as subgenus of Leptothorax: Forel, 1914d: 233; Forel, 1917: 244.
Caulomyrma as junior synonym of Goniothorax: Emery, 1924d: 249 (incorrect procedure as Goniothorax is a junior homonym for which Nesomyrmex is the first available replacement name; see Smith, M.R. 1950: 30).]
Caulomyrma as junior synonym of Leptothorax: Brown, 1973b: 182 [provisional].
Limnomyrmex Arnold, 1948: 222. Type-species: Limnomyrmex stramineus, by original designation.
Taxonomic history
Limnomyrmex as junior synonym of Nesomyrmex: Brown, 1971a: 4.
Ireneopone Donisthorpe, 1946d: 242. Type-species: Ireneopone gibber, by original designation.
Taxonomic history
Ireneopone in Myrmicinae, Tetramoriini: Donisthorpe, 1946d: 243; Wheeler, G.C. & Wheeler, J. 1985: 257 (anachronism).
Ireneopone incertae sedis in Myrmicinae: Bolton, 1976: 292; Dlussky & Fedoseeva, 1988: 81.
Ireneopone in Myrmicinae, Formicoxenini: Bolton, 1994: 105; Bolton, 1995a: 1050; Bolton, 1995b: 217.
Ireneopone as junior synonym of Nesomyrmex: Bolton, 2003: 250, 272.
Meia Pagliano & Scaramozzino, 1990: 5.
Taxonomic history
[Unnecessary replacement name for Goniothorax Emery, 1896g: 58 (junior homonym of Goniothorax Milne-Edwards, 1879: 103). Nesomyrmex is the first available replacement name for Goniothorax Emery, therefore Meia is automatically a junior synonym of Nesomyrmex: Bolton, 1994: 105.]
Genus Nesomyrmex references
Emery, 1896g: 58 (Neotropical species key); Arnold, 1916: 258 (South Africa species key); Arnold, 1917: 358 (Tetramyrma diagnosis); Wheeler, W.M. 1922a: 890, 908 (Afrotropical L. (Goniothorax), Tetramyrma catalogues); Wheeler, W.M. 1922a: 1029 (Malagasy L. (Goniothorax) catalogue); Emery, 1924d: 249 (L. (Goniothorax) diagnosis, catalogue); Emery, 1924d: 291 (Tetramyrma review, catalogue); Kempf, 1959c: 394 (Neotropical Leptothorax (Nesomyrmex) species revision, key); Kempf, 1972a: 133 (Neotropical Leptothorax (Nesomyrmex) catalogue); Kempf, 1975c: 367 (additions to 1959b key); Bolton, 1982: 319 (diagnosis, review of genus, Afrotropical species key); Hölldobler & Wilson, 1990: 13 (synoptic classification); Brandão, 1991: 349 (Neotropical catalogue); Bolton, 1995b: 235 (catalogue); Bolton, 2003: 272 (genus diagnosis); Mbanyana & Robertson, 2008: 36 (southern Africa species revision, key).
Homonym replaced by NESOMYRMEX
Goniothorax Emery, 1896g: 58 (diagnosis in key) [as subgenus of Leptothorax]. Type-species: Leptothorax vicinus, by subsequent designation of Wheeler, W.M. 1911f: 164.
Taxonomic history
[Junior homonym of Goniothorax Milne-Edwards, 1879: 103 (Crustacea).]
Goniothorax in Myrmicinae, Leptothoracini: Forel, 1917: 244.
Goniothorax as subgenus of Leptothorax: Wheeler, W.M. 1910g: 139; Forel, 1917: 244; Wheeler, W.M. 1922a: 679, 890, 1029; Emery, 1924d: 249; Creighton, 1950a: 259.
[Goniomyrmex Ashmead, 1905b: 383, incorrect subsequent spelling.]
57 species
Pristomyrmex Mayr, 1866b: 903. Type-species: Pristomyrmex pungens (junior synonym of Myrmica punctata) by monotypy.
Taxonomic history
Pristomyrmex in Myrmicidae: Mayr, 1866b: 903.
Pristomyrmex in Myrmicidae, Myrmicidae: Emery, 1877a: 81.
Pristomyrmex in Myrmicinae: Dalla Torre, 1893: 62.
Pristomyrmex in Myrmicinae, Myrmicini: Emery, 1895j: 769; Ashmead, 1905b: 383; Wheeler, W.M. 1910g: 139.
Pristomyrmex in Myrmicinae, Myrmecinini: Emery, 1912b: 105; Emery, 1914a: 41; Forel, 1917: 244; Wheeler, W.M. 1922a: 663; Emery, 1924d: 233; all subsequent authors except the entries below; Bolton, 1994: 105; Bolton, 2003: 71.
Pristomyrmex in Myrmicinae, Tetramoriini: Bernard, 1953b: 251.
Pristomyrmex in Myrmicinae, Pheidolini: Hölldobler & Wilson, 1990: 16.
Junior synonyms of PRISTOMYRMEX
Odontomyrmex André, 1905: 207. Type-species: Odontomyrmex quadridentatus, by monotypy.
Taxonomic history
Odontomyrmex in Myrmicinae, Myrmicini: Wheeler, W.M. 1910g: 139.
Odontomyrmex as subgenus of Pristomyrmex: Forel, 1917: 244; Wheeler, W.M. 1922a: 682; Chapman & Capco, 1951: 121 (anachronism).
Odontomyrmex as junior synonym of Pristomyrmex: Mann, 1919: 341; Emery, 1924d: 233; Donisthorpe, 1943f: 676; Brown, 1953c: 9; Taylor, 1965b: 35; Bolton, 1981b: 282.
Hylidris Weber, 1941a: 190. Type-species: Hylidris myersi (junior synonym of Pristomyrmex africanus), by original designation.
Taxonomic history
Hylidris in Myrmicinae, Myrmecinini: Weber, 1941a: 183; Donisthorpe, 1943f: 651.
Hylidris as junior synonym of Pristomyrmex: Brown, 1953c: 9.
Dodous Donisthorpe, 1946f: 145. Type-species: Dodous trispinosus, by original designation.
Taxonomic history
Dodous in Myrmicinae, Solenopsidini: Donisthorpe, 1946f: 145.
Dodous as junior synonym of Pristomyrmex: Brown, 1971a: 3.
Genus Pristomyrmex references
Dalla Torre, 1893: 62 (catalogue); Forel, 1903a: 696 (India & Sri Lanka species key); Bingham, 1903: 193 (diagnosis); Emery, 1924d: 233 (diagnosis, catalogue); Chapman & Capco, 1951: 121, 161 (Asia Pristomyrmex, Dodous checklists); Taylor, 1965b: 37 (Australia species revision, key); Taylor, 1968c: 63 (supplement to previous); Bolton, 1981b: 282 (diagnosis, Afrotropical species revision, key); Taylor & Brown, D.R. 1985: 85 (Australia catalogue); Taylor, 1987a: 64 (Australia checklist); Morisita, Kubota, et al. 1992: 57 (Japan species key); Bolton, 1994: 105 (synoptic classification); Bolton, 1995a: 1052 (census); Bolton, 1995b: 365 (catalogue); Lin & Wu, 1998: 94 (Taiwan species key); Shattuck, 1999: 159 (Australia synopsis); Xu & Zhang, 2002: 69 (China species key); Wang, M. 2003: 393 (all species revision, key); Zettell, 2006: 65 (Philippines species, key); Terayama, 2009: 190 (Taiwan species key).
Genera incertae sedis in Myrmecinini
Ponerinae Ponerini Anochetus TRUE TRUE FALSEExtant: 94 species
Fossil: 8 species
Anochetus Mayr, 1861: 53. Type-species: Odontomachus ghilianii, by monotypy.
Taxonomic history
Anochetus in Formicidae, Odontomachidae: Mayr, 1862: 712; Mayr, 1865: 11.
Anochetus in Odontomachidae: Ashmead, 1905b: 382.
Anochetus in Ponerinae: Mayr, 1861: 53 [Poneridae]; Emery & Forel, 1879a: 455 [Poneridae]; Dalla Torre, 1893: 47.
Anochetus in Ponerinae, Odontomachini: Forel, 1893a: 163; Emery, 1895j: 768; Forel, 1899c: 18; Wheeler, W.M. 1910g: 136; Emery, 1911d: 106; Arnold, 1915: 103; Forel, 1917: 238; Gallardo, 1918b: 89; Wheeler, W.M. 1922a: 653; Donisthorpe, 1943f: 623; Chapman & Capco, 1951: 31; Wilson, 1959a: 483; Wheeler, G.C. & Wheeler, J. 1985: 256; Dlussky & Fedoseeva, 1988: 78; Hölldobler & Wilson, 1990: 11; Jaffe, 1993: 8.
Anochetus in Ponerinae, Ponerini: Brown, 1976a: 71; Brown, 1978c: 550 [subtribe Odontomachiti]; Bolton, 1994: 164; Bolton, 2003: 159.
Junior synonyms of ANOCHETUS
Stenomyrmex Mayr, 1862: 711. Type-species: Myrmecia emarginata, by subsequent designation of Wheeler, W.M. 1911f: 173.
Taxonomic history
Stenomyrmex in Formicidae, Odontomachidae: Mayr, 1862: 711; Mayr, 1865: 11.
Stenomyrmex in Odontomachidae: Ashmead, 1905b: 382.
Stenomyrmex as subgenus of Anochetus: Dalla Torre, 1893: 47; Forel, 1893a: 163; Dalla Torre, 1893: 47; Wheeler, W.M. 1910g: 136; Emery, 1911d: 110; Forel, 1917: 238; Wheeler, W.M. 1922a: 653; Borgmeier, 1923: 76; Donisthorpe, 1943g: 728; Kempf, 1972a: 21.
Stenomyrmex as junior synonym of Anochetus: Forel, 1887: 382; Brown, 1978c: 552; all subsequent authors.
Myrmapatetes Wheeler, W.M. 1929b: 6. Type-species: Myrmapatetes filicornis, by original designation.
Taxonomic history
Myrmapatetes in Dolichoderinae: Wheeler, W.M. 1929b: 7; Chapman & Capco, 1951: 186.
Myrmapatetes as junior synonym of Anochetus: Brown, 1953h: 2.
Genus Anochetus references
Roger, 1863b: 21, 22 (catalogue); Mayr, 1863: 394, 454 (catalogue); Mayr, 1865: 11 (Anochetus, Stenomyrmex diagnoses); André, 1882b: 230 (Europe & Algeria species); Emery, 1884a: 378 (all species key); Dalla Torre, 1893: 47 (catalogue); Emery, 1894c: 185 (Neotropical species key); Forel, 1900c: 58 (India & Sri Lanka species key); Bingham, 1903: 39 (India, Sri Lanka & Burma species key); Emery, 1911d: 107 (diagnosis, catalogue); Arnold, 1915: 103 (South Africa species key); Gallardo, 1918b: 90 (Argentina species key); Mann, 1919: 303 (New Guinea & Solomon Is species key); Wheeler, W.M. 1922a: 96, 790, 1012 (diagnosis, Afrotropical, Malagasy catalogues); Wheeler, W.M. 1925a: 9 (Stenomyrmex species key); Chapman & Capco, 1951: 39 (Asia checklist); Wilson, 1959a: 503 (Melanesia & Moluccas species, revision, key); Kempf, 1964f: 244 (A. (Stenomyrmex) species key); Kempf, 1972a: 20 (Neotropical catalogue); Brown, 1978c: 552, 565, 569, 571 (diagnosis & review of genus, Malesian species, Austral, Palaearctic, Afrotropical & Malagasy species, Neotropical species keys); Taylor & Brown, D.R. 1985: 20 (Australia catalogue); Taylor, 1987a: 7 (Australia checklist); Terayama, 1989a: 26 (Taiwan species key); Brandão, 1991: 324 (Neotropical catalogue); Wang, M. 1993a: 226 (China species key); De Andrade, 1994: 24 (†Dominican Amber species key); Bolton, 1995a: 1048 (census); Bolton, 1995b: 63 (catalogue); Gronenberg & Ehmer, 1996: 183 (mandible morphology); Collingwood & Agosti, 1996: 309 (Saudi Arabia species key); Shattuck, 1999: 181 (Australia synopsis); Zhou, 2001b: 29 (China, Guangxi species key); Kugler, J. & Ionescu, 2007: 298 (Mediterraean & Middle East species key); Fisher & Smith, 2008: 4 (Malagasy species revision, key); González-Campero & Elizalde, 2008: 98 (Argentina & Paraguay species key); Zabala, in Fernández, 2008: 125 (Neotropical species key); Terayama, 2009: 117 (Taiwan species key).
Ponerinae Ponerini Leptogenys TRUE TRUE FALSEExtant: 216 species, 31 subspecies
Fossil: 1 species
Leptogenys Roger, 1861a: 41. Type-species: Leptogenys falcigera, by subsequent designation of Bingham, 1903: 52.
Taxonomic history
Leptogenys in Ponerinae: Dalla Torre, 1893: 46; Mayr, 1862: 714 (in key) [Poneridae]; Mayr, 1865: 14 [Poneridae].
Leptogenys in Ponerinae, Leptogenyini: Forel, 1893a: 162; Forel, 1899c: 17; Ashmead, 1905b: 382; Emery, 1911d: 97; Arnold, 1915: 89; Forel, 1917: 238; Gallardo, 1918b: 84; Wheeler, W.M. 1922a: 652; Donisthorpe, 1943f: 656; Chapman & Capco, 1951: 31; Dlussky & Fedoseeva, 1988: 78.
Leptogenys in Ponerinae, Ponerini: Emery, 1895j: 767; Forel, 1900d: 304; Wheeler, W.M. 1910g: 135; Brown, 1963: 3; Kempf, 1972a: 129; Jaffe, 1993: 8; Bolton, 1994: 164; Bolton, 2003: 163.
Junior synonyms of LEPTOGENYS
Lobopelta Mayr, 1862: 714 (diagnosis in key), 733. Type-species: Ponera diminuta, by subsequent designation of Bingham, 1903: 54.
Taxonomic history
Lobopelta in Ponerinae: Mayr, 1862: 714 (in key) [Poneridae]; Mayr, 1865: 15 [Poneridae]; Dalla Torre, 1893: 43.
Lobopelta in Ponerinae, Leptogenyini: Forel, 1893a: 162; Ashmead, 1905b: 382; Emery, 1911d: 101; Wheeler, W.M. 1922a: 653; Donisthorpe, 1943f: 658.
Lobopelta as genus: Mayr, 1862: 733; Cresson, 1887: 258; Dalla Torre, 1893: 43; Bingham, 1903: 54.
Lobopelta as subgenus of Leptogenys: Forel, 1892k: 520; Forel, 1899c: 18; Forel, 1900d: 304; Wheeler, W.M. 1910g: 135; Emery, 1911d: 101; Arnold, 1915: 95; Forel, 1917: 238; Gallardo, 1918b: 85; Wheeler, W.M. 1922a: 653; Wheeler, W.M. 1923d: 1; Borgmeier, 1923: 75; Creighton, 1950a: 50; Kempf, 1972a: 131; Smith, D.R. 1979: 1343 (anachronism).
Lobopelta as junior synonym of Leptogenys: Emery, 1896e: 177 (footnote); Brown, 1973b: 181 [provisional]; Bolton, 1975a: 240; Bolton, 2003: 163.
Prionogenys Emery, 1895g: 348. Type-species: Prionogenys podenzanai, by monotypy.
Taxonomic history
Prionogenys in Ponerinae, Ponerini: Emery, 1895j: 767; Wheeler, W.M. 1910g: 135;
Prionogenys in Ponerinae, Leptogenyini: Ashmead, 1905b: 382; Emery, 1911d: 106; Forel, 1917: 238; Wheeler, W.M. 1922a: 652; Donisthorpe, 1943f: 685; Dlussky & Fedoseeva, 1988: 78.
Prionogenys as junior synonym of Leptogenys: Taylor, 1988: 33.
Machaerogenys Emery, 1911d: 100 [as subgenus of Leptogenys]. Type-species: Leptogenys truncatirostris, by original designation.
Taxonomic history
Machaerogenys as subgenus of Leptogenys: Emery, 1911d: 100; Forel, 1917: 238; Wheeler, W.M. 1922a: 653; subsequent authors to the following.
Machaerogenys as junior synonym of Leptogenys: Brown, 1973b: 181 [provisional]; Bolton, 1975a: 240.
Odontopelta Emery, 1911d: 101 [as subgenus of Leptogenys]. Type-species: Leptogenys turneri, by monotypy.
Taxonomic history
Odontopelta as subgenus of Leptogenys: Emery, 1911d: 101; Forel, 1917: 238; Wheeler, W.M. 1922a: 653; Donisthorpe, 1943f: 677.
Odontopelta as junior synonym of Leptogenys: Brown, 1973b: 183 [provisional]; Snelling, R.R. 1981: 390; Taylor & Brown, D.R. 1985: 32.
Dorylozelus Forel, 1915b: 24. Type-species: Dorylozelus mjobergi (junior secondary homonym in Leptogenys, replaced by Leptogenys tricosa), by monotypy.
Taxonomic history
Dorylozelus in Ponerinae, Ponerini: Forel, 1917: 238.
Dorylozelus in Ponerinae, Dorylozelini: Wheeler, W.M. 1922a: 646; Donisthorpe, 1943f: 640.
Dorylozelus incertae sedis in Ponerinae, Amblyoponini: Brown, 1960a: 181.
Dorylozelus as junior synonym of Leptogenys: Taylor, 1969: 132; Bolton, 1975a: 240.
Microbolbos Donisthorpe, 1948f: 170. Type-species: Microbolbos testaceus, by original designation.
Taxonomic history
Microbolbos as junior synonym of Leptogenys: Wilson, 1955c: 136; Bolton, 1975a: 240.
Genus Leptogenys references
Roger, 1863b: 19 (Lobopelta, Leptogenys catalogues); Mayr, 1863: 426, 427 (Leptogenys, Lobopelta catalogues); Mayr, 1865: 14, 15 (Leptogenys, Lobopelta diagnoses); Mayr, 1867a: 89 (diagnosis); Mayr, 1879: 664 (Lobopelta species key); Cresson, 1887: 258 (U.S.A. catalogue); Dalla Torre, 1893: 43, 46 (Lobopelta, Leptogenys catalogues); Forel, 1900d: 304 (India & Sri Lanka species key); Bingham, 1903: 52, 55 (India, Sri Lanka & Burma Leptogenys, Lobopelta species keys); Emery, 1911d: 97 (diagnosis, subgenera key, catalogue); Arnold, 1915: 89, 95 (diagnosis, South Africa L. (Leptogenys) & L. (Lobopelta) species keys); Mann, 1919: 298 (Papuasia L. (Leptogenys) species key); Mann, 1921: 426 (Fiji L. (Lobopelta) species key); Wheeler, W.M. 1922a: 93, 786, 1010 (diagnosis, Afrotropical, Malagasy catalogues); Wheeler, W.M. 1923d: 14 (New World L. (Lobopelta) species key); Creighton, 1950a: 51 (North America L. (Lobopelta) species key); Chapman & Capco, 1951: 31 (Asia checklist); Wilson, 1958a: 113 (Melanesia & Moluccas species revision, key); Wilson, 1958a: 136 (New Caledonia species key); Kempf, 1972a: 129 (Neotropical catalogue); Alayo, 1974: 8 (Cuba species key); Bolton, 1975a: 239 (diagnosis, review of genus); Bolton, 1975a: 244 (Afrotropical species revision, key); Bolton, 1975a: 293 (Malagasy species key); Smith, D.R. 1979: 1343 (North America catalogue); Taylor & Brown, D.R. 1985: 32, 39 (Australia catalogue); Taylor, 1987a: 34, 64 (Australia, New Caledonia checklists); Trager & Johnson, 1988: 30 (U.S.A. species key); Brandão, 1991: 349 (Neotropical catalogue); Bolton, 1995a: 1050 (census); Bolton, 1995b: 229 (catalogue); Wu, J. & Wang, 1995: 41 (China species key); Shattuck, 1999: 189 (Australia synopsis); Xu, 2000a: 118 (China species key); Zhou, 2001b: 39 (China, Guangxi species key); Bolton, 2003: 163 (synopsis); Lattke, in Fernández, 2008: 145 (Colombia species, review); Terayama, 2009: 115 (Taiwan species key); Heterick, 2009: 133 (south-western Australia species key).
Ponerinae Ponerini Pachycondyla TRUE TRUE FALSEExtant: 252 species, 62 subspecies
Fossil: 12 species
Pachycondyla Smith, F. 1858b: 105. Type-species: Formica crassinoda, by subsequent designation of Emery, 1901a: 42.
Taxonomic history
Pachycondyla in Poneridae: Smith, F. 1858b: 105; Smith, 1871a: 324.
Pachycondyla in Ponerinae: Mayr, 1862: 713 [Poneridae]; Mayr, 1865: 13 [Poneridae]; Dalla Torre, 1893: 32.
Pachycondyla in Pachycondylinae, Pachycondylini: Ashmead, 1905b: 382.
Pachycondyla in Ponerinae, Ponerini: Emery, 1895j: 767; Forel, 1895b: 114; Forel, 1899c: 10; Wheeler, W.M. 1910g: 135; Emery, 1911d: 73 [subtribe Pachycondylini]; Arnold, 1915: 53; Forel, 1917: 237; Wheeler, W.M. 1922a: 648; all subsequent authors.
Junior synonyms of PACHYCONDYLA
Bothroponera Mayr, 1862: 713 (diagnosis in key), 717. Type-species: Ponera pumicosa, by subsequent designation of Emery, 1901a: 42.
Taxonomic history
Bothroponera in Ponerinae: Mayr, 1862: 713 (in key) [Poneridae]; Mayr, 1865: 13 [Poneridae]; Dalla Torre, 1893: 35.
Bothroponera in Pachycondylinae, Pachycondylini: Ashmead, 1905b: 382.
Bothroponera in Ponerinae, Ponerini: Emery, 1895j: 767; Emery, 1911d: 75 [subtribe Pachycondylini]; Wheeler, W.M. 1922a: 648; all subsequent authors.
Bothroponera as subgenus of Ponera: Emery, 1895j: 767; Forel, 1900d: 325.
Bothroponera as subgenus of Pachycondyla: Emery, 1901a: 42; Wheeler, W.M. 1910g: 135; Emery, 1911d: 75; Arnold, 1915: 55; Forel, 1917: 237; Donisthorpe, 1943f: 628.
Bothroponera as genus: Mayr, 1862: 717; Dalla Torre, 1893: 35; Bingham, 1903: 95; Ashmead, 1905b: 382; Wheeler, W.M. 1918c: 299 (footnote); Wheeler, W.M. 1922a: 648; Wheeler, W.M. & Chapman, 1925: 67; Chapman & Capco, 1951: 49; Bernard, 1953b: 187; Taylor, 1987a: 9; Taylor & Brown, D.R. 1985: 21; Wheeler, G.C. & Wheeler, J. 1985: 256; Dlussky & Fedoseeva, 1988: 78.
Bothroponera as junior synonym of Pachycondyla: Brown, 1973b: 179 [provisional]; Snelling, R.R. 1981: 389; Hölldobler & Wilson, 1990: 11; Brown, in Bolton, 1994: 164; Bolton, 2003: 166.
Megaponera Mayr, 1862: 714 (diagnosis in key), 734. Type-species: Formica foetens (junior primary homonym in Formica, replaced by Formica analis), by monotypy.
Taxonomic history
Megaponera in Ponerinae: Dalla Torre, 1893: 30; Mayr, 1862: 714 [Poneridae]; Mayr, 1865: 15 [Poneridae].
Megaponera in Pachycondylinae, Ectatommini: Ashmead, 1905b: 382.
Megaponera in Ponerinae, Ponerini: Emery, 1895j: 767; Wheeler, W.M. 1910g: 135; Emery, 1911d: 67 [subtribe Pachycondylini]; Arnold, 1915: 46; Wheeler, W.M. 1922a: 647; all subsequent authors to the following.
Megaponera as junior synonym of Pachycondyla: Brown, in Bolton, 1994: 164; Bolton, 2003: 166.
[Megaloponera Roger, 1863b: 17, incorrect subsequent spelling; misspelling repeated by several authors, for example Emery, 1877b: 368; Forel, 1917: 237.]
Paltothyreus Mayr, 1862: 714 (diagnosis in key), 735. Type-species: Formica tarsata, by monotypy.
Taxonomic history
Paltothyreus in Ponerinae: Mayr, 1862: 714 [Poneridae]; Mayr, 1865: 15 [Poneridae]; Dalla Torre, 1893: 32.
Paltothyreus in Pachycondylinae, Ectatommini: Ashmead, 1905b: 382.
Paltothyreus in Ponerinae, Ponerini: Emery, 1895j: 767; Wheeler, W.M. 1910g: 135; Emery, 1911d: 62 [subtribe Pachycondylini]; Arnold, 1915: 43; Forel, 1917: 237; Wheeler, W.M. 1922a: 647; all subsequent authors.
Paltothyreus as junior synonym of Pachycondyla: Brown, in Bolton, 1994: 164; Bolton, 2003: 166.
Ectomomyrmex Mayr, 1867a: 83. Type-species: Ectomomyrmex javanus, by subsequent designation of Bingham, 1903: 85.
Taxonomic history
Ectomomyrmex in Ponerinae: Dalla Torre, 1893: 30.
Ectomomyrmex in Pachycondylinae, Pachycondylini: Ashmead, 1905b: 382.
Ectomomyrmex in Ponerinae, Ponerini: Emery, 1895j: 767; Forel, 1900d: 321; Emery, 1911d: 78 [subtribe Pachycondylini]; Wheeler, W.M. 1922a: 648; all subsequent authors.
Ectomomyrmex as subgenus of Pachycondyla: Emery, 1901a: 42; Wheeler, W.M. 1910g: 135; Emery, 1911d: 78; Arnold, 1915: 53; Forel, 1917: 237.
Ectomomyrmex as genus: Mayr, 1867a: 83; Dalla Torre, 1893: 30; Ashmead, 1905b: 382; Forel, 1900d: 321; Bingham, 1903: 85; Wheeler, W.M. 1922a: 648; Chapman & Capco, 1951: 60; Brown, 1963: 9; Taylor & Brown, D.R. 1985: 29; Taylor, 1987a: 26; Ogata, 1987: 112.
Ectomomyrmex as junior synonym of Pachycondyla: Brown, 1973b: 180 [provisional]; Snelling, R.R. 1981: 389; Hölldobler & Wilson, 1990: 11; Brown, in Bolton, 1994: 164; Bolton, 2003: 166.
[Ectomyrmex Donisthorpe, 1943f: 641, incorrect subsequent spelling.]
Ophthalmopone Forel, 1890b: cxii. Type-species: Ophthalmopone berthoudi, by monotypy.
Taxonomic history
Ophthalmopone in Ponerinae: Dalla Torre, 1893: 31.
Ophthalmopone in Pachycondylinae, Pachycondylini: Ashmead, 1905b: 382.
Ophthalmopone in Ponerinae, Ponerini: Emery, 1895j: 767; Wheeler, W.M. 1910g: 135; Emery, 1911d: 69 [subtribe Pachycondylini]; Arnold, 1915: 49; Forel, 1917: 237; Wheeler, W.M. 1922a: 647; all subsequent authors.
Ophthalmopone as junior synonym of Pachycondyla: Brown, in Bolton, 1994: 164; Bolton, 2003: 166.
Euponera Forel, 1891b: 126 [as subgenus of Ponera]. Type-species: Ponera (Euponera) sikorae, by monotypy.
Taxonomic history
Euponera in Ponerinae, Ponerini: Emery, 1895j: 767; Emery, 1911d: 79 [subtribe Pachycondylini]; Wheeler, W.M. 1915h: 37; Arnold, 1915: 63; Forel, 1917: 237; Wheeler, W.M. 1922a: 648; all subsequent authors.
Euponera as subgenus of Ponera: Forel, 1891b: 126; Emery, 1895j: 767.
Euponera as genus: Emery, 1901a: 46; Emery, 1909c: 364; Wheeler, W.M. 1910g: 135; Emery, 1911d: 79; Arnold, 1915: 63; Forel, 1917: 237; Gallardo, 1918b: 64; Wheeler, W.M. 1922a: 648; Borgmeier, 1923: 70; Clark, 1934b: 30; Creighton, 1950a: 44; Chapman & Capco, 1951: 63; Bernard, 1953b: 189; Wheeler, G.C. & Wheeler, J. 1985: 256; Dlussky & Fedoseeva, 1988: 78; Hölldobler & Wilson, 1990: 11.
Euponera as junior synonym of Pachycondyla: Brown, in Bolton, 1994: 164; Bolton, 2003: 166.
Cryptopone Emery, 1893a: cclxxv. Type-species: Cryptopone testacea, by monotypy.
Taxonomic history
[Cryptopone also described as new by Emery, 1893f: 240. Type-species not Amblyopone testacea, unjustified subsequent designation by Wheeler, W.M. 1911f: 161, repeated in Emery, 1911d: 88, Wheeler, W.M. 1922a: 780, Donisthorpe, 1943f: 636, Kempf, 1972a: 90 and Taylor & Brown, D.R. 1985: 28; see discussion in Wilson, 1958d: 360.]
Cryptopone in Ponerinae, Ponerini: Emery, 1895j: 767; Ashmead, 1905b: 382; Wheeler, W.M. 1910g: 135; Emery, 1911d: 88 [subtribe Ponerini]; Forel, 1917: 238; Wheeler, W.M. 1922a: 650; all subsequent authors.
Cryptopone as junior synonym of Pachycondyla: Mackay & Mackay, 2010: 3.
Pseudoponera Emery, 1900c: 314 [as subgenus of Pachycondyla]. Type-species: Ponera quadridentata (junior synonym of Formica stigma), by monotypy.
Taxonomic history
[Pseudoponera also described as new by Emery, 1901a: 42. Type-species not Ponera amblyops, unjustified subsequent designation by Emery, 1901a: 42; repeated in Wheeler, W.M. 1911f: 171, Wheeler, W.M. 1922a: 779; Donisthorpe, 1943g: 723.]
Pseudoponera in Pachycondylinae, Pachycondylini: Ashmead, 1905b: 382.
Pseudoponera in Ponerinae, Ponerini: Emery, 1911d: 86 [subtribe Pachycondylini]; Forel, 1917: 238; all subsequent authors.
Pseudoponera as subgenus of Pachycondyla: Emery, 1900c: 314; Emery, 1901a: 42.
Pseudoponera as subgenus of Euponera: Forel, 1900c: 141; Forel, 1900e: 398; Emery, 1909c: 364; Wheeler, W.M. 1910g: 135.
Pseudoponera as genus: Bingham, 1903: 91; Ashmead, 1905b: 382; Emery, 1911d: 86; Forel, 1917: 238; Wheeler, W.M. 1922a: 649; Donisthorpe, 1943g: 722; Chapman & Capco, 1951: 74.
Pseudoponera as junior synonym of Pachycondyla: Brown, 1973b: 184 [provisional]; Snelling, R.R. 1981: 389; Hölldobler & Wilson, 1990: 11; Brown, in Bolton, 1994: 164; Bolton, 2003: 167.
Brachyponera Emery, 1900c: 315 [as subgenus of Euponera]. Type-species: Euponera (Brachyponera) croceicornis, by monotypy.
Taxonomic history
[Brachyponera also described as new by Emery, 1901a: 43. Type-species not Ponera sennaarensis, unjustified subsequent designation by Emery, 1901a: 43, repeated in Wheeler, W.M. 1911f: 160, Emery, 1911d: 84, Wheeler, W.M. 1922a: 777, Donisthorpe, 1943f: 628, Wilson, 1958d: 346; Bolton, 1973a: 335; Taylor & Brown, D.R. 1985: 23.]
Brachyponera in Pachycondylinae, Pachycondylini: Ashmead, 1905b: 382.
Brachyponera in Ponerinae, Ponerini: Emery, 1911d: 83; all subsequent authors.
Brachyponera as subgenus of Euponera: Emery, 1900c: 315; Emery, 1909c: 366; Wheeler, W.M. 1910g: 135; Emery, 1911d: 83; Arnold, 1915: 72; Forel, 1917: 237; Wheeler, W.M. 1922a: 649; subsequent authors to the following.
Brachyponera as genus: Bingham, 1903: 101; Ashmead, 1905b: 382; Wilson, 1958d: 346; Taylor & Brown, D.R. 1985: 23; Wheeler, G.C. & Wheeler, J. 1985: 256; Taylor, 1987a: 9; Ogata, 1987: 116; Dlussky & Fedoseeva, 1988: 78; Hölldobler & Wilson, 1990: 10.
Brachyponera as junior synonym of Pachycondyla: Snelling, R.R. 1981: 389 [provisional]; Brown, in Bolton, 1994: 164; Bolton, 2003: 167.
Mesoponera Emery, 1900d: 668 [as subgenus of Euponera]. Type-species: Ponera melanaria, by monotypy.
Taxonomic history
[Mesoponera also described as new by Emery, 1901a: 43. Type-species not Ponera caffraria, unjustified subsequent designation by Emery, 1901a: 43, repeated by Emery, 1911d: 81, Wheeler, W.M. 1911f: 167, Wheeler, W.M. 1922a: 775, Donisthorpe, 1943f: 661, Wilson, 1958d: 349; Kempf, 1972a: 141, Bolton, 1973a: 338; Taylor & Brown, D.R. 1985: 35.]
Mesoponera in Pachycondylinae, Pachycondylini: Ashmead, 1905b: 382.
Mesoponera in Ponerinae, Ponerini: Emery, 1911d: 80; all subsequent authors.
Mesoponera as subgenus of Euponera: Emery, 1900d: 668; Wheeler, W.M. 1910g: 135; Emery, 1911d: 80; Arnold, 1915: 64; Forel, 1917: 237; Gallardo, 1918b: 65; Wheeler, W.M. 1922a: 649; Borgmeier, 1923: 71; Donisthorpe, 1943f: 661; Chapman & Capco, 1951: 64; Bernard, 1953b: 190.
Mesoponera as genus: Bingham, 1903: 99; Ashmead, 1905b: 382; Wilson, 1958d: 349; Kempf, 1972a: 141; Taylor & Brown, D.R. 1985: 34; Taylor, 1987a: 39; Dlussky & Fedoseeva, 1988: 78.
Mesoponera as junior synonym of Pachycondyla: Brown, 1973b: 182 [provisional]; Snelling, R.R. 1981: 389; Hölldobler & Wilson, 1990: 11; Brown, in Bolton, 1994: 164; Bolton, 2003: 167.
Eumecopone Forel, 1901e: 335 [as subgenus of Neoponera]. Type-species: Neoponera (Eumecopone) agilis, by monotypy.
Taxonomic history
Eumecopone as subgenus of Neoponera: Forel, 1901e: 335; Wheeler, W.M. 1910g: 135; Emery, 1911d: 71; Forel, 1917: 237; Wheeler, W.M. 1922a: 648; subsequent authors to the following.
Eumecopone junior synonym of Pachycondyla: Brown, 1973b: 180 [provisional]; Snelling, R.R. 1981: 389; Hölldobler & Wilson, 1990: 11; Brown, in Bolton, 1994: 164; Bolton, 2003: 167.
Hagensia Forel, 1901f: 333 [as subgenus of Megaloponera (sic)]. Type-species: Megaloponera (Hagensia) havilandi, by monotypy.
Taxonomic history
Hagensia in Ponerinae, Ponerini: Emery, 1911d: 69 [subtribe Pachycondylini]; all subsequent authors.
Hagensia as subgenus of Megaponera: Forel, 1901f: 333; Wheeler, W.M. 1910g: 135; Emery, 1911d: 61.
Hagensia as subgenus of Euponera: Forel, 1917: 237.
Hagensia as genus: Arnold, 1926: 202; Donisthorpe, 1943f: 648; Wheeler, G.C. & Wheeler, J. 1985: 256; Dlussky & Fedoseeva, 1988: 78; Hölldobler & Wilson, 1990: 11.
Hagensia as junior synonym of Pachycondyla: Brown, in Bolton, 1994: 164; Bolton, 2003: 167.
Neoponera Emery, 1901a: 43. Type-species: Formica villosa, by original designation.
Taxonomic history
Neoponera in Pachycondylinae, Pachycondylini: Ashmead, 1905b: 382.
Neoponera in Ponerinae, Ponerini: Wheeler, W.M. 1910g: 135; Emery, 1911d: 70 [subtribe Pachycondylini]; Forel, 1917: 237; Wheeler, W.M. 1922a: 648; all subsequent authors.
Neoponera as junior synonym of Pachycondyla: Brown, 1973b: 183 [provisional]; Snelling, R.R. 1981: 389; Hölldobler & Wilson, 1990: 11; Brown, in Bolton, 1994: 164; Bolton, 2003: 168.
Trachymesopus Emery, 1911d: 84 [as subgenus of Euponera]. Type-species: Formica stigma, by original designation.
Taxonomic history
Trachymesopus in Ponerinae, Ponerini: Emery, 1911d: 84; all subsequent authors.
Trachymesopus as subgenus of Euponera: Emery, 1911d: 84; Arnold, 1915: 74; Forel, 1917: 237; Wheeler, W.M. 1922a: 649; Borgmeier, 1923: 72; Donisthorpe, 1943g: 733; Creighton, 1950a: 46; Chapman & Capco, 1951: 65; Smith, M.R. 1951a: 786; Bernard, 1953b: 195.
Trachymesopus as genus: Wilson, 1958d: 352; Kempf, 1960f: 423; Brown, 1963: 6; Kempf, 1972a: 251; Taylor & Brown, D.R. 1985: 52; Ogata, 1987: 114; Dlussky & Fedoseeva, 1988: 78.
Trachymesopus as junior synonym of Pachycondyla: Brown, 1973b: 185 [provisional]; Snelling, R.R. 1981: 389; Hölldobler & Wilson, 1990: 11; Brown, in Bolton, 1994: 164.
Trachymesopus as junior synonym of Pseudoponera: Bolton, 1995b: 48 (because of synonymous type-species); Bolton, 2003: 168.
[Trachyponera Santschi, 1928a: 43, incorrect subsequent spelling.]
Xiphopelta Forel, 1913a: 108 [as subgenus of Ponera]. Type-species: Ponera (Xiphopelta) arnoldi, by monotypy.
Taxonomic history
Xiphopelta as subgenus of Ponera: Forel, 1913a: 108; Donisthorpe, 1943g: 737.
Xiphopelta as subgenus of Euponera: Forel, 1917: 237; Emery, 1919c: 106; Bernard, 1953b: 191.
Xiphopelta as junior synonym of Mesoponera: Wheeler, W.M. 1922a: 775; all subsequent authors to the following.
Xiphopelta as junior synonym of Pachycondyla: Brown, 1973b: 185 [provisional]; Snelling, R.R. 1981: 389; Hölldobler & Wilson, 1990: 11; Brown, in Bolton, 1994: 164; Bolton, 2003: 168.
[Hiphopelta Forel, 1913a: 108, incorrect subsequent spelling.]
Termitopone Wheeler, W.M. 1936d: 159. Type-species: Ponera laevigata, by original designation.
Taxonomic history
Termitopone in Ponerinae, Ponerini: Donisthorpe, 1943g: 731.
Termitopone as junior synonym of Pachycondyla: Brown, 1973b: 185 [provisional]; Snelling, R.R. 1981: 389; Hölldobler & Wilson, 1990: 11; Brown, in Bolton, 1994: 164; Bolton, 2003: 168.
