subfamily tribe genus species species author date country valid available current valid name original combination fossil taxonomic history Dolichoderinae TRUE TRUE FALSE

Extant: 28 genera, 692 species, 130 subspecies

Fossil: 16 genera, 103 species

Dolichoderidae Forel, 1878: 364. Type-genus: Dolichoderus.

Taxonomic history

Dolichoderinae as family: Emery, 1894g: 378 [Dolichoderidae]; Ashmead, 1905b: 384 [Dolichoderidae]; Novák & Sadil, 1941: 94 [Dolichoderidae]; Bernard, 1951: 1071 [Dolichoderidae]; Bernard, 1953b: 255 [Dolichoderidae].

Dolichoderinae as tribe of Formicidae: André, 1882a: 127 [Dolichoderidae].

Dolichoderinae as subfamily of Formicidae: Forel, 1878: 364 [Dolichoderidae]; Emery & Forel, 1879a: 454 [Dolichoderidae]; Nasonov, 1889: 26 [Dolichoderidae]; Forel, 1892j: 220 [Dolichoderidae]; Forel, 1893a: 165; Dalla Torre, 1893: 156; Forel, 1895b: 107 [Dolichoderidae]; Emery, 1895j: 771 [subfamily spelled Dolichoderini]; Emery, 1896e: 186; Forel, 1899c: 98; Bingham, 1903: 288; Wheeler, W.M. 1910g: 142; Emery, 1913a: 2; Arnold, 1915: 143; Wheeler, W.M. 1915h: 71; Wheeler, 1915g: 811 [Dolichoderides]; Arnold, 1915: 143; Donisthorpe, 1915d: 178; Forel, 1917: 247; Escherich, 1917: 2 [Dolichoderini]; Bondroit, 1918: 86 [Dolichoderitae]; Wheeler, W.M. 1920: 53; Wheeler, W.M. 1922a: 199; Borgmeier, 1923: 80; Karavaiev, 1936: 163; Smith, M.R. 1951a: 833; Clark, 1951: 16; Brown, 1954e: 29; Kempf, 1972a: 266; Wheeler, G.C. & Wheeler, J. 1972a: 41; Brown, 1973b: 169; all subsequent authors.

Dolichoderinae as formicomorph subfamily of Formicidae: Bolton, 2003: 18, 80.

Dolichoderinae as formicoid subfamily of Formicidae: Brady, Schultz, et al. 2006: 18173; Moreau, Bell, et al. 2006: 102.

Dolichoderinae as formicoid dolichoderomorph subfamily of Formicidae: Ward, 2007a: 556.

Tribes (extant) of Dolichoderinae: Bothriomyrmecini, Dolichoderini, Leptomyrmecini, Tapinomini.

Tribes (extinct) incertae sedis in Dolichoderinae: †Miomyrmecini, †Zherichiniini.

Genera (extinct) incertae sedis in Dolichoderinae: †Alloiomma, †Asymphylomyrmex, †Elaeomyrmex, †Elaphrodites, †Eldermyrmex, †Emplastus, †Eotapinoma, †Eurymyrmex, †Kotshkorkia, †Leptomyrmula, †Miomyrmex, †Petraeomyrmex, †Proiridomyrmex, †Protazteca, †Zherichinius.

Subfamily Dolichoderinae and tribes references, world

Forel, 1878: 364, 380 (diagnosis, genera); Dalla Torre, 1893: 156 (catalogue); Emery, 1895j: 771 (synoptic classification); Emery, 1896e: 186 (genera key); Handlirsch, 1907: 869 (†fossil taxa catalogue); Wheeler, W.M. 1910g: 142 (diagnosis); Emery, 1913a: 2, 6 (diagnosis, tribe key, catalogue); Emery, 1913a: 17 (Tapinomini diagnosis, genera key, catalogue); Arnold, 1915: 144, (diagnosis); Gallardo, 1916a: 3 (diagnosis); Forel, 1917: 247 (synoptic classification); Forel, 1921c: 136 (diagnosis); Wheeler, W.M. 1922a: 199, 687, 688 (diagnosis, tribe key, Tapinomini genera key); Brown & Nutting, 1950: 127 (venation, phylogeny); Brown, 1954e: 29 (phylogeny); Pavan, 1955: 135 (gastral organs); Eisner, 1957: 453 (proventriculus morphology); Bernard, 1967: 246 (diagnosis); Gotwald, 1969: 118 (mouthparts morphology); Wheeler, G.C. & Wheeler, J. 1972a: 41 (diagnosis); Brown, 1973b: 169 (genera & distribution); Wheeler, G.C. & Wheeler, J. 1976b: 61 (larvae, review & synthesis); Snelling, R.R. 1981: 401 (synoptic classification); Dazzini Valcurone & Fanfani, 1985: 1 (gastral glands); Wheeler, G.C. & Wheeler, J. 1985: 258 (synoptic classification); Billen, 1986: 173 (Dufour's gland); Billen, 1987: 278 (abdominal glands); Baroni Urbani & Wilson, 1987: 1 (†fossil Leptomyrmecini); Dlussky & Fedoseeva, 1988: 77 (synoptic classification); Hölldobler & Wilson, 1990: 9 (synoptic classification, genera keys); Baroni Urbani, Bolton & Ward, 1992: 316 (phylogeny); Shattuck, 1992b: 199 (higher classification, phylogeny); Shattuck, 1992c: 20 (revision of subfamily, genera key); Shattuck, 1994: 3 (catalogue); Bolton, 1994: 22 (diagnosis, synoptic classification, genera keys); Shattuck, 1995: 217 (phylogeny, genera); Bolton, 1995a: 1038 (census); Bolton, 1995b: 10 (catalogue); Wenseleers, Schoeters, et al., 1998: 121 (cloacal gland); Brandão, Baroni Urbani, et al. 1999: 411 (phylogeny, genera); Chiotis, Jermiin & Crozier, 2000: 108 (phylogeny, genera); Dlussky & Rasnitsyn, 2002: 416 (diagnosis for impression fossils); Bolton, 2003: 18, 80 (diagnosis, synopsis); Dubovikov, 2005a: 91 (tribes of Dolichoderinae); Brady, Schultz, et al. 2006: 18173 (phylogeny); Moreau, Bell, et al. 2006: 102 (phylogeny); Ward, Brady, et al. 2010: 342 (phylogeny, tribe-rank classification).

Regional and national faunas with keys

André, 1882a: 127 (Europe & Algeria); Nasonov, 1889: 50 (Russia); Forel, 1891b: 9 (Madagascar genera); Forel, 1895e: 460 (India & Sri Lanka); Bingham, 1903: 288 (India, Sri Lanka & Burma); Ruzsky, 1905b: 102 (Russian Empire); Bondroit, 1910: 488 (Belgium); Wheeler, W.M. 1910g: 560 (North America genera); Stitz, 1914: 77 (Central Europe); Gallardo, 1915: 34 (Argentina genera); Forel, 1915d: 40 (Switzerland); Arnold, 1915: 145 (South Africa); Donisthorpe, 1915d: 178 (Britain); Gallardo, 1916a: 12 (Argentina); Emery, 1916b: 208 (Italy); Wheeler, W.M. 1916m: 589 (U.S.A., Connecticut); Bondroit, 1918: 86 (France & Belgium); Gallardo, 1919b: 253 (Argentina genera); Soudek, 1922: 58 (Czechoslovakia); Stärcke, 1926: 117 (Netherlands); Karavaiev, 1927c: 271 (Ukraine); Donisthorpe, 1927b: 199 (Britain); Menozzi & Russo, 1930: 172 (Dominican Republic); Arnol'di, 1933b: 600 (Russia); Menozzi, 1933b: 90 (Israel genera); Karavaiev, 1936: 164 (Ukraine); Smith, M.R. 1937: 861 (Puerto Rico); Stitz, 1939: 209 (Germany); Kratochvíl, 1941: 94 (Central Europe); Novák & Sadil, 1941: 94 (Central Europe); Cole, 1942: 370 (U.S.A., Utah); Smith, M.R. 1943f: 309 (U.S.A. males); Buren, 1944a: 290 (U.S.A., Iowa); Smith, M.R. 1947f: 592 (U.S.A. genera); Creighton, 1950a: 330 (North America); Kusnezov, 1956: 27 (Argentina); Brown, 1958h: 29 (New Zealand); Kusnezov, 1959: 40 (Neotropical genera); Gregg, 1963: 341 (U.S.A., Colorado); Wheeler, G.C. & Wheeler, J. 1963: 149 (U.S.A., North Dakota); Bernard, 1967: 246 (Western Europe); Wilson & Taylor, 1967: 17 (Polynesia); Bolton, 1973a: 329 (West Africa genera); Bolton & Collingwood, 1975: 3 (Britain); Boven, 1970b: 24 (Netherlands); Snelling, R.R. & Hunt, 1976: 90 (Chile); Tarbinsky, 1976: 118 (Kyrghyzstan); Boven, 1977: 123 (Belgium); Kutter, 1977c: 170 (Switzerland); Arnol'di & Dlussky, 1978: 546 (former European U.S.S.R.); Collingwood, 1978: 80 (Iberian Peninsula); Collingwood, 1979: 32 (Fennoscandia & Denmark); Greenslade, 1979: 28 (South Australia genera); Schembri & Collingwood, 1981: 420 (Malta); Allred, 1982: 443 (U.S.A., Utah); Baroni Urbani, 1984: 80 (Neotropical genera); Gösswald, 1985: 314 (Germany); Collingwood, 1985: 242 (Saudi Arabia); Wheeler, G.C. & Wheeler, J. 1986g: 55 (U.S.A., Nevada); Agosti & Collingwood, 1987b: 278 (Balkans); Dlussky, Soyunov & Zabelin, 1990: 168 (Turkmenistan); Kupyanskaya, 1990: 152 (Far Eastern Russia); Morisita, Kubota, Onoyama, et al., 1991: 2 (Japan); Atanasov & Dlussky, 1992: 49 (Bulgaria); Lattke, in Jaffe, 1993: 147 (Neotropical genera); Arakelian, 1994: 70 (Armenia); Wu, J. & Wang, 1995: 115 (China genera); Kupyanskaya, 1995: 331 (Far Eastern Russia); Collingwood & Agosti, 1996: 359 (Saudi Arabia); Seifert, 1996b: 161 (Central Europe); Skinner & Allen, 1996: 40 (Britain); Collingwood & Prince, 1998: 20 (Portugal); Kim & Kim, 1999: 18 (Korea); Shattuck, 1999: 26, 64 (Australia genera, synopsis); Andersen, 2000: 59 (northern Australia genera); Zhou, 2001b: 149 (China, Guangxi); Czechowski, Radchenko & Czechowska, 2002: 134 (Poland); Aktaç & Radchenko, 2002: 54 (Turkey genera); Yoshimura & Onoyama, 2002: 425 (Japan genera, males); Mackay & Mackay, 2002: 221 (U.S.A., New Mexico); Palacio & Fernández, in Fernández, 2003d: 239 (Neotropical genera); Radchenko, 2005b: 186 (North Korea); Coovert, 2005: 102 (U.S.A., Ohio); Clouse, 2007b: 190 (Micronesia); Seifert, 2007: 147 (North and Central Europe); Fisher, 2009: 51 (Malagasy genera); Terayama, 2009: 196 (Taiwan); Heterick, 2009: 28 (south-western Australia genera); Boer, 2010: 14 (Benelux); Yoshimura & Fisher, 2011: 7 (Malagasy males key).

Formicinae TRUE TRUE FALSE

Extant: 49 genera, 2,831 species, 839 subspecies

Fossil: 29 genera, 193 species, 1 subspecies

Formicariae Latreille, 1809: 124. Type-genus: Formica.

Taxonomic history

Formicinae as group name: Latreille, 1809: 124 [Formicariae]; Lepeletier de Saint-Fargeau, 1835: 197 [Formicites]; Nylander, 1846a: 877 [Formicae].

Formicinae as family: Smith, F. 1858b: 1 [Formicidae]; Smith, F. 1861b: 36 [Formicidae]; Smith, 1871a: 302 [Formicidae]; André, 1882a: 125 [Formicidae]; Saunders, 1896: 18 [Formicidae]; Novák & Sadil, 1941: 97 [Formicidae]; Bernard, 1951: 1073 [Formicidae]; Bernard, 1953b: 256 [Formicidae].

Formicinae as tribe of Formicidae: André, 1874: 167 [Formicidae].

Formicinae as subfamily of Formicidae: Mayr, 1855: 286, 299 [Formicidae]; Mayr, 1861: 21 [Formicidae]; Mayr, 1862: 651 [Formicidae]; Mayr, 1865: 6 [Formicidae]; Mayr, 1868c: 24 [Formicidae]; Forel, 1870: 307 [Formicidae]; Forel, 1874: 21 [Formicidae]; Emery, 1877a: 70 [Formicidae]; André, 1881b: 54 [Formicidae]; Ashmead, 1905b: 384; Bondroit, 1918: 17 [Formicitae]; Wheeler, W.M. 1920: 53; Wheeler, W.M. 1922a: 210; Emery, 1925b: 2; Karavaiev, 1936: 172; Clark, 1951: 16; Brown, 1954e: 29; Kempf, 1972a: 266; Brown, 1973b: 169; all subsequent authors.

Formicinae as formicomorph subfamily of Formicidae: Bolton, 2003: 20, 93.

Formicinae as formicoid subfamily of Formicidae: Brady, Schultz, et al. 2006: 18173; Moreau, Bell, et al. 2006: 102.

Formicinae as formicoid formicomorph subfamily of Formicidae: Ward, 2007a: 556.

Tribes of Formicinae: Camponotini, Formicini, Gesomyrmecini, Gigantiopini, Lasiini, Melophorini, Myrmecorhynchini, Myrmoteratini, Notostigmatini, Oecophyllini, Plagiolepidini.

Genera (extinct) incertae sedis in Formicinae: †Camponotites, †Drymomyrmex, †Imhoffia, †Kyromyrma, †Leucotaphus, †Protrechina, †Zhangidris.

Hong (2002) genera (extinct) incertae sedis in Formicinae: †Curtipalpulus, †Eoleptocerites, †Eurytarsites, †Fushuniformica, †Huaxiaformica, †Leptogasteritus, †Liaoformica, †Longiformica, †Magnogasterites, †Orbicapitia, †Ovalicapito, †Ovaligastrula, †Sinoformica, †Sinotenuicapito, †Wilsonia (unresolved junior homonym).

Subfamily Formicinae references, world

Mayr, 1862: 651 (genera key); Mayr, 1865: 6 (diagnosis); Handlirsch, 1907: 859 (†fossil taxa catalogue); Dalla Torre, 1893: 171 (catalogue); Emery, 1895j: 772 (synoptic classification); Emery, 1896e: 187 (genera key); Wheeler, W.M. 1910g: 143 (diagnosis); Forel, 1912i: 88 (tribes key); Forel, 1917: 248 (synoptic classification); Arnold, 1920a: 551 (diagnosis); Forel, 1921c: 139 (diagnosis); Wheeler, W.M. 1922a: 210, 691 (diagnosis, tribes key); Emery, 1925b: 2 (diagnosis, tribe key, catalogue); Brown & Nutting, 1950: 127 (venation, phylogeny); Eisner, 1957: 465 (proventriculus morphology); Hung & Brown, 1966: 198 (gastric apex, structure); Bernard, 1967: 267 (diagnosis); Gotwald, 1969: 120 (mouthparts morphology); Wheeler, G.C. & Wheeler, J. 1972a: 41 (diagnosis); Brown, 1973b: 169 (genera, distribution); Wheeler, G.C. & Wheeler, J. 1976b: 62 (larvae, review & synthesis); Snelling, R.R. 1981: 402 (synoptic classification); Wheeler, G.C. & Wheeler, J. 1985: 258 (synoptic classification); Billen, 1986: 173 (Dufour's gland); Dlussky & Fedoseeva, 1988: 77 (synoptic classification); Hölldobler & Wilson, 1990: 9 (synoptic classification, genera keys); Agosti, 1991: 295 (genus group diagnoses); Shattuck, 1992b: 201 (phylogeny); Baroni Urbani, Bolton & Ward, 1992: 317 (phylogeny); Bolton, 1994: 42 (diagnosis, synoptic classification, genera keys); Bolton, 1995a: 1039 (census); Bolton, 1995b: 11 (catalogue); Wenseleers, Schoeters, et al. 1998: 121 (cloacal gland); Dlussky & Rasnitsyn, 2002: 417 (diagnosis for impression fossils); Bolton, 2003: 20, 93 (diagnosis, synopsis); Brady, Schultz, et al. 2006: 18173 (phylogeny); Moreau, Bell, et al. 2006: 102 (phylogeny).

Regional and national faunas with keys

Mayr, 1855: 299 (Austria); Mayr, 1861: 25 (Europe); Mayr, 1868c: 25 (†Baltic Amber); André, 1874: 167 (Europe); Forel, 1874: 22 (Switzerland); Saunders, E. 1880: 203 (Britain); André, 1882a: 126 (Europe & Algeria); Provancher, 1887: 225 (Canada); Cresson, 1887: 94 (U.S.A. genera); Nasonov, 1889: 50 (Russia); Forel, 1891b: 8 (Madagascar genera); Lameere, 1892: 62 (Belgium); Forel, 1892j: 220 (India & Sri Lanka); Bingham, 1903: 308 (India, Sri Lanka & Burma); Ruzsky, 1905b: 100 (Russian Empire); Wasmann, 1906: 7 (Luxemburg); Bondroit, 1910: 481 (Belgium); Wheeler, W.M. 1910g: 560 (North America genera); Stitz, 1914: 80 (Central Europe); Gallardo, 1915: 35 (Argentina genera); Forel, 1915d: 45 (Switzerland); Donisthorpe, 1915d: 184 (Britain); Emery, 1916b: 216 (Italy); Wheeler, W.M. 1916m: 590 (U.S.A., Connecticut); Bondroit, 1918: 17 (France & Belgium); Arnold, 1920a: 552 (South Africa); Kutter, 1920b: 134 (Switzerland); Soudek, 1922: 61 (Czechoslovakia); Lomnicki, 1925a: 160 (Poland); Stärcke, 1926: 118, 146 (Netherlands); Karavaiev, 1927c: 273 (Ukraine); Donisthorpe, 1927b: 205 (Britain); Menozzi & Russo, 1930: 172 (Dominican Republic); Arnol'di, 1933b: 601 (Russia); Menozzi, 1933b: 90 (Israel genera); Karavaiev, 1936: 173 (Ukraine); Smith, M.R. 1937: 865 (Puerto Rico); Stitz, 1939: 230 (Germany); Kratochvíl, 1941: 97 (Central Europe); Novák & Sadil, 1941: 97 (Central Europe); Cole, 1942: 373 (U.S.A., Utah); Smith, M.R. 1943f: 309 (U.S.A., males); Buren, 1944a: 292 (U.S.A., Iowa); Holgersen, 1943b: 173 (Norway); Holgersen, 1944: 199 (Norway); Smith, M.R. 1947f: 599 (U.S.A. genera); Boven, 1947: 181 (Belgium); Creighton, 1950a: 355 (North America); Kusnezov, 1956: 31 (Argentina); Brown, 1958h: 42 (New Zealand); Boven, 1959: 11 (Netherlands); Gregg, 1963: 447 (U.S.A., Colorado); Wheeler, G.C. & Wheeler, J. 1963: 160 (U.S.A., North Dakota); Collingwood, 1964: 104 (Britain); Bernard, 1967: 268 (Western Europe); Wilson & Taylor, 1967: 17 (Polynesia); Boven, 1970b: 26 (Netherlands); Kempf, 1972a: 266 (Neotropical, synoptic classification); Bolton, 1973a: 329 (West Africa genera); Bolton & Collingwood, 1975: 3 (Britain); Snelling & Hunt, 1976: 104 (Chile); Tarbinsky, 1976: 126 (Kyrghyzstan); Boven, 1977: 126 (Belgium); Kutter, 1977c: 183 (Switzerland); Arnol'di & Dlussky, 1978: 548 (former European U.S.S.R.); Collingwood, 1978: 88 (Iberian Peninsula); Collingwood, 1979: 85 (Fennoscandia & Denmark); Greenslade, 1979: 32 (South Australia genera); Schembri & Collingwood, 1981: 436 (Malta); Prins, 1983: 8 (Southern Africa genera); Allred, 1982: 444 (U.S.A., Utah); Verhaeghe, Deligne, et al., 1984: 106 (Belgium genera); Baroni Urbani, 1984: 81 (Neotropical genera); Gösswald, 1985: 263 (Germany); Collingwood, 1985: 273 (Saudi Arabia); Wheeler, G.C. & Wheeler, J. 1986g: 58 (U.S.A., Nevada); Nilsson & Douwes, 1987: 68 (Norway); Agosti & Collingwood, 1987b: 279 (Balkans); Dlussky, Soyunov & Zabelin, 1990: 124 (Turkmenistan); Kupyanskaya, 1990: 162 (Far Eastern Russia); Morisita, Kubota, Onoyama, et al., 1991: 10 (Japan); Atanasov & Dlussky, 1992: 49 (Bulgaria); Shattuck, 1992b: 199 (higher classification, phylogeny); Lattke, in Jaffe, 1993: 150 (Neotropical genera); Arakelian, 1994: 76 (Armenia); Wu, J. & Wang, 1995: 125 (China genera); Kupyanskaya, 1995: 332 (Far Eastern Russia); Collingwood & Agosti, 1996: 361 (Saudi Arabia); Seifert, 1996b: 166 (Central Europe); Skinner & Allen, 1996: 41 (Britain); Collingwood & Prince, 1998: 21 (Portugal); Shattuck, 1999: 25, 86 (Australia genera, synopsis); Andersen, 2000: 68 (northern Australia genera); Zhou, 2001b: 165 (China, Guangxi); Czechowski, Radchenko & Czechowska, 2002: 147 (Poland); Aktaç & Radchenko, 2002: 54 (Turkey genera); Yoshimura & Onoyama, 2002: 425 (Japan genera, males); Mackay & Mackay, 2002: 236 (U.S.A., New Mexico); Palacio & Fernández, in Fernández, 2003d: 242 (Neotropical genera); Radchenko, 2005b: 187 (North Korea); Coovert, 2005: 113 (U.S.A., Ohio); Clouse, 2007b: 190 (Micronesia); Seifert, 2007: 150 (North and Central Europe); Terayama, 2009: 202 (Taiwan); Heterick, 2009: 30 (south-western Australia genera); Boer, 2010: 17 (Benelux).

Myrmicinae TRUE TRUE FALSE

Extant: 141 genera, 6,087 species, 864 subspecies

Fossil: 31 genera, 130 species

Myrmicites Lepeletier de Saint-Fargeau, 1835: 169. Type-genus: Myrmica.

Taxonomic history

Myrmicinae as group name: Lepeletier de Saint-Fargeau, 1835: 169 [Myrmicites]; Nylander, 1846a: 877 [Myrmicae].

Myrmicinae as family: Smith, F. 1851: 4 [Myrmicidae]; Smith, F. 1861b: 45 [Myrmicidae]; Smith, F. 1871a: 324 [Myrmicidae]; André, 1882a: 125 [Myrmicidae]; Cresson, 1887: 93 [Myrmicidae]; Emery, 1894g: 383 [Myrmicidae]; Saunders, 1896: 18 [Myrmicidae]; Ashmead, 1905b: 383 [Myrmicidae]; Novák & Sadil, 1941: 71 [Myrmicidae]; Bernard, 1951: 1058 [Myrmicidae]; Bernard, 1953b: 222 [Myrmicidae].

Myrmicinae as subfamily of Poneridae: Smith, F. 1858b: 114 [Myrmicidae].

Myrmicinae as tribe of Formicidae: André, 1874: 167 [Myrmicidae].

Myrmicinae as subfamily of Myrmicidae: Ashmead, 1905b: 383.

Myrmicinae as subfamily of Formicidae: Mayr, 1855: 290, 299 [Myrmicidae]; Smith, F. 1857a: 70 [Myrmicidae]; Mayr, 1861: 21 [Myrmicidae]; Smith, F. 1862b: 33 [Myrmicidae]; Mayr, 1862: 738 [Myrmicidae]; Mayr, 1865: 17 [Myrmicidae]; Mayr, 1868c: 24 [Myrmicidae]; Forel, 1870: 307 [Myrmicidae]; Forel, 1874: 22 [Myrmicidae]; Emery, 1877a: 70 [Myrmicidae]; Forel, 1878: 367 [Myrmicidae]; Emery & Forel, 1879a: 456 [Myrmicidae]; André, 1881b: 64 [Myrmicidae]; Nasonov, 1889: 28 [Myrmicidae]; Forel, 1891b: 11 [Myrmicidae]; Forel, 1892j: 220 [Myrmicidae]; Forel, 1893a: 163; Dalla Torre, 1893: 53; Emery, 1895j: 768 [subfamily spelled Myrmicini]; Emery, 1896e: 179; Forel, 1899c: 30; Forel, 1902f: 520; Bingham, 1903: 105; Wheeler, W.M. 1910g: 138; Emery, 1914a: 29; Wheeler, 1915g: 806 [Myrmicides]; Wheeler, W.M. 1915h: 40; Donisthorpe, 1915d: 74; Arnold, 1916: 166; Escherich, 1917: 2 [Myrmicini]; Forel, 1917: 240 [subfamily spelled Myrmicini]; Bondroit, 1918: 14 [Myrmicitae]; Wheeler, W.M. 1920: 53; Wheeler, W.M. 1922a: 124; Emery, 1921f: 3; Karavaiev, 1934: 59; Clark, 1951: 16; Brown, 1954e: 28; Wheeler, G.C. & Wheeler, J. 1972a: 40; Brown, 1973b: 166; all subsequent authors.

Myrmicinae as myrmicomorph subfamily of Formicidae: Bolton, 2003: 52, 182.

Myrmicinae as formicoid subfamily of Formicidae: Moreau, Bell, et al. 2006: 102; Brady, Schultz, et al. 2006: 18173.

Myrmicinae as formicoid myrmicomorph subfamily of Formicidae: Ward, 2007a: 556.

Tribes of Myrmicinae: Adelomyrmecini, Ankylomyrmini, Attini, Basicerotini, Blepharidattini, Cataulacini, Cephalotini, Crematogastrini, Dacetini, Formicoxenini, Lenomyrmecini, Liomyrmecini, Melissotarsini, Meranoplini, Metaponini, Myrmecinini, Myrmicariini, Myrmicini, Paratopulini, Phalacromyrmecini, Pheidolini, Solenopsidini, Stegomyrmecini, Stenammini, Tetramoriini.

Genera (extant) incertae sedis in Myrmicinae: Diaphoromyrma, Tyrannomyrmex.

Genera (extinct) incertae sedis in Myrmicinae: †Afromyrma, †Attopsis, †Cephalomyrmex, †Electromyrmex, †Eocenidris, †Eomyrmex, †Fallomyrma, †Miosolenopsis, †Myrmecites, †Solenopsites.

Hong (2002) genera (extinct) incertae sedis in Myrmicinae: †Brachytarsites, †Clavipetiola, †Fushunomyrmex, †Orbigastrula, †Quadrulicapito, †Quineangulicapito, †Sinomyrmex, †Sphaerogasterites, †Wumyrmex.

Collective group name in Myrmicinae: †Myrmicites.

Subfamily Myrmicinae references, world

Mayr, 1865: 17 (diagnosis); Mayr, 1867a: 91 (diagnosis); Forel, 1878: 367 (diagnosis); Handlirsch, 1907: 872 (†fossil taxa catalogue); Dalla Torre, 1893: 53 (catalogue); Emery, 1895j: 768 (diagnosis); Emery, 1896e: 179 (genera key); Wheeler, W.M. 1910g: 138 (diagnosis); Emery, 1912b: 101 (phylogeny); Emery, 1914a: 34 (phylogeny, tribe key); Arnold, 1916: 164 (diagnosis); Forel, 1917: 240 (synoptic classification); Forel, 1921c: 139 (diagnosis); Emery, 1921f: 3 (diagnosis, tribes & genera key, catalogue); Wheeler, W.M. 1922a: 124, 655 (diagnosis, tribes key); Brown & Nutting, 1950: 126 (venation, phylogeny); Brown, 1954e: 28 (phylogeny); Eisner, 1957: 477 (proventriculus morphology); Bernard, 1967: 93 (diagnosis); Gotwald, 1969: 99 (mouthparts morphology); Wheeler, G.C. & Wheeler, J. 1972a: 40 (diagnosis); Brown, 1973b: 166 (genera, distribution); Wheeler, G.C. & Wheeler, J. 1976b: 52 (larvae, review & synthesis); Kugler, C. 1978a: 413 (sting structure); Kugler, C. 1978b: 267 (pygidial glands); Kugler, C. 1979c: 117 (sting, evolution); Snelling, R.R. 1981: 393 (synoptic classification); Caetano, F.H. 1984: 257 (digestive tract, morphology); Wheeler, G.C. & Wheeler, J. 1985: 257 (synoptic classification); Billen, 1986: 167 (Dufour's gland); Dlussky & Fedoseeva, 1988: 79 (synoptic classification); Hölldobler & Wilson, 1990: 9 onward (synoptic classification, genera keys); Baroni Urbani, Bolton & Ward, 1992: 317 (phylogeny); Bolton, 1994: 75 (diagnosis, synoptic classification, genera keys); Bolton, 1995a: 1040 (census); Bolton, 1995b: 13 (catalogue); Hashimoto, 1996: 354 (phylogenetic position); Baroni Urbani, 2000: 480 (phylogeny); Dlussky & Rasnitsyn, 2002: 422 (diagnosis for wingless fossils); Bolton, 2003: 52, 182 (diagnosis, synopsis); Moreau, Bell, et al. 2006: 102 (phylogeny); Brady, Schultz, et al. 2006: 18173 (phylogeny); Ward, 2007a: 556 (classification).

Regional and national faunas with keys

Mayr, 1855: 391 (Austria); Mayr, 1861: 29 (Europe); Mayr, 1868c: 79 (†Baltic Amber); André, 1874: 171 (Europe); Forel, 1874: 29 (Switzerland); Saunders, E. 1880: 213 (Britain); André, 1882c: 256 (Europe & Algeria); Cresson, 1887: 98 (U.S.A. genera); Provancher, 1887: 243 (Canada); Nasonov, 1889: 54 (Russia); Forel, 1891b: 11 (Madagascar genera); Lameere, 1892: 66 (Belgium); Forel, 1902f: 520 (India & Sri Lanka genera); Bingham, 1903: 105 (India, Sri Lanka & Burma); Ruzsky, 1905b: 103 (Russian Empire); Wasmann, 1906: 13 (Luxemburg); Bondroit, 1910: 490 (Belgium); Wheeler, W.M. 1910g: 558 (North America genera); Stitz, 1914: 55 (Central Europe); Gallardo, 1915: 32 (Argentina genera); Forel, 1915d: 8 (Switzerland); Donisthorpe, 1915d: 74 (Britain); Arnold, 1916: 166, 170 (South Africa tribes, genera); Emery, 1916b: 112 (Italy); Wheeler, W.M. 1916m: 581 (U.S.A., Connecticut); Bondroit, 1918: 90 (France & Belgium); Kutter, 1920b: 144 (Switzerland); Soudek, 1922: 20 (Czechoslovakia); Stärcke, 1926: 84 (Netherlands); Karavaiev, 1927c: 256 (Ukraine); Donisthorpe, 1927b: 77 (Britain); Menozzi & Russo, 1930: 170 (Dominican Republic); Gallardo, 1932b: 91 (Argentina, tribes); Arnol'di, 1933b: 596 (Russia); Menozzi, 1933b: 88 (Israel genera); Karavaiev, 1934: 60 (Ukraine); Smith, M.R. 1937: 829 (Puerto Rico); Stitz, 1939: 63 (Germany); Kratochvíl, 1941: 71 (Central Europe); Novák & Sadil, 1941: 71 (Central Europe); Cole, 1942: 360 (U.S.A., Utah); Smith, M.R. 1943f: 291 (U.S.A., males); Holgersen, 1943b: 166 (Norway); Holgersen, 1944: 198 (Norway); Buren, 1944a: 281 (U.S.A., Iowa); Smith, M.R. 1947f: 543 (U.S.A. genera); Boven, 1947: 170 (Belgium); Creighton, 1950a: 83 (North America); Kusnezov, 1956: 15 (Argentina); Brown, 1958h: 25 (New Zealand); Boven, 1959: 7 (Netherlands); Gregg, 1963: 288 (U.S.A., Colorado); Wheeler, G.C. & Wheeler, J. 1963: 92 (U.S.A., North Dakota); Collingwood, 1964: 94 (Britain); Bernard, 1967: 95 (Western Europe); Wilson & Taylor, 1967: 13 (Polynesia); Boven, 1970b: 9 (Netherlands); Kempf, 1972a: 263 (Neotropical, synoptic classification); Bolton, 1973a: 325 (West Africa genera); Bolton & Collingwood, 1975: 3 (Britain); Snelling, R.R. & Hunt, 1976: 70 (Chile); Tarbinsky, 1976: 19 (Kyrghyzstan); Boven, 1977: 69 (Belgium); Kutter, 1977c: 31 (Switzerland); Arnol'di & Dlussky, 1978: 524 (former European U.S.S.R.); Collingwood, 1978: 75 (Iberian Peninsula); Collingwood, 1979: 36 (Fennoscandia & Denmark); Greenslade, 1979: 20 (South Australia genera); Schembri & Collingwood, 1981: 423 (Malta); Allred, 1982: 438 (U.S.A., Utah); Baroni Urbani, 1984: 76 (Neotropical genera); Verhaeghe, Deligne, et al. 1984: 112 (Belgium genera); Gösswald, 1985: 289 (Germany); Collingwood, 1985: 245 (Saudi Arabia); Wheeler, G.C. & Wheeler, J. 1986g: 20 (U.S.A., Nevada); Nilsson & Douwes, 1987: 57 (Norway); Agosti & Collingwood, 1987b: 265 (Balkans); Dlussky, Soyunov & Zabelin, 1990: 181 (Turkmenistan); Kupyanskaya, 1990: 89 (Far Eastern Russia); Ogata, 1991b: 61 (Japan genera); Morisita, Kubota, et al. 1992: 1 (Japan); Atanasov & Dlussky, 1992: 51 (Bulgaria); Lattke, in Jaffe, 1993: 153 (Neotropical genera); Arakelian, 1994: 15 (Armenia); Wu, J. & Wang, 1995: 57 (China genera); Kupyanskaya, 1995: 327 (Far Eastern Russia); Collingwood & Agosti, 1996: 308 (Saudi Arabia); Seifert, 1996b: 108 (Central Europe); Skinner & Allen, 1996: 43 (Britain); Collingwood & Prince, 1998: 10 (Portugal); Shattuck, 1999: 39, 122 (Australia genera, synopsis); Andersen, 2000: 37 (northern Australia genera); Zhou, 2001b: 69 (China, Guangxi); Czechowski, Radchenko & Czechowska, 2002: 135 (Poland); Aktaç & Radchenko, 2002: 55 (Turkey genera); Yoshimura & Onoyama, 2002: 424 (Japan genera, males); Mackay & Mackay, 2002: 58 (U.S.A., New Mexico); Palacio & Fernández, in Fernández, 2003d: 244 (Neotropical genera and synopsis); Coovert, 2005: 32 (U.S.A., Ohio); Radchenko, 2005b: 184 (North Korea); Clouse, 2007b: 190 (Micronesia); Seifert, 2007: 110 (North and Central Europe); Terayama, 2009: 131 (Taiwan); Heterick, 2009: 36 (south-western Australia genera); Boer, 2010: 46 (Benelux); Eguchi, Bui & Yamane, 2011: 8 (Vietnam genera).

Ponerinae TRUE TRUE FALSE

Extant: 27 genera, 1,033 species, 155 subspecies

Fossil: 9 genera, 68 species, 2 subspecies

Ponerites Lepeletier de Saint-Fargeau, 1835: 185. Type-genus: Ponera.

Taxonomic history

Ponerinae as group name: Lepeletier de Saint-Fargeau, 1835: 185 [Ponérites].

Ponerinae as family: Smith, F. 1851: 6 [Poneridae]; Smith, F. 1858b: 76 [Poneridae]; Smith, F. 1861b: 44 [Poneridae]; Smith, F. 1871a: 320 [Poneridae]; André, 1882a: 125 [Poneridae]; Cresson, 1887: 93 [Poneridae]; Emery, 1894g: 379 [Poneridae]; Saunders, 1896: 18 [Poneridae]; Ashmead, 1905b: 382 [Poneridae]; Novák & Sadil, 1941: 70 [Poneridae]; Bernard, 1951: 1042 [Poneridae]; Bernard, 1953b: 185 [Poneridae].

Ponerinae as tribe of Formicidae: André, 1874: 167 [Poneridae].

Ponerinae as subfamily of Poneridae: Ashmead, 1905b: 382.

Ponerinae as subfamily of Formicidae: Mayr, 1855: 289, 299 [Poneridae]; Mayr, 1861: 21 [Poneridae]; Mayr, 1862: 712 [Poneridae]; Smith, F. 1857a: 64 [Poneridae]; Smith, F. 1862b: 31 [Poneridae]; Mayr, 1865: 11 [Poneridae]; Mayr, 1868c: 24 [Poneridae]; Forel, 1870: 307 [Poneridae]; Forel, 1874: 21 [Poneridae]; Emery, 1877a: 70 [Poneridae]; Forel, 1878: 366 [Poneridae]; Emery & Forel, 1879a: 455 [Poneridae]; André, 1881b: 64 [Poneridae]; Forel, 1892j: 220 [Poneridae]; Dalla Torre, 1893: 13; Forel, 1893a: 162; Forel, 1895b: 111 [Poneridae]; Nasonov, 1889: 27 [Poneridae]; Emery, 1895j: 766 [subfamily spelled Ponerini]; Emery, 1896e: 176; Forel, 1899c: 2; Emery, 1901a: 36; Bingham, 1903: 23; Wheeler, W.M. 1910g: 134; Emery, 1911d: 2; Wheeler, 1915g: 805 [Ponerides]; Donisthorpe, 1915d: 65; Wheeler, W.M. 1915h: 25; Arnold, 1915: 9; Escherich, 1917: 2 [Ponerini]; Forel, 1917: 235; Bondroit, 1918: 78 [Poneritae]; Wheeler, W.M. 1920: 53; Wheeler, W.M. 1922a: 56, 632, 640; Borgmeier, 1923: 37; Karavaiev, 1934: 49; Clark, 1951: 15; Brown, 1954e: 24; Wheeler, G.C. & Wheeler, J. 1972a: 39; Brown, 1973b: 165; all subsequent authors.

Ponerinae as poneromorph subfamily of Formicidae: Bolton, 2003: 42, 156.

Ponerinae as poneroid subfamily of Formicidae: Ouellette, Fisher, et al. 2006: 365; Brady, Schultz, et al. 2006: 18173; Moreau, Bell et al. 2006: 102; Ward, 2007a: 555.

Tribes of Ponerinae: Platythyreini, Ponerini, Thaumatomyrmecini.

Genera (extinct) incertae sedis in Ponerinae: †Afropone, †Eogorgites, †Eoponerites, †Furcisutura, †Longicapitia.