Syntermitopone Wheeler, W.M. 1936d: 169 [as subgenus of Termitopone.] Type-species: Ponera commutata, by original designation.
Taxonomic history
Syntermitopone as subgenus of Termitopone: Wheeler, W.M. 1936d: 169.
Syntermitopone as genus: Kusnezov, 1956: 15.
Syntermitopone as junior synonym of Termitopone: Borgmeier, 1959a: 312.
Syntermitopone as junior synonym of Pachycondyla: Brown, 1973b: 185 [provisional]; Snelling, R.R. 1981: 389; Hölldobler & Wilson, 1990: 11; Brown, in Bolton, 1994: 164; Bolton, 2003: 168.
Wadeura Weber, 1939a: 102. Type-species: Wadeura guianensis, by original designation.
Taxonomic history
Wadeura in Ponerinae, Ponerini: Donisthorpe, 1943g: 735.
Wadeura as junior synonym of Pachycondyla: Brown, 1973b: 185 [provisional]; Snelling, R.R. 1981: 389; Hölldobler & Wilson, 1990: 11; Brown, in Bolton, 1994: 164; Bolton, 2003: 168.
Pseudoneoponera Donisthorpe, 1943d: 439. Type-species: Pseudoneoponera verecundae, by original designation.
Taxonomic history
Pseudoneoponera in Ponerinae, Pseudoneoponerini: Chapman & Capco, 1951: 77.
Pseudoneoponera as junior synonym of Bothroponera: Wilson, 1958d: 361.
Pseudoneoponera as junior synonym of Pachycondyla: Brown, 1973b: 184 [provisional]; Snelling, R.R. 1981: 389; Hölldobler & Wilson, 1990: 11; Brown, in Bolton, 1994: 164; Bolton, 2003: 168.
Genus Pachycondyla references
Mayr, 1862: 719 (diagnosis); Roger, 1863b: 16, 17, 18 (catalogue); Mayr, 1863: 397, 428, 439, 440 (catalogue); Mayr, 1865: 13, 15 (Bothroponera, Pachycondyla, Megaponera, Paltothyreus diagnoses); Mayr, 1867a: 82 (Bothroponera diagnosis); Mayr, 1870a: 396 (Colombia + Panama (= New Grenada) species key); Emery, 1887b: 18 (footnote) (Asia & Australia Bothroponera species key); Emery, 1890a: 71 (Neotropical species key); Dalla Torre, 1893: 30, 31, 32, 35 (Ectomomyrmex, Megaponera, Ophthalmopone, Paltothyreus, Pachycondyla, Bothroponera catalogues); Forel, 1900d: 321 (India & Sri Lanka Ectomomyrmex species key); Bingham, 1903: 86, 91, 96, 101 (India, Sri Lanka & Burma Ectomomyrmex, Pseudoponera, Bothroponera, Brachyponera species keys); Emery, 1911d: 62 (Paltothyreus diagnosis, catalogue); Emery, 1911d: 67 (Megaponera diagnosis, catalogue); Emery, 1911d: 69 (Megaponera (Hagensia) and Ophthalmopone diagnoses, catalogues); Emery, 1911d: 70 (Neoponera diagnosis, catalogue); Emery, 1911d: 73 (diagnosis, subgenera key, catalogue); Emery, 1911d: 75 (P. (Bothroponera) diagnosis, catalogue); Emery, 1911d: 78 (P. (Ectomomyrmex) diagnosis, catalogue); Emery, 1911d: 80 (Euponera diagnosis, subgenera key, catalogue); Emery, 1911d: 80 (Euponera (Mesoponera) diagnosis, catalogue); Emery, 1911d: 83 (Euponera (Brachyponera) diagnosis, catalogue); Emery, 1911d: 84 (Euponera (Trachymesopus) diagnosis, catalogue); Emery, 1911d: 86 (Pseudoponera diagnosis, catalogue); Emery, 1911d: 88 (Cryptopone diagnosis, catalogue); Arnold, 1915: 43, 46, 49, 53, 63 (Paltothyreus, Megaponera, Ophthalmopone, Pachycondyla, Euponera diagnoses); Arnold, 1915: 50, 55, 64 (South Africa Ophthalmopone, Bothroponera, Euponera (Mesoponera) species keys); Gallardo, 1918b: 56, 66 (Argentina Neoponera, Mesoponera, keys); Wheeler, W.M. 1922a: 72 (Afrotropical Bothroponera species key); Wheeler, W.M. 1922a: 60, 63, 69, 81 (Paltothyreus, Megaponera, Bothroponera, Euponera diagnoses); Wheeler, W.M. 1922a: 763, 766, 768, 769, 774, 780 (Afrotropical Paltothyreus, Megaponera, Ophthalmopone, Bothroponera, Ectomomyrmex, Euponera, Pseudoponera, Cryptopone catalogues); Wheeler, W.M. 1922a: 1007, 1008 (Malagasy Bothroponera, Euponera catalogues); Arnold, 1926: 203 (South Africa Hagensia species key); Wheeler, W.M. 1933g: 5 (Cryptopone all species key); Santschi, 1935b: 259 (Africa Mesoponera, key); Creighton, 1950a: 45 (North America Euponera species key); Arnold, 1951: 53 (Hagensia species key); Chapman & Capco, 1951: 49, 52, 60, 63, 74 (Asia Bothroponera, Cryptopone, Ectmomomyrmex, Euponera, Pseudoponera checklists); Arnold, 1952a: 461 (South Africa Bothroponera, key); Bernard, 1953b: 191 (Guinea Euponera (Xiphopelta) species key); Brown, 1958h: 20 (New Zealand species); Wilson, 1958d: 346, 357, 362 (Melanesia Brachyponera, Cryptopone, Ectomomyrmex species keys); Kempf, 1961b: 192 (Brazil species key); Brown, 1963: 6, 9 (reviews of Cryptopone, Trachymesopus & Ectomomyrmex); Kempf, 1964e: 51 (Neotropical Pachycondyla species key); Kusnezov, 1969: 36 (Argentina Neoponera species key); Kempf, 1972a: 90, 141, 160, 174, 248, 251, 257 (Neotropical Cryptopone, Mesoponera, Neoponera, Pachycondyla, Termitopone, Trachymesopus, Wadeura catalogues); Alayo, 1974: 7 (Cuba species key); Smith, D.R. 1979: 1339, 1341 (North America, Pachycondyla, Brachyponera, Cryptopone catalogues); Taylor & Brown, D.R. 1985: 21, 23, 28, 29, 34, 34, 52 (Australia, catalogue); Taylor, 1987a: 9, 23, 26, 39, 80 (Australia, New Caledonia & New Zealand checklists); Morisita, Kubota, et al. 1989: 18 (Japan Ectomomyrmex, Cryptopone, Brachyponera, Trachymesopus species key); Brandão, 1991: 354, 360, 365, 381, 382 (Neotropical catalogue); Xu, 1994b: 182 (China Brachyponera species key); Bolton, 1994: 164 (synoptic classification); Bolton, 1995a: 1051 (census); Bolton, 1995b: 166, 302 (Cryptopone, Pachycondyla catalogues); Wu, J. & Wang, 1995: 45 (China species key); Xu, 1995b: 104 (China species key); Collingwood & Agosti, 1996: 310 (Cryptopone, Pachycondyla Saudi Arabia species key); Kim, Kim & Kim, 1998: 147 (Korea species key); Shattuck, 1999: 182, 196 (Australia Cryptopone, Pachycondyla fauna synopsis); Zhou, 2001b: 35, 49 (China, Guangxi Cryptopone, Pachycondyla species keys); Dlussky & Rasnitsyn, 2002: 424 (wingless female fossils, diagnosis); Wild, 2002: 4 (Paraguay species key); Bolton, 2003: 161, 165 (Cryptopone, Pachycondyla synopses); Wild, 2005: 5 (P. apicalis complex key); Yamane, 2007: 652 (P. nigrita species group key); Mackay, Mackay, Fernández & Arias-Penna, in Fernández, 2008: 172 (Neotropical species key); Terayama, 2009: 105, 108 (Taiwan Pachycondyla, Cryptopone species keys); Heterick, 2009: 134 (south-western Australia species key); Mackay & Mackay, 2010: 26, 107 (New World Pachycondyla species revision, species complexes key and species key).
Ponerinae Ponerini Plectroctena TRUE TRUE FALSE16 species
Plectroctena Smith, F. 1858b: 101. Type-species: Plectroctena mandibularis, by monotypy.
Taxonomic history
Plectroctena in Poneridae: Smith, F. 1858b: 101.
Plectroctena in Ponerinae: Mayr, 1862: 713 [Poneridae]; Mayr, 1865: 12 [Poneridae]; Dalla Torre, 1893: 31.
Plectroctena in Pachycondylinae, Pachycondylini: Ashmead, 1905b: 382.
Plectroctena in Ponerinae, Ponerini: Emery, 1895j: 767; Wheeler, W.M. 1910g: 135; Emery, 1911d: 94 [subtribe Plectroctenini]; Arnold, 1915: 84; Forel, 1917: 238; Wheeler, W.M. 1922a: 650; all subsequent authors.
Junior synonym of PLECTROCTENA
Cacopone Santschi, 1914d: 325. Type-species: Cacopone hastifer, by monotypy.
Taxonomic history
Cacopone in Ponerinae, Ponerini: Forel, 1917: 238 [subtribe Plectroctenini]; Wheeler, W.M. 1922a: 650; Santschi, 1924a: 159; Donisthorpe, 1943f: 629.
Cacopone as junior synonym of Plectroctena: Bolton, 1974c: 313.
Genus Plectroctena references
Roger, 1863b: 19 (catalogue); Mayr, 1863: 442 (catalogue); Mayr, 1865: 12 (diagnosis); Dalla Torre, 1893: 31 (catalogue); Emery, 1911d: 94 (diagnosis, catalogue); Arnold, 1915: 84 (diagnosis, South Africa species); Wheeler, W.M. 1922a: 85, 783, 785, 786 (diagnosis, Plectroctena, Myopias, Cacopone catalogues); Santschi, 1924a: 172 (all species revision, key); Bolton, 1974c: 309 (diagnosis, all species revision, key); Bolton, 1994: 164 (synoptic classification); Bolton, 1995a: 1052 (census); Bolton, 1995b: 337 (catalogue); Bolton & Brown, 2002: 11 (all species key); Bolton, 2003: 169 (synopsis).
Ponerinae Ponerini Ponera TRUE TRUE FALSEExtant: 52 species
Fossil: 5 species
Ponera Latreille, 1804: 179. Type-species: Formica coarctata, by subsequent designation of Westwood, 1840a: 83 (see discussion in Taylor, 1967a: 6).
Taxonomic history
Ponera in Ponérites: Lepeletier de Saint-Fargeau, 1835: 188.
Ponera in Poneridae: Smith, F. 1858b: 83; Smith, F. 1871a: 320.
Ponera in Ponerinae: Mayr, 1855: 386 [Poneridae]; Smith, F. 1857a: 65 [Poneridae]; Mayr, 1861: 54 [Poneridae]; Mayr, 1862: 713 [Poneridae]; Mayr, 1865: 13 [Poneridae]; Emery & Forel, 1879a: 455 [Poneridae]; Dalla Torre, 1893: 37.
Ponera in Ponerinae, Ponerini: Emery, 1895j: 767; Forel, 1899c: 15; Ashmead, 1905b: 382; Wheeler, W.M. 1910g: 135; Emery, 1911d: 88 [subtribe Ponerini]; Wheeler, W.M. 1915h: 38; Arnold, 1915: 77; Forel, 1917: 238; Wheeler, W.M. 1922a: 650; all subsequent authors.
Junior synonyms of PONERA
Pseudocryptopone Wheeler, W.M. 1933g: 12. Type-species: Cryptopone tenuis, by original designation.
Taxonomic history
Pseudocryptopone in Ponerinae, Ponerini: Donisthorpe, 1943g: 722.
Pseudocryptopone as junior synonym of Ponera: Wilson, 1957b: 356.
Selenopone Wheeler, W.M. 1933g: 19. Type-species: Ponera selenophora, by original designation.
Taxonomic history
Selenopone in Ponerinae, Ponerini: Donisthorpe, 1943g: 725.
Selenopone as junior synonym of Ponera: Wilson, 1957b: 356.
Pteroponera Bernard, 1950a: 3. Type-species: Pteroponera sysphinctoides, by monotypy.
Taxonomic history
Pteroponera as junior synonym of Ponera: Brown, 1973b: 184 [provisional]; Brown, in Bolton, 1994: 164.
Genus Ponera references
[References before Taylor, 1967a refer to both Ponera and Hypoponera.]
Mayr, 1855: 386 (diagnosis); Smith, F. 1858b: 83 (diagnosis); Mayr, 1862: 721 (diagnosis); Roger, 1863b: 16 (catalogue); Mayr, 1863: 447 (catalogue); Mayr, 1865: 13 (diagnosis); Mayr, 1867a: 85 (diagnosis); Forel, 1874: 64 (Switzerland species key); Mayr, 1879: 661 (all species key); André, 1882c: 239 (Europe & Algeria species key); Cresson, 1887: 258 (U.S.A. catalogue); Dalla Torre, 1893: 37 (catalogue); Emery, 1895b: 60 (Mediterranean species key); Emery, 1896g: 53 (New World species key); Emery, 1900c: 316 (Australia, Papuasia & Oceania species key); Forel, 1900d: 322 (India & Sri Lanka species key); Bingham, 1903: 89 (India, Sri Lanka & Burma species key); Emery, 1909c: 367 (Palaearctic species key); Emery, 1911d: 88 (diagnosis, catalogue); Arnold, 1915: 76 (diagnosis, South Africa species key); Emery, 1916b: 107 (Italy species key); Bondroit, 1918: 81 (France & Belgium species key); Gallardo, 1918b: 73 (Argentina species key); Mann, 1921: 419 (Fiji species key); Wheeler, W.M. 1922a: 780, 1009 (Afrotropical, Malagasy catalogues); Menozzi, 1931b: 266 (Costa Rica species key); Wheeler, W.M. 1933g: 13 (Pseudocryptopone species key); Smith, M.R. 1936: 420 (North America species key); Stitz, 1939: 58 (Germany species key); Creighton, 1950a: 47 (North America species key); Chapman & Capco, 1951: 68, 73, 74 (Asia Ponera, Pseudocryptopone, Selenopone checklists); Bernard, 1953b: 197 (West Africa species key); Wilson, 1957b: 359, 376 (P. tenuis, P. selenophora groups revisions, keys); Wilson, 1958d: 323, 343 (Melanesia & Moluccas species, Fiji species revisions, keys); Brown, 1958h: 22 (New Zealand species); Taylor, 1960: 180 (P. leae complex, key); Taylor, 1964: 139 (†fossil species, review); Taylor, 1967a: 18 (diagnosis, all species revision, key); Bernard, 1967: 83 (diagnosis, Western Europe species key); Smith, D.R. 1979: 1341 (North America catalogue); Taylor & Brown, D.R. 1985: 38 (Australia catalogue); Taylor, 1987a: 63 (Australia, New Caledonia & New Zealand, checklist); Morisita, Kubota, et al. 1989: 22 (Japan species key); Perrault, 1993: 337 (addition to Taylor, 1967a key); Bolton, 1994: 164 (synoptic classification); Bolton, 1995a: 1052 (census); Bolton, 1995b: 360 (catalogue); Terayama, 1996: 14 (Japan species key); Kim, Kim & Kim, 1998: 149 (Korea species key); Shattuck, 1999: 199 (Australia synopsis); Xu, 2001a: 52 and Xu, 2001c: 218 (China species keys); Zhou, 2001b: 37 (China, Guangxi species key); Bolton, 2003: 170 (synopsis); Yoshimura, Hosoishi, et al. 2009: 200 (Japan species key); Terayama, 2009: 109 (Taiwan species key).
Formicinae Lasiini Acropyga arnoldi TRUE TRUE FALSEarnoldi. Acropyga (Malacomyrma) arnoldi Santschi, 1926b: 245 (w.q.) SOUTH AFRICA. Prins, 1982: 245 (m.). Senior synonym of rhodesiana: Prins, 1982: 238. See also: LaPolla, 2004a: 35.
Aenictinae Aenictini Aenictus decolor TRUE TRUE FALSEdecolor. Typhlatta decolor Mayr, 1879: 668 (diagnosis in key) (w.) EAST AFRICA. Gotwald & Leroux, 1980: 602 (q.). Combination in Aenictus: Dalla Torre, 1893: 7. Senior synonym of batesi, bidentatus: Gotwald & Leroux, 1980: 600.
Aenictinae Aenictini Aenictus eugenii TRUE TRUE FALSE2 subspecies
eugenii. Aenictus eugenii Emery, 1895h: 17, pl. 2, figs. 1-4 (w.) SOUTH AFRICA. Gotwald & Cunningham-Van Someren, 1976: 183 (q.). Senior synonym of kenyensis: Gotwald & Cunningham-Van Someren, 1976: 183. Current subspecies: nominal plus caroli, henrii.
Aenictinae Aenictini Aenictus inconspicuus TRUE TRUE FALSEinconspicuus. Aenictus inconspicuus Westwood, 1845: 85 (m.) SOUTH AFRICA. [Also described as new by Westwood, 1847a: 238.]
Aenictinae Aenictini Aenictus mariae TRUE TRUE FALSE1 subspecies
mariae. Aenictus mariae Emery, 1895h: 18, pl. 2, figs. 5-7 (w.) SOUTH AFRICA. Current subspecies: nominal plus natalensis.
Aenictinae Aenictini Aenictus rixator TRUE TRUE FALSErixator. Aenictus rixator Forel, in Emery, 1901d: 48 (w.) SOUTH AFRICA.
Aenictinae Aenictini Aenictus rotundatus TRUE TRUE FALSE2 subspecies
rotundatus. Aenictus rotundatus Mayr, 1901b: 1 (w.) SOUTH AFRICA. Current subspecies: nominal plus guineensis, merwei.
Aenictinae Aenictini Aenictus steindachneri TRUE TRUE FALSEsteindachneri. Aenictus steindachneri Mayr, 1901b: 2 (w.) SOUTH AFRICA.
Aenictinae Aenictini Aenictus mariae natalensis TRUE TRUE FALSEnatalensis. Aenictus mariae var. natalensis Forel, in Emery, 1901d: 49 (w.) SOUTH AFRICA.
Aenictinae Aenictini Aenictus rotundatus merwei TRUE TRUE FALSEmerwei. Aenictus rotundatus var. merwei Santschi, 1932a: 382 (w.) SOUTH AFRICA.
Formicinae Plagiolepidini Agraulomyrmex meridionalis TRUE TRUE FALSEmeridionalis. Agraulomyrmex meridionalis Prins, 1983: 2, fig. 1 (w.) SOUTH AFRICA.
Ponerinae Ponerini Anochetus africanus TRUE TRUE FALSEafricanus. Stenomyrmex africanus Mayr, 1865: 11 (footnote) (w.) GHANA. André, 1892a: 47 (q.). Combination in Anochetus: Forel, 1887: 382. Senior synonym of camerunensis: Mayr, 1904b: 2; of pasteuri: Brown, 1978c: 556 (see also p. 602).
Ponerinae Ponerini Anochetus faurei TRUE TRUE FALSEfaurei. Anochetus faurei Arnold, 1948: 215, fig. 4 (w.) SOUTH AFRICA. See also: Brown, 1978c: 609.
Ponerinae Ponerini Anochetus fuliginosus TRUE TRUE FALSEfuliginosus. Anochetus fuliginosus Arnold, 1948: 214, fig. 3 (w.) SOUTH AFRICA. See also: Brown, 1978c: 605.
Ponerinae Ponerini Anochetus jonesi TRUE TRUE FALSEjonesi. Anochetus jonesi Arnold, 1926: 216, fig. 61 (w.q.m.) SOUTH AFRICA.
Ponerinae Ponerini Anochetus levaillanti TRUE TRUE FALSElevaillanti. Anochetus levaillanti Emery, 1895h: 21 (w.) SOUTH AFRICA. Arnold, 1915: 108 (q.). See also: Brown, 1978c: 594.
Ponerinae Ponerini Anochetus madagascarensis TRUE TRUE FALSEmadagascarensis. Anochetus africanus var. madagascarensis Forel, 1887: 382 (w.) MADAGASCAR. Fisher & Smith, 2008: 12 (q.m.). Raised to species and senior synonym of friederichsi: Brown, 1978c: 557.
Ponerinae Ponerini Anochetus natalensis TRUE TRUE FALSEnatalensis. Anochetus natalensis Arnold, 1926: 215 (w.) SOUTH AFRICA. See also: Brown, 1978c: 603.
Ponerinae Ponerini Anochetus obscuratus TRUE TRUE FALSEobscuratus. Anochetus madagascarensis var. obscurata Santschi, 1911c: 351 (w.q.) TANZANIA. Subspecies of africanus: Emery, 1911d: 108; Santschi, 1914b: 56. Raised to species: Santschi, 1923e: 265. Senior synonym of schoutedeni, ustus: Brown, 1978c: 558 (see also p. 602).
Ponerinae Ponerini Anochetus punctaticeps TRUE TRUE FALSEpunctaticeps. Anochetus punctaticeps Mayr, 1901b: 4 (w.) SOUTH AFRICA. See also: Brown, 1978c: 606.
Ponerinae Ponerini Anochetus talpa TRUE TRUE FALSEtalpa. Anochetus talpa Forel, 1901f: 351 (w.) SOUTH AFRICA.
Formicinae Lasiini Anoplolepis custodiens TRUE TRUE FALSE3 subspecies
custodiens. Formica custodiens Smith, F. 1858b: 33 (w.) SOUTH AFRICA. Mayr, 1895: 148 (m.); Emery, 1895h: 44 (q.); Wheeler, G.C. & Wheeler, J. 1953c: 135 (l.). Combination in Plagiolepis: Emery, 1892a: 117; in Plagiolepis (Anoplolepis): Santschi, 1914b: 123; in Plagiolepis (Zealleyella): Arnold, 1922: 586; in Anoplolepis: Emery, 1925b: 17. Senior synonym of hendecarthrus: Mayr, 1865: 54; of berthoudi: Forel, 1879a: 91. See also: Prins, 1982: 218. Current subspecies: nominal plus detrita, hirsuta, pilipes.
Formicinae Lasiini Anoplolepis fallax TRUE TRUE FALSEfallax. Plagiolepis fallax Mayr, 1865: 54 (w.) SOUTH AFRICA. Forel, 1910f: 24 (q.). Combination in Plagiolepis (Zealleyella): Arnold, 1922: 589; in Anoplolepis: Emery, 1925b: 17. Subspecies of custodiens: Mayr, 1893: 5; Arnold, 1922: 589. Revived status as species: Emery, 1925b: 17.
Formicinae Lasiini Anoplolepis nuptialis TRUE TRUE FALSEnuptialis. Plagiolepis (Anoplolepis) nuptialis Santschi, 1917b: 289 (m.) SOUTH AFRICA. Prins, 1982: 222 (q.). Combination in Anoplolepis: Emery, 1925b: 18; in Anoplolepis (Zealleyella): Santschi, 1926a: 14.
Formicinae Lasiini Anoplolepis rufescens TRUE TRUE FALSErufescens. Plagiolepis (Anoplolepis) rufescens Santschi, 1917b: 289 (s.w.) SOUTH AFRICA. Combination in Anoplolepis: Emery, 1925b: 18; in Anoplolepis (Zealleyella): Santschi, 1926a: 14.
Formicinae Lasiini Anoplolepis steingroeveri parsonsi TRUE TRUE FALSEparsonsi. Anoplolepis (Zealleyella) braunsi st. parsonsi Santschi, 1937g: 83 (w.) SOUTH AFRICA. Currently subspecies of steingroeveri: Bolton, 1995b: 67.
Ponerinae Ponerini Asphinctopone differens TRUE TRUE FALSEdifferens. Asphinctopone differens Bolton & Fisher, 2008a: 59, figs. 1c,d; 2b,d (w.) CENTRAL AFRICAN REPUBLIC.
Dolichoderinae Tapinomini Axinidris lignicola TRUE TRUE FALSElignicola. Axinidris lignicola Snelling, R.R. 2007: 564, figs. 5, 15, 25 (w.) SOUTH AFRICA.
Dolichoderinae Tapinomini Axinidris mlalu TRUE TRUE FALSEmlalu. Axinidris mlalu Snelling, R.R. 2007: 566 (w.) CENTRAL AFRICAN REPUBLIC.
Ponerinae Ponerini Boloponera vicans TRUE TRUE FALSEvicans. Boloponera vicans Fisher, 2006: 113, figs. 1-4, 24 (w.) CENTRAL AFRICAN REPUBLIC.
Myrmicinae Stenammini Calyptomyrmex brunneus TRUE TRUE FALSEbrunneus. Calyptomyrmex brunneus Arnold, 1948: 221 (w.) SOUTH AFRICA. See also: Bolton, 1981a: 70.
Myrmicinae Stenammini Calyptomyrmex claviseta TRUE TRUE FALSEclaviseta. Dicroaspis claviseta Santschi, 1914e: 27 (w.) SOUTH AFRICA. Combination in Calyptomyrmex (Dicroaspis): Emery, 1915g: 15; in Calyptomyrmex: Bolton, 1981a: 71.
Formicinae Camponotini Camponotus angusticeps TRUE TRUE FALSEangusticeps. Camponotus angusticeps Emery, 1886: 355, pl. 17, fig. 1 (w.) SOUTH AFRICA. Arnold, 1924: 688 (s.). Combination in C. (Myrmosericus): Forel, 1914a: 268.
Formicinae Camponotini Camponotus arnoldinus TRUE TRUE FALSEarnoldinus. Camponotus (Myrmoturba) arnoldinus Forel, 1914d: 251 (s.w.) SOUTH AFRICA. Combination in C. (Tanaemyrmex): Emery, 1925b: 91. Subspecies of maculatus: Santschi, 1923e: 290. Revived status as species: Emery, 1925b: 91. See also: Arnold, 1922: 639.
Formicinae Camponotini Camponotus baynei TRUE TRUE FALSEbaynei. Camponotus maculatus r. baynei Arnold, 1922: 632 (s.w.) SOUTH AFRICA. Combination in C. (Tanaemyrmex): Emery, 1925b: 92. Raised to species: Emery, 1925b: 92.
Formicinae Camponotini Camponotus berthoudi TRUE TRUE FALSEberthoudi. Camponotus berthoudi Forel, 1879a: 108, pl. 1, fig. 6 (w.) SOUTH AFRICA. Arnold, 1924: 700. Combination in C. (Myrmamblys): Forel, 1914a: 271; in C. (Myrmopiromis): Emery, 1925b: 128.
Formicinae Camponotini Camponotus bertolonii TRUE TRUE FALSEbertolonii. Camponotus bertolonii Emery, 1895h: 51 (s.w.q.) SOUTH AFRICA. Mayr, 1901b: 28 (m.). Combination in C. (Myrmoturba): Forel, 1914a: 267; Arnold, 1922: 651; in C. (Myrmotemnus): Emery, 1920b: 258; in C. (Myrmamblys): Santschi, 1914b: 134; Wheeler, W.M. 1922a: 989; Emery, 1925b: 141; in C. (Paramyrmamblys): Santschi, 1926c: 604.
Formicinae Camponotini Camponotus bianconii TRUE TRUE FALSEbianconii. Camponotus bianconii Emery, 1895h: 52 (s.) SOUTH AFRICA. Combination in C. (Tanaemyrmex): Emery, 1925b: 92.
Formicinae Camponotini Camponotus bifossus TRUE TRUE FALSEbifossus. Camponotus (Colobopsis) bifossus Santschi, 1917b: 293 (s.) SOUTH AFRICA.
Formicinae Camponotini Camponotus brevisetosus TRUE TRUE FALSEbrevisetosus. Camponotus fulvopilosus subsp. brevisetosus Forel, 1910f: 25 (w.q.) SOUTH AFRICA. Combination in C. (Myrmopiromis): Wheeler, W.M. 1922a: 986. Raised to species: Robertson, 1990: 327. See also: Robertson & Zachariades, 1997: 3.
Formicinae Camponotini Camponotus brookei TRUE TRUE FALSEbrookei. Camponotus (Myrmamblys) brookei Forel, 1914d: 259 (s.w.) SOUTH AFRICA. Arnold, 1924: 706 (s.q.). Combination in C. (Paramyrmamblys): Santschi, 1926c: 604.
Formicinae Camponotini Camponotus buttikeri TRUE TRUE FALSEbuttikeri. Camponotus (Tanaemyrmex) buttikeri Arnold, 1958: 126, fig. 7 (s.w.) SOUTH AFRICA.
Formicinae Camponotini Camponotus caffer TRUE TRUE FALSEcaffer. Camponotus caffer Emery, 1895h: 47, pl. 2, figs. 25, 26 (s.w.) SOUTH AFRICA. Combination in C. (Myrmoturba): Forel, 1914a: 267; in C. (Dinomyrmex): Emery, 1920b: 256; in C. (Tanaemyrmex): Emery, 1925b: 84. Subspecies of maculatus: Forel, 1901h: 69. Revived status as species: Forel, 1914a: 267; Emery, 1920b: 256; Arnold, 1922: 650; Wheeler, W.M. 1922a: 963.
Formicinae Camponotini Camponotus cleobulus TRUE TRUE FALSEcleobulus. Camponotus (Myrmoturba) cleobulus Santschi, 1919b: 238 (s.w.) SOUTH AFRICA. Combination in C. (Tanaemyrmex): Emery, 1925b: 92.
Formicinae Camponotini Camponotus cognatocompressus TRUE TRUE FALSEcognatocompressus. Camponotus maculatus r. cognatocompressus Forel, 1904b: 379 (w.q.m.) "AFRICA" (no locality). [First available use of Camponotus rubripes r. cognatus var. cognatocompressus Forel, 1886f: 151; unavailable name.] Junior synonym of xerxes: Karavaiev, 1910b: 9. Revived from synonymy and raised to species: Radchenko, 1997d: 810.
Formicinae Camponotini Camponotus cosmicus TRUE TRUE FALSEcosmicus. Formica cosmica Smith, F. 1858b: 34 (s.w.) SOUTH AFRICA. Combination in Camponotus: Roger, 1863b: 2; in C. (Myrmosericus): Emery, 1925b: 106. Subspecies of rufoglaucus: Emery, 1895h: 49 (in key); Stitz, 1916: 401; Arnold, 1924: 693. Revived status as species: Santschi, 1935a: 280.
Formicinae Camponotini Camponotus cuneiscapus TRUE TRUE FALSEcuneiscapus. Camponotus cuneiscapus Forel, 1910f: 29, pl. 1, fig. 2 (s.w.) SOUTH AFRICA. Arnold, 1924: 679 (q.). Combination in C. (Myrmopsamma): Forel, 1914a: 270; Wheeler, W.M. 1922a: 988; Arnold, 1924: 679; in C. (Camponotus): Santschi, 1923e: 292; in C. (Myrmamblys): Emery, 1925b: 142; in C. (Myrmespera): Santschi, 1926b: 248.
Formicinae Camponotini Camponotus detritus TRUE TRUE FALSEdetritus. Camponotus detritus Emery, 1886: 357 (w.) SOUTH AFRICA. Forel, 1910f: 26 (s.). Combination in C. (Myrmepomis): Forel, 1914a: 273; in C. (Myrmopiromis): Wheeler, W.M. 1922a: 986. Subspecies of fulvopilosus: Forel, 1910f: 26; Wheeler, W.M. 1922a: 986; Stitz, 1923: 166; Arnold, 1924: 739. Revived status as species: Forel, 1914a: 273; Emery, 1920b: 258; Emery, 1925b: 128; Robertson, 1990: 327. See also: Robertson & Zachariades, 1997: 5.
Formicinae Camponotini Camponotus dicksoni TRUE TRUE FALSEdicksoni. Camponotus (Tanaemyrmex) dicksoni Arnold, 1948: 227, fig. 13 (s.w.) SOUTH AFRICA.
Formicinae Camponotini Camponotus emarginatus TRUE TRUE FALSEemarginatus. Camponotus emarginatus Emery, 1886: 358, pl. 17, fig. 2 (s.w.q.m.) SOUTH AFRICA. Combination in C. (Myrmophyma): Forel, 1914a: 269; Arnold, 1922: 663; in C. (Myrmotemnus): Emery, 1920b: 258; in C. (Myrmamblys): Wheeler, W.M. 1922a: 989; in C. (Myrmespera): Santschi, 1926c: 604.
Formicinae Camponotini Camponotus eugeniae TRUE TRUE FALSE1 subspecies
eugeniae. Camponotus micans r. eugeniae Forel, 1879a: 78 (s.w.) SOUTH AFRICA. Forel, 1886f: 174 (q.); Arnold, 1924: 687 (m.). Combination in C. (Myrmosericus): Forel, 1912i: 91. Subspecies of rufoglaucus: Mayr, 1893: 195. Raised to species: Forel, 1886f: 174; Emery, 1895h: 50. Current subspecies: nominal plus amplior.
Formicinae Camponotini Camponotus fulvopilosus TRUE TRUE FALSEfulvopilosus. Formica fulvopilosa De Geer, 1778: 613, pl. 45, figs. 13, 14 (w.) SOUTH AFRICA. Forel, 1879a: 108 (s.); Forel, 1910f: 25 (m.); Arnold, 1924: 737 (q.). Combination in Camponotus: Mayr, 1862: 668; in C. (Myrmepomis): Forel, 1912i: 92; in C. (Myrmopiromis): Wheeler, W.M. 1922a: 985. Senior synonym of pilosa, rufiventris: Lepeletier, 1835: 213; Smith, F. 1858b: 32; Wheeler, W.M. 1922a: 985; of detritoides, flavopilosus: Robertson, 1990: 327. See also: Robertson & Zachariades, 1997: 8.
Formicinae Camponotini Camponotus guanchus TRUE TRUE FALSEguanchus. Camponotus sicheli var. guancha Santschi, 1908: 534 (w.q.) "North Africa". Cagniant & Espadaler, 1993: 427 (m.). Combination in C. (Myrmentoma): Emery, 1925b: 121. Subspecies of lateralis: Emery, 1925a: 64; Emery, 1925b: 121. Raised to species: Cagniant & Espadaler, 1993: 426.
Formicinae Camponotini Camponotus havilandi TRUE TRUE FALSEhavilandi. Camponotus (Myrmophyma) havilandi Arnold, 1922: 666 (s.w.) SOUTH AFRICA. Combination in C. (Myrmamblys): Emery, 1925b: 140; in C. (Myrmespera): Santschi, 1926c: 604.
Formicinae Camponotini Camponotus importunus TRUE TRUE FALSEimportunus. Camponotus maculatus subsp. importunus Forel, 1911e: 288 (s.w.q.m.) SOUTH AFRICA. [First available use of Camponotus maculatus subsp. sexpunctatus var. importuna Forel, 1910f: 27; unavailable name.] Combination in C. (Myrmoturba): Emery, 1920c: 5; in C. (Tanaemyrmex): Emery, 1925b: 87. Subspecies of maculatus: Emery, 1920c: 5; Arnold, 1922: 635. Raised to species: Baroni Urbani, 1972: 131.
Formicinae Camponotini Camponotus klugii TRUE TRUE FALSEklugii. Camponotus bedoti var. klugii Emery, 1895h: 51 (w.) SOUTH AFRICA. Combination in C. (Myrmamblys): Wheeler, W.M. 1922a: 989. Raised to species: Emery, 1925b: 141.
Formicinae Camponotini Camponotus knysnae TRUE TRUE FALSEknysnae. Camponotus (Myrmoturba) maculatus r. knysnae Arnold, 1922: 642 (s.w.q.) SOUTH AFRICA. Combination in C. (Tanaemyrmex): Emery, 1925b: 100. Raised to species: Emery, 1925b: 100 (misspelled as kaysnae).
Formicinae Camponotini Camponotus liogaster TRUE TRUE FALSEliogaster. Camponotus (Orthonotomyrmex) liogaster Santschi, 1932a: 390 (w.) SOUTH AFRICA.
Formicinae Camponotini Camponotus maculatus TRUE TRUE FALSE5 subspecies
maculatus. Formica maculata Fabricius, 1782: 491 (w.) WEST AFRICA. Mayr, 1862: 654 (q.m.); Forel, 1891b: 29 (s.w.q.m.); Donisthorpe, 1915a: 221 (redescription of holotype). Combination in Camponotus: Mayr, 1862: 654; in C. (Myrmoturba): Forel, 1912i: 91; in C. (Tanaemyrmex): Emery, 1925b: 86. Subspecies of sylvaticus: Forel, 1879a: 64; of rubripes: Forel, 1886f: 143. Revived status as species: Emery, 1891b: 17. Senior synonym of atramentarius, ballioni, boera, cavallus, cluisoides, cognata, conakryensis, erythraea, flavifemur, flavominor, hannae, hieroglyphicus, intonsus, lacteipennis, liengmei, liocnemis, lividior, lohieri, manzer, mathildae, melanocnemis (and its junior synonym schultzei), nubis, proletaria, radamoides, sarmentus, schereri, semispicatus, sudanicus, thomensis, tuckeri, zumpti and material of the unavailable names calceatus, cataractae, citinus, cluis, contaminatus, diffusus, hansingi, importunoides, madecassa, pessimus referred here: Baroni Urbani, 1972: 125-127. Current subspecies: nominal plus foveolatus (unresolved junior homonym), miserabilis (unresolved junior homonym), obfuscatus, subnudus, sylvaticomaculatus, ugandensis.
Formicinae Camponotini Camponotus mayri TRUE TRUE FALSE1 subspecies
mayri. Camponotus mayri Forel, 1879a: 92, pl. 1, fig. 1 (w.) SOUTH AFRICA. Combination in C. (Myrmentoma): Forel, 1913a: 146; in C. (Orthonotomyrmex): Forel, 1913b: 347; Emery, 1925b: 125. Current subspecies: nominal plus chimporensis.
Formicinae Camponotini Camponotus mystaceus TRUE TRUE FALSE2 subspecies
mystaceus. Camponotus mystaceus Emery, 1886: 356, pl. 17, fig. 3 (s.w.m.) SOUTH AFRICA. Arnold, 1924: 675 (q.). Combination in C. (Myrmopsamma): Forel, 1914a: 261. Current subspecies: nominal plus exsanguis, kamae.
Formicinae Camponotini Camponotus nasutus TRUE TRUE FALSE5 subspecies
nasutus. Camponotus nasutus Emery, 1895h: 53, pl. 2, fig. 27 (w.) SOUTH AFRICA. Combination in C. (Myrmophyma): Forel, 1914a: 269; in C. (Myrmotemnus): Emery, 1920b: 258; in C. (Myrmamblys): Santschi, 1921f: 311; in C. (Myrmespera): Santschi, 1926c: 604. Current subspecies: nominal plus fenestralis, pretiosus, quadridentatus, quinquedentatus, subnasutus.
Formicinae Camponotini Camponotus natalensis TRUE TRUE FALSE4 subspecies
natalensis. Formica natalensis Smith, F. 1858b: 33 (s.w.q.m.) SOUTH AFRICA. Combination in Camponotus: Roger, 1863b: 2; in C. (Myrmoturba): Forel, 1914a: 267; in C. (Tanaemyrmex): Emery, 1925b: 103. See also: Arnold, 1922: 652. Current subspecies: nominal plus corvus, diabolus, fulvipes, politiceps.