Subfamily Ponerinae references, world

Smith, F. 1858b: 76 (diagnosis); Mayr, 1862: 712 (genera key); Mayr, 1865: 11 (Odontomachidae, Poneridae diagnoses), Mayr, 1867a: 79, 81 (Odontomachidae, Poneridae diagnoses); Forel, 1878: 366 (diagnosis); Dalla Torre, 1893: 13 (world catalogue); Emery, 1895j: 766 (diagnosis); Emery, 1896e: 176 (genera key); Handlirsch, 1907: 879 (†fossil taxa catalogue); Wheeler, W.M. 1910g: 134 (diagnosis); Emery, 1911d: 3 (diagnosis, tribes key); Forel, 1917: 235 (synoptic classification); Forel, 1921c: 133 (diagnosis); Wheeler, W.M. 1922a: 56, 636 (diagnosis, tribes key); Brown & Nutting, 1950: 124 (venation, phylogeny); Brown, 1954e: 24 (phylogeny); Eisner, 1957: 475 (proventriculus morphology); Bernard, 1967: 79 (diagnosis); Gotwald, 1969: 25 (mouthparts morphology); Wheeler, G.C. & Wheeler, J. 1972a: 39 (diagnosis); Brown, 1973b: 165 (genera, distribution); Wheeler, G.C. & Wheeler, J. 1976b: 48 (larvae, review & synthesis); Snelling, R.R. 1981: 387 (synoptic classification); Wheeler, G.C. & Wheeler, J. 1985: 256 (synoptic classification); Billen, 1986: 168 (Dufour's gland); Fanfani & Dazzini Valcurone, 1986: 115 (exocrine glands synopsis); Dlussky & Fedoseeva, 1988: 78 (synoptic classification); Hölldobler & Wilson, 1990: 9 onward (synoptic classification, genera keys); Baroni Urbani, Bolton & Ward, 1992: 317 (phylogeny); Bolton, 1994: 153 (diagnosis, synoptic classification, genera keys); Bolton, 1995a: 1042 (census); Bolton, 1995b: 14 (catalogue); Grimaldi, Agosti & Carpenter, 1997: 20 (†Cretaceous genera); Dlussky & Rasnitsyn, 2002: 422 (diagnosis for wingless fossils); Bolton, 2003: 42, 156 (diagnosis, synopsis); Ouellette, Fisher et al. 2006: 359 (phylogeny); Brady, Schultz, et al. 2006: 18173 (phylogeny); Moreau, Bell et al. 2006: 102 (phylogeny); Ward, 2007a: 555 (classification).

.

Regional and national faunas with keys

Mayr, 1861: 28 (Europe); Mayr, 1868c: 69 (†Baltic Amber); André, 1874: 170 (Europe); Forel, 1874: 29 (Switzerland); André, 1882b: 227 (Europe & Algeria); Cresson, 1887: 97 (U.S.A., genera); Provancher, 1887: 238 (Canada); Nasonov, 1889: 54 (Russia); Forel, 1891b: 9 (Madagascar genera); Forel, 1900c: 52 (India & Sri Lanka); Bingham, 1903: 23 (India, Sri Lanka & Burma); Ruzsky, 1905b: 108 (Russian Empire); Wheeler, W.M. 1910g: 557 (North America genera); Bondroit, 1910: 489 (Belgium); Stitz, 1914: 54 (Central Europe); Gallardo, 1915: 31 (Argentina genera); Forel, 1915d: 6 (Switzerland); Arnold, 1915: 10 (South Africa); Donisthorpe, 1915d: 65 (Britain); Emery, 1916b: 98 (Italy); Wheeler, W.M. 1916m: 580 (U.S.A., Connecticut); Bondroit, 1918: 79 (France & Belgium); Gallardo, 1918b: 4 (Argentina); Soudek, 1922: 18 (Czechoslovakia); Stärcke, 1926: 82 (Netherlands); Donisthorpe, 1927b: 68 (Britain); Menozzi & Russo, 1930: 169 (Dominican Republic); Arnol'di, 1933b: 596 (Russia); Karavaiev, 1934: 50 (Ukraine); Smith, M.R. 1937: 823 (Puerto Rico); Stitz, 1939: 58 (Germany); Smith, M.R. 1943f: 278 (U.S.A. males); Buren, 1944a: 279 (U.S.A., Iowa); Smith, M.R. 1947f: 530 (U.S.A. genera); Creighton, 1950a: 31 (North America); Kusnezov, 1956: 11 (Argentina); Brown, 1958h: 11 (New Zealand); Gregg, 1963: 280 (U.S.A., Colorado); Bernard, 1967: 80 (Western Europe); Wilson & Taylor, 1967: 10 (Polynesia); Kempf, 1972a: 262 (Neotropical synoptic classification); Bolton, 1973a: 323 (West Africa genera); Bolton & Collingwood, 1975: 3 (Britain); Boven, 1977: 66 (Belgium); Kutter, 1977c: 21 (Switzerland); Arnol'di & Dlussky, 1978: 522 (former European U.S.S.R.); Collingwood, 1978: 74 (Iberian Peninsula); Collingwood, 1979: 29 (Fennoscandia & Denmark); Greenslade, 1979: 14 (South Australia genera); Francoeur, 1979: 30 (Canada, Québec); Schembri & Collingwood, 1981: 417 (Malta); Allred, 1982: 438 (U.S.A., Utah); Baroni Urbani, 1984: 75 (Neotropical genera); Collingwood, 1985: 236 (Saudi Arabia); Wheeler, G.C. & Wheeler, J. 1986g: 17 (U.S.A., Nevada); Agosti & Collingwood, 1987b: 264 (Balkans); Ogata, 1987: 101 (Japan genera); Morisita, Kubota, et al. 1989: 8 (Japan); Dlussky, Soyunov & Zabelin, 1990: 173 (Turkmenistan); Kupyanskaya, 1990: 85 (Far Eastern Russia); Atanasov & Dlussky, 1992: 51 (Bulgaria); Lattke, in Jaffe, 1993: 166 (Neotropical genera); Arakelian, 1994: 10 (Armenia); Wu, J. & Wang, 1995: 31 (China genera); Collingwood & Agosti, 1996: 308 (Saudi Arabia); Seifert, 1996b: 107 (Central Europe); Collingwood & Prince, 1998: 10 (Portugal); Kim, Kim & Kim, 1998: 145 (Korea); Shattuck, 1999: 52, 179 (Australia genera synopsis); Andersen, 2000: 19 (northern Australia genera); Zhou, 2001b: 22 (China, Guangxi); Czechowski, Radchenko & Czechowska, 2002: 133 (Poland); Aktaç & Radchenko, 2002: 53 (Turkey genera); Yoshimura & Onoyama, 2002: 436 (Japan genera, males); Mackay & Mackay, 2002: 27 (U.S.A., New Mexico); Csösz, 2003: 147 (Carpathian Basin); Palacio & Fernández, in Fernández, 2003d: 256 (Neotropical genera); Radchenko, 2005b: 184 (North Korea); Coovert, 2005: 23 (U.S.A., Ohio); Yoshimura & Fisher, 2007: 27 (Malagasy genera, males); Clouse, 2007b: 190 (Micronesia); Seifert, 2007: 108 (North and Central Europe); Bolton & Fisher, 2008c: 32 (Afrotropical genera); Fernández & Arias-Penna, 2008: 32 (Neotropical genera); Dlussky, 2009: 1071 (Eocene amber); Terayama, 2009: 101 (Taiwan genera); Heterick, 2009: 34 (south-western Australia genera); Boer, 2010: 68 (Benelux).

Dolichoderinae Tapinomini Tapinoma TRUE TRUE FALSE

Extant: 64 species, 26 subspecies

Fossil: 4 species

Tapinoma Foerster, 1850a: 43. Type-species: Tapinoma collina (junior synonym of Formica erratica), by monotypy.

Taxonomic history

Tapinoma in Formicidae: Smith, F. 1858b: 55.

Tapinoma in Formicinae: Mayr, 1855: 372 [Formicidae]; Mayr, 1861: 41 [Formicidae]; Mayr, 1862: 653 (in key) [Formicidae]; Mayr, 1865: 10 [Formicidae]; André, 1874: 168 [Formicidae]; André, 1881b: 63 [Formicidae].

Tapinoma in Dolichoderinae: Forel, 1878: 385 [Dolichoderidae]; Emery & Forel, 1879a: 454 [Dolichoderidae]; Forel, 1895b: 109 [Dolichoderidae]; Dalla Torre, 1893: 164; Emery, 1895j: 771; Forel, 1899c: 101; Wheeler, W.M. 1910g: 142.

Tapinoma in Dolichoderidae: Ashmead, 1905b: 384.

Tapinoma in Dolichoderinae, Tapinomini: Emery, 1913a: 38; Arnold, 1915: 152; Forel, 1917: 248; Bondroit, 1918: 89; Wheeler, W.M. 1922a: 690; subsequent authors except those below; then Jaffe, 1993: 9; Dubovikov, 2005a: 91 (implied); Ward, Brady, et al. 2010: 361.

Tapinoma in Dolichoderinae, Dolichoderini: Shattuck, 1992c: 146; Bolton, 1994: 26; Bolton, 2003: 91.

Junior synonyms of TAPINOMA

Micromyrma Dufour, 1857: 60. Type-species: Micromyrma pygmaea, by monotypy.

Taxonomic history

[Type-species not Formica erratica, unjustified subsequent designation by Wheeler, W.M. 1911f: 167.]

Micromyrma as subgenus of Tapinoma: Menozzi, 1925a: 19; Santschi, 1928e: 472; Chapman & Capco, 1951: 192.

Micromyrma as junior synonym of Tapinoma: Mayr, 1863: 455; Forel, 1878: 385; Emery & Forel, 1879a: 454; Dalla Torre, 1893: 164; Emery, 1913a: 38; Wheeler, W.M. 1922a: 923; Smith, M.R. 1951a: 837; Shattuck, 1992c: 146; Shattuck, 1994: 140; Bolton, 1994: 26; Bolton, 2003: 91; Fisher & Bolton, 2007: 65.

Semonius Forel, 1910f: 21. Type-species: Semonius schultzei, by monotypy.

Taxonomic history

Semonius in Dolichoderinae: Wheeler, W.M. 1910g: 142.

Semonius in Dolichoderinae, Tapinomini: Emery, 1913a: 44; Arnold, 1915: 157; Forel, 1917: 247; Wheeler, W.M. 1922a: 688; all subsequent authors.

Semonius as junior synonym of Tapinoma: Shattuck, 1992c: 146; Shattuck, 1994: 140; Bolton, 1994: 26; Bolton, 2003: 91; Fisher & Bolton, 2007: 66.

Pseudaphomomyrmex Wheeler, W.M. 1920: 53. Type-species: Aphomomyrmex emeryi, by original designation.

Taxonomic history

Pseudaphomomyrmex in Formicinae, Dimorphomyrmecini: Donisthorpe, 1943g: 721.

Pseudaphomomyrmex incertae sedis in Formicinae: Dlussky & Fedoseeva, 1988: 77.

Pseudaphomomyrmex in Formicinae, Myrmelachistini: Hölldobler & Wilson, 1990: 18.

Pseudaphomomyrmex in Formicinae, Brachymyrmecini: Bolton, 1994: 50.

Pseudaphomomyrmex in Formicinae, Plagiolepidini: Wheeler, W.M. 1922a: 695; Chapman & Capco, 1951: 214; Wheeler, G.C. & Wheeler, J. 1985: 258; Bolton, 2003: 106.

Pseudaphomomyrmex in Dolichoderinae: LaPolla & Longino, 2006: 305.

Pseudaphomomyrmex as junior synonym of Aphomomyrmex: Emery, 1925b: 44; Donisthorpe, 1943g: 721.

Pseudaphomomyrmex as genus: Wheeler, W.M. 1920: 53; Chapman & Capco, 1951: 214; Bolton, 1994: 50; Bolton, 1995b: 369; Bolton, 2003: 106.

Pseudaphomomyrmex as junior synonym of Tapinoma: Fisher & Bolton, 2007: 66.

Zatapinoma Wheeler, W.M. 1928a: 20. Type-species: Zatapinoma annandalei, by original designation.

Taxonomic history

Zatapinoma in Dolichoderinae, Tapinomini: Donisthorpe, 1943g: 737; all subsequent authors.

Zatapinoma as junior synonym of Tapinoma: Shattuck, 1992c: 146; Shattuck, 1994: 140; Bolton, 1994: 26; Bolton, 2003: 91; Fisher & Bolton, 2007: 66.

Neoclystopsenella Kurian, 1955: 133. Type-species: Neoclystopsenella luffae, by monotypy.

Taxonomic history

Neoclystopsenella in Bethylidae: Kurian, 1955: 133.

Neoclystopsenella in Dolichoderinae: Brown, 1988a: 337.

Neoclystopsenella as junior synonym of Tapinoma: Brown, 1988a: 337; Shattuck, 1992c: 146; Shattuck, 1994: 140; Bolton, 1994: 26; Bolton, 2003: 91; Fisher & Bolton, 2007: 66.

Genus Tapinoma references

Mayr, 1855: 372 (diagnosis); Smith, F. 1858b: 55 (diagnosis); Roger, 1863b: 13, 14 (Tapinoma, Micromyrma catalogues); Mayr, 1863: 455 (catalogue); Mayr, 1865: 10 (diagnosis); Mayr, 1867a: 78 (diagnosis); André, 1882b: 222 (Europe & Algeria species key); Cresson, 1887: 258 (U.S.A. catalogue); Dalla Torre, 1893: 164 (catalogue); Bingham, 1903: 304 (India, Sri Lanka & Burma species key); Emery, 1913a: 38 (diagnosis, catalogue); Emery, 1913a: 44 (Semonius diagnosis, catalogue); Arnold, 1915: 152, 157 (diagnosis, South Africa species key, Semonius diagnosis); Wheeler, W.M. 1922a: 695 (Pseudaphomomyrmex diagnosis, in key). Wheeler, W.M. 1922a: 923, 927 (Afrotropical Tapinoma, Semonius catalogues); Wheeler, W.M. 1922a: 1034 (Malagasy catalogue); Emery, 1925d: 63 (Palaearctic species key); Kuznetsov-Ugamsky, 1927d: 33 (Turkestan species key); Smith, M.R. 1928a: 311 (North America species key); Kratochvíl, Novák & Snoflák, 1944: 79 (Czechoslovakia species key); Creighton, 1950a: 352 (North America species key); Chapman & Capco, 1951: 192, 196 (Asia Semonius, Tapinoma, Zatapinoma checklists); Bernard, 1967: 254 (diagnosis, Western Europe species key); Boven, 1970b: 24 (Netherlands species key); Kempf, 1972a: 246 (Neotropical catalogue); Tarbinsky, 1976: 118 (Kyrghyzstan species key); Kutter, 1977c: 178 (Switzerland species key); Arnol'di & Dlussky, 1978: 547 (former European U.S.S.R. species key); Collingwood, 1978: 87 (Iberian Peninsula species key); Smith, D.R. 1979: 1421 (North America catalogue); Schembri & Collingwood, 1981: 421 (Malta species key); Gösswald, 1985: 317 (Germany species key); Taylor & Brown, D.R. 1985: 106 (Australia catalogue); Agosti & Collingwood, 1987b: 279 (Balkans species key); Taylor, 1987a: 77 (Australia, New Caledonia & New Zealand checklist); Dlussky, Soyunov & Zabelin, 1990: 168 (Turkmenistan species key); Brandão, 1991: 380 (Neotropical catalogue); Shattuck, 1992c: 146 (diagnosis, review of genus); Arakelian, 1994: 72 (Armenia species key); Shattuck, 1994: 140 (catalogue); Bolton, 1995a: 1053 (census); Bolton, 1995b: 399 (catalogue); Wu, J. & Wang, 1995: 116 (China species key); Collingwood & Agosti, 1996: 360 (Saudi Arabia species key); Seifert, 1996b: 165 (Central Europe species key); Skinner & Allen, 1996: 40 (Britain species key); Collingwood & Prince, 1998: 20 (Portugal species key); Shattuck, 1999: 81 (Australia synopsis); Zhou, 2001b: 151 (China, Guangxi species key); Czechowski, Radchenko & Czechowska, 2002: 135 (Poland species key); Seifert, 2007: 149 (North and Central Europe species key); Heterick, 2009: 58 (south-western Australia species key); Yoshimura & Fisher, 2011: 13 (Malagasy males).

Dolichoderinae Tapinomini Technomyrmex TRUE TRUE FALSE

Extant: 93 species

Fossil: 4 species

Technomyrmex Mayr, 1872: 147. Type-species: Technomyrmex strenuus, by monotypy.

Taxonomic history

Technomyrmex in Dolichoderinae: Forel, 1878: 380 [Dolichoderidae]; Dalla Torre, 1893: 166; Emery, 1895j: 771; Wheeler, W.M. 1910g: 142.

Technomyrmex in Dolichoderinae, Tapinomini: Emery, 1913a: 42; Arnold, 1915: 147; Forel, 1917: 248; Wheeler, W.M. 1922a: 690; subsequent authors except those below; then Jaffe, 1993: 9; Ward, Brady, et al. 2010: 361.

Technomyrmex in Dolichoderinae, Dolichoderini: Shattuck, 1992c: 153; Bolton, 1994: 26; Bolton, 2003: 92.

Junior synonyms of TECHNOMYRMEX

Engramma Forel, 1905b: 180. Type-species: Engramma lujae, by monotypy.

Taxonomic history

Engramma in Dolichoderinae: Wheeler, W.M. 1910g: 142.

Engramma in Dolichoderinae, Tapinomini: Emery, 1913a: 38; Forel, 1917: 248; Wheeler, W.M. 1922a: 690; all subsequent authors.

Engramma as junior synonym of Technomyrmex: Shattuck, 1992c: 153; Shattuck, 1994: 156; Bolton, 1994: 26; Bolton, 2003: 92; Bolton, 2007a: 4.

Tapinoptera Santschi, 1925g: 348 [as subgenus of Tapinoma]. Type-species: Tapinoma vexatum, by monotypy.

Taxonomic history

Tapinoptera as junior synonym of Tapinoma: Shattuck, 1992c: 146; Shattuck, 1994: 140; Bolton, 1994: 26; Bolton, 2003: 91.

Tapinoptera as junior synonym of Technomyrmex: Fisher & Bolton, 2007: 66 (in text); Bolton, 2007a: 4.

Aphantolepis Wheeler, W.M. 1930d: 44. Type-species: Aphantolepis quadricolor, by monotypy.

Taxonomic history

Aphantolepis in Formicinae, Lasiini: Wheeler, W.M. 1930d: 44.

Aphantolepis in Formicinae, Prenolepidini: Donisthorpe, 1943f: 624.

Aphantolepis as junior synonym of Technomyrmex: Brown, 1953h: 5; Shattuck, 1992c: 153; Shattuck, 1994: 156; Bolton, 1994: 26; Bolton, 2003: 92; Bolton, 2007a: 4.

Genus Technomyrmex references

Forel, 1878: 380 (diagnosis); Dalla Torre, 1893: 166 (catalogue); Bingham, 1903: 301 (India, Sri Lanka & Burma species key); Emery, 1913a: 38 (Engramma diagnosis, catalogue); Emery, 1913a: 42 (diagnosis, catalogue); Arnold, 1915: 147 (diagnosis, South Africa species key); Wheeler, W.M. 1922a: 201, 208, 209 (Engramma, Tapinoma, Technomyrmex diagnoses); Wheeler, W.M. 1922a: 202 (Engramma species key); Wheeler, W.M. 1922a: 922, 925 (Afrotropical Engramma, Technomyrmex catalogues); Wheeler, W.M. 1922a: 1034 (Malagasy catalogue); Taylor & Brown, D.R. 1985: 106 (Australia catalogue); Taylor, 1987a: 77 (Australia, New Caledonia & New Zealand checklist); Shattuck, 1992c: 153 (diagnosis, review of genus); Shattuck, 1994: 156 (catalogue); Bolton, 1995a: 1053 (census); Bolton, 1995b: 402 (catalogue); Wu, J. & Wang, 1995: 118 (China species key); Collingwood & Agosti, 1996: 360 (Saudi Arabia species key); Shattuck, 1999: 83 (Australia synopsis); Zhou, 2001b: 158 (China, Guangxi species key); Bolton, 2007a: 1 (all species revision; Afrotropical & West Palaearctic species key: 13; Malagasy species key: 43; East Palaearctic, Oriental, Malesian & Polynesian species key: 59; Austral species key: 108; New World species key: 119); Fernández & Guerrero, 2008: 110 (New World synopsis, key); Terayama, 2009: 200 (Taiwan species key); Yoshimura & Fisher, 2011: 14 (Malagasy males); Sharaf, et al. 2011: 15 (Arabian Peninsula species key).

Tribes (extinct) incertae sedis in DOLICHODERINAE

Formicinae Lasiini Anoplolepis TRUE TRUE FALSE

9 species, 5 subspecies

Anoplolepis Santschi, 1914b: 123 [as subgenus of Plagiolepis]. Type-species: Formica longipes (junior primary homonym in Formica; Formica gracilipes first available replacement name: Bolton, 1995b: 67), by monotypy.

Taxonomic history

Anoplolepis in Camponotinae, Plagiolepidini: Forel, 1917: 249; Arnold, 1922: 593.

Anoplolepis in Formicinae, Plagiolepidini: Wheeler, W.M. 1922a: 696; Emery, 1925b: 16; Santschi, 1926a: 13; all subsequent authors except the following.

Anoplolepis in Formicinae [part in Oecophylla genus group, part in Pseudolasius genus group]: Agosti, 1991: 295, 296.

Anoplolepis in Formicinae, Lasiini: Bolton, 2003: 22, 95.

Anoplolepis as subgenus of Plagiolepis: Santschi, 1914b: 123; Forel, 1917: 249; Arnold, 1922: 593; Wheeler, W.M. 1922a: 696.

Anoplolepis as genus: Emery, 1925b: 16; Santschi, 1926a: 13; all subsequent authors.

Junior synonym of ANOPLOLEPIS

Zealleyella Arnold, 1922: 579 (diagnosis in key) [as subgenus of Plagiolepis]. Type-species: Formica custodiens, by subsequent designation of Santschi, 1926a: 14.

Taxonomic history

Zealleyella as subgenus of Plagiolepis: Arnold, 1922: 579.

Zealleyella as subgenus of Anoplolepis: Santschi, 1926a: 14.

Zealleyella as junior synonym of Anoplolepis: Brown, 1973b: 185 [provisional]; Bolton, 2003: 96, 267.

Genus Anoplolepis references

Arnold, 1922: 593 (South Africa, species); Wheeler, W.M. 1922a: 931, 1036 (Afrotropical, Malagasy catalogues); Emery, 1925b: 16 (diagnosis, catalogue); Santschi, 1926a: 13 (subgenera diagnoses); Santschi, 1937g: 83 (subgenera key); Chapman & Capco, 1951: 213 (Asia checklist); Kempf, 1972a: 22 (Neotropical catalogue); Prins, 1982: 235 (South Africa A. (Zealleyella) partial revision, key); Taylor & Brown, D.R. 1985: 108 (Australia catalogue); Dlussky & Fedoseeva, 1988: 77 (synoptic classification); Bolton, 1994: 51 (synoptic classification); Bolton, 1995a: 1048 (census); Bolton, 1995b: 66 (catalogue); Collingwood & Agosti, 1996: 362 (Saudi Arabia species key); Shattuck, 1999: 88 (Australia synopsis); Bolton, 2003: 267 (genus diagnosis).

Formicinae Plagiolepidini Aphomomyrmex TRUE TRUE FALSE

1 species

Aphomomyrmex Emery, 1899e: 493. Type-species: Aphomomyrmex afer, by subsequent designation of Wheeler, W.M. 1911f: 159.

Taxonomic history

Aphomomyrmex in Formicinae, Plagiolepidini: Ashmead, 1905b: 384.

Aphomomyrmex in Camponotinae, Plagiolepidini: Wheeler, W.M. 1910g: 143.

Aphomomyrmex in Camponotinae, Myrmelachistini: Forel, 1912i: 89; Forel, 1917: 249; Arnold, 1920a: 552.

Aphomomyrmex in Formicinae, Dimorphomyrmecini: Emery, 1925b: 44 [subtribe Brachymyrmecini]; Donisthorpe, 1943f: 624.

Aphomomyrmex incertae sedis in Formicinae: Dlussky & Fedoseeva, 1988: 77.

Aphomomyrmex in Formicinae, Myrmelachistini: Wheeler, W.M. 1922a: 697; Hölldobler & Wilson, 1990: 18.

Aphomomyrmex in Formicinae, Pseudolasius genus group: Agosti, 1991: 296.

Aphomomyrmex in Formicinae, Brachymyrmecini: Wheeler, W.M. 1929a: 12; Wheeler, G.C. & Wheeler, J. 1985: 258; Bolton, 1994: 50.

Aphomomyrmex in Formicinae, Plagiolepidini: Bolton, 2003: 23, 101.

[Aphomyrmex Ashmead, 1905a: 111, incorrect subsequent spelling.]

Genus Aphomomyrmex references

Arnold, 1920a: 552 (South Africa species); Wheeler, W.M. 1922a: 940 (Afrotropical catalogue); Emery, 1925b: 44 (diagnosis, catalogue); Snelling, R.R. 1979b: 3 (review of genus); Bolton, 1995a: 1048 (census); Bolton, 1995b: 74 (catalogue).

Formicinae Plagiolepidini Lepisiota TRUE TRUE FALSE

81 species, 51 subspecies

Lepisiota Santschi, 1926a: 15 [as subgenus of Acantholepis]. Type-species: Plagiolepis rothneyi, by original designation.

Taxonomic history

[Lepisiota first available replacement name for Acantholepis Mayr, 1861 (junior homonym of Acantholepis Kroyer, 1846: 98 (Pisces)), hence valid name of genus: Bolton, 1995b: 33.]

Lepisiota in Formicinae, Plagiolepidini: Bolton, 1994: 51; Bolton, 2003: 23.

[Lepisiota incorrectly as junior synonym of Acanthomyrmex: Brown, 1973b: 181; Snelling, R.R. 1981: 397. In both publications Acanthomyrmex is an error for Acantholepis.]

Junior synonym of LEPISIOTA

Baroniurbania Pagliano & Scaramozzino, 1990: 4. Unnecessary replacement name for Acantholepis Mayr (junior homonym).

Taxonomic history

Baroniurbania as junior synonym of Lepisiota: Bolton, 1994: 51; Bolton, 2003: 102.

Baroniurbania as subgenus of Lepisiota: Dietrich, 2004: 324.

Genus Lepisiota references

[Note. Entries prior to Bolton, 1995b: 44, refer to genus as Acantholepis.]

Roger, 1863b: 11 (catalogue); Mayr, 1863: 394 (catalogue); Mayr, 1865: 9 (diagnosis); Forel, 1878: 378 (diagnosis); André, 1882b: 210 (Europe & Algeria species key); Forel, 1892a: 41 (all species key); Dalla Torre, 1893: 171 (catalogue); Forel, 1894c: 411 (India & Sri Lanka species key); Bingham, 1903: 315 (India, Sri Lanka & Burma species key); Arnold, 1920a: 554 (diagnosis, South Africa species key); Wheeler, W.M. 1922a: 214, 934, 1036 (diagnosis, Afrotropical, Malagasy catalogues); Emery, 1925b: 23 (diagnosis, catalogue); Kuznetsov-Ugamsky, 1929c: 480 (Turkestan species key); Finzi, 1936: 188 (Egypt species key); Chapman & Capco, 1951: 209 (Asia checklist); Collingwood, 1985: 292 (Saudi Arabia species key); Agosti & Collingwood, 1987b: 281 (Balkans species key); Atanasov & Dlussky, 1992: 205 (Bulgaria species key); Xu, 1994c: 232 (China species key); Arakelian, 1994: 80 (Armenia species key); Bolton, 1994: 51 (synoptic classification); Bolton, 1995a: 1050 (census); Bolton, 1995b: 226 (catalogue); Wu, J. & Wang, 1995: 128 (China species key); Collingwood & Agosti, 1996: 363 (Saudi Arabia species key); Zhou, 2001b: 167 (China, Guangxi species key).

Homonym replaced by LEPISIOTA

Acantholepis Mayr, 1861: 42. Type-species: Hypoclinea frauenfeldi, by monotypy.

Taxonomic history

[Junior homonym of Acantholepis Kroyer, 1846: 98 (Pisces)].

Acantholepis in Formicinae: Mayr, 1861: 42 [Formicinae]; Mayr, 1862: 652 [Formicidae]; Mayr, 1865: 9 [Formicidae].

Acantholepis in Camponotinae: Forel, 1878: 378 [Camponotidae]; Emery & Forel, 1879a: 453 [Camponotidae]; Dalla Torre, 1893: 171.

Acantholepis in Camponotinae, Plagiolepidini: Forel, 1886f: 212; Forel, 1893a: 165; Emery, 1895j: 771; Wheeler, W.M. 1910g: 143; Forel, 1912i: 88; Forel, 1917: 249; Arnold, 1920a: 554.

Acantholepis in Formicinae, Plagiolepidini: Ashmead, 1905b: 384; Wheeler, W.M. 1922a: 696; Emery, 1925b: 23; Santschi, 1926a: 15; all subsequent authors except the following.

Acantholepis in Formicinae, Pseudolasius genus group: Agosti, 1991: 296.

[Achantilepis Santschi, 1935b: 274, incorrect subsequent spelling.]

[Pseudacantholepis Bernard, 1953b: 256 (attributed to Santschi) [as subgenus of Acantholepis]. Unavailable name. Proposed without designation of type-species and therefore unavailable. Species included by Bernard (1953) are all referable to Lepisiota: Bolton, 1995b: 44.]

Formicinae Plagiolepidini Plagiolepis TRUE TRUE FALSE

Extant: 57 species, 19 subspecies

Fossil: 10 species

Plagiolepis Mayr, 1861: 42. Type-species: Formica pygmaea, by monotypy.

Taxonomic history

Plagiolepis in Formicinae: Mayr, 1861: 42 [Formicinae]; Mayr, 1862: 652 [Formicidae]; Mayr, 1865: 7 [Formicidae]; Mayr, 1868c: 36 [Formicidae].

Plagiolepis in Camponotinae: Forel, 1878: 378 [Camponotidae]; Emery & Forel, 1879a: 453 [Camponotidae]; Dalla Torre, 1893: 172.

Plagiolepis in Camponotinae, Plagiolepidini: Forel, 1886f: 212; Forel, 1893a: 165; Emery, 1895j: 771; Wheeler, W.M. 1910g: 143; Forel, 1912i: 88; Wheeler, W.M. 1915h: 100; Forel, 1917: 249; Arnold, 1920a: 578.

Plagiolepis in Formicinae, Plagiolepidini: Ashmead, 1905b: 384; Bondroit, 1918: 18; Wheeler, W.M. 1922a: 696; Emery, 1925b: 19; Santschi, 1926a: 14; all subsequent authors except the one below; Bolton, 2003: 23, 105.

Plagiolepis in Formicinae, Pseudolasius genus group: Agosti, 1991: 296.

Junior synonyms of PLAGIOLEPIS

†Rhopalomyrmex Mayr, 1868c: 41. Type-species: †Rhopalomyrmex pygmaeus, by monotypy.

Taxonomic history

†Rhopalomyrmex in Formicinae: Mayr, 1868c: 41 [Formicidae].

†Rhopalomyrmex in Camponotinae: Forel, 1878: 376 [Camponotidae]; Dalla Torre, 1893: 175.

†Rhopalomyrmex in Camponotinae, Myrmelachistini: Forel, 1912i: 89.

†Rhopalomyrmex in Camponotinae, Plagiolepidini: Wheeler, W.M. 1910g: 143; Wheeler, W.M. 1915h: 103.

†Rhopalomyrmex in Formicinae, Plagiolepidini: Donisthorpe, 1943g: 724; Bolton, 1994: 51.

†Rhopalomyrmex as junior synonym of Plagiolepis: Dlussky, 1997: 624; Bolton, 2003: 105.

Anacantholepis Santschi, 1914e: 36 [as subgenus of Plagiolepis]. Type-species: Plagiolepis (Anacantholepis) decora, by original designation.

Taxonomic history

Anacantholepis as genus: Chapman & Capco, 1951: 212.

Anacantholepis as subgenus of Plagiolepis: Santschi, 1914e: 36 Forel, 1917: 249; Arnold, 1922: 581; Wheeler, W.M. 1922a: 696; Emery, 1925b: 22; Bolton, 1994: 51.

Anacantholepis as junior synonym of Plagiolepis: Brown, 1973b: 178 [provisional]; Bolton, 2003: 105, 268.

Aporomyrmex Faber, 1969: 52. Type-species: Aporomyrmex ampeloni, by original designation.

Taxonomic history

Aporomyrmex in Formicinae, Plagiolepidini: Faber, 1969: 52 (by implication).

Aporomyrmex as junior synonym of Plagiolepis: Brown, 1973b: 178 [provisional]; Bolton, 1994: 51; all subsequent authors.

Paraplagiolepis Faber, 1969: 65 [as subgenus of Plagiolepis]. Type-species: Plagiolepis xene, by monotypy.

Taxonomic history

Paraplagiolepis as junior synonym of Plagiolepis: Brown, 1973b: 183 [provisional]; Bolton, 1994: 51; all subsequent authors.

Genus Plagiolepis references

Roger, 1863b: 11 (catalogue); Mayr, 1863: 442 (catalogue); Mayr, 1865: 7 (diagnosis); Mayr, 1867a: 73 (diagnosis); Forel, 1878: 376, 378 (†Rhopalomyrmex, Plagiolepis diagnoses); André, 1882b: 208 (Europe & Algeria species key); Dalla Torre, 1893: 172, 175 (Plagiolepis, †Rhopalomyrmex, catalogue); Forel, 1894c: 414 (India & Sri Lanka species key); Bingham, 1903: 320 (India, Sri Lanka & Burma species key); Arnold, 1920a: 578 (diagnosis, subgenera key); Arnold, 1922: 579 (South Africa species key); Wheeler, W.M. 1922a: 211, 696, 928, 1035  (diagnosis, subgenera key, Afrotropical, Malagasy catalogues); Emery, 1925b: 19 (diagnosis, catalogue); Emery, 1925b: 22 (P. (Anacantholepis) diagnosis, catalogue); Stitz, 1939: 231 (Germany species key); Kratochvíl, 1941: 98 (Central Europe species key); Novák & Sadil, 1941: 98 (Central Europe species key); Chapman & Capco, 1951: 212, 213 (Asia Anacantholepis, Plagiolepis checklists); Bernard, 1967: 272 (diagnosis, Western Europe species key); Kempf, 1972a: 205 (Neotropical catalogue); Kutter, 1977c: 185 (Switzerland species key); Collingwood, 1978: 89 (Iberian Peninsula species key); Smith, D.R. 1979: 1422 (North America, catalogue); Gösswald, 1985: 264 (Germany species key); Collingwood, 1985: 297 (Saudi Arabia species key); Taylor & Brown, D.R. 1985: 130 (Australia catalogue); Taylor, 1987a: 55 (Australia, New Caledonia checklist); Agosti & Collingwood, 1987b: 280 (Balkans species key); Radchenko, 1989b: 153 (European former U.S.S.R. species key); Dlussky, Soyunov & Zabelin, 1990: 161 (Turkmenistan species key); Morisita, Kubota, et al. 1991: 16 (Japan species key); Atanasov & Dlussky, 1992: 201 (Bulgaria species key); Arakelian, 1994: 78 (Armenia species key); Bolton, 1994: 51 (synoptic classification); Bolton, 1995a: 1052 (census); Bolton, 1995b: 334 (catalogue); Wu, J. & Wang, 1995: 130 (China species key); Radchenko, 1996a: 178 (Central & Southern Palaearctic species key); Collingwood & Agosti, 1996: 362 (Saudi Arabia species key); Seifert, 1996b: 178 (Central Europe species key); Collingwood & Prince, 1998: 22 (Portugal species key); Shattuck, 1999: 106 (Australia synopsis); Seifert, 2007: 158 (North and Central Europe species key); Terayama, 2009: 213 (Taiwan species key); Heterick, 2009: 106 (south-western Australia species key); Dlussky, 2010: 72 (Eocene European amber species key).

Formicinae Camponotini Camponotus TRUE TRUE FALSE

Extant: 1,058 species, 495 subspecies

Fossil: 31 species

Camponotus Mayr, 1861: 35. Type-species: Formica ligniperda, by subsequent designation of Bingham, 1903: 347.

Taxonomic history

[Type-species not Formica herculeana, unjustified subsequent designation by Forel, 1914a: 259, repeated in Arnold, 1922: 612.]

Camponotus in Formicinae: Mayr, 1861: 35 [Formicidae]; Mayr, 1862: 652 (in key) [Formicidae]; Mayr, 1865: 6 [Formicidae]; Mayr, 1868c: 26 [Formicidae].

Camponotus in Camponotinae: Forel, 1878: 367 [Camponotidae]; Emery & Forel, 1879a: 447 [Camponotidae]; Forel, 1892j: 223 [Camponotidae]; Dalla Torre, 1893: 171.

Camponotus in Camponotinae, Camponotini: Forel, 1886f: 141; Forel, 1893a: 165; Forel, 1895b: 101; Emery, 1895j: 772; Forel, 1899c: 130; Ashmead, 1905b: 384; Wheeler, W.M. 1910g: 144; Forel, 1912i: 89; Wheeler, W.M. 1915h: 138; Forel, 1917: 250; Arnold, 1922: 610.

Camponotus in Formicinae, Camponotini: Bondroit, 1918: 65; Wheeler, W.M. 1922a: 703; Emery, 1925b: 59; all subsequent authors except the following.

Camponotus in Formicinae, Formica genus group: Agosti, 1991: 295.

Camponotus as subgenus of Formica: Smith, F. 1871a: 306.

Camponotus as genus: all authors except the entry above.

Junior synonym of CAMPONOTUS

†Paleosminthurus Pierce & Gibron, 1962: 146. Type-species: †Paleosminthurus juliae, by monotypy.

Taxonomic history

†Paleosminthurus in Collembola, family †Paleosminthuridae: Pierce & Gibron, 1962: 146.

†Paleosminthurus incertae sedis in Formicidae: Najt, 1987: 152.

†Paleosminthurus incertae sedis in Formicinae: Hölldobler & Wilson, 1990: 18.

†Paleosminthurus in †Paleosmithurinae: Bolton, 1994: 187 [provisional subfamily status]; Bolton, 1995b: 14, 311; Bolton, 2003: 76, 263.

†Paleosminthurus as junior synonym of Camponotus: Snelling (pers. comm., 2004). [Synonymy remained unpublished at his death in 2008.]

†Shanwangella Zhang, 1989: 307. Type-species: †Shanwangella palaeoptera, by original designation.

Taxonomic history

†Shanwangella as junior synonym of Camponotus: Hong & Wu, 2000: 20.

Subgenera of CAMPONOTUS include the nominal plus the following.

All the subgenera were given as provisional junior synonyms of Camponotus by Brown, 1973b: 179 - 185. Brown's list was repeated in Hölldobler & Wilson, 1990: 18 but with all subgenera listed as junior synonyms; subgeneric status was reinstated in Bolton, 1994: 50. These consecutive entries are not repeated individually below.

Subgenus CAMPONOTUS (COLOBOPSIS)

Colobopsis Mayr, 1861: 38. Type-species: Formica truncata, by subsequent designation of Bingham, 1903: 342.

Taxonomic history

Colobopsis in Formicinae: Mayr, 1861: 38 [Formicidae]; Mayr, 1862: 652 (in key) [Formicidae]; Mayr, 1865: 7 [Formicidae].

Colobopsis in Camponotinae: Forel, 1878: 368 [Camponotidae]; Emery & Forel, 1897: 449 [Camponotidae].

Colobopsis in Camponotinae, Camponotini: Forel, 1886f: 193; Emery, 1895j: 772; Ashmead, 1905b: 384.

Colobopsis in Formicinae, Camponotini: Bondroit, 1918: 65; Emery, 1925b: 144; all subsequent authors.

Colobopsis as genus: Mayr, 1861: 38; Mayr, 1862: 652; Mayr, 1865: 7; Mayr, 1870b: 940; Forel, 1878: 368; Forel, 1879a: 125; Emery & Forel, 1879a: 449; André, 1882b: 159; Bingham, 1903: 342; Forel, 1886f: 193; Ashmead, 1905b: 384; Bondroit, 1918: 66; Jaffe, 1993: 14.

Colobopsis as subgenus of Camponotus: Emery, 1889b: 517; Dalla Torre, 1893: 219; Forel, 1893a: 165; Forel, 1893b: 435; Emery, 1895j: 772; Wheeler, W.M. 1904b: 139; Wheeler, W.M. 1910g: 144; Forel, 1912i: 90; Forel, 1914a: 263; Forel, 1917: 251; Emery, 1920b: 247; Arnold, 1922: 613; Wheeler, W.M. 1922a: 708; Emery, 1925b: 144; Donisthorpe, 1943f: 634; Creighton, 1950a: 390; Chapman & Capco, 1951: 222; Kempf, 1972a: 42; Smith, D.R., 1979: 1433; Bolton, 1994: 50.

Junior synonyms of CAMPONOTUS (COLOBOPSIS)

Condylomyrma Santschi, 1928c: 72 [as subgenus of Camponotus]. Type-species: Camponotus (Condylomyrma) bryani, by monotypy.

Taxonomic history

Condylomyrma as subgenus of Camponotus: Santschi, 1928c: 72.

Condylomyrma as junior synonym of Colobopsis: Wheeler, W.M. 1934e: 422 (in text).

[Campylomyrma Wheeler, W.M. 1934e: 421 (in text), incorrect subsequent spelling.]

Dolophra Wu, J. & Wang, 1994: 35. Type-species: Dolophra politae, by original designation.

Taxonomic history

Dolophra as genus: Wu, J. & Wang, 1994: 35; Wu, J. & Wang, 1995: 158.

Dolophra as junior synonym of Camponotus: Bolton, 1995b: 27.

Dolophra as junior synonym of Camponotus (Colobopsis): Bolton, 2003: 113, 268.

Subgenus CAMPONOTUS (MAYRIA)

Mayria Forel, 1878: 369. Type-species: Mayria madagascarensis (junior secondary homonym in Camponotus, replaced by Camponotus repens), by monotypy.

Taxonomic history

Mayria in Camponotinae: Forel, 1878: 369 [Camponotidae]; Dalla Torre, 1893: 219.

Mayria in Camponotinae, Camponotini: Forel, 1893a: 165; Emery, 1895j: 772; Ashmead, 1905b: 384; Wheeler, W.M. 1910g: 144.

Mayria as subgenus of Camponotus: Forel, 1894e: 227; Forel, 1914a: 262; Forel, 1917: 251; Wheeler, W.M. 1922a: 706; Emery, 1925b: 121; all subsequent authors.

Junior synonym of CAMPONOTUS (MAYRIA)

Myrmosaga Forel, 1912i: 92 [as subgenus of Camponotus]. Type-species: Camponotus kelleri, by subsequent designation of Wheeler, W.M. 1913a: 81.

Taxonomic history

[Type-species not Camponotus quadrimaculatus, unjustified subsequent designation by Forel, 1914a: 260; repeated in Arnold, 1922: 613.]

Myrmosaga as subgenus of Camponotus Forel, 1912i: 92; Forel, 1914a: 260; Forel, 1917: 250; Emery, 1920b: 241; Arnold, 1922: 613; Wheeler, W.M. 1922a: 706; all subsequent authors.

Myrmosaga as junior synonym of Mayria: Emery, 1925b: 121.

Subgenus CAMPONOTUS (RHINOMYRMEX)

Rhinomyrmex Forel, 1886f: 192. Type-species: Rhinomyrmex klaesii, by monotypy.

Taxonomic history

Rhinomyrmex in Camponotinae: Forel, 1886f: 192 [Camponotii]; Dalla Torre, 1893: 219.

Rhinomyrmex in Camponotinae, Camponotini: Forel, 1893a: 165; Emery, 1895j: 772; Ashmead, 1905b: 384; Wheeler, W.M. 1910g: 144.

Rhinomyrmex as genus: Forel, 1886f: 192; Forel, 1893a: 165; Dalla Torre, 1893: 219; Emery, 1895j: 772; Wheeler, W.M. 1910g: 144.

Rhinomyrmex as subgenus of Camponotus: Emery, 1896d: 374; Forel, 1914a: 264; Forel, 1917: 251; Emery, 1920b: 247; Wheeler, W.M. 1922a: 708; Emery, 1925b: 142; all subsequent authors.

Subgenus CAMPONOTUS (DENDROMYRMEX)

Dendromyrmex Emery, 1895j: 772. Type-species: Formica chartifex, by subsequent designation of Wheeler, W.M. 1911f: 161.

Taxonomic history

Dendromyrmex in Camponotinae, Camponotini: Emery, 1895j: 772; Ashmead, 1905b: 384; Forel, 1912i: 89; Wheeler, W.M. 1910g: 144; Forel, 1917: 250.

Dendromyrmex in Formicinae, Camponotini: Wheeler, W.M. 1922a: 700; Emery, 1925b: 172; all subsequent authors except the following.

Dendromyrmex in Formicinae, Formica genus group: Agosti, 1991: 295.

Dendromyrmex as subgenus of Camponotus: Fernández, 2002: 51.

Subgenus CAMPONOTUS (DINOMYRMEX)

Dinomyrmex Ashmead, 1905b: 384. Type-species: Formica gigas, by original designation.

Taxonomic history

Dinomyrmex in Camponotinae, Camponotini: Ashmead, 1905b: 384.

Dinomyrmex as genus: Ashmead, 1905b: 384.

Dinomyrmex as subgenus of Camponotus: Forel, 1913b: 350; Forel, 1914a: 259; Forel, 1917: 250; Emery, 1920b: 236; Arnold, 1922: 612; Wheeler, W.M. 1922a: 704; Emery, 1925b: 69; all subsequent authors.

Junior synonym of CAMPONOTUS (DINOMYRMEX)

Myrmogigas Forel, 1912i: 91 [as subgenus of Camponotus]. Type-species: Formica gigas, by subsequent designation of Wheeler, W.M. 1913a: 81.

Taxonomic history

Myrmogigas as subgenus of Camponotus: Forel, 1912i: 91.

Myrmogigas as junior synonym of Dinomyrmex: Forel, 1913b: 350 (in text); Forel, 1914a: 259; Wheeler, W.M. 1922a: 704. [Dinomyrmex and Myrmogigas share the same type-species, synonymy is therefore absolute.]

Subgenus CAMPONOTUS (TANAEMYRMEX)

Tanaemyrmex Ashmead, 1905b: 384. Type-species: Formica longipes (junior primary homonym in Formica, replaced by Camponotus etiolipes), by original designation.

Taxonomic history

Tanaemyrmex in Camponotinae, Camponotini: Ashmead, 1905b: 384.

Tanaemyrmex as genus: Ashmead, 1905b: 384.

Tanaemyrmex as subgenus of Camponotus: Emery, 1925b: 75; all subsequent authors.

Junior synonym of CAMPONOTUS (TANAEMYRMEX)

Myrmoturba Forel, 1912i: 91 [as subgenus of Camponotus]. Type-species: Formica maculata, by subsequent designation of Wheeler, W.M. 1913a: 82.

Taxonomic history

Myrmoturba as subgenus of Camponotus: Forel, 1912i: 91; Forel, 1914a: 259; Forel, 1917: 250; Emery, 1920b: 235; Arnold, 1922: 612; Wheeler, W.M. 1922a: 704.

Myrmoturba as junior synonym of Tanaemyrmex: Emery, 1925b: 75; all subsequent authors.

Subgenus CAMPONOTUS (ORTHONOTOMYRMEX)

Orthonotomyrmex Ashmead, 1906: 31.

Taxonomic history

[Replacement name for Orthonotus Ashmead, 1905b: 384; junior homonym of Orthonotus Westwood, in Stephens, 1829: 344 (Hemiptera). Type-species not Formica lateralis, unjustified subsequent designation by Forel, 1914a: 264; repeated in Arnold, 1922: 613.]

Orthonotomyrmex as subgenus of Camponotus: Forel, 1913i: 350 (in text); Forel, 1914a: 264; Forel, 1917: 251; Emery, 1920b: 244; Arnold, 1922: 613; Wheeler, W.M. 1922a: 707; Emery, 1925b: 124; all subsequent authors.

Homonym replaced by ORTHONOTOMYRMEX

Orthonotus Ashmead, 1905b: 384. Type-species: Formica sericea, by original designation.

Taxonomic history

[Junior homonym of Orthonotus Westwood, in Stephens, 1829: 344 (Hemiptera).]

Orthonotus in Camponotinae, Camponotini: Ashmead, 1905b: 384.

Subgenus CAMPONOTUS (MYRMAMBLYS)

Myrmamblys Forel, 1912i: 90 [as subgenus of Camponotus]. Type-species: Camponotus reticulatus, by subsequent designation of Wheeler, W.M. 1913a: 80.

Taxonomic history

[Type-species not Camponotus fastigatus, unjustified subsequent designation by Forel, 1914a: 263; repeated in Arnold, 1922: 613.]

Myrmamblys as subgenus of Camponotus: Forel, 1912i: 90; Forel, 1914a: 263; Forel, 1917: 251; Emery, 1920b: 248; Arnold, 1922: 613; Wheeler, W.M. 1922a: 708; Emery, 1925b: 136; Santschi, 1926c: 601; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOTHRIX)

Myrmothrix Forel, 1912i: 91 [as subgenus of Camponotus]. Type-species: Formica abdominalis (junior primary homonym; Formica atriceps is first available replacement name), by subsequent designation of Wheeler, W.M. 1913a: 81.

Taxonomic history

[Type-species not Formica rufipes, unjustified subsequent designation by Forel, 1914a: 260.]

Myrmothrix as subgenus of Camponotus: Forel, 1912i: 91; Forel, 1914a: 260; Forel, 1917: 250; Emery, 1920b: 236; Emery, 1925b: 107; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOSERICUS)

Myrmosericus Forel, 1912i: 91 [as subgenus of Camponotus]. Type-species: Formica rufoglauca, by subsequent designation of Wheeler, W.M. 1913a: 81.

Taxonomic history

Myrmosericus as subgenus of Camponotus: Forel, 1912i: 91; Forel, 1914a: 259; Forel, 1917: 250; Emery, 1920b: 236; Arnold, 1922: 613; Wheeler, W.M. 1922a: 704; Emery, 1925b: 104; all subsequent authors.

[See note under Myrmosaulus.]

Subgenus CAMPONOTUS (MYRMOPHYMA)

Myrmophyma Forel, 1912i: 91 [as subgenus of Camponotus]. Type-species: Camponotus capito, by subsequent designation of Wheeler, W.M. 1913a: 81.

Taxonomic history

[Type-species not Camponotus quadrisectus, unjustified subsequent designation by Forel, 1914a: 261; repeated in Arnold, 1922: 612.]

Myrmophyma as subgenus of Camponotus: Forel, 1912i: 91; Forel, 1914a: 261; Forel, 1917: 250; Emery, 1920b: 239; Arnold, 1922: 612; Wheeler, W.M. 1922a: 706; Emery, 1925b: 109; all subsequent authors.

Junior synonym of CAMPONOTUS (MYRMOPHYMA)

Myrmocamelus Forel, 1914a: 261 [as subgenus of Camponotus]. Type-species: Formica ephippium, by original designation.

Taxonomic history

[Type-species not Camponotus gambeyi, unjustified subsequent designation by Forel, 1922: 101. Myrmocamelus also described as new by Forel, 1915b: 102.]

Myrmocamelus as subgenus of Camponotus: Forel, 1914a: 261; Forel, 1915b: 102; Forel, 1917: 250; Forel, 1922: 101.

Myrmocamelus as junior synonym of Myrmophyma: Emery, 1920b: 257; Wheeler, W.M. 1921a: 18; Emery, 1925b: 109.

Subgenus CAMPONOTUS (MYRMOTREMA)

Myrmotrema Forel, 1912i: 91 [as subgenus of Camponotus]. Type-species: Camponotus foraminosus, by subsequent designation of Wheeler, W.M. 1913a: 81.

Taxonomic history

Myrmotrema as subgenus of Camponotus: Forel, 1912i: 91; Forel, 1914a: 262; Forel, 1917: 250; Emery, 1920b: 245; Arnold, 1922: 613; Wheeler, W.M. 1922a: 707; Emery, 1925b: 130; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOBRACHYS)

Myrmobrachys Forel, 1912i: 91 [as subgenus of Camponotus]. Type-species: Formica senex, by subsequent designation of Wheeler, W.M. 1913a: 81.

Taxonomic history

Myrmobrachys as subgenus of Camponotus: Forel, 1912i: 91; Forel, 1914a: 262; Forel, 1917: 251; Emery, 1920b: 251; Wheeler, W.M. 1922a: 709; Emery, 1925b: 161; all subsequent authors.

Subgenus CAMPONOTUS (MYRMENTOMA)

Myrmentoma Forel, 1912i: 92 [as subgenus of Camponotus]. Type-species: Formica lateralis, by subsequent designation of Wheeler, W.M. 1913a: 80.

Taxonomic history

Myrmentoma as junior synonym of Orthonotomyrmex: Forel, 1913b: 350 (in text); Forel, 1914a: 264; Wheeler, W.M. 1922a: 970.

Myrmentoma as subgenus of Camponotus: Forel, 1912i: 92; Emery, 1920b: 243; Wheeler, W.M. 1922a: 707; Emery, 1925a: 62; Emery, 1925b: 116; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOSPHINCTA)

Myrmosphincta Forel, 1912i: 92 [as subgenus of Camponotus]. Type-species: Formica sexguttata, by subsequent designation of Wheeler, W.M. 1913a: 81.

Taxonomic history

[Type-species not Formica cinerascens, unjustified subsequent designation by Forel, 1914a: 265; repeated in Arnold, 1922: 614.]

Myrmosphincta as subgenus of Camponotus: Forel, 1912i: 92; Forel, 1914a: 265; Forel, 1917: 251; Emery, 1920b: 238; Arnold, 1922: 614; Wheeler, W.M. 1922a: 708; Emery, 1925b: 151; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOTARSUS)

Myrmotarsus Forel, 1912i: 92 [as subgenus of Camponotus]. Type-species: Formica mistura, by subsequent designation of Wheeler, W.M. 1913a: 81.

Taxonomic history

[Type-species not Formica irritabilis, unjustified subsequent designation by Forel, 1914a: 260.]

Myrmotarsus as subgenus of Camponotus: Forel, 1912i: 92; Forel, 1914a: 260; Forel, 1917: 250; Emery, 1920b: 238; Wheeler, W.M. 1922a: 705; Emery, 1925b: 134; all subsequent authors.

Subgenus CAMPONOTUS (MYRMEPOMIS)

Myrmepomis Forel, 1912i: 92 [as subgenus of Camponotus]. Type-species: Formica sericeiventris, by subsequent designation of Wheeler, W.M. 1913a: 81.

Taxonomic history

[Type-species not Formica fulvopilosa, unjustified subsequent designation by Forel, 1914a: 265; repeated in Arnold, 1922: 614.]

Myrmepomis as subgenus of Camponotus: Forel, 1912i: 92; Forel, 1914a: 265; Forel, 1917: 251; Emery, 1920b: 245; Arnold, 1922: 614; Wheeler, W.M. 1922a: 705; Emery, 1925b: 170; all subsequent authors.

Junior synonym of CAMPONOTUS (MYRMEPOMIS)

Myrmolophus Emery, 1920b: 237 [as subgenus of Camponotus]. Type-species: Formica sericeiventris, by original designation.

Taxonomic history

Myrmolophus as subgenus of Camponotus: Emery, 1920b: 237.

Myrmolophus as junior synonym of Myrmepomis: Wheeler, W.M. 1921a: 17; Wheeler, W.M. 1922a: 705; Emery, 1925b: 170. [Myrmepomis and Myrmolophus share the same type-species, synonymy is therefore absolute.]

Subgenus CAMPONOTUS (MYRMOGONIA)

Myrmogonia Forel, 1912i: 92 [as subgenus of Camponotus]. Type-species: Camponotus laminatus, by subsequent designation of Wheeler, W.M. 1913a: 81.

Taxonomic history

Myrmogonia as subgenus of Camponotus: Forel, 1912i: 92; Forel, 1914a: 261; Forel, 1917: 250; Emery, 1920b: 240; Wheeler, W.M. 1922a: 706; Emery, 1925b: 143; all subsequent authors.

Subgenus CAMPONOTUS (MYRMEURYNOTA)

Myrmeurynota Forel, 1912i: 92 [as subgenus of Camponotus]. Type-species: Camponotus eurynotus, by subsequent designation of Wheeler, W.M. 1913a: 81.

Taxonomic history

[Type-species not Camponotus gilviventris, unjustified subsequent designation by Forel, 1914a: 266.]

Myrmeurynota as subgenus of Camponotus: Forel, 1912i: 92; Forel, 1914a: 266; Forel, 1917: 251; Emery, 1920b: 251; Wheeler, W.M. 1922a: 710; Emery, 1925b: 167; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOPSAMMA)

Myrmopsamma Forel, 1914a: 261 [as subgenus of Camponotus]. Type-species: Camponotus mystaceus, by original designation.

Taxonomic history

Myrmopsamma as subgenus of Camponotus: Forel, 1914a: 261; Forel, 1917: 250; Emery, 1920b: 246; Arnold, 1922: 612; Wheeler, W.M. 1922a: 708; Emery, 1925b: 70; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOMALIS)

Myrmomalis Forel, 1914a: 263 [as subgenus of Camponotus]. Type-species: Camponotus depressus, by original designation.

Taxonomic history

Myrmomalis as subgenus of Camponotus: Forel, 1914a: 263; Forel, 1917: 251; Emery, 1920b: 251; Wheeler, W.M. 1922a: 710; Emery, 1925b: 169; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOPLATYS)

Myrmoplatys Forel, 1916: 460 [as subgenus of Camponotus]. Type-species: Camponotus korthalsiae, by original designation.

Taxonomic history

Myrmoplatys as genus: Wheeler, W.M. 1936e: 219 (in text); Donisthorpe, 1943f: 670.

Myrmoplatys as subgenus of Camponotus: Forel, 1916: 460; Forel, 1917: 251; Emery, 1920b: 238; Wheeler, W.M. 1922a: 705; Emery, 1925b: 135; Bolton, 1994: 50.

Subgenus CAMPONOTUS (MYRMAPHAENUS)

Myrmaphaenus Emery, 1920b: 237 [as subgenus of Camponotus]. Type-species: Camponotus leydigi, by original designation.

Taxonomic history

Myrmaphaenus as subgenus of Camponotus: Emery, 1920b: 237; Wheeler, W.M. 1922a: 705; Emery, 1925b: 152; all subsequent authors.

Junior synonyms of CAMPONOTUS (MYRMAPHAENUS)

Paracolobopsis Emery, 1920b: 249 [as subgenus of Camponotus]. Type-species: Camponotus salvini, by original designation.

Taxonomic history

Paracolobopsis as subgenus of Camponotus: Emery, 1920b: 249; Wheeler, W.M. 1922a: 709.

Paracolobopsis as junior synonym of Myrmaphaenus: Emery, 1925b: 152; Kempf, 1972a: 44.

Neomyrmamblys Wheeler, W.M. 1921a: 19 [as subgenus of Camponotus]. Type-species: Camponotus fastigatus, by subsequent designation of Santschi, 1921f: 311.

Taxonomic history

Neomyrmamblys as subgenus of Camponotus: Wheeler, W.M. 1921a: 19; Wheeler, W.M. 1922a: 708.

Neomyrmamblys as junior synonym of Myrmaphaenus: Emery, 1925b: 152; Kempf, 1972a: 44.

Subgenus CAMPONOTUS (MYRMONESITES)

Myrmonesites Emery, 1920b: 242 [as subgenus of Camponotus]. Type-species: Camponotus putatus, by original designation.

Taxonomic history

Myrmonesites as subgenus of Camponotus: Emery, 1920b: 242; Wheeler, W.M. 1922a: 706; Emery, 1925b: 123.

[Myrmensites Donisthorpe, 1943f: 666, incorrect subsequent spelling.]

Subgenus CAMPONOTUS (MYRMOPYTIA)

Myrmopytia Emery, 1920b: 243 [as subgenus of Camponotus]. Type-species: Camponotus imitator, by original designation.

Taxonomic history

Myrmopytia as subgenus of Camponotus: Emery, 1920b: 243; Wheeler, W.M. 1922a: 707; Emery, 1925b: 114; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOTEMNUS)

Myrmotemnus Emery, 1920b: 246 [as subgenus of Camponotus]. Type-species: Camponotus moeschi, by original designation.

Taxonomic history

Myrmotemnus as junior synonym of Myrmamblys: Wheeler, W.M. 1921a: 19; Wheeler, W.M. 1922a: 708; Emery, 1925b: 137; Donisthorpe, 1943f: 671 (anachronism).

Myrmotemnus as subgenus of Camponotus: Emery, 1920b: 246; Santschi, 1926c: 601.

Subgenus CAMPONOTUS (PSEUDOCOLOBOPSIS)

Pseudocolobopsis Emery, 1920b: 249 [as subgenus of Camponotus]. Type-species: Camponotus macrocephalus, by original designation.

Taxonomic history

Pseudocolobopsis as subgenus of Camponotus: Emery, 1920b: 249; Wheeler, W.M. 1922a: 709; Emery, 1925b: 157; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOSTENUS)

Myrmostenus Emery, 1920b: 250 [as subgenus of Camponotus]. Type-species: Camponotus mirabilis, by original designation.

Taxonomic history

Myrmostenus as subgenus of Camponotus: Emery, 1920b: 250; Wheeler, W.M. 1922a: 709; Emery, 1925b: 161; all subsequent authors.

Subgenus CAMPONOTUS (HYPERCOLOBOPSIS)

Hypercolobopsis Emery, 1920b: 250 [as subgenus of Camponotus]. Type-species: Colobopsis paradoxa, by original designation.

Taxonomic history

Hypercolobopsis as subgenus of Camponotus: Emery, 1920b: 250; Wheeler, W.M. 1922a: 709; Emery, 1925b: 160; all subsequent authors.

Junior synonym of CAMPONOTUS (HYPERCOLOBOPSIS)

Neocolobopsis Borgmeier, 1928b: 65 [as subgenus of Camponotus]. Type-species: Camponotus (Neocolobopsis) scrobifer (junior synonym of Camponotus coriolanus), by original designation.

Taxonomic history

Neocolobopsis as subgenus of Camponotus Borgmeier, 1928b: 65.

Neocolobopsis as junior synonym of Hypercolobopsis: Kempf, 1968b: 411; Kempf, 1972a: 43.

Subgenus CAMPONOTUS (MYRMOPIROMIS)

Myrmopiromis Wheeler, W.M. 1921a: 17 [as subgenus of Camponotus]. Type-species: Formica fulvopilosa, by subsequent designation of Wheeler, W.M. 1922a: 707.

Taxonomic history

Myrmopiromis as subgenus of Camponotus: Wheeler, W.M. 1921a: 17; Wheeler, W.M. 1922a: 707; Emery, 1925b: 1127.

Subgenus CAMPONOTUS (MYRMOSAULUS)

Myrmosaulus Wheeler, W.M. 1921a: 18 [as subgenus of Camponotus]. Type-species: Formica cinerascens, by subsequent designation of Wheeler, W.M. 1922a: 705.

Taxonomic history

[Type-species not Formica singularis, unjustified subsequent designation by Emery, 1925b: 113, repeated in Donisthorpe, 1943f: 670. Note that Donisthorpe, 1932c: 445, states that C. cinerascens belongs in subgenus Myrmosericus. If so then Myrmosaulus would fall as a junior synonym of Myrmosericus.]

Myrmosaulus as subgenus of Camponotus: Wheeler, W.M. 1921a: 18; Wheeler, W.M. 1922a: 705; Emery, 1925b: 112; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOCLADOECUS)

Myrmocladoecus Wheeler, W.M. 1921a: 19 [as subgenus of Camponotus]. Type-species: Camponotus latangulus, by subsequent designation of Wheeler, W.M. 1922a: 709.

Taxonomic history

Myrmocladoecus as subgenus of Camponotus: Wheeler, W.M. 1921a: 19; Wheeler, W.M. 1922a: 709; Emery, 1925b: 166; all subsequent authors.

Subgenus CAMPONOTUS (MANNIELLA)

Manniella Wheeler, W.M. 1921a: 19 [as subgenus of Camponotus]. Type-species: Camponotus sphaericus, by original designation.

Taxonomic history

Manniella subgenus of Camponotus: Wheeler, W.M. 1921a: 19; Wheeler, W.M. 1922a: 710; Emery, 1925b: 159; all subsequent authors.

Subgenus CAMPONOTUS (MYRMISOLEPIS)

Myrmisolepis Santschi, 1921f: 310 [as subgenus of Camponotus]. Type-species: Camponotus epinotalis, by original designation.

Taxonomic history

Myrmisolepis as subgenus of Camponotus: Santschi, 1921f: 310; Emery, 1925b: 133; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOPELTA)

Myrmopelta Santschi, 1921f: 310 [as subgenus of Camponotus]. Type-species: Camponotus arminius, by original designation.

Taxonomic history

Myrmopelta as junior synonym of Myrmamblys: Emery, 1925b: 137.

Myrmopelta as subgenus of Camponotus: Santschi, 1921f: 310; Wheeler, W.M. 1922a: 704 (footnote); Santschi, 1926a: 16; Santschi, 1926c: 603; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOPLATYPUS)

Myrmoplatypus Santschi, 1921f: 311 [as subgenus of Camponotus]. Type-species: Camponotus platytarsus, by original designation.

Taxonomic history

Myrmoplatypus as subgenus of Camponotus: Santschi, 1921f: 311; Emery, 1925b: 169; all subsequent authors.

Subgenus CAMPONOTUS (MYRMEPINOTUS)

Myrmepinotus Santschi, 1921f: 312 [as subgenus of Camponotus]. Type-species: Camponotus echinoploides, by original designation.

Taxonomic history

Myrmepinotus as subgenus of Camponotus: Santschi, 1921f: 312; Emery, 1925b: 126; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOXYGENYS)

Myrmoxygenys Emery, 1925b: 70 [as subgenus of Camponotus]. Type-species: Camponotus caesar, by monotypy.

Taxonomic history

Myrmoxygenys as subgenus of Camponotus: Emery, 1925b: 70; all subsequent authors.

Subgenus CAMPONOTUS (KARAVAIEVIA)

Karavaievia Emery, 1925b: 115 [as subgenus of Camponotus]. Type-species: Camponotus exsectus, by original designation.

Taxonomic history

Karavaievia as subgenus of Camponotus: Emery, 1925b: 115; all subsequent authors.

Subgenus CAMPONOTUS (MYRMODIRACHIS)

Myrmodirachis Emery, 1925b: 168 [as subgenus of Camponotus]. Type-species: Camponotus heathi, by original designation.

Taxonomic history

Myrmodirachis as subgenus of Camponotus: Emery, 1925b: 168; all subsequent authors.

Subgenus CAMPONOTUS (MYRMESPERA)

Myrmespera Santschi, 1926b: 247 [as subgenus of Camponotus]. Type-species: Camponotus (Myrmespera) debellator, by subsequent designation of Santschi, 1926c: 604.

Taxonomic history

Myrmespera as subgenus of Camponotus: Santschi, 1926b: 247; Santschi, 1926c: 603; all subsequent authors.

Subgenus CAMPONOTUS (PARAMYRMAMBLYS)

Paramyrmamblys Santschi, 1926c: 604 [as subgenus of Camponotus].Type-species: Camponotus (Myrmamblys) ostiarius, by original designation.

Taxonomic history

Paramyrmamblys as subgenus of Camponotus: Santschi, 1926c: 604; all subsequent authors.

Subgenus CAMPONOTUS (MYRMACRHAPHE)

Myrmacrhaphe Santschi, 1926c: 607 [as subgenus of Camponotus]. Type-species: Camponotus conradti, by original designation.

Taxonomic history

Myrmacrhaphe as subgenus of Camponotus: Santschi, 1926c: 607; all subsequent authors.

Subgenus CAMPONOTUS (THLIPSEPINOTUS)

Thlipsepinotus Santschi, 1928e: 483 [as subgenus of Camponotus]. Type-species: Camponotus claripes, by original designation.

Taxonomic history

Thlipsepinotus as subgenus of Camponotus: Santschi, 1928e: 483; all subsequent authors.

Subgenus CAMPONOTUS (MYRMOPALPELLA)

Myrmopalpella Stärcke, 1934: 30 [as subgenus of Camponotus]. Type-species: Camponotus megalonyx, by monotypy.

Taxonomic history

Myrmopalpella as genus: Wheeler, W.M. 1936e: 219 (in text); Donisthorpe, 1943f: 669.

Myrmopalpella as subgenus of Camponotus: Bolton, 1995b: 37.

Genus Camponotus references

Mayr, 1861: 35 (Europe species key); Roger, 1863b: 1, 9 (Camponotus, Colobopsis catalogues); Mayr, 1863: 397, 403 (Camponotus, Colobopsis catalogues); Mayr, 1865: 6, 7 (Camponotus, Colobopsis diagnosis); Mayr, 1867a: 67 (Colobopsis diagnois); Mayr, 1868c: 27 (†Baltic Amber species key); Mayr, 1870a: 373 (Colombia + Panama (= New Grenada) species key); Mayr, 1870b: 941 (Colobopsis species key); André, 1874: 175 (Europe species key); Forel, 1874: 38 (Switzerland species key); Mayr, 1876: 58 (Australia species key); Mayr, 1878: 869 (all Dendromyrmex species key (as part of Camponotus)); Forel, 1878: 367, 368 (Camponotus, Colobopsis diagnoses); André, 1882a: 138 (Europe & Algeria species key); André, 1882b: 159 (Europe & Algeria Colobopsis species key); Cresson, 1887: 255 (U.S.A. Camponotus, Colobopsis catalogue); Provancher, 1887: 228 (Canada species key); Nasonov, 1889: 57 (Russia species key); Forel, 1891b: 71, 216 (Madagascar species key); Forel, 1892j: 223 (India & Sri Lanka species key); Dalla Torre, 1893: 219 (Rhinomyrmex, Mayria, Camponotus catalogues); Emery, 1893i: 667 (North America species key); Forel, 1893b: 435 (India & Sri Lanka C. (Colobopsis) species key); Emery, 1895j: 772 (synoptic classification); Emery, 1896d: 370, 378 (species groups, catalogue); Emery, 1898a: 225 (additions to 1896a catalogue); Emery, 1903 (South America species key); Bingham, 1903: 343, 348 (India, Sri Lanka & Burma Colobopsis, Camponotus species keys); Ruzsky, 1905b: 185 (Russian Empire species key); Emery, 1908a: 183 (Palaearctic C. herculeanus & C. maculatus groups species key); Bondroit, 1910: 486 (Belgium species key); Wheeler, W.M. 1910d: 297 (North America species key); Forel, 1914a: 259, 266 (subgenera diagnoses, species listed by subgenus); Stitz, 1914: 94 (Central Europe species key); Santschi, 1915c: 266 (C. (Myrmotrema) species key); Emery, 1916b: 227 (Italy species key); Wheeler, W.M. 1916m: 600 (U.S.A., Connecticut species key); Mann, 1916: 489 (Dendromyrmex, all species key); Bondroit, 1918 (France & Belgium species key); Emery, 1920b: 235 (subgenera, diagoses); Emery, 1920c: 8 (Old World C. maculatus group, checklist & key); Wheeler, W.M. 1921a: 16 (subgenera, notes); Santschi, 1921f: 310 (subgenera, notes); Mann, 1921: 489, 499 (Fiji Is C. (Myrmogonia) & C. (Colobopsis) species keys); Wheeler, W.M. 1922a: 704 (subgenera synopsis); Wheeler, W.M. 1922a: 232, 948, 1039 (diagnosis, Afrotropical, Malagasy catalogues); Arnold, 1922: 610, 612, 619, 662, 673 (diagnosis, South Africa subgenera synopsis, South Africa C. (Tanaemyrmex), C. (Myrmophyma), C. (Myrmopsamma) species keys); Arnold, 1924: 683, 700, 709, 724, 735 (South Africa C. (Myrmosericus), C. (Myrmamblys), C. (Orthonotomyrmex), C. (Myrmotrema), C. (Myrmepomis) species keys); Emery, 1925a: 62 (Palaearctic C. lateralis group, key); Emery, 1925b: 59 (diagnosis, subgenera key, catalogue); Emery, 1925b: 69 (C. (Dinomyrmex) diagnosis, catalogue); Emery, 1925b: 70 (C. (Myrmoxygenys) & C. (Myrmopsamma) diagnoses, catalogues); Emery, 1925b: 71 (C. (Camponotus) diagnosis, catalogue); Emery, 1925b: 75 (C. (Tanaemyrmex) diagnosis, catalogue); Emery, 1925b: 104 (C. (Myrmosericus) diagnosis, catalogue); Emery, 1925b: 107 (C. (Myrmothrix) diagnosis, catalogue); Emery, 1925b: 109 (C. (Myrmophyma) diagnosis, catalogue); Emery, 1925b: 112 (C. (Myrmosaulus) diagnosis, catalogue); Emery, 1925b: 114 (C. (Myrmopytia) diagnosis, catalogue); Emery, 1925b: 115 (C. (Karavaievia) diagnosis, catalogue); Emery, 1925b: 116 (C. (Myrmentoma) diagnosis, catalogue); Emery, 1925b: 121 (C. (Mayria) diagnosis, catalogue); Emery, 1925b: 123 (C. (Myrmonesites) diagnosis, catalogue); Emery, 1925b: 124 (C. (Orthonotomyrmex) diagnosis, catalogue); Emery, 1925b: 126 (C. (Myrmepinotus) diagnosis, catalogue); Emery, 1925b: 127 (C. (Myrmopiromis) diagnosis, catalogue); Emery, 1925b: 130 (C. (Myrmotrema) diagnosis, catalogue); Emery, 1925b: 133 (C. (Myrmisolepis) diagnosis, catalogue); Emery, 1925b: 134 (C. (Myrmotarsus) diagnosis, catalogue); Emery, 1925b: 135 (C. (Myrmoplatys) diagnosis, catalogue); Emery, 1925b: 136 (C. (Myrmamblys) diagnosis, catalogue); Emery, 1925b: 142 (C. (Rhinomyrmex) diagnosis, catalogue); Emery, 1925b: 143 (C. (Myrmogonia) diagnosis, catalogue); Emery, 1925b: 144 (C. (Colobopsis) diagnosis, catalogue); Emery, 1925b: 151 (C. (Myrmosphincta) diagnosis, catalogue); Emery, 1925b: 152 (C. (Myrmaphaenus) diagnosis, catalogue); Emery, 1925b: 157 (C. (Pseudocolobopsis) diagnosis, catalogue); Emery, 1925b: 159 (C. (Manniella) diagnosis, catalogue); Emery, 1925b: 160 (C. (Hypercolobopsis) diagnosis, catalogue); Emery, 1925b: 161 (C. (Myrmostenus) & C. (Myrmobrachys) diagnoses, catalogues); Emery, 1925b: 166 (C. (Myrmocladoecus) diagnosis, catalogue); Emery, 1925b: 167 (C. (Myrmeurynota) diagnosis, catalogue); Emery, 1925b: 168 (C. (Myrmodirachis) diagnosis, catalogue); Emery, 1925b: 169 (C. (Myrmomalis) & C. (Myrmoplatypus) diagnoses, catalogues); Emery, 1925b: 170 (C. (Myrmepomis) diagnosis, catalogue); Emery, 1925b: 172 (Dendromyrmex diagnosis, catalogue); Santschi, 1926a: 16, 21 (C. (Myrmopelta) diagnosis, species key); Karavaiev, 1927c: 275 (Ukraine species key); Arnol'di, 1933b: 601 (Russia species key); Menozzi, 1933b: 81 (Israel species key); Menozzi, 1935c: 330 (Chile species key); Karavaiev, 1936: 175 (Ukraine species key); Finzi, 1936: 190 (Egypt species key); Menozzi, 1939a: 316 (Himalaya & Tibet species key); Stitz, 1939: 236 (Germany species key); Kratochvíl, 1941: 109 (Central Europe species key); Novák & Sadil, 1941: 109 (Central Europe species key); Cole, 1942: 387 (U.S.A., Utah species key); Donisthorpe, 1932g: 248 (C. (Orthonotomyrmex) notes); Holgersen, 1943b: 174 (Norway species key); Holgersen, 1944: 199 (Norway species key); Buren, 1944a: 293 (U.S.A., Iowa species key); Creighton, 1950a: 362 (North America species keys); Chapman & Capco, 1951: 220 (Asia checklist); Yasumatsu & Brown, 1951: 29 (C. herculeanus complex); Kusnezov, 1952d: 192 (Argentina species key); Yasumatsu & Brown, 1957: 45 (C. herculeanus complex); Pisarski, 1961a: 175 (Poland species key); Gregg, 1963: 655 (U.S.A., Colorado species key); Wheeler, G.C. & Wheeler, J. 1963: 163 (U.S.A., North Dakota species key); Arnol'di, 1967: 1817 (former U.S.S.R. C. (Camponotus) species key); Bernard, 1967: 330 (diagnosis, Western Europe species key); Kempf, 1972a: 42, 95 (Neotropical catalogue); Hashmi, 1973: 1 (C. (Myrmothrix) all species revision, key); Alayo, 1974: 27 (Cuba species key); Snelling & Hunt, 1976: 117 (Chile species key); Tarbinsky, 1976: 144 (Kyrghyzstan species key); Boven, 1977: 130 (Belgium species key); Kutter, 1977c: 191 (Switzerland species key); Arnol'di & Dlussky, 1978: 551 (former European U.S.S.R. species key); Collingwood, 1978: 90 (Iberian Peninsula species key); Collingwood, 1979: 87 (Fennoscandia & Denmark species key); Smith, D.R. 1979: 1424 (North America catalogue); Allred, 1982: 445 (U.S.A., Utah species key); Gösswald, 1985: 264 (Germany species key); Collingwood, 1985: 275 (Saudi Arabia species key); Taylor & Brown, D.R. 1985: 109 (Australia catalogue); Dumpert, 1986: 571 (C. (Karavaievia) species key); Wheeler, G.C. & Wheeler, J. 1986g: 60 (U.S.A., Nevada species key); Agosti & Collingwood, 1987b: 282 (Balkans species key); Taylor, 1987a: 10, 16 (Australia, New Caledonia, New Zealand checklists); Snelling, R.R. 1988: 57 (Nearctic C. (Myrmentoma) species key); Wang, C., Xiao & Wu, 1989a: 222 (China species key); Dlussky, Soyunov & Zabelin, 1990: 125 (Turkmenistan species key); Kupyanskaya, 1990: 164 (Far Eastern Russia species key); Brandão, 1991: 332 (Neotropical catalogue); Morisita, Kubota, Onoyama, et al. 1991: 38 (Japan species key); Terayama, 1991: 165 (Japan C. (Paramyrmamblys) species); Atanasov & Dlussky, 1992: 207 (Bulgaria species key); Wang, C. & Wu, 1994: 28 (China species key); Radchenko, 1994b: 116 (South Siberia species key); Arakelian, 1994: 82 (Armenia species key); Bolton, 1994: 50 (synoptic classification); Bolton, 1995a: 1048 (census); Bolton, 1995b: 83, 169 (catalogue); Douwes, 1995: 92 (Sweden species key); Dumpert, Maschwitz, Weissflog et al. 1995: 104 (C. (Karavaievia) species key); Kupyanskaya, 1995: 352 (Far Eastern Russia species key); Wu, J. & Wang, 1995: 168 (China species key); Collingwood & Agosti, 1996: 370 (Saudi Arabia species key); Seifert, 1996b: 170 (Central Europe species key); Radchenko, 1996b: 1195 (Asiatic Palaearctic species key); Cagniant, 1996b: 87 (Morocco species key); McArthur & Adams, 1996: 18 (Australia C. nigriceps group, key); Mackay, 1997: 194 (C. (Myrmostenus) species key); Mackay & Mackay, 1997b: 319 and Snelling, R.R. 2000: 607 (C. montivagus complex, keys); Robertson & Zachariades, 1997: 2 (C. fulvopilosus group, key); Snelling & Torres, 1998: 9 (Puerto Rico & Virgin Is species key); Collingwood & Prince, 1998: 24 (Portugal species key); Terayama, 1999b: 27 (Japan species key); Shattuck, 1999: 91 (Australia synopsis); Zhou, 2001b: 200 (China, Guangxi species key); McArthur & Shattuck, 2001: 27 (Australia C. macrocephalus group, key); Czechowski, Radchenko & Czechowska, 2002: 153 (Poland species key); Fernández, 2002: 58 (C. (Dendromyrmex) diagnosis, all species revision, key); Mackay & Mackay, 2002: 248 (U.S.A., New Mexico species key); Shattuck & McArthur, 2002: 65 (Australia C. wiederkehri group key); Bolton, 2003: 112 (synopsis); Shattuck, 2005: 3 (C. aureopilus group key); Radchenko, 2005b: 188 (North Korea species key); Hansen & Klotz, 2005: 75 (North American “Carpenter Ants” key); Snelling, R.R. 2006: 96 (U.S.A. C. festinatus complex, key); Karmaly & Narendran, 2006: 29 (India species, key); Dumpert, Maschwitz & Weissflog, 2006: 77 (C. (Karavaievia) species, keys); Seifert, 2007: 152 (North and Central Europe species key); McArthur, 2007a: 290 (Australia species key); Shattuck & Janda, 2009: 252 (C. aureopilus group key); Mackay & Delsinne, 2009: 496 (New World C. maculatus complex, key); Terayama, 2009: 216 (Taiwan species key); Heterick, 2009: 60 (south-western Australia species key); Ionescu-Hirsch, 2009: 59 (Israel species key); McArthur, 2010: 18 (south Australia species key).