Formicinae Camponotini Camponotus niveosetosus TRUE TRUE FALSE2 subspecies
niveosetosus. Camponotus niveosetosus Mayr, 1862: 665 (w.) SOUTH AFRICA. Mayr, 1866b: 885 (q.); Forel, 1891b: 41 (s.); Arnold, 1922: 659 (m.). Combination in C. (Myrmoturba): Forel, 1914a: 267; in C. (Myrmepomis): Emery, 1920b: 259; in C. (Myrmopiromis): Wheeler, W.M. 1922a: 986. Current subspecies: nominal plus irredux, madagascarensis.
Formicinae Camponotini Camponotus orites TRUE TRUE FALSEorites. Camponotus (Myrmamblys) orites Santschi, 1919a: 346, fig. 4 (w.) SOUTH AFRICA. Arnold, 1926: 290 (s.). Combination in C. (Paramyrmamblys): Santschi, 1926c: 604. Senior synonym of orinodromus: Arnold, 1926: 291.
Formicinae Camponotini Camponotus ostiarius TRUE TRUE FALSEostiarius. Camponotus (Myrmamblys) ostiarius Forel, 1914d: 258 (s.w.) SOUTH AFRICA. Arnold, 1924: 702 (q.m.). Combination in C. (Paramyrmamblys): Santschi, 1926c: 604.
Formicinae Camponotini Camponotus petersii TRUE TRUE FALSE1 subspecies
petersii. Camponotus petersii Emery, 1895h: 50, pl. 2, fig. 28 (w.) SOUTH AFRICA. Forel, 1914d: 254 (s.); Arnold, 1924: 684 (q.m.). Combination in C. (Myrmosericus): Forel, 1912i: 91. Current subspecies: nominal plus janus.
Formicinae Camponotini Camponotus postoculatus TRUE TRUE FALSEpostoculatus. Camponotus (Myrmotrema) postoculatus Forel, 1914d: 254 (s.w.) SOUTH AFRICA.
Formicinae Camponotini Camponotus reevei TRUE TRUE FALSEreevei. Camponotus (Myrmophyma) reevei Arnold, 1922: 671, fig. 58 (s.w.) SOUTH AFRICA. Combination in C. (Myrmamblys): Emery, 1925b: 141; in C. (Myrmespera): Santschi, 1926c: 604.
Formicinae Camponotini Camponotus scalaris TRUE TRUE FALSEscalaris. Camponotus scalaris Forel, 1901d: 308 (s.w.q.) SOUTH AFRICA. Combination in C. (Myrmosphincta): Forel, 1912i: 92; in C. (Orthonotomyrmex): Emery, 1920b: 258; in C. (Myrmisolepis): Santschi, 1921f: 310; in C. (Myrmopiromis): Emery, 1925b: 129.
Formicinae Camponotini Camponotus sellidorsatus TRUE TRUE FALSEsellidorsatus. Camponotus (Mayria) sellidorsatus Prins, 1973: 21, figs. 25, 26, 34 (w.) SOUTH AFRICA.
Formicinae Camponotini Camponotus storeatus TRUE TRUE FALSEstoreatus. Camponotus fulvopilosus subsp. storeatus Forel, 1910f: 26 (w.) SOUTH AFRICA. Arnold, 1924: 739 (s.q.). Combination in C. (Myrmopiromis): Wheeler, W.M. 1922a: 986. Raised to species: Robertson, 1990: 327. See also: Robertson & Zachariades, 1997: 15.
Formicinae Camponotini Camponotus traegaordhi TRUE TRUE FALSE1 subspecies
traegaordhi. Camponotus (Myrmoturba) maculatus st. traegaordhi Santschi, 1914e: 38 (s.w.q.m.) SOUTH AFRICA. Combination in C. (Tanaemyrmex): Emery, 1925b: 85. Subspecies of guttatus: Emery, 1920c: 6; of maculatus: Arnold, 1922: 627. Raised to species: Santschi, 1926b: 252. Current subspecies: nominal plus fumeus.
Formicinae Camponotini Camponotus transvaalensis TRUE TRUE FALSE2 subspecies
transvaalensis. Camponotus (Myrmespera) transvaalensis Arnold, 1948: 228, fig. 14 (s.w.) SOUTH AFRICA. Current subspecies: nominal plus prinsi, griqua.
Formicinae Camponotini Camponotus trifasciatus TRUE TRUE FALSEtrifasciatus. Camponotus (Myrmespera) debellator var. trifasciatus Santschi, 1926b: 248 (q.) SOUTH AFRICA. Raised to species: Santschi, 1926c: 604.
Formicinae Camponotini Camponotus valdeziae TRUE TRUE FALSEvaldeziae. Camponotus valdeziae Forel, 1879a: 90 (w.q.) SOUTH AFRICA. Arnold, 1924: 696 (s.). Combination in C. (Myrmosericus): Forel, 1914a: 268.
Formicinae Camponotini Camponotus vestitus TRUE TRUE FALSE8 subspecies
vestitus. Formica vestita Smith, F. 1858b: 32 (q.) SOUTH AFRICA. Arnold, 1924: 694 (s.w.). Combination in Camponotus: Roger, 1863b: 2; in C. (Myrmosericus): Emery, 1925b: 106. Subspecies of rufoglaucus: Emery, 1894i: 47; Emery, 1895h: 48; Forel, 1907e: 16; Santschi, 1914d: 381; Arnold, 1924: 694. Revived status as species: Santschi, 1926b: 258; Santschi, 1930a: 74; Bernard, 1953b: 262. Current subspecies: nominal plus anthracinus, bombycinus, comptus, intuens, lujai, pectitus, perpectitus, strophiatus.
Formicinae Camponotini Camponotus victoriae TRUE TRUE FALSEvictoriae. Camponotus (Myrmopiromis) victoriae Arnold, 1959: 338, fig. 24 (s.w.) SOUTH AFRICA.
Formicinae Camponotini Camponotus rufoglaucus zulu TRUE TRUE FALSEzulu. Camponotus rufoglaucus subsp. zulu Emery, 1895h: 50 (s.w.) SOUTH AFRICA. Combination in C. (Myrmosericus): Emery, 1925b: 107.
Formicinae Camponotini Camponotus olivieri tenuipilis TRUE TRUE FALSEtenuipilis. Camponotus (Myrmotrema) olivieri var. tenuipilis Santschi, 1915c: 270 (in key) (w.) SOUTH AFRICA. [First available use of Camponotus foraminosus subsp. olivieri var. tenuipilis Emery, 1895h: 54; unavailable name.]
Formicinae Camponotini Camponotus nasutus subnasutus TRUE TRUE FALSEsubnasutus. Camponotus (Myrmophyma) nasutus var. subnasutus Arnold, 1922: 670 (s.w.) SOUTH AFRICA. Combination in C. (Myrmamblys): Emery, 1925b: 141.
Formicinae Camponotini Camponotus vestitus strophiatus TRUE TRUE FALSEstrophiatus. Camponotus (Myrmosericus) vestitus var. strophiatus Santschi, 1926b: 259 (w.) SOUTH AFRICA.
Formicinae Camponotini Camponotus werthi skaifei TRUE TRUE FALSEskaifei. Camponotus (Tanaemyrmex) werthi var. skaifei Arnold, 1959: 336, fig. 22 (s.w.) SOUTH AFRICA.
Formicinae Camponotini Camponotus transvaalensis griqua TRUE TRUE FALSEgriqua. Camponotus (Myrmespera) transvaalensis subsp. griqua Arnold, 1952a: 465 (s.w.q.m.) SOUTH AFRICA.
Formicinae Camponotini Camponotus natalensis fulvipes TRUE TRUE FALSEfulvipes. Camponotus (Myrmoturba) fulvipes Forel, 1914a: 267 (w.) SOUTH AFRICA. [First available use of Camponotus natalensis subsp. corvus var. fulvipes Emery, 1895h: 51; unavailable name.] Santschi, 1914e: 38 (q.m.). Currently subspecies of natalensis: Santschi, 1914e: 38; Arnold, 1922: 654; Emery, 1925b: 103.
Formicinae Camponotini Camponotus cubangensis dofleini TRUE TRUE FALSEdofleini. Camponotus dofleini Forel, 1911e: 291 (w.) SOUTH AFRICA. Combination in C. (Myrmosphincta): Forel, 1914a: 273; in C. (Orthonotomyrmex): Emery, 1920b: 257. Arnold, 1924: 714 (s.). Senior synonym of ledieui: Emery, 1920c: 29. Currently subspecies of cubangensis: Santschi, 1930b: 81 (in text).
Formicinae Camponotini Camponotus natalensis diabolus TRUE TRUE FALSEdiabolus. Camponotus natalensis r. diabolus Forel, 1879a: 81 (s.w.) SOUTH AFRICA. Combination in C. (Myrmoturba): Forel, 1914a: 267; in C. (Tanaemyrmex): Emery, 1925b: 103. Raised to species: Dalla Torre, 1893: 228; Forel, 1914a: 267. Currently subspecies of natalensis: Emery, 1896d: 373; Wheeler, W.M. 1922a: 961; Emery, 1925b: 103. See also: Arnold, 1922: 646.
Formicinae Camponotini Camponotus arminius bicontractus TRUE TRUE FALSEbicontractus. Camponotus (Orthonotomyrmex) arminius var. bicontractus Wheeler, W.M. 1922a: 971 (w.) SOUTH AFRICA. [First available use of Camponotus (Orthonotomyrmex) barbarossa r. arminius var. bicontracta Forel, 1914d: 256; unavailable name.] Arnold, 1924: 721 (s.). Combination in C. (Myrmamblys): Emery, 1925b: 142; in C. (Myrmopelta): Santschi, 1926c: 603 (misspelled as biconstrictus).
Formicinae Camponotini Camponotus thraso agricola TRUE TRUE FALSEagricola. Camponotus maculatus subsp. agricola Forel, 1910e: 454 (s.w.) SOUTH AFRICA. Combination in C. (Tanaemyrmex): Emery, 1925b: 92. Subspecies of maculatus: Arnold, 1922: 647. Currently subspecies of thraso: Emery, 1920c: 6; Emery, 1925b: 92.
Formicinae Camponotini Camponotus auropubens absalon TRUE TRUE FALSEabsalon. Camponotus (Myrmotrema) auropubens var. absalon Santschi, 1915c: 274 (s.w.q.) SOUTH AFRICA.
Formicinae Camponotini Camponotus natalensis corvus TRUE TRUE FALSEcorvus. Camponotus natalensis r. corvus Forel, 1879a: 79 (s.w.q.m.) SOUTH AFRICA. Combination in C. (Tanaemyrmex): Emery, 1925b: 103. Raised to species: Dalla Torre, 1893: 227. Currently subspecies of natalensis: Emery, 1895h: 51; Arnold, 1922: 654; Wheeler, W.M. 1922a: 960.
Formicinae Camponotini Camponotus olivieri infelix TRUE TRUE FALSEinfelix. Camponotus (Myrmotrema) olivieri st. infelix Santschi, 1915c: 270 (s.w.) SOUTH AFRICA. [First available use of Camponotus (Myrmotrema) foraminosus r. lemma var. infelix Forel, 1914d: 254; unavailable name.]
Formicinae Camponotini Camponotus niveosetosus irredux TRUE TRUE FALSEirredux. Camponotus irredux Forel, 1910f: 30 (s.w.) SOUTH AFRICA. Combination in C. (Myrmopiromis): Wheeler, W.M. 1922a: 987. Currently subspecies of niveosetosus: Forel, 1910e: 451; Arnold, 1922: 659.
Formicinae Camponotini Camponotus petersii janus TRUE TRUE FALSEjanus. Camponotus petersi var. janus Forel, 1911e: 228 (w.) SOUTH AFRICA. Combination in C. (Myrmosericus): Emery, 1925b: 105.
Formicinae Camponotini Camponotus hova pictiventris TRUE TRUE FALSEpictiventris. Camponotus maculatus subsp. pictiventris Mayr, 1901b: 28 (s.w.) SOUTH AFRICA. Combination in C. (Tanaemyrmex): Emery, 1925b: 86. Subspecies of maculatus: Arnold, 1922: 629; Wheeler, W.M. 1922a: 958. Currently subspecies of hova: Emery, 1920c: 6; Emery, 1925b: 86.
Formicinae Camponotini Camponotus natalensis politiceps TRUE TRUE FALSEpoliticeps. Camponotus (Myrmoturba) natalensis var. politiceps Santschi, 1914e: 38 (w.m.) SOUTH AFRICA. Combination in C. (Tanaemyrmex): Emery, 1925b: 103.
Formicinae Camponotini Camponotus nasutus quinquedentatus TRUE TRUE FALSEquinquedentatus. Camponotus nasutus var. quinquedentata Forel, 1910e: 451 (w.) SOUTH AFRICA. Combination in C. (Myrmamblys): Wheeler, W.M. 1922a: 990.
Myrmicinae Solenopsidini Carebara africana TRUE TRUE FALSEafricana. Aeromyrma africana Forel, 1910f: 15 (s.) BOTSWANA. Combination in Oligomyrmex (Aeromyrma): Emery, 1924d: 216; in Carebara: Fernández, 2004a: 235.
Myrmicinae Solenopsidini Carebara arnoldiella TRUE TRUE FALSEarnoldiella. Oligomyrmex (Aeromyrma) arnoldiella Santschi, 1919a: 340, fig. 3 (s.w.) SOUTH AFRICA. Combination in Aeromyrma: Wheeler, W.M. 1922a: 882; in Oligomyrmex: Ettershank, 1966: 123; in Carebara: Fernández, 2004a: 235.
Myrmicinae Solenopsidini Carebara donisthorpei TRUE TRUE FALSEdonisthorpei. Oligomyrmex donisthorpei Weber, 1950a: 7, fig. (s.w.) CENTRAL AFRICAN REPUBLIC. Combination in Carebara: Fernández, 2004a: 235.
Myrmicinae Solenopsidini Carebara junodi TRUE TRUE FALSEjunodi. Carebara junodi Forel, 1904a: 154 (q.) SOUTH AFRICA. Forel, 1909b: 53 (m.); Forel, 1913b: 336 (w.). Subspecies of vidua: Forel, 1909b: 53; Forel, 1913b: 336; Arnold, 1916: 252. Revived status as species: Santschi, 1914d: 364 (in key); Wheeler, W.M. 1922a: 883; Emery, 1924d: 220; Santschi, 1935a: 263.
Myrmicinae Solenopsidini Carebara perpusilla TRUE TRUE FALSE3 subspecies
perpusilla. Pheidologeton perpusillum Emery, 1895h: 26, pl. 2, figs. 8-11 (s.w.) SOUTH AFRICA. Combination in P. (Aneleus): Emery, 1900c: 327; in Aneleus: Santschi, 1914b: 77; in Oligomyrmex: Ettershank, 1966: 124; in Carebara: Fernández, 2004a: 235. Current subspecies: nominal plus arnoldiana, concedens, spinosa. See also: Arnold, 1916: 254.
Myrmicinae Solenopsidini Carebara robertsoni TRUE TRUE FALSErobertsoni. Paedalgus robertsoni Bolton & Belshaw, 1993: 188, fig. 1 (w.q.) SOUTH AFRICA. Combination in Carebara: Fernández, 2004a: 235.
Myrmicinae Solenopsidini Carebara semilaevis TRUE TRUE FALSEsemilaevis. Solenopsis semilaevis Mayr, 1901b: 15 (w.) SOUTH AFRICA. Combination in Oligomyrmex (Aeromyrma): Emery, 1915c: 59 (in text); in Aeromyrma: Wheeler, W.M. 1922a: 882; in Oligomyrmex: Ettershank, 1966: 124; in Carebara: Fernández, 2004a: 235. Senior synonym of hewitti: Arnold, 1926: 236.
Myrmicinae Solenopsidini Carebara traegaordhi TRUE TRUE FALSEtraegaordhi. Aeromyrma traegaordhi Santschi, 1914e: 20, fig. 2 (w.) SOUTH AFRICA. Combination in Oligomyrmex (Aeromyrma): Santschi, 1919a: 338; in Carebara: Fernández, 2004a: 235. Subspecies of semilaevis: Santschi, 1917b: 285. Revived status as species: Santschi, 1919a: 338; Emery, 1924d: 216; Arnold, 1926: 233.
Myrmicinae Solenopsidini Carebara vidua TRUE TRUE FALSE1 subspecies
vidua. Carebara vidua Smith, F. 1858b: 179 (m.) SOUTH AFRICA. Mayr, 1866b: 904 (q.); Forel, 1901g: 392 (w.). Senior synonym of colossus, dux: Mayr, 1866b: 904; Santschi, 1913c: 307; of abdominalis: Forel, 1913b: 337; Arnold, 1916: 252; Wheeler, W.M. 1922a: 884. See also: Arnold, 1916: 249; Robertson & Villet, 1989: 121. Current subspecies: nominal plus fur.
Formicinae Formicini Cataglyphis albicans TRUE TRUE FALSE6 subspecies
albicans. Formica albicans Roger, 1859: 235 (w.) NORTH AFRICA. André, 1882b: 174 (m.); Forel, 1894d: 11 (q.); Santschi, 1921d: 76 (gynandromorph). Combination in Cataglyphis: Mayr, 1862: 701; Santschi, 1912c: 149; Menozzi, 1922b: 332; Santschi, 1929b: 37; in Myrmecocystus: Emery, 1891b: 17; in Myrmecocystus (Cataglyphis): Karavaiev, 1924: 305. Subspecies of viatica: Emery & Forel, 1879: 449. Revived status as species: André, 1882b: 168; Emery, 1891b: 17; Emery, 1906d: 51. Senior synonym of armena: Radchenko, 1997c: 425. Current subspecies: nominal plus cubica, fezzanensis, franchettii, kairuana, opaca, targuia.
Formicinae Formicini Cataglyphis bicolor TRUE TRUE FALSE7 subspecies
bicolor. Formica bicolor Fabricius, 1793: 351 (m.) NORTH AFRICA. Wheeler, G.C. & Wheeler, J. 1980: 538 (l.). Combination in Cataglyphis: Roger, 1863b: 12; Mayr, 1863: 402; Emery, 1912f: 98; Emery, 1915g: 22; in Myrmecocystus: Emery & Forel, 1879: 449. Junior synonym of viatica: Smith, F. 1861a: 32; Nasonov, 1889: 14. Revived from synonymy: Emery & Forel, 1879: 449; Emery, 1906d: 58; Forel, 1907e: 15. Subspecies of viatica: Forel, 1907e: 15. Revived status as species: Karavaiev, 1912b: 590; Emery, 1925b: 264; Santschi, 1929b: 41. Senior synonym of fairmairei: Santschi, 1929b: 41; of megalocola: Mayr, 1855: 382; Santschi, 1929b: 41; of rotundinodis: Santschi, 1929b: 41. Biogeography of bicolor group: Wehner, R., Wehner, S. & Agosti, 1994: 163. Current subspecies: nominal plus adusta, basalis, congolensis, pubens, rufiventris, seticornis, sudanica.
Myrmicinae Cataulacini Cataulacus fricatidorsus TRUE TRUE FALSEfricatidorsus. Cataulacus otii st. fricatidorsus Santschi, 1914e: 26 (w.) SOUTH AFRICA. Raised to species: Bolton, 1974a: 42.
Myrmicinae Cataulacini Cataulacus guineensis TRUE TRUE FALSEguineensis. Cataulacus guineensis Smith, F. 1853: 225, pl. 20, fig. 5 (w.) "TROPICAL WEST AFRICA". Santschi, 1910c: 390 (q.); Wheeler, W.M. 1922a: 199 (m.). Senior synonym of sulcatus: Forel, 1910e: 421; of alenensis, fernandensis, parallelus, sulcatus, sulcinodis: Bolton, 1974a: 55.
Myrmicinae Cataulacini Cataulacus intrudens TRUE TRUE FALSEintrudens. Meranoplus intrudens Smith, F. 1876d: 609, pl. 11, fig. 7 (w.q.m.) SOUTH AFRICA. Combination in Cataulacus: Mayr, 1886c: 364. Senior synonym of batonga, baumi, densipunctatus, foveosquamosus, gazanus, hararicus, intermedius, johannae, pseudotrema, rugosus, subrugosus, umbilicatus and material of the unavailable names krugeri, tangana referred here: Bolton, 1974a: 43; of bulawayensis: Bolton, 1995b: 138.
Myrmicinae Cataulacini Cataulacus micans TRUE TRUE FALSEmicans. Cataulacus rugosus subsp. micans Mayr, 1901b: 27 (w.q.m.) SOUTH AFRICA. Raised to species: Forel, 1914d: 219. Senior synonym of tristiculus: Bolton, 1974a: 46.
Myrmicinae Cataulacini Cataulacus traegaordhi TRUE TRUE FALSEtraegaordhi. Cataulacus traegaordhi Santschi, 1914e: 24, fig. 3 (w.q.m.) SOUTH AFRICA. Junior synonym of pygmaeus: Bolton, 1974a: 48. Revived from synonymy and senior synonym of marleyi, suddensis, ugandensis: Bolton, 1982: 359.
Ponerinae Ponerini Centromyrmex ereptor TRUE TRUE FALSEereptor. Centromyrmex ereptor Bolton & Fisher, 2008c: 18, figs. 13, 14 (w.) CENTRAL AFRICAN REPUBLIC.
Cerapachyinae Cerapachyini Cerapachys arnoldi TRUE TRUE FALSEarnoldi. Cerapachys arnoldi Forel, 1914d: 211 (w.) SOUTH AFRICA. Senior synonym of hewitti: Brown, 1975: 22. See also: Arnold, 1926: 191.
Cerapachyinae Cerapachyini Cerapachys braunsi TRUE TRUE FALSEbraunsi. Phyracaces braunsi Emery, 1902c: 27 (w.) SOUTH AFRICA. Combination in Cerapachys: Brown, 1975: 22.
Cerapachyinae Cerapachyini Cerapachys faurei TRUE TRUE FALSEfaurei. Cerapachys faurei Arnold, 1949: 262, fig. 2 (w.) SOUTH AFRICA.
Cerapachyinae Cerapachyini Cerapachys natalensis TRUE TRUE FALSEnatalensis. Cerapachys cribrinodis var. natalensis Forel, 1901f: 335 (w.) SOUTH AFRICA. [Also described as new by Forel, 1910e: 423.] Raised to species: Brown, 1975: 23.
Cerapachyinae Cerapachyini Cerapachys peringueyi TRUE TRUE FALSEperingueyi. Parasyscia peringueyi Emery, 1886: 360 (w.) SOUTH AFRICA. Emery, 1895j: 757 (q.). Combination in Cerapachys: Emery, 1902c: 24. Senior synonym of latiuscula: Brown, 1975: 23. See also: Arnold, 1915: 13.
Cerapachyinae Cerapachyini Cerapachys validus TRUE TRUE FALSEvalidus. Cerapachys validus Arnold, 1960a: 79, fig. 1 (w.) SOUTH AFRICA.
Cerapachyinae Cerapachyini Cerapachys wroughtoni TRUE TRUE FALSEwroughtoni. Cerapachys wroughtoni Forel, 1910e: 422 (w.) SOUTH AFRICA. Senior synonym of rhodesiana, roberti: Brown, 1975: 24.
Myrmicinae Crematogastrini Crematogaster africana TRUE TRUE FALSE10 subspecies
africana. Crematogaster africana Mayr, 1895: 142 (w.) CAMEROUN. Santschi, 1937g: 78 (q.). Combination in C. (Atopogyne): Santschi, 1916b: 501. Subspecies of buchneri: Forel, 1913a: 125. Revived status as species: Santschi, 1918d: 183. Current subspecies: nominal plus alligatrix, biemarginata, camena, polymorphica, schumanni, stanleyi, stolonis, thoracica, tibialis, variegata.
Myrmicinae Crematogastrini Crematogaster arnoldi TRUE TRUE FALSE1 subspecies
arnoldi. Crematogaster arnoldi Forel, 1914d: 238 (w.) SOUTH AFRICA. Combination in C. (Acrocoelia): Emery, 1922e: 145; in C. (Crematogaster): Bolton, 1995b: 166. Subspecies of godfreyi: Arnold, 1920a: 504. Revived status as species: Santschi, 1926b: 214. Current subspecies: nominal plus loveridgei.
Myrmicinae Crematogastrini Crematogaster capensis TRUE TRUE FALSE2 subspecies
capensis. Crematogaster capensis Mayr, 1862: 764 (w.q.) SOUTH AFRICA. Combination in C. (Crematogaster): Wheeler, W.M. 1922a: 830; in C. (Acrocoelia): Emery, 1922e: 145; in C. (Crematogaster): Bolton, 1995b: 166. Current subspecies: nominal plus calens, tropicorum.
Myrmicinae Crematogastrini Crematogaster castanea TRUE TRUE FALSE19 subspecies
castanea. Crematogaster castaneus Smith, F. 1858b: 136, pl. 9, fig. 2 (w.q.) SOUTH AFRICA (not Albania). Combination in C. (Crematogaster): Wheeler, W.M. 1922a: 830; in C. (Acrocoelia): Emery, 1922e: 145; in C. (Crematogaster): Bolton, 1995b: 166. Subspecies of tricolor: Forel, 1911f: 279; Santschi, 1913a: 412 (in key). Revived status as species: Emery, 1915g: 12; Arnold, 1920a: 488; Santschi, 1920d: 378. Senior synonym of decolor: Forel, 1907g: 80. Current subspecies: nominal plus adusta, analis, aquila, arborea, bruta, busschodtsi, durbanensis, ferruginea, hararica, insidiosa, inversa, mediorufa, museisapientiae, rufimembrum, rufonigra, simia, tricolor, ulugurensis, yambatensis.
Myrmicinae Crematogastrini Crematogaster cicatriculosa TRUE TRUE FALSEcicatriculosa. Crematogaster cicatriculosa Roger, 1863a: 207 (w.) SOUTH AFRICA. Mayr, 1866b: 901 (q.). Combination in C. (Crematogaster): Wheeler, W.M. 1922a: 836; in C. (Acrocoelia): Emery, 1922e: 146; in C. (Crematogaster): Bolton, 1995b: 166.
Myrmicinae Crematogastrini Crematogaster constructor TRUE TRUE FALSE1 subspecies
constructor. Crematogaster constructor Emery, 1895h: 29, pl. 2, figs. 12, 13 (w.) SOUTH AFRICA. [Also described as new by Emery, in Mayr, 1895: 135 (footnote).] Combination in C. (Crematogaster): Wheeler, W.M. 1922a: 836; in C. (Acrocoelia): Emery, 1922e: 146; in C. (Crematogaster): Bolton, 1995b: 166. Current subspecies: nominal plus kirbyella.
Myrmicinae Crematogastrini Crematogaster desperans TRUE TRUE FALSEdesperans. Crematogaster gabonensis var. desperans Forel, 1914d: 233 (w.) SOUTH AFRICA. Combination in C. (Atopogyne): Wheeler, W.M. 1922a: 854; in C. (Acrocoelia): Emery, 1922e: 146; in C. (Crematogaster): Bolton, 1995b: 166. Subspecies of bulawayensis: Arnold, 1920a: 523; Wheeler, W.M. 1922a: 854. Raised to species: Santschi, 1918d: 185; Emery, 1922e: 146.
Myrmicinae Crematogastrini Crematogaster hottentota TRUE TRUE FALSE1 subspecies
hottentota. Crematogaster kneri subsp. hottentota Emery, 1899e: 482 (w.) SOUTH AFRICA. Combination in C. (Sphaerocrema): Wheeler, W.M. 1922a: 847. Raised to species: Santschi, 1925h: 161 (footnote); Santschi, 1926b: 224. Current subspecies: nominal plus bassuto.
Myrmicinae Crematogastrini Crematogaster laurenti TRUE TRUE FALSE1 subspecies
laurenti. Crematogaster africana subsp. laurenti Forel, 1909b: 69 (w.) DEMOCRATIC REPUBLIC OF CONGO. Combination in C. (Atopogyne): Wheeler, W.M. 1922a: 852. Subspecies of buchneri: Forel, 1913a: 125; Forel, 1913c: 352; Forel, 1916: 410; of africana: Wheeler, W.M. 1922a: 852. Raised to species: Emery, 1922e: 155; Santschi, 1935a: 262. Current subspecies: nominal plus zeta.
Myrmicinae Crematogastrini Crematogaster magitae TRUE TRUE FALSEmagitae. Crematogaster (Oxygyne) magitae Forel, 1910f: 9 (q.) WEST AFRICA. Combination in C. (Nematocrema): Emery, 1922e: 157.
Myrmicinae Crematogastrini Crematogaster melanogaster TRUE TRUE FALSE1 subspecies
melanogaster. Crematogaster arborea subsp. melanogaster Emery, 1895h: 29 (w.q.) SOUTH AFRICA. Combination in C. (Crematogaster): Wheeler, W.M. 1922a: 840; in C. (Acrocoelia): Emery, 1922e: 147; in C. (Crematogaster): Bolton, 1995b: 166. Raised to species: Wheeler, W.M. 1922a: 840. Current subspecies: nominal plus homonyma.
Myrmicinae Crematogastrini Crematogaster monticola TRUE TRUE FALSEmonticola. Crematogaster monticola Arnold, 1920a: 517, fig. 40 (w.) SOUTH AFRICA. Combination in C. (Crematogaster): Wheeler, W.M. 1922a: 841.
Myrmicinae Crematogastrini Crematogaster natalensis TRUE TRUE FALSE2 subspecies
natalensis. Crematogaster sordidula subsp. natalensis Forel, 1910e: 431 (w.q.m.) SOUTH AFRICA. Combination in C. (Orthocrema): Wheeler, W.M. 1922a: 850. Raised to species: Emery, 1912e: 668. Subspecies of sordidula: Arnold, 1920a: 539; Wheeler, W.M. 1922a: 850. Revived status as species: Emery, 1922e: 131; Santschi, 1926b: 212; Menozzi, 1930b: 85. Current subspecies: nominal plus braunsi, dulcis.
Myrmicinae Crematogastrini Crematogaster opaciceps TRUE TRUE FALSE3 subspecies
opaciceps. Crematogaster opaciceps Mayr, 1901b: 12 (w.) SOUTH AFRICA. Combination in C. (Crematogaster): Wheeler, W.M. 1922a: 842; in C. (Acrocoelia): Emery, 1922e: 148; in C. (Crematogaster): Bolton, 1995b: 166. Current subspecies: nominal plus cacodaemon, clepens, defleta.
Myrmicinae Crematogastrini Crematogaster orobia TRUE TRUE FALSEorobia. Crematogaster orobia Santschi, 1919a: 343 (w.) SOUTH AFRICA. Combination in C. (Crematogaster): Wheeler, W.M. 1922a: 842.
Myrmicinae Crematogastrini Crematogaster oscaris TRUE TRUE FALSEoscaris. Crematogaster (Oxygyne) oscaris Forel, 1910f: 7, pl. 1, figs. 5, 6 (w.q.m.) SOUTH AFRICA.
Myrmicinae Crematogastrini Crematogaster peringueyi TRUE TRUE FALSE4 subspecies
peringueyi. Crematogaster peringueyi Emery, 1895h: 27, figs. 16, 17 (w.q.m.) SOUTH AFRICA. Combination in C. (Crematogaster): Wheeler, W.M. 1922a: 842; in C. (Acrocoelia): Emery, 1922e: 148; in C. (Crematogaster): Bolton, 1995b: 166. Current subspecies: nominal plus angustior, cacochyma, dentulata, gedeon.
Myrmicinae Crematogastrini Crematogaster schultzei TRUE TRUE FALSEschultzei. Crematogaster schultzei Forel, 1910f: 6 (w.q.) SOUTH AFRICA. Combination in C. (Crematogaster): Wheeler, W.M. 1922a: 843; in C. (Acrocoelia): Emery, 1922e: 148; in C. (Crematogaster): Bolton, 1995b: 166.
Myrmicinae Crematogastrini Crematogaster theta TRUE TRUE FALSEtheta. Crematogaster africana var. theta Forel, 1911f: 278 (w.) DEMOCRATIC REPUBLIC OF CONGO. Forel, 1916: 410 (q.). Combination in C. (Atopogyne): Wheeler, W.M. 1922a: 857. Raised to species: Wheeler, W.M. 1922a: 857; Emery, 1922e: 155; Santschi, 1935a: 262.
Myrmicinae Crematogastrini Crematogaster transvaalensis TRUE TRUE FALSE1 subspecies
transvaalensis. Crematogaster transvaalensis Forel, 1894b: 96 (w.) SOUTH AFRICA. Combination in C. (Orthocrema): Wheeler, W.M. 1922a: 851. Current subspecies: nominal plus hammi.
Myrmicinae Crematogastrini Crematogaster gerstaeckeri zulu TRUE TRUE FALSEzulu. Crematogaster (Atopogyne) bulawayensis var. zulu Wheeler, W.M. 1922a: 855 (w.q.) SOUTH AFRICA. [First available use of Crematogaster sjostedti st. bulawayensis var. zulu Santschi, 1920b: 15; unavailable name.] Subspecies of bulawayensis: Arnold, 1926: 289. Currently subspecies of gerstaeckeri: Bolton, 1995b: 165.
Myrmicinae Crematogastrini Crematogaster laurenti zeta TRUE TRUE FALSEzeta. Crematogaster (Atopogyne) laurenti var. zeta Emery, 1922e: 155 (w.) DEMOCRATIC REPUBLIC OF CONGO. [First available use of Crematogaster africana subsp. laurenti var. zeta Forel, 1909b: 70; unavailable name.] Currently subspecies of laurenti: Santschi, 1937g: 79.
Myrmicinae Crematogastrini Crematogaster kohli winkleri TRUE TRUE FALSEwinkleri. Crematogaster africana var. winkleri Forel, 1909b: 69 (w.) CAMEROUN. Forel, 1913a: 124 (q.). Combination in C. (Atopogyne): Wheeler, W.M. 1922a: 158; Santschi, 1937g: 80. Subspecies of buchneri: Forel, 1913a: 124; Forel, 1913b: 324; Forel, 1913e: 670; of africana: Wheeler, W.M. 1922a: 158. Currently subspecies of kohli: Santschi, 1937g: 80.
Myrmicinae Crematogastrini Crematogaster liengmei weitzeckeri TRUE TRUE FALSEweitzeckeri. Crematogaster weitzeckeri Emery, 1895h: 28, pl. 2, figs. 14, 15 (w.) SOUTH AFRICA. Combination in C. (Decacrema): Forel, 1910a: 18; Forel, 1910f: 9. Senior synonym of kloofensis: Arnold, 1920a: 549. Currently subspecies of liengmei: Forel, 1910a: 18; Santschi, 1914e: 21; Arnold, 1920a: 548.
Myrmicinae Crematogastrini Crematogaster africana variegata TRUE TRUE FALSEvariegata. Crematogaster africana var. variegata Mayr, 1902: 294 (w.) CAMEROUN. Combination in C. (Atopogyne): Wheeler, W.M. 1922a: 852.
Myrmicinae Crematogastrini Crematogaster africana tibialis TRUE TRUE FALSEtibialis. Crematogaster (Atopogyne) africana subsp. tibialis Santschi, in Wheeler, W.M. 1922a: 157 (w.) DEMOCRATIC REPUBLIC OF CONGO.
Myrmicinae Crematogastrini Crematogaster africana thoracica TRUE TRUE FALSEthoracica. Crematogaster (Atopogyne) africana st. thoracica Santschi, 1921c: 118 (w.) DEMOCRATIC REPUBLIC OF CONGO.
Myrmicinae Crematogastrini Crematogaster africana stolonis TRUE TRUE FALSEstolonis. Crematogaster (Atopogyne) africana st. stolonis Santschi, 1937g: 78, fig. 4 (w.) DEMOCRATIC REPUBLIC OF CONGO.
Myrmicinae Crematogastrini Crematogaster africana stanleyi TRUE TRUE FALSEstanleyi. Crematogaster (Atopogyne) africana var. stanleyi Wheeler, W.M. 1922a: 852 (w.) CONGO. [First available use of Crematogaster buchneri st. africana var. stanleyi Santschi, 1915c: 253; unavailable name.] [Crematogaster africana var. stanleyi Forel, 1911f: 278. Nomen nudum.] Santschi, 1937g: 79 (q.).
Myrmicinae Crematogastrini Crematogaster africana schumanni TRUE TRUE FALSEschumanni. Crematogaster africana subsp. schumanni Mayr, 1895: 144 (w.) CAMEROUN. Combination in C. (Atopogyne): Wheeler, W.M. 1922a: 852.
Myrmicinae Crematogastrini Crematogaster castanea rufonigra TRUE TRUE FALSErufonigra. Crematogaster tricolor subsp. rufonigra Emery, 1895h: 27 (w.) SOUTH AFRICA. Arnold, 1920a: 492 (q.m.). Combination in C. (Acrocoelia): Emery, 1922e: 146. Currently subspecies of castanea: Santschi, 1916b: 502.
Myrmicinae Crematogastrini Crematogaster gerstaeckeri rufescens TRUE TRUE FALSErufescens. Crematogaster (Crematogaster) sjostedti var. rufescens Wheeler, W.M. 1922a: 844 (w.) SOUTH AFRICA. [First available use of Crematogaster sjostedti st. bulawayensis var. rufescens Santschi, 1919b: 236; unavailable name.] Subspecies of bulawayensis: Arnold, 1926: 289. Currently subspecies of gerstaeckeri: Bolton, 1995b: 161.
Myrmicinae Crematogastrini Crematogaster acaciae generosa TRUE TRUE FALSEgenerosa. Crematogaster acaciae var. generosa Santschi, 1919a: 343 (w.q.) SOUTH AFRICA.
Myrmicinae Crematogastrini Crematogaster peringueyi gedeon TRUE TRUE FALSEgedeon. Crematogaster peringueyi var. gedeon Arnold, 1920a: 512 (w.) SOUTH AFRICA. [First available use of Crematogaster peringueyi r. cacodaemon var. gedeon Forel, 1915c: 341; unavailable name.] Combination in C. (Acrocoelia): Emery, 1922e: 148.
Myrmicinae Crematogastrini Crematogaster castanea durbanensis TRUE TRUE FALSEdurbanensis. Crematogaster ferruginea var. durbanensis Forel, 1914d: 234 (w.) SOUTH AFRICA. Arnold, 1920a: 491 (q.). Combination in C. (Acrocoelia): Emery, 1922e: 145. Currently subspecies of castanea: Bolton, 1995b: 152.
Myrmicinae Crematogastrini Crematogaster natalensis dulcis TRUE TRUE FALSEdulcis. Crematogaster (Orthocrema) natalensis var. dulcis Santschi, 1926b: 212 (w.) SOUTH AFRICA.
Myrmicinae Crematogastrini Crematogaster peringueyi dentulata TRUE TRUE FALSEdentulata. Crematogaster peringueyi r. dentulata Stitz, 1923: 156 (w.) SOUTH AFRICA.
Myrmicinae Crematogastrini Crematogaster opaciceps defleta TRUE TRUE FALSEdefleta. Crematogaster opaciceps subsp. defletus Forel, 1910e: 434 (w.) SOUTH AFRICA. Combination in C. (Acrocoelia): Emery, 1922e: 148.