Formicinae Camponotini Polyrhachis TRUE TRUE FALSE

Extant: 614 species, 86 subspecies

Fossil: 1 species

Polyrhachis Smith, F. 1857a: 58. Type-species: Formica bihamata, by original designation.

Taxonomic history

[Polyrhachis Shuckard, in Swainson & Shuckard, 1840: 172, nomen nudum.]

Polyrhachis in Formicidae: Smith, F. 1858b: 58.

Polyrhachis in Formicinae: Mayr, 1862: 652 [Formicidae]; Mayr, 1865: 6 [Formicidae].

Polyrhachis in Camponotinae: Forel, 1878: 367 [Camponotidae]; Dalla Torre, 1893: 257.

Polyrhachis in Camponotinae, Polyrhachidini: Ashmead, 1905b: 384.

Polyrhachis in Camponotinae, Camponotini: Forel, 1886f: 194; Forel, 1893a: 165; Emery, 1895j: 772; Wheeler, W.M. 1910g: 144; Forel, 1912i: 89; Forel, 1917: 251; Arnold, 1924: 741.

Polyrhachis in Formicinae, Camponotini: Wheeler, W.M. 1922a: 701; Emery, 1925b: 175; all subsequent authors except the following.

Polyrhachis in Formicinae, Formica genus group: Agosti, 1991: 295.

Polyrhachis as subgenus of Myrma: Wheeler, W.M. 1911c: 860; Wheeler, W.M. 1911f: 170. [Myrma as subgenus of Polyrhachis: Forel, 1915b: 106; Forel, 1917: 251; Wheeler, W.M. 1922a: 702, 993; Emery, 1925b: 198; all subsequent authors (see note under Myrma).]

[Polyrachis Arnold, 1924: 741, incorrect subsequent spelling.]

Subgenera of POLYRHACHIS include the nominal plus the following.

All the earlier subgenera were given as provisional junior synonyms of Polyrhachis by Brown, 1973b: 178 - 184. Brown's list was repeated in Hölldobler & Wilson, 1990: 19 but with all subgenera listed as junior synonyms; subgeneric status was reinstated in Bolton, 1994: 50. These consecutive entries are not repeated individually below.

Subgenus POLYRHACHIS (MYRMA)

Myrma Billberg, 1820: 104. Type-species: Formica militaris, by subsequent designation of Wheeler, W.M. 1911c: 859.

Taxonomic history

[Although Myrma antedates Polyrhachis, the former has been treated as a subgenus of the latter since Forel, 1917: 251 (see Hung, 1967b: 396 for history). Dorow, Kohout & Taylor, 1997: 236 proposed the precedence of Polyrhachis over Myrma and that precedence was established by the ICZN (Opinion 1919) 1999: 92.]

Polyrhachis as subgenus of Myrma: Wheeler, W.M. 1911c: 860; Wheeler, W.M. 1911f: 170.

Myrma as subgenus of Polyrhachis: Forel, 1915b: 106; Forel, 1917: 251; Wheeler, W.M. 1922a: 702, 993; Emery, 1925b: 198; all subsequent authors, see note above.

Junior synonyms of POLYRHACHIS (MYRMA)

Hoplomyrmus Gerstäcker, 1859: 262. Type-species: Hoplomyrmus schistaceus, by monotypy.

Taxonomic history

[Hoplomyrmus also described as new by Gerstäcker, 1862: 508.]

Hoplomyrmus as junior synonym of Polyrhachis: Roger, 1861b: 174; Roger, 1863b: 6; Mayr, 1863: 446; Dalla Torre, 1893: 257.

Hoplomyrmus as junior synonym of Myrma: Wheeler, W.M. 1911c: 860; Wheeler, W.M. 1922a: 993; Emery, 1925b: 198; Bolton, 1973b: 288.

Pseudocyrtomyrma Emery, 1921e: 18 [as subgenus of Polyrhachis]. Type-species: Polyrhachis revoili, by original designation.

Taxonomic history

Pseudocyrtomyrma as subgenus of Polyrhachis: Emery, 1921e: 18; Emery, 1925b: 206.

Pseudocyrtomyrma as junior synonym of Myrma: Bolton, 1973b: 288.

Anoplomyrma Chapman, 1963: 258 [as subgenus of Polyrhachis]. Type-species: Polyrhachis (Anoplomyrma) parabiotica, by monotypy.

Taxonomic history

Anoplomyrma as junior synonym of Myrma: Dorow, 1995: 30.

Subgenus POLYRHACHIS (HEMIOPTICA)

Hemioptica Roger, 1862a: 238. Type-species: Hemioptica scissa, by monotypy.

Taxonomic history

Hemioptica in Formicinae: Mayr, 1862: 769 [Formicidae]; Mayr, 1865: 6 [Formicidae].

Hemioptica in Camponotinae: Dalla Torre, 1893: 271.

Hemioptica in Camponotinae, Camponotini: Emery, 1895j: 772; Wheeler, W.M. 1910g: 144; all subsequent authors except the following.

Hemioptica in Camponotinae, Polyrhachidini: Ashmead, 1905b: 384.

Hemioptica as junior synonym of Polyrhachis: Forel, 1878: 368.

Hemioptica as subgenus of Myrma: Wheeler, W.M. 1911c: 860; Wheeler, W.M. 1911f: 164.

Hemioptica as genus: Roger, 1862a: 238; Mayr, 1862: 769; Mayr, 1865: 6; Dalla Torre, 1893: 271; Emery, 1895j: 772; Ashmead, 1905b: 384; Wheeler, W.M. 1910g: 144; Emery, 1925b: 209; Donisthorpe, 1943f: 649; Chapman & Capco, 1951: 255.

Hemioptica as subgenus of Polyrhachis: Mayr, 1872: 138; Forel, 1915b: 107; Forel, 1917: 251; Wheeler, W.M. 1922a: 701; Dorow & Kohout, 1995: 95; Bolton, 1995b: 31; Dorow, 1995: 29.

Subgenus POLYRHACHIS (HAGIOMYRMA)

Hagiomyrma Wheeler, W.M. 1911c: 860 [as subgenus of Polyrhachis]. Type-species: Formica ammon, by original designation.

Taxonomic history

Hagiomyrma as subgenus of Polyrhachis: Wheeler, W.M. 1911c: 860; Forel, 1917: 251; Wheeler, W.M. 1922a: 702; Emery, 1925b: 184; all subsequent authors.

Subgenus POLYRHACHIS (HIRTOMYRMA)

Hirtomyrma Kohout, 2010: 199 [as subgenus of Polyrhachis]. Type-species: Polyrhachis hirta, by original designation.

Taxonomic history

Hirtomyrma as subgenus of Polyrhachis: Kohout, 2010: 199.

Subgenus POLYRHACHIS (CAMPOMYRMA)

Campomyrma Wheeler, W.M. 1911g: 860 [as subgenus of Polyrhachis]. Type-species: Polyrhachis clypeata (junior synonym of Formica exercita), by original designation.

Taxonomic history

Campomyrma as subgenus of Polyrhachis: Forel, 1917: 251; Wheeler, W.M. 1922a: 702; Emery, 1925b: 178; all subsequent authors.

Subgenus POLYRHACHIS (MYRMOTHRINAX)

Myrmothrinax Forel, 1915b: 107 [as subgenus of Polyrhachis]. Type-species: Polyrhachis thrinax, by original designation.

Taxonomic history

Myrmothrinax as subgenus of Polyrhachis: Forel, 1915b: 107; Forel, 1917: 251; Wheeler, W.M. 1922a: 704; Emery, 1925b: 178; all subsequent authors.

Junior synonym of POLYRHACHIS (MYRMOTHRINAX)

Evelyna Donisthorpe, 1937c: 273 [as subgenus of Polyrhachis]. Type-species: Polyrhachis (Evelyna) cheesmanae, by original designation.

Taxonomic history

Evelyna as junior synonym of Myrmothrinax: Hung, 1967b: 402.

Subgenus POLYRHACHIS (CYRTOMYRMA)

Cyrtomyrma Forel, 1915b: 107 [as subgenus of Polyrhachis]. Type-species: Formica rastellata, by original designation.

Taxonomic history

Cyrtomyrma as subgenus of Polyrhachis: Forel, 1915b: 107; Forel, 1917: 251; Wheeler, W.M. 1922a: 701; Emery, 1925b: 207; all subsequent authors.

Subgenus POLYRHACHIS (MYRMHOPLA)

Myrmhopla Forel, 1915b: 107 [as subgenus of Polyrhachis]. Type-species: Formica armata, by original designation.

Taxonomic history

Myrmhopla as subgenus of Polyrhachis: Forel, 1915b: 107; Forel, 1917: 251; Wheeler, W.M. 1922a: 701; Emery, 1925b: 190; all subsequent authors.

Junior synonyms of POLYRHACHIS (MYRMHOPLA)

Cephalomyrma Karavaiev, 1935a: 115 [as subgenus of Polyrhachis]. Type-species: Polyrhachis (Cephalomyrma) stylifera, by monotypy.

Taxonomic history

Cephalomyrma as junior synonym of Myrmhopla: Hung, 1967b: 402.

Florencea Donisthorpe, 1937a: 624 [as subgenus of Polyrhachis]. Type-species: Polyrhachis (Florencea) kirkae (junior synonym of Polyrhachis nigriceps), by original designation.

Taxonomic history

Florencea as junior synonym of Myrmhopla: Hung, 1967b: 402.

Subgenus POLYRHACHIS (CHARIOMYRMA)

Chariomyrma Forel, 1915b: 107 [as subgenus of Polyrhachis]. Type-species: Polyrhachis guerini, by original designation.

Taxonomic history

Chariomyrma as subgenus of Polyrhachis: Forel, 1915b: 107; Forel, 1917: 251; Wheeler, W.M. 1922a: 702; Emery, 1925b: 185; all subsequent authors.

Subgenus POLYRHACHIS (HEDOMYRMA)

Hedomyrma Forel, 1915b: 107 [as subgenus of Polyrhachis]. Type-species: Polyrhachis ornata, by original designation.

Taxonomic history

Hedomyrma as subgenus of Polyrhachis: Forel, 1915b: 107; Forel, 1917: 251; Wheeler, W.M. 1922a: 702; Emery, 1925b: 189; all subsequent authors.

Junior synonyms of POLYRHACHIS (HEDOMYRMA)

Dolichorhachis Mann, 1919: 386 [as subgenus of Polyrhachis]. Type-species: Polyrhachis (Dolichorhachis) malaensis, by monotypy.

Taxonomic history

Dolichorhachis as subgenus of Polyrhachis: Mann, 1919: 386; Wheeler, W.M. 1922a: 702; Emery, 1925b: 188.

Dolichorhachis as junior synonym of Hedomyrma: Dorow, 1995: 26.

Morleyidris Donisthorpe, 1944c: 64 [as subgenus of Polyrhachis]. Type-species: Polyrhachis (Morleyidris) trina, by original designation.

Taxonomic history

Morleyidris as junior synonym of Hedomyrma: Hung, 1967b: 402.

Subgenus POLYRHACHIS (MYRMATOPA)

Myrmatopa Forel, 1915b: 107 [as subgenus of Polyrhachis]. Type-species: Polyrhachis schang, by original designation.

Taxonomic history

Myrmatopa as subgenus of Polyrhachis: Forel, 1917: 251; Wheeler, W.M. 1922a: 702; Emery, 1925b: 180; all subsequent authors.

Junior synonym of POLYRHACHIS (MYRMATOPA)

Irenea Donisthorpe, 1938e: 502 [as subgenus of Dolichoderus]. Type-species: Dolichoderus (Irenea) omyrmex, by original designation.

Taxonomic history

Irenea in Dolichoderinae, Dolichoderini: Donisthorpe, 1938e: 502; Donisthorpe, 1943f: 653; Chapman & Capco, 1951: 186.

Irenea as subgenus of Dolichoderus: Donisthorpe, 1938e: 502.

Irenea as genus: Donisthorpe, 1941h: 59; Chapman & Capco, 1951: 186.

Irenea as junior synonym of Polyrhachis (Myrmatopa): Bolton, 1994: 50; Dorow, 1995: 42.

Subgenus POLYRHACHIS (AULACOMYRMA)

Aulacomyrma Emery, 1921e: 17 [as subgenus of Polyrhachis]. Type-species: Polyrhachis porcata, by original designation.

Taxonomic history

Aulacomyrma as subgenus of Polyrhachis: Emery, 1921e: 17; Emery, 1925b: 197; all subsequent authors.

Junior synonym of POLYRHACHIS (AULACOMYRMA)

Johnia Karavaiev, 1927e: 43 [as subgenus of Polyrhachis]. Type-species: Polyrhachis (Johnia) schizospina, by monotypy.

Taxonomic history

Johnia as junior synonym of Aulacomyrma: Hung, 1967b: 402.

Genus Polyrhachis references

Smith, F. 1858b: 58 (diagnosis); Roger, 1863b: 6 (catalogue); Mayr, 1863: 422, 443 (Hemioptica, Polyrhachis catalogues); Mayr, 1865: 6 (Polyrhachis, Hemioptica diagnoses); Mayr, 1867a: 40, 41 (diagnosis, species groups (as "turmae")); Mayr, 1876: 68 (Australia species key); Forel, 1878: 368 (diagnosis); Mayr, 1879: 648 (species groups); André, 1887: 286 (Afrotropical species key); Dalla Torre, 1893: 257, 271 (Polyrhachis, Hemioptica catalogues); Forel, 1893c: 17 (India & Sri Lanka species key); Emery, 1896d: 378 (catalogue); Emery, 1897d: 579, 582 (P. relucens & P. guerini groups, keys); Emery, 1898a: 227 (additions to 1896a catalogue); Bingham, 1903: 383 (India, Sri Lanka & Burma species key); Santschi, 1910h: 284 (P. rastellata group, key); Viehmeyer, 1914b: 50, 54 (P. rastellata & P. hostilis groups, keys); Wheeler, W.M. 1922a: 256, 701, 992 (diagnosis, subgenera key, Afrotropical catalogue); Arnold, 1924: 741 (diagnosis, South Africa species key); Emery, 1925b: 175 (diagnosis, subgenera key, catalogue); Emery, 1925b: 178 (P. (Campomyrma) diagnosis, catalogue); Emery, 1925b: 180 (P. (Myrmatopa) diagnosis, catalogue); Emery, 1925b: 181 (P. (Polyrhachis) diagnosis, catalogue); Emery, 1925b: 182 (P. (Myrmothrinax) diagnosis, catalogue); Emery, 1925b: 184 (P. (Hagiomyrma) diagnosis, catalogue); Emery, 1925b: 185 (P. (Chariomyrma) diagnosis, catalogue); Emery, 1925b: 188 (P. (Dolichorhachis) diagnosis, catalogue); Emery, 1925b: 189 (P. (Hedomyrma) diagnosis, catalogue); Emery, 1925b: 190 (P. (Myrmhopla) diagnosis, catalogue); Emery, 1925b: 197 (P. (Aulacomyrma) diagnosis, catalogue); Emery, 1925b: 198 (P. (Myrma) diagnosis, catalogue); Emery, 1925b: 206 (P. (Pseudocyrtomyrma) diagnosis, catalogue); Emery, 1925b: 207 (P. (Cyrtomyrma) diagnosis, catalogue); Emery, 1925b: 209 (Hemioptica diagnosis, catalogue); Donisthorpe, 1938c: 246 (P. (Cyrtomyrma) review); Santschi, 1939c: 13 (P. (Pseudocyrtomyrma) species key); Chapman & Capco, 1951: 186, 255, 256 (Asia Irenea, Hemioptica, Polyrhachis checklists); Hung, 1962: 24 (Taiwan species key); Hung, 1967b: 395 (review of subgenera); Hung, 1970: 5 (P. (Polyrhachis) species revision, key); Bolton, 1973b: 289 (Afrotropical species revision, key); Bolton, 1975c: 1 (P. sexspinosa group, key); Taylor & Brown, D.R. 1985: 131 (Australia catalogue); Taylor, 1987a: 59 (Australia, New Caledonia checklist); Kohout, 1987: 169 (Philippines P. sexspinosa group, key); Kohout, 1989: 515 (Australia P. relucens group, key); Kohout, 1990: 500 (P. viehmeyeri group, key); Kohout & Taylor, 1990: 509 (Australia checklist); Morisita, Kubota, Onoyama, et al. 1991: 45 (Japan species key); Wang C. & Wu, 1991: 596 (China species key); Dorow & Kohout, 1995: 96 (P. (Hemioptica) species key); Bolton, 1994: 50 (synoptic classification); Bolton, 1995a: 1052 (census); Bolton, 1995b: 342 (catalogue); Dorow, 1995: 9 (subgenera revision, subgenera key, catalogue); Dorow, 1995: 10-11 (P. (Myrmhopla) species groups, key; P. arachne-, P. cryptoceroides- P. flavoflagellata-, P. furcata- and P. hector species groups, keys); Dorow, 1995: 12 (P. (Aulacomyrma) synopsis); Dorow, 1995: 13 (P. (Campomoyrma) synopsis); Dorow, 1995: 16 (P. (Chariomyrma) synopsis); Dorow, 1995: 21 (P. (Cyrtomyrma) synopsis); Dorow, 1995: 24 (P. (Hagiomyrma) synopsis); Dorow, 1995: 26 (P. (Hedomyrma) synopsis); Dorow, 1995: 28 (P. (Hemioptica) synopsis); Dorow, 1995: 30 (P. (Myrma) synopsis); Dorow, 1995: 42 (P. (Myrmatopa) synopsis); Dorow, 1995: 45 (P. (Myrmhopla) synopsis); Dorow, 1995: 61 (P. (Myrmothrinax) synopsis); Dorow, 1995: 63 (P. (Polyrhachis) synopsis); Wu, J. & Wang, 1995: 159 (China species key); Shattuck, 1999: 107 (Australia synopsis); Zhou, 2001b: 190 (China, Guangxi species key); Xu, 2002b: 523 (China P. (Cyrtomyrma) species key; Bolton, 2003: 122 (synopsis); Karmaly, 2004: 542 (India species key); Kohout, 2006a: 146 (P. cryptoceroides group, key); Kohout, 2006b: 87 (P. Cyrtomyrma) of Australia, Borneo, New Guinea, Solomon Is, keys); Kohout, 2007a: 189 (P. Aulacomyrma) diagnosis, all species revision, key); Kohout, 2007c: 8 (P. (Campomyrma) Borneo, key); Kohout, 2008a: 255 (Sulawesi species, keys); Kohout, 2008b: 298 (P. vestita group key); Kohout, 2008d: 12 (flavoflagellata group key); Kohout, 2009: 40, 44 (modifications to Kohout 2006b key); Terayama, 2009: 218 (Taiwan species key); Heterick, 2009: 106 (south-western Australia species key); Kohout, 2010: 169, 171, 200 (Australia subgenera key, Australia P. (Myrmhopla) species key, Australia P. (Hirtomyrma) species key).

Pseudomyrmecinae Pseudomyrmecini Tetraponera TRUE TRUE FALSE

Extant: 89 species, 19 subspecies

Fossil: 7 species

Tetraponera Smith, F. 1852: 44. Type-species: Tetraponera atrata (junior synonym of Eciton nigrum), by subsequent designation of Wheeler, W.M. 1911f: 173.

Taxonomic history

Tetraponera in Formicidae, Poneridae: Smith, F. 1857a: 70.

Tetraponera in Myrmicidae: Smith, F. 1877b: 68.

Tetraponera in Myrmicinae, Pseudomyrmini: Forel, 1917: 240; Emery, 1921f: 24.

Tetraponera in Pseudomyrminae: Wheeler, W.M. 1922a: 654; subsequent authors to the following.

Tetraponera in Pseudomyrmecinae: Taylor & Brown, D.R. 1985: 17; all subsequent authors.

Tetraponera as junior synonym of Pseudomyrma: Roger, 1863b: 24; Mayr, 1863: 451.

Tetraponera as junior synonym of Sima: Dalla Torre, 1893: 53; Bingham, 1903: 107; Emery, 1917a: 95.

Tetraponera as subgenus of Sima: Emery, 1900d: 673; Emery, 1915b: 266; Arnold, 1916: 180; Forel, 1917: 240; Emery, 1921f: 24.

Tetraponera as genus: Smith, F. 1852: 44; Smith, F. 1857a: 70; Smith, F. 1877b: 68; Donisthorpe, 1916b: 244; Wheeler, W.M. 1922a: 654; Taylor & Brown, D.R. 1985: 17; Ward, 1990: 470; Bolton, 1994: 184.

Junior synonyms of TETRAPONERA

Sima Roger, 1863a: 178. Type-species: Sima compressa (junior synonym of Pseudomyrma allaborans), by monotypy.

Taxonomic history

[Type-species not Eciton rufonigrum, unjustified subsequent designation by Emery, 1915b: 266, repeated in Emery, 1917a: 95 and Emery, 1921f: 22.]

Sima in Myrmicidae: Smith, F. 1871a: 328.

Sima in Myrmicidae, Myrmeciidae: Emery, 1877a: 81.

Sima in Myrmicinae: Mayr, 1865: 25 [Myrmicidae]; Mayr, 1866b: 906 [Myrmicidae]; Dalla Torre, 1893: 53.

Sima in Myrmicidae, Pseudomyrminae: Ashmead, 1905b: 383.

Sima in Myrmicinae, Pseudomyrmini: Forel, 1893a: 164; Emery, 1895j: 768; Emery, 1914a: 40; Forel, 1917: 240; Wheeler, W.M. 1915h: 40; Arnold, 1916: 173; Emery, 1921f: 21.

Sima as genus: Roger, 1863a: 178; Mayr, 1863: 453; Mayr, 1868c: 100; Dalla Torre, 1893: 53; Emery, 1900d: 672; Bingham, 1903: 107; Emery, 1914a: 40; Emery, 1915b: 266; Arnold, 1916: 173; Emery, 1917a: 95; Forel, 1917: 240; Emery, 1921f: 21.

Sima as junior synonym of Tetraponera: Smith, F. 1877b: 68; Donisthorpe, 1916b: 244; Wheeler, W.M. 1920: 47; Wheeler, W.M. 1922a: 105; Donisthorpe, 1943g: 726; Ward, 1990: 470.

Pachysima Emery, 1912b: 97 [as subgenus of Sima]. Type-species: Tetraponera aethiops, by monotypy.

Taxonomic history

Pachysima in Myrmicinae, Pseudomyrmini: Forel, 1917: 240; Emery, 1921f: 22.

Pachysima in Pseudomyrminae: Wheeler, W.M. 1922a: 112, 654.

Pachysima as genus: Wheeler, W.M. 1918c: 308 (footnote); Wheeler, W.M. 1922a: 112, 654.

Pachysima as subgenus of Sima: Emery, 1912b: 97; Arnold, 1916: 174; Emery, 1921f: 22.

Pachysima as junior synonym of Tetraponera: Brown, 1973b: 183 [provisional]; Ward, 1990: 470.

Viticicola Wheeler, W.M. 1919i: 130 (in text). Type-species: Sima tessmanni, by original designation.

Taxonomic history

[Viticicola also as new in Wheeler, W.M. 1920: 53.]

Viticicola in Pseudomyrminae: Wheeler, W.M. 1922a: 107, 654.

Viticicola as junior synonym of Tetraponera: Brown, 1973b: 185 [provisional]; Ward, 1990: 470.

[Viticola Donisthorpe, 1943g: 735, incorrect subsequent spelling.]

Parasima Donisthorpe, 1948d: 592 [as subgenus of Tetraponera]. [Unnecessary replacement name for Sima in the sense of Emery, 1921f: 23.]

Taxonomic history

Parasima as junior synonym of Tetraponera: Ward, 1990: 470.

Genus Tetraponera references

Mayr, 1863: 453 (catalogue); Mayr, 1865: 25 (Sima diagnosis); Mayr, 1867a: 82 (Sima diagnosis); Mayr, 1868c: 101 (†Baltic Amber species key); Smith, F. 1877b: 68 (checklist); Dalla Torre, 1893: 53 (catalogue); Emery, 1900d: 17 (Asia & Australia species key); Forel, 1903a: 708 (India & Sri Lanka species key); Bingham, 1903: 107 (India, Sri Lanka & Burma species key); Arnold, 1916: 173 (Sima diagnosis, South Africa species key); Emery, 1921f: 21 (Sima diagnosis, subgenera key, catalogue); Emery, 1921f: 22 (S. (Pachysima) diagnosis, catalogue); Emery, 1921f: 24 (S. (Tetraponera) diagnosis, catalogue); Wheeler, W.M. 1922a: 112 (Pachysima species key); Wheeler, W.M. 1922a: 104, 107, 112 (Tetraponera, Viticicola, Pachysima diagnoses); Wheeler, W.M. 1922a: 795, 801 (Afrotropical Tetraponera, Viticicola, Pachysima catalogues); Wheeler, W.M. 1922a: 1014 (Malagasy catalogue); Taylor & Brown, D.R. 1985: 17 (Australia catalogue); Taylor, 1987a: 79 (Australia checklist); Wu, J. & Wang, 1990: 516 (China species key); Dlussky & Radchenko, 1990: 120 (Vietnam species key); Ward, 1990: 470 (diagnosis, review of genus, phylogeny); Bolton, 1994: 184 (synoptic classification); Bolton, 1995a: 1053 (census); Bolton, 1995b: 416 (catalogue); Wu, J. & Wang, 1995: 55 (China species key); Collingwood & Agosti, 1996: 312 (Saudi Arabia species key); Shattuck, 1999: 208 (Australia synopsis); Zhou, 2001b: 64 (China, Guangxi species key); Ward, 2001: 596 (Oriental & Australian, all species, revision, key); Xu & Chai, 2004: 64 (China species key); Ward, 2006: 120 (Afrotropical species groups key); Ward, 2006: 123 (T. ambigua group, key); Ward, 2009: 288 (T. grandidieri group, key); Dlussky, 2009: 1084 (Eocene amber species key); Terayama, 2009: 128 (Taiwan species key).

Cerapachyinae Cerapachyini Cerapachys TRUE TRUE FALSE

146 species

Cerapachys Smith, F. 1857a: 74. Type-species: Cerapachys antennatus, by subsequent designation of Bingham, 1903: 28.

Taxonomic history

Cerapachys in Myrmicinae: Smith, F. 1857a: 74 [Myrmicidae]; Smith, F. 1861b: 47 [Myrmicidae]; Smith, F. 1871a: 329 [Myrmicidae].

Cerapachys in Ponerinae: Dalla Torre, 1893: 17.

Cerapachys in Dorylinae, Cerapachyini: Emery, 1895j: 765; Emery, 1901a: 34.

Cerapachys in Ponerinae, Cerapachyini: Forel, 1893a: 162 (also misspelled Orapachys, same page); Forel, 1900d: 331; Ashmead, 1905b: 382; Wheeler, W.M. 1910g: 137; Emery, 1911d: 8; Arnold, 1915: 11; Forel, 1917: 239; Brown, 1975: 18; Snelling, R.R. 1981: 389; Dlussky & Fedoseeva, 1988: 79; Hölldobler & Wilson, 1990: 10.

Cerapachys in Cerapachyinae, Cerapachyini: Wheeler, W.M. 1902d: 185; Wheeler, W.M. 1922a: 639; Donisthorpe, 1943f: 632; Chapman & Capco, 1951: 17; Kempf, 1972a: 76; Wheeler, G.C. & Wheeler, J. 1985: 256; Bolton, 1990a: 67; Bolton, 1990c: 1357; Jaffe, 1993: 9; Bolton, 1994: 19; Bolton, 2003: 34.

Junior synonyms of CERAPACHYS

Syscia Roger, 1861a: 19. Type-species: Syscia typhla, by monotypy.

Taxonomic history

Syscia in Ponerinae: Mayr, 1862: 714 (in key) [Poneridae]; Mayr, 1865: 15 [Poneridae]; Dalla Torre, 1893: 17.

Syscia in Dorylinae, Cerapachyini: Emery, 1895j: 765.

Syscia in Ponerinae, Cerapachyini: Forel, 1893a: 162; Forel, 1900d: 329; Ashmead, 1905b: 382.

Syscia as genus: Dalla Torre, 1893: 17; Forel, 1900d: 329; Bingham, 1903: 31.

Syscia as subgenus of Cerapachys: Wheeler, W.M. 1902d: 185; Emery, 1902c: 24; Wheeler, W.M. 1910g: 137; Emery, 1911d: 10; Forel, 1917: 239; Wheeler, W.M. 1922a: 639; Donisthorpe, 1943g: 730.

Syscia as junior synonym of Cerapachys: Kempf, 1972a: 76; Brown, 1975: 18; all subsequent authors.

Ooceraea Roger, 1862a: 248. Type-species: Ooceraea fragosa, by monotypy.

Taxonomic history

Ooceraea in Myrmicinae: Mayr, 1865: 24 [Myrmicidae].

Ooceraea in Myrmicidae, Pheidolidae: Emery, 1877a: 81.

Ooceraea in Poneridae: Smith, F. 1871a: 324.

Ooceraea in Ponerinae: Dalla Torre, 1893: 17.

Ooceraea in Ponerinae, Cerapachyini: Forel, 1893a: 162.

Ooceraea in Dorylinae, Cerapachyini: Emery, 1895j: 765; Emery, 1901a: 34.

Ooceraea in Ecitoninae, Ecitonini: Ashmead, 1905b: 381; Ashmead, 1906: 25.

Ooceraea as genus: Roger, 1862a: 248; Smith, F. 1871a: 324; Dalla Torre, 1893: 17; Forel, 1900d: 329; Bingham, 1903: 31.

Ooceraea as subgenus of Cerapachys: Emery, 1902c: 24; Wheeler, W.M. 1902d: 185; Wheeler, W.M. 1910g: 137; Emery, 1911d: 10; Forel, 1917: 239; Wheeler, W.M. 1922a: 639; Donisthorpe, 1943f: 678.

Ooceraea as junior synonym of Cerapachys: Brown, 1973b: 183 [provisional]; Brown, 1975: 19; all subsequent authors.

Lioponera Mayr, 1879: 666. Type-species: Lioponera longitarsus, by monotypy.

Taxonomic history

Lioponera in Ponerinae: Dalla Torre, 1893: 17.

Lioponera in Pachycondylinae, Lioponerini: Ashmead, 1905b: 382.

Lioponera in Ponerinae, Cerapachyini: Forel, 1893a: 162; Forel, 1900d: 329; Wheeler, W.M. 1910g: 137; Emery, 1911d: 11; Forel, 1917: 239.

Lioponera in Dorylinae, Cerapachyini: Emery, 1895j: 765.

Lioponera in Cerapachyinae, Cerapachyini: Wheeler, W.M. 1902d: 185; Wheeler, W.M. 1922a: 639; Donisthorpe, 1943f: 657.

Lioponera as subgenus of Cerapachys: Forel, 1892l: 244.

Lioponera as genus: Mayr, 1879: 666; Dalla Torre, 1893: 17; Forel, 1900d: 329; Bingham, 1903: 26; Emery, 1911d: 11; Wheeler, W.M. 1922a: 639; Donisthorpe, 1943f: 657.

Lioponera as junior synonym of Cerapachys: Brown, 1975: 19; all subsequent authors.

Parasyscia Emery, in André, 1882c: 235. Type-species: Parasyscia piochardi, by monotypy.

Taxonomic history

Parasyscia in Ponerinae: Dalla Torre, 1893: 17.

Parasyscia in Dorylinae, Cerapachyini: Emery, 1895j: 765.

Parasyscia in Ponerinae, Cerapachyini: Forel, 1893a: 162 [Cerapachysii]; Ashmead, 1905b: 382.

Parasyscia as genus: Dalla Torre, 1893: 17.

Parasyscia as subgenus of Cerapachys: Forel, 1892l: 243; Wheeler, W.M. 1902d: 185; Wheeler, W.M. 1910g: 137; Emery, 1911d: 9; Forel, 1917: 239; Wheeler, W.M. 1922a: 639; Donisthorpe, 1943f: 681; Creighton, 1950a: 57.

Parasyscia as junior synonym of Cerapachys: Kempf, 1972a: 76; Brown, 1975: 18; all subsequent authors.

Phyracaces Emery, 1902c: 23. Type-species: Cerapachys mayri, by original designation.

Taxonomic history

Phyracaces in Ponerinae, Cerapachyini: Ashmead, 1905b: 382; Wheeler, W.M. 1910g: 137; Emery, 1911d: 10; Arnold, 1915: 17; Forel, 1917: 239.

Phyracaces in Cerapachyinae, Cerapachyini: Wheeler, W.M. 1902d: 185; Wheeler, W.M. 1922a: 639; Donisthorpe, 1943f: 683.

Phyracaces as junior synonym of Cerapachys: Brown, 1973b: 183 [provisional]; Brown, 1975: 19; all subsequent authors.

Cysias Emery, 1902c: 24 [as subgenus of Cerapachys]. Type-species: Ooceraea papuana, by original designation.

Taxonomic history

Cysias in Ponerinae, Cerapachyini: Ashmead, 1905b: 382.

Cysias as genus: Ashmead, 1905b: 382.

Cysias as subgenus of Cerapachys: Emery, 1902c: 24; Wheeler, W.M. 1910g: 137; Donisthorpe, 1943f: 637.

Cysias as junior synonym of Syscia: Emery, 1911d: 10; Brown, 1975: 19; all subsequent authors.

Ceratopachys Schulz, W.A. 1906: 155, unjustified emendation of Cerapachys.

Taxonomic history

Ceratopachys as junior synonym of Cerapachys: Wheeler, W.M. 1922a: 755.

Chrysapace Crawley, 1924: 380. Type-species: Chrysapace jacobsoni (junior secondary homonym in Cerapachys, replaced by Cerapachys crawleyi), by monotypy.

Taxonomic history

Chrysapace in Ponerinae: Crawley, 1924: 380.

Chrysapace in Cerapachyinae, Cerapachyini: Donisthorpe, 1943f: 633.

Chrysapace as junior synonym of Cerapachys: Brown, 1975: 19; all subsequent authors.

Neophyracaces Clark, 1941: 76. Type-species: Phyracaces clarus, by original designation.

Taxonomic history

Neophyracaces in Cerapachyinae: Clark, 1941: 76.

Neophyracaces as junior synonym of Cerapachys: Brown, 1973b: 183 [provisional]; Brown, 1975: 19; all subsequent authors.

Yunodorylus Xu, 2000b: 297. Type-species: Yunodorylus sexspinus, by original designation.

Taxonomic history

Yunodorylus in Dorylinae: Xu, 2000b: 297.

Yunodorylus in Cerapachyinae, Cerapachyini: Bolton, 2003: 141, 268.

Yunodorylus as junior synonym of Cerapachys: Bolton, 2003: 141, 268; Borowiec, 2009: 57.

Genus Cerapachys references

Roger, 1863b: 21, 25 (catalogue); Mayr, 1863: 403, 439, 455 (catalogue); Mayr, 1865: 15, 24 (Syscia, Ooceraea diagnoses); André, 1882c: 236 (Europe & Algeria species); Dalla Torre, 1893: 17 (Lioponera, Cerapachys, Parasyscia, Ooceraea, Syscia catalogues); Forel, 1900d: 331 (India & Sri Lanka species key); Bingham, 1903: 27, 28 (India, Sri Lanka & Burma Lioponera, Cerapachys species keys); Emery, 1911d: 8 (diagnosis, subgenera key, catalogue); Emery, 1911d: 9 (C. (Parasyscia) diagnosis, catalogue); Emery, 1911d: 10 (C. (Ooceraea) & C. (Syscia) diagnoses, catalogues); Emery, 1911d: 10 (Phyracaces diagnosis, catalogue); Emery, 1911d: 11 (Lioponera diagnosis, catalogue); Arnold, 1915: 11 (diagnosis, South Africa species key); Wheeler, W.M. 1918a: 239 (Australia Phyracaces species key); Mann, 1921: 408 (Fiji species key); Wheeler, W.M. 1922a: 52, 53, 639 (Cerapachys & Phyracaces diagnoses, subgenera key); Wheeler, W.M. 1924d: 224 (Chrysapace, comments); Arnold, 1926: 193 (South Africa species, revised key); Donisthorpe, 1939a: 253 (Lioponera diagnosis); Chapman & Capco, 1951: 17, 21, 22 (Asia Cerapachys, Lioponera, Phyracaces checklists); Wilson, 1959b: 44, 52, 55, 56 (Melanesia, Fiji Is, New Guinea & New Caledonia species keys); Kempf, 1972a: 76 (Neotropical catalogue); Brown, 1975: 18, 26 (diagnosis, keys); Smith, D.R. 1979: 1333 (North America catalogue); Taylor & Brown, D.R. 1985: 23 (Australia catalogue); Taylor, 1987a: 16 (Australia, New Caledonia checklist); Terayama, Kubota, Sakai & Kawazoe, 1988: 35 (Taiwan species key); Morisita, Kubota, Onoyama, et al. 1989: 30 (Japan species key); Bolton, 1995a: 1048 (census); Bolton, 1995b: 142 (catalogue); Wu, J. & Wang, 1995: 47 (China species key); Collingwood & Agosti, 1996: 312 (Saudi Arabia species key); Terayama, 1996: 18 (Japan species key); Shattuck, 1999: 60 (Australia synopsis); Mackay & Mackay, 2002: 36 (U.S.A., New Mexico species key); Borowiec, 2009: 46 (C. sexspinus group key); Terayama, 2009: 122 (Taiwan species key); Heterick, 2009: 126 (south-western Australia species key).

Myrmicinae Dacetini Pyramica TRUE TRUE FALSE

Extant: 344 species

Fossil: 4 species

Pyramica Roger, 1862a: 251. Type-species: Pyramica gundlachi, by monotypy.

Taxonomic history

Pyramica in Myrmicinae, Dacetini: Emery, 1924d: 320; all subsequent authors.

Pyramica as junior synonym of Strumigenys: Roger, 1863b: 40; Emery & Forel, 1879a: 464; Dalla Torre, 1893: 145; Bingham, 1903: 147; Emery, 1924d: 320; Donisthorpe, 1943g: 723; Brown & Wilson, 1959b: 281; Brown, 1960b: 37; Kempf, 1972a: 242; Bolton, 1983: 358; Bolton, 1994: 105; Baroni Urbani & De Andrade, 2007: 101.

Pyramica as subgenus of Strumigenys: Brown, 1948e: 110.

Pyramica as genus: Mayr, 1863: 453; Bolton, 1999: 1667; Bolton, 2000: 90; Eguchi, Bui & Yamane, 2011: 26.

Junior synonyms of PYRAMICA

Epitritus Emery, 1869a: 136. Type-species: Epitritus argiolus, by monotypy.

Taxonomic history

Epitritus in Myrmicidae, Cryptoceridae: Emery, 1877a: 81.

Epitritus in Myrmicinae: Emery & Forel, 1879a: 465 [Myrmicidae]; Dalla Torre, 1893: 148.

Epitritus in Cryptoceridae, Dacetonini: Ashmead, 1905b: 384.

Epitritus in Myrmicinae, Dacetini: Forel, 1892d: 344; Forel, 1893a: 164; Emery, 1895j: 770; Wheeler, W.M. 1910g: 141; Emery, 1914a: 42; Forel, 1917: 246; Arnold, 1917: 384; Wheeler, W.M. 1922a: 666; Emery, 1924d: 326; Donisthorpe, 1943f: 643; Brown, 1948e: 122; Brown, 1949b: 44; all subsequent authors.

Epitritus as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 12; Baroni Urbani & De Andrade, 2007: 101.

Epitritus as junior synonym of Pyramica: Bolton, 1999: 1667; Bolton, 2000: 90.

Trichoscapa Emery, 1869b: 24 [as subgenus of Strumigenys]. Type-species: Strumigenys (Trichoscapa) membranifera, by monotypy.

Taxonomic history

Trichoscapa in Myrmicinae: Emery & Forel, 1879a: 464 [Myrmicidae]; Dalla Torre, 1893: 145.

Trichoscapa in Myrmicinae, Dacetini: Forel, 1917: 246; Wheeler, W.M. 1922a: 668; Emery, 1924d: 323; Donisthorpe, 1943g: 734; Brown, 1948e: 112; all subsequent authors.

Trichoscapa as subgenus of Strumigenys: Emery & Forel, 1879a: 464; Forel, 1917: 246; Creighton, 1950a: 304.

Trichoscapa as junior synonym of Strumigenys: Dalla Torre, 1893: 145; Baroni Urbani & De Andrade, 1994: 13; Baroni Urbani & De Andrade, 2007: 101.

Trichoscapa as junior synonym of Cephaloxys: Wheeler, W.M. 1922a: 668; Emery, 1924d: 323; Donisthorpe, 1943g: 734.

Trichoscapa as genus: Brown, 1948e: 112; Kempf, 1972a: 255; Bolton, 1983: 319.

Trichoscapa as junior synonym of Pyramica: Bolton, 1999: 1667; Bolton, 2000: 90.

Pentastruma Forel, 1912a: 50. Type-species: Pentastruma sauteri, by monotypy.

Taxonomic history

Pentastruma in Myrmicinae, Dacetini: Emery, 1914a: 42; Forel, 1917: 246; Wheeler, W.M. 1922a: 666; Emery, 1924d: 326; Donisthorpe, 1943f: 682; Brown, 1948e: 102.

Pentastruma as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 15; Baroni Urbani & De Andrade, 2007: 101.

Pentastruma as junior synonym of Pyramica: Bolton, 1999: 1667; Bolton, 2000: 91.

Glamyromyrmex Wheeler, W.M. 1915e: 487. Type-species: Glamyromyrmex beebei, by monotypy.

Taxonomic history

Glamyromyrmex in Myrmicinae, Dacetini: Wheeler, W.M. 1915e: 491; Forel, 1917: 246; Wheeler, W.M. 1922a: 667; Emery, 1924d: 326; Donisthorpe, 1943f: 647; Brown, 1948e: 116; all subsequent authors.

Glamyromyrmex as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 14; Baroni Urbani & De Andrade, 2007: 101.

Glamyromyrmex as junior synonym of Pyramica: Bolton, 1999: 1667; Bolton, 2000: 91.

Codiomyrmex Wheeler, W.M. 1916d: 326. Type-species: Codiomyrmex thaxteri, by monotypy.

Taxonomic history

Codiomyrmex in Myrmicinae, Dacetini: Wheeler, W.M. 1922a: 667; Emery, 1924d: 325; Donisthorpe, 1943f: 633; Brown, 1948e: 114; all subsequent authors.

Codiomyrmex as subgenus of Strumigenys: Emery, 1924d: 325; Donisthorpe, 1943f: 633.

Codiomyrmex as genus: Wheeler, W.M. 1916d: 326; Brown, 1948e: 114.

Codiomyrmex as junior synonym of Glamyromyrmex: Brown, 1973b: 179 [provisional].

Codiomyrmex as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 14; Baroni Urbani & De Andrade, 2007: 101.

Codiomyrmex as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Tingimyrmex Mann, 1926: 104 [as subgenus of Strumigenys]. Type-species: Strumigenys (Tingimyrmex) mirabilis, by monotypy.

Taxonomic history

Tingimyrmex in Myrmicinae, Dacetini: Donisthorpe, 1943g: 732; Brown, 1948e: 111; all subsequent authors.

Tingimyrmex as subgenus of Strumigenys: Mann, 1926: 104; Donisthorpe, 1943g: 732.

Tingimyrmex as genus: Brown, 1948e: 111; Kempf, 1972a: 250.

Tingimyrmex as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 15; Baroni Urbani & De Andrade, 2007: 101.

Tingimyrmex as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Codioxenus Santschi, 1931c: 278 [as subgenus of Epitritus]. Type-species: Epitritus (Codioxenus) simulans, by monotypy.

Taxonomic history

Codioxenus in Myrmicinae, Dacetini: Donisthorpe, 1943f: 633; Brown, 1948e: 123; all subsequent authors.

Codioxenus as genus: Brown, 1948e: 123.

Codioxenus as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 14; Baroni Urbani & De Andrade, 2007: 101.

Codioxenus as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Smithistruma Brown, 1948e: 104.

Taxonomic history

[Replacement name for Cephaloxys Smith, F. 1865: 76; junior homonym of Cephaloxys Signoret, 1847: 294 (Hemiptera). Type-species not Strumigenys pulchella, unjustified subsequent designation by Brown, 1948e: 104, repeated in Kempf, 1972a: 230, Bolton, 1983: 274 and Bolton, 1995b: 46.]

Smithistruma in Myrmicinae, Dacetini: Brown, 1948e: 104.

Smithistruma as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 14; Baroni Urbani & De Andrade, 2007: 102.

Smithistruma as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Homonym replaced by Smithistruma

Cephaloxys Smith, F. 1865: 76. Type-species: Cephaloxys capitata, by monotypy.

Taxonomic history

[Cephaloxys Smith, F. junior homonym of Cephaloxys Signoret, 1847: 294 (Hemiptera).]

Cephaloxys in Cryptoceridae: Smith, F. 1871a: 335.

Cephaloxys in Myrmicinae, Dacetini: Wheeler, W.M. 1922a: 668.

Cephaloxys as junior synonym of Strumigenys: Dalla Torre, 1893: 15.

Cephaloxys as subgenus of Strumigenys: Wheeler, W.M. 1922a: 668; Emery, 1924d: 323; Bernard, 1953b: 254 (anachronism).

Weberistruma Brown, 1948e: 106 [as subgenus of Smithistruma]. Type-species: Strumigenys (Cephaloxys) leptothrix, by original designation.

Taxonomic history

Weberistruma in Myrmicinae, Dacetini: Brown, 1948e: 106.

Weberistruma as genus: Brown, 1949d: 7; Brown, 1953g: 24.

Weberistruma as junior synonym of Smithistruma: Brown, 1973a: 35.

Weberistruma as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Weberistruma as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 2007: 102.

Wessonistruma Brown, 1948e: 106 [as subgenus of Smithistruma]. Type-species: Strumigenys pergandei, by original designation.

Taxonomic history

Wessonistruma in Myrmicinae, Dacetini: Brown, 1948e: 106.

Wessonistruma junior synonym of Smithistruma: Brown, 1973a: 35.

Wessonistruma as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Wessonistruma as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 2007: 102.

Serrastruma Brown, 1948e: 107 [as subgenus of Smithistruma]. Type-species: Strumigenys simoni, by original designation.

Taxonomic history

Serrastruma in Myrmicinae, Dacetini: Brown, 1948e: 107, and all subsequent authors.

Serrastruma as genus: Brown, 1949d: 6; Brown, 1952e: 70; Bolton, 1983: 335; Bolton, 1994: 105.

Serrastruma as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 14; Baroni Urbani & De Andrade, 2007: 102.

Serrastruma as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Neostruma Brown, 1948e: 111. Type-species: Strumigenys crassicornis, by original designation.

Taxonomic history

Neostruma in Myrmicinae, Dacetini: Brown, 1948e: 111; all subsequent authors.

Neostruma as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 12; Baroni Urbani & De Andrade, 2007: 102.

Neostruma as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Dorisidris Brown, 1948e: 116. Type-species: Strumigenys (Codiomyrmex) nitens, by original designation.

Taxonomic history

Dorisidris in Myrmicinae, Dacetini: Brown, 1948e: 116; all subsequent authors.

Dorisidris as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 14; Baroni Urbani & De Andrade, 2007: 102.

Dorisidris as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Miccostruma Brown, 1948e: 123. Type-species: Epitritus mandibularis, by original designation.

Taxonomic history

Miccostruma in Myrmicinae, Dacetini: Brown, 1948e: 123.

Miccostruma as junior synonym of Smithistruma: Bolton, 1983: 274.

Miccostruma as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Miccostruma as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 2007: 102.

Kyidris Brown, 1949d: 3. Type-species: Kyidris mutica, by original designation.

Taxonomic history

Kyidris in Myrmicinae, Dacetini: Brown, 1949d: 3.

Kyidris as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 15; Baroni Urbani & De Andrade, 2007: 103.

Kyidris as junior synonymof Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Chelystruma Brown, 1950a: 33 [as subgenus of Glamyromyrmex]. Type-species: Glamyromyrmex (Chelystruma) lilloana, by monotypy.

Taxonomic history

Chelystruma in Myrmicinae, Dacetini: Brown, 1950a: 33; all subsequent authors.

Chelystruma as subgenus of Glamyromyrmex: Brown, 1953g: 16.

Chelystruma as genus: Kempf, 1959d: 338; Kempf, 1972a: 77.

Chelystruma as junior synonym of Glamyromyrmex: Brown, 1973b: 179 [provisional].

Chelystruma as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 14; Baroni Urbani & De Andrade, 2007: 103.

Chelystruma as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Polyhomoa Azuma, 1950: 36. Type-species: Polyhomoa itoi (junior synonym of Kyidris mutica), by monotypy.

Taxonomic history

Polyhomoa as junior synonym of Kyidris: Creighton, 1950b: 93; Brown & Yasumatsu, 1951: 93.

Polyhomoa as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Polyhomoa as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 2007: 103.

Borgmeierita Brown, 1953g: 23. Type-species: Codiomyrmex excisus, by original designation.

Taxonomic history

Borgmeierita in Myrmicinae, Dacetini: Brown, 1953g: 23.

Borgmeierita as junior synonym of Glamyromyrmex: Brown, 1973a: 35.

Borgmeierita as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Borgmeierita as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 2007: 103.

Platystruma Brown, 1953g: 112 [as subgenus of Smithistruma]. Type-species: Strumigenys (Cephaloxys) depressiceps, by original designation.

Taxonomic history

Platystruma in Myrmicinae, Dacetini: Brown, 1953g: 112; all subsequent authors.

Platystruma as junior synonym of Smithistruma: Brown, 1973a: 35.

Platystruma as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Platystruma as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 2007: 103.

Gymnomyrmex Borgmeier, 1954b: 279. Type-species: Gymnomyrmex splendens, by original designation.

Taxonomic history

Gymnomyrmex in Myrmicinae, Dacetini: Borgmeier, 1954b: 279; all subsequent authors.

Gymnomyrmex as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 14; Baroni Urbani & De Andrade, 2007: 103.

Gymnomyrmex as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Dysedrognathus Taylor, 1968d: 132. Type-species: Dysedrognathus extemenus, by original designation.

Taxonomic history

Dysedrognathus in Myrmicinae, Dacetini: Taylor, 1968d: 132.

Dysedrognathus as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 13; Baroni Urbani & De Andrade, 2007: 103.

Dysedrognathus as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Asketogenys Brown, 1972: 23. Type-species: Asketogenys acubecca, by original designation.

Taxonomic history

Asketogenys in Myrmicinae, Dacetini: Brown, 1972: 23.

Asketogenys as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 15; Baroni Urbani & De Andrade, 2007: 103.

Asketogenys as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Cladarogenys Brown, 1976b: 33. Type-species: Cladarogenys lasia, by original designation.

Taxonomic history

Cladarogenys in Myrmicinae, Dacetini: Brown, 1976b: 33.

Cladarogenys as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 14; Baroni Urbani & De Andrade, 2007: 103.

Cladarogenys as junior synonym of Pyramica: Bolton, 1999: 1668; Bolton, 2000: 91.

Genus Pyramica references

Mayr, 1863: 453 (catalogue); André, 1883b: 398, 402 (Europe & Algeria Strumigenys (Trichoscapa), Epitritus species); Dalla Torre, 1893: 145, 148 (Strumigenys, Epitritus catalogues); Emery, 1895c: 325 (North America species key); Wheeler, W.M. 1908a: 146 (New World species key); Santschi, 1913b: 257 (Afrotropical species key); Emery, 1916b: 205 (Italy species key); Arnold, 1917: 373 (South Africa species key); Wheeler, W.M. 1922a: 918, 920 (Afrotropical Strumigenys (Cephaloxys), Epitritus catalogues); Emery, 1924d: 323, 325, 326 (Strumigenys (Cephaloxys), Strumigenys (Codiomyrmex), Glamyromyrmex, Pentastruma, Epitritus diagnoses, catalogues); Smith, M.R. 1931c: 691 (North America Strumigenys (Cephaloxys) species key); Buren, 1944a: 290 (U.S.A., Iowa species key); Brown, 1948e: 112 - 124 (minor genera reviews); Brown, 1949b: 43 (Epitritus diagnosis, review); Brown, 1949d: 8 (Weberistruma species, review); Brown, 1949d: 21 (Japan, China & Taiwan species key); Creighton, 1950a: 301 (North America Strumigenys (Trichoscapa) species key); Chapman & Capco, 1951: 104, 105, 106 (Asia Epitritus, Kyidris, Pentastruma, Smithistruma, Strumigenys (Cephaloxys) checklists); Brown, 1950a: 27 (Glamyromyrmex review); Brown, 1952e: 86 (Serrastruma diagnosis, species revision, key); Brown, 1953g: 20, 24 (Codiomyrmex, Weberistruma reviews); Brown, 1953g: 31, 46, 92, 114, 125, 130 (Smithistruma diagnosis, Nearctic, Neotropical, Malesian, Afrotropical, Palaearctic species revisions, keys); Wilson & Brown, 1956b: 439 (Kyidris species review); Brown, 1959b: 1 (Neostruma species review); Kempf, 1959d: 339 (Gymnomyrmex species key); Brown, 1960b: 37 (Strumigenys gundlachi group revision, key); Kempf, 1960f: 447 (Neotropical Glamyromyrmex species key); Kempf, 1960f: 451 (Gymnomyrmex species key); Brown, 1962b: 79 (Epitritus species key); Brown, 1964a: 183 (supplement to 1953e Smithistruma revision); Bernard, 1967: 243, 245 (Smithistruma, Epitritus diagnoses); Kempf, 1972a: 37, 77, 98, 110, 116, 163, 230, 250, 255 (Neotropical Borgmeierita, Chelystruma, Codiomyrmex, Codioxenus, Dorisidris, Glamyromyrmex, Gymnomyrmex, Neostruma, Smithistruma, Tingimyrmex, Trichoscapa catalogues); Bolton, 1972: 208 (Epitritus species key); Alayo, 1974: 19 (Cuba species key); Smith, D.R. 1979: 1405, 1408 (North America Smithistruma, Trichoscapa catalogues); Taylor, 1987a: 28 (Australia checklist); Brown & Boisvert, 1979: 201 (Pentastruma species review); Bolton, 1983: 274, 319, 320, 335, 353, 354 (Afrotropical Smithistruma, Trichoscapa, Glamyromyrmex, Serrastruma, Cladarogenys, Epitritus species revisions, keys); Taylor & Brown, D.R. 1985: 64 (Australia Glamyromyrmex catalogue); Perrault, 1986a: 3 (Gymnomyrmex species key); Ward, 1988: 122 (Western Nearctic Smithistruma species key); Terayama & Kubota, 1989: 787 (Taiwan Smithistruma species key); Hölldobler & Wilson, 1990: 15 (synoptic classification); Brandão, 1991: 344, 346, 360, 378 (Neotropical catalogue); Arakelian & Dlussky, 1991: 150 (former U.S.S.R. species key); Morisita, Kubota, et al. 1992: 68 (Japan Smithistruma species key); Morisita, Kubota, et al. 1992: 76 (Japan Epitritus species key); Bolton, 1994: 105 (synoptic classification); Bolton, 1995a: 1048 - 1053 (census); Bolton, 1995b: 75, 145, 146, 177, 183, 188, 207, 211, 219, 292, 316, 382, 383, 420, 421 (catalogue); Terayama, Lin & Wu, 1996: 328 (Taiwan Smithistruma species key); Ogata & Onoyama, 1998: 277 (Japan Smithistruma species key); Shattuck, 1999: 136, 170 (Australia Glamyromyrmex, Trichoscapa synopses); Bolton, 1999: 1657, 1667 (morphology, diagnosis, review of genus, phylogeny); Xu, 2000e: 297 (China Epitritus species key); Bolton, 2000: 95, 137, 272, 285, 342, 372, 472 (Nearctic, Neotropical, Afrotropical, West Palaearctic, Malagasy, Malesian-Oriental-East Palaearctic, Austral species revisions, keys); Xu & Zhou, 2004: 441 (China species, key); Longino, 2006b: 139 (Costa Rica species, additions to key); Terayama, 2009: 138 (Taiwan species key).

Myrmicinae Dacetini Strumigenys TRUE TRUE FALSE

Extant: 487 species

Fossil:

Strumigenys Smith, F. 1860c: 72. Type-species: Strumigenys mandibularis, by monotypy.

Taxonomic history

Strumigenys in Myrmicidae: Smith, F. 1871a: 334; Cresson, 1887: 262.

Strumigenys in Myrmicidae, Cryptoceridae: Emery, 1877a: 81.

Strumigenys in Myrmicinae: Mayr, 1865: 26 [Myrmicidae]; Emery & Forel, 1879a: 464 [Myrmicidae]; Dalla Torre, 1893: 145.

Strumigenys in Cryptoceridae, Dacetini: Ashmead, 1905b: 384.

Strumigenys in Myrmicinae, Dacetini: Forel, 1892d: 344; Forel, 1893a: 164; Forel, 1895b: 133; Emery, 1895j: 770; Forel, 1899c: 42; Wheeler, W.M. 1910g: 141; Emery, 1914a: 42; Forel, 1917: 246; Arnold, 1917: 372; Wheeler, W.M. 1922a: 668; Emery, 1924d: 319; Brown, 1948e: 108; all subsequent authors.

Junior synonyms of STRUMIGENYS

Labidogenys Roger, 1862a: 249. Type-species: Labidogenys lyroessa, by monotypy.

Taxonomic history

Labidogenys in Myrmicinae, Dacetini: Donisthorpe, 1943f: 654; Brown, 1948e: 102.

Labidogenys as genus: Roger, 1862a: 249; Brown, 1948e: 102.

Labidogenys as junior synonym of Strumigenys: Roger, 1863b: 40; Emery & Forel, 1879a: 464; Dalla Torre, 1893: 145; Bingham, 1903: 147; Emery, 1924d: 320; Brown & Wilson, 1959b: 281; Brown, 1960b: 38; Bolton, 1983: 358; Bolton, 1999: 1673; Bolton, 2000: 492.

Proscopomyrmex Patrizi, 1946: 294. Type-species: Proscopomyrmex londianensis, by monotypy.

Taxonomic history

Proscopomyrmex in Myrmicinae, Dacetini: Patrizi, 1946: 294 [Dacetonini].

Proscopomyrmex as junior synonym of Strumigenys: Brown, 1949d: 15; Bolton, 1999: 1673; Bolton, 2000: 492.

Eneria Donisthorpe, 1948d: 598. Type-species: Eneria excisa (junior synonym of Strumigenys loriae), by original designation.

Taxonomic history

Eneria as genus: Chapman & Capco, 1951: 104 (anachronism).

Eneria as junior synonym of Strumigenys: Brown, 1949d: 15; Bolton, 1999: 1673; Bolton, 2000: 492.

Quadristruma Brown, 1949b: 47. Type-species: Epitritus emmae, by original designation.

Taxonomic history

Quadristruma in Myrmicinae, Dacetini: Brown, 1949b: 47; all subsequent authors.

Quadristruma as junior synonym of Strumigenys: Baroni Urbani & De Andrade, 1994: 12; Bolton, 1999: 1673; Bolton, 2000: 492.

Genus Strumigenys references

Roger, 1863b: 40 (catalogue); Mayr, 1863: 424, 455 (catalogue); Mayr, 1865: 26 (diagnosis); Forel, 1886a: 216 (all species key); Mayr, 1887: 568 (all species key); Dalla Torre, 1893: 145 (catalogue); Emery, 1895c: 325 (North America species key); Emery, 1897c: 574 (New Guinea species key); Forel, 1903a: 707 (India & Sri Lanka species key); Bingham, 1903: 147 (India, Sri Lanka & Burma species key); Wheeler, W.M. 1908a: 146 (New World species key); Santschi, 1913b: 257 (Afrotropical species key); Arnold, 1917: 373 (diagnosis, South Africa species key); Mann, 1921: 467 (Fiji Is species key); Wheeler, W.M. 1922a: 917, 1034 (Afrotropical, Malagasy catalogues); Emery, 1924d: 319 (diagnosis, catalogue); Brown, 1948e: 108 (diagnosis, review of genus); Brown, 1949d: 21 (Japan, China & Taiwan species key); Creighton, 1950a: 301 (North America species key); Chapman & Capco, 1951: 104, 107 (Asia Eneria, Strumigenys (Strumigenys) checklists); Brown, 1954k: 10 (Afrotropical species revision, key); Brown, 1958h: 38 (New Zealand species); Brown, 1962c: 238 (Neotropical species synopsis, key); Kempf, 1972a: 226, 243 (Neotropical Quadristruma, Strumigenys catalogues); Alayo, 1974: 19 (Cuba species key); Kempf, 1976b: 43 (additions to Brown, 1962c key); Smith, D.R. 1979: 1403, 1409 (North America Strumigenys, Quadristruma catalogues); Bolton, 1983: 358, 400 (Afrotropical Strumigenys, Quadristruma species revisions, keys); Taylor & Brown, D.R. 1985: 85, 87 (Australia Quadristruma, Strumigenys catalogues); Taylor, 1987a: 66, 76 (Australia, New Caledonia & New Zealand checklists); Terayama & Kubota, 1989: 780 (Taiwan species key); Brandão, 1991: 380 (Neotropical catalogue); Arakelian & Dlussky, 1991: 150 (former U.S.S.R. species key); Morisita, Kubota, et al. 1992: 63 (Japan species key); Dlussky, 1993: 53 (Fiji Is, Tonga and Samoa species key); Bolton, 1994: 105 (synoptic classification); Bolton, 1995a: 1053 (census); Bolton, 1995b: 377, 395 (catalogue); Gronenberg, 1996: 2021 (mandible morphology); Lin & Wu, 1996: 139 (Taiwan species key); Lattke & Goitía, 1997: 370 (Venezuela species key); Shattuck, 1999: 160, 167 (Australia synopsis); Bolton, 1999: 1657, 1673 (morphology, diagnosis, review of genus, phylogeny); Bolton, 2000: 496, 583, 716, 942 (Neotropical-Nearctic, Afrotropical, Malesian-Oriental-East Palaearctic, Austral species revisions, keys); Fisher, 2000: 614 (Malagasy species revision, key); Zhou, 2001b: 78 (China, Guangxi species key); Lin & Wu, 2001: 166 (Taiwan species key); Zhou & Xu, 2003: 737 (China species key); Yoshimura & Onoyama, 2007: 676 (Japan species key); Terayama, 2009: 142 (Taiwan species key); Heterick, 2009: 176 (south-western Australia species key).

Myrmicinae Solenopsidini Carebara TRUE TRUE FALSE

Extant: 160 species, 13 subspecies

Fossil: 6 species

Carebara Westwood, 1840b: 86. Type-species: Carebara lignata, by monotypy.

Taxonomic history

Carebara in Poneridae, Attidae: Smith, F. 1858b: 178.

Carebara in Myrmicinae: Mayr, 1865: 23 [Myrmicidae]; Smith, F. 1871a: 334 [Myrmicidae]; Dalla Torre, 1893: 74.

Carebara in Myrmicinae, Pheidolini: Emery, 1877a: 81 [Myrmicidae, Pheidolidae].

Carebara in Myrmicinae, Myrmicariini: Ashmead, 1905b: 383.

Carebara in Myrmicinae, Solenopsidini: Forel, 1893a: 164; Emery, 1895j: 770; Wheeler, W.M. 1910g: 140; Wheeler, W.M. 1922a: 663; Kusnezov, 1964: 61; Kempf, 1972a: 74; Jaffe, 1993: 10; Bolton, 2003: 61.

Carebara in Myrmicinae, Pheidologeton genus group: Ettershank, 1966: 81; Bolton, 1987: 265.

Carebara in Myrmicinae, Pheidologetini: Emery, 1914a: 41; Arnold, 1916: 248; Forel, 1917: 244; Emery, 1924d: 219; Dlussky & Fedoseeva, 1988: 80; Hölldobler & Wilson, 1990: 16; Bolton, 1994: 106 [Pheidologetonini].

Junior synonyms of CAREBARA

Oligomyrmex Mayr, 1867a: 110. Type-species: Oligomyrmex concinnus, by monotypy.

Taxonomic history

Oligomyrmex in Myrmicinae: Dalla Torre, 1893: 74.

Oligomyrmex in Myrmicinae, Stenammini: Ashmead, 1905b: 383.

Oligomyrmex in Myrmicinae, Solenopsidini: Forel, 1893a: 164; Emery, 1895j: 770; Wheeler, W.M. 1910g: 140; Wheeler, W.M. 1922a: 663; Kempf, 1972a: 172; Smith, D.R. 1979: 1389; Jaffe, 1993: 10.

Oligomyrmex in Myrmicinae, Pheidologeton genus group: Ettershank, 1966: 81; Bolton, 1987: 265.

Oligomyrmex in Myrmicinae, Pheidologetini: Emery, 1914a: 41; Forel, 1917: 243; Emery, 1924d: 215; Dlussky & Fedoseeva, 1988: 80; Hölldobler & Wilson, 1990: 16; Bolton, 1994: 106 [Pheidologetonini].

Oligomyrmex in Myrmicinae, Solenopsidini: Bolton, 2003: 61.

Oligomyrmex as junior synonym of Carebara: Fernández, 2004a: 194.

Aeromyrma Forel, 1891b: 198. Type-species: Aeromyrma nosindambo, by monotypy.

Taxonomic history

[Aeromyrma also described as new by Forel, 1891a: cccvii; no species-rank taxa named.]

Aeromyrma in Myrmicinae: Dalla Torre, 1893: 78.

Aeromyrma in Myrmicinae, Solenopsidini: Forel, 1893a: 164; Emery, 1895j: 770; Ashmead, 1905b: 383; Wheeler, W.M. 1910g: 140; Wheeler, W.M. 1922a: 663.

Aeromyrma in Myrmicinae, Pheidologetini: Emery, 1914a: 41; Arnold, 1916: 256; Forel, 1917: 243; Emery, 1924d: 215; all subsequent authors.

Aeromyrma as genus: Forel, 1891b: 198; Arnold, 1916: 256; Forel, 1917: 243; Wheeler, W.M. 1922a: 663, 882.

Aeromyrma as subgenus of Oligomyrmex: Emery, 1915c: 59 (footnote); Emery, 1924d: 215; Donisthorpe, 1943f: 620.

Aeromyrma as junior synonym of Oligomyrmex: Ettershank, 1966: 119; all subsequent authors to the following.

Aeromyrma as junior synonym of Carebara: Fernández, 2004a: 194.

Aneleus Emery, 1900c: 327 [as subgenus of Pheidologeton]. Type-species: Solenopsis similis, by subsequent designation of Wheeler, W.M. 1911f: 158.

Taxonomic history

[Type-species not Pheidologeton pygmaeus, unjustified subsequent designation by Wheeler, W.M. 1913a: 77; repeated in Emery, 1924d: 214.]

Aneleus in Myrmicinae, Solenopsidini: Wheeler, W.M. 1910g: 140; Wheeler, W.M. 1922a: 663.

Aneleus in Myrmicinae, Pheidologetini: Emery, 1914a: 41; Arnold, 1916: 254; Forel, 1917: 243; Emery, 1924d: 213; all subsequent authors.

Aneleus as genus: Emery, 1914a: 41; Arnold, 1916: 254; Forel, 1917: 243; Wheeler, W.M. 1922a: 663; Emery, 1924d: 213.

Aneleus as junior synonym of Oligomyrmex: Ettershank, 1966: 119; all subsequent authors to the following.

Aneleus as junior synonym of Carebara: Fernández, 2004a: 194.

Erebomyrma Wheeler, W.M. 1903a: 138. Type-species: Erebomyrma longii, by monotypy.

Taxonomic history

Erebomyrma in Myrmicinae, Solenopsidini: Wheeler, W.M. 1903a: 145; Wheeler, W.M. 1910g: 140; Wheeler, W.M. 1915h: 46; Wheeler, W.M. 1922a: 663.

Erebomyrma in Myrmicinae, Stenammini: Ashmead, 1905b: 383.

Erebomyrma in Myrmicinae, Pheidologetini: Emery, 1914a: 41; Forel, 1917: 244; Emery, 1924d: 218; all subsequent authors.

Erebomyrma as genus: Wheeler, W.M. 1922a: 663; Emery, 1924d: 218; Creighton, 1950a: 245; Kusnezov, 1964: 62; Wilson, 1986b: 61; Hölldobler & Wilson, 1990: 13.

Erebomyrma as junior synonym of Oligomyrmex: Ettershank, 1966: 119; Kempf, 1972a: 172; Smith, D.R. 1979: 1389; Bolton, 1994: 106; Bolton, 1995b: 298.

Erebomyrma as junior synonym of Carebara: Fernández, 2004a: 194.

Paedalgus Forel, 1911i: 217. Type-species: Paedalgus escherichi, by monotypy.

Taxonomic history

Paedalgus in Myrmicinae, Solenopsidini: Forel, 1911i: 218; Wheeler, W.M. 1922a: 663.

Paedalgus in Myrmicinae, Pheidologeton genus group: Ettershank, 1966: 81; Bolton, 1987: 265.

Paedalgus in Myrmicinae, Pheidologetini: Emery, 1914a: 41; Forel, 1917: 244; Emery, 1924d: 221; Dlussky & Fedoseeva, 1988: 80; Bolton, 1994: 106 [Pheidologetonini].

Paedalgus in Myrmicinae, Solenopsidini: Bolton, 2003: 61.

Paedalgus as junior synonym of Carebara: Fernández, 2004a: 194.

Lecanomyrma Forel, 1913k: 56 [as subgenus of Pheidologeton]. Type-species: Pheidologeton (Lecanomyrma) butteli, by monotypy.

Taxonomic history

Lecanomyrma as subgenus of Pheidologeton: Forel, 1913k: 56; Forel, 1917: 243.

Lecanomyrma as subgenus of Aneleus: Emery, 1924d: 215.

Lecanomyrma as junior synonym of Oligomyrmex: Ettershank, 1966: 119; all subsequent authors to the following.

Lecanomyrma as junior synonym of Carebara: Fernández, 2004a: 194.

Spelaeomyrmex Wheeler, W.M. 1922c: 9. Type-species: Spelaeomyrmex urichi, by original designation.

Taxonomic history

Spelaeomyrmex in Myrmicinae, Pheidologitonini: Wheeler, W.M. 1922c: 11; Donisthorpe, 1944b: 59.

Spelaeomyrmex in Myrmicinae, Solenopsidini: Donisthorpe, 1943g: 727; Kusnezov, 1964: 61.

Spelaeomyrmex as junior synonym of Erebomyrma: Wilson, 1962a: 63.

Spelaeomyrmex as junior synonym of Carebara: Fernández, 2004a: 194.

Hendecatella Wheeler, W.M. 1927h: 93 [as subgenus of Oligomyrmex]. Type-species: Oligomyrmex (Hendecatella) capreolus, by monotypy.

Taxonomic history

Hendecatella as junior synonym of Oligomyrmex: Ettershank, 1966: 119; all subsequent authors.

Hendecatella as junior synonym of Carebara: Fernández, 2004a: 194.

Solenops Karavaiev, 1930a: 207 [as subgenus of Solenopsis]. Type-species: Solenopsis (Solenops) weyeri, by monotypy.

Taxonomic history

Solenops as junior synonym of Oligomyrmex: Ettershank, 1966: 119; all subsequent authors.

Solenops as junior synonym of Carebara: Fernández, 2004a: 194.

Crateropsis Patrizi, 1948: 174 [as subgenus of Solenopsis]. Type-species: Solenopsis (Crateropsis) elmenteitae, by original designation.

Taxonomic history

Crateropsis as junior synonym of Oligomyrmex: Ettershank, 1966: 120; all subsequent authors.

Crateropsis as junior synonym of Carebara: Fernández, 2004a: 194.

Sporocleptes Arnold, 1948: 219. Type-species: Sporocleptes nicotianae, by original designation.

Taxonomic history

Sporocleptes as junior synonym of Aneleus: Consani, 1951: 169; Arnold, 1952a: 460.

Sporocleptes as junior synonym of Carebara: Fernández, 2004a: 194.

Nimbamyrma Bernard, 1953b: 240. Type-species: Nimbamyrma villiersi, by monotypy.

Taxonomic history

Nimbamyrma in Myrmicinae, Pheidologetini: Bernard, 1953b: 240.

Nimbamyrma as junior synonym of Oligomyrmex: Ettershank, 1966: 120; all subsequent authors.

Nimbamyrma as junior synonym of Carebara: Fernández, 2004a: 194; Fernández, 2010: 201.

Afroxyidris Belshaw & Bolton, 1994: 631. Type-species: Afroxyidris crigensis, by original designation.

Taxonomic history

Afroxyidris in Myrmicinae, Pheidologetonini: Belshaw & Bolton, 1994: 632; Bolton, 1994: 106.

Afroxyidris in Myrmicinae, Solenopsidini: Bolton, 2003: 61.

Afroxyidris as junior synonym of Carebara: Fernández, 2004a: 194.

Neoblepharidatta Sheela & Narendran, 1997: 88. Type-species: Neoblepharidatta nayana, by original designation.

Taxonomic history

Neoblepharidatta incertae sedis in Myrmicinae: Sheela & Narendran, 1997: 89.

Neoblepharidatta as junior synonym of Oligomyrmex: Bolton, 2003: 216, 273.

Neoblepharidatta as junior synonym of Carebara: Fernández, 2004a: 194.

Parvimyrma Eguchi & Bui, 2007: 40. Type-species: Parvimyrma sangi, by original designation.

Taxonomic history

Parvimyrma in Myrmicinae, Solenopsis genus group: Eguchi & Bui, 2007: 40.

Parvimyrma as junior synonym of Carebara: Fernández, 2010: 195.

Genus Carebara references

Smith, F. 1858b: 178 (Carebara diagnosis); Roger, 1863b: 32 (Carebara catalogue); Mayr, 1863: 402 (Carebara catalogue); Mayr, 1865: 23 (Carebara diagnosis); Mayr, 1867a: 106 (Carebara diagnosis); Dalla Torre, 1893: 74, 78 (Carebara, Oligomyrmex, Aeromyrma catalogues); Bingham, 1903: 150 (Carebara diagnosis); Bingham, 1903: 154 (Oligomyrmex India, Sri Lanka & Burma species key); Santschi, 1913e: 459 (African Oligomyrmex, key); Santschi, 1914d: 363 (Afrotropical Carebara species key); Arnold, 1916: 248, 254, 256 (Carebara, Aneleus, Aeromyrma diagnoses); Wheeler, W.M. 1922a: 165, 687 (Aeromyrma diagnosis, Oligomyrmex subgenera key); Wheeler, W.M. 1922a: 168, 172, 883 (Carebara diagnosis, Afrotropical species key, Afrotropical catalogue); Wheeler, W.M. 1922a: 177, 884 (Paedalgus diagnosis, Afrotropical catalogue); Wheeler, W.M. 1922a: 880, 881, 882 (Afrotropical Aneleus, Oligomyrmex, Aeromyrma catalogues); Wheeler, W.M. 1922a: 1028 (Malagasy Oligomyrmex, Aeromyrma catalogues); Emery, 1924d: 213 (Aneleus diagnosis, catalogue); Emery, 1924d: 215 (Aneleus (Lecanomyrma) diagnosis, catalogue); Emery, 1924d: 215 (Oligomyrmex (Aeromyrma) diagnosis, catalogue); Emery, 1924d: 217 (O. (Oligomyrex) diagnosis, catalogue); Emery, 1924d: 218 (Erebomyrma diagnosis, catalogue); Emery, 1924d: 219 (Carebara diagnosis, catalogue); Emery, 1924d: 221 (Paedalgus diagnosis, catalogue); Weber, 1950a: 1 (Africa Oligomyrmex species, synopsis); Chapman & Capco, 1951: 151, 154, 156, 158 (Asia Carebara, Aeromyrma, Aneleus, Oligomyrmex, Paedalgus checklists); Weber, 1952: 11 (Africa O. (Aeromyrma) species key); Ettershank, 1966: 119 (Oligomyrmex diagnosis, review of genus, checklist); Ettershank, 1966: 125 (Carebara diagnosis, review of genus, checklist); Ettershank, 1966: 128 (Paedalgus diagnosis, review of genus, checklist); Kempf, 1972a: 74 (Carebara Neotropical catalogue); Kempf, 1972a: 172 (Oligomyrmex Neotropical catalogue); Smith, D.R. 1979: 1389 (Oligomyrmex North America catalogue); Taylor & Brown, D.R. 1985: 71 (Oligomyrmex Australia catalogue); Taylor, 1987a: 50 (Oligomyrmex Australia, New Caledonia checklist); Taylor, 1991db: 605 (Oligomyrmex Australia species, partial key); Brandão, 1991: 342 (Oligomyrmex Neotropical catalogue); Morisita, Kubota, et al. 1992: 45 (Oligomyrmex Japan species key); Bolton & Belshaw, 1993: 181 (Paedalgus diagnosis, all species revision, key); Bolton, 1995a: 1048, 1051 (Carebara, Oligomyrmex, Paedalgus census); Bolton, 1995b: 133, 298, 311 (Carebara, Oligomyrmex, Paedalgus catalogues); Wu, J. & Wang, 1995: 74 (Oligomyrmex China species key); Terayama, 1996: 24 (Oligomyrmex Japan species key); Shattuck, 1999: 148 (Oligomyrmex Australia synopsis); Xu, 2003: 311 (Oligomyrmex China species key); Terayama, 2009: 151 (Taiwan species key).

Myrmicinae Solenopsidini Monomorium TRUE TRUE FALSE

Extant: 386 species, 30 subspecies

Fossil: 3 species

Monomorium Mayr, 1855: 452. Type-species: Monomorium monomorium (replacement name for Monomorium minutum, junior secondary homonym in Monomorium), by monotypy.

Taxonomic history

Monomorium in Myrmicidae, Myrmicidae: Emery, 1877a: 81.

Monomorium in Myrmicinae: Mayr, 1855: 452 [Myrmicidae]; Mayr, 1861: 71 [Myrmicidae]; Mayr, 1865: 22 [Myrmicidae]; Emery & Forel, 1879a: 456 [Myrmicidae]; Dalla Torre, 1893: 65.

Monomorium in Myrmicinae, Myrmicini: Forel, 1895b: 124; Emery, 1895j: 769; Forel, 1899c: 78; Wheeler, W.M. 1910g: 139.