Myrmicinae Crematogastrini Crematogaster neuvillei cooperi TRUE TRUE FALSEcooperi. Crematogaster neuvillei r. cooperi Forel, 1914d: 233 (w.) SOUTH AFRICA. Arnold, 1920a: 498 (q.). Combination in C. (Acrocoelia): Emery, 1922e: 148.
Myrmicinae Crematogastrini Crematogaster santschii clymene TRUE TRUE FALSEclymene. Crematogaster santschii var. clymene Forel, 1915c: 337 (w.) SOUTH AFRICA. Combination in C. (Oxygyne): Wheeler, W.M. 1922a: 858; in C. (Acrocoelia): Emery, 1922e: 148.
Myrmicinae Crematogastrini Crematogaster delagoensis acutidens TRUE TRUE FALSEacutidens. Crematogaster delagoense var. acutidens Arnold, 1944: 7, fig. 7 (w.) SOUTH AFRICA.
Myrmicinae Crematogastrini Crematogaster africana alligatrix TRUE TRUE FALSEalligatrix. Crematogaster buchneri subsp. alligatrix Forel, 1911e: 271 (w.) NIGERIA. Combination in C. (Atopogyne): Wheeler, W.M. 1922a: 852. Currently subspecies of africana: Wheeler, W.M. 1922a: 852.
Myrmicinae Crematogastrini Crematogaster peringueyi angustior TRUE TRUE FALSEangustior. Crematogaster peringueyi var. angustior Arnold, 1920a: 510, fig. 40 (w.q.m.) SOUTH AFRICA.
Myrmicinae Crematogastrini Crematogaster castanea arborea TRUE TRUE FALSEarborea. Crematogaster arboreus Smith, F. 1858b: 138, pl. 14 (nest) (w.) SOUTH AFRICA. Combination in C. (Acrocoelia): Emery, 1922e: 145. Junior synonym of tricolor: Mayr, 1895: 136 (footnote); Arnold, 1920a: 506 (note that arborea has priority); of castanea: Wheeler, W.M. 1922a: 830. Revived from synonymy as subspecies of castanea: Emery, 1922e: 145. Senior synonym of decolorata: Santschi, 1930b: 69. Currently subspecies of castanea: Santschi, 1930b: 69.
Myrmicinae Crematogastrini Crematogaster africana biemarginata TRUE TRUE FALSEbiemarginata. Crematogaster africana var. biemarginata Forel, 1910e: 433 (w.) CAMEROUN. Combination in C. (Atopogyne): Wheeler, 1922a: 852.
Myrmicinae Crematogastrini Crematogaster natalensis braunsi TRUE TRUE FALSEbraunsi. Crematogaster (Orthocrema) natalensis var. braunsi Emery, 1922e: 131 (w.) SOUTH AFRICA. [First available use of Crematogaster sordidula subsp. natalensis var. braunsi Forel, 1911f: 277; unavailable name.]
Myrmicinae Crematogastrini Crematogaster castanea bruta TRUE TRUE FALSEbruta. Crematogaster castanea st. bruta Santschi, 1926b: 213 (w.) SOUTH AFRICA. [First available use of Crematogaster tricolor st. ferruginea var. bruta Santschi, 1913a: 413; unavailable name.]
Myrmicinae Crematogastrini Crematogaster amita bushimana TRUE TRUE FALSEbushimana. Crematogaster (Sphaerocrema) amita var. bushimana Santschi, 1926b: 226 (w.q.) SOUTH AFRICA.
Myrmicinae Crematogastrini Crematogaster peringueyi cacochyma TRUE TRUE FALSEcacochyma. Crematogaster peringueyi r. cacochyma Forel, 1914d: 235 (w.q.) SOUTH AFRICA. Combination in C. (Acrocoelia): Emery, 1922e: 512.
Myrmicinae Crematogastrini Crematogaster opaciceps cacodaemon TRUE TRUE FALSEcacodaemon. Crematogaster peringueyi r. cacodaemon Forel, 1914d: 234 (w.q.m.) SOUTH AFRICA. Combination in C. (Acrocoelia): Emery, 1922e: 148. Currently subspecies of opaciceps: Santschi, 1926b: 216.
Myrmicinae Crematogastrini Crematogaster liengmei caculata TRUE TRUE FALSEcaculata. Crematogaster peringueyi r. caculata Forel, 1915c: 341 (w.) SOUTH AFRICA. Combination in C. (Decacrema): Arnold, 1920a: 550; Wheeler, W.M. 1922a: 861; in C. (Acrocoelia): Emery, 1922e: 148. Currently subspecies of liengmei: Arnold, 1920a: 550; Wheeler, W.M. 1922a: 861.
Myrmicinae Crematogastrini Crematogaster amita caffra TRUE TRUE FALSEcaffra. Crematogaster (Sphaerocrema) amita var. caffra Santschi, 1926b: 226 (w.q.m.) SOUTH AFRICA. [First available use of Crematogaster (Sphaerocrema) kneri st. amita var. caffra Santschi, in Forel, 1922: 94 (footnote); unavailable name.]
Myrmicinae Crematogastrini Crematogaster africana camena TRUE TRUE FALSEcamena. Crematogaster (Atopogyne) africana var. camena Wheeler, W.M. 1922a: 852 (w.) DEMOCRATIC REPUBLIC OF CONGO. [First available use of Crematogaster buchneri r. africana var. camena Forel, 1916: 410; unavailable name.]
Myrmicinae Crematogastrini Crematogaster gerstaeckeri godefreyi TRUE TRUE FALSEgodefreyi. Crematogaster foraminiceps r. godefreyi Forel, 1914d: 237 (w.) SOUTH AFRICA. Combination in C. (Crematogaster): Wheeler, W.M. 1922a: 839; in C. (Acrocoelia): Emery, 1922e: 148. Subspecies of vulcania: Forel, 1916: 406; of sjostedti: Santschi, 1918d: 184; Emery, 1922e: 148. Raised to species: Arnold, 1920a: 503; Wheeler, W.M. 1922a: 839. Currently subspecies of gerstaeckeri: Santschi, 1930b: 69. Senior synonym of carininotum: Santschi, 1918d: 184.
Myrmicinae Crematogastrini Crematogaster melanogaster homonyma TRUE TRUE FALSEhomonyma. Crematogaster (Acrocoelia) melanogaster var. homonyma Emery, 1922e: 147 (w.q.) SOUTH AFRICA.
Myrmicinae Crematogastrini Crematogaster censor junodi TRUE TRUE FALSEjunodi. Crematogaster censor r. junodi Forel, 1916: 406 (w.) SOUTH AFRICA. Combination in C. (Acrocoelia): Emery, 1922e: 146.
Myrmicinae Crematogastrini Crematogaster gambiensis krantziana TRUE TRUE FALSEkrantziana. Crematogaster gambiensis var. krantziana Forel, 1914d: 234 (w.m.) SOUTH AFRICA. Arnold, 1920a: 530 (q.). Combination in C. (Sphaerocrema): Wheeler, W.M. 1922a: 846.
Myrmicinae Crematogastrini Crematogaster castanea mediorufa TRUE TRUE FALSEmediorufa. Crematogaster tricolor var. mediorufa Forel, 1907g: 81 (footnote) (w.m.) SOUTH AFRICA. Forel, 1913a: 124 (q.); Arnold, 1920a: 493 (w.q.m.). Currently subspecies of castanea: Bolton, 1995b: 157.
Myrmicinae Crematogastrini Crematogaster delagoensis merwei TRUE TRUE FALSEmerwei. Crematogaster (Acrocoelia) delagoensis st. merwei Santschi, 1932a: 385 (w.) SOUTH AFRICA.
Myrmicinae Crematogastrini Crematogaster castanea museisapientiae TRUE TRUE FALSEmuseisapientiae. Crematogaster ferruginea subsp. museisapientiae Forel, 1911e: 272 (w.q.m.) "Tropical East Africa". Combination in C. (Acrocoelia): Emery, 1922e: 146. Subspecies of tricolor: Santschi, 1913a: 413 (in key). Currently subspecies of castanea: Wheeler, W.M. 1922a: 833.
Myrmicinae Crematogastrini Crematogaster brunneipennis omniparens TRUE TRUE FALSEomniparens. Crematogaster brunneipennis r. omniparens Forel, 1914d: 237 (w.) SOUTH AFRICA. Combination in C. (Acrocoelia): Emery, 1922e: 145.
Myrmicinae Crematogastrini Crematogaster africana polymorphica TRUE TRUE FALSEpolymorphica. Crematogaster (Atopogyne) africana subsp. polymorphica Weber, 1943c: 357 (w.) SUDAN.
Myrmicinae Solenopsidini Diplomorium longipenne TRUE TRUE FALSElongipenne. Diplomorium longipenne Mayr, 1901b: 18 (w.q.) SOUTH AFRICA. See also: Arnold, 1916: 240; Bolton, 1987: 278.
Proceratiinae Proceratiini Discothyrea hewitti TRUE TRUE FALSEhewitti. Discothyrea hewitti Arnold, 1916: 160 (q.) SOUTH AFRICA.
Proceratiinae Proceratiini Discothyrea traegaordhi TRUE TRUE FALSEtraegaordhi. Discothyrea traegaordhi Santschi, 1914e: 3 (w.) SOUTH AFRICA. See also: Brown, 1958g: 340.
Dorylinae Dorylini Dorylus erraticus TRUE TRUE FALSEerraticus. Anomma erratica Smith, F. 1865: 71 (w.) AFRICA (not New Guinea as stated; see: Wilson, 1964a: 443). Combination in Dorylus: Emery, 1910b: 15; in D. (Anomma): Donisthorpe, 1932c: 473.
Dorylinae Dorylini Dorylus faurei TRUE TRUE FALSEfaurei. Dorylus faurei Arnold, 1946: 60, fig. 12 (q.) SOUTH AFRICA.
Dorylinae Dorylini Dorylus fulvus TRUE TRUE FALSE11 subspecies
fulvus. Typhlopone fulva Westwood, 1839: 219, fig. 86 (w.) NORTH AFRICA. Emery, 1895j: 723 (m.). Combination in Dorylus: Dalla Torre, 1893: 10; in D. (Typhlopone): Emery, 1895j: 723. Senior synonym of kirbii: Smith, F. 1858b: 110; Mayr, 1863: 457; of clausii, dahlbomii, shuckardi, spinolae, thwaitsii: Emery, 1895j: 723. Current subspecies: nominal plus badius, crosi, dentifrons, eurous, glabratus, juvenculus, mordax, obscurior, punicus, ruficeps, saharensis.
Dorylinae Dorylini Dorylus furcatus TRUE TRUE FALSEfurcatus. Dichthadia furcata Gerstäcker, 1872: 267 (q.) SOUTH AFRICA. Combination in Dorylus: Emery, 1888b: 345.
Dorylinae Dorylini Dorylus helvolus TRUE TRUE FALSE1 subspecies
helvolus. Vespa helvola Linnaeus, 1764: 412 (m.) SOUTH AFRICA. Linnaeus, 1767: 968 (w.); Emery, 1888b: 344 (q.). Combination in Dorylus: Fabricius, 1793: 365. Senior synonym of dorylus, europaea, planifrons, punctata: Emery, 1888b: 350; Emery, 1895j: 714. Current subspecies: nominal plus pretoriae. See also: Arnold, 1915: 116.
Dorylinae Dorylini Dorylus attenuatus acuminatus TRUE TRUE FALSEacuminatus. Dorylus (Alaopone) attenuatus var. acuminata Emery, 1899e: 462 (m.) SOUTH AFRICA.
Dorylinae Dorylini Dorylus attenuatus australis TRUE TRUE FALSEaustralis. Dorylus (Alaopone) montanum st. australis Santschi, 1919b: 231 (w.) SOUTH AFRICA. [Also described as new, but as species, by Santschi, 1920d: 362; based on same specimens.] Subspecies of montanus: Wheeler, W.M. 1922a: 750. Currently subspecies of attenuatus: Santschi, 1939a: 147.
Dorylinae Dorylini Dorylus attenuatus bondroiti TRUE TRUE FALSEbondroiti. Dorylus (Alaopone) montanus var. bondroiti Santschi, 1912b: 162 (m.) SOUTH AFRICA. Subspecies of montanus: Santschi, 1915c: 248. Currently subspecies of attenuatus: Santschi, 1939a: 148.
Dorylinae Dorylini Dorylus fimbriatus poweri TRUE TRUE FALSEpoweri. Dorylus (Rhogmus) fimbriatus var. poweri Forel, 1914d: 217 (w.) SOUTH AFRICA.
Dorylinae Dorylini Dorylus helvolus pretoriae TRUE TRUE FALSEpretoriae. Dorylus helvolus subsp. pretoriae Arnold, 1946: 59, fig. 11 (w.q.m.) SOUTH AFRICA.
Dorylinae Dorylini Dorylus emeryi pulsi TRUE TRUE FALSEpulsi. Anomma emeryi var. pulsi Forel, 1904d: 170 (w.) "West Africa". Santschi, 1923e: 273 (s.). Combination in Dorylus (Anomma): Emery, 1910b: 11.
Dorylinae Dorylini Dorylus nigricans rubellus TRUE TRUE FALSErubellus. Anomma rubella Savage, 1849: 196 (w.) "West Africa". Combination in Dorylus (Anomma): Emery, 1895j: 710. Subspecies of burmeisteri: Emery, 1892d: 554. Currently subspecies of nigricans: Emery, 1895j: 701; Emery, 1910b: 12; Forel, 1913h: 348.
Ponerinae Ponerini Hypoponera austra TRUE TRUE FALSEaustra. Hypoponera austra Bolton & Fisher, 2011: 27, figs. 7-9 (w.q.,ergatoid q.) SOUTH AFRICA.
Ponerinae Ponerini Hypoponera blanda TRUE TRUE FALSEblanda. Hypoponera blanda Bolton & Fisher, 2011: 29, figs. 10-12 (w.q.) CENTRAL AFRICAN REPUBLIC.
Ponerinae Ponerini Hypoponera boerorum TRUE TRUE FALSEboerorum. Ponera coarctata r. boerorum Forel, 1901f: 339 (w.) SOUTH AFRICA. Combination in Hypoponera: Taylor, 1967a: 12. Raised to species: Santschi, 1938b: 78. See also: Bolton & Fisher, 2011: 30.
Ponerinae Ponerini Hypoponera meridia TRUE TRUE FALSEmeridia. Hypoponera meridia Bolton & Fisher, 2011: 70, figs. 73-75 (w.q.) SOUTH AFRICA.
Ponerinae Ponerini Hypoponera natalensis TRUE TRUE FALSEnatalensis. Ponera coarctata st. natalensis Santschi, 1914e: 7 (w.) SOUTH AFRICA. Combination in P. (Hypoponera): Santschi, 1938b: 79; in Hypoponera: Taylor, 1967a: 12. Raised to species: Santschi, 1938b: 79; Taylor, 1967a: 12. See also: Bolton & Fisher, 2011: 74.
Ponerinae Ponerini Hypoponera perparva TRUE TRUE FALSEperparva. Hypoponera perparva Bolton & Fisher, 2011: 82, figs. 91-93 (w.) SOUTH AFRICA.
Ponerinae Ponerini Hypoponera spei TRUE TRUE FALSEspei. Ponera spei Forel, 1910e: 423 (w.) SOUTH AFRICA. Combination in Hypoponera: Bolton, 1995b: 216. Senior synonym of fidelis and material of the unavailable name sancta referred here: Bolton & Fisher, 2011: 101. See also: Arnold, 1915: 77.
Ponerinae Ponerini Hypoponera sulcatinasis TRUE TRUE FALSEsulcatinasis. Ponera sulcatinasis Santschi, 1914e: 4 (w.) SOUTH AFRICA. Combination in Hypoponera: Bolton, 1995b: 216. Senior synonym of devota: Bolton & Fisher, 2011: 103.
Ponerinae Ponerini Hypoponera traegaordhi TRUE TRUE FALSEtraegaordhi. Ponera traegaordhi Santschi, 1914e: 6 (w.) SOUTH AFRICA. Combination in P. (Hypoponera): Santschi, 1938b: 79; in Hypoponera: Bolton, 1995b: 216. See also: Bolton & Fisher, 2011: 108.
Ponerinae Ponerini Hypoponera transvaalensis TRUE TRUE FALSEtransvaalensis. Ponera transvaalensis Arnold, 1947: 132, fig. 2 (w.) SOUTH AFRICA. Combination in Hypoponera: Bolton, 1995b: 216. See also: Bolton & Fisher, 2011: 108.
Formicinae Plagiolepidini Lepisiota capensis TRUE TRUE FALSE13 subspecies
capensis. Acantholepis capensis Mayr, 1862: 699 (w.) SOUTH AFRICA. Mayr, 1865: 57 (m.); Emery, 1877b: 366 (q.); Wheeler, G.C. & Wheeler, J. 1980: 542 (l.). Combination in Lepisiota: Xu, 1994c: 233. Current subspecies: nominal plus acholli, anceps, guineensis, issore, junodi, laevis, lunaris, minuta, simplex, simplicoides, specularis, subopaciceps, thoth. See also: Bingham, 1903: 316; Arnold, 1920a: 568.
Formicinae Plagiolepidini Lepisiota crinita TRUE TRUE FALSEcrinita. Acantholepis crinita Mayr, 1895: 149 (w.) SOUTH AFRICA. Combination in Lepisiota: Bolton, 1995b: 227.
Formicinae Plagiolepidini Lepisiota capensis simplicoides TRUE TRUE FALSEsimplicoides. Acantholepis capensis var. simplicoides Forel, 1907g: 86 (footnote) (w.) SOUTH AFRICA. Combination in Lepisiota: Bolton, 1995b: 228.
Formicinae Plagiolepidini Lepisiota spinosior natalensis TRUE TRUE FALSEnatalensis. Acantholepis spinosior var. natalensis Arnold, 1920a: 559 (w.) SOUTH AFRICA. Combination in Lepisiota: Bolton, 1995b: 228.
Formicinae Plagiolepidini Lepisiota capensis minuta TRUE TRUE FALSEminuta. Acantholepis simplex var. minuta Forel, 1916: 438 (w.) SOUTH AFRICA. Combination in Lepisiota: Bolton, 1995b: 228. Currently subspecies of capensis: Arnold, 1920a: 572.
Formicinae Plagiolepidini Lepisiota capensis junodi TRUE TRUE FALSEjunodi. Acantholepis capensis r. junodi Forel, 1916: 438 (w.) SOUTH AFRICA. Combination in Lepisiota: Bolton, 1995b: 227.
Leptanillinae Leptanillini Leptanilla africana TRUE TRUE FALSEafricana. Leptanilla africana Baroni Urbani, 1977c: 468, figs. 34-37, 43 (m.) NIGERIA.
Leptanillinae Leptanillini Leptanilla australis TRUE TRUE FALSEaustralis. Leptanilla australis Baroni Urbani, 1977c: 471, figs. 38, 40-42, 44 (m.) SOUTH AFRICA.
Ponerinae Ponerini Leptogenys attenuata TRUE TRUE FALSEattenuata. Ponera attenuata Smith, F. 1858b: 91 (w.) SOUTH AFRICA. Arnold, 1915: 98 (m.). Combination in Lobopelta: Roger, 1863b: 19; in Leptogenys: Mayr, 1895: 124. Senior synonym of jaegerskjoeldi: Arnold, 1926: 210. See also: Bolton, 1975a: 274.
Ponerinae Ponerini Leptogenys castanea TRUE TRUE FALSEcastanea. Lobopelta castanea Mayr, 1862: 734 (w.) SOUTH AFRICA. Wheeler, G.C. & Wheeler, J. 1986c: 89 (l.). Combination in Leptogenys: Emery, 1895h: 20. Senior synonym of bellua, dispar, hewitti, parva: Bolton, 1975a: 285.
Ponerinae Ponerini Leptogenys crassinoda TRUE TRUE FALSEcrassinoda. Leptogenys attenuata var. crassinoda Arnold, 1926: 210 (w.) SOUTH AFRICA. Raised to species: Bolton, 1975a: 276.
Ponerinae Ponerini Leptogenys crustosa TRUE TRUE FALSEcrustosa. Leptogenys crustosa Santschi, 1914d: 326, fig. 12 (w.) GUINEA. Senior synonym of africanus, rufipes: Bolton, 1975a: 251.
Ponerinae Ponerini Leptogenys havilandi TRUE TRUE FALSEhavilandi. Leptogenys (Lobopelta) havilandi Forel, 1901f: 332 (w.) SOUTH AFRICA. See also: Bolton, 1975a: 282.
Ponerinae Ponerini Leptogenys intermedia TRUE TRUE FALSEintermedia. Leptogenys intermedia Emery, 1902c: 32 (w.) SOUTH AFRICA. See also: Forel, 1914d: 214. Junior synonym of nitida Smith: Bolton, 1975a: 289. First available replacement name for nitida Smith, F. 1858b: 92 [Junior primary homonym of †nitida Heer, 1849: 149.]; designated by Bolton, 1995b: 231. Hence senior synonym of tenuis: Forel, 1912n: 52; Bolton, 1975a: 289; of adpressa, aena, capensis, deflocata, gracilis, grandior, insinuata, intermedia, speculans: Bolton, 1975a: 289.
Ponerinae Ponerini Leptogenys mactans TRUE TRUE FALSEmactans. Leptogenys mactans Bolton, 1975a: 288, fig. 17 (w.) SOUTH AFRICA.
Ponerinae Ponerini Leptogenys peringueyi TRUE TRUE FALSEperingueyi. Leptogenys (Lobopelta) havilandi st. peringueyi Forel, 1913j: 210 (w.) SOUTH AFRICA. Raised to species: Arnold, 1915: 96. See also: Bolton, 1975a: 282.
Myrmicinae Meranoplini Meranoplus inermis TRUE TRUE FALSEinermis. Meranoplus inermis Emery, 1895h: 41, pl. 2, fig. 24 (w.) SOUTH AFRICA. Subspecies of nanus: Wheeler, W.M. 1922a: 888. Revived status as species and senior synonym of affinis, kiboshana, nanior, soriculus: Bolton, 1981a: 52.
Myrmicinae Meranoplini Meranoplus peringueyi TRUE TRUE FALSEperingueyi. Meranoplus peringueyi Emery, 1886: 365, pl. 17, fig. 12 (w.) SOUTH AFRICA. Senior synonym of excisus: Bolton, 1981a: 56.
Myrmicinae Pheidolini Messor arenarius TRUE TRUE FALSE4 subspecies
arenarius. Formica arenaria Fabricius, 1787: 310 (w.) NORTH AFRICA. André, 1883b: 366 (q.); Emery, 1891b: 12 (q.m.); Forel, 1902a: 148 (m.). Combination in Aphaenogaster: Roger, 1863b: 29; in A. (Messor): Forel, 1890a: lxx; in Messor: Forel, 1894d: 45; in Stenamma (Messor): Emery, 1898c: 125; in Messor: Emery, 1908e: 439. Senior synonym of amaurocyclia, scalpturata: Roger, 1862c: 292; of bugnioni: Forel, 1904f: 424. Current subspecies: nominal plus alfierii, diabolus, ratus, thebaicus. See also: Emery, 1908e: 439; Santschi, 1938a: 32.
Myrmicinae Pheidolini Messor capensis TRUE TRUE FALSEcapensis. Atta capensis Mayr, 1862: 743 (w.) SOUTH AFRICA. Mayr, 1866b: 896 (q.m.). Combination in Aphaenogaster: Roger, 1863b: 30; in Stenamma (Messor): Emery, 1895h: 35; in Messor: Forel, 1910e: 444. Subspecies of barbarus: Emery, 1891b: 12; Forel, 1910e: 444; Arnold, 1920a: 405; Emery, 1922c: 98; Stitz, 1923: 150; Prins, 1963: 100. Status as species: Santschi, 1917e: 94; Wheeler, W.M. 1922a: 804; Bolton, 1982: 345. Senior synonym of braunsi, donisthorpei, pseudoaegyptiaca, schencki: Bolton, 1982: 345.
Myrmicinae Pheidolini Messor decipiens TRUE TRUE FALSEdecipiens. Messor capensis st. decipiens Santschi, 1917e: 94 (w.q.) SOUTH AFRICA. [First available use of Stenamma (Messor) barbarum r. capense var. decipiens Forel, 1905b: 177; unavailable name.] Raised to species and senior synonym of arcistriatus, probus: Bolton, 1982: 348.
Myrmicinae Pheidolini Messor piceus TRUE TRUE FALSEpiceus. Messor piceus Stitz, 1923: 150 (w.q.) SOUTH AFRICA. See also: Bolton, 1982: 352.
Myrmicinae Dacetini Microdaceton exornatum TRUE TRUE FALSEexornatum. Microdaceton exornatum Santschi, 1913g: 478 (w.) SOUTH AFRICA. [Also described as new by Santschi, 1914e: 33.] Arnold, 1948: 225 (q.). Senior synonym of laevior, leakeyi: Bolton, 1983: 402. See also: Bolton, 2000: 28.
Myrmicinae Solenopsidini Monomorium albopilosum TRUE TRUE FALSEalbopilosum. Monomorium albopilosum Emery, 1895h: 24 (w.) SOUTH AFRICA. Combination in M. (Xeromyrmex): Santschi, 1919b: 235. Senior synonym of clarithorax, fingo, paucipilosa, thales: Bolton, 1987: 335.
Myrmicinae Solenopsidini Monomorium anceps TRUE TRUE FALSEanceps. Monomorium subopacum var. anceps Emery, 1895h: 24 (w.) SOUTH AFRICA. Combination in M. (Xeromyrmex): Wheeler, W.M. 1922a: 872. Raised to species: Bolton, 1987: 336.
Myrmicinae Solenopsidini Monomorium australe TRUE TRUE FALSEaustrale. Monomorium subopacum r. australe Emery, 1886: 363 (w.q.) SOUTH AFRICA. Forel, 1913j: 216 (q.). Combination in M. (Parholcomyrmex): Santschi, 1917b: 282; Wheeler, W.M. 1922a: 873. Subspecies of salomonis: Forel, 1913j: 216; Arnold, 1916: 225. Raised to species: Dalla Torre, 1893: 65; Santschi, 1917b: 282; Bolton, 1987: 337. Material of the unavailable name laeviceps referred here by Bolton, 1987: 337.
Myrmicinae Solenopsidini Monomorium boerorum TRUE TRUE FALSEboerorum. Monomorium minutum var. boerorum Forel, 1910e: 442 (w.q.) SOUTH AFRICA. Raised to species: Bolton, 1987: 381.
Myrmicinae Solenopsidini Monomorium braunsi TRUE TRUE FALSEbraunsi. Monomorium braunsi Mayr, 1901b: 7 (w.) SOUTH AFRICA. See also: Bolton, 1987: 382.
Myrmicinae Solenopsidini Monomorium excelsior TRUE TRUE FALSEexcelsior. Monomorium tchelichofi var. excelsior Arnold, 1926: 227 (w.m.) SOUTH AFRICA. Raised to species and senior synonym of speculiceps: Bolton, 1987: 344.
Myrmicinae Solenopsidini Monomorium excensurae TRUE TRUE FALSEexcensurae. Monomorium oscaris var. excensurae Forel, 1915c: 342 (w.) SOUTH AFRICA. Raised to species: Bolton, 1987: 386.
Myrmicinae Solenopsidini Monomorium fastidium TRUE TRUE FALSEfastidium. Monomorium fastidium Bolton, 1987: 389 (w.q.) SOUTH AFRICA.
Myrmicinae Solenopsidini Monomorium fridae TRUE TRUE FALSEfridae. Monomorium medinae r. fridae Forel, 1905b: 183 (w.) SOUTH AFRICA. Combination in M. (Xeromyrmex): Wheeler, W.M. 1922a: 870. Raised to species: Bolton, 1987: 344.
Myrmicinae Solenopsidini Monomorium havilandi TRUE TRUE FALSEhavilandi. Monomorium havilandi Forel, 1910e: 443 (w.q.) SOUTH AFRICA. Combination in M. (Parholcomyrmex): Santschi, 1917b: 282. Subspecies of australe: Santschi, 1917b: 282. Revived status as species and senior synonym of distinctum, leviceps: Bolton, 1987: 368.
Myrmicinae Solenopsidini Monomorium junodi TRUE TRUE FALSEjunodi. Monomorium salomonis subsp. junodi Forel, 1910e: 441 (w.) SOUTH AFRICA. Combination in M. (Xeromyrmex): Santschi, 1919b: 235. Subspecies of delagoense: Santschi, 1928f: 192; Arnold, 1944: 14. Raised to species and senior synonym of pretoriensis: Bolton, 1987: 346.
Myrmicinae Solenopsidini Monomorium kelapre TRUE TRUE FALSEkelapre. Monomorium kelapre Bolton, 1987: 395 (w.) SOUTH AFRICA.
Myrmicinae Solenopsidini Monomorium lubricum TRUE TRUE FALSElubricum. Monomorium lubricum Arnold, 1948: 218, fig. 7 (w.) SOUTH AFRICA. See also: Bolton, 1987: 398.
Myrmicinae Solenopsidini Monomorium macrops TRUE TRUE FALSEmacrops. Monomorium mediocre subsp. macrops Arnold, 1944: 11, fig. 17 (w.) SOUTH AFRICA. Raised to species: Bolton, 1987: 369.
Myrmicinae Solenopsidini Monomorium mavide TRUE TRUE FALSEmavide. Monomorium mavide Bolton, 1987: 400 (w.) SOUTH AFRICA.
Myrmicinae Solenopsidini Monomorium mediocre TRUE TRUE FALSEmediocre. Monomorium mediocre Santschi, 1920d: 376, fig. 13 (w.) SOUTH AFRICA. See also: Bolton, 1987: 349.
Myrmicinae Solenopsidini Monomorium micropacum TRUE TRUE FALSEmicropacum. Monomorium micropacum Bolton, 1987: 350 (w.) SOUTH AFRICA.
Myrmicinae Solenopsidini Monomorium modestum TRUE TRUE FALSEmodestum. Monomorium modestum Santschi, 1914e: 17 (w.) SOUTH AFRICA. Combination in Syllophopsis: Santschi, 1915c: 259; in Monomorium: Bolton, 1987: 423. Senior synonym of smutsi, transwaalensis: Bolton, 1987: 423. See also: Heterick, 2006: 177.
Myrmicinae Solenopsidini Monomorium musicum TRUE TRUE FALSEmusicum. Monomorium oscaris subsp. musicum Forel, 1910e: 442 (w.q.) SOUTH AFRICA. Raised to species: Emery, 1922e: 173; Bolton, 1987: 402.
Myrmicinae Solenopsidini Monomorium notulum TRUE TRUE FALSEnotulum. Monomorium setuliferum var. notula Forel, 1910e: 441 (w.m.) SOUTH AFRICA. Raised to species and senior synonym of dolichops, latior: Bolton, 1987: 370.
Myrmicinae Solenopsidini Monomorium ocellatum TRUE TRUE FALSEocellatum. Monomorium (Xeromyrmex) salomonis st. ocelletum Arnold, in Santschi, 1920d: 377 (w.) SOUTH AFRICA. Arnold, 1926: 229 (q.). Raised to species: Santschi, 1936a: 39; Arnold, 1944: 14; Bolton, 1987: 352.
Myrmicinae Solenopsidini Monomorium orangiae TRUE TRUE FALSEorangiae. Monomorium (Xeromyrmex) orangiae Arnold, 1956: 67, fig. 16 (w.) SOUTH AFRICA. See also: Bolton, 1987: 354.
Myrmicinae Solenopsidini Monomorium pacis TRUE TRUE FALSEpacis. Monomorium pacis Forel, 1915c: 343 (w.) SOUTH AFRICA. See also: Bolton, 1987: 404.
Myrmicinae Solenopsidini Monomorium paternum TRUE TRUE FALSEpaternum. Monomorium paternum Bolton, 1987: 405, fig. 89 (w.) SOUTH AFRICA.
Myrmicinae Solenopsidini Monomorium rhopalocerum TRUE TRUE FALSErhopalocerum. Monomorium rhopalocerum Emery, 1895h: 25, pl. 2, fig 29 (w.) SOUTH AFRICA. Senior synonym of hottentota, leimbachi: Bolton, 1987: 407.
Myrmicinae Solenopsidini Monomorium rotundatum TRUE TRUE FALSErotundatum. Monomorium rotundatum Santschi, 1920b: 14, fig. 2 (w.) SOUTH AFRICA. See also: Bolton, 1987: 409.
Myrmicinae Solenopsidini Monomorium schultzei TRUE TRUE FALSEschultzei. Monomorium schultzei Forel, 1910f: 18 (w.q.) SOUTH AFRICA. See also: Bolton, 1987: 410.
Myrmicinae Solenopsidini Monomorium tablense TRUE TRUE FALSEtablense. Monomorium altinode st. tablensis Santschi, 1932a: 384, figs. 6, 7 (w.q.) SOUTH AFRICA. Raised to species: Bolton, 1987: 414.
Myrmicinae Solenopsidini Monomorium taedium TRUE TRUE FALSEtaedium. Monomorium taedium Bolton, 1987: 415 (w.) SOUTH AFRICA.
Myrmicinae Solenopsidini Monomorium tchelichofi TRUE TRUE FALSEtchelichofi. Monomorium tchelichofi Forel, 1914d: 244 (w.) SOUTH AFRICA. See also: Bolton, 1987: 362.
Myrmicinae Solenopsidini Monomorium torvicte TRUE TRUE FALSEtorvicte. Monomorium torvicte Bolton, 1987: 416 (w.) SOUTH AFRICA.
Myrmicinae Solenopsidini Monomorium willowmorense TRUE TRUE FALSEwillowmorense. Monomorium willowmorense Bolton, 1987: 364, fig. 54 (w.) SOUTH AFRICA. Material of the unavailable names willowmorensis, belli referred here by Bolton, 1987: 364. See also: Heterick, 2006: 107.
Myrmicinae Solenopsidini Monomorium xanthognathum TRUE TRUE FALSExanthognathum. Monomorium xanthognathum Arnold, 1944: 9, fig. 15 (w.) SOUTH AFRICA. See also: Bolton, 1987: 371.
Myrmicinae Solenopsidini Monomorium zulu TRUE TRUE FALSEzulu. Monomorium zulu Santschi, 1914e: 18 (w.) SOUTH AFRICA. See also: Bolton, 1987: 365.
Myrmicinae Myrmicariini Myrmicaria anomala TRUE TRUE FALSEanomala. Myrmicaria anomala Arnold, 1960a: 81, fig. 2 (w.) SOUTH AFRICA.
Myrmicinae Myrmicariini Myrmicaria faurei TRUE TRUE FALSEfaurei. Myrmicaria faurei Arnold, 1947: 134, fig. 3 (w.) SOUTH AFRICA.
Myrmicinae Myrmicariini Myrmicaria foreli TRUE TRUE FALSE1 subspecies
foreli. Myrmicaria foreli Santschi, 1925c: 163 (w.) SOUTH AFRICA. Current subspecies: nominal plus pallida.
Myrmicinae Myrmicariini Myrmicaria laevior TRUE TRUE FALSElaevior. Myrmicaria laevior Forel, 1910e: 445 (w.) SOUTH AFRICA.
Myrmicinae Myrmicariini Myrmicaria natalensis TRUE TRUE FALSE6 subspecies
natalensis. Physatta natalensis Smith, F. 1858b: 172 (q.) SOUTH AFRICA. Santschi, 1925c: 143 (q.m.). Combination in Myrmicaria: Mayr, 1866b: 905. Junior synonym of eumenoides: Mayr, 1866b: 905 (note that natalensis has priority). Revived from synonymy and senior synonym of sulcatus: Emery, 1922e: 123. Current subspecies: nominal plus eumenoides, navicula, nigriventris, obscuriceps, taeniata, verticalis. [M. natalensis referred by error to Myrmica by Santschi, 1925c: 143.]
Myrmicinae Myrmicariini Myrmicaria nigra TRUE TRUE FALSEnigra. Heptacondylus niger Mayr, 1862: 755 (w.) SOUTH AFRICA. Arnold, 1916: 270 (m.). Combination in Myrmicaria: Dalla Torre, 1893: 155. See also: Mayr, 1865: 110.
Myrmicinae Myrmicariini Myrmicaria foreli pallida TRUE TRUE FALSEpallida. Myrmicaria foreli var. pallida Arnold, in Santschi, 1925c: 165 (w.m.) SOUTH AFRICA.
Myrmicinae Myrmicariini Myrmicaria natalensis obscuriceps TRUE TRUE FALSEobscuriceps. Myrmicaria natalensis var. obscuriceps Santschi, 1937d: 228 (w.) SOUTH AFRICA. Currently subspecies of natalensis: Santschi, 1939b: 238.
Myrmicinae Formicoxenini Myrmoxenus africana TRUE TRUE FALSEafricana. Epimyrma (Gonepimyrma) africana Bernard, 1948: 148, fig. 7 (w.) ALGERIA. Combination in Myrmoxenus (Gonepimyrma): new combination (unpublished).
Myrmicinae Formicoxenini Nesomyrmex antoinetteae TRUE TRUE FALSEantoinetteae. Nesomyrmex antoinetteae Mbanyana & Robertson, 2008: 39, figs. 2g-i (w.) SOUTH AFRICA.
Myrmicinae Formicoxenini Nesomyrmex braunsi TRUE TRUE FALSEbraunsi. Dilobocondyla (Tetramyrma) braunsi Forel, 1912m: 767 (w.) SOUTH AFRICA. Forel, 1913a: 122 (pseudogyne); Arnold, 1917: 359 (q.). Combination in Tetramyrma: Forel, 1913a: 122; in Leptothorax: Bolton, 1982: 325; in Nesomyrmex: Bolton, 2003: 272. See also: Mbanyana & Robertson, 2008: 42.
Myrmicinae Formicoxenini Nesomyrmex cederbergensis TRUE TRUE FALSEcederbergensis. Nesomyrmex cederbergensis Mbanyana & Robertson, 2008: 42, figs. 4g-i (w.) SOUTH AFRICA.
Myrmicinae Formicoxenini Nesomyrmex denticulatus TRUE TRUE FALSEdenticulatus. Leptothorax denticulatus Mayr, 1901b: 5 (w.q.) SOUTH AFRICA. Combination in L. (Goniothorax): Wheeler, W.M. 1922a: 891; in Nesomyrmex: Bolton, 2003: 272. See also: Bolton, 1982: 328; Mbanyana & Robertson, 2008: 38.
Myrmicinae Formicoxenini Nesomyrmex entabeni TRUE TRUE FALSEentabeni. Nesomyrmex entabeni Mbanyana & Robertson, 2008: 43, figs. 3d-i (w.) SOUTH AFRICA.
Myrmicinae Formicoxenini Nesomyrmex ezantsi TRUE TRUE FALSEezantsi. Nesomyrmex ezantsi Mbanyana & Robertson, 2008: 43, figs. 4m-o (w.) SOUTH AFRICA.
Myrmicinae Formicoxenini Nesomyrmex humerosus TRUE TRUE FALSEhumerosus. Leptothorax (Goniothorax) humerosus Emery, 1896g: 62 (w.) EAST AFRICA (no locality given, probably KENYA). Combination in Nesomyrmex: Bolton, 2003: 272. See also: Bolton, 1982: 329.