Monomorium in Myrmicinae, Tetramoriini: Ashmead, 1905b: 383.

Monomorium in Myrmicinae, Monomoriini: Wheeler, W.M. 1915h: 45; Dlussky & Fedoseeva, 1988: 80.

Monomorium in Myrmicinae, Monomorium genus group: Ettershank, 1966: 81; in Solenopsis genus group: Bolton, 1987: 271.

Monomorium in Myrmicinae, Solenopsidini: Emery, 1914a: 41 [subtribe Monomoriini]; Arnold, 1916: 203; Forel, 1917: 242; Emery, 1922e: 166; Wheeler, W.M. 1922a: 662; Hölldobler & Wilson, 1990: 16; Jaffe, 1993: 10; Bolton, 1994: 106.

Junior synonyms of MONOMORIUM

Phacota Roger, 1862b: 260. Type-species: Phacota sichelii, by monotypy.

Taxonomic history

Phacota in Myrmicidae, Pheidolidae: Emery, 1877a: 81.

Phacota in Myrmicinae: Mayr, 1865: 22 [Myrmicidae]; Emery & Forel, 1879a: 464 [Myrmicidae]; Dalla Torre, 1893: 71.

Phacota in Myrmicinae, Myrmicini: Emery, 1895j: 769; Wheeler, W.M. 1910g: 139.

Phacota in Myrmicinae, Stenammini: Ashmead, 1905b: 383.

Phacota in Myrmicinae, Solenopsidini: Emery, 1914a: 41 [subtribe Monomoriini]; Forel, 1917: 243; Emery, 1922e: 187; Wheeler, W.M. 1922a: 662; all subsequent authors except the entry below; Hölldobler & Wilson, 1990: 16; Bolton, 1994: 106.

Phacota in Myrmicinae, Solenopsis genus group: Bolton, 1987: 271.

Phacota as genus: Bolton, 1987: 281.

Phacota as junior synonym of Monomorium: Ettershank, 1966: 82; Fernández, in Heterick, 2006: 79; Fernández, 2007b: 130.

Trichomyrmex Mayr, 1865: 19. Type-species: Trichomyrmex rogeri, by monotypy.

Taxonomic history

Trichomyrmex in Myrmicidae, Myrmicidae: Emery, 1877a: 81.

Trichomyrmex in Myrmicinae: Mayr, 1865: 19 [Myrmicidae]; Dalla Torre, 1893: 72.

Trichomyrmex in Myrmicinae, Myrmicini: Wheeler, W.M. 1910g: 140.

Trichomyrmex in Myrmicinae, Solenopsidini: Emery, 1914a: 41 [subtribe Monomoriini]; Forel, 1917: 243; Emery, 1922e: 185; Wheeler, W.M. 1922a: 662; Donisthorpe, 1943g: 733.

Trichomyrmex as junior synonym of Monomorium: Ettershank, 1966: 82; Smith, D.R. 1979: 1381; Bolton, 1987: 296.

†Lampromyrmex Mayr, 1868c: 93. Type-species: †Monomorium mayrianum (replacement name for †Lampromyrmex gracillimus, junior secondary homonym in Monomorium), by monotypy.

Taxonomic history

†Lampromyrmex in Myrmicidae, Pheidolidae: Emery, 1877a: 81.

†Lampromyrmex in Myrmicinae: Mayr, 1868c: 93 [Myrmicidae]; Dalla Torre, 1893: 78.

†Lampromyrmex in Myrmicinae, Solenopsidini: Emery, 1922e: 183 [subtribe Monomoriini]; Donisthorpe, 1943f: 655.

†Lampromyrmex as junior synonym of Monomorium: Wheeler, W.M. 1915h: 45.

†Lampromyrmex as subgenus of Monomorium: Emery, 1922e: 183; Wheeler, W.M. 1922a: 676.

†Lampromyrmex as junior synonym of Monomorium: Wheeler, W.M. 1915h: 45; Ettershank, 1966: 82; Smith, D.R. 1979: 1381; Bolton, 1987: 299.

Holcomyrmex Mayr, 1879: 671. Type-species: Holcomyrmex scabriceps, by subsequent designation of Bingham, 1903: 280.

Taxonomic history

[Holcomyrmex Smith, F. 1873: ix, nomen nudum, attributed to Mayr.]

Holcomyrmex in Myrmicinae, Myrmicini: Emery, 1895j: 769; Ashmead, 1905b: 383.

Holcomyrmex in Myrmicinae, Solenopsidini: Emery, 1922e: 181 [subtribe Monomoriini]; Donisthorpe, 1943f: 650.

Holcomyrmex as genus: Dalla Torre, 1893: 65; Forel, 1903a: 692; Bingham, 1903: 280.

Holcomyrmex as subgenus of Monomorium: Wheeler, W.M. 1910g: 139; Emery, 1915i: 190; Forel, 1917: 242; Emery, 1922e: 181; Wheeler, W.M. 1922a: 676.

Holcomyrmex as junior synonym of Monomorium: Emery, 1908h: 667; Ettershank, 1966: 82; Smith, D.R. 1979: 1381; Bolton, 1987: 295.

Epoecus Emery, 1893a: cclxxvi. Type-species: Epoecus pergandei, by monotypy.

Taxonomic history

Epoecus in Myrmicinae, Myrmicini: Emery, 1895j: 769; Wheeler, W.M. 1910g: 139.

Epoecus in Myrmicinae, Stenammini: Ashmead, 1905b: 383.

Epoecus in Myrmicinae, Solenopsidini: Emery, 1914a: 41 [subtribe Anergatini]; Forel, 1917: 243; Emery, 1922e: 204; Wheeler, W.M. 1922a: 663; Donisthorpe, 1943f: 643.

Epoecus as junior synonym of Monomorium: Ettershank, 1966: 82; Smith, D.R. 1979: 1381; Bolton, 1987: 287.

Wheeleriella Forel, 1907f: 145.

Taxonomic history

[Replacement name for Wheeleria Forel, 1905b: 171; junior homonym of Wheeleria Tutt, 1905: 37 (Lepidoptera).]

Wheeleriella in Myrmicinae, Myrmicini: Wheeler, W.M. 1910g: 139.

Wheeleriella in Myrmicinae, Solenopsidini: Emery, 1914a: 41 [subtribe Monomoriini]; Forel, 1917: 243; Emery, 1922e: 186; Wheeler, W.M. 1922a: 662; Donisthorpe, 1943g: 735.

Wheeleriella as junior synonym of Monomorium: Ettershank, 1966: 82; Smith, D.R. 1979: 1381; Bolton, 1987: 291.

Homonym replaced by Wheeleriella

Wheeleria Forel, 1905b: 171. Type-species: Wheeleria santschii, by monotypy.

Taxonomic history

[Junior homonym of Wheeleria Tutt, 1905: 37 (Lepidoptera).]

Epixenus Emery, 1908f: 556. Type-species: Epixenus andrei (junior secondary homonym in Monomorium, replaced by Monomorium advena), by subsequent designation of Wheeler, W.M. 1911f: 163.

Taxonomic history

Epixenus in Myrmicinae, Solenopsidini: Emery, 1914a: 41 [subtribe Monomoriini]; Forel, 1917: 243; Emery, 1922e: 184; Wheeler, W.M. 1922a: 662; Donisthorpe, 1943f: 643.

Epixenus as junior synonym of Monomorium: Brown & Wilson, 1957bb: 244; Ettershank, 1966: 82; Smith, D.R. 1979: 1381; Bolton, 1987: 287; Tinaut & Ortiz, 1988: 167.

Mitara Emery, 1913b: 261 [as subgenus of Monomorium]. Type-species: Monomorium laeve, by original designation.

Taxonomic history

Mitara in Myrmicinae, Solenopsidini: Arnold, 1916: 238; Forel, 1917: 242 [subtribe Monomoriini]; all subsequent authors.

Mitara as subgenus of Monomorium: Emery, 1913b: 261; Emery, 1915i: 190; Arnold, 1916: 238; Forel, 1917: 242.

Mitara as junior synonym of †Lampromyrmex: Emery, 1922e: 183; Wheeler, W.M. 1922a: 162, 676.

Mitara as genus: Donisthorpe, 1943f: 662.

Mitara as junior synoynm of Monomorium: Smith, D.R. 1979: 1381; Bolton, 1987: 299.

Chelaner Emery, 1914f: 410 [as subgenus of Monomorium]. Type-species: Monomorium (Chelaner) forcipatum, by subsequent designation of Emery, 1922e: 168.

Taxonomic history

Chelaner in Myrmicinae, Solenopsidini: Forel, 1917: 242 [subtribe Monomoriini]; Emery, 1922e: 168; all subsequent authors except the following.

Chelaner in Myrmicinae, Monomorium genus group: Ettershank, 1966: 81.

Chelaner in Myrmicinae, Monomoriini: Dlussky & Fedoseeva, 1988: 80.

Chelaner as subgenus of Monomorium: Emery, 1914f: 410; Forel, 1917: 242; Emery, 1922e: 168; Wheeler, W.M. 1922a: 676.

Chelaner as genus: Ettershank, 1966: 93; Taylor & Brown, D.R. 1985: 55; Wheeler, G.C. & Wheeler, J. 1985: 257; Dlussky & Fedoseeva, 1988: 80 (anachronism).

Chelaner (and its junior synonyms Notomyrmex, Protholcomyrmex, Schizopelta) as junior synonym of Monomorium: Bolton, 1987: 300; Heterick, 2001: 354.

Notomyrmex Emery, 1915i: 190 [as subgenus of Monomorium]. Type-species: Atta antarctica, by original designation.

Taxonomic history

Notomyrmex as subgenus of Monomorium: Emery, 1915i: 190; Forel, 1917: 242; Emery, 1922e: 168; Wheeler, W.M. 1922a: 676; Kusnezov, 1956: 20.

Notomyrmex as genus: Kusnezov, 1957a: 269 (in key); Kusnezov, 1960b: 343; Kusnezov, 1962b: 160; Kusnezov, 1964: 60.

Notomyrmex as junior synonym of Chelaner: Ettershank, 1966: 93.

Parholcomyrmex Emery, 1915i: 190 [as subgenus of Monomorium]. Type-species: Myrmica gracillima (junior synonym of Atta destructor), by original designation.

Taxonomic history

Parholcomyrmex as subgenus of Monomorium: Emery, 1915i: 190; Forel, 1917: 242; Emery, 1922e: 179; Wheeler, W.M. 1922a: 676; subsequent authors to the following.

Parholcomyrmex as junior synonym of Monomorium: Ettershank, 1966: 82; Smith, D.R. 1979: 1381; Bolton, 1987: 296.

[Paraholcomyrmex Emery, 1915i: 191, Forel, 1917: 242, incorrect subsequent spellings.]

Xeromyrmex Emery, 1915i: 190 [as subgenus of Monomorium]. Type-species: Formica salomonis, by original designation.

Taxonomic history

Xeromyrmex as subgenus of Monomorium: Forel, 1917: 242; Emery, 1922e: 175; Wheeler, W.M. 1922a: 677.

Xeromyrmex as junior synonym of Monomorium: Ettershank, 1966: 82; Smith, D.R. 1979: 1381; Bolton, 1987: 294.

Syllophopsis Santschi, 1915c: 259 [as subgenus of Monomorium.] Type-species: Monomorium modestum, by monotypy.

Taxonomic history

Syllophopsis in Myrmicinae, Solenopsidini: Forel, 1917: 242 [subtribe Monomoriini]; Emery, 1922e: 175; all subsequent authors to the following.

Syllophopsis in Myrmicinae, Monomorium genus group: Ettershank, 1966: 81.

Syllophopsis in Myrmicinae, Monomoriini: Dlussky & Fedoseeva, 1988: 80.

Syllophopsis as subgenus of Monomorium: Santschi, 1915c: 259; Forel, 1917: 242; Emery, 1922e: 175; Wheeler, W.M. 1922a: 676; Donisthorpe, 1943g: 729; Arnold, 1952a: 465.

Syllophopsis as genus: Santschi, 1921c: 119; Santschi, 1936a: 32; Ettershank, 1966: 100.

Syllophopsis as junior synonym of Monomorium: Bolton, 1987: 287.

[Syllopsis Santschi, 1921c: 120, incorrect subsequent spelling.]

Corynomyrmex Viehmeyer, 1916a: 133 [as subgenus of Monomorium]. Type-species: Monomorium (Corynomyrmex) hospitum, by monotypy.

Taxonomic history

Corynomyrmex as subgenus of Monomorium: Viehmeyer, 1916a: 133; Forel, 1917: 242; Emery, 1922e: 174.

Corynomyrmex as junior synonym of Monomorium: Ettershank, 1966: 82 [provisional]; Smith, D.R. 1979: 1381; Bolton, 1987: 287.

Isolcomyrmex Santschi, 1917b: 296 [as subgenus of Monomorium]. Type-species: Holcomyrmex santschii (junior secondary homonym in Monomorium, replaced by Monomorium santschianum), by monotypy.

Taxonomic history

Isolcomyrmex as subgenus of Monomorium: Santschi, 1917b: 296; Wheeler, W.M. 1922a: 676.

Isolcomyrmex as junior synonym of Monomorium: Ettershank, 1966: 82; Smith, D.R. 1979: 1381; Bolton, 1987: 296.

[Isholcomyrmex Santschi, 1936a: 32, incorrect subsequent spelling.]

Paraphacota Santschi, 1919d: 90. Type-species: Paraphacota surcoufi, by monotypy.

Taxonomic history

[Type-species not Phacota noualhieri, unjustified subsequent designation by Santschi, 1927d: 245; repeated in Ettershank, 1966: 82 and Bolton, 1973a: 352.]

Paraphacota in Myrmicinae, Solenopsidini: Wheeler, W.M. 1922a: 663; Donisthorpe, 1943f: 681.

Paraphacota as junior synonym of Monomorium: Santschi, 1927d: 243; Ettershank, 1966: 82; Smith, D.R. 1979: 1381; Bolton, 1987: 293.

Equestrimessor Santschi, 1919d: 92 [as subgenus of Monomorium]. Type-species: Holcomyrmex chobauti, by subsequent designation of Donisthorpe, 1943f: 644.

Taxonomic history

Equestrimessor as subgenus of Monomorium: Santschi, 1919d: 92;Wheeler, W.M. 1922a: 677; Santschi, 1936a: 32.

Equestrimessor as junior synonym of Monomorium: Ettershank, 1966: 82; Smith, D.R. 1979: 1381; Bolton, 1987: 287, 297.

[Equessimessor Santschi, 1936a: 32, incorrect subsequent spelling.]

Xenhyboma Santschi, 1919g: 405. Type-species: Xenhyboma mystes (junior synonym of Monomorium medinae), by monotypy.

Taxonomic history

Xenhyboma in Myrmicinae, Solenopsidini: Donisthorpe, 1943g: 736.

Xenhyboma as junior synonym of Monomorium: Ettershank, 1966: 82 [provisional]; Smith, D.R. 1979: 1381; Espadaler, 1982b: 112; Bolton, 1987: 293.

Protholcomyrmex Wheeler, W.M. 1922a: 162 [as subgenus of Monomorium]. Type-species: Monomorium rothsteini, by original designation.

Taxonomic history

Protholcomyrmex as junior synonym of Chelaner: Ettershank, 1966: 93.

Ireneidris Donisthorpe, 1943c: 81. Type-species: Ireneidris myops (junior synonym of Monomorium talpa), by original designation.

Taxonomic history

Ireneidris in Myrmicinae, Solenopsidini [subtribe Monomoriini]: Donisthorpe, 1943c: 82.

Ireneidris as junior synonym of Monomorium: Ettershank, 1966: 82; Smith, D.R. 1979: 1381; Bolton, 1987: 298.

Schizopelta McAreavey, 1949: 14. Type-species: Schizopelta falcata, by original designation.

Taxonomic history

Schizopelta as junior synonym of Chelaner: Ettershank, 1966: 93.

Nothidris Ettershank, 1966: 105. Type-species: Monomorium latastei, by original designation.

Taxonomic history

Nothidris in Myrmicinae, Megalomyrmex genus group: Ettershank, 1966: 81; in Solenopsis genus group: Bolton, 1987: 271.

Nothidris in Myrmicinae, Megalomyrmecini: Dlussky & Fedoseeva, 1988: 80.

Nothidris in Myrmicinae, Solenopsidini: Kempf, 1972a: 165; Hölldobler & Wilson, 1990: 16; Jaffe, 1993: 10; Bolton, 1994: 106.

Nothidris as junior synonym of Monomorium: Fernández, in Heterick, 2006: 79; Fernández, 2007b: 130.

Pharaophanes Bernard, 1967: 168 (attributed to Santschi) [as subgenus of Monomorium]. Type-species: Formica pharaonis, by monotypy.

Taxonomic history

[Pharaophanes Bernard, 1953b: 238 (attributed to Santschi) [as subgenus of Monomorium]. Unavailable name; proposed without designation of type-species. Species included by Bernard (1953) are all referable to Monomorium: Bolton, 1987: 288; Bolton, 1995b: 42 (see also note in Baroni Urbani, 1971c: 90).]

Pharaophanes as junior synonym of Monomorium: Bolton, 2003: 213.

Antichthonidris Snelling, R.R. 1975: 5. Type-species: Monomorium denticulatum, by original designation.

Taxonomic history

Antichthonidris in Myrmicinae, Solenopsis genus group: Bolton, 1987: 271.

Antichthonidris in Myrmicinae, Solenopsidini: Hölldobler & Wilson, 1990: 16; Jaffe, 1993: 10; Bolton, 1994: 106.

Antichthonidris as junior synonym of Monomorium: Heterick, 2001: 361.

Epelysidris Bolton, 1987: 279. Type-species: Epelysidris brocha, by original designation.

Taxonomic history

Epelysidris in Myrmicinae, Solenopsis genus group: Bolton, 1987: 271.

Epelysidris in Myrmicinae, Solenopsidini: Hölldobler & Wilson, 1990: 16; Bolton, 1994: 106; Bolton, 1995a: 1049; Bolton, 1995b: 188.

Epelysidris as junior synonym of Monomorium: Fernández, in Heterick, 2006: 79; Fernández, 2007b: 130.

Genus Monomorium references

Roger, 1863b: 30, 31 (Phacota, Monomorium catalogues); Mayr, 1863: 429, 440 (Monomorium, Phacota catalogues); Mayr, 1865: 19, 22 (Trichomyrmex, Monomorium, Phacota diagnoses); Mayr, 1867a: 95 (Monomorium diagnosis); Mayr, 1876: 100 (Australia Monomorium species key); Emery, 1881b: 530 (Mediterranean & Red Sea Monomorium species key); André, 1883a: 331, 343 (Europe & Algeria Monomorium, Holcomyrmex species keys); André, 1883b: 397 (Phacota); Cresson, 1887: 262 (U.S.A. Monomorium catalogue); Nasonov, 1889: 69 (Russia Monomorium species key); Dalla Torre, 1893: 65, 71, 72, 78 (Holcomyrmex, Monomorium, Phacota, Trichomyrmex, †Lampromyrmex catalogues); Forel, 1903a: 685, 692 (India & Sri Lanka Monomorium, Holcomyrmex species keys); Bingham, 1903: 200, 282 (India, Sri Lanka & Burma Monomorium, Holcomyrmex species keys); Ruzsky, 1905b: 633 (Russian Empire Monomorium species key); Emery, 1908h: 664 (Palaearctic Monomorium species key); Emery, 1915i: 190 (Monomorium subgenera key); Arnold, 1916: 203, 204 (Monomorium diagnosis, South Africa species key); Emery, 1916b: 161 (Italy Monomorium species key); Santschi, 1921c: 121 (Syllophopsis species key); Emery, 1922e: 166 (Monomorium diagnosis, subgenera key, catalogue); Emery, 1922e: 168 (M. (Chelaner) & M. (Notomyrmex) diagnoses, catalogues); Emery, 1922e: 170 (M. (Monomorium) diagnosis, catalogue); Emery, 1922e: 174 (M. (Corynomyrmex) diagnosis, catalogue); Emery, 1922e: 175 (M. (Syllophopsis) & M. (Xeromyrmex) diagnoses, catalogues); Emery, 1922e: 179 (M. (Parholcomyrmex) diagnosis, catalogue); Emery, 1922e: 181 (M. (Holcomyrmex) diagnosis, catalogue); Emery, 1922e: 183 (M. (Lampromyrmex) diagnosis, catalogue); Emery, 1922e: 184 (Epixenus diagnosis, catalogue); Emery, 1922e: 185 (Trichomyrmex diagnosis, catalogue); Emery, 1922e: 186 (Wheeleriella diagnosis, catalogue); Emery, 1922e: 187 (Phacota diagnosis, catalogue); Emery, 1922e: 204 (Epoecus diagnosis, catalogue); Wheeler, W.M. 1922a: 161, 675, 862, 1026 (Monomorium diagnosis, subgenera key, Afrotropical, Malagasy catalogues); Menozzi, 1933b: 64 (Israel Monomorium species key); Santschi, 1936a: 33 (Mediterranean & Africa M. (Xeromyrmex) key); Buren, 1944a: 289 (U.S.A., Iowa Monomorium species key); Kusnezov, 1949a: 431 (Argentina Monomorium species); Creighton, 1950a: 217 (North America Monomorium species key); Creighton, 1950a: 239 (Epoecus, review of genus); Chapman & Capco, 1951: 162, 169 (Asia Ireneidris, Monomorium, Trichomyrmex, Wheeleriella checklists); Bernard, 1955b: 282 (Epixenus species key); Brown, 1958h: 28 (New Zealand Monomorium species key); Gregg, 1963: 367 (U.S.A., Colorado Monomorium species key); Ettershank, 1966: 82, 93, 100 (Monomorium, Chelaner, Syllophopsis diagnoses, reviews of genera, checklists); Bernard, 1967: 165 (Monomorium diagnosis, Western Europe species key); Kempf, 1972a: 143 (Neotropical Monomorium catalogue); Kempf, 1972a: 165 (Nothidris catalogue); Alayo, 1974: 14 (Cuba Monomorium species key); Snelling, R.R. 1975: 5 (Nothidris all species key); Snelling, R.R. & Hunt, 1976: 79 (Nothidris all species key); Arnol'di & Dlussky, 1978: 538 (former European U.S.S.R. Monomorium species key); Collingwood, 1978: 82 (Iberian Peninsula Monomorium species key); Smith, D.R. 1979: 1381 (North America Monomorium catalogue); Tohmé & Tohmé, 1980a: 1087 (Lebanon & Syria Epixenus species key); Collingwood, 1985: 267 (Saudi Arabia Monomorium species key); Taylor & Brown, D.R. 1985: 55, 70 (Australia Chelaner, Monomorium catalogues); DuBois, 1986: 74 (Nearctic Monomorium species revision, key); Bolton, 1987: 281 (Phacota diagnosis, review of genus); Bolton, 1987: 283 (Antichthonidris, review of genus); Bolton, 1987: 284 (Nothidris review of genus); Bolton, 1987: 287 (Monomorium diagnosis, review of genus, Afrotropical species revision, key); Taylor, 1987a: 19, 40 and Taylor, 1987b: 2 (Australia, New Caledonia & New Zealand Monomorium checklist); Agosti & Collingwood, 1987b: 272 (Balkans Monomorium species key); Hölldobler & Wilson, 1990: 14 (synoptic classification); Dlussky, Soyunov & Zabelin, 1990: 231 (Turkmenistan Monomorium species key); Brandão, 1991: 326, 357 (Neotropical Monomorium catalogue); Brandão, 1991: 361 (Nothidris catalogue); Morisita, Kubota, et al. 1992: 38 (Japan Monomorium species key); Arakelian, 1994: 46 (Armenia Monomorium species key); Bolton, 1995a: 1048, 1050, 1051 (census); Bolton, 1995b: 67, 258, 292, 316 (Antichthonidris, Monomorium, Nothidris, Phacota catalogues); Wu, J. & Wang, 1995: 88 (China Monomorium species key); Collingwood & Agosti, 1996: 337 (Saudi Arabia Monomorium species key); Radchenko, 1997f: 211 (M. scabriceps group revision, key); Collingwood & Prince, 1998: 19 (Portugal Monomorium species key); Shattuck, 1999: 145 (Australia synopsis); Heterick, 2001: 367 (Australia Monomorium species revision, key); Zhou, 2001b: 112 (China, Guangxi Monomorium species key); Mackay & Mackay, 2002: 127 (U.S.A., New Mexico Monomorium species key); Heterick, 2006: 84 (Malagasy Monomorium species revision, key); Fernández, 2007b: 139 (Neotropical Monomorium species key); Terayama, 2009: 154 (Taiwan species key); Heterick, 2009: 158 (south-western Australia species key); Aldawood & Sharaf, 2011: 52 (Saudi Arabia M. monomorium group key).

Myrmicinae Tetramoriini Rhoptromyrmex TRUE TRUE FALSE

10 species

Rhoptromyrmex Mayr, 1901b: 18. Type-species: Rhoptromyrmex globinodis, by subsequent designation of Wheeler, W.M. 1911f: 172.

Taxonomic history

Rhoptromyrmex in Myrmicinae, Stenammini: Ashmead, 1905b: 383.

Rhoptromyrmex in Myrmicinae, Tetramoriini: Wheeler, W.M. 1910g: 141; Emery, 1914a: 42; Forel, 1917: 245; Arnold, 1917: 351; Wheeler, W.M. 1922a: 664; Emery, 1924d: 289; all subsequent authors.

Junior synonyms of RHOPTROMYRMEX

Hagioxenus Forel, 1910a: 8. Type-species: Hagioxenus schmitzi, by monotypy.

Taxonomic history

Hagioxenus in Myrmicinae, Solenopsidini: Emery, 1914a: 41 [subtribe Monomoriini]; Forel, 1917: 243; Emery, 1922e: 186; Wheeler, W.M. 1922a: 662; subsequent authors to the following.

Hagioxenus in Myrmicinae, Monomorium genus group: Ettershank, 1966: 81.

Hagioxenus as junior synonym of Rhoptromyrmex: Bolton, 1986a: 2.

Acidomyrmex Emery, 1915i: 191 [as subgenus of Rhoptromyrmex]. Type-species: Rhoptromyrmex wroughtonii, by original designation.

Taxonomic history

Acidomyrmex in Myrmicinae, Tetramoriini: Forel, 1917: 245; Wheeler, W.M. 1922a: 664; Emery, 1924d: 290; all subsequent authors.

Acidomyrmex as genus: Wheeler, W.M. 1922a: 664, 672; Chapman & Capco, 1951: 173.

Acidomyrmex as subgenus of Rhoptromyrmex: Emery, 1915i: 191; Forel, 1917: 245; Emery, 1924d: 290; Donisthorpe, 1943f: 619.

Acidomyrmex as junior synonym of Rhoptromyrmex: Brown, 1964c: 11; Bolton, 1976: 298; Bolton, 1986a: 2.

Ireneella Donisthorpe, 1941i: 175. Type-species: Ireneella papuensis (junior synonym of Tetramorium melleum), by original designation.

Taxonomic history

Ireneella in Myrmicinae, Tetramoriini: Donisthorpe, 1941i: 175. Donisthorpe, 1943f: 653.

Ireneella incertae sedis in Formicidae: Wheeler, G.C. & Wheeler, J. 1985: 259 (incomprehensible entry).

Ireneella as junior synonym of Rhoptromyrmex: Bolton, 1976: 298; Bolton, 1986a: 2.

Genus Rhoptromyrmex references

Arnold, 1917: 351 (diagnosis, South Africa species key); Wheeler, W.M. 1922a: 194, 908 (diagnosis, Afrotropical catalogue); Emery, 1922e: 186 (Hagioxenus diagnosis, catalogue); Emery, 1924d: 289 (Rhoptromyrmex diagnosis, catalogue); Emery, 1924d: 290 (R. (Acidomyrmex) diagnosis, catalogue); Chapman & Capco, 1951: 173, 174 (Asia Acidomyrmex, Ireneela, Rhoptromyrmex checklists); Brown, 1964c: 11 (diagnosis, all species revision, key); Bolton, 1976: 298 (diagnosis, all species key); Taylor & Brown, D.R. 1985: 86 (Australia catalogue); Bolton, 1986a: 5 (diagnosis, review of genus, all species key); Taylor, 1987a: 67 (Australia checklist); Bolton, 1994: 106 (synoptic classification); Bolton, 1995a: 1052 (census); Bolton, 1995b: 377 (catalogue); Shattuck, 1999: 163 (Australia synopsis).

Myrmicinae Tetramoriini Tetramorium TRUE TRUE FALSE

Extant: 461 species, 16 subspecies

Fossil:

Tetramorium Mayr, 1855: 423. Type-species: Formica caespitum, by subsequent designation of Girard, 1879: 1016.

Taxonomic history

Tetramorium in Myrmicidae, Myrmicidae: Emery, 1877a: 81.

Tetramorium in Myrmicinae: Mayr, 1855: 423 [Myrmicidae]; Mayr, 1861: 61 [Myrmicidae]; Mayr, 1865: 20 [Myrmicidae]; Emery & Forel, 1879a: 457 [Myrmicidae]; Dalla Torre, 1893: 130.

Tetramorium in Myrmicinae, Myrmicini: Forel, 1895b: 125; Forel, 1899c: 52; Kusnezov, 1964: 57 (anachronism).

Tetramorium in Myrmicinae, Tetramoriini: Emery, 1895j: 770; Ashmead, 1905b: 383; Wheeler, W.M. 1910g: 141; Emery, 1914a: 42; Arnold, 1917: 271; Forel, 1917: 245; Wheeler, W.M. 1922a: 664; Emery, 1924d: 275; all subsequent authors.

Junior synonyms of TETRAMORIUM

Tetrogmus Roger, 1857: 10. Type-species: Tetrogmus caldarius, by monotypy.

Taxonomic history

Tetrogmus as subgenus of Tetramorium: Wheeler, W.M. 1910g: 141.

Tetrogmus as junior synonym of Tetramorium: Roger, 1862c: 297; all subsequent authors except the above; Bolton, 1976: 359; Bolton, 1980: 195.

Xiphomyrmex Forel, 1887: 385 [as subgenus of Tetramorium]. Type-species: Tetramorium (Xiphomyrmex) kelleri, by subsequent designation of Wheeler, W.M. 1911f: 175.

Taxonomic history

Xiphomyrmex in Myrmicinae: Dalla Torre, 1893: 130.

Xiphomyrmex in Myrmicinae, Myrmicini: Ashmead, 1905b: 383.

Xiphomyrmex in Myrmicinae, Tetramoriini: Emery, 1895j: 770; Emery, 1914a: 42; Forel, 1917: 245; Wheeler, W.M. 1922a: 664; Emery, 1924d: 286; all subsequent authors.

Xiphomyrmex as subgenus of Tetramorium: Dalla Torre, 1893: 130; Forel, 1899c: 53; Forel, 1903a: 700; Wheeler, W.M. 1910g: 141; Arnold, 1917: 274; Arnold, 1926: 277.

Xiphomyrmex as genus: Emery, 1895j: 770; Emery, 1896e: 183; Emery, 1914a: 42; Forel, 1917: 245; Wheeler, W.M. 1922a: 193; Emery, 1924d: 286; Smith, M.R. 1938a: 126; Donisthorpe, 1943g: 736; Creighton, 1950a: 293; Smith, M.R. 1951a: 824; Chapman & Capco, 1951: 179; Bernard, 1953b: 250; Wheeler, G.C. & Wheeler, J. 1985: 257 (anachronism); Dlussky & Fedoseeva, 1988: 80 (anachronism).

Xiphomyrmex as junior synonym of Tetramorium: Bingham, 1903: 175; Bolton, 1976: 359; Bolton, 1980: 195; Bolton, 1994: 106.

Triglyphothrix Forel, 1890b: cvi. Type-species: Triglyphothrix walshi, by monotypy.

Taxonomic history

Triglyphothrix in Myrmicinae: Dalla Torre, 1893: 135.

Triglyphothrix in Myrmicinae, Tetramoriini: Emery, 1895j: 770; Ashmead, 1905b: 383; Wheeler, W.M. 1910g: 141; Emery, 1914a: 42; Forel, 1917: 245; Wheeler, W.M. 1922a: 664; Emery, 1924d: 272; all subsequent authors.

Triglyphothrix as subgenus of Tetramorium: Arnold, 1917: 274; Arnold, 1926: 272.

Triglyphothrix as genus: Forel, 1890b: cvi. Dalla Torre, 1893: 135; Emery, 1895j: 770; Forel, 1903a: 703; Bingham, 1903: 171; Wheeler, W.M. 1910g: 141; Emery, 1914a: 42; Forel, 1917: 245; Wheeler, W.M. 1922a: 196; Emery, 1924d: 272; Donisthorpe, 1943g: 734; Creighton, 1950a: 285; Smith, M.R. 1951a: 823; Chapman & Capco, 1951: 178; Bernard, 1953b: 248; Kempf, 1972a: 255; Bolton, 1976: 310; Smith, D.R. 1979: 1399; Wheeler, G.C. & Wheeler, J. 1985: 257; Taylor & Brown, D.R. 1985: 91.

Triglyphothrix as junior synonym of Tetramorium: Bolton, 1985: 247; Bolton, 1994: 106.

Atopula Emery, 1912b: 104. Type-species: Atopomyrmex nodifer, by original designation.

Taxonomic history

Atopula in Myrmicinae, Myrmecinini: Emery, 1912b: 105; Emery, 1914a: 41; Forel, 1917: 244; Wheeler, W.M. 1922a: 663; Emery, 1924d: 242 [subtribe Podomyrmini]; all subsequent authors to Bolton, 1976: 359.

Atopula in Myrmicinae, Podomyrmini: Dlussky & Fedoseeva, 1988: 79.

Atopula as subgenus of Vollenhovia: Emery, 1912d: 273.

Atopula as subgenus of Terataner: Arnold, 1952b: 129.

Atopula as genus: Emery, 1912b: 104; Emery, 1914a: 41; Wheeler, W.M. 1922a: 663; Emery, 1924d: 242; Bernard, 1948: 177; Wheeler, G.C. & Wheeler, J. 1985: 257 (anachronism); Dlussky & Fedoseeva, 1988: 79 (anachronism).

Atopula as junior synonym of Tetramorium: Bolton, 1976: 359; Bolton, 1980: 195; Bolton, 1994: 106.

Macromischoides Wheeler, W.M. 1920: 53. Type-species: Macromischa aculeata, by original designation.

Taxonomic history

Macromischoides in Myrmicinae, Leptothoracini: Wheeler, W.M. 1922a: 664; Wheeler, G.C. & Wheeler, J. 1985: 257 (anachronism).

Macromischoides in Myrmicinae, Tetramoriini: Santschi, 1924b: 207; all subsequent authors.

Macromischoides as genus: Wheeler, W.M. 1922a: 187; Santschi, 1924b: 206; Bernard, 1953b: 248; Dlussky & Fedoseeva, 1988: 80 (anachronism).

Macromischoides as junior synonym of Tetramorium: Bolton, 1976: 359; Bolton, 1980: 196.

[Macromichoides Santschi, 1924b: 206, incorrect subsequent spelling.]

Lobomyrmex Kratochvíl, 1941: 84 [as subgenus of Tetramorium]. Type-species: Tetramorium (Lobomyrmex) ferox silhavyi (junior synonym of Tetramorium ferox), by monotypy.

Taxonomic history

Lobomyrmex as junior synonym of Tetramorium: Brown, 1973b: 181 [provisional]; Bolton, 1976: 359; Bolton, 1980: 196.

Sulcomyrmex Kratochvíl, 1941: 84 [as subgenus of Tetramorium]. Unavailable name. Proposed without designation of type-species and therefore unavailable. Species included by Kratochvíl (1941) are all referable to Tetramorium: Bolton, 1976: 359.]

Apomyrmex Calilung, 2000: 66. Type-species: Apomyrmex manobo, by original designation.

Taxonomic history

Apomyrmex incertae sedis in Myrmicinae: Calilung, 2000: 66.

Apomyrmex as junior synonym of Tetramorium: Bolton, 2003: 227, 269.

Genus Tetramorium references

Roger, 1863b: 26, 27 (catalogue); Mayr, 1863: 456 (catalogue); Mayr, 1865: 20 (diagnosis); Mayr, 1870b: 972 (all species key); André, 1883a: 285 (Europe & Algeria species key); Cresson, 1887: 261 (U.S.A. catalogue); Emery, 1893e: 214 (Triglyphothrix species key); Dalla Torre, 1893: 130, 135 (Tetramorium, Triglyphothrix catalogues); Forel, 1903a: 700, 703 (India & Sri Lanka Tetramorium, Triglyphothrix species keys); Bingham, 1903: 172, 175 (India, Sri Lanka & Burma Triglyphothrix, Tetramorium species keys); Ruzsky, 1905b: 517 (Russian Empire species key); Emery, 1909d: 695 (Palaearctic species key); Emery, 1916b: 195 (Italy species key); Arnold, 1917: 271, 275, 334, 345 (diagnosis, South Africa Tetramorium (Tetramorium), T. (Triglyphothrix), T. (Xiphomyrmex) species keys); Bondroit, 1918: 106 (France & Belgium species key); Mann, 1919: 353 (Papuasia Triglyphothrix species key); Wheeler, W.M. 1922a: 187, 190, 193, 196 (Macromischoides, Tetramorium, Xiphomyrmex, Triglyphothrix diagnoses); Wheeler, W.M. 1922a: 886, 889, 893, 906, 909 (Afrotropical Atopula, Macromischoides, Tetramorium, Xiphomyrmex, Triglyphothrix catalogues); Wheeler, W.M. 1922a: 1030, 1031, 1032 (Malagasy Tetramorium, Xiphomyrmex, Triglyphothrix catalogues); Emery, 1924d: 242 (Atopula diagnosis, catalogue); Emery, 1924d: 272 (Triglyphothrix diagnosis, catalogue); Emery, 1924d: 275 (Tetramorium diagnosis, catalogue); Emery, 1924d: 286 (Xiphomyrmex diagnosis, catalogue); Santschi, 1924b: 210 (Macromischoides species key); Arnold, 1926: 241 (South Africa Tetramorium (Tetramorium) species key); Menozzi, 1933b: 74 (Israel species key); Finzi, 1936: 183 (Egypt species key); Smith, M.R. 1938a: 127 (North America Xiphomyrmex species key); Kratochvíl, 1941: 84 (Central Europe species key); Novák & Sadil, 1941: 84 (Central Europe species key); Smith, M.R. 1943b: 2 (North America species key); Kratochvíl, Novák & Snoflák, 1944: 76, 95 (Central Europe species key); Creighton, 1950a: 290, 293 (North America Tetramorium, Xiphomyrmex species key); Chapman & Capco, 1951: 174, 178, 179 (Asia Tetramorium, Triglyphothrix, Xiphomyrmex checklists); Brown, 1958h: 27 (New Zealand species); Bernard, 1967: 227 (diagnosis, Western Europe species key); Kempf, 1972a: 249, 255 (Neotropical Tetramorium, Triglyphothrix catalogues); Alayo, 1974: 18 (Cuba species key); Tarbinsky, 1976: 105 (Kyrghyzstan species key); Bolton, 1976: 310, 314, 341 (Triglyphothrix diagnosis, Afrotropical, Oriental & Malesian species revisions, keys); Bolton, 1976: 359 (Tetramorium diagnosis, review of genus, synonymy); Bolton, 1977: 72, 133 (Oriental & Malesian, Australia species revisions, keys); Kutter, 1977c: 150 (Switzerland species key); Arnol'di & Dlussky, 1978: 544 (former European U.S.S.R. species key); Collingwood, 1978: 86 (Iberian Peninsula species key); Collingwood, 1979: 82 (Fennoscandia & Denmark species key); Smith, D.R. 1979: 1399, 1400 (North America Triglyphothrix, Tetramorium catalogues); Bolton, 1979: 132, 159 (Malagasy, New World species revisions, keys); Bolton, 1980: 196, 205 (diagnosis, review of genus, Afrotropical species revision, key); Collingwood, 1985: 262 (Saudi Arabia species key); Gösswald, 1985: 311 (Germany species key); Taylor & Brown, D.R. 1985: 88 (Australia catalogue); Taylor, 1987a: 78 (Australia, New Caledonia & New Zealand checklist); Agosti & Collingwood, 1987b: 277 (Balkans species key); Wang, M., Xiao & Wu, 1988: 264 (China species key); Radchenko & Arakelian, 1990: 371 (Crimea & Caucasus T. ferox complex, key); Dlussky, Soyunov & Zabelin, 1990: 197 (Turkmenistan species key); Atanasov & Dlussky, 1992: 149 (Bulgaria species key); Radchenko, 1992a: 44 (former U.S.S.R. species key); Morisita, Kubota, et al. 1992: 33 (Japan species key); Arakelian, 1994: 60 (Armenia species key); Radchenko, 1994b: 109 (South Siberia species key); Bolton, 1994: 106 (synoptic classification); Bolton, 1995a: 1053 (census); Bolton, 1995b: 403 (catalogue); Wu, J. & Wang, 1995: 79 (China species key); Collingwood & Agosti, 1996: 332 (Saudi Arabia species key); Seifert, 1996b: 158 (Central Europe species key); Cagniant, 1997: 89 (Morocco species key); Radchenko, Czechowski & Czechowsa, 1998: 107 (Poland species key); Collingwood & Prince, 1998: 17 (Portugal species key); Sanetra, Güsten & Schulz, 1999: 317 (Italy, species); Shattuck, 1999: 168 (Australia synopsis); Chang & He, 2001a: 1 (northwest China species key); Zhou, 2001b: 100 (China, Guangxi species key); Czechowski, Radchenko & Czechowska, 2002: 145 (Poland species key); Mackay & Mackay, 2002: 214 (U.S.A., New Mexico species key); Seifert, 2007: 145 (North and Central Europe species key); Csösz, Radchenko & Schulz, 2007: 8 (T. chefketi complex key); Terayama, 2009: 162 (Taiwan species key); Heterick, 2009: 177 (south-western Australia species key); Csösz & Schulz, 2010: 12 (T. ferox complex key); Hita Garcia, Fischer & Peters, 2010b: 12 (Afrotropical T. weitzeckeri species group key).

Myrmicinae Pheidolini Ocymyrmex TRUE TRUE FALSE

37 species

Ocymyrmex Emery, 1886: 364. Type-species: Ocymyrmex barbiger, by monotypy.

Taxonomic history

Ocymyrmex in Myrmicinae: Dalla Torre, 1893: 72.

Ocymyrmex in Myrmicinae, Myrmicini: Emery, 1895j: 769; Wheeler, W.M. 1910g: 139.

Ocymyrmex in Myrmicinae, Ocymyrmecini: Emery, 1914a: 41; Arnold, 1916: 194; Forel, 1917: 245; Wheeler, W.M. 1922a: 664; Emery, 1924d: 270; all subsequent authors to the following.

Ocymyrmex in Myrmicinae, Pheidolini: Bolton & Marsh, 1989: 1281; Hölldobler & Wilson, 1990: 16; Bolton, 1994: 106

Genus Ocymyrmex references

Emery, 1892a: 116 (all species key); Arnold, 1916: 194 (diagnosis, South Africa species key); Wheeler, W.M. 1922a: 891 (catalogue); Emery, 1924d: 270 (diagnosis, catalogue); Bolton, 1981b: 260 (diagnosis, all species revision, key); Bolton & Marsh, 1989: 1267 (diagnosis, revised key); Bolton, 1995a: 1051 (census); Bolton, 1995b: 294 (catalogue).

Myrmicinae Pheidolini Pheidole TRUE TRUE FALSE

Extant: 970 species, 142 subspecies

Fossil: 4 species

Pheidole Westwood, 1839: 219. Type-species: Atta providens, by monotypy.

Taxonomic history

Pheidole in Poneridae, Attidae: Smith, F. 1858b: 172.

Pheidole in Attidae: Smith, F. 1860a: 74; Smith, F. 1862a: 49.

Pheidole in Myrmicidae: Smith, F. 1871a: 331; Cresson, 1887: 262.

Pheidole in Myrmicidae, Pheidolidae: Emery, 1877a: 81.

Pheidole in Myrmicinae: Mayr, 1861: 69 [Myrmicidae]; Mayr, 1865: 22 [Myrmicidae]; Emery & Forel, 1879a: 463 [Myrmicidae]; Dalla Torre, 1893: 88.

Pheidole in Myrmicinae, Myrmicini: Forel, 1895b: 127; Emery, 1895j: 769; Forel, 1899c: 64; Ashmead, 1905b: 383; Wheeler, W.M. 1910g: 139; Kempf, 1972a: 183 (anachronism).

Pheidole in Myrmicinae, Pheidolini: Forel, 1893a: 165; Emery, 1914a: 40; Forel, 1917: 241; Arnold, 1920a: 414; Emery, 1921f: 77; Wheeler, W.M. 1922a: 661; all subsequent authors except Kempf, 1972a: 183, above.

Junior synonyms of PHEIDOLE

Oecophthora Heer, 1852: 15. Type-species: Oecophthora pusilla (junior synonym of Formica megacephala), by monotypy.

Taxonomic history

Oecophthora in Myrmicinae: Mayr, 1855: 453 [Myrmicidae].

Oecophthora as junior synonym of Pheidole: Smith, F. 1858a: 282; Smith, F. 1858b: 172; Roger, 1863b: 30.

Ischnomyrmex Mayr, 1862: 738. Type-species: Myrmica longipes, by monotypy.

Taxonomic history

Ischnomyrmex in Myrmicinae: Mayr, 1865: 18 [Myrmicidae]; Dalla Torre, 1893: 98.

Ischnomyrmex in Myrmicinae, Myrmicini: Emery, 1895j: 769; Ashmead, 1905b: 383; Wheeler, W.M. 1910g: 140.

Ischnomyrmex in Myrmicinae, Pheidolini: Emery, 1914a: 40; Forel, 1917: 241; Emery, 1922e: 113; Wheeler, W.M. 1922a: 661; all subsequent authors.

Ischnomyrmex as junior synonym of Aphaenogaster: Emery & Forel, 1879a: 461; Bingham, 1903: 270.

Ischnomyrmex as subgenus of Aphaenogaster: Forel, 1891b: 166; Dalla Torre, 1893: 98; Forel, 1899c: 59; Emery, 1908c: 313; Wheeler, W.M. 1910g: 140.

Ischnomyrmex as subgenus of Stenamma: Emery, 1895c: 298; Emery, 1895j: 769; Forel, 1902h: 439; Forel, 1903a: 693.

Ischnomyrmex as subgenus of Pheidole: Forel, 1913k: 49.

Ischnomyrmex as genus: Mayr, 1862: 738; Emery, 1914a: 40; Viehmeyer, 1914d: 601; Forel, 1917: 241; Wheeler, W.M. 1922a: 661; Emery, 1922e: 113. Stitz, 1911a: 367; Kusnezov, 1952b: 10 (in key).

Ischnomyrmex as junior synonym of Pheidole: Brown, 1973b: 181 [provisional]; Smith, D.R. 1979: 1365; Snelling, R.R. 1981: 395.

Leptomyrma Motschoulsky, 1863: 17. Type-species: Leptomyrma gracilipes, by monotypy.

Taxonomic history

Leptomyrma in Myrmicidae: Smith, F. 1871a: 329.

Leptomyrma as junior synonym of Pheidole: Emery, 1892b: 166; all subsequent authors.

Pheidolacanthinus Smith, F. 1865: 75. Type-species: Pheidolacanthinus armatus (junior synonym of Myrmica quadrispinosa), by monotypy.

Taxonomic history

Pheidolacanthinus in Myrmicidae: Smith, F. 1871a: 333.

Pheidolacanthinus in Myrmicinae: Dalla Torre, 1893: 98.

Pheidolacanthinus as subgenus of Pheidole: Emery, 1915i: 190; Forel, 1917: 241; Emery, 1921f: 81; Wheeler, W.M. 1922a: 673; all subsequent authors to the following.

Pheidolacanthinus as junior synonym of Pheidole: Brown, 1973b: 183 [provisional]; Smith, D.R. 1979: 1365; Snelling, R.R. 1981: 395.

Ceratopheidole Pergande, 1896: 889 [as subgenus of Pheidole]. Type-species: Pheidole (Ceratopheidole) granulata, by monotypy.

Taxonomic history

Ceratopheidole in Myrmicinae, Pheidolini: Emery, 1914a: 40; Forel, 1917: 241; Emery, 1922e: 112; Wheeler, W.M. 1922a: 661; all subsequent authors.

Ceratopheidole as genus: Emery, 1914a: 40; Forel, 1917: 241; Emery, 1922e: 112; Wheeler, W.M. 1922a: 661; Chapman & Capco, 1951: 135; Wu, J. & Wang, 1995: 104 (anachronism).

Ceratopheidole as subgenus of Pheidole: Pergande, 1896: 889; Kempf, 1972a: 183.

Ceratopheidole as junior synonym of Pheidole: Brown, 1973b: 179 [provisional]; Smith, D.R. 1979: 1365; Snelling, R.R. 1981: 395; Wilson, 2003: 6.

Epipheidole Wheeler, W.M. 1903f: 664. Type-species: Epipheidole inquilina, by monotypy.

Taxonomic history

[Epipheidole also described as new by Wheeler, W.M. 1904a: 14.]

Epipheidole in Myrmicinae, Stenammini: Ashmead, 1905b: 383.

Epipheidole in Myrmicinae, Myrmicini: Wheeler, W.M. 1910g: 140.

Epipheidole in Myrmicinae, Pheidolini: Emery, 1914a: 40; Forel, 1917: 241; Emery, 1922e: 114; Wheeler, W.M. 1922a: 661; Creighton, 1950a: 192.

Epipheidole as junior synonym of Pheidole: Cole, 1965: 174; Smith, D.R. 1979: 1366.

Phidole Bingham, 1903: 220, unjustified emendation of Pheidole.

Taxonomic history

Phidole as junior synonym of Pheidole: Wheeler, 1922a: 806.

Sympheidole Wheeler, W.M. 1904a: 7. Type-species: Sympheidole elecebra, by monotypy.

Taxonomic history

Sympheidole in Myrmicinae, Myrmicini: Wheeler, W.M. 1910g: 140.

Sympheidole in Myrmicinae, Stenammini: Ashmead, 1905b: 383.

Sympheidole in Myrmicinae, Pheidolini: Emery, 1914a: 40; Forel, 1917: 241; Emery, 1922e: 115; Wheeler, W.M. 1922a: 661; all subsequent authors to the following.

Sympheidole as junior synonym of Pheidole: Brown, 1973b: 185 [provisional]; Smith, D.R. 1979: 1365; Snelling, R.R. 1981: 395; Wilson, 2003: 6.

Phidola Schulz, W.A. 1906: 155, unjustified emendation of Pheidole.

Taxonomic history

Phidola as junior synonym of Pheidole: Smith, D.R. 1979: 1365.

Allopheidole Forel, 1912f: 237 [as subgenus of Pheidole]. Type-species: Pheidole kingi (junior synonym of Pheidole tepicana), by subsequent designation of Wheeler, W.M. 1913a: 79.

Taxonomic history

Allopheidole as subgenus of Pheidole: Forel, 1912f: 237; Forel, 1917: 241.

Allopheidole as junior synonym of Pheidole: Emery, 1915i: 190; Emery, 1921f: 84; Smith, D.R. 1979: 1366.

Decapheidole Forel, 1912f: 237 [as subgenus of Pheidole]. Type-species: Pheidole perpusilla, by subsequent designation of Wheeler, W.M. 1913a: 80.

Taxonomic history

Decapheidole in Myrmicinae, Pheidolini: Wheeler, W.M. 1922a: 661.

Decapheidole as genus: Wheeler, W.M. 1922a: 661; Borgmeier, 1927c: 78.

Decapheidole as subgenus of Pheidole: Forel, 1912f: 237, Forel, 1917: 241; Emery, 1922e: 112; Kempf, 1972a: 183.

Decapheidole as junior synonym of Pheidole: Brown, 1973b: 180 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395; Wilson, 2003: 6.

Isopheidole Forel, 1912m: 765 [as subgenus of Pheidole]. Type-species: Myrmica longipes, by monotypy.

Taxonomic history

Isopheidole as junior synonym of Ischnomyrmex: Forel, 1913b: 350 (in text); Forel, 1913k: 49; Emery, 1922e: 113; Wheeler, W.M. 1922a: 680 (in key). [Ischnomyrmex and Isopheidole share the same type-species, synonymy is therefore absolute.]

Elasmopheidole Forel, 1913k: 43 [as subgenus of Pheidole]. Type-species: Pheidole aberrans, by subsequent designation of Emery, 1915i: 190.

Taxonomic history

Elasmopheidole as subgenus of Pheidole: Forel, 1913k: 43; Emery, 1915i: 190; Forel, 1917: 241; Emery, 1922e: 110; Wheeler, W.M. 1922a: 673; all subsequent authors to the following.

Elasmopheidole as junior synonym of Pheidole: Brown, 1973b: 180 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395.

Cardiopheidole Wheeler, W.M. 1914b: 48 [as subgenus of Pheidole]. Type-species: Pheidole vaslitii, by original designation.

Taxonomic history

Cardiopheidole as subgenus of Pheidole: Wheeler, W.M. 1914b: 48; Forel, 1917: 241.

Cardiopheidole as junior synonym of Pheidole: Emery, 1915i: 190; Emery, 1921f: 84; Smith, D.R. 1979: 1366.

Anergatides Wasmann, 1915b: 281. Type-species: Anergatides kohli (junior secondary homonym in Pheidole, replaced by Pheidole neokohli), by monotypy.

Taxonomic history

Anergatides in Myrmicinae, Pheidolini: Forel, 1917: 243 [subtribe Anergatini]; Emery, 1922e: 116 [subtribe Anergatidini]; Donisthorpe, 1943f: 623.

Anergatides in Myrmicinae, Solenopsidini: Wheeler, W.M. 1922a: 663; Wheeler, G.C. & Wheeler, J. 1985: 257 (anachronism).

Anergatides as junior synonym of Pheidole: Brown, 1973b: 178 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395; Wilson, 1984: 328; Bolton, 1994: 106.

Parapheidole Emery, 1915d: 68. Type-species: Aphaenogaster oculata, by monotypy.

Taxonomic history

Parapheidole in Myrmicinae, Pheidolini: Forel, 1917: 241; Emery, 1922e: 115; Wheeler, W.M. 1922a: 661; all subsequent authors.

Parapheidole as junior synonym of Pheidole: Brown, 1973b: 183 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395.

Macropheidole Emery, 1915i: 190 [as subgenus of Pheidole]. Type-species: Pheidole fimbriata, by monotypy.

Taxonomic history

Macropheidole as subgenus of Pheidole: Emery, 1915i: 190; Forel, 1917: 241; Emery, 1921f: 81; Wheeler, W.M. 1922a: 673; all subsequent authors to the following.

Macropheidole as junior synonym of Pheidole: Smith, M.R. 1951a: 800; Brown, 1973b: 181 [provisional]; Smith, D.R. 1979: 1366.

Scrobopheidole Emery, 1915i: 190 [as subgenus of Pheidole]. Type-species: Pheidole scrobifera, by monotypy.

Taxonomic history

Scrobopheidole as subgenus of Pheidole: Forel, 1917: 241; Emery, 1922e: 112; Wheeler, W.M. 1922a: 673; all subsequent authors to the following.

Scrobopheidole as junior synonym of Pheidole: Brown, 1973b: 184 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395; Wilson, 2003: 6.

Stegopheidole Emery, 1915i: 190 [as subgenus of Pheidole]. Type-species: Pheidole (Elasmopheidole) upeneci, by monotypy.

Taxonomic history

Stegopheidole as subgenus of Pheidole: Emery, 1921f: 83; Wheeler, W.M. 1922a: 673; all subsequent authors to the following.

Stegopheidole as junior synonym of Pheidole: Brown, 1973b: 185 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395.

Trachypheidole Emery, 1915i: 190 [as subgenus of Pheidole]. Type-species: Pheidole bicornis, by original designation.

Taxonomic history

Trachypheidole as subgenus of Pheidole: Forel, 1917: 241; Emery, 1922e: 111; Wheeler, W.M. 1922a: 673; all subsequent authors to the following.

Trachypheidole as junior synonym of Pheidole: Brown, 1973b: 185 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395; Wilson, 2003: 6.

Electropheidole Mann, 1921: 438 [as subgenus of Pheidole]. Type-species: Pheidole (Electropheidole) roosevelti, by subsequent designation of Donisthorpe, 1943f: 642.

Taxonomic history

Electropheidole as subgenus of Pheidole: Mann, 1921: 438; Wheeler, W.M. 1922a: 672; all subsequent authors to the following.

Electropheidole as junior synonym of Pheidole: Brown, 1973b: 180 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395.

Bruchomyrma Santschi, 1922d: 248. Type-species: Bruchomyrma acutidens, by monotypy.

Taxonomic history

Bruchomyrma in Myrmicinae, Pheidolini: Donisthorpe, 1943f: 629.

Bruchomyrma in Myrmicinae, Myrmicini: Kempf, 1972a: 42.

Bruchomyrma as junior synonym of Pheidole: Brown, 1973b: 179 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395; Wilson, 1984: 327; Bolton, 1994: 106; Wilson, 2003: 6.

Bruchomyrma as genus: Dlussky & Fedoseeva, 1988: 80 (anachronism).

Cephalomorium Forel, 1922: 91 [as subgenus of Tetramorium]. Type-species: Tetramorium (Cephalomorium) bahai, by monotypy.

Taxonomic history

Cephalomorium as junior synonym of Hendecapheidole: Santschi, 1925d: 228.

Hendecapheidole Wheeler, W.M. 1922d: 3 [as subgenus of Pheidole]. Type-species: Pheidole tachigaliae, by original designation.

Taxonomic history

Hendecapheidole as junior synonym of Pheidole: Brown, 1973b: 181 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395; Wilson, 2003: 6.

Gallardomyrma Bruch, 1932: 271. Type-species: Gallardomyrma argentina, by original designation.

Taxonomic history

Gallardomyrma in Myrmicinae, Pheidolini: Donisthorpe, 1943f: 646.

Gallardomyrma in Myrmicinae, Myrmicini: Kempf, 1972a: 110.

Gallardomyrma as junior synonym of Pheidole: Brown, 1973b: 180 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395; Wilson, 1984: 327; Wilson, 2003: 6.

Gallardomyrma as genus: Kempf, 1972a: 110; Dlussky & Fedoseeva, 1988: 80 (anachronism).

Conothoracoides Strand, 1935: 176.

Taxonomic history

[Replacement name for Conothorax Karavaiev; junior homonym of Conothorax Jekel, 1854: 9 bis (foldout pages) (Coleoptera).]

Conothoracoides as junior synonym of Pheidole: Brown, 1973b: 179 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395.

Homonym replaced by Conothoracoides

Conothorax Karavaiev, 1935a: 75. Type-species: Conothorax bilobum, by monotypy.

Taxonomic history

[Conothorax Karavaiev junior homonym of Conothorax Jekel, 1854: 9 bis (foldout pages) (Coleoptera).]

Conothorax in Myrmicinae, Pheidolini: Karavaiev, 1935a: 75.

Eriopheidole Kusnezov, 1952a: 10. Type-species: Eriopheidole symbiotica (unresolved junior secondary homonym in Pheidole), by monotypy.

Taxonomic history

Eriopheidole in Myrmicinae, Pheidolini: Kusnezov, 1952a: 10.

Eriopheidole in Myrmicinae, Myrmicini: Kempf, 1972a: 107.

Eriopheidole junior synonym of Pheidole: Brown, 1973b: 180 [provisional]; Smith, D.R. 1979: 1366; Snelling, R.R. 1981: 395; Bolton, 1994: 106.

Eriopheidole as genus: Kempf, 1972a: 107; Dlussky & Fedoseeva, 1988: 80 (anachronism).

Xenoaphaenogaster Baroni Urbani, 1964b: 50. Type-species: Xenoaphaenogaster inquilina (junior synonym of Myrmica pallidula), by original designation.

Taxonomic history

Xenoaphaenogaster as junior synonym of Monomorium: Brown, 1973b: 185 [provisional].

Xenoaphaenogaster as junior synonym of Pheidole: Bolton, 1987: 291.

Genus Pheidole references

Smith, F. 1858b: 172 (diagnosis); Mayr, 1861: 70 (Europe species key); Roger, 1863b: 27, 30 (catalogue); Mayr, 1863: 424, 440 (Ischnomyrmex, Pheidole catalogues); Mayr, 1865: 18, 22 (Ischnomyrmex, Pheidole diagnoses); Mayr, 1867a: 92, 96 (Ischnomyrmex, Pheidole diagnoses); Mayr, 1870b: 977, 979 (Australia & New World species keys); Mayr, 1876: 102 (Australia species key); André, 1883b: 382 (Europe & Algeria species key); Mayr, 1879: 674 (Asia species key); Mayr, 1887: 582 (New World species key); Cresson, 1887: 262 (U.S.A., catalogue); Nasonov, 1889: 75 (Russia species key); Dalla Torre, 1893: 88, 98 (Pheidole, Pheidolacanthinus catalogues); Emery, 1895c: 288 (North America species key); Mayr, 1896: 241 (Africa species, partial key); Emery, 1896g: 80 (P. flavens group, key); Forel, 1902c: 165 (India species key); Forel, 1902f: 523 (India & Sri Lanka species key); Bingham, 1903: 223 (India, Sri Lanka & Burma species key); Wheeler, W.M. 1908a: 133 (Puerto Rico species key); Arnold, 1920a: 414 (diagnosis, South Africa species key); Emery, 1921f: 78 (diagnosis, subgenera key, catalogue); Emery, 1921f: 81 (P. (Macropheidole) & P. (Pheidolacanthinus) diagnoses, catalogues); Emery, 1921f: 83 (P. (Stegopheidole) diagnosis, catalogue); Mann, 1921: 442 (Fiji Is species key); Emery, 1922e: 110 (P. (Elasmopheidole) diagnosis, catalogue); Emery, 1922e: 111 (P. (Trachypheidole) diagnosis, catalogue); Emery, 1922e: 112 (P. (Scrobopheidole), P. (Decapheidole) & P. (Ceratopheidole) diagnoses, catalogues); Emery, 1922e: 113 (Ischnomyrmex diagnosis, catalogue); Emery, 1922e: 114 (Epipheidole diagnosis, catalogue); Emery, 1922e: 115 (Sympheidole, Parapheidole diagnoses, catalogues); Emery, 1922e: 116 (Anergatides diagnosis, catalogue); Wheeler, W.M. 1922a: 126, 672 (diagnosis, subgenera key); Wheeler, W.M. 1922a: 806, 879 (Afrotropical Pheidole, Anergatides catalogues); Wheeler, W.M. 1922a: 1017, 1021 (Malagasy Pheidole, Parapheidole catalogues); Gallardo, 1932c: 179 (Argentina Elasmopheidole species key); Finzi, 1936: 165 (Egypt species key); Menozzi, 1939a: 298 (Himalaya species key); Cole, 1942: 362 (U.S.A., Utah species key); Buren, 1944a: 285 (U.S.A., Iowa species key); Creighton, 1950a: 163 (North America species key); Creighton, 1950a: 192, 194 (North America Epipheidole, Sympheidole species); Chapman & Capco, 1951: 135, 137 (Asia Ceratopheidole, Conothorax, Ischnomyrmex, Pheidole checklists); Kusnezov, 1952b: 61 (Argentina species key); Smith, M.R. 1955c: 302 (P. (Hendecapheidole) species key); Gregg, 1959: 9 (U.S.A. species key); Gregg, 1963: 407 (U.S.A., Colorado species key); Wheeler, G.C. & Wheeler, J. 1963: 126 (U.S.A., North Dakota species key); Bernard, 1967: 149 (diagnosis, Western Europe species key); Gregg, 1969: 95 (addition to Gregg, 1959 U.S.A. key); Kempf, 1972a: 183 (Neotropical catalogue); Alayo, 1974: 11 (Cuba species key); Smith, D.R. 1979: 1365 (North America catalogue); Allred, 1982: 441 (U.S.A., Utah species key); Ogata, 1982: 191 (Japan species key); Collingwood, 1985: 253 (Saudi Arabia species key); Naves, 1985: 54 (U.S.A., Florida species key); Taylor & Brown, D.R. 1985: 74 (Australia catalogue); Wheeler, G.C. & Wheeler, J. 1986g: 41 (U.S.A., Nevada species key); Taylor, 1987a: 53 (Australia, New Caledonia & New Zealand checklists); Agosti & Collingwood, 1987b: 271 (Balkans species key); Hölldobler & Wilson, 1990: 15 (synoptic classification); Kupyanskaya, 1990: 123 (Far Eastern Russia species key); Brandão, 1991: 368 (Neotropical catalogue); Morisita, Kubota, et al. 1992: 22 (Japan species key); Bolton, 1994: 106 (synoptic classification); Kupyanskaya, 1995: 345 (Far Eastern Russia species key); Bolton, 1995a: 1051 (census); Bolton, 1995b: 316 (catalogue); Wu, J. & Wang, 1995: 99 (China species key); Collingwood & Agosti, 1996: 322 (Saudi Arabia species key); Park & Kim, 2000: 109 (Korea species key); Shattuck, 1999: 152 (Australia synopsis); Eguchi, 1999: 103 (Borneo P. longipes group, key); Ward, 2000: 96 (U.S.A. P. hyatti complex, key); Zhou, 2001b: 121 (China, Guangxi species key); Eguchi, 2001b: 10 (Borneo species key); ; Mackay & Mackay, 2002: 149 (U.S.A., New Mexico species key)Wilson, 2003: 27 (New World, all species revision, keys); Sarnat, 2008: 8 (Fiji Is P. roosevelti group, key); Eguchi, 2008: 8 (North Vietnam species key); Terayama, 2009: 171 (Taiwan species key); Heterick, 2009: 169 (south-western Australia species key); Sarnat & Moreau, 2011: 114 (P. roosevelti group phylogeny and radiation).

Genera incertae sedis in Pheidolini

Myrmicinae Formicoxenini Leptothorax TRUE TRUE FALSE

Extant: 18 species, 2 subspecies

Fossil:

Leptothorax Mayr, 1855: 431. Type-species: Formica acervorum, by subsequent designation of Bingham, 1903: 214.

Taxonomic history

[Type-species not Myrmica clypeata, unjustified subsequent designation by Emery, 1912d: 271; repeated in Wheeler, W.M. 1913a: 79 and Emery, 1924d: 248.]

Leptothorax in Myrmicidae, Myrmicidae: Emery, 1877a: 81.

Leptothorax in Myrmicinae: Mayr, 1855: 431 [Myrmicidae]; Mayr, 1861: 57 [Myrmicidae]; Mayr, 1865: 20 [Myrmicidae]; Emery & Forel, 1879a: 458 [Myrmicidae]; Dalla Torre, 1893: 122.

Leptothorax in Myrmicinae, Myrmicini: Forel, 1895b: 125; Emery, 1895j: 769; Forel, 1899c: 54; Wheeler, W.M. 1910g: 139; Wheeler, W.M. 1915h: 63; Kusnezov, 1964: 57 (anachronism).

Leptothorax in Myrmicinae, Stenammini: Ashmead, 1905b: 383.

Leptothorax in Myrmicinae, Leptothoracini: Emery, 1914a: 42; Arnold, 1916: 257; Forel, 1917: 244; Wheeler, W.M. 1922a: 664; Emery, 1924d: 247; subsequent authors to the following.

Leptothorax in Myrmicinae, Formicoxenini: Bolton, 1994: 105; Mackay, 2000: 267.

Junior synonyms of LEPTOTHORAX

Mychothorax Ruzsky, 1904a: 288. Type-species: Formica acervorum, by original designation.

Taxonomic history

Mychothorax as subgenus of Leptothorax: Ruzsky, 1905b: 609; Emery, 1915g: 24; Emery, 1916b: 176; Forel, 1917: 245; Wheeler, W.M. 1922a: 679; Emery, 1924d: 260; Creighton, 1950a: 274.

Mychothorax as junior synonym of Leptothorax: Smith, M.R. 1950: 29; Brown, 1973b: 182 [provisional]; Bolton, 1982: 319. [Leptothorax and Mychothorax share the same type-species, synonymy is therefore absolute.]

Doronomyrmex Kutter, 1945: 485. Type-species: Doronomyrmex pacis, by monotypy.

Taxonomic history

Doronomyrmex in Myrmicinae, Leptothoracini:Dlussky & Fedoseeva, 1988: 79.

Doronomyrmex in Myrmicinae, Formicoxenini: Bolton, 1994: 105.

Doronomyrmex as junior synonym of Leptothorax: Brown, 1973b: 180 [provisional]; Heinze, 1998: 195; Bolton, 2003: 270.

Genus Leptothorax references

Mayr, 1855: 433 (Austria species key); Mayr, 1861: 58 (Europe species key); Roger, 1863b: 26, 28, 30 (catalogue); Mayr, 1863: 426, 428, 456 (Leptothorax, Macromischa, Temnothorax catalogues); Mayr, 1865: 19, 20, 21 (Macromischa, Leptothorax, Temnothorax diagnoses); Mayr, 1868c: 83 (†Baltic Amber species key); André, 1874: 188 (Europe species key); Forel, 1874: 84 (Switzerland species key); André, 1883a: 293 (Europe & Algeria species key); Mayr, 1886d: 451 (U.S.A. species key); Cresson, 1887: 261 (U.S.A. catalogue); Nasonov, 1889: 70 (Russia species key); Emery, 1891b: 5 (North Africa species key); Dalla Torre, 1893: 120, 122 (Macromischa, Leptothorax catalogues); Emery, 1895c: 317 (North America species key); Emery, 1896g: 58 (Neotropical species key); Bingham, 1903: 215 (India, Sri Lanka & Burma species key); Wheeler, W.M. 1903c: 223 (North America species key); Ruzsky, 1905b: 570 (Russian Empire species key); Wasmann, 1906: 16 (Luxemburg species key); Wheeler, W.M. 1908a: 141 (Macromischa species key); Santschi, 1909b: 460 (L. rottenbergi group key); Bondroit, 1910: 496 (Belgium species key); Stitz, 1914: 60 (Central Europe species key); Crawley, 1914: 91 (Britain species key); Donisthorpe, 1915d: 147 (Britain species key); Emery, 1916b: 176 (Italy species key); Wheeler, W.M. 1916m: 581 (U.S.A., Connecticut species key); Arnold, 1916: 257 (diagnosis, South Africa species key); Bondroit, 1918: 117 (France & Belgium species key); Mann, 1920: 408 (Macromischa species key); Wheeler, W.M. 1922a: 677 (Macromischa subgenera key); Wheeler, W.M. 1922a: 679 (Leptothorax subgenera key); Wheeler, W.M. 1922a: 890 (Afrotropical catalogue); Wheeler, W.M. 1922a: 1029 (Malagasy catalogue); Emery, 1924d: 245 (Macromischa catalogue); Emery, 1924d: 247 (diagnosis, subgenera key, catalogue); Emery, 1924d: 249 (L. (Goniothorax) diagnosis, catalogue); Emery, 1924d: 251 (L. (Leptothorax) diagnosis, catalogue); Emery, 1924d: 259 (L. (Temnothorax) diagnosis, catalogue); Emery, 1924d: 260 (L. (Dichothorax) & L. (Mychothorax) diagnoses, catalogues); Emery, 1924d: 264 (Symmyrmica review, catalogue); Karavaiev, 1927c: 266 (Ukraine species key); Donisthorpe, 1927b: 163 (Britain species key); Kuznetsov-Ugamsky, 1927d: 38 (Turkestan species key); Wheeler, W.M. 1931b: 31 (Macromischa, Croesomyrmex, Antillaemyrmex checklists); Arnol'di, 1933b: 598 (Russia species key); Menozzi, 1933b: 68 (Israel species key); Karavaiev, 1934: 134 (Ukraine species key); Wheeler, W.M. 1937b: 463 (Macromischa checklist); Menozzi, 1939a: 307 (Himalaya & Tibet species key); Stitz, 1939: 158 (Germany species key); Smith, M.R. 1939e: 503 (U.S.A. Macromischa species key); Kratochvíl, 1941: 89 (Central Europe species key); Novák & Sadil, 1941: 89 (Central Europe species key); Cole, 1942: 369 (U.S.A., Utah species key); Holgersen, 1943b: 172 (Norway species key); Holgersen, 1944: 198 (Norway species key); Kratochvíl, Novák & Snoflák, 1944: 123 (Czechoslovakia males, key); Buren, 1944a: 286 (U.S.A., Iowa species key); Boven, 1947: 178 (Belgium species key); Creighton, 1950a: 251, 256 (North America Macromischa, Leptothorax species keys); Chapman & Capco, 1951: 110 (Asia checklist); Smith, M.R. 1952b: 97 (U.S.A. L. tricarinatus complex, key); Bernard, 1956a: 151 (Western Europe species groups); Kusnezov, 1958c: 266 (subgenera key); Boven, 1959: 8 (Netherlands species key); Gregg, 1963: 380 (U.S.A., Colorado species key); Wheeler, G.C. & Wheeler, J. 1963: 139 (U.S.A., North Dakota species key); Collingwood, 1964: 99 (Britain species key); Bernard, 1967: 185 (diagnosis, Western Europe species key); Bernard, 1967: 187 (diagnosis, Western Europe species key); Bernard, 1967: 224 (Doronomyrmex diagnosis); Boven, 1970b: 20 (Netherlands species key); Arnol'di, 1971: 1824 (Kazakhstan species key); Kempf, 1972a: 132, 135 (Neotropical Leptothorax, Macromischa catalogues); Alayo, 1974: 15 (Macromischa Cuba species key); Bolton & Collingwood, 1975: 5 (Britain species key); Tarbinsky, 1976: 83 (Kyrghyzstan species key); Boven, 1977: 94 (Belgium species key); Kutter, 1977c: 103 (Switzerland species key); Arnol'di & Dlussky, 1978: 540 (former European U.S.S.R. species key); Collingwood, 1978: 83 (Iberian Peninsula species key); Collingwood, 1979: 68 (Fennoscandia & Denmark species key); Baroni Urbani, 1978b: 533 (L. (Macromischa) all species revision, key); Smith, D.R. 1979: 1391 (North America catalogue); Buschinger, 1981: 211 (Doronomyrmex review); Allred, 1982: 440 (U.S.A., Utah species key); Bolton, 1982: 319 (diagnosis, review of genus, Afrotropical species key); Gösswald, 1985: 303 (Germany species key); Wheeler, G.C. & Wheeler, J. 1986g: 51 (U.S.A., Nevada species key); Nilsson & Douwes, 1987: 60 (Norway species key); Agosti & Collingwood, 1987b: 273 (Balkans species key); Dlussky & Soyunov, 1988: 29 (former U.S.S.R. Temnothorax species); Hölldobler & Wilson, 1990: 13 (synoptic classification); Dlussky, Soyunov & Zabelin, 1990: 184, 188 (Turkmenistan species key); Kupyanskaya, 1990: 136 (Far Eastern Russia species key); Brandão, 1991: 349 (Neotropical catalogue); Morisita, Kubota, et al. 1992: 26 (Japan species key); Atanasov & Dlussky, 1992: 129 (Bulgaria species key); Arakelian, 1994: 52 (Armenia species key); Radchenko, 1994b: 111 (South Siberia species key); Radchenko, 1994d: 146 (Central & Eastern Palaearctic species key); Bolton, 1995a: 1050 (census); Bolton, 1995b: 235 (catalogue); Douwes, 1995: 89 (Sweden species key); Kupyanskaya, 1995: 347 (Far Eastern Russia species key); Wu, J. & Wang, 1995: 108 (China species key); Collingwood & Agosti, 1996: 325 (Saudi Arabia species key); Seifert, 1996b: 13 (Central Europe Doronomyrmex species key); Seifert, 1996b: 119 (Central Europe species key); Skinner & Allen, 1996: 45 (Britain species key); Cagniant & Espadaler, 1997: 260 (Morocco species key); Collingwood & Prince, 1998: 16 (Portugal species key); Terayama & Onoyama, 1999: 73 (Japan species key); Mackay, 2000: 265, 288, 297 (New World L. (Myrafant) species revision key, Mexico & Guatemala species key, species complexes key); Fontenla Rizo, 2001: 15 (Cuba Macromischa species key); Chang & He, 2001b: 1 (northwest China species key); Czechowski, Radchenko & Czechowska, 2002: 143 (Poland species key); Mackay & Mackay, 2002: 93 (U.S.A., New Mexico species key); Bolton, 2003: 270 (genus diagnosis); Lyu & Cho, 2003b: 269 (Korea species key); Radchenko, 2004: 131 (Eastern Palaearctic species, key); Deyrup & Cover, 2004a: 56 (southeast U.S.A. species key); Radchenko, 2005b: 192 (North Korea species key); Seifert, 2007: 118 (North and Central Europe species key).

Myrmicinae Formicoxenini Myrmoxenus TRUE TRUE FALSE

12 species

Myrmoxenus Ruzsky, 1902b: 474. Type-species: Myrmoxenus gordiagini, by monotypy.

Taxonomic history

Myrmoxenus in Myrmicinae, Myrmicini: Wheeler, W.M. 1910g: 139.

Myrmoxenus in Myrmicinae, Leptothoracini: Emery, 1914a: 42; Forel, 1917: 245; Wheeler, W.M. 1922a: 664; Emery, 1924d: 262; all subsequent authors to the following.

Myrmoxenus in Myrmicinae, Formicoxenini: Bolton, 1994: 105.

Myrmoxenus as junior synonym of Epimyrma: Buschinger, 1990: 245 (provisional synoynym); Bolton, 1994: 105 [seniority incorrect, Myrmoxenus has priority; see below.]

Junior synonyms of MYRMOXENUS

Epimyrma Emery, 1915a: 262. Type-species: Epimyrma kraussei, by original designation.

Taxonomic history

Epimyrma in Myrmicinae, Leptothoracini: Forel, 1917: 245; Wheeler, W.M. 1922a: 664; Emery, 1924d: 263; all subsequent authors to the following.

Epimyrma in Myrmicinae, Formicoxenini: Bolton, 1994: 105.

Epimyrma as junior synonym of Leptothorax: Brown, 1973b: 180 [provisional].

Epimyrma as junior synonym of Myrmoxenus: Schulz & Sanetra, 2002: 162.

Myrmetaerus Soudek, 1925a: 33. Type-species: Myrmetaerus microcellatus (junior synonym of Myrmoxenus gordiagini), by monotypy.

Taxonomic history

[Myrmetaerus also described as new by Soudek, 1925b: 14.]

Myrmetaerus in Myrmicinae, Leptothoracini: Donisthorpe, 1943f: 667.

Myrmetaerus as junior synonym of Leptothorax: Brown, 1973b: 182 [provisional].

Myrmetaerus as junior synonym of Camponotus: Snelling, R.R. 1981: 404 (error).

Myrmetaerus as junior synonym of Myrmoxenus: Buschinger, Winter & Faber, 1984: 336.

Subgenera of MYRMOXENUS include the nominal plus the following.

Subgenus MYRMOXENUS (GONEPIMYRMA)

Gonepimyrma Bernard, 1948: 146 [as subgenus of Epimyrma]. Type-species: Epimyrma (Gonepimyrma) africana, by monotypy.

Taxonomic history

Gonepimyrma junior synonym of Epimyrma: Brown, 1973b: 181 [provisional].