Myrmicinae Formicoxenini Nesomyrmex inye TRUE TRUE FALSEinye. Nesomyrmex inye Mbanyana & Robertson, 2008: 46, figs. 4j-l (w.) SOUTH AFRICA.
Myrmicinae Formicoxenini Nesomyrmex karooensis TRUE TRUE FALSEkarooensis. Nesomyrmex karooensis Mbanyana & Robertson, 2008: 47, figs. 4d-f (w.) SOUTH AFRICA.
Myrmicinae Formicoxenini Nesomyrmex koebergensis TRUE TRUE FALSEkoebergensis. Nesomyrmex koebergensis Mbanyana & Robertson, 2008: 48, figs. 2d-f (w.) SOUTH AFRICA.
Myrmicinae Formicoxenini Nesomyrmex larsenae TRUE TRUE FALSElarsenae. Nesomyrmex larsenae Mbanyana & Robertson, 2008: 48, figs. 2m-o (w.) SOUTH AFRICA.
Myrmicinae Formicoxenini Nesomyrmex mcgregori TRUE TRUE FALSEmcgregori. Nesomyrmex mcgregori Mbanyana & Robertson, 2008: 49, figs. 2j-l (w.) SOUTH AFRICA.
Myrmicinae Formicoxenini Nesomyrmex nanniae TRUE TRUE FALSEnanniae. Nesomyrmex nanniae Mbanyana & Robertson, 2008: 50, figs. 3g-i (w.) SOUTH AFRICA.
Myrmicinae Formicoxenini Nesomyrmex njengelanga TRUE TRUE FALSEnjengelanga. Nesomyrmex njengelanga Mbanyana & Robertson, 2008: 51, figs. 3m-o (w.) SOUTH AFRICA.
Myrmicinae Formicoxenini Nesomyrmex ruani TRUE TRUE FALSEruani. Nesomyrmex ruani Mbanyana & Robertson, 2008: 51, figs. 3a-c (w.) SOUTH AFRICA.
Myrmicinae Formicoxenini Nesomyrmex saasveldensis TRUE TRUE FALSEsaasveldensis. Nesomyrmex saasveldensis Mbanyana & Robertson, 2008: 52, figs. 2a-c (w.) SOUTH AFRICA.
Myrmicinae Formicoxenini Nesomyrmex simoni TRUE TRUE FALSEsimoni. Tetramorium simoni Emery, 1895h: 35, pl. 2, fig. 22 (w.) SOUTH AFRICA. Combination in Tetramyrma: Emery, 1924d: 291; in Leptothorax: Bolton, 1982: 331; in Nesomyrmex: Bolton, 2003: 272. See also: Mbanyana & Robertson, 2008: 53.
Myrmicinae Formicoxenini Nesomyrmex stramineus TRUE TRUE FALSEstramineus. Limnomyrmex stramineus Arnold, 1948: 223, fig. 10 (w.) SOUTH AFRICA. Combination in Leptothorax: Brown, 1971a: 4; in Nesomyrmex: Bolton, 2003: 272. See also: Bolton, 1982: 332; Mbanyana & Robertson, 2008: 38.
Myrmicinae Formicoxenini Nesomyrmex tshiguvhoae TRUE TRUE FALSEtshiguvhoae. Nesomyrmex tshiguvhoae Mbanyana & Robertson, 2008: 53, figs. 4a-c (w.) SOUTH AFRICA.
Myrmicinae Formicoxenini Nesomyrmex vannoorti TRUE TRUE FALSEvannoorti. Nesomyrmex vannoorti Mbanyana & Robertson, 2008: 54, figs. 3j-l (w.) SOUTH AFRICA.
Formicinae Plagiolepidini Nylanderia traegaordhi natalensis TRUE TRUE FALSEnatalensis. Prenolepis (Nylanderia) traegaordhi var. natalensis Forel, 1915c: 348 (w.) SOUTH AFRICA. Combination in Paratrechina (Nylanderia): Emery, 1925b: 218; in Nylanderia: LaPolla, Brady & Shattuck, 2010a: 127.
Myrmicinae Pheidolini Ocymyrmex alacer TRUE TRUE FALSEalacer. Ocymyrmex alacer Bolton & Marsh, 1989: 1290, fig. 9 (w.q.) SOUTH AFRICA.
Myrmicinae Pheidolini Ocymyrmex barbiger TRUE TRUE FALSEbarbiger. Ocymyrmex barbiger Emery, 1886: 364, pl. 17, figs. 9-11 (w.m.) SOUTH AFRICA. [Misspelled as barbatus by Emery, 1892a: 114.] Bolton, 1981b: 261 (q.). See also: Bolton & Marsh, 1989: 1291.
Myrmicinae Pheidolini Ocymyrmex cavatodorsatus TRUE TRUE FALSEcavatodorsatus. Ocymyrmex cavatodorsatus Prins, 1965c: 1021, figs. 1, 2 (w.) SOUTH AFRICA. Bolton & Marsh, 1989: 1278 (q.). See also: Bolton, 1981b: 266.
Myrmicinae Pheidolini Ocymyrmex cilliei TRUE TRUE FALSEcilliei. Ocymyrmex cilliei Prins & Roux, in Bolton & Marsh, 1989: 1292 (m.) SOUTH AFRICA.
Myrmicinae Pheidolini Ocymyrmex gariepensis TRUE TRUE FALSEgariepensis. Ocymyrmex gariepensis Prins & Roux, in Bolton & Marsh, 1989: 1282 (diagnosis in key) (w.) SOUTH AFRICA.
Myrmicinae Pheidolini Ocymyrmex gordoni TRUE TRUE FALSEgordoni. Ocymyrmex gordoni Prins & Roux, in Bolton & Marsh, 1989: 1282 (diagnosis in key) (w.) SOUTH AFRICA.
Myrmicinae Pheidolini Ocymyrmex ignotus TRUE TRUE FALSEignotus. Ocymyrmex ignotus Bolton & Marsh, 1989: 1297 (w.q.) SOUTH AFRICA.
Myrmicinae Pheidolini Ocymyrmex resekhes TRUE TRUE FALSEresekhes. Ocymyrmex resekhes Bolton & Marsh, 1989: 1301, fig. 17 (w.q.) SOUTH AFRICA.
Ponerinae Ponerini Odontoponera denticulata TRUE TRUE FALSEdenticulata. Ponera denticulata Smith, F. 1858b: 90, pl. 6, figs. 13, 14 (q.) SINGAPORE (not South Africa as originally stated; see Donisthorpe, 1943f: 677). Roger, 1861a: 12 (w.). Combination in Odontoponera: Mayr, 1862: 717. Junior synonym of transversa: Dalla Torre, 1893: 30; Forel, 1900d: 314. Revived from synonymy: Yamane, 2009: 5.
Ponerinae Ponerini Pachycondyla aenigmatica TRUE TRUE FALSEaenigmatica. Ponera aenigmatica Arnold, 1949: 265, fig. 5 (w.) SOUTH AFRICA. Combination in Pachycondyla: Brown, in Bolton, 1995b: 302.
Ponerinae Ponerini Pachycondyla berthoudi TRUE TRUE FALSE1 subspecies
berthoudi. Ophthalmopone berthoudi Forel, 1890b: cxiii (w.) SOUTH AFRICA. Forel, 1894b: 76 (m.); Wheeler, G.C. & Wheeler, J. 1971b: 1203 (l.). Combination in Pachycondyla: Brown, in Bolton, 1995b: 303. Current subspecies: nominal plus pubescens. See also: Arnold, 1915: 50; Peeters & Crewe, 1985: 29; Peeters & Crewe, 1987: 201.
Ponerinae Ponerini Pachycondyla caffraria TRUE TRUE FALSE2 subspecies
caffraria. Ponera caffraria Smith, F. 1858b: 91 (q.) SOUTH AFRICA. Forel, 1913a: 108 (w.); Arnold, 1915: 65 (w.m.); Emery, 1899c: 5 (l.); Wheeler, G.C. & Wheeler, J. 1971b: 1206 (l.). Combination in Euponera (Mesoponera): Emery, 1901a: 46; in Mesoponera: Wheeler, G.C. & Wheeler, J. 1971b: 1206; in Pachycondyla: Brown, in Bolton, 1995b: 303. Senior synonym of guineensis: Mayr, 1895: 125; Emery, 1911d: 81; Wheeler, W.M. 1922a: 775. Current subspecies: nominal plus affinis, caffra.
Ponerinae Ponerini Pachycondyla cavernosa TRUE TRUE FALSE1 subspecies
cavernosa. Ponera cavernosa Roger, 1860: 288 (w.) SOUTH AFRICA. Combination in Bothroponera: Mayr, 1862: 717; Wheeler, W.M. 1922a: 72; in Pachycondyla (Bothroponera): Emery, 1901a: 45; in Pachycondyla: Brown, in Bolton, 1995b: 304. Current subspecies: nominal plus montivaga.
Ponerinae Ponerini Pachycondyla darwinii TRUE TRUE FALSE3 subspecies
darwinii. Belonopelta darwinii Forel, 1893d: 460 (q.) AUSTRALIA. Terayama, 1990: 897 (w.). Combination in Pachycondyla (Pseudoponera): Emery, 1900d: 668; Emery, 1901a: 46; in Pseudoponera: Bingham, 1903: 93; in Euponera (Trachymesopus): Emery, 1911d: 86; in Trachymesopus: Wilson, 1958d: 353; in Pachycondyla: Brown, in Bolton, 1995b: 304. Senior synonym of lamarki, rufotestaceus: Brown, 1963: 6. Current subspecies: nominal plus africana, indica, madecassa.
Ponerinae Ponerini Pachycondyla fossigera TRUE TRUE FALSEfossigera. Euponera fossigera Mayr, 1901b: 3 (w.) SOUTH AFRICA. Combination in E. (Mesoponera): Arnold, 1915: 71; in Pachycondyla: Brown, in Bolton, 1995b: 305.
Ponerinae Ponerini Pachycondyla granosa TRUE TRUE FALSEgranosa. Ponera granosa Roger, 1860: 290 (w.) SOUTH AFRICA. Arnold, 1926: 201 (m.). Combination in Bothroponera: Mayr, 1862: 717; in Pachycondyla (Bothroponera): Emery, 1901a: 45; in Bothroponera: Wheeler, W.M. 1922a: 72; in Pachycondyla: Brown, in Bolton, 1995b: 305.
Ponerinae Ponerini Pachycondyla hartwigi TRUE TRUE FALSEhartwigi. Cryptopone hartwigi Arnold, 1948: 213, fig. 2 (w.) SOUTH AFRICA. Combination in Hypoponera: Taylor, 1967a: 12; in Cryptopone: Bolton, 1995b: 166; in Pachycondyla: Mackay & Mackay, 2010: 3 (by implication as Cryptopone synonymised with Pachycondyla).
Ponerinae Ponerini Pachycondyla havilandi TRUE TRUE FALSE3 subspecies
havilandi. Megaloponera (Hagensia) havilandi Forel, 1901f: 333 (w.) SOUTH AFRICA. Arnold, 1915: 68 (m.). Combination in Euponera (Mesoponera): Arnold, 1915: 67; in Hagensia: Arnold, 1926: 203; in Pachycondyla: Brown, in Bolton, 1995b: 305. Senior synonym of sulcigera: Arnold, 1915: 67. Current subspecies: nominal plus fochi, godfreyi, marleyi.
Ponerinae Ponerini Pachycondyla hottentota TRUE TRUE FALSEhottentota. Pachycondyla hottentota Emery, 1886: 360 (w.) SOUTH AFRICA. Arnold, 1915: 52 (m.). Combination in Ophthalmopone: Emery, 1902c: 30; in Pachycondyla: Brown, in Bolton, 1995b: 306. Senior synonym of lanceolata: Emery, 1902c: 30.
Ponerinae Ponerini Pachycondyla kruegeri TRUE TRUE FALSE2 subspecies
kruegeri. Pachycondyla (Bothroponera) kruegeri Forel, 1910a: 28 (w.) SOUTH AFRICA. Combination in Bothroponera: Wheeler, W.M. 1922a: 22; in Pachycondyla: Brown, in Bolton, 1995b: 306. Current subspecies: nominal plus asina, rhodesiana. See also: Arnold, 1915: 56.
Ponerinae Ponerini Pachycondyla laevissima TRUE TRUE FALSE1 subspecies
laevissima. Pachycondyla (Bothroponera) laevissima Arnold, 1915: 58, pl. 2, fig. 15 (w.) SOUTH AFRICA. Combination in Bothroponera: Wheeler, W.M. 1922a: 73; in Pachycondyla: Brown, in Bolton, 1995b: 306. Current subspecies: nominal plus aspera.
Ponerinae Ponerini Pachycondyla peringueyi TRUE TRUE FALSE1 subspecies
peringueyi. Ponera peringueyi Emery, 1899e: 474 (w.) SOUTH AFRICA. Arnold, 1951: 53 (m.); Wheeler, G.C. & Wheeler, J. 1971b: 1203 (l.). Combination in Euponera (Brachyponera): Emery, 1901a: 47; in E. (Mesoponera): Emery, 1911d: 81; in Hagensia: Arnold, 1926: 202; in Pachycondyla: Brown, in Bolton, 1995b: 308. Current subspecies: nominal plus saldanhae. See also: Arnold, 1915: 66.
Ponerinae Ponerini Pachycondyla pumicosa TRUE TRUE FALSE1 subspecies
pumicosa. Ponera pumicosa Roger, 1860: 290 (w.) SOUTH AFRICA. Forel, 1901f: 344 (m.); Wheeler, G.C. & Wheeler, J. 1971c: 390 (l.). Combination in Bothroponera: Mayr, 1862: 717; in Pachycondyla (Bothroponera): Emery, 1901a: 45; in Bothroponera: Wheeler, W.M. 1922a: 72; in Pachycondyla: Brown, in Bolton, 1995b: 308. [Misspelled as pumicata by Raffray, 1887: 21.] Current subspecies: nominal plus sculpturata. See also: Arnold, 1915: 62.
Ponerinae Ponerini Pachycondyla strigulosa TRUE TRUE FALSEstrigulosa. Bothroponera strigulosa Emery, 1895h: 19 (w.) SOUTH AFRICA. Combination in Pachycondyla (Bothroponera): Emery, 1901a: 45; in Bothroponera: Wheeler, W.M. 1922a: 72; in Pachycondyla: Brown, in Bolton, 1995b: 310. Senior synonym of berthoudi: Arnold, 1952a: 460.
Ponerinae Ponerini Pachycondyla variolosa TRUE TRUE FALSEvariolosa. Bothroponera variolosa Arnold, 1947: 131, fig. 1 (w.) SOUTH AFRICA. Combination in Pachycondyla: Brown, in Bolton, 1995b: 311.
Ponerinae Ponerini Pachycondyla wroughtonii TRUE TRUE FALSE1 subspecies
wroughtonii. Euponera (Pseudoponera) wroughtonii Forel, 1901f: 341 (w.m.) SOUTH AFRICA. Arnold, 1926: 204 (q.); Wheeler, G.C. & Wheeler, J. 1971b: 1206 (l.). Combination in E. (Trachymesopus): Emery, 1911d: 85; in Mesoponera: Wheeler, G.C. & Wheeler, J. 1971b: 1206; in Pachycondyla: Brown, in Bolton, 1995b: 311. Current subspecies: nominal plus crudelis.
Ponerinae Ponerini Pachycondyla peringueyi saldanhae TRUE TRUE FALSEsaldanhae. Hagensia peringueyi r. saldanhae Arnold, 1951: 53, fig. 4 (w.) SOUTH AFRICA. Combination in Pachycondyla: Brown, in Bolton, 1995b: 309.
Ponerinae Ponerini Pachycondyla elisae rotundata TRUE TRUE FALSErotundata. Ponera elisae var. rotundata Emery, 1895h: 20 (w.) SOUTH AFRICA. Arnold, 1915: 71 (q.). Combination in Euponera (Mesoponera): Emery, 1901a: 46; in E. (Xiphopelta): Forel, 1913j: 206; in Pachycondyla: Brown, in Bolton, 1995b: 309. Senior synonym of arnoldi: Wheeler, W.M. 1922a: 776. Currently subspecies of elisae: Santschi, 1937d: 217.
Ponerinae Ponerini Pachycondyla cavernosa montivaga TRUE TRUE FALSEmontivaga. Bothroponera cavernosa var. montivaga Arnold, 1947: 132 (w.) SOUTH AFRICA. Combination in Pachycondyla: Brown, in Bolton, 1995b: 307.
Ponerinae Ponerini Pachycondyla havilandi marleyi TRUE TRUE FALSEmarleyi. Hagensia havilandi r. marleyi Arnold, 1926: 203 (w.) SOUTH AFRICA. Combination in Pachycondyla: Brown, in Bolton, 1995b: 307.
Ponerinae Ponerini Pachycondyla havilandi godfreyi TRUE TRUE FALSEgodfreyi. Hagensia havilandi r. godfreyi Arnold, 1926: 203 (w.m.) SOUTH AFRICA. Combination in Pachycondyla: Brown, in Bolton, 1995b: 305.
Ponerinae Ponerini Pachycondyla havilandi fochi TRUE TRUE FALSEfochi. Euponera (Hagensia) havilandi var. fochi Forel, 1918b: 155 (w.) SOUTH AFRICA. Combination in Pachycondyla: Brown, in Bolton, 1995b: 305.
Ponerinae Ponerini Pachycondyla wroughtonii crudelis TRUE TRUE FALSEcrudelis. Euponera (Pseudoponera) wroughtonii var. crudelis Forel, 1901f: 343 (w.m.) SOUTH AFRICA. Combination in E. (Trachymesopus): Emery, 1911d: 85; in Pachycondyla: Brown, in Bolton, 1995b: 304.
Ponerinae Ponerini Pachycondyla laevissima aspera TRUE TRUE FALSEaspera. Bothroponera laevissima var. aspera Arnold, 1962: 844 (w.) SOUTH AFRICA. Combination in Pachycondyla: Brown, in Bolton, 1995b: 303.
Ponerinae Ponerini Pachycondyla darwinii africana TRUE TRUE FALSEafricana. Euponera (Pseudoponera) darwini var. africana Forel, 1909b: 51 (q.) DEMOCRATIC REPUBLIC OF CONGO. Santschi, 1914b: 51 (m.). Combination in Euponera (Trachymesopus): Emery, 1911d: 86; in Pachycondyla: Brown, in Bolton, 1995b: 302. Currently subspecies of darwinii: Santschi, 1937b: 95.
Myrmicinae Pheidolini Pheidole akermani TRUE TRUE FALSEakermani. Pheidole akermani Arnold, 1920a: 471 (s.w.) SOUTH AFRICA.
Myrmicinae Pheidolini Pheidole aspera TRUE TRUE FALSEaspera. Pheidole aspera Mayr, 1862: 746 (s.w.) SOUTH AFRICA. See also: Mayr, 1865: 97.
Myrmicinae Pheidolini Pheidole caffra TRUE TRUE FALSE5 subspecies
caffra. Pheidole caffra Emery, 1895h: 33 (s.) SOUTH AFRICA. Santschi, 1939b: 239 (w.). Current subspecies: nominal plus abyssinica, amoena, bayeri, montivaga, senilifrons.
Myrmicinae Pheidolini Pheidole capensis TRUE TRUE FALSE2 subspecies
capensis. Pheidole capensis Mayr, 1862: 746 (s.w.) SOUTH AFRICA. Forel, 1913a: 130 (q.). Subspecies of megacephala: Emery, 1895h: 34. Revived status as species: Forel, 1913a: 130; Arnold, 1920a: 441. Current subspecies: nominal plus dregei, modestior, reddenburgensis.
Myrmicinae Pheidolini Pheidole crassinoda TRUE TRUE FALSE3 subspecies
crassinoda. Pheidole crassinoda Emery, 1895h: 32 (s.w.) SOUTH AFRICA. Mayr, 1907b: 12 (q.m.). [P. crassinoda Forel, 1894b: 94. Nomen nudum, attributed to Emery.] Current subspecies: nominal plus pluto, ruspolii, sordidula.
Myrmicinae Pheidolini Pheidole foreli TRUE TRUE FALSE2 subspecies
foreli. Pheidole foreli Mayr, 1901b: 8 (s.w.) SOUTH AFRICA. Forel, 1910f: 12 (q.). Current subspecies: nominal plus pubens. See also: Arnold, 1920a: 433.
Myrmicinae Pheidolini Pheidole hewitti TRUE TRUE FALSEhewitti. Pheidole hewitti Santschi, 1932a: 382, figs. 1-4 (s.w.) SOUTH AFRICA. Senior synonym of hewitti Arnold: Arnold, 1952a: 462.
Myrmicinae Pheidolini Pheidole irritans TRUE TRUE FALSEirritans. Myrmica irritans Smith, F. 1858b: 127 (w.) SOUTH AFRICA. Combination in Pheidole: Dixey & Longstaff, 1907: 331.
Myrmicinae Pheidolini Pheidole kitschneri TRUE TRUE FALSEkitschneri. Pheidole kitschneri Forel, 1910e: 436 (s.w.) SOUTH AFRICA. Santschi, 1914b: 76 (q.).
Myrmicinae Pheidolini Pheidole punctulata TRUE TRUE FALSE3 subspecies
punctulata. Pheidole punctulata Mayr, 1866b: 899 (s.) SOUTH AFRICA. Arnold, 1920a: 421 (w.q.m.). Subspecies of megacephala: Forel, 1894b: 94; Stitz, 1911b: 385; Emery, 1915j: 235; Emery, 1921f: 85; Wheeler, W.M. 1922a: 132; Santschi, 1930b: 67. Status as species: Forel, 1910e: 441; Santschi, 1914b: 76; Forel, 1914d: 240; Arnold, 1920a: 420; Santschi, 1935a: 256. Current subspecies: nominal plus atrox, spinosa, subatrox.
Myrmicinae Pheidolini Pheidole sculpturata TRUE TRUE FALSE7 subspecies
sculpturata. Pheidole sculpturata Mayr, 1866b: 897 (s.) SOUTH AFRICA. Emery, 1895h: 31 (w.). Current subspecies: nominal plus areolata, berthoudi, dignata, particeps, rhodesiana, welgelegenensis, zambesiana. See also: Arnold, 1920a: 462.
Myrmicinae Pheidolini Pheidole tenuinodis TRUE TRUE FALSE2 subspecies
tenuinodis. Pheidole tenuinodis Mayr, 1901b: 10 (s.w.) SOUTH AFRICA. Arnold, 1920a: 430 (q.). Current subspecies: nominal plus bothae, robusta, sipapomae.
Myrmicinae Pheidolini Pheidole capensis dregei TRUE TRUE FALSEdregei. Pheidole megacephala subsp. dregei Emery, 1895h: 34 (s.) SOUTH AFRICA. Currently subspecies of capensis: Emery, 1915j: 246.
Myrmicinae Pheidolini Pheidole spinulosa conigera TRUE TRUE FALSEconigera. Pheidole spinulosa subsp. conigera Forel, 1910e: 440 (s.) SOUTH AFRICA.
Myrmicinae Pheidolini Pheidole foreli bothae TRUE TRUE FALSEbothae. Pheidole foreli r. bothae Forel, 1901h: 62 (s.w.) SOUTH AFRICA. Santschi, 1914e: 22 (q.). Subspecies of tenuinodis: Santschi, 1914e: 22. Junior synonym of tenuinodis: Emery, 1921f: 87. Revived from synonymy as subspecies of tenuinodis: Wheeler, W.M. 1922a: 822.
Myrmicinae Pheidolini Pheidole sculpturata berthoudi TRUE TRUE FALSEberthoudi. Pheidole sculpturata r. berthoudi Forel, 1894b: 89 (s.w.) SOUTH AFRICA. Raised to species: Santschi, 1921c: 116. Currently subspecies of sculpturata: Emery, 1921f: 89; Wheeler, W.M. 1922a: 819.
Myrmicinae Pheidolini Pheidole sculpturata areolata TRUE TRUE FALSEareolata. Pheidole sculpturata var. areolata Forel, 1911e: 269 (s.w.) SOUTH AFRICA.
Myrmicinae Pheidolini Pheidole capensis modestior TRUE TRUE FALSEmodestior. Pheidole capensis var. modestior Santschi, 1919a: 337 (s.w.) SOUTH AFRICA.
Myrmicinae Pheidolini Pheidole foreli pubens TRUE TRUE FALSEpubens. Pheidole foreli var. pubens Forel, 1910f: 12 (s.w.) SOUTH AFRICA. Forel, 1910e: 436 (q.m.).
Myrmicinae Pheidolini Pheidole cuitensis reddenburgensis TRUE TRUE FALSEreddenburgensis. Pheidole (Allopheidole) cuitensis r. reddenburgensis Forel, 1913a: 135 (s.w.q.m.) SOUTH AFRICA. Subspecies of capensis: Emery, 1915j: 246; Wheeler, W.M. 1922a: 809. [Spelling emended to reddersbergensis by Arnold, 1920a: 450; emendation not accepted by Wheeler, W.M. 1922a: 809.]
Myrmicinae Solenopsidini Pheidologeton hostilis TRUE TRUE FALSEhostilis. Atta hostilis Smith, F. 1858b: 165 (w.q.) SOUTH AFRICA. Combination in Pheidologeton: Mayr, 1866b: 899.
Formicinae Plagiolepidini Plagiolepis capensis TRUE TRUE FALSEcapensis. Plagiolepis capensis Mayr, 1865: 55 (q.) SOUTH AFRICA.
Formicinae Plagiolepidini Plagiolepis decora TRUE TRUE FALSEdecora. Plagiolepis (Anacantholepis) decora Santschi, 1914e: 36, fig. 9 (w.) SOUTH AFRICA. Arnold, 1922: 582 (q.).
Formicinae Plagiolepidini Plagiolepis deweti TRUE TRUE FALSEdeweti. Plagiolepis deweti Forel, 1904f: 423 (w.q.) SOUTH AFRICA. Combination in P. (Anacantholepis): Forel, 1914d: 249.
Formicinae Plagiolepidini Plagiolepis funicularis TRUE TRUE FALSEfunicularis. Plagiolepis funicularis Santschi, 1919a: 344 (w.) SOUTH AFRICA.
Formicinae Plagiolepidini Plagiolepis fuscula TRUE TRUE FALSEfuscula. Plagiolepis fuscula Emery, 1895h: 45 (w.) SOUTH AFRICA. Combination in P. (Anacantholepis): Emery, 1925b: 22.
Formicinae Plagiolepidini Plagiolepis intermedia TRUE TRUE FALSE1 subspecies
intermedia. Plagiolepis pygmaea var. intermedia Emery, 1895h: 45 (w.q.) SOUTH AFRICA. Raised to species: Emery, 1921d: 316. Current subspecies: nominal plus minutula.
Formicinae Plagiolepidini Plagiolepis jouberti TRUE TRUE FALSEjouberti. Plagiolepis jouberti Forel, 1910f: 23 (w.) SOUTH AFRICA. Combination in P. (Anacantholepis): Santschi, 1914b: 122.
Formicinae Plagiolepidini Plagiolepis puncta TRUE TRUE FALSEpuncta. Plagiolepis pygmaea var. punctum Forel, 1910e: 448 (w.m.) SOUTH AFRICA. Arnold, 1922: 602 (q.). Subspecies of pygmaea: Arnold, 1922: 602; Wheeler, W.M. 1922a: 930. Raised to species: Emery, 1921d: 317; Emery, 1925b: 21.
Formicinae Plagiolepidini Plagiolepis vanderkelleni polita TRUE TRUE FALSEpolita. Plagiolepis (Anacantholepis) vanderkeleni st. polita Santschi, 1914e: 37, fig. 10 (w.) SOUTH AFRICA.
Formicinae Plagiolepidini Plagiolepis intermedia minutula TRUE TRUE FALSEminutula. Plagiolepis intermedia var. minutula Emery, 1921d: 316, fig. 2 (w.q.) SOUTH AFRICA.
Ponerinae Platythyreini Platythyrea cooperi TRUE TRUE FALSEcooperi. Platythyrea cooperi Arnold, 1915: 29 (w.m.) SOUTH AFRICA.
Ponerinae Platythyreini Platythyrea lamellosa TRUE TRUE FALSElamellosa. Ponera lamellosa Roger, 1860: 295 (w.) SOUTH AFRICA. Wheeler, G.C. & Wheeler, J. 1989b: 51 (l.). Combination in Platythyrea: Roger, 1863a: 173. Senior synonym of grisea: Forel, 1913b: 307; of longinoda, rhodesiana, suturalis: Brown, 1975: 8.
Ponerinae Ponerini Plectroctena mandibularis TRUE TRUE FALSEmandibularis. Plectroctena mandibularis Smith, F. 1858b: 101, pl. 7, figs. 1-5 (m., ergatoid q., not w.) SOUTH AFRICA. Forel, 1913a: 108 (w.); Arnold, 1915: 86 (w.). Material of the nomen nudum caffra referred here: Roger, 1861a: 41. Senior synonym of major: Emery, 1899e: 469; Bolton, 1974c: 330; of integra and material of the unavailable name strialiventris referred here: Bolton, 1974c: 330; of conjugata: Villet, McKitterick & Robertson, 1999: 282. .
Ponerinae Ponerini Plectroctena strigosa TRUE TRUE FALSEstrigosa. Plectroctena mandibularis var. strigosa Emery, 1899e: 469 (w.) SOUTH AFRICA. Subspecies of cristata: Santschi, 1924a: 161. Raised to species: Bolton, 1974c: 332.
Formicinae Camponotini Polyrhachis arnoldi TRUE TRUE FALSEarnoldi. Polyrhachis (Myrma) arnoldi Forel, 1914d: 263 (w.) SOUTH AFRICA. See also: Arnold, 1924: 750; Bolton, 1973b: 324.
Formicinae Camponotini Polyrhachis cubaensis TRUE TRUE FALSEcubaensis. Polyrhachis cubaensis Mayr, 1862: 687 (w.) SOUTH AFRICA (not Cuba; see Mayr, 1893: 195). Combination in P. (Myrma): Wheeler, W.M. 1922a: 994. Senior synonym of gerstaeckeri, striolatorugosa, wilmsi: Bolton, 1973b: 325.
Formicinae Camponotini Polyrhachis durbanensis TRUE TRUE FALSEdurbanensis. Polyrhachis (Myrma) cubaensis r. durbanensis Forel 1914d: 262 (w.q.) SOUTH AFRICA. Raised to species: Bolton, 1973b: 327.
Formicinae Camponotini Polyrhachis gagates TRUE TRUE FALSEgagates. Polyrhachis gagates Smith, F. 1858b: 71, pl. 4, fig. 14 (w.) SOUTH AFRICA. Forel, 1886f: 194 (q.); Wheeler, G.C. & Wheeler, J. 1953e: 207 (l.). Combination in P. (Myrma): Santschi, 1914b: 139. Subspecies of schistacea: Arnold, 1924: 745. Revived status as species: Emery, 1925b: 199; Bolton, 1973b: 305. Senior synonym of clariseta, congolensis (and its junior synonym indefinita), nigriseta, obsidiana and material of the unavailable name acheron referred here: Bolton, 1973b: 305.
Formicinae Camponotini Polyrhachis gamaii TRUE TRUE FALSEgamaii. Polyrhachis (Myrma) gamaii Santschi, 1917b: 295 (q.) SOUTH AFRICA. Arnold, 1947: 136 (w.). See also: Bolton, 1973b: 351.
Formicinae Camponotini Polyrhachis militaris TRUE TRUE FALSEmilitaris. Formica militaris Fabricius, 1782: 493 (q.) "Tropical Africa". Smith, F. 1858b: 72 (m.); Mayr, 1866b: 886 (w.). Combination in Myrma: Billberg, 1820: 104; in Polyrhachis: Smith, F. 1858b: 72; in P. (Myrma): Santschi, 1914b: 140. Senior synonym of bruta, calabarica, cupreopubescens, ssibangensis, striativentris: Bolton, 1973b: 313; of epinotalis, nkomoensis: Dorow, 1995: 36, 37 respectively. Material of the unavailable names argentatus, dido, pleurata, sankisiana, transversaria referred here by Bolton, 1973b: 313.
Formicinae Camponotini Polyrhachis phidias TRUE TRUE FALSEphidias. Polyrhachis phidias Forel, 1910e: 450 (w.) "Equatorial Africa". Combination in P. (Myrma): Wheeler, W.M. 1922a: 999. See also: Bolton, 1973b: 316.
Formicinae Camponotini Polyrhachis schlueteri TRUE TRUE FALSEschlueteri. Polyrhachis militaris r. schlueteri Forel, 1886f: 195 (w.) EAST AFRICA. Arnold, 1924: 748 (q.). Combination in P. (Myrma): Santschi, 1914b: 143. Raised to species: Dalla Torre, 1893: 268. Subspecies of schistacea: Santschi, 1914e: 42; Wheeler, W.M. 1922a: 1003; Arnold, 1924: 747. Status as species: Forel, 1907g: 92; Forel, 1914d: 261; Santschi, 1914b: 143; Emery, 1925b: 200; Bolton, 1973b: 321. Senior synonym of indigens: Arnold, 1924: 747; of plebeia: Bolton, 1973b: 321.
Formicinae Camponotini Polyrhachis viscosa TRUE TRUE FALSEviscosa. Polyrhachis viscosus Smith, F. 1858b: 71, pl. 4, fig. 41 (w.) SOUTH AFRICA. Combination in P. (Myrma): Santschi, 1914b: 143. Senior synonym of antinorii: Dalla Torre, 1893: 271; of imatongica, spretula: Bolton, 1973b: 330. See also: Arnold, 1924: 749.
Amblyoponinae Amblyoponini Prionopelta aethiopica TRUE TRUE FALSEaethiopica. Prionopelta descarpentriesi subsp. aethiopica Arnold, 1949: 263, fig. 4 (w.) SOUTH AFRICA. Raised to species: Brown, 1960a: 177.
Myrmicinae Myrmecinini Pristomyrmex africanus TRUE TRUE FALSEafricanus. Pristomyrmex africanus Karavaiev, 1931d: 47, fig. 5 (w.) KENYA. Wang, M. 2003: 424 (q. and ergatoid q.). Combination in Hylidris: Weber, 1952: 17; in Pristomyrmex: Bolton, 1981b: 284. Senior synonym of beni, mbomu, myersi, primus: Bolton, 1981b: 284.
Myrmicinae Myrmecinini Pristomyrmex fossulatus TRUE TRUE FALSEfossulatus. Tetramorium (Xiphomyrmex) fossulatum Forel, 1910e: 428 (w.) SOUTH AFRICA. Menozzi, 1942: 172 (q.). Combination in Pristomyrmex: Santschi, 1916a: 51; in Hylidris: Weber, 1952: 17; in Pristomyrmex: Bolton, 1981b: 285. See also: Wang, M. 2003: 406.
Proceratiinae Probolomyrmecini Probolomyrmex filiformis TRUE TRUE FALSEfiliformis. Probolomyrmex filiformis Mayr, 1901b: 3 (w.) SOUTH AFRICA. See also: Taylor, 1965d: 353.
Myrmicinae Dacetini Pyramica africana TRUE TRUE FALSEafricana. Glamyromyrmex africanus Bolton, 1983: 322, fig. 27 (w.) GABON. Combination in Pyramica: Bolton, 1999: 1672. See also: Bolton, 2000: 329.
Myrmicinae Dacetini Pyramica agnosta TRUE TRUE FALSEagnosta. Pyramica agnosta Bolton, 2000: 293, figs. 199, 227 (w.) SOUTH AFRICA.
Myrmicinae Dacetini Pyramica anarta TRUE TRUE FALSEanarta. Smithistruma anarta Bolton, 1983: 314, fig. 18 (w.) SOUTH AFRICA. Combination in Pyramica: Bolton, 1999: 1673. See also: Bolton, 2000: 323.
Myrmicinae Dacetini Pyramica dictynna TRUE TRUE FALSEdictynna. Pyramica dictynna Bolton, 2000: 294 (w.) SOUTH AFRICA.
Myrmicinae Dacetini Pyramica emarginata TRUE TRUE FALSEemarginata. Strumigenys emarginata Mayr, 1901b: 26 (w.) SOUTH AFRICA. Combination in S. (Trichoscapa): Santschi, 1913b: 257; in S. (Cephaloxys): Wheeler, W.M. 1922a: 919; Emery, 1924d: 324; in Smithistruma: Brown, 1948e: 105; in Pyramica: Bolton, 1999: 1673. See also: Brown, 1953g: 126; Bolton, 1983: 291; Bolton, 2000: 301.
Myrmicinae Dacetini Pyramica mira TRUE TRUE FALSEmira. Pyramica mira Bolton, 2000: 324 (w.) SOUTH AFRICA.
Myrmicinae Dacetini Pyramica noara TRUE TRUE FALSEnoara. Pyramica noara Bolton, 2000: 304 (w.) SOUTH AFRICA.
Myrmicinae Dacetini Pyramica ogyga TRUE TRUE FALSEogyga. Pyramica ogyga Bolton, 2000: 324, fig. 213 (w.m.) SOUTH AFRICA.
Myrmicinae Dacetini Pyramica oxysma TRUE TRUE FALSEoxysma. Smithistruma oxysma Bolton, 1983: 315, fig. 19 (w.q.) SOUTH AFRICA. Combination in Pyramica: Bolton, 1999: 1673. See also: Bolton, 2000: 325.
Myrmicinae Dacetini Pyramica sardonica TRUE TRUE FALSEsardonica. Pyramica sardonica Bolton, 2000: 326 (w.) SOUTH AFRICA.
Myrmicinae Dacetini Pyramica simoni TRUE TRUE FALSEsimoni. Strumigenys simoni Emery, 1895h: 42, pl. 2, fig. 21 (w.) SOUTH AFRICA. Combination in S. (Trichoscapa): Santschi, 1913b: 258 (in key); in S. (Cephaloxys): Wheeler, W.M. 1922a: 920; Emery, 1924d: 324; in Smithistruma (Serrastruma): Brown, 1948e: 107; in Serrastruma: Brown, 1952e: 82; in Pyramica: Bolton, 1999: 1673. Senior synonym of biconvexa, boerorum, cliens, cognata, escherichi, fusciventris, limbata and material of the unavailable name obscuriventris referred here: Brown, 1952e: 82; of raymondi: Bolton, 1983: 350. See also: Bolton, 2000: 318.
Myrmicinae Dacetini Pyramica transversa TRUE TRUE FALSEtransversa. Strumigenys (Trichoscapa) transversa Santschi, 1913b: 258 (diagnosis in key) (w.) SOUTH AFRICA. Santschi, 1914e: 31 (w.). Combination in S. (Cephaloxys): Wheeler, W.M. 1922a: 920; Emery, 1924d: 324; in Smithistruma: Brown, 1948e: 105; in Pyramica: Bolton, 1999: 1673. See also: Brown, 1953g: 127; Bolton, 1983: 297; Bolton, 2000: 334.
Myrmicinae Tetramoriini Rhoptromyrmex globulinodis TRUE TRUE FALSEglobulinodis. Rhoptromyrmex globulinodis Mayr, 1901b: 20 (w.q.m.) SOUTH AFRICA. Senior synonym of alberti, obscurus: Brown, 1964c: 17. See also: Bolton, 1986a: 13.
Myrmicinae Tetramoriini Rhoptromyrmex transversinodis TRUE TRUE FALSEtransversinodis. Rhoptromyrmex transversinodis Mayr, 1901b: 22 (w.) SOUTH AFRICA. Arnold, 1917: 356 (q.); Bolton, 1986a: 16 (m.). Senior synonym of pretoriae, steini: Brown, 1964c: 16.
Cerapachyinae Cerapachyini Simopone marleyi TRUE TRUE FALSEmarleyi. Simopone marleyi Arnold, 1915: 20 (w.) SOUTH AFRICA.
Myrmicinae Solenopsidini Solenopsis africana TRUE TRUE FALSEafricana. Solenopsis fugax st. africana Santschi, 1914b: 81 (w.m.) KENYA. Raised to species: Emery, 1915c: 59 (in text); Emery, 1922e: 201.
Myrmicinae Solenopsidini Solenopsis capensis TRUE TRUE FALSEcapensis. Solenopsis capensis Mayr, 1866b: 905 (q.) SOUTH AFRICA.
Myrmicinae Solenopsidini Solenopsis punctaticeps TRUE TRUE FALSE7 subspecies
punctaticeps. Solenopsis punctaticeps Mayr, 1865: 109 (w.) SOUTH AFRICA. Forel, 1910e: 431 (q.m.). Current subspecies: nominal plus caffra, cleptomana, erythraea, fur, indocilis, juba, kibaliensis. See also: Arnold, 1916: 243.
Myrmicinae Solenopsidini Solenopsis punctaticeps caffra TRUE TRUE FALSEcaffra. Solenopsis punctaticeps var. caffra Forel, 1894b: 99 (w.q.) SOUTH AFRICA. Senior synonym of cyclops, diversipilosa: Emery, 1915c: 65.
Cerapachyinae Cerapachyini Sphinctomyrmex chariensis TRUE TRUE FALSEchariensis. Sphinctomyrmex rufiventris st. chariensis Santschi, 1915c: 245 (m.) CENTRAL AFRICAN REPUBLIC. Combination in Aethiopopone: Santschi, 1930a: 50; in Sphinctomyrmex: Brown, 1975: 33. Raised to species: Brown, 1975: 33.
Myrmicinae Stenammini Stenamma msilanum TRUE TRUE FALSEmsilanum. Stenamma (Stenamma) westwoodi var. msilanum Forel, 1901e: 347 (q.) ALGERIA. Raised to species and senior synonym of africanum, submuticum: DuBois, 1998b: 254.
Ponerinae Ponerini Streblognathus aethiopicus TRUE TRUE FALSEaethiopicus. Ponera aethiopica Smith, F. 1858b: 91, pl. 6, figs. 19, 20 (w.) SOUTH AFRICA. Emery, 1899e: 468 (m.); Wheeler, G.C. & Wheeler, J. 1989b: 53 (l.). Combination in Streblognathus: Mayr, 1862: 716. See also: Arnold, 1915: 41; Robertson, 2002: 10.
Ponerinae Ponerini Streblognathus peetersi TRUE TRUE FALSEpeetersi. Streblognathus peetersi Robertson, 2002: 11, figs. 1, 2, 3b, 3d, 4 (w.m.) SOUTH AFRICA.
Myrmicinae Dacetini Strumigenys faurei TRUE TRUE FALSEfaurei. Strumigenys faurei Arnold, 1948: 226, fig. 12 (w.) SOUTH AFRICA. Junior synonym of rufobrunea: Brown, 1954k: 17. Revived from synonymy: Bolton, 1983: 371. See also: Bolton, 2000: 602.
Myrmicinae Dacetini Strumigenys havilandi TRUE TRUE FALSEhavilandi. Strumigenys havilandi Forel, 1905c: 13 (footnote) (w.q.) SOUTH AFRICA. See also: Brown, 1954k: 25; Bolton, 1983: 373; Bolton, 2000: 592.
Myrmicinae Dacetini Strumigenys irrorata TRUE TRUE FALSEirrorata. Strumigenys (Strumigenys) irrorata Santschi, 1913b: 257 (diagnosis in key) (w.) SOUTH AFRICA. Santschi, 1914e: 29 (w.). See also: Brown, 1954k: 33; Bolton, 1983: 375; Bolton, 2000: 593.
Myrmicinae Dacetini Strumigenys marleyi TRUE TRUE FALSEmarleyi. Strumigenys havilandi r. marleyi Arnold, 1914: 31, fig. 10 (w.) SOUTH AFRICA. Raised to species: Arnold, 1926: 286. See also: Brown, 1954k: 25; Bolton, 1983: 378; Bolton, 2000: 599.
Myrmicinae Dacetini Strumigenys pretoriae TRUE TRUE FALSEpretoriae. Strumigenys pretoriae Arnold, 1949: 267, fig. 8 (w.) SOUTH AFRICA. See also: Brown, 1954k: 15; Bolton, 1983: 385; Bolton, 2000: 603.
Myrmicinae Dacetini Strumigenys rakkota TRUE TRUE FALSErakkota. Strumigenys rakkota Bolton, 2000: 595 (w.) SOUTH AFRICA.
Myrmicinae Dacetini Strumigenys syntacta TRUE TRUE FALSEsyntacta. Strumigenys syntacta Bolton, 2000: 597, fig. 364 (w.) SOUTH AFRICA.
Myrmicinae Dacetini Strumigenys traegaordhi TRUE TRUE FALSEtraegaordhi. Strumigenys (Strumigenys) traegaordhi Santschi, 1913b: 257 (diagnosis in key) (w.) SOUTH AFRICA. Santschi, 1914e: 28 (w.); Emery, 1915g: 19 (q.). See also: Brown, 1954k: 27; Bolton, 1983: 396; Bolton, 2000: 598.
Formicinae Plagiolepidini Tapinolepis candida TRUE TRUE FALSEcandida. Anoplolepis (Tapinolepis) candida Santschi, 1928f: 210 (w.) SOUTH AFRICA. Combination in Tapinolepis: Bolton, 2003: 268.
Formicinae Plagiolepidini Tapinolepis decolor TRUE TRUE FALSEdecolor. Plagiolepis decolor Emery, 1895h: 44 (w.) SOUTH AFRICA. Forel, 1910f: 23 (q.m.). Combination in Plagiolepis (Anoplolepis): Arnold, 1922: 599; in Anoplolepis: Emery, 1925b: 17; in Anoplolepis (Mesanoplolepis): Santschi, 1926a: 14; in Tapinolepis: Bolton, 2003: 268.
Formicinae Plagiolepidini Tapinolepis trimenii TRUE TRUE FALSE2 subspecies
trimenii. Plagiolepis trimenii Forel, 1895g: 430 (w.) SOUTH AFRICA. Arnold, 1922: 596 (q.). Combination in Plagiolepis (Anoplolepis): Arnold, 1922: 595; in Anoplolepis: Emery, 1925b: 18; in Anoplolepis (Mesanoplolepis): Santschi, 1930b: 72; in Tapinolepis: Bolton, 2003: 268. Current subspecies: nominal plus angolensis, karrooensis.
Formicinae Plagiolepidini Tapinolepis trimenii karrooensis TRUE TRUE FALSEkarrooensis. Plagiolepis (Anoplolepis) trimeni var. karrooensis Arnold, 1922: 597 (w.q.) SOUTH AFRICA. Combination in Anoplolepis: Emery, 1925b: 18.
Dolichoderinae Tapinomini Tapinoma albinase TRUE TRUE FALSEalbinase. Technomyrmex nigriventris subsp. albinasis Forel, 1910f: 22 (w.) SOUTH AFRICA. Combination in Tapinoma: Bolton, 2007a: 121. Raised to species: Bolton, 2007a: 121.
Dolichoderinae Tapinomini Tapinoma danitschi TRUE TRUE FALSE1 subspecies
danitschi. Tapinoma danitschi Forel, 1915c: 345 (w.) SOUTH AFRICA. Current subspecies: nominal plus bevisi.
Dolichoderinae Tapinomini Tapinoma luteum TRUE TRUE FALSE1 subspecies
luteum. Technomyrmex luteus Emery, 1895h: 43 (w.) SOUTH AFRICA. Mayr, 1907b: 18 (q.); Wheeler, G.C. & Wheeler, J. 1966: 730 (l.). Combination in Tapinoma: Mayr, 1907b: 18. Current subspecies: nominal plus natalicum.
Dolichoderinae Tapinomini Tapinoma arnoldi tectum TRUE TRUE FALSEtectum. Tapinoma tenue var. tectum Santschi, 1917b: 288 (w.) SOUTH AFRICA. Currently subspecies of arnoldi: Wheeler, W.M. 1922a: 923.
Dolichoderinae Tapinomini Tapinoma danitschi bevisi TRUE TRUE FALSEbevisi. Tapinoma danitschi var. bevisi Forel, 1915c: 346 (w.q.m.) SOUTH AFRICA.
Dolichoderinae Tapinomini Technomyrmex metandrei TRUE TRUE FALSEmetandrei. Technomyrmex metandrei Bolton, 2007a: 28 (w.) CENTRAL AFRICAN REPUBLIC.
Myrmicinae Formicoxenini Terataner transvaalensis TRUE TRUE FALSEtransvaalensis. Terataner (Tranetera) transvaalensis Arnold, 1952b: 130, fig. (w.) SOUTH AFRICA. See also: Bolton, 1981b: 294.
Myrmicinae Tetramoriini Tetramorium africanum TRUE TRUE FALSEafricanum. Macromischa africana Mayr, 1866a: 507 (w.) GHANA. Mayr, 1895: 131 (q.). Combination in Tetramorium: Emery, 1896a: 103; in Macromischoides: Wheeler, W.M. 1922a: 188; in Tetramorium: Bolton, 1976: 363 (in text); Bolton, 1980: 355. Senior synonym of tessmanni: Brown, 1964c: 12; of lamottei: Bolton, 1980: 355.
Myrmicinae Tetramoriini Tetramorium akermani TRUE TRUE FALSEakermani. Tetramorium akermani Arnold, 1926: 265, fig. 73 (w.) SOUTH AFRICA. Senior synonym of myersi: Bolton, 1980: 253.
Myrmicinae Tetramoriini Tetramorium avium TRUE TRUE FALSEavium. Tetramorium avium Bolton, 1980: 276, figs. 58, 63, 64 (w.q.) SOUTH AFRICA.
Myrmicinae Tetramoriini Tetramorium bothae TRUE TRUE FALSEbothae. Tetramorium simillimum subsp. bothae Forel, 1910e: 425 (w.q.m.) SOUTH AFRICA. Raised to species and senior synonym of guillarmodi: Bolton, 1980: 309.
Myrmicinae Tetramoriini Tetramorium buthrum TRUE TRUE FALSEbuthrum. Tetramorium buthrum Bolton, 1980: 309 (w.) CENTRAL AFRICAN REPUBLIC.
Myrmicinae Tetramoriini Tetramorium capense TRUE TRUE FALSEcapense. Tetramorium capense Mayr, 1865: 89 (w.) SOUTH AFRICA. Emery, 1895h: 36 (q.m.). Senior synonym of braunsi: Santschi, 1914a: 435; Bolton, 1980: 359; of popovici: Bolton, 1980: 359. See also: Prins, 1973: 8.
Myrmicinae Tetramoriini Tetramorium clunum TRUE TRUE FALSEclunum. Tetramorium setuliferum st. cluna Forel, 1913j: 218 (w.) SOUTH AFRICA. Raised to species: Bolton, 1980: 244.
Myrmicinae Tetramoriini Tetramorium dichroum TRUE TRUE FALSEdichroum. Tetramorium solidum st. dichroum Santschi, 1932a: 388 (w.) SOUTH AFRICA. Subspecies of peringueyi: Prins, 1973: 11. Raised to species: Bolton, 1980: 245.
Myrmicinae Tetramoriini Tetramorium dominum TRUE TRUE FALSEdominum. Tetramorium dominum Bolton, 1980: 360, fig. 136, 139 (w.) SOUTH AFRICA.
Myrmicinae Tetramoriini Tetramorium emeryi TRUE TRUE FALSEemeryi. Tetramorium emeryi Mayr, 1901b: 23 (w.q.m.) SOUTH AFRICA. Senior synonym of cristulatum: Bolton, 1980: 269.
Myrmicinae Tetramoriini Tetramorium erectum TRUE TRUE FALSEerectum. Tetramorium guineense var. erectum Emery, 1895h: 37 (w.) SOUTH AFRICA. Raised to species and senior synonym of bacchus: Bolton, 1980: 269.
Myrmicinae Tetramoriini Tetramorium frenchi TRUE TRUE FALSEfrenchi. Tetramorium frenchi Forel, 1914d: 229 (w.) SOUTH AFRICA. See also: Bolton, 1980: 278.
Myrmicinae Tetramoriini Tetramorium frigidum TRUE TRUE FALSEfrigidum. Tetramorium akermani var. frigidum Arnold, 1926: 266 (w.) SOUTH AFRICA. Raised to species and senior synonym of drakensbergensis: Bolton, 1980: 256.
Myrmicinae Tetramoriini Tetramorium grandinode TRUE TRUE FALSEgrandinode. Tetramorium grandinode Santschi, 1913c: 308 (w.) SOUTH AFRICA. Senior synonym of hopensis: Bolton, 1980: 247.
Myrmicinae Tetramoriini Tetramorium grassii TRUE TRUE FALSEgrassii. Tetramorium grassii Emery, 1895h: 37 (w.) SOUTH AFRICA. Mayr, 1901b: 25 (q.). Senior synonym of algoa, joffrei, laevigatum, mayri, simulans: Bolton, 1980: 262.
Myrmicinae Tetramoriini Tetramorium inezulae TRUE TRUE FALSEinezulae. Triglyphothrix inezulae Forel, 1914d: 221 (w.m.) SOUTH AFRICA. Combination in Tetramorium: Arnold, 1917: 343. Senior synonym of hepburni, mashonana: Bolton, 1976: 328. See also: Bolton, 1985: 247.
Myrmicinae Tetramoriini Tetramorium isipingense TRUE TRUE FALSEisipingense. Tetramorium simillimum r. isipingense Forel, 1914d: 225 (w.) SOUTH AFRICA. Raised to species: Bolton, 1980: 347.
Myrmicinae Tetramoriini Tetramorium jauresi TRUE TRUE FALSEjauresi. Tetramorium jauresi Forel, 1914d: 226 (w.) SOUTH AFRICA. Senior synonym of latens: Bolton, 1980: 348. See also: Prins, 1973: 17.
Myrmicinae Tetramoriini Tetramorium laevithorax TRUE TRUE FALSElaevithorax. Tetramorium laevithorax Emery, 1895h: 39 (w.) SOUTH AFRICA. Senior synonym of jeanae: Bolton, 1980: 279.
Myrmicinae Tetramoriini Tetramorium longoi TRUE TRUE FALSElongoi. Tetramorium longoi Forel, 1915c: 344 (w.) SOUTH AFRICA. See also: Bolton, 1980: 362.
Myrmicinae Tetramoriini Tetramorium microgyna TRUE TRUE FALSEmicrogyna. Tetramorium microgyna Santschi, 1918b: 132 (q., not w.) SOUTH AFRICA. See also: Arnold, 1926: 253; Santschi, 1928f: 207; Bolton, 1980: 329.
Myrmicinae Tetramoriini Tetramorium microps TRUE TRUE FALSEmicrops. Triglyphothrix microps Mayr, 1901b: 25 (w.q.) SOUTH AFRICA. Combination in Tetramorium: Arnold, 1917: 337; Bolton, 1985: 247. Senior synonym of auropunctatus, bulawayensis, fusciventris, pallens, rhodesiana: Bolton, 1976: 329.
Myrmicinae Tetramoriini Tetramorium parasiticum TRUE TRUE FALSEparasiticum. Tetramorium parasiticum Bolton, 1980: 281, figs. 65, 66 (q.) SOUTH AFRICA.
Myrmicinae Tetramoriini Tetramorium peringueyi TRUE TRUE FALSEperingueyi. Tetramorium peringueyi Arnold, 1926: 260 (w.) SOUTH AFRICA. See also: Prins, 1973: 9; Bolton, 1980: 248.
Myrmicinae Tetramoriini Tetramorium phasias TRUE TRUE FALSEphasias. Tetramorium guineense var. phasias Forel, 1914d: 226 (w.) SOUTH AFRICA. Raised to species and senior synonym of hertigi: Bolton, 1980: 273.
Myrmicinae Tetramoriini Tetramorium platynode TRUE TRUE FALSEplatynode. Tetramorium platynode Bolton, 1980: 258, figs. 35, 38 (w.) SOUTH AFRICA.
Myrmicinae Tetramoriini Tetramorium poweri TRUE TRUE FALSEpoweri. Tetramorium simillimum var. poweri Forel, 1914d: 225 (w.) SOUTH AFRICA. Raised to species: Bolton, 1980: 317.
Myrmicinae Tetramoriini Tetramorium pusillum TRUE TRUE FALSEpusillum. Tetramorium pusillum Emery, 1895h: 38 (w.q.) SOUTH AFRICA. Senior synonym of ladismithensis, tablensis: Bolton, 1980: 318.
Myrmicinae Tetramoriini Tetramorium quadrispinosum TRUE TRUE FALSEquadrispinosum. Tetramorium quadrispinosum Emery, 1886: 362, pl. 17, fig. 8 (w.) SOUTH AFRICA. Senior synonym of montanum: Bolton, 1979: 155; of angolense, beirae, calvum, eudoxia (and its junior synonym elegans), otaviensis, repertum and material of the unavailable name benguelense referred here: Bolton, 1980: 330.
Myrmicinae Tetramoriini Tetramorium regulare TRUE TRUE FALSEregulare. Tetramorium regulare Bolton, 1980: 263, fig. 45 (w.m.) SOUTH AFRICA.
Myrmicinae Tetramoriini Tetramorium rotundatum TRUE TRUE FALSErotundatum. Macromischoides africanus var. rotundatus Santschi, 1924b: 209 (q.) DEMOCRATIC REPUBLIC OF CONGO. Combination in Tetramorium: Bolton, 1980: 357. Raised to species: Bolton, 1980: 357.
Myrmicinae Tetramoriini Tetramorium setigerum TRUE TRUE FALSEsetigerum. Tetramorium setigerum Mayr, 1901b: 22 (w.) SOUTH AFRICA. Forel, 1910e: 424 (q.m.). Senior synonym of anteversa, quaerens: Bolton, 1980: 283.
Myrmicinae Tetramoriini Tetramorium setuliferum TRUE TRUE FALSEsetuliferum. Tetramorium setuliferum Emery, 1895h: 36 (w.) SOUTH AFRICA. Forel, 1910f: 19 (q.); Arnold, 1917: 290 (m.). Material of the nomen nudum squamiferum referred here by Wheeler, W.M. 1922a: 903. Senior synonym of cucalense (and its junior syonym triptolemus): Bolton, 1980: 250.
Myrmicinae Tetramoriini Tetramorium signatum TRUE TRUE FALSEsignatum. Tetramorium solidum var. signatum Emery, 1895h: 35 (w.) SOUTH AFRICA. Raised to species and senior synonym of grootensis, lugubre, tuckeri: Bolton, 1980: 251.
Myrmicinae Tetramoriini Tetramorium solidum TRUE TRUE FALSEsolidum. Tetramorium solidum Emery, 1886: 362, pl. 17, fig. 7 (w.q.) SOUTH AFRICA. See also: Prins, 1973: 4; Bolton, 1980: 252.
Myrmicinae Tetramoriini Tetramorium thoth TRUE TRUE FALSEthoth. Triglyphothrix thoth Bolton, 1976: 337 (w.) SOUTH AFRICA. Combination in Tetramorium: Bolton, 1985: 247.
Myrmicinae Tetramoriini Tetramorium titus TRUE TRUE FALSEtitus. Tetramorium titus Forel, 1910e: 427 (w.) SOUTH AFRICA. See also: Bolton, 1980: 264.
Myrmicinae Tetramoriini Tetramorium traegaordhi TRUE TRUE FALSEtraegaordhi. Tetramorium traegaordhi Santschi, 1914e: 23 (w.q.) SOUTH AFRICA. See also: Bolton, 1980: 292.
Myrmicinae Tetramoriini Tetramorium trimeni TRUE TRUE FALSEtrimeni. Triglyphothrix trimeni Emery, 1895h: 40 (w.q.) SOUTH AFRICA. Combination in Tetramorium: Arnold, 1917: 337. See also: Bolton, 1976: 338; Bolton, 1985: 247.
Myrmicinae Tetramoriini Tetramorium vexator TRUE TRUE FALSEvexator. Tetramorium vexator Arnold, 1926: 269, fig. 76 (w.) SOUTH AFRICA. See also: Bolton, 1980: 265.
Myrmicinae Tetramoriini Tetramorium warreni TRUE TRUE FALSEwarreni. Tetramorium warreni Arnold, 1926: 268, fig. 75 (w.) SOUTH AFRICA. See also: Bolton, 1980: 294.
Myrmicinae Tetramoriini Tetramorium weitzeckeri TRUE TRUE FALSEweitzeckeri. Tetramorium (Xiphomyrmex) weitzeckeri Emery, 1895h: 39 (w.) SOUTH AFRICA. Arnold, 1917: 346 (q.). Combination in Xiphomyrmex: Wheeler, W.M. 1922a: 908; in Tetramorium: Bolton, 1980: 233. Senior synonym of ebeninum, edithae, escherichi, nigellus: Bolton, 1980: 233. See also: Hita Garcia, Fischer & Peters, 2010b: 85.
Pseudomyrmecinae Pseudomyrmecini Tetraponera aethiops TRUE TRUE FALSEaethiops. Tetraponera aethiops Smith, F. 1877b: 71 (w.) SOUTH AFRICA. Santschi, 1911g: 207 (m.); Forel, 1913c: 352 (q.); Wheeler, G.C. & Wheeler, J. 1956: 392 (l.). Combination in Sima: Dalla Torre, 1893: 53; in Sima (Pachysima): Emery, 1912b: 97; in Pachysima: Wheeler, W.M. 1918c: 305; in Tetraponera: Ward, 1990: 487. Senior synonym of spininoda: Emery, 1912b: 97; Wheeler, W.M. 1922a: 801. See also: Arnold, 1916: 175; Wheeler, W.M. 1922a: 114.
Pseudomyrmecinae Pseudomyrmecini Tetraponera ambigua TRUE TRUE FALSEambigua. Sima ambigua Emery, 1895h: 23 (w.q.) SOUTH AFRICA. Forel, 1910f: 5 (m.). Combination in S. (Tetraponera): Arnold, 1916: 185; in Tetraponera: Wheeler, W.M. 1922a: 796. [Sima ambigua Forel, 1894b: 89. Nomen nudum, attributed to Emery.] Senior synonym of angolensis, bifoveolata, encephala, erythraea, maculifrons, occidentalis, rhodesiana, syriaca: Ward, 2006: 123.
Pseudomyrmecinae Pseudomyrmecini Tetraponera braunsi TRUE TRUE FALSE1 subspecies
braunsi. Sima braunsi Forel, 1913a: 112 (w.q.) SOUTH AFRICA. Combination in S. (Tetraponera): Arnold, 1916: 184; in Tetraponera: Wheeler, W.M. 1922a: 797. Subspecies of clypeata: Arnold, 1916: 184; Wheeler, W.M. 1922a: 797. Revived status as species: Ward, 1990: 488. Current subspecies: nominal plus durbanensis.
Pseudomyrmecinae Pseudomyrmecini Tetraponera capensis TRUE TRUE FALSEcapensis. Pseudomyrma capensis Smith, F. 1858b: 160 (w.) SOUTH AFRICA. Combination in Sima: Mayr, 1866b: 906; in Tetraponera: Wheeler, W.M. 1922a: 797.
Pseudomyrmecinae Pseudomyrmecini Tetraponera clypeata TRUE TRUE FALSEclypeata. Sima clypeata Emery, 1886: 361, pl. 17, figs. 4, 5 (q.) SOUTH AFRICA. Emery, 1895h: 24 (w.); Emery, 1899c: 6 (l.); Wheeler, G.C. & Wheeler, J. 1956: 390 (l.). Combination in S. (Tetraponera): Arnold, 1916: 182; in Tetraponera: Wheeler, W.M. 1922a: 797.
Pseudomyrmecinae Pseudomyrmecini Tetraponera emeryi TRUE TRUE FALSEemeryi. Sima emeryi Forel, 1911d: 367 (w.) SOUTH AFRICA. Combination in S. (Tetraponera): Arnold, 1916: 187; in Tetraponera: Wheeler, W.M. 1922a: 797.
Pseudomyrmecinae Pseudomyrmecini Tetraponera natalensis TRUE TRUE FALSE4 subspecies
natalensis. Pseudomyrma natalensis Smith, F. 1858b: 160 (q.) SOUTH AFRICA. Emery, 1895h: 22 (w.); Emery, 1899c: 6 (l.); Wheeler, G.C. & Wheeler, J. 1973d: 208 (l.). Combination in Sima: Emery, 1892e: 237; in Tetraponera: Smith, F. 1877b: 69; Wheeler, W.M. 1922a: 798. Junior synonym of capensis: Gerstäcker, 1871: 355. Revived from synonymy: Smith, F. 1877b: 69. Current subspecies: nominal plus caffra, cuitensis, obscurata, usambarensis. See also: Arnold, 1916: 176.
Pseudomyrmecinae Pseudomyrmecini Tetraponera natalensis obscurata TRUE TRUE FALSEobscurata. Sima natalensis subsp. obscurata Emery, 1895h: 22 (w.) SOUTH AFRICA. Arnold, 1916: 177 (q.m.). Combination in Tetraponera: Wheeler, W.M. 1922a: 799.
Pseudomyrmecinae Pseudomyrmecini Tetraponera braunsi durbanensis TRUE TRUE FALSEdurbanensis. Sima braunsi var. durbanensis Forel, 1914d: 218 (w.) SOUTH AFRICA. Combination in Tetraponera: Wheeler, W.M. 1922a: 797.
Pseudomyrmecinae Pseudomyrmecini Tetraponera natalensis caffra TRUE TRUE FALSEcaffra. Sima natalensis st. caffra Santschi, 1914e: 15 (w.q.m.) SOUTH AFRICA. Combination in Tetraponera: Wheeler, W.M. 1922a: 799.
Dolichoderinae TRUE TRUE FALSEExtant: 28 genera, 692 species, 130 subspecies
Fossil: 16 genera, 103 species
Dolichoderidae Forel, 1878: 364. Type-genus: Dolichoderus.
Taxonomic history
Dolichoderinae as family: Emery, 1894g: 378 [Dolichoderidae]; Ashmead, 1905b: 384 [Dolichoderidae]; Novák & Sadil, 1941: 94 [Dolichoderidae]; Bernard, 1951: 1071 [Dolichoderidae]; Bernard, 1953b: 255 [Dolichoderidae].
Dolichoderinae as tribe of Formicidae: André, 1882a: 127 [Dolichoderidae].
Dolichoderinae as subfamily of Formicidae: Forel, 1878: 364 [Dolichoderidae]; Emery & Forel, 1879a: 454 [Dolichoderidae]; Nasonov, 1889: 26 [Dolichoderidae]; Forel, 1892j: 220 [Dolichoderidae]; Forel, 1893a: 165; Dalla Torre, 1893: 156; Forel, 1895b: 107 [Dolichoderidae]; Emery, 1895j: 771 [subfamily spelled Dolichoderini]; Emery, 1896e: 186; Forel, 1899c: 98; Bingham, 1903: 288; Wheeler, W.M. 1910g: 142; Emery, 1913a: 2; Arnold, 1915: 143; Wheeler, W.M. 1915h: 71; Wheeler, 1915g: 811 [Dolichoderides]; Arnold, 1915: 143; Donisthorpe, 1915d: 178; Forel, 1917: 247; Escherich, 1917: 2 [Dolichoderini]; Bondroit, 1918: 86 [Dolichoderitae]; Wheeler, W.M. 1920: 53; Wheeler, W.M. 1922a: 199; Borgmeier, 1923: 80; Karavaiev, 1936: 163; Smith, M.R. 1951a: 833; Clark, 1951: 16; Brown, 1954e: 29; Kempf, 1972a: 266; Wheeler, G.C. & Wheeler, J. 1972a: 41; Brown, 1973b: 169; all subsequent authors.
Dolichoderinae as formicomorph subfamily of Formicidae: Bolton, 2003: 18, 80.
Dolichoderinae as formicoid subfamily of Formicidae: Brady, Schultz, et al. 2006: 18173; Moreau, Bell, et al. 2006: 102.
Dolichoderinae as formicoid dolichoderomorph subfamily of Formicidae: Ward, 2007a: 556.
Tribes (extant) of Dolichoderinae: Bothriomyrmecini, Dolichoderini, Leptomyrmecini, Tapinomini.
Tribes (extinct) incertae sedis in Dolichoderinae: †Miomyrmecini, †Zherichiniini.
Genera (extinct) incertae sedis in Dolichoderinae: †Alloiomma, †Asymphylomyrmex, †Elaeomyrmex, †Elaphrodites, †Eldermyrmex, †Emplastus, †Eotapinoma, †Eurymyrmex, †Kotshkorkia, †Leptomyrmula, †Miomyrmex, †Petraeomyrmex, †Proiridomyrmex, †Protazteca, †Zherichinius.
Subfamily Dolichoderinae and tribes references, world
Forel, 1878: 364, 380 (diagnosis, genera); Dalla Torre, 1893: 156 (catalogue); Emery, 1895j: 771 (synoptic classification); Emery, 1896e: 186 (genera key); Handlirsch, 1907: 869 (†fossil taxa catalogue); Wheeler, W.M. 1910g: 142 (diagnosis); Emery, 1913a: 2, 6 (diagnosis, tribe key, catalogue); Emery, 1913a: 17 (Tapinomini diagnosis, genera key, catalogue); Arnold, 1915: 144, (diagnosis); Gallardo, 1916a: 3 (diagnosis); Forel, 1917: 247 (synoptic classification); Forel, 1921c: 136 (diagnosis); Wheeler, W.M. 1922a: 199, 687, 688 (diagnosis, tribe key, Tapinomini genera key); Brown & Nutting, 1950: 127 (venation, phylogeny); Brown, 1954e: 29 (phylogeny); Pavan, 1955: 135 (gastral organs); Eisner, 1957: 453 (proventriculus morphology); Bernard, 1967: 246 (diagnosis); Gotwald, 1969: 118 (mouthparts morphology); Wheeler, G.C. & Wheeler, J. 1972a: 41 (diagnosis); Brown, 1973b: 169 (genera & distribution); Wheeler, G.C. & Wheeler, J. 1976b: 61 (larvae, review & synthesis); Snelling, R.R. 1981: 401 (synoptic classification); Dazzini Valcurone & Fanfani, 1985: 1 (gastral glands); Wheeler, G.C. & Wheeler, J. 1985: 258 (synoptic classification); Billen, 1986: 173 (Dufour's gland); Billen, 1987: 278 (abdominal glands); Baroni Urbani & Wilson, 1987: 1 (†fossil Leptomyrmecini); Dlussky & Fedoseeva, 1988: 77 (synoptic classification); Hölldobler & Wilson, 1990: 9 (synoptic classification, genera keys); Baroni Urbani, Bolton & Ward, 1992: 316 (phylogeny); Shattuck, 1992b: 199 (higher classification, phylogeny); Shattuck, 1992c: 20 (revision of subfamily, genera key); Shattuck, 1994: 3 (catalogue); Bolton, 1994: 22 (diagnosis, synoptic classification, genera keys); Shattuck, 1995: 217 (phylogeny, genera); Bolton, 1995a: 1038 (census); Bolton, 1995b: 10 (catalogue); Wenseleers, Schoeters, et al., 1998: 121 (cloacal gland); Brandão, Baroni Urbani, et al. 1999: 411 (phylogeny, genera); Chiotis, Jermiin & Crozier, 2000: 108 (phylogeny, genera); Dlussky & Rasnitsyn, 2002: 416 (diagnosis for impression fossils); Bolton, 2003: 18, 80 (diagnosis, synopsis); Dubovikov, 2005a: 91 (tribes of Dolichoderinae); Brady, Schultz, et al. 2006: 18173 (phylogeny); Moreau, Bell, et al. 2006: 102 (phylogeny); Ward, Brady, et al. 2010: 342 (phylogeny, tribe-rank classification).
Regional and national faunas with keys
André, 1882a: 127 (Europe & Algeria); Nasonov, 1889: 50 (Russia); Forel, 1891b: 9 (Madagascar genera); Forel, 1895e: 460 (India & Sri Lanka); Bingham, 1903: 288 (India, Sri Lanka & Burma); Ruzsky, 1905b: 102 (Russian Empire); Bondroit, 1910: 488 (Belgium); Wheeler, W.M. 1910g: 560 (North America genera); Stitz, 1914: 77 (Central Europe); Gallardo, 1915: 34 (Argentina genera); Forel, 1915d: 40 (Switzerland); Arnold, 1915: 145 (South Africa); Donisthorpe, 1915d: 178 (Britain); Gallardo, 1916a: 12 (Argentina); Emery, 1916b: 208 (Italy); Wheeler, W.M. 1916m: 589 (U.S.A., Connecticut); Bondroit, 1918: 86 (France & Belgium); Gallardo, 1919b: 253 (Argentina genera); Soudek, 1922: 58 (Czechoslovakia); Stärcke, 1926: 117 (Netherlands); Karavaiev, 1927c: 271 (Ukraine); Donisthorpe, 1927b: 199 (Britain); Menozzi & Russo, 1930: 172 (Dominican Republic); Arnol'di, 1933b: 600 (Russia); Menozzi, 1933b: 90 (Israel genera); Karavaiev, 1936: 164 (Ukraine); Smith, M.R. 1937: 861 (Puerto Rico); Stitz, 1939: 209 (Germany); Kratochvíl, 1941: 94 (Central Europe); Novák & Sadil, 1941: 94 (Central Europe); Cole, 1942: 370 (U.S.A., Utah); Smith, M.R. 1943f: 309 (U.S.A. males); Buren, 1944a: 290 (U.S.A., Iowa); Smith, M.R. 1947f: 592 (U.S.A. genera); Creighton, 1950a: 330 (North America); Kusnezov, 1956: 27 (Argentina); Brown, 1958h: 29 (New Zealand); Kusnezov, 1959: 40 (Neotropical genera); Gregg, 1963: 341 (U.S.A., Colorado); Wheeler, G.C. & Wheeler, J. 1963: 149 (U.S.A., North Dakota); Bernard, 1967: 246 (Western Europe); Wilson & Taylor, 1967: 17 (Polynesia); Bolton, 1973a: 329 (West Africa genera); Bolton & Collingwood, 1975: 3 (Britain); Boven, 1970b: 24 (Netherlands); Snelling, R.R. & Hunt, 1976: 90 (Chile); Tarbinsky, 1976: 118 (Kyrghyzstan); Boven, 1977: 123 (Belgium); Kutter, 1977c: 170 (Switzerland); Arnol'di & Dlussky, 1978: 546 (former European U.S.S.R.); Collingwood, 1978: 80 (Iberian Peninsula); Collingwood, 1979: 32 (Fennoscandia & Denmark); Greenslade, 1979: 28 (South Australia genera); Schembri & Collingwood, 1981: 420 (Malta); Allred, 1982: 443 (U.S.A., Utah); Baroni Urbani, 1984: 80 (Neotropical genera); Gösswald, 1985: 314 (Germany); Collingwood, 1985: 242 (Saudi Arabia); Wheeler, G.C. & Wheeler, J. 1986g: 55 (U.S.A., Nevada); Agosti & Collingwood, 1987b: 278 (Balkans); Dlussky, Soyunov & Zabelin, 1990: 168 (Turkmenistan); Kupyanskaya, 1990: 152 (Far Eastern Russia); Morisita, Kubota, Onoyama, et al., 1991: 2 (Japan); Atanasov & Dlussky, 1992: 49 (Bulgaria); Lattke, in Jaffe, 1993: 147 (Neotropical genera); Arakelian, 1994: 70 (Armenia); Wu, J. & Wang, 1995: 115 (China genera); Kupyanskaya, 1995: 331 (Far Eastern Russia); Collingwood & Agosti, 1996: 359 (Saudi Arabia); Seifert, 1996b: 161 (Central Europe); Skinner & Allen, 1996: 40 (Britain); Collingwood & Prince, 1998: 20 (Portugal); Kim & Kim, 1999: 18 (Korea); Shattuck, 1999: 26, 64 (Australia genera, synopsis); Andersen, 2000: 59 (northern Australia genera); Zhou, 2001b: 149 (China, Guangxi); Czechowski, Radchenko & Czechowska, 2002: 134 (Poland); Aktaç & Radchenko, 2002: 54 (Turkey genera); Yoshimura & Onoyama, 2002: 425 (Japan genera, males); Mackay & Mackay, 2002: 221 (U.S.A., New Mexico); Palacio & Fernández, in Fernández, 2003d: 239 (Neotropical genera); Radchenko, 2005b: 186 (North Korea); Coovert, 2005: 102 (U.S.A., Ohio); Clouse, 2007b: 190 (Micronesia); Seifert, 2007: 147 (North and Central Europe); Fisher, 2009: 51 (Malagasy genera); Terayama, 2009: 196 (Taiwan); Heterick, 2009: 28 (south-western Australia genera); Boer, 2010: 14 (Benelux); Yoshimura & Fisher, 2011: 7 (Malagasy males key).
Formicinae TRUE TRUE FALSEExtant: 49 genera, 2,831 species, 839 subspecies
Fossil: 29 genera, 193 species, 1 subspecies
Formicariae Latreille, 1809: 124. Type-genus: Formica.
Taxonomic history
Formicinae as group name: Latreille, 1809: 124 [Formicariae]; Lepeletier de Saint-Fargeau, 1835: 197 [Formicites]; Nylander, 1846a: 877 [Formicae].
Formicinae as family: Smith, F. 1858b: 1 [Formicidae]; Smith, F. 1861b: 36 [Formicidae]; Smith, 1871a: 302 [Formicidae]; André, 1882a: 125 [Formicidae]; Saunders, 1896: 18 [Formicidae]; Novák & Sadil, 1941: 97 [Formicidae]; Bernard, 1951: 1073 [Formicidae]; Bernard, 1953b: 256 [Formicidae].
Formicinae as tribe of Formicidae: André, 1874: 167 [Formicidae].
Formicinae as subfamily of Formicidae: Mayr, 1855: 286, 299 [Formicidae]; Mayr, 1861: 21 [Formicidae]; Mayr, 1862: 651 [Formicidae]; Mayr, 1865: 6 [Formicidae]; Mayr, 1868c: 24 [Formicidae]; Forel, 1870: 307 [Formicidae]; Forel, 1874: 21 [Formicidae]; Emery, 1877a: 70 [Formicidae]; André, 1881b: 54 [Formicidae]; Ashmead, 1905b: 384; Bondroit, 1918: 17 [Formicitae]; Wheeler, W.M. 1920: 53; Wheeler, W.M. 1922a: 210; Emery, 1925b: 2; Karavaiev, 1936: 172; Clark, 1951: 16; Brown, 1954e: 29; Kempf, 1972a: 266; Brown, 1973b: 169; all subsequent authors.
Formicinae as formicomorph subfamily of Formicidae: Bolton, 2003: 20, 93.
Formicinae as formicoid subfamily of Formicidae: Brady, Schultz, et al. 2006: 18173; Moreau, Bell, et al. 2006: 102.
Formicinae as formicoid formicomorph subfamily of Formicidae: Ward, 2007a: 556.
Tribes of Formicinae: Camponotini, Formicini, Gesomyrmecini, Gigantiopini, Lasiini, Melophorini, Myrmecorhynchini, Myrmoteratini, Notostigmatini, Oecophyllini, Plagiolepidini.
Genera (extinct) incertae sedis in Formicinae: †Camponotites, †Drymomyrmex, †Imhoffia, †Kyromyrma, †Leucotaphus, †Protrechina, †Zhangidris.
Hong (2002) genera (extinct) incertae sedis in Formicinae: †Curtipalpulus, †Eoleptocerites, †Eurytarsites, †Fushuniformica, †Huaxiaformica, †Leptogasteritus, †Liaoformica, †Longiformica, †Magnogasterites, †Orbicapitia, †Ovalicapito, †Ovaligastrula, †Sinoformica, †Sinotenuicapito, †Wilsonia (unresolved junior homonym).
Subfamily Formicinae references, world
Mayr, 1862: 651 (genera key); Mayr, 1865: 6 (diagnosis); Handlirsch, 1907: 859 (†fossil taxa catalogue); Dalla Torre, 1893: 171 (catalogue); Emery, 1895j: 772 (synoptic classification); Emery, 1896e: 187 (genera key); Wheeler, W.M. 1910g: 143 (diagnosis); Forel, 1912i: 88 (tribes key); Forel, 1917: 248 (synoptic classification); Arnold, 1920a: 551 (diagnosis); Forel, 1921c: 139 (diagnosis); Wheeler, W.M. 1922a: 210, 691 (diagnosis, tribes key); Emery, 1925b: 2 (diagnosis, tribe key, catalogue); Brown & Nutting, 1950: 127 (venation, phylogeny); Eisner, 1957: 465 (proventriculus morphology); Hung & Brown, 1966: 198 (gastric apex, structure); Bernard, 1967: 267 (diagnosis); Gotwald, 1969: 120 (mouthparts morphology); Wheeler, G.C. & Wheeler, J. 1972a: 41 (diagnosis); Brown, 1973b: 169 (genera, distribution); Wheeler, G.C. & Wheeler, J. 1976b: 62 (larvae, review & synthesis); Snelling, R.R. 1981: 402 (synoptic classification); Wheeler, G.C. & Wheeler, J. 1985: 258 (synoptic classification); Billen, 1986: 173 (Dufour's gland); Dlussky & Fedoseeva, 1988: 77 (synoptic classification); Hölldobler & Wilson, 1990: 9 (synoptic classification, genera keys); Agosti, 1991: 295 (genus group diagnoses); Shattuck, 1992b: 201 (phylogeny); Baroni Urbani, Bolton & Ward, 1992: 317 (phylogeny); Bolton, 1994: 42 (diagnosis, synoptic classification, genera keys); Bolton, 1995a: 1039 (census); Bolton, 1995b: 11 (catalogue); Wenseleers, Schoeters, et al. 1998: 121 (cloacal gland); Dlussky & Rasnitsyn, 2002: 417 (diagnosis for impression fossils); Bolton, 2003: 20, 93 (diagnosis, synopsis); Brady, Schultz, et al. 2006: 18173 (phylogeny); Moreau, Bell, et al. 2006: 102 (phylogeny).
Regional and national faunas with keys
Mayr, 1855: 299 (Austria); Mayr, 1861: 25 (Europe); Mayr, 1868c: 25 (†Baltic Amber); André, 1874: 167 (Europe); Forel, 1874: 22 (Switzerland); Saunders, E. 1880: 203 (Britain); André, 1882a: 126 (Europe & Algeria); Provancher, 1887: 225 (Canada); Cresson, 1887: 94 (U.S.A. genera); Nasonov, 1889: 50 (Russia); Forel, 1891b: 8 (Madagascar genera); Lameere, 1892: 62 (Belgium); Forel, 1892j: 220 (India & Sri Lanka); Bingham, 1903: 308 (India, Sri Lanka & Burma); Ruzsky, 1905b: 100 (Russian Empire); Wasmann, 1906: 7 (Luxemburg); Bondroit, 1910: 481 (Belgium); Wheeler, W.M. 1910g: 560 (North America genera); Stitz, 1914: 80 (Central Europe); Gallardo, 1915: 35 (Argentina genera); Forel, 1915d: 45 (Switzerland); Donisthorpe, 1915d: 184 (Britain); Emery, 1916b: 216 (Italy); Wheeler, W.M. 1916m: 590 (U.S.A., Connecticut); Bondroit, 1918: 17 (France & Belgium); Arnold, 1920a: 552 (South Africa); Kutter, 1920b: 134 (Switzerland); Soudek, 1922: 61 (Czechoslovakia); Lomnicki, 1925a: 160 (Poland); Stärcke, 1926: 118, 146 (Netherlands); Karavaiev, 1927c: 273 (Ukraine); Donisthorpe, 1927b: 205 (Britain); Menozzi & Russo, 1930: 172 (Dominican Republic); Arnol'di, 1933b: 601 (Russia); Menozzi, 1933b: 90 (Israel genera); Karavaiev, 1936: 173 (Ukraine); Smith, M.R. 1937: 865 (Puerto Rico); Stitz, 1939: 230 (Germany); Kratochvíl, 1941: 97 (Central Europe); Novák & Sadil, 1941: 97 (Central Europe); Cole, 1942: 373 (U.S.A., Utah); Smith, M.R. 1943f: 309 (U.S.A., males); Buren, 1944a: 292 (U.S.A., Iowa); Holgersen, 1943b: 173 (Norway); Holgersen, 1944: 199 (Norway); Smith, M.R. 1947f: 599 (U.S.A. genera); Boven, 1947: 181 (Belgium); Creighton, 1950a: 355 (North America); Kusnezov, 1956: 31 (Argentina); Brown, 1958h: 42 (New Zealand); Boven, 1959: 11 (Netherlands); Gregg, 1963: 447 (U.S.A., Colorado); Wheeler, G.C. & Wheeler, J. 1963: 160 (U.S.A., North Dakota); Collingwood, 1964: 104 (Britain); Bernard, 1967: 268 (Western Europe); Wilson & Taylor, 1967: 17 (Polynesia); Boven, 1970b: 26 (Netherlands); Kempf, 1972a: 266 (Neotropical, synoptic classification); Bolton, 1973a: 329 (West Africa genera); Bolton & Collingwood, 1975: 3 (Britain); Snelling & Hunt, 1976: 104 (Chile); Tarbinsky, 1976: 126 (Kyrghyzstan); Boven, 1977: 126 (Belgium); Kutter, 1977c: 183 (Switzerland); Arnol'di & Dlussky, 1978: 548 (former European U.S.S.R.); Collingwood, 1978: 88 (Iberian Peninsula); Collingwood, 1979: 85 (Fennoscandia & Denmark); Greenslade, 1979: 32 (South Australia genera); Schembri & Collingwood, 1981: 436 (Malta); Prins, 1983: 8 (Southern Africa genera); Allred, 1982: 444 (U.S.A., Utah); Verhaeghe, Deligne, et al., 1984: 106 (Belgium genera); Baroni Urbani, 1984: 81 (Neotropical genera); Gösswald, 1985: 263 (Germany); Collingwood, 1985: 273 (Saudi Arabia); Wheeler, G.C. & Wheeler, J. 1986g: 58 (U.S.A., Nevada); Nilsson & Douwes, 1987: 68 (Norway); Agosti & Collingwood, 1987b: 279 (Balkans); Dlussky, Soyunov & Zabelin, 1990: 124 (Turkmenistan); Kupyanskaya, 1990: 162 (Far Eastern Russia); Morisita, Kubota, Onoyama, et al., 1991: 10 (Japan); Atanasov & Dlussky, 1992: 49 (Bulgaria); Shattuck, 1992b: 199 (higher classification, phylogeny); Lattke, in Jaffe, 1993: 150 (Neotropical genera); Arakelian, 1994: 76 (Armenia); Wu, J. & Wang, 1995: 125 (China genera); Kupyanskaya, 1995: 332 (Far Eastern Russia); Collingwood & Agosti, 1996: 361 (Saudi Arabia); Seifert, 1996b: 166 (Central Europe); Skinner & Allen, 1996: 41 (Britain); Collingwood & Prince, 1998: 21 (Portugal); Shattuck, 1999: 25, 86 (Australia genera, synopsis); Andersen, 2000: 68 (northern Australia genera); Zhou, 2001b: 165 (China, Guangxi); Czechowski, Radchenko & Czechowska, 2002: 147 (Poland); Aktaç & Radchenko, 2002: 54 (Turkey genera); Yoshimura & Onoyama, 2002: 425 (Japan genera, males); Mackay & Mackay, 2002: 236 (U.S.A., New Mexico); Palacio & Fernández, in Fernández, 2003d: 242 (Neotropical genera); Radchenko, 2005b: 187 (North Korea); Coovert, 2005: 113 (U.S.A., Ohio); Clouse, 2007b: 190 (Micronesia); Seifert, 2007: 150 (North and Central Europe); Terayama, 2009: 202 (Taiwan); Heterick, 2009: 30 (south-western Australia genera); Boer, 2010: 17 (Benelux).
Myrmicinae TRUE TRUE FALSEExtant: 141 genera, 6,087 species, 864 subspecies
Fossil: 31 genera, 130 species
Myrmicites Lepeletier de Saint-Fargeau, 1835: 169. Type-genus: Myrmica.
Taxonomic history
Myrmicinae as group name: Lepeletier de Saint-Fargeau, 1835: 169 [Myrmicites]; Nylander, 1846a: 877 [Myrmicae].
Myrmicinae as family: Smith, F. 1851: 4 [Myrmicidae]; Smith, F. 1861b: 45 [Myrmicidae]; Smith, F. 1871a: 324 [Myrmicidae]; André, 1882a: 125 [Myrmicidae]; Cresson, 1887: 93 [Myrmicidae]; Emery, 1894g: 383 [Myrmicidae]; Saunders, 1896: 18 [Myrmicidae]; Ashmead, 1905b: 383 [Myrmicidae]; Novák & Sadil, 1941: 71 [Myrmicidae]; Bernard, 1951: 1058 [Myrmicidae]; Bernard, 1953b: 222 [Myrmicidae].
Myrmicinae as subfamily of Poneridae: Smith, F. 1858b: 114 [Myrmicidae].
Myrmicinae as tribe of Formicidae: André, 1874: 167 [Myrmicidae].
Myrmicinae as subfamily of Myrmicidae: Ashmead, 1905b: 383.
Myrmicinae as subfamily of Formicidae: Mayr, 1855: 290, 299 [Myrmicidae]; Smith, F. 1857a: 70 [Myrmicidae]; Mayr, 1861: 21 [Myrmicidae]; Smith, F. 1862b: 33 [Myrmicidae]; Mayr, 1862: 738 [Myrmicidae]; Mayr, 1865: 17 [Myrmicidae]; Mayr, 1868c: 24 [Myrmicidae]; Forel, 1870: 307 [Myrmicidae]; Forel, 1874: 22 [Myrmicidae]; Emery, 1877a: 70 [Myrmicidae]; Forel, 1878: 367 [Myrmicidae]; Emery & Forel, 1879a: 456 [Myrmicidae]; André, 1881b: 64 [Myrmicidae]; Nasonov, 1889: 28 [Myrmicidae]; Forel, 1891b: 11 [Myrmicidae]; Forel, 1892j: 220 [Myrmicidae]; Forel, 1893a: 163; Dalla Torre, 1893: 53; Emery, 1895j: 768 [subfamily spelled Myrmicini]; Emery, 1896e: 179; Forel, 1899c: 30; Forel, 1902f: 520; Bingham, 1903: 105; Wheeler, W.M. 1910g: 138; Emery, 1914a: 29; Wheeler, 1915g: 806 [Myrmicides]; Wheeler, W.M. 1915h: 40; Donisthorpe, 1915d: 74; Arnold, 1916: 166; Escherich, 1917: 2 [Myrmicini]; Forel, 1917: 240 [subfamily spelled Myrmicini]; Bondroit, 1918: 14 [Myrmicitae]; Wheeler, W.M. 1920: 53; Wheeler, W.M. 1922a: 124; Emery, 1921f: 3; Karavaiev, 1934: 59; Clark, 1951: 16; Brown, 1954e: 28; Wheeler, G.C. & Wheeler, J. 1972a: 40; Brown, 1973b: 166; all subsequent authors.
Myrmicinae as myrmicomorph subfamily of Formicidae: Bolton, 2003: 52, 182.
Myrmicinae as formicoid subfamily of Formicidae: Moreau, Bell, et al. 2006: 102; Brady, Schultz, et al. 2006: 18173.
Myrmicinae as formicoid myrmicomorph subfamily of Formicidae: Ward, 2007a: 556.
Tribes of Myrmicinae: Adelomyrmecini, Ankylomyrmini, Attini, Basicerotini, Blepharidattini, Cataulacini, Cephalotini, Crematogastrini, Dacetini, Formicoxenini, Lenomyrmecini, Liomyrmecini, Melissotarsini, Meranoplini, Metaponini, Myrmecinini, Myrmicariini, Myrmicini, Paratopulini, Phalacromyrmecini, Pheidolini, Solenopsidini, Stegomyrmecini, Stenammini, Tetramoriini.
Genera (extant) incertae sedis in Myrmicinae: Diaphoromyrma, Tyrannomyrmex.
Genera (extinct) incertae sedis in Myrmicinae: †Afromyrma, †Attopsis, †Cephalomyrmex, †Electromyrmex, †Eocenidris, †Eomyrmex, †Fallomyrma, †Miosolenopsis, †Myrmecites, †Solenopsites.
Hong (2002) genera (extinct) incertae sedis in Myrmicinae: †Brachytarsites, †Clavipetiola, †Fushunomyrmex, †Orbigastrula, †Quadrulicapito, †Quineangulicapito, †Sinomyrmex, †Sphaerogasterites, †Wumyrmex.
Collective group name in Myrmicinae: †Myrmicites.
Subfamily Myrmicinae references, world
Mayr, 1865: 17 (diagnosis); Mayr, 1867a: 91 (diagnosis); Forel, 1878: 367 (diagnosis); Handlirsch, 1907: 872 (†fossil taxa catalogue); Dalla Torre, 1893: 53 (catalogue); Emery, 1895j: 768 (diagnosis); Emery, 1896e: 179 (genera key); Wheeler, W.M. 1910g: 138 (diagnosis); Emery, 1912b: 101 (phylogeny); Emery, 1914a: 34 (phylogeny, tribe key); Arnold, 1916: 164 (diagnosis); Forel, 1917: 240 (synoptic classification); Forel, 1921c: 139 (diagnosis); Emery, 1921f: 3 (diagnosis, tribes & genera key, catalogue); Wheeler, W.M. 1922a: 124, 655 (diagnosis, tribes key); Brown & Nutting, 1950: 126 (venation, phylogeny); Brown, 1954e: 28 (phylogeny); Eisner, 1957: 477 (proventriculus morphology); Bernard, 1967: 93 (diagnosis); Gotwald, 1969: 99 (mouthparts morphology); Wheeler, G.C. & Wheeler, J. 1972a: 40 (diagnosis); Brown, 1973b: 166 (genera, distribution); Wheeler, G.C. & Wheeler, J. 1976b: 52 (larvae, review & synthesis); Kugler, C. 1978a: 413 (sting structure); Kugler, C. 1978b: 267 (pygidial glands); Kugler, C. 1979c: 117 (sting, evolution); Snelling, R.R. 1981: 393 (synoptic classification); Caetano, F.H. 1984: 257 (digestive tract, morphology); Wheeler, G.C. & Wheeler, J. 1985: 257 (synoptic classification); Billen, 1986: 167 (Dufour's gland); Dlussky & Fedoseeva, 1988: 79 (synoptic classification); Hölldobler & Wilson, 1990: 9 onward (synoptic classification, genera keys); Baroni Urbani, Bolton & Ward, 1992: 317 (phylogeny); Bolton, 1994: 75 (diagnosis, synoptic classification, genera keys); Bolton, 1995a: 1040 (census); Bolton, 1995b: 13 (catalogue); Hashimoto, 1996: 354 (phylogenetic position); Baroni Urbani, 2000: 480 (phylogeny); Dlussky & Rasnitsyn, 2002: 422 (diagnosis for wingless fossils); Bolton, 2003: 52, 182 (diagnosis, synopsis); Moreau, Bell, et al. 2006: 102 (phylogeny); Brady, Schultz, et al. 2006: 18173 (phylogeny); Ward, 2007a: 556 (classification).
Regional and national faunas with keys
Mayr, 1855: 391 (Austria); Mayr, 1861: 29 (Europe); Mayr, 1868c: 79 (†Baltic Amber); André, 1874: 171 (Europe); Forel, 1874: 29 (Switzerland); Saunders, E. 1880: 213 (Britain); André, 1882c: 256 (Europe & Algeria); Cresson, 1887: 98 (U.S.A. genera); Provancher, 1887: 243 (Canada); Nasonov, 1889: 54 (Russia); Forel, 1891b: 11 (Madagascar genera); Lameere, 1892: 66 (Belgium); Forel, 1902f: 520 (India & Sri Lanka genera); Bingham, 1903: 105 (India, Sri Lanka & Burma); Ruzsky, 1905b: 103 (Russian Empire); Wasmann, 1906: 13 (Luxemburg); Bondroit, 1910: 490 (Belgium); Wheeler, W.M. 1910g: 558 (North America genera); Stitz, 1914: 55 (Central Europe); Gallardo, 1915: 32 (Argentina genera); Forel, 1915d: 8 (Switzerland); Donisthorpe, 1915d: 74 (Britain); Arnold, 1916: 166, 170 (South Africa tribes, genera); Emery, 1916b: 112 (Italy); Wheeler, W.M. 1916m: 581 (U.S.A., Connecticut); Bondroit, 1918: 90 (France & Belgium); Kutter, 1920b: 144 (Switzerland); Soudek, 1922: 20 (Czechoslovakia); Stärcke, 1926: 84 (Netherlands); Karavaiev, 1927c: 256 (Ukraine); Donisthorpe, 1927b: 77 (Britain); Menozzi & Russo, 1930: 170 (Dominican Republic); Gallardo, 1932b: 91 (Argentina, tribes); Arnol'di, 1933b: 596 (Russia); Menozzi, 1933b: 88 (Israel genera); Karavaiev, 1934: 60 (Ukraine); Smith, M.R. 1937: 829 (Puerto Rico); Stitz, 1939: 63 (Germany); Kratochvíl, 1941: 71 (Central Europe); Novák & Sadil, 1941: 71 (Central Europe); Cole, 1942: 360 (U.S.A., Utah); Smith, M.R. 1943f: 291 (U.S.A., males); Holgersen, 1943b: 166 (Norway); Holgersen, 1944: 198 (Norway); Buren, 1944a: 281 (U.S.A., Iowa); Smith, M.R. 1947f: 543 (U.S.A. genera); Boven, 1947: 170 (Belgium); Creighton, 1950a: 83 (North America); Kusnezov, 1956: 15 (Argentina); Brown, 1958h: 25 (New Zealand); Boven, 1959: 7 (Netherlands); Gregg, 1963: 288 (U.S.A., Colorado); Wheeler, G.C. & Wheeler, J. 1963: 92 (U.S.A., North Dakota); Collingwood, 1964: 94 (Britain); Bernard, 1967: 95 (Western Europe); Wilson & Taylor, 1967: 13 (Polynesia); Boven, 1970b: 9 (Netherlands); Kempf, 1972a: 263 (Neotropical, synoptic classification); Bolton, 1973a: 325 (West Africa genera); Bolton & Collingwood, 1975: 3 (Britain); Snelling, R.R. & Hunt, 1976: 70 (Chile); Tarbinsky, 1976: 19 (Kyrghyzstan); Boven, 1977: 69 (Belgium); Kutter, 1977c: 31 (Switzerland); Arnol'di & Dlussky, 1978: 524 (former European U.S.S.R.); Collingwood, 1978: 75 (Iberian Peninsula); Collingwood, 1979: 36 (Fennoscandia & Denmark); Greenslade, 1979: 20 (South Australia genera); Schembri & Collingwood, 1981: 423 (Malta); Allred, 1982: 438 (U.S.A., Utah); Baroni Urbani, 1984: 76 (Neotropical genera); Verhaeghe, Deligne, et al. 1984: 112 (Belgium genera); Gösswald, 1985: 289 (Germany); Collingwood, 1985: 245 (Saudi Arabia); Wheeler, G.C. & Wheeler, J. 1986g: 20 (U.S.A., Nevada); Nilsson & Douwes, 1987: 57 (Norway); Agosti & Collingwood, 1987b: 265 (Balkans); Dlussky, Soyunov & Zabelin, 1990: 181 (Turkmenistan); Kupyanskaya, 1990: 89 (Far Eastern Russia); Ogata, 1991b: 61 (Japan genera); Morisita, Kubota, et al. 1992: 1 (Japan); Atanasov & Dlussky, 1992: 51 (Bulgaria); Lattke, in Jaffe, 1993: 153 (Neotropical genera); Arakelian, 1994: 15 (Armenia); Wu, J. & Wang, 1995: 57 (China genera); Kupyanskaya, 1995: 327 (Far Eastern Russia); Collingwood & Agosti, 1996: 308 (Saudi Arabia); Seifert, 1996b: 108 (Central Europe); Skinner & Allen, 1996: 43 (Britain); Collingwood & Prince, 1998: 10 (Portugal); Shattuck, 1999: 39, 122 (Australia genera, synopsis); Andersen, 2000: 37 (northern Australia genera); Zhou, 2001b: 69 (China, Guangxi); Czechowski, Radchenko & Czechowska, 2002: 135 (Poland); Aktaç & Radchenko, 2002: 55 (Turkey genera); Yoshimura & Onoyama, 2002: 424 (Japan genera, males); Mackay & Mackay, 2002: 58 (U.S.A., New Mexico); Palacio & Fernández, in Fernández, 2003d: 244 (Neotropical genera and synopsis); Coovert, 2005: 32 (U.S.A., Ohio); Radchenko, 2005b: 184 (North Korea); Clouse, 2007b: 190 (Micronesia); Seifert, 2007: 110 (North and Central Europe); Terayama, 2009: 131 (Taiwan); Heterick, 2009: 36 (south-western Australia genera); Boer, 2010: 46 (Benelux); Eguchi, Bui & Yamane, 2011: 8 (Vietnam genera).
Ponerinae TRUE TRUE FALSEExtant: 27 genera, 1,033 species, 155 subspecies
Fossil: 9 genera, 68 species, 2 subspecies
Ponerites Lepeletier de Saint-Fargeau, 1835: 185. Type-genus: Ponera.
Taxonomic history
Ponerinae as group name: Lepeletier de Saint-Fargeau, 1835: 185 [Ponérites].
Ponerinae as family: Smith, F. 1851: 6 [Poneridae]; Smith, F. 1858b: 76 [Poneridae]; Smith, F. 1861b: 44 [Poneridae]; Smith, F. 1871a: 320 [Poneridae]; André, 1882a: 125 [Poneridae]; Cresson, 1887: 93 [Poneridae]; Emery, 1894g: 379 [Poneridae]; Saunders, 1896: 18 [Poneridae]; Ashmead, 1905b: 382 [Poneridae]; Novák & Sadil, 1941: 70 [Poneridae]; Bernard, 1951: 1042 [Poneridae]; Bernard, 1953b: 185 [Poneridae].
Ponerinae as tribe of Formicidae: André, 1874: 167 [Poneridae].
Ponerinae as subfamily of Poneridae: Ashmead, 1905b: 382.
Ponerinae as subfamily of Formicidae: Mayr, 1855: 289, 299 [Poneridae]; Mayr, 1861: 21 [Poneridae]; Mayr, 1862: 712 [Poneridae]; Smith, F. 1857a: 64 [Poneridae]; Smith, F. 1862b: 31 [Poneridae]; Mayr, 1865: 11 [Poneridae]; Mayr, 1868c: 24 [Poneridae]; Forel, 1870: 307 [Poneridae]; Forel, 1874: 21 [Poneridae]; Emery, 1877a: 70 [Poneridae]; Forel, 1878: 366 [Poneridae]; Emery & Forel, 1879a: 455 [Poneridae]; André, 1881b: 64 [Poneridae]; Forel, 1892j: 220 [Poneridae]; Dalla Torre, 1893: 13; Forel, 1893a: 162; Forel, 1895b: 111 [Poneridae]; Nasonov, 1889: 27 [Poneridae]; Emery, 1895j: 766 [subfamily spelled Ponerini]; Emery, 1896e: 176; Forel, 1899c: 2; Emery, 1901a: 36; Bingham, 1903: 23; Wheeler, W.M. 1910g: 134; Emery, 1911d: 2; Wheeler, 1915g: 805 [Ponerides]; Donisthorpe, 1915d: 65; Wheeler, W.M. 1915h: 25; Arnold, 1915: 9; Escherich, 1917: 2 [Ponerini]; Forel, 1917: 235; Bondroit, 1918: 78 [Poneritae]; Wheeler, W.M. 1920: 53; Wheeler, W.M. 1922a: 56, 632, 640; Borgmeier, 1923: 37; Karavaiev, 1934: 49; Clark, 1951: 15; Brown, 1954e: 24; Wheeler, G.C. & Wheeler, J. 1972a: 39; Brown, 1973b: 165; all subsequent authors.
Ponerinae as poneromorph subfamily of Formicidae: Bolton, 2003: 42, 156.
Ponerinae as poneroid subfamily of Formicidae: Ouellette, Fisher, et al. 2006: 365; Brady, Schultz, et al. 2006: 18173; Moreau, Bell et al. 2006: 102; Ward, 2007a: 555.
Tribes of Ponerinae: Platythyreini, Ponerini, Thaumatomyrmecini.
Genera (extinct) incertae sedis in Ponerinae: †Afropone, †Eogorgites, †Eoponerites, †Furcisutura, †Longicapitia.
Subfamily Ponerinae references, world
Smith, F. 1858b: 76 (diagnosis); Mayr, 1862: 712 (genera key); Mayr, 1865: 11 (Odontomachidae, Poneridae diagnoses), Mayr, 1867a: 79, 81 (Odontomachidae, Poneridae diagnoses); Forel, 1878: 366 (diagnosis); Dalla Torre, 1893: 13 (world catalogue); Emery, 1895j: 766 (diagnosis); Emery, 1896e: 176 (genera key); Handlirsch, 1907: 879 (†fossil taxa catalogue); Wheeler, W.M. 1910g: 134 (diagnosis); Emery, 1911d: 3 (diagnosis, tribes key); Forel, 1917: 235 (synoptic classification); Forel, 1921c: 133 (diagnosis); Wheeler, W.M. 1922a: 56, 636 (diagnosis, tribes key); Brown & Nutting, 1950: 124 (venation, phylogeny); Brown, 1954e: 24 (phylogeny); Eisner, 1957: 475 (proventriculus morphology); Bernard, 1967: 79 (diagnosis); Gotwald, 1969: 25 (mouthparts morphology); Wheeler, G.C. & Wheeler, J. 1972a: 39 (diagnosis); Brown, 1973b: 165 (genera, distribution); Wheeler, G.C. & Wheeler, J. 1976b: 48 (larvae, review & synthesis); Snelling, R.R. 1981: 387 (synoptic classification); Wheeler, G.C. & Wheeler, J. 1985: 256 (synoptic classification); Billen, 1986: 168 (Dufour's gland); Fanfani & Dazzini Valcurone, 1986: 115 (exocrine glands synopsis); Dlussky & Fedoseeva, 1988: 78 (synoptic classification); Hölldobler & Wilson, 1990: 9 onward (synoptic classification, genera keys); Baroni Urbani, Bolton & Ward, 1992: 317 (phylogeny); Bolton, 1994: 153 (diagnosis, synoptic classification, genera keys); Bolton, 1995a: 1042 (census); Bolton, 1995b: 14 (catalogue); Grimaldi, Agosti & Carpenter, 1997: 20 (†Cretaceous genera); Dlussky & Rasnitsyn, 2002: 422 (diagnosis for wingless fossils); Bolton, 2003: 42, 156 (diagnosis, synopsis); Ouellette, Fisher et al. 2006: 359 (phylogeny); Brady, Schultz, et al. 2006: 18173 (phylogeny); Moreau, Bell et al. 2006: 102 (phylogeny); Ward, 2007a: 555 (classification).
.
Regional and national faunas with keys
Mayr, 1861: 28 (Europe); Mayr, 1868c: 69 (†Baltic Amber); André, 1874: 170 (Europe); Forel, 1874: 29 (Switzerland); André, 1882b: 227 (Europe & Algeria); Cresson, 1887: 97 (U.S.A., genera); Provancher, 1887: 238 (Canada); Nasonov, 1889: 54 (Russia); Forel, 1891b: 9 (Madagascar genera); Forel, 1900c: 52 (India & Sri Lanka); Bingham, 1903: 23 (India, Sri Lanka & Burma); Ruzsky, 1905b: 108 (Russian Empire); Wheeler, W.M. 1910g: 557 (North America genera); Bondroit, 1910: 489 (Belgium); Stitz, 1914: 54 (Central Europe); Gallardo, 1915: 31 (Argentina genera); Forel, 1915d: 6 (Switzerland); Arnold, 1915: 10 (South Africa); Donisthorpe, 1915d: 65 (Britain); Emery, 1916b: 98 (Italy); Wheeler, W.M. 1916m: 580 (U.S.A., Connecticut); Bondroit, 1918: 79 (France & Belgium); Gallardo, 1918b: 4 (Argentina); Soudek, 1922: 18 (Czechoslovakia); Stärcke, 1926: 82 (Netherlands); Donisthorpe, 1927b: 68 (Britain); Menozzi & Russo, 1930: 169 (Dominican Republic); Arnol'di, 1933b: 596 (Russia); Karavaiev, 1934: 50 (Ukraine); Smith, M.R. 1937: 823 (Puerto Rico); Stitz, 1939: 58 (Germany); Smith, M.R. 1943f: 278 (U.S.A. males); Buren, 1944a: 279 (U.S.A., Iowa); Smith, M.R. 1947f: 530 (U.S.A. genera); Creighton, 1950a: 31 (North America); Kusnezov, 1956: 11 (Argentina); Brown, 1958h: 11 (New Zealand); Gregg, 1963: 280 (U.S.A., Colorado); Bernard, 1967: 80 (Western Europe); Wilson & Taylor, 1967: 10 (Polynesia); Kempf, 1972a: 262 (Neotropical synoptic classification); Bolton, 1973a: 323 (West Africa genera); Bolton & Collingwood, 1975: 3 (Britain); Boven, 1977: 66 (Belgium); Kutter, 1977c: 21 (Switzerland); Arnol'di & Dlussky, 1978: 522 (former European U.S.S.R.); Collingwood, 1978: 74 (Iberian Peninsula); Collingwood, 1979: 29 (Fennoscandia & Denmark); Greenslade, 1979: 14 (South Australia genera); Francoeur, 1979: 30 (Canada, Québec); Schembri & Collingwood, 1981: 417 (Malta); Allred, 1982: 438 (U.S.A., Utah); Baroni Urbani, 1984: 75 (Neotropical genera); Collingwood, 1985: 236 (Saudi Arabia); Wheeler, G.C. & Wheeler, J. 1986g: 17 (U.S.A., Nevada); Agosti & Collingwood, 1987b: 264 (Balkans); Ogata, 1987: 101 (Japan genera); Morisita, Kubota, et al. 1989: 8 (Japan); Dlussky, Soyunov & Zabelin, 1990: 173 (Turkmenistan); Kupyanskaya, 1990: 85 (Far Eastern Russia); Atanasov & Dlussky, 1992: 51 (Bulgaria); Lattke, in Jaffe, 1993: 166 (Neotropical genera); Arakelian, 1994: 10 (Armenia); Wu, J. & Wang, 1995: 31 (China genera); Collingwood & Agosti, 1996: 308 (Saudi Arabia); Seifert, 1996b: 107 (Central Europe); Collingwood & Prince, 1998: 10 (Portugal); Kim, Kim & Kim, 1998: 145 (Korea); Shattuck, 1999: 52, 179 (Australia genera synopsis); Andersen, 2000: 19 (northern Australia genera); Zhou, 2001b: 22 (China, Guangxi); Czechowski, Radchenko & Czechowska, 2002: 133 (Poland); Aktaç & Radchenko, 2002: 53 (Turkey genera); Yoshimura & Onoyama, 2002: 436 (Japan genera, males); Mackay & Mackay, 2002: 27 (U.S.A., New Mexico); Csösz, 2003: 147 (Carpathian Basin); Palacio & Fernández, in Fernández, 2003d: 256 (Neotropical genera); Radchenko, 2005b: 184 (North Korea); Coovert, 2005: 23 (U.S.A., Ohio); Yoshimura & Fisher, 2007: 27 (Malagasy genera, males); Clouse, 2007b: 190 (Micronesia); Seifert, 2007: 108 (North and Central Europe); Bolton & Fisher, 2008c: 32 (Afrotropical genera); Fernández & Arias-Penna, 2008: 32 (Neotropical genera); Dlussky, 2009: 1071 (Eocene amber); Terayama, 2009: 101 (Taiwan genera); Heterick, 2009: 34 (south-western Australia genera); Boer, 2010: 68 (Benelux).
Dolichoderinae Bothriomyrmecini Bothriomyrmex TRUE TRUE FALSE34 species, 11 subspecies
Bothriomyrmex Emery, 1869c: 117. Type-species: Bothriomyrmex costae, by monotypy.
Taxonomic history
Bothriomyrmex in Formicinae: André, 1881b: 64 [Formicidae].
Bothriomyrmex in Dolichoderinae: Forel, 1878: 380 [Dolichoderidae]; Emery & Forel, 1879a: 454 [Dolichoderidae]; Dalla Torre, 1893: 170; Emery, 1895j: 771; Forel, 1895e: 469; Wheeler, W.M. 1910g: 142.
Bothriomyrmex in Dolichoderidae: Ashmead, 1905b: 384.
Bothriomyrmex in Dolichoderinae, Tapinomini: Emery, 1913a: 27; Forel, 1917: 248; Bondroit, 1918: 88; Wheeler, W.M. 1922a: 689; all subsequent authors to the following.
Bothriomyrmex in Dolichoderinae, Dolichoderini: Shattuck, 1992c: 54; Bolton, 1994: 26; Bolton, 2003: 84.
Bothriomyrmex in Dolichoderinae, Iridomyrmecini, Bothriomyrmecina: Dubovikov, 2005a: 92.
Bothriomyrmex in Dolichoderinae, Bothriomyrmecini: Ward, Brady, et al. 2010: 361.
Genus Bothriomyrmex references
Forel, 1878: 380 (diagnosis); André, 1882b: 218 (Europe & Algeria species); Dalla Torre, 1893: 170 (catalogue); Forel, 1895e: 469 (India & Sri Lanka species key); Bingham, 1903: 305 (India, Sri Lanka & Burma species key); Emery, 1913a: 27 (diagnosis, catalogue); Wheeler, W.M. 1922a: 690 (subgenera key); Emery, 1925e: 5 (European and Oriental species); Kratochvíl, 1941: 96 (Central Europe species key); Novák & Sadil, 1941: 96 (Central Europe species key); Chapman & Capco, 1951: 187 (Asia checklist); Bernard, 1967: 260 (diagnosis, Western Europe species key); Kutter, 1977c: 175 (Switzerland species key); Arnol'di & Dlussky, 1978: 548 (former European U.S.S.R. species key); Collingwood, 1978: 87 (Iberian Peninsula species key); Gösswald, 1985: 316 (Germany species key); Taylor & Brown, D.R. 1985: 92 (Australia catalogue); Taylor, 1987a: 8 (Australia checklist); Agosti & Collingwood, 1987b: 279 (Balkans species key); Shattuck, 1992c: 54 (diagnosis, review of genus); Shattuck, 1994: 31 (catalogue); Bolton, 1995a: 1048 (census); Bolton, 1995b: 80 (catalogue); Wu, J. & Wang, 1995: 119 (China species key); Seifert, 1996b: 163 (Central Europe species key); Shattuck, 1999: 66 (Australia synopsis); Seifert, 2007: 148 (North and Central Europe species key).
Dolichoderinae Dolichoderini Dolichoderus TRUE TRUE FALSEExtant: 122 species, 21 subspecies
Fossil: 53 species
Dolichoderus Lund, 1831a: 130. Type-species: Formica attelaboides, by monotypy.
Taxonomic history
Dolichoderus in Formicidae: Smith, F. 1858b: 75.
Dolichoderus in Formicinae: Mayr, 1862: 652 (in key) [Formicidae]; Mayr, 1865: 9 [Formicidae].
Dolichoderus in Dolichoderidae: Ashmead, 1905b: 384.
Dolichoderus in Dolichoderinae: Forel, 1878: 386 [Dolichoderidae]; Emery & Forel, 1879a: 455 [Dolichoderidae]; Dalla Torre, 1893: 156; Forel, 1895b: 107 [Dolichoderidae]; Emery, 1895j: 771; Forel, 1895e: 462; Forel, 1899c: 98; Wheeler, W.M. 1910g: 142.
Dolichoderus in Dolichoderinae, Dolichoderini: Emery, 1913a: 7; Wheeler, W.M. 1915h: 77; Forel, 1917: 247; Wheeler, W.M. 1922a: 688; all subsequent authors.
Junior synonyms of DOLICHODERUS
Hypoclinea Mayr, 1855: 377 (attributed to Foerster). Type-species: Formica quadripunctata, by subsequent designation of Wheeler, W.M. 1911f: 165.
Taxonomic history
Hypoclinea in Formicidae: Smith, F. 1858b: 57.
Hypoclinea in Formicinae: Mayr, 1855: 377 [Formicidae]; Mayr, 1861: 40 [Formicidae]; Mayr, 1862: 652 (in key) [Formicidae]; Mayr, 1865: 9 [Formicidae]; Mayr, 1868c: 53 [Formicidae]; André, 1874: 168 [Formicidae].
Hypoclinea in Dolichoderidae: Ashmead, 1905b: 384.
Hypoclinea in Dolichoderinae, Dolichoderini: Emery, 1913a: 18; Forel, 1917: 247; all subsequent authors.
Hypoclinea as genus: Mayr, 1855: 377; Mayr, 1865: 9; Mayr, 1868c: 53; Mayr, 1870b: 953; Smith, 1871a: 310; Ashmead, 1905b: 384; Kempf, 1972a: 118; Lattke, 1987b: 259; Dlussky & Fedoseeva, 1988: 78; Mackay & Vinson, 1989b: 17; Hölldobler & Wilson, 1990: 17; Wu, J. & Wang, 1995: 120.
Hypoclinea as subgenus of Dolichoderus: Emery, 1894c: 234; Wheeler, W.M. 1910g: 142; Emery, 1913a: 10; Wheeler, W.M. 1915h: 77; Forel, 1917: 247; Borgmeier, 1923: 84; Clark, 1930b: 252; McAreavey, 1949: 17; Creighton, 1950a: 331; Chapman & Capco, 1951: 182; Kusnezov, 1956: 28; Kusnezov, 1959: 50; Smith, D.R. 1979: 1415; Kupyanskaya, 1990: 154.
Hypoclinea as junior synonym of Dolichoderus: Forel, 1878: 386; Emery & Forel, 1879a: 455; Dalla Torre, 1893: 156; Bingham, 1903: 291; Brown, 1973b: 181 [provisional]; Shattuck, 1992c: 66; Mackay, 1993b: 6; Shattuck, 1994: 39; Bolton, 1994: 26; Bolton, 1995b: 31; Bolton, 2003: 85.
Monacis Roger, 1862a: 233. Type-species: Formica bispinosa, by subsequent designation of Wheeler, W.M. 1911f: 167.
Taxonomic history
Monacis in Dolichoderidae: Ashmead, 1905b: 384.
Monacis in Dolichoderinae, Dolichoderini: Emery, 1913a: 9; Forel, 1917: 247; all subsequent authors.
Monacis as genus: Roger, 1862a: 233; Ashmead, 1905b: 384; Brown, 1950e: 249; Kempf, 1959b: 227; Kempf, 1972a: 141; Kempf, 1972c: 253; Dlussky & Fedoseeva, 1988: 78; Mackay & Vinson, 1989b: 20; Hölldobler & Wilson, 1990: 17.
Monacis as subgenus of Dolichoderus: Emery, 1894c: 228; Wheeler, W.M. 1910g: 142; Emery, 1913a: 9; Forel, 1917: 247; Wheeler, W.M. 1922a: 688; Borgmeier, 1923: 81; Kusnezov, 1956: 27; Kusnezov, 1959: 50.
Monacis as junior synonym of Hypoclinea: Mayr, 1862: 704.
Monacis as junior synonym of Dolichoderus: Forel, 1878: 386; Emery & Forel, 1879a: 455; Dalla Torre, 1893: 156; Brown, 1973b: 182 [provisional]; Shattuck, 1992c: 66; Mackay, 1993b: 6; Shattuck, 1994: 39; Bolton, 1994: 26; Bolton, 1995b: 35; Bolton, 2003: 85.
Monoceratoclinea Wheeler, W.M. 1935c: 68 [as subgenus of Dolichoderus]. Type-species: Dolichoderus (Hypoclinea) monoceros, by original designation.
Taxonomic history
Monoceratoclinea as subgenus of Dolichoderus: Wheeler, W.M. 1935c: 68.
Monoceratoclinea as genus: Brown, 1950e: 249; Dlussky & Fedoseeva, 1988: 78; Hölldobler & Wilson, 1990: 17.
Monoceratoclinea as junior synonym of Dolichoderus: Shattuck, 1992c: 66; Shattuck, 1994: 40; Bolton, 1994: 26; Bolton, 1995b: 35; Bolton, 2003: 85.
Diceratoclinea Wheeler, W.M. 1935c: 69 [as subgenus of Dolichoderus]. Type-species: Dolichoderus scabridus, by original designation.
Taxonomic history
Diceratoclinea as subgenus of Dolichoderus: Wheeler, W.M. 1935c: 69; Chapman & Capco, 1951: 181.
Diceratoclinea as genus: Brown, 1950e: 249.
Diceratoclinea as junior synonym of Dolichoderus: Brown, 1973b: 180 [provisional]; Taylor & Brown, D.R. 1985: 93; Shattuck, 1992c: 66; Shattuck, 1994: 40; Bolton, 1994: 26; Bolton, 1995b: 26; Bolton, 2003: 85.
Acanthoclinea Wheeler, W.M. 1935c: 69 [as subgenus of Dolichoderus]. Type-species: Dolichoderus doriae, by original designation.
Taxonomic history
Acanthoclinea in Dolichoderinae, Dolichoderini: Donisthorpe, 1943f: 618.
Acanthoclinea as subgenus of Dolichoderus: Wheeler, W.M. 1935c: 69.
Acanthoclinea as genus: Brown, 1950e: 249.
Acanthoclinea as junior synonym of Dolichoderus: Taylor & Brown, D.R. 1985: 93; Shattuck, 1992c: 66; Shattuck, 1994: 40; Bolton, 1994: 26; Bolton, 1995b: 18; Bolton, 2003: 85.
Karawajewella Donisthorpe, 1944b: 59.
Taxonomic history
[Replacement name for Diabolus Karavaiev, 1926d: 424; junior homonym of Diabolus Gray, J.E. 1841: 400 (Mammalia).]
Karawajewella as junior synonym of Dolichoderus: Brown, 1973b: 181 [provisional]; Shattuck, 1992c: 66; Shattuck, 1994: 39, 40; Bolton, 1994: 26; Bolton, 1995b: 32; Bolton, 2003: 85.
Homonym replaced by Karawajewella
Diabolus Karavaiev, 1926d: 424 [as subgenus of Dolichoderus]. Type-species: Dolichoderus (Diabolus) bifurcatus (junior synonym of Dolichoderus cuspidatus), by original designation.
Taxonomic history
[Diabolus Karavaiev junior homonym of Diabolus Gray, J.E. 1841: 400 (Mammalia).]
Diabolus as subgenus of Dolichoderus: Wheeler, W.M. 1935c: 68.
Genus Dolichoderus references
Roger, 1863b: 10, 14, 15 (Dolichoderus, Hypoclinea, Monacis catalogues); Mayr, 1863: 407, 423 (Dolichoderus, Hypoclinea catalogues); Mayr, 1865: 9 (Dolichoderus, Hypoclinea diagnoses); Mayr, 1867a: 75 (Hypoclinea diagnosis); Mayr, 1868c: 54 (†Baltic Amber species key); Mayr, 1870b: 955 (Hypoclinea species key); Mayr, 1876: 79 (Australia species key); André, 1882b: 225 (Europe & Algeria species); Mayr, 1886d: 435 (U.S.A. species key); Cresson, 1887: 258 (U.S.A. catalogue); Dalla Torre, 1893: 156 (catalogue); Forel, 1895e: 462 (India & Sri Lanka species key); Bingham, 1903: 292 (India, Sri Lanka & Burma species key); Wheeler, W.M. 1905d: 306 (North America species key); Emery, 1913a: 7 (diagnosis, subgenera key, catalogue); Emery, 1913a: 9, 10 (D. (Monacis), D. (Hypoclinea), catalogues); Mann, 1916: 460 (Brazil D. (Dolichoderus) species key); Wheeler, W.M. 1922a: 688 (subgenera key); Clark, 1930b: 253 (Australia D. (Hypoclinea) species key); Creighton, 1950a: 333 (North America D. (Hypoclinea) species key); Chapman & Capco, 1951: 181 (Asia checklist); Kempf, 1959b: 230 (Monacis, all species revision, key); Wheeler, J. 1963: 149 (U.S.A., North Dakota species key); Bernard, 1967: 249 (diagnosis); Kempf, 1969: 292 (Neotropical D. (Dolichoderus) species key); Kempf, 1972a: 98, 118, 141 (Neotropical Dolichoderus, Hypoclinea, Monacis, catalogues); Kempf, 1972c: 253 (Monacis species, additions to 1959a key); Smith, D.R. 1979: 1415 (North America catalogue); Taylor & Brown, D.R. 1985: 93 (Australia catalogue); Taylor, 1987a: 24 (Australia, New Caledonia checklist); Harada, 1987: 602 (Monacis species key); Lattke, 1987b: 263 (Neotropical D. bispinosus group, key); Dlussky & Fedoseeva, 1988: 78 (synoptic classification); Johnson, 1989a: 5 (U.S.A. species key); Hölldobler & Wilson, 1990: 17 (synoptic classification); Brandão, 1991: 340, 347, 356 (Neotropical catalogue); Shattuck, 1992c: 66 (diagnosis, review of genus); Mackay, 1993b: 17 (New World, all species revision, key); Radchenko, 1994b: 112 (South Siberia species key); Shattuck, 1994: 39 (catalogue); Bolton, 1995a: 1049 (census); Bolton, 1995b: 172 (catalogue); Wu, J. & Wang, 1995: 120 (China species key); Xu, 1995a: 34 (China species key); Shattuck, 1999: 68 (Australia synopsis); Xu, 2001b: 356 (China species key); Zhou, 2001b: 153 (China, Guangxi species key); Dill, 2002: 21, 30 (southeast Asia species groups key, D. cuspidatus group revision, key); Dlussky & Rasnitsyn, 2002: 416 (diagnosis for impression fossils); Dlussky, 2002b: 54 (Baltic & Rovno amber species); Dlussky, 2008b: 509 (Europe Eocene amber species key); Heterick, 2009: 45 (south-western Australia species key).
Dolichoderinae Tapinomini Liometopum TRUE TRUE FALSEExtant: 7 species
Fossil: 9 species
Liometopum Mayr, 1861: 38. Type-species: Formica microcephala, by monotypy.
Taxonomic history
Liometopum in Formicinae: Mayr, 1861: 38 [Formicidae]; Mayr, 1862: 653 (in key) [Formicidae]; Mayr, 1865: 9 [Formicidae]; André, 1874: 168 [Formicidae].
Liometopum in Dolichoderinae: Forel, 1878: 383 [Dolichoderidae]; Emery & Forel, 1879a: 454 [Dolichoderidae]; Dalla Torre, 1893: 163; Emery, 1895j: 771; Forel, 1899c: 104; Wheeler, W.M. 1910g: 142.
Liometopum in Dolichoderidae: Ashmead, 1905b: 384.
Liometopum in Dolichoderinae, Tapinomini: Emery, 1913a: 19; Wheeler, W.M. 1915h: 95; Forel, 1917: 248; Bondroit, 1918: 87; Wheeler, W.M. 1922a: 688; Carpenter, 1930: 46; subsequent authors except those below; then Jaffe, 1993: 9; Ward, Brady, et al. 2010: 361.
Liometopum in Dolichoderinae, Liometopini: Dlussky & Fedoseeva, 1988: 77.
Liometopum in Dolichoderinae, Dolichoderini: Shattuck, 1992c: 121; Bolton, 1994: 26; Bolton, 2003: 89.
Genus Liometopum references
Roger, 1863b: 14 (catalogue); Mayr, 1863: 427 (catalogue); Mayr, 1865: 9 (diagnosis); Forel, 1878: 383 (diagnosis); André, 1882b: 220 (Europe & Algeria species); Cresson, 1887: 258 (U.S.A. catalogue); Dalla Torre, 1893: 162 (catalogue); Bingham, 1903: 289 (diagnosis); Wheeler, W.M. 1905e: 322 (North America species); Emery, 1913a: 19 (diagnosis, catalogue); Creighton, 1950a: 338 (North America species key); Chapman & Capco, 1951: 192 (Asia checklist); Gregg, 1963: 438 (U.S.A., Colorado species key); Bernard, 1967: 253 (diagnosis); Smith, D.R. 1979: 1417 (North America catalogue); Wheeler, G.C. & Wheeler, J. 1986g: 55 (U.S.A., Nevada species key); Shattuck, 1992c: 121 (diagnosis, review of genus); Shattuck, 1994: 128 (catalogue); Bolton, 1995a: 1050 (census); Bolton, 1995b: 247 (catalogue); Mackay & Mackay, 2002: 232 (U.S.A., New Mexico species key); Del Toro, Pacheco & Mackay, 2009: 306 (all species revision, key).
Dolichoderinae Tapinomini Tapinoma TRUE TRUE FALSEExtant: 64 species, 26 subspecies
Fossil: 4 species
Tapinoma Foerster, 1850a: 43. Type-species: Tapinoma collina (junior synonym of Formica erratica), by monotypy.
Taxonomic history
Tapinoma in Formicidae: Smith, F. 1858b: 55.
Tapinoma in Formicinae: Mayr, 1855: 372 [Formicidae]; Mayr, 1861: 41 [Formicidae]; Mayr, 1862: 653 (in key) [Formicidae]; Mayr, 1865: 10 [Formicidae]; André, 1874: 168 [Formicidae]; André, 1881b: 63 [Formicidae].
Tapinoma in Dolichoderinae: Forel, 1878: 385 [Dolichoderidae]; Emery & Forel, 1879a: 454 [Dolichoderidae]; Forel, 1895b: 109 [Dolichoderidae]; Dalla Torre, 1893: 164; Emery, 1895j: 771; Forel, 1899c: 101; Wheeler, W.M. 1910g: 142.
Tapinoma in Dolichoderidae: Ashmead, 1905b: 384.
Tapinoma in Dolichoderinae, Tapinomini: Emery, 1913a: 38; Arnold, 1915: 152; Forel, 1917: 248; Bondroit, 1918: 89; Wheeler, W.M. 1922a: 690; subsequent authors except those below; then Jaffe, 1993: 9; Dubovikov, 2005a: 91 (implied); Ward, Brady, et al. 2010: 361.
Tapinoma in Dolichoderinae, Dolichoderini: Shattuck, 1992c: 146; Bolton, 1994: 26; Bolton, 2003: 91.
Junior synonyms of TAPINOMA
Micromyrma Dufour, 1857: 60. Type-species: Micromyrma pygmaea, by monotypy.
Taxonomic history
[Type-species not Formica erratica, unjustified subsequent designation by Wheeler, W.M. 1911f: 167.]
Micromyrma as subgenus of Tapinoma: Menozzi, 1925a: 19; Santschi, 1928e: 472; Chapman & Capco, 1951: 192.
Micromyrma as junior synonym of Tapinoma: Mayr, 1863: 455; Forel, 1878: 385; Emery & Forel, 1879a: 454; Dalla Torre, 1893: 164; Emery, 1913a: 38; Wheeler, W.M. 1922a: 923; Smith, M.R. 1951a: 837; Shattuck, 1992c: 146; Shattuck, 1994: 140; Bolton, 1994: 26; Bolton, 2003: 91; Fisher & Bolton, 2007: 65.
Semonius Forel, 1910f: 21. Type-species: Semonius schultzei, by monotypy.
Taxonomic history
Semonius in Dolichoderinae: Wheeler, W.M. 1910g: 142.
Semonius in Dolichoderinae, Tapinomini: Emery, 1913a: 44; Arnold, 1915: 157; Forel, 1917: 247; Wheeler, W.M. 1922a: 688; all subsequent authors.
Semonius as junior synonym of Tapinoma: Shattuck, 1992c: 146; Shattuck, 1994: 140; Bolton, 1994: 26; Bolton, 2003: 91; Fisher & Bolton, 2007: 66.
Pseudaphomomyrmex Wheeler, W.M. 1920: 53. Type-species: Aphomomyrmex emeryi, by original designation.
Taxonomic history
Pseudaphomomyrmex in Formicinae, Dimorphomyrmecini: Donisthorpe, 1943g: 721.
Pseudaphomomyrmex incertae sedis in Formicinae: Dlussky & Fedoseeva, 1988: 77.
Pseudaphomomyrmex in Formicinae, Myrmelachistini: Hölldobler & Wilson, 1990: 18.
Pseudaphomomyrmex in Formicinae, Brachymyrmecini: Bolton, 1994: 50.
Pseudaphomomyrmex in Formicinae, Plagiolepidini: Wheeler, W.M. 1922a: 695; Chapman & Capco, 1951: 214; Wheeler, G.C. & Wheeler, J. 1985: 258; Bolton, 2003: 106.
Pseudaphomomyrmex in Dolichoderinae: LaPolla & Longino, 2006: 305.
Pseudaphomomyrmex as junior synonym of Aphomomyrmex: Emery, 1925b: 44; Donisthorpe, 1943g: 721.
Pseudaphomomyrmex as genus: Wheeler, W.M. 1920: 53; Chapman & Capco, 1951: 214; Bolton, 1994: 50; Bolton, 1995b: 369; Bolton, 2003: 106.
Pseudaphomomyrmex as junior synonym of Tapinoma: Fisher & Bolton, 2007: 66.
Zatapinoma Wheeler, W.M. 1928a: 20. Type-species: Zatapinoma annandalei, by original designation.
Taxonomic history
Zatapinoma in Dolichoderinae, Tapinomini: Donisthorpe, 1943g: 737; all subsequent authors.
Zatapinoma as junior synonym of Tapinoma: Shattuck, 1992c: 146; Shattuck, 1994: 140; Bolton, 1994: 26; Bolton, 2003: 91; Fisher & Bolton, 2007: 66.
Neoclystopsenella Kurian, 1955: 133. Type-species: Neoclystopsenella luffae, by monotypy.
Taxonomic history
Neoclystopsenella in Bethylidae: Kurian, 1955: 133.
Neoclystopsenella in Dolichoderinae: Brown, 1988a: 337.
Neoclystopsenella as junior synonym of Tapinoma: Brown, 1988a: 337; Shattuck, 1992c: 146; Shattuck, 1994: 140; Bolton, 1994: 26; Bolton, 2003: 91; Fisher & Bolton, 2007: 66.
Genus Tapinoma references
Mayr, 1855: 372 (diagnosis); Smith, F. 1858b: 55 (diagnosis); Roger, 1863b: 13, 14 (Tapinoma, Micromyrma catalogues); Mayr, 1863: 455 (catalogue); Mayr, 1865: 10 (diagnosis); Mayr, 1867a: 78 (diagnosis); André, 1882b: 222 (Europe & Algeria species key); Cresson, 1887: 258 (U.S.A. catalogue); Dalla Torre, 1893: 164 (catalogue); Bingham, 1903: 304 (India, Sri Lanka & Burma species key); Emery, 1913a: 38 (diagnosis, catalogue); Emery, 1913a: 44 (Semonius diagnosis, catalogue); Arnold, 1915: 152, 157 (diagnosis, South Africa species key, Semonius diagnosis); Wheeler, W.M. 1922a: 695 (Pseudaphomomyrmex diagnosis, in key). Wheeler, W.M. 1922a: 923, 927 (Afrotropical Tapinoma, Semonius catalogues); Wheeler, W.M. 1922a: 1034 (Malagasy catalogue); Emery, 1925d: 63 (Palaearctic species key); Kuznetsov-Ugamsky, 1927d: 33 (Turkestan species key); Smith, M.R. 1928a: 311 (North America species key); Kratochvíl, Novák & Snoflák, 1944: 79 (Czechoslovakia species key); Creighton, 1950a: 352 (North America species key); Chapman & Capco, 1951: 192, 196 (Asia Semonius, Tapinoma, Zatapinoma checklists); Bernard, 1967: 254 (diagnosis, Western Europe species key); Boven, 1970b: 24 (Netherlands species key); Kempf, 1972a: 246 (Neotropical catalogue); Tarbinsky, 1976: 118 (Kyrghyzstan species key); Kutter, 1977c: 178 (Switzerland species key); Arnol'di & Dlussky, 1978: 547 (former European U.S.S.R. species key); Collingwood, 1978: 87 (Iberian Peninsula species key); Smith, D.R. 1979: 1421 (North America catalogue); Schembri & Collingwood, 1981: 421 (Malta species key); Gösswald, 1985: 317 (Germany species key); Taylor & Brown, D.R. 1985: 106 (Australia catalogue); Agosti & Collingwood, 1987b: 279 (Balkans species key); Taylor, 1987a: 77 (Australia, New Caledonia & New Zealand checklist); Dlussky, Soyunov & Zabelin, 1990: 168 (Turkmenistan species key); Brandão, 1991: 380 (Neotropical catalogue); Shattuck, 1992c: 146 (diagnosis, review of genus); Arakelian, 1994: 72 (Armenia species key); Shattuck, 1994: 140 (catalogue); Bolton, 1995a: 1053 (census); Bolton, 1995b: 399 (catalogue); Wu, J. & Wang, 1995: 116 (China species key); Collingwood & Agosti, 1996: 360 (Saudi Arabia species key); Seifert, 1996b: 165 (Central Europe species key); Skinner & Allen, 1996: 40 (Britain species key); Collingwood & Prince, 1998: 20 (Portugal species key); Shattuck, 1999: 81 (Australia synopsis); Zhou, 2001b: 151 (China, Guangxi species key); Czechowski, Radchenko & Czechowska, 2002: 135 (Poland species key); Seifert, 2007: 149 (North and Central Europe species key); Heterick, 2009: 58 (south-western Australia species key); Yoshimura & Fisher, 2011: 13 (Malagasy males).
Formicinae Lasiini Lasius TRUE TRUE FALSEExtant: 100 species, 2 subspecies
Fossil: 22 species, 1 subspecies
Lasius Fabricius, 1804: 415. Type-species: Formica nigra, by subsequent designation of Bingham, 1903: 338.
Taxonomic history
[Lasius Jurine, 1801: 164 (Apidae), resurrected by Morice & Durrant, 1915: 421, later suppressed by Act of Commission.]
Lasius in Formicinae: Mayr, 1861: 49 [Formicidae]; Mayr, 1862: 652 (in key) [Formicidae]; Mayr, 1865: 8 [Formicidae]; Mayr, 1868c: 42 [Formicidae].
Lasius in Camponotinae: Forel, 1878: 374 [Camponotidae]; Emery & Forel, 1879a: 452 [Camponotidae]; Forel, 1886f: 206; Dalla Torre, 1893: 181.
Lasius in Camponotinae, Camponotini: Forel, 1886f: 206.
Lasius in Camponotinae, Formicini: Forel, 1893a: 165; Forel, 1895b: 103; Emery, 1895j: 772; Forel, 1899c: 127; Wheeler, W.M. 1910g: 143; Forel, 1912i: 89; Wheeler, W.M. 1915h: 120.
Lasius in Formicinae, Formicini: Wheeler, W.M. 1922a: 698; Carpenter, 1930: 58; Chapman & Capco, 1951: 201; Wheeler, G.C. & Wheeler, J. 1970: 651; Wheeler, G.C. & Wheeler, J. 1976b: 101; Wheeler, G.C. & Wheeler, J. 1985: 258.
Lasius in Formicinae, Acanthomyopsini: Donisthorpe, 1943f: 655.
Lasius in Formicinae, Lasius genus group: Agosti, 1991: 296.
Lasius in Formicinae, Lasiini: Ashmead, 1905b: 384; Emery, 1925b: 226; all subsequent authors except the entries above; Bolton, 1994: 50; Bolton, 2003: 22, 96.
Lasius as junior synonym of Acanthomyops: Forel, 1916: 460; Donisthorpe, 1916c: 276; Morice & Durrant, 1917: 442. Donisthorpe, 1927b: 206; Donisthorpe, 1937d: 132; Donisthorpe, 1943f: 655; Donisthorpe, 1946g: 91.
Lasius as junior synonym of Formicina: Emery, 1916a: 61; Emery, 1916b: 239; Emery, 1917a: 96; Wheeler, W.M. 1916k: 170; Bondroit, 1918: 19.
Lasius as genus: Fabricius, 1804: 415; Wheeler, W.M. 1916k: 172; Emery, 1925b: 226; Wilson, 1955a: 11; all subsequent authors.
Junior synonyms of LASIUS
Donisthorpea Morice & Durrant, 1915: 423. Type-species: Formica nigra, by original designation.
Taxonomic history
[Unnecessary replacement name for Lasius Fabricius.]
Donisthorpea in Formicinae, Acanthomyopsini: Donisthorpe, 1943f: 640; Donisthorpe, 1946g: 91.
Donisthorpea as genus: Morice & Durrant, 1915: 423; Donisthorpe, 1915d: 184.
Donisthorpea as subgenus of Formicina: Emery, 1916a: 11; Emery, 1916b: 240.
Donisthorpea as subgenus of Acanthomyops: Donisthorpe, 1927b: 229; Donisthorpe, 1943f: 640; Donisthorpe, 1946g: 91.
Donisthorpea as junior synonym of Acanthomyops: Forel, 1916: 460; Morice & Durrant, 1917: 442; Donisthorpe, 1927b: 209.
Donisthorpea as junior synonym of Lasius: Wheeler, W.M. 1916k: 172; Emery, 1925b: 226; Wilson, 1955a: 11. [Lasius and Donisthorpea share the same type-species; synonymy is therefore absolute.]
†Tylolasius Zhang, 1989: 295. Type-species: †Tylolasius inflatus, by original designation.
Taxonomic history
†Tylolasius incertae sedis in Formicinae: Bolton, 2003: 28, 131.
†Tylolasius as junior synonym of Lasius: Hong, Wu & Ren, 2001: 348.
Subgenera of LASIUS include the nominal plus the following.
Subgenus LASIUS (ACANTHOMYOPS)
Acanthomyops Mayr, 1862: 652 (diagnosis in key), 699. Type-species: Formica clavigera, by monotypy.
Taxonomic history
Acanthomyops in Formicinae: Mayr, 1862: 652 (in key) [Formicidae]; Mayr, 1865: 8 [Formicidae].
Acanthomyops in Camponotinae, Formicini: Wheeler, W.M. 1910g: 143; Forel, 1917: 249.
Acanthomyops in Formicinae, Formicini: Wheeler, W.M. 1922a: 698; Wheeler, G.C. & Wheeler, J. 1970: 651; Wheeler, G.C. & Wheeler, J. 1976b: 101; Wheeler, G.C. & Wheeler, J. 1985: 258; Jaffe, 1993: 14.
Acanthomyops in Formicinae, Acanthomyopsini: Donisthorpe, 1943f: 618.
Acanthomyops in Formicinae, Lasius genus group: Agosti, 1991: 296.
Acanthomyops in Formicinae, Lasiini: Ashmead, 1905b: 384 (misspelled as Acanthomyrmex); Emery, 1925b: 236; all subsequent authors except for those above.
Acanthomyops as junior synonym of Lasius: Mayr, 1866b: 888; Dalla Torre, 1893: 181.
Acanthomyops as subgenus of Formicina: Emery, 1916a: 11; Forel, 1917: 249.
Acanthomyops as genus: Mayr, 1862: 699; Donisthorpe, 1927b: 209; Donisthorpe, 1943f: 618; Creighton, 1950a: 426; Smith, M.R. 1958c: 150; Wing, 1968: 47; Bolton, 1994: 50; Bolton, 2003: 94.
Acanthomyops as subgenus of Lasius: Emery, 1893i: 642; Wheeler, W.M. 1910g: 143; Wheeler, W.M. 1916k: 172; Wheeler, W.M. 1922a: 698; Emery, 1925b: 236; Buren, 1950: 188; Smith, M.R. 1951a: 852; Gregg, 1963: 453; Ward, 2005: 13.
Subgenus LASIUS (DENDROLASIUS)
Dendrolasius Ruzsky, 1912: 630 [as subgenus of Lasius]. Type-species: Formica fuliginosa, by monotypy.
Taxonomic history
[Dendrolasius also described as new by Ruzsky, 1914a: 59.]
Dendrolasius as subgenus of Formicina: Forel, 1917: 249.
Dendrolasius as subgenus of Acanthomyops: Donisthorpe, 1927b: 211; Donisthorpe, 1943f: 637.
Dendrolasius as subgenus of Lasius: Ruzsky, 1912: 630; Wheeler, W.M. 1922a: 698; Emery, 1925b: 235; Wilson, 1955a: 14; all subsequent authors except the following.
Dendrolasius as junior synonym of Lasius: Brown, 1973b: 180 [provisional].
Subgenus LASIUS (CHTHONOLASIUS)
Chtonolasius Ruzsky, 1912: 630 [as subgenus of Lasius]. Type-species: Formica umbrata, by subsequent designation of Emery, 1925b: 232.
Taxonomic history
[Chtonolasius also described as new by Ruzsky, 1914a: 59. The claim by some authors (e.g. Emery, 1916a: 11; Donisthorpe, 1941f: 37; Donisthorpe, 1943f: 633) that Ruzsky nominated Lasius flavus as type-species is incorrect. Ruzsky (1912: 630) included those species "with the appearance of the yellow Lasius (Las. flavus D.G.) and its closely related species, and races (Las. carniolicus, Las. umbratus, Las. myops, Las. mixtus)". This is not a direct nomination of a type-species, nor is De Geer (= D.G.) the author of flavus: Bolton, 1995b: 24.]
Chthonolasius justified emendation of spelling: Wheeler, W.M. 1916k: 170.
Chthonolasius as junior synonym of Formicina: Wheeler, W.M. 1916k: 172; Emery, 1916a: 11.
Chthonolasius as subgenus of Acanthomyops: Donisthorpe, 1927b: 254.
Chthonolasius as subgenus of Lasius: Ruzsky, 1912: 630; Ruzsky, 1914a: 59; Wheeler, W.M. 1922a: 698; Emery, 1925b: 231; Wilson, 1955a: 13; all subsequent authors except the following.
Chthonolasius as junior synonym of Lasius: Brown, 1973b: 179 [provisional].
Subgenus LASIUS (CAUTOLASIUS)
Cautolasius Wilson, 1955a: 13 [as subgenus of Lasius]. Type-species: Formica flava, by original designation.
Taxonomic history
Cautolasius as junior synonym of Lasius: Brown, 1973b: 179 [provisional].
Cautolasius as subgenus of Lasius: Bolton, 2003: 97.
Subgenus LASIUS (AUSTROLASIUS)
Austrolasius Faber, 1967: 74 [as subgenus of Lasius]. Type-species: Lasius carniolicus, by original designation.
Taxonomic history
Austrolasius as junior synonym of Lasius: Brown, 1973b: 179 [provisional].
Autrolasius as subgenus of Lasius: Bolton, 2003: 97.
Genus Lasius references
Mayr, 1861: 49 (Europe species key); Roger, 1863b: 11 (catalogue); Mayr, 1863: 394, 425 (catalogue); Mayr, 1865: 8 (diagnosis); Mayr, 1867a: 74 (diagnosis); André, 1874: 179 (Europe species key); Forel, 1874: 46 (Switzerland species key); Forel, 1878: 374 (diagnosis); André, 1882b: 191 (Europe & Algeria species key); Cresson, 1887: 257 (U.S.A. catalogue); Provancher, 1887: 236 (Canada species key); Nasonov, 1889: 64 (Russia species key); Dalla Torre, 1893: 181 (catalogue); Emery, 1893i: 637 (North America species key); Bingham, 1903: 339 (India species key); Ruzsky, 1905b: 264 (Russian Empire species key); Wasmann, 1906: 10 (Luxemburg species key); Bondroit, 1910: 484 (Belgium species key); Wheeler, W.M. 1910e: 237 (North America L. umbratus forms, key); Stitz, 1914: 82 (Central Europe species key); Donisthorpe, 1915d: 188 (Britain species key); Emery, 1916b: 243 (Italy species key); Wheeler, W.M. 1916m: 591 (U.S.A., Connecticut species key); Bondroit, 1918: 19 (France & Belgium species key); Wheeler, W.M. 1922a: 698 (subgenera key); Emery, 1925b: 226 (diagnosis, subgenera key, catalogue); Emery, 1925b: 231 (L. (Chthonolasius) diagnosis, catalogue); Emery, 1925b: 235 (L. (Dendrolasius) diagnosis, catalogue); Emery, 1925b: 236 (L. (Acanthomyops) diagnosis, catalogue); Kuznetsov-Ugamsky, 1927e: 186 (Turkestan species key); Karavaiev, 1927c: 277 (Ukraine species key); Donisthorpe, 1927b: 206 (Britain species key); Arnol'di, 1933b: 602 (Russia species key); Karavaiev, 1936: 193 (Ukraine species key); Stärcke, 1937: 57 (Europe L. umbratus group, key); Menozzi, 1939a: 313 (Himalaya & Tibet species key); Stitz, 1939: 267 (Germany species key); Kratochvíl, 1941: 100 (Central Europe species key); Novák & Sadil, 1941: 100 (Central Europe species key); Cole, 1942: 373 (U.S.A., Utah species key); Holgersen, 1943b: 174 (Norway species key); Holgersen, 1944: 199 (Norway species key); Buren, 1944a: 296 (U.S.A., Iowa species key); Stärcke, 1944a: 157 (Europe L. niger group, key); Boven, 1947: 184 (Belgium species key); Creighton, 1950a: 418 (North America Lasius species key); Creighton, 1950a: 428 (North America Acanthomyops species key); Buren, 1950: 188 (Acanthomyops queens, key); Chapman & Capco, 1951: 201 (Asia checklist); Wilson, 1955a: 13 (subgenera review); Wilson, 1955a: 26 (Nearctic Lasius species revision, key); Wilson, 1955a: 28 (Palaearctic species revision, key); Boven, 1959: 11 (Netherlands species key); Wheeler, G.C. & Wheeler, J. 1963: 177 (U.S.A., North Dakota Lasius species key); Wheeler, G.C. & Wheeler, J. 1963: 195 (U.S.A., North Dakota Acanthomyops species key); Gregg, 1963: 453 (U.S.A., Colorado species key); Collingwood, 1963: 155 (Europe L. umbratus group species key); Collingwood, 1964: 107 (Britain species key); Bernard, 1967: 349 (diagnosis, Western Europe species key); Wing, 1968: 53 (Acanthomyops diagnosis, all species revision, key); Bourne, 1973: 19 (Britain species key); Boven, 1970b: 28 (Netherlands species key); Bolton & Collingwood, 1975: 7 (Britain species key); Tarbinsky, 1976: 133 (Kyrghyzstan species key); Boven, 1977: 135 (Belgium species key); Kutter, 1977c: 208 (Switzerland species key); Arnol'di & Dlussky, 1978: 555 (former European U.S.S.R. species key); Collingwood, 1978: 89 (Iberian Peninsula species key); Collingwood, 1979: 92 (Fennoscandia & Denmark species key); Smith, D.R. 1979: 1435 (North America Lasius catalogue); Smith, D.R. 1979: 1440 (North America Acanthomyops catalogue); Yamauchi, 1979: 147 (Japan species keys); Allred, 1982: 444 (U.S.A., Utah Acanthomyops species key); Allred, 1982: 450 (U.S.A., Utah Lasius species key); Collingwood, 1982: 283 (Himalayan species key); Gösswald, 1985: 268 (Germany species key); Wheeler, G.C. & Wheeler, J. 1986g: 64 (U.S.A., Nevada Lasius species key); Wheeler, G.C. & Wheeler, J. 1986g: 68 (U.S.A., Nevada Acanthomyops species key); Nilsson & Douwes, 1987: 70 (Norway species key); Agosti & Collingwood, 1987b: 281 (Balkans species key); Dlussky & Fedoseeva, 1988: 77 (synoptic classification); Seifert, 1988a: 143 (Europe L. (Chthonolasius) species key); Kupyanskaya, 1989: 781 (East Palaearctic L. (Dendrolasius) species key); Dlussky, Soyunov & Zabelin, 1990: 158 (Turkmenistan species key); Seifert, 1990: 1 (supplement to European L. (Chthonolasius)); Agosti & Bolton, 1990b: 149 (characters); Kupyanskaya, 1990: 211 (Far Eastern Russia species key); Morisita, Kubota, Onoyama, et al., 1991: 24 (Japan species key); Agosti, 1991: 295 (Lasius genus group diagnosis); Seifert, 1992b: 48 (Palaearctic L. (Lasius) species key); Atanasov & Dlussky, 1992: 232 (Bulgaria species key); Arakelian, 1994: 111 (Armenia species key); Bolton, 1994: 50 (synoptic classification); Radchenko, 1994b: 114 (South Siberia species key); Bolton, 1995a: 1050 (census); Bolton, 1995b: 53 (Acanthomyops catalogue); Bolton, 1995b: 221 (Lasius catalogue); Douwes, 1995: 92 (Sweden species key); Kupyanskaya, 1995: 361 (Far Eastern Russia species key); Wu, J. & Wang, 1995: 153 (China species key); Seifert, 1996b: 179 (Central Europe species key); Skinner & Allen, 1996: 42 (Britain species key); Collingwood & Prince, 1998: 22 (Portugal species key); Zhou, 2001b: 187 (China, Guangxi species key); Blacker & Collingwood, 2002: 44 (British species key); Czechowski, Radchenko & Czechowska, 2002: 154 (Poland species key); Mackay & Mackay, 2002: 237 (U.S.A., New Mexico Acanthomyops species key); Mackay & Mackay, 2002: 331 (U.S.A., New Mexico Lasius species key); Radchenko, 2005a: 93 (East Palaearctic L. (Dendrolasius) species key); Radchenko, 2005b: 190 (North Korea species key); Seifert, 2007: 159 (North and Central Europe species key); Terayama, 2009: 207 (Taiwan species key).
Formicinae Plagiolepidini Brachymyrmex TRUE TRUE FALSE41 species, 17 subspecies
Brachymyrmex Mayr, 1868b: 163. Type-species: Brachymyrmex patagonicus, by monotypy.
Taxonomic history
Brachymyrmex in Camponotinae: Forel, 1878: 375 [Camponotidae]; Dalla Torre, 1893: 174.
Brachymyrmex in Camponotinae, Plagiolepidini: Forel, 1893a: 165; Forel, 1895b: 106; Emery, 1895j: 771; Forel, 1899c: 123; Wheeler, W.M. 1910g: 143.
Brachymyrmex in Formicinae, Plagiolepidini: Ashmead, 1905b: 384.
Brachymyrmex in Camponotinae, Myrmelachistini: Forel, 1912i: 89.
Brachymyrmex in Formicinae, Dimorphomyrmecini: Emery, 1925b: 40 [subtribe Brachymyrmecini]; Donisthorpe, 1943f: 628.
Brachymyrmex incertae sedis in Formicinae: Dlussky & Fedoseeva, 1988: 77.
Brachymyrmex in Formicinae, Myrmelachistini: Wheeler, W.M. 1922a: 697; Hölldobler & Wilson, 1990: 18.
Brachymyrmex in Formicinae, Pseudolasius genus group: Agosti, 1991: 296.
Brachymyrmex in Formicinae, Brachymyrmecini: Wheeler, W.M. 1929a: 12; Kempf, 1972a: 37; Wheeler, G.C. & Wheeler, J. 1985: 258; Jaffe, 1993: 13; Bolton, 1994: 50.
Brachymyrmex in Formicinae, Plagiolepidini: Bolton, 2003: 23, 101.
Junior synonym of BRACHYMYRMEX
Bryscha Santschi, 1923b: 652 [as subgenus of Brachymyrmex]. Type-species: Brachymyrmex pilipes, by original designation.
Taxonomic history
Bryscha as junior synonym of Brachymyrmex: Brown, 1973b: 179 [provisional]; Smith, D.R. 1979: 1424; all subsequent authors.
Genus Brachymyrmex references
Forel, 1878: 375 (diagnosis); André, 1882b: 214 (Europe & Algeria species); Cresson, 1887: 255 (U.S.A. catalogue); Dalla Torre, 1893: 174 (catalogue); Wheeler, W.M. 1922a: 1036 (Malagasy catalogue); Santschi, 1923b: 652 (diagnosis, all species key); Emery, 1925b: 40 (diagnosis, catalogue); Emery, 1925b: 43 (B. (Bryscha) diagnosis, catalogue); Creighton, 1950a: 356 (North America species, review); Bernard, 1967: 279 (diagnosis); Kempf, 1972a: 37 (Neotropical catalogue); Alayo, 1974: 26 (Cuba species key); Smith, D.R. 1979: 1424 (North America catalogue); Brandão, 1991: 331 (catalogue); Bolton, 1995a: 1048 (census); Bolton, 1995b: 81 (catalogue); Quirán, et al. 2004: 275 (diagnosis).
Formicinae Plagiolepidini Lepisiota TRUE TRUE FALSE81 species, 51 subspecies
Lepisiota Santschi, 1926a: 15 [as subgenus of Acantholepis]. Type-species: Plagiolepis rothneyi, by original designation.
Taxonomic history
[Lepisiota first available replacement name for Acantholepis Mayr, 1861 (junior homonym of Acantholepis Kroyer, 1846: 98 (Pisces)), hence valid name of genus: Bolton, 1995b: 33.]
Lepisiota in Form