Genus Myrmoxenus references

Emery, 1916b: 189 (Italy species key); Bondroit, 1918: 139 (France & Belgium species key); Emery, 1924d: 262 (Myrmoxenus diagnosis, catalogue); Emery, 1924d: 263 (Epimyma diagnosis, catalogue); Menozzi, 1931c: 36 (diagnosis, all species key); Bernard, 1967: 216 (diagnosis, Western Europe species key); Kutter, 1973e: 281 (species, notes); Kutter, 1977c: 138 (Switzerland species key); Buschinger, Fischer, et al. 1987: 253 (partial revision); Agosti & Collingwood, 1987b: 276 (Balkans species key); Buschinger, 1989: 265 (evolution, speciation); Bolton, 1995a: 1049 (census); Bolton, 1995b: 188 (catalogue); Seifert, 1996b: 118 (Central Europe species key); Cagniant & Espadaler, 1997: 268 (Morocco species key); Seifert, 2007: 117 (North and Central Europe species key).

Myrmicinae Formicoxenini Nesomyrmex TRUE TRUE FALSE

Extant: 51 species, 1 subspecies

Fossil: 2 species

Nesomyrmex Wheeler, W.M. 1910a: 259. Type-species: Nesomyrmex clavipilis, by monotypy.

Taxonomic history

[Nesomyrmex is the oldest synonym of Goniothorax Emery (a junior homonym, see below) and hence is the first available replacement name for Goniothorax Emery: Smith, M.R. 1950: 30.]

Nesomyrmex in Myrmicinae, Leptothoracini: Forel, 1917: 244; subsequent authors to the following.

Nesomyrmex in Myrmicinae, Formicoxenini: Bolton, 1994: 105.

Nesomyrmex as subgenus of Leptothorax: Emery, 1915g: 24; Forel, 1917: 244; Wheeler, W.M. 1922a: 679; Smith, M.R. 1950: 30; Smith, M.R. 1951a: 816; Kempf, 1959c: 393; Brown, 1971a: 4; Kempf, 1972a: 133; Smith, D.R. 1979: 1391; Mackay, 2000: 272 (in key).

Nesomyrmex as junior synonym of Goniothorax: Emery, 1924d: 249; Donisthorpe, 1943f: 674. [Goniothorax is a junior homonym and hence Nesomyrmex takes priority as first available replacement name.]

Nesomyrmex as junior synonym of Leptothorax: Brown, 1973b: 183 [provisional]; Bolton, 1982: 319; Bolton, 1994: 105; Bolton, 1995b: 39.

Nesomyrmex as genus: Wheeler, W.M. 1910a: 259; Francoeur & Loiselle, 1988b: 43 (possibly valid); Hölldobler & Wilson, 1990: 14 [junior synonyms of Nesomyrmex left behind under Leptothorax]; Jaffe, 1993: 11; Bolton, 2003: 70.

Junior synonyms of NESOMYRMEX

Tetramyrma Forel, 1912m: 766 [as subgenus of Dilobocondyla]. Type-species: Dilobocondyla (Tetramyrma) braunsi, by monotypy.

Taxonomic history

Tetramyrma in Myrmicinae, Tetramoriini: Emery, 1914a: 42; Forel, 1917: 245; Arnold, 1917: 358; Wheeler, W.M. 1922a: 664; Emery, 1924d: 291; Donisthorpe, 1943g: 732; Wheeler, G.C. & Wheeler, J. 1985: 257 (anachronism); Dlussky & Fedoseeva, 1988: 80 (anachronism).

Tetramyrma incertae sedis in Myrmicinae: Bolton, 1976: 291.

Tetramyrma as subgenus of Dilobocondyla: Forel, 1912m: 766.

Tetramyrma as genus: Forel, 1913a: 122; Arnold, 1917: 358; Wheeler, W.M. 1922a: 664; Emery, 1924d: 291; Donisthorpe, 1943g: 732; Bolton, 1976: 291.

Tetramyrma as junior synonym of Leptothorax: Bolton, 1982: 319; Bolton, 1994: 105.

Tetramyrma as junior synonym of Nesomyrmex: Bolton, 2003: 249, 272.

Caulomyrma Forel, 1914d: 233 [as subgenus of Leptothorax]. Type-species: Leptothorax echinatinodis, by original designation.

Taxonomic history

Caulomyrma as junior synonym of Nesomyrmex: Forel, 1915c: 364 [provisional]; Emery, 1915g: 24; Wheeler, W.M. 1922a: 679.

Caulomyrma as subgenus of Leptothorax: Forel, 1914d: 233; Forel, 1917: 244.

Caulomyrma as junior synonym of Goniothorax: Emery, 1924d: 249 (incorrect procedure as Goniothorax is a junior homonym for which Nesomyrmex is the first available replacement name; see Smith, M.R. 1950: 30).]

Caulomyrma as junior synonym of Leptothorax: Brown, 1973b: 182 [provisional].

Limnomyrmex Arnold, 1948: 222. Type-species: Limnomyrmex stramineus, by original designation.

Taxonomic history

Limnomyrmex as junior synonym of Nesomyrmex: Brown, 1971a: 4.

Ireneopone Donisthorpe, 1946d: 242. Type-species: Ireneopone gibber, by original designation.

Taxonomic history

Ireneopone in Myrmicinae, Tetramoriini: Donisthorpe, 1946d: 243; Wheeler, G.C. & Wheeler, J. 1985: 257 (anachronism).

Ireneopone incertae sedis in Myrmicinae: Bolton, 1976: 292; Dlussky & Fedoseeva, 1988: 81.

Ireneopone in Myrmicinae, Formicoxenini: Bolton, 1994: 105; Bolton, 1995a: 1050; Bolton, 1995b: 217.

Ireneopone as junior synonym of Nesomyrmex: Bolton, 2003: 250, 272.

Meia Pagliano & Scaramozzino, 1990: 5.

Taxonomic history

[Unnecessary replacement name for Goniothorax Emery, 1896g: 58 (junior homonym of Goniothorax Milne-Edwards, 1879: 103). Nesomyrmex is the first available replacement name for Goniothorax Emery, therefore Meia is automatically a junior synonym of Nesomyrmex: Bolton, 1994: 105.]

Genus Nesomyrmex references

Emery, 1896g: 58 (Neotropical species key); Arnold, 1916: 258 (South Africa species key); Arnold, 1917: 358 (Tetramyrma diagnosis); Wheeler, W.M. 1922a: 890, 908 (Afrotropical L. (Goniothorax), Tetramyrma catalogues); Wheeler, W.M. 1922a: 1029 (Malagasy L. (Goniothorax) catalogue); Emery, 1924d: 249 (L. (Goniothorax) diagnosis, catalogue); Emery, 1924d: 291 (Tetramyrma review, catalogue); Kempf, 1959c: 394 (Neotropical Leptothorax (Nesomyrmex) species revision, key); Kempf, 1972a: 133 (Neotropical Leptothorax (Nesomyrmex) catalogue); Kempf, 1975c: 367 (additions to 1959b key); Bolton, 1982: 319 (diagnosis, review of genus, Afrotropical species key); Hölldobler & Wilson, 1990: 13 (synoptic classification); Brandão, 1991: 349 (Neotropical catalogue); Bolton, 1995b: 235 (catalogue); Bolton, 2003: 272 (genus diagnosis); Mbanyana & Robertson, 2008: 36 (southern Africa species revision, key).

Homonym replaced by NESOMYRMEX

Goniothorax Emery, 1896g: 58 (diagnosis in key) [as subgenus of Leptothorax]. Type-species: Leptothorax vicinus, by subsequent designation of Wheeler, W.M. 1911f: 164.

Taxonomic history

[Junior homonym of Goniothorax Milne-Edwards, 1879: 103 (Crustacea).]

Goniothorax in Myrmicinae, Leptothoracini: Forel, 1917: 244.

Goniothorax as subgenus of Leptothorax: Wheeler, W.M. 1910g: 139; Forel, 1917: 244; Wheeler, W.M. 1922a: 679, 890, 1029; Emery, 1924d: 249; Creighton, 1950a: 259.

[Goniomyrmex Ashmead, 1905b: 383, incorrect subsequent spelling.]

Myrmicinae Myrmecinini Pristomyrmex TRUE TRUE FALSE

57 species

Pristomyrmex Mayr, 1866b: 903. Type-species: Pristomyrmex pungens (junior synonym of Myrmica punctata) by monotypy.

Taxonomic history

Pristomyrmex in Myrmicidae: Mayr, 1866b: 903.

Pristomyrmex in Myrmicidae, Myrmicidae: Emery, 1877a: 81.

Pristomyrmex in Myrmicinae: Dalla Torre, 1893: 62.

Pristomyrmex in Myrmicinae, Myrmicini: Emery, 1895j: 769; Ashmead, 1905b: 383; Wheeler, W.M. 1910g: 139.

Pristomyrmex in Myrmicinae, Myrmecinini: Emery, 1912b: 105; Emery, 1914a: 41; Forel, 1917: 244; Wheeler, W.M. 1922a: 663; Emery, 1924d: 233; all subsequent authors except the entries below; Bolton, 1994: 105; Bolton, 2003: 71.

Pristomyrmex in Myrmicinae, Tetramoriini: Bernard, 1953b: 251.

Pristomyrmex in Myrmicinae, Pheidolini: Hölldobler & Wilson, 1990: 16.

Junior synonyms of PRISTOMYRMEX

Odontomyrmex André, 1905: 207. Type-species: Odontomyrmex quadridentatus, by monotypy.

Taxonomic history

Odontomyrmex in Myrmicinae, Myrmicini: Wheeler, W.M. 1910g: 139.

Odontomyrmex as subgenus of Pristomyrmex: Forel, 1917: 244; Wheeler, W.M. 1922a: 682; Chapman & Capco, 1951: 121 (anachronism).

Odontomyrmex as junior synonym of Pristomyrmex: Mann, 1919: 341; Emery, 1924d: 233; Donisthorpe, 1943f: 676; Brown, 1953c: 9; Taylor, 1965b: 35; Bolton, 1981b: 282.

Hylidris Weber, 1941a: 190. Type-species: Hylidris myersi (junior synonym of Pristomyrmex africanus), by original designation.

Taxonomic history

Hylidris in Myrmicinae, Myrmecinini: Weber, 1941a: 183; Donisthorpe, 1943f: 651.

Hylidris as junior synonym of Pristomyrmex: Brown, 1953c: 9.

Dodous Donisthorpe, 1946f: 145. Type-species: Dodous trispinosus, by original designation.

Taxonomic history

Dodous in Myrmicinae, Solenopsidini: Donisthorpe, 1946f: 145.

Dodous as junior synonym of Pristomyrmex: Brown, 1971a: 3.

Genus Pristomyrmex references

Dalla Torre, 1893: 62 (catalogue); Forel, 1903a: 696 (India & Sri Lanka species key); Bingham, 1903: 193 (diagnosis); Emery, 1924d: 233 (diagnosis, catalogue); Chapman & Capco, 1951: 121, 161 (Asia Pristomyrmex, Dodous checklists); Taylor, 1965b: 37 (Australia species revision, key); Taylor, 1968c: 63 (supplement to previous); Bolton, 1981b: 282 (diagnosis, Afrotropical species revision, key); Taylor & Brown, D.R. 1985: 85 (Australia catalogue); Taylor, 1987a: 64 (Australia checklist); Morisita, Kubota, et al. 1992: 57 (Japan species key); Bolton, 1994: 105 (synoptic classification); Bolton, 1995a: 1052 (census); Bolton, 1995b: 365 (catalogue); Lin & Wu, 1998: 94 (Taiwan species key); Shattuck, 1999: 159 (Australia synopsis); Xu & Zhang, 2002: 69 (China species key); Wang, M. 2003: 393 (all species revision, key); Zettell, 2006: 65 (Philippines species, key); Terayama, 2009: 190 (Taiwan species key).

Genera incertae sedis in Myrmecinini

Ponerinae Ponerini Anochetus TRUE TRUE FALSE

Extant: 94 species

Fossil: 8 species

Anochetus Mayr, 1861: 53. Type-species: Odontomachus ghilianii, by monotypy.

Taxonomic history

Anochetus in Formicidae, Odontomachidae: Mayr, 1862: 712; Mayr, 1865: 11.

Anochetus in Odontomachidae: Ashmead, 1905b: 382.

Anochetus in Ponerinae: Mayr, 1861: 53 [Poneridae]; Emery & Forel, 1879a: 455 [Poneridae]; Dalla Torre, 1893: 47.

Anochetus in Ponerinae, Odontomachini: Forel, 1893a: 163; Emery, 1895j: 768; Forel, 1899c: 18; Wheeler, W.M. 1910g: 136; Emery, 1911d: 106; Arnold, 1915: 103; Forel, 1917: 238; Gallardo, 1918b: 89; Wheeler, W.M. 1922a: 653; Donisthorpe, 1943f: 623; Chapman & Capco, 1951: 31; Wilson, 1959a: 483; Wheeler, G.C. & Wheeler, J. 1985: 256; Dlussky & Fedoseeva, 1988: 78; Hölldobler & Wilson, 1990: 11; Jaffe, 1993: 8.

Anochetus in Ponerinae, Ponerini: Brown, 1976a: 71; Brown, 1978c: 550 [subtribe Odontomachiti]; Bolton, 1994: 164; Bolton, 2003: 159.

Junior synonyms of ANOCHETUS

Stenomyrmex Mayr, 1862: 711. Type-species: Myrmecia emarginata, by subsequent designation of Wheeler, W.M. 1911f: 173.

Taxonomic history

Stenomyrmex in Formicidae, Odontomachidae: Mayr, 1862: 711; Mayr, 1865: 11.

Stenomyrmex in Odontomachidae: Ashmead, 1905b: 382.

Stenomyrmex as subgenus of Anochetus: Dalla Torre, 1893: 47; Forel, 1893a: 163; Dalla Torre, 1893: 47; Wheeler, W.M. 1910g: 136; Emery, 1911d: 110; Forel, 1917: 238; Wheeler, W.M. 1922a: 653; Borgmeier, 1923: 76; Donisthorpe, 1943g: 728; Kempf, 1972a: 21.

Stenomyrmex as junior synonym of Anochetus: Forel, 1887: 382; Brown, 1978c: 552; all subsequent authors.

Myrmapatetes Wheeler, W.M. 1929b: 6. Type-species: Myrmapatetes filicornis, by original designation.

Taxonomic history

Myrmapatetes in Dolichoderinae: Wheeler, W.M. 1929b: 7; Chapman & Capco, 1951: 186.

Myrmapatetes as junior synonym of Anochetus: Brown, 1953h: 2.

Genus Anochetus references

Roger, 1863b: 21, 22 (catalogue); Mayr, 1863: 394, 454 (catalogue); Mayr, 1865: 11 (Anochetus, Stenomyrmex diagnoses); André, 1882b: 230 (Europe & Algeria species); Emery, 1884a: 378 (all species key); Dalla Torre, 1893: 47 (catalogue); Emery, 1894c: 185 (Neotropical species key); Forel, 1900c: 58 (India & Sri Lanka species key); Bingham, 1903: 39 (India, Sri Lanka & Burma species key); Emery, 1911d: 107 (diagnosis, catalogue); Arnold, 1915: 103 (South Africa species key); Gallardo, 1918b: 90 (Argentina species key); Mann, 1919: 303 (New Guinea & Solomon Is species key); Wheeler, W.M. 1922a: 96, 790, 1012 (diagnosis, Afrotropical, Malagasy catalogues); Wheeler, W.M. 1925a: 9 (Stenomyrmex species key); Chapman & Capco, 1951: 39 (Asia checklist); Wilson, 1959a: 503 (Melanesia & Moluccas species, revision, key); Kempf, 1964f: 244 (A. (Stenomyrmex) species key); Kempf, 1972a: 20 (Neotropical catalogue); Brown, 1978c: 552, 565, 569, 571 (diagnosis & review of genus, Malesian species, Austral, Palaearctic, Afrotropical & Malagasy species, Neotropical species keys); Taylor & Brown, D.R. 1985: 20 (Australia catalogue); Taylor, 1987a: 7 (Australia checklist); Terayama, 1989a: 26 (Taiwan species key); Brandão, 1991: 324 (Neotropical catalogue); Wang, M. 1993a: 226 (China species key); De Andrade, 1994: 24 (†Dominican Amber species key); Bolton, 1995a: 1048 (census); Bolton, 1995b: 63 (catalogue); Gronenberg & Ehmer, 1996: 183 (mandible morphology); Collingwood & Agosti, 1996: 309 (Saudi Arabia species key); Shattuck, 1999: 181 (Australia synopsis); Zhou, 2001b: 29 (China, Guangxi species key); Kugler, J. & Ionescu, 2007: 298 (Mediterraean & Middle East species key); Fisher & Smith, 2008: 4 (Malagasy species revision, key); González-Campero & Elizalde, 2008: 98 (Argentina & Paraguay species key); Zabala, in Fernández, 2008: 125 (Neotropical species key); Terayama, 2009: 117 (Taiwan species key).

Ponerinae Ponerini Leptogenys TRUE TRUE FALSE

Extant: 216 species, 31 subspecies

Fossil: 1 species

Leptogenys Roger, 1861a: 41. Type-species: Leptogenys falcigera, by subsequent designation of Bingham, 1903: 52.

Taxonomic history

Leptogenys in Ponerinae: Dalla Torre, 1893: 46; Mayr, 1862: 714 (in key) [Poneridae]; Mayr, 1865: 14 [Poneridae].

Leptogenys in Ponerinae, Leptogenyini: Forel, 1893a: 162; Forel, 1899c: 17; Ashmead, 1905b: 382; Emery, 1911d: 97; Arnold, 1915: 89; Forel, 1917: 238; Gallardo, 1918b: 84; Wheeler, W.M. 1922a: 652; Donisthorpe, 1943f: 656; Chapman & Capco, 1951: 31; Dlussky & Fedoseeva, 1988: 78.

Leptogenys in Ponerinae, Ponerini: Emery, 1895j: 767; Forel, 1900d: 304; Wheeler, W.M. 1910g: 135; Brown, 1963: 3; Kempf, 1972a: 129; Jaffe, 1993: 8; Bolton, 1994: 164; Bolton, 2003: 163.

Junior synonyms of LEPTOGENYS

Lobopelta Mayr, 1862: 714 (diagnosis in key), 733. Type-species: Ponera diminuta, by subsequent designation of Bingham, 1903: 54.

Taxonomic history

Lobopelta in Ponerinae: Mayr, 1862: 714 (in key) [Poneridae]; Mayr, 1865: 15 [Poneridae]; Dalla Torre, 1893: 43.

Lobopelta in Ponerinae, Leptogenyini: Forel, 1893a: 162; Ashmead, 1905b: 382; Emery, 1911d: 101; Wheeler, W.M. 1922a: 653; Donisthorpe, 1943f: 658.

Lobopelta as genus: Mayr, 1862: 733; Cresson, 1887: 258; Dalla Torre, 1893: 43; Bingham, 1903: 54.

Lobopelta as subgenus of Leptogenys: Forel, 1892k: 520; Forel, 1899c: 18; Forel, 1900d: 304; Wheeler, W.M. 1910g: 135; Emery, 1911d: 101; Arnold, 1915: 95; Forel, 1917: 238; Gallardo, 1918b: 85; Wheeler, W.M. 1922a: 653; Wheeler, W.M. 1923d: 1; Borgmeier, 1923: 75; Creighton, 1950a: 50; Kempf, 1972a: 131; Smith, D.R. 1979: 1343 (anachronism).

Lobopelta as junior synonym of Leptogenys: Emery, 1896e: 177 (footnote); Brown, 1973b: 181 [provisional]; Bolton, 1975a: 240; Bolton, 2003: 163.

Prionogenys Emery, 1895g: 348. Type-species: Prionogenys podenzanai, by monotypy.

Taxonomic history

Prionogenys in Ponerinae, Ponerini: Emery, 1895j: 767; Wheeler, W.M. 1910g: 135;

Prionogenys in Ponerinae, Leptogenyini: Ashmead, 1905b: 382; Emery, 1911d: 106; Forel, 1917: 238; Wheeler, W.M. 1922a: 652; Donisthorpe, 1943f: 685; Dlussky & Fedoseeva, 1988: 78.

Prionogenys as junior synonym of Leptogenys: Taylor, 1988: 33.

Machaerogenys Emery, 1911d: 100 [as subgenus of Leptogenys]. Type-species: Leptogenys truncatirostris, by original designation.

Taxonomic history

Machaerogenys as subgenus of Leptogenys: Emery, 1911d: 100; Forel, 1917: 238; Wheeler, W.M. 1922a: 653; subsequent authors to the following.

Machaerogenys as junior synonym of Leptogenys: Brown, 1973b: 181 [provisional]; Bolton, 1975a: 240.

Odontopelta Emery, 1911d: 101 [as subgenus of Leptogenys]. Type-species: Leptogenys turneri, by monotypy.

Taxonomic history

Odontopelta as subgenus of Leptogenys: Emery, 1911d: 101; Forel, 1917: 238; Wheeler, W.M. 1922a: 653; Donisthorpe, 1943f: 677.

Odontopelta as junior synonym of Leptogenys: Brown, 1973b: 183 [provisional]; Snelling, R.R. 1981: 390; Taylor & Brown, D.R. 1985: 32.

Dorylozelus Forel, 1915b: 24. Type-species: Dorylozelus mjobergi (junior secondary homonym in Leptogenys, replaced by Leptogenys tricosa), by monotypy.

Taxonomic history

Dorylozelus in Ponerinae, Ponerini: Forel, 1917: 238.

Dorylozelus in Ponerinae, Dorylozelini: Wheeler, W.M. 1922a: 646; Donisthorpe, 1943f: 640.

Dorylozelus incertae sedis in Ponerinae, Amblyoponini: Brown, 1960a: 181.

Dorylozelus as junior synonym of Leptogenys: Taylor, 1969: 132; Bolton, 1975a: 240.

Microbolbos Donisthorpe, 1948f: 170. Type-species: Microbolbos testaceus, by original designation.

Taxonomic history

Microbolbos as junior synonym of Leptogenys: Wilson, 1955c: 136; Bolton, 1975a: 240.

Genus Leptogenys references

Roger, 1863b: 19 (Lobopelta, Leptogenys catalogues); Mayr, 1863: 426, 427 (Leptogenys, Lobopelta catalogues); Mayr, 1865: 14, 15 (Leptogenys, Lobopelta diagnoses); Mayr, 1867a: 89 (diagnosis); Mayr, 1879: 664 (Lobopelta species key); Cresson, 1887: 258 (U.S.A. catalogue); Dalla Torre, 1893: 43, 46 (Lobopelta, Leptogenys catalogues); Forel, 1900d: 304 (India & Sri Lanka species key); Bingham, 1903: 52, 55 (India, Sri Lanka & Burma Leptogenys, Lobopelta species keys); Emery, 1911d: 97 (diagnosis, subgenera key, catalogue); Arnold, 1915: 89, 95 (diagnosis, South Africa L. (Leptogenys) & L. (Lobopelta) species keys); Mann, 1919: 298 (Papuasia L. (Leptogenys) species key); Mann, 1921: 426 (Fiji L. (Lobopelta) species key); Wheeler, W.M. 1922a: 93, 786, 1010 (diagnosis, Afrotropical, Malagasy catalogues); Wheeler, W.M. 1923d: 14 (New World L. (Lobopelta) species key); Creighton, 1950a: 51 (North America L. (Lobopelta) species key); Chapman & Capco, 1951: 31 (Asia checklist); Wilson, 1958a: 113 (Melanesia & Moluccas species revision, key); Wilson, 1958a: 136 (New Caledonia species key); Kempf, 1972a: 129 (Neotropical catalogue); Alayo, 1974: 8 (Cuba species key); Bolton, 1975a: 239 (diagnosis, review of genus); Bolton, 1975a: 244 (Afrotropical species revision, key); Bolton, 1975a: 293 (Malagasy species key); Smith, D.R. 1979: 1343 (North America catalogue); Taylor & Brown, D.R. 1985: 32, 39 (Australia catalogue); Taylor, 1987a: 34, 64 (Australia, New Caledonia checklists); Trager & Johnson, 1988: 30 (U.S.A. species key); Brandão, 1991: 349 (Neotropical catalogue); Bolton, 1995a: 1050 (census); Bolton, 1995b: 229 (catalogue); Wu, J. & Wang, 1995: 41 (China species key); Shattuck, 1999: 189 (Australia synopsis); Xu, 2000a: 118 (China species key); Zhou, 2001b: 39 (China, Guangxi species key); Bolton, 2003: 163 (synopsis); Lattke, in Fernández, 2008: 145 (Colombia species, review); Terayama, 2009: 115 (Taiwan species key); Heterick, 2009: 133 (south-western Australia species key).

Ponerinae Ponerini Pachycondyla TRUE TRUE FALSE

Extant: 252 species, 62 subspecies

Fossil: 12 species

Pachycondyla Smith, F. 1858b: 105. Type-species: Formica crassinoda, by subsequent designation of Emery, 1901a: 42.

Taxonomic history

Pachycondyla in Poneridae: Smith, F. 1858b: 105; Smith, 1871a: 324.

Pachycondyla in Ponerinae: Mayr, 1862: 713 [Poneridae]; Mayr, 1865: 13 [Poneridae]; Dalla Torre, 1893: 32.

Pachycondyla in Pachycondylinae, Pachycondylini: Ashmead, 1905b: 382.

Pachycondyla in Ponerinae, Ponerini: Emery, 1895j: 767; Forel, 1895b: 114; Forel, 1899c: 10; Wheeler, W.M. 1910g: 135; Emery, 1911d: 73 [subtribe Pachycondylini]; Arnold, 1915: 53; Forel, 1917: 237; Wheeler, W.M. 1922a: 648; all subsequent authors.

Junior synonyms of PACHYCONDYLA

Bothroponera Mayr, 1862: 713 (diagnosis in key), 717. Type-species: Ponera pumicosa, by subsequent designation of Emery, 1901a: 42.

Taxonomic history

Bothroponera in Ponerinae: Mayr, 1862: 713 (in key) [Poneridae]; Mayr, 1865: 13 [Poneridae]; Dalla Torre, 1893: 35.

Bothroponera in Pachycondylinae, Pachycondylini: Ashmead, 1905b: 382.

Bothroponera in Ponerinae, Ponerini: Emery, 1895j: 767; Emery, 1911d: 75 [subtribe Pachycondylini]; Wheeler, W.M. 1922a: 648; all subsequent authors.

Bothroponera as subgenus of Ponera: Emery, 1895j: 767; Forel, 1900d: 325.

Bothroponera as subgenus of Pachycondyla: Emery, 1901a: 42; Wheeler, W.M. 1910g: 135; Emery, 1911d: 75; Arnold, 1915: 55; Forel, 1917: 237; Donisthorpe, 1943f: 628.

Bothroponera as genus: Mayr, 1862: 717; Dalla Torre, 1893: 35; Bingham, 1903: 95; Ashmead, 1905b: 382; Wheeler, W.M. 1918c: 299 (footnote); Wheeler, W.M. 1922a: 648; Wheeler, W.M. & Chapman, 1925: 67; Chapman & Capco, 1951: 49; Bernard, 1953b: 187; Taylor, 1987a: 9; Taylor & Brown, D.R. 1985: 21; Wheeler, G.C. & Wheeler, J. 1985: 256; Dlussky & Fedoseeva, 1988: 78.

Bothroponera as junior synonym of Pachycondyla: Brown, 1973b: 179 [provisional]; Snelling, R.R. 1981: 389; Hölldobler & Wilson, 1990: 11; Brown, in Bolton, 1994: 164; Bolton, 2003: 166.

Megaponera Mayr, 1862: 714 (diagnosis in key), 734. Type-species: Formica foetens (junior primary homonym in Formica, replaced by Formica analis), by monotypy.

Taxonomic history

Megaponera in Ponerinae: Dalla Torre, 1893: 30; Mayr, 1862: 714 [Poneridae]; Mayr, 1865: 15 [Poneridae].

Megaponera in Pachycondylinae, Ectatommini: Ashmead, 1905b: 382.

Megaponera in Ponerinae, Ponerini: Emery, 1895j: 767; Wheeler, W.M. 1910g: 135; Emery, 1911d: 67 [subtribe Pachycondylini]; Arnold, 1915: 46; Wheeler, W.M. 1922a: 647; all subsequent authors to the following.

Megaponera as junior synonym of Pachycondyla: Brown, in Bolton, 1994: 164; Bolton, 2003: 166.

[Megaloponera Roger, 1863b: 17, incorrect subsequent spelling; misspelling repeated by several authors, for example Emery, 1877b: 368; Forel, 1917: 237.]

Paltothyreus Mayr, 1862: 714 (diagnosis in key), 735. Type-species: Formica tarsata, by monotypy.

Taxonomic history

Paltothyreus in Ponerinae: Mayr, 1862: 714 [Poneridae]; Mayr, 1865: 15 [Poneridae]; Dalla Torre, 1893: 32.

Paltothyreus in Pachycondylinae, Ectatommini: Ashmead, 1905b: 382.

Paltothyreus in Ponerinae, Ponerini: Emery, 1895j: 767; Wheeler, W.M. 1910g: 135; Emery, 1911d: 62 [subtribe Pachycondylini]; Arnold, 1915: 43; Forel, 1917: 237; Wheeler, W.M. 1922a: 647; all subsequent authors.

Paltothyreus as junior synonym of Pachycondyla: Brown, in Bolton, 1994: 164; Bolton, 2003: 166.

Ectomomyrmex Mayr, 1867a: 83. Type-species: Ectomomyrmex javanus, by subsequent designation of Bingham, 1903: 85.

Taxonomic history

Ectomomyrmex in Ponerinae: Dalla Torre, 1893: 30.

Ectomomyrmex in Pachycondylinae, Pachycondylini: Ashmead, 1905b: 382.

Ectomomyrmex in Ponerinae, Ponerini: Emery, 1895j: 767; Forel, 1900d: 321; Emery, 1911d: 78 [subtribe Pachycondylini]; Wheeler, W.M. 1922a: 648; all subsequent authors.

Ectomomyrmex as subgenus of Pachycondyla: Emery, 1901a: 42; Wheeler, W.M. 1910g: 135; Emery, 1911d: 78; Arnold, 1915: 53; Forel, 1917: 237.

Ectomomyrmex as genus: Mayr, 1867a: 83; Dalla Torre, 1893: 30; Ashmead, 1905b: 382; Forel, 1900d: 321; Bingham, 1903: 85; Wheeler, W.M. 1922a: 648; Chapman & Capco, 1951: 60; Brown, 1963: 9; Taylor & Brown, D.R. 1985: 29; Taylor, 1987a: 26; Ogata, 1987: 112.

Ectomomyrmex as junior synonym of Pachycondyla: Brown, 1973b: 180 [provisional]; Snelling, R.R. 1981: 389; Hölldobler & Wilson, 1990: 11; Brown, in Bolton, 1994: 164; Bolton, 2003: 166.

[Ectomyrmex Donisthorpe, 1943f: 641, incorrect subsequent spelling.]

Ophthalmopone Forel, 1890b: cxii. Type-species: Ophthalmopone berthoudi, by monotypy.

Taxonomic history

Ophthalmopone in Ponerinae: Dalla Torre, 1893: 31.

Ophthalmopone in Pachycondylinae, Pachycondylini: Ashmead, 1905b: 382.

Ophthalmopone in Ponerinae, Ponerini: Emery, 1895j: 767; Wheeler, W.M. 1910g: 135; Emery, 1911d: 69 [subtribe Pachycondylini]; Arnold, 1915: 49; Forel, 1917: 237; Wheeler, W.M. 1922a: 647; all subsequent authors.

Ophthalmopone as junior synonym of Pachycondyla: Brown, in Bolton, 1994: 164; Bolton, 2003: 166.

Euponera Forel, 1891b: 126 [as subgenus of Ponera]. Type-species: Ponera (Euponera) sikorae, by monotypy.

Taxonomic history

Euponera in Ponerinae, Ponerini: Emery, 1895j: 767; Emery, 1911d: 79 [subtribe Pachycondylini]; Wheeler, W.M. 1915h: 37; Arnold, 1915: 63; Forel, 1917: 237; Wheeler, W.M. 1922a: 648; all subsequent authors.

Euponera as subgenus of Ponera: Forel, 1891b: 126; Emery, 1895j: 767.

Euponera as genus: Emery, 1901a: 46; Emery, 1909c: 364; Wheeler, W.M. 1910g: 135; Emery, 1911d: 79; Arnold, 1915: 63; Forel, 1917: 237; Gallardo, 1918b: 64; Wheeler, W.M. 1922a: 648; Borgmeier, 1923: 70; Clark, 1934b: 30; Creighton, 1950a: 44; Chapman & Capco, 1951: 63; Bernard, 1953b: 189; Wheeler, G.C. & Wheeler, J. 1985: 256; Dlussky & Fedoseeva, 1988: 78; Hölldobler & Wilson, 1990: 11.

Euponera as junior synonym of Pachycondyla: Brown, in Bolton, 1994: 164; Bolton, 2003: 166.

Cryptopone Emery, 1893a: cclxxv. Type-species: Cryptopone testacea, by monotypy.

Taxonomic history

[Cryptopone also described as new by Emery, 1893f: 240. Type-species not Amblyopone testacea, unjustified subsequent designation by Wheeler, W.M. 1911f: 161, repeated in Emery, 1911d: 88, Wheeler, W.M. 1922a: 780, Donisthorpe, 1943f: 636, Kempf, 1972a: 90 and Taylor & Brown, D.R. 1985: 28; see discussion in Wilson, 1958d: 360.]

Cryptopone in Ponerinae, Ponerini: Emery, 1895j: 767; Ashmead, 1905b: 382; Wheeler, W.M. 1910g: 135; Emery, 1911d: 88 [subtribe Ponerini]; Forel, 1917: 238; Wheeler, W.M. 1922a: 650; all subsequent authors.

Cryptopone as junior synonym of Pachycondyla: Mackay & Mackay, 2010: 3.

Pseudoponera Emery, 1900c: 314 [as subgenus of Pachycondyla]. Type-species: Ponera quadridentata (junior synonym of Formica stigma), by monotypy.

Taxonomic history

[Pseudoponera also described as new by Emery, 1901a: 42. Type-species not Ponera amblyops, unjustified subsequent designation by Emery, 1901a: 42; repeated in Wheeler, W.M. 1911f: 171, Wheeler, W.M. 1922a: 779; Donisthorpe, 1943g: 723.]

Pseudoponera in Pachycondylinae, Pachycondylini: Ashmead, 1905b: 382.

Pseudoponera in Ponerinae, Ponerini: Emery, 1911d: 86 [subtribe Pachycondylini]; Forel, 1917: 238; all subsequent authors.

Pseudoponera as subgenus of Pachycondyla: Emery, 1900c: 314; Emery, 1901a: 42.

Pseudoponera as subgenus of Euponera: Forel, 1900c: 141; Forel, 1900e: 398; Emery, 1909c: 364; Wheeler, W.M. 1910g: 135.

Pseudoponera as genus: Bingham, 1903: 91; Ashmead, 1905b: 382; Emery, 1911d: 86; Forel, 1917: 238; Wheeler, W.M. 1922a: 649; Donisthorpe, 1943g: 722; Chapman & Capco, 1951: 74.

Pseudoponera as junior synonym of Pachycondyla: Brown, 1973b: 184 [provisional]; Snelling, R.R. 1981: 389; Hölldobler & Wilson, 1990: 11; Brown, in Bolton, 1994: 164; Bolton, 2003: 167.

Brachyponera Emery, 1900c: 315 [as subgenus of Euponera]. Type-species: Euponera (Brachyponera) croceicornis, by monotypy.

Taxonomic history

[Brachyponera also described as new by Emery, 1901a: 43. Type-species not Ponera sennaarensis, unjustified subsequent designation by Emery, 1901a: 43, repeated in Wheeler, W.M. 1911f: 160, Emery, 1911d: 84, Wheeler, W.M. 1922a: 777, Donisthorpe, 1943f: 628, Wilson, 1958d: 346; Bolton, 1973a: 335; Taylor & Brown, D.R. 1985: 23.]

Brachyponera in Pachycondylinae, Pachycondylini: Ashmead, 1905b: 382.

Brachyponera in Ponerinae, Ponerini: Emery, 1911d: 83; all subsequent authors.

Brachyponera as subgenus of Euponera: Emery, 1900c: 315; Emery, 1909c: 366; Wheeler, W.M. 1910g: 135; Emery, 1911d: 83; Arnold, 1915: 72; Forel, 1917: 237; Wheeler, W.M. 1922a: 649; subsequent authors to the following.

Brachyponera as genus: Bingham, 1903: 101; Ashmead, 1905b: 382; Wilson, 1958d: 346; Taylor & Brown, D.R. 1985: 23; Wheeler, G.C. & Wheeler, J. 1985: 256; Taylor, 1987a: 9; Ogata, 1987: 116; Dlussky & Fedoseeva, 1988: 78; Hölldobler & Wilson, 1990: 10.

Brachyponera as junior synonym of Pachycondyla: Snelling, R.R. 1981: 389 [provisional]; Brown, in Bolton, 1994: 164; Bolton, 2003: 167.

Mesoponera Emery, 1900d: 668 [as subgenus of Euponera]. Type-species: Ponera melanaria, by monotypy.

Taxonomic history

[Mesoponera also described as new by Emery, 1901a: 43. Type-species not Ponera caffraria, unjustified subsequent designation by Emery, 1901a: 43, repeated by Emery, 1911d: 81, Wheeler, W.M. 1911f: 167, Wheeler, W.M. 1922a: 775, Donisthorpe, 1943f: 661, Wilson, 1958d: 349; Kempf, 1972a: 141, Bolton, 1973a: 338; Taylor & Brown, D.R. 1985: 35.]

Mesoponera in Pachycondylinae, Pachycondylini: Ashmead, 1905b: 382.

Mesoponera in Ponerinae, Ponerini: Emery, 1911d: 80; all subsequent authors.

Mesoponera as subgenus of Euponera: Emery, 1900d: 668; Wheeler, W.M. 1910g: 135; Emery, 1911d: 80; Arnold, 1915: 64; Forel, 1917: 237; Gallardo, 1918b: 65; Wheeler, W.M. 1922a: 649; Borgmeier, 1923: 71; Donisthorpe, 1943f: 661; Chapman & Capco, 1951: 64; Bernard, 1953b: 190.

Mesoponera as genus: Bingham, 1903: 99; Ashmead, 1905b: 382; Wilson, 1958d: 349; Kempf, 1972a: 141; Taylor & Brown, D.R. 1985: 34; Taylor, 1987a: 39; Dlussky & Fedoseeva, 1988: 78.

Mesoponera as junior synonym of Pachycondyla: Brown, 1973b: 182 [provisional]; Snelling, R.R. 1981: 389; Hölldobler & Wilson, 1990: 11; Brown, in Bolton, 1994: 164; Bolton, 2003: 167.

Eumecopone Forel, 1901e: 335 [as subgenus of Neoponera]. Type-species: Neoponera (Eumecopone) agilis, by monotypy.

Taxonomic history

Eumecopone as subgenus of Neoponera: Forel, 1901e: 335; Wheeler, W.M. 1910g: 135; Emery, 1911d: 71; Forel, 1917: 237; Wheeler, W.M. 1922a: 648; subsequent authors to the following.

Eumecopone junior synonym of Pachycondyla: Brown, 1973b: 180 [provisional]; Snelling, R.R. 1981: 389; Hölldobler & Wilson, 1990: 11; Brown, in Bolton, 1994: 164; Bolton, 2003: 167.

Hagensia Forel, 1901f: 333 [as subgenus of Megaloponera (sic)]. Type-species: Megaloponera (Hagensia) havilandi, by monotypy.

Taxonomic history

Hagensia in Ponerinae, Ponerini: Emery, 1911d: 69 [subtribe Pachycondylini]; all subsequent authors.

Hagensia as subgenus of Megaponera: Forel, 1901f: 333; Wheeler, W.M. 1910g: 135; Emery, 1911d: 61.

Hagensia as subgenus of Euponera: Forel, 1917: 237.

Hagensia as genus: Arnold, 1926: 202; Donisthorpe, 1943f: 648; Wheeler, G.C. & Wheeler, J. 1985: 256; Dlussky & Fedoseeva, 1988: 78; Hölldobler & Wilson, 1990: 11.

Hagensia as junior synonym of Pachycondyla: Brown, in Bolton, 1994: 164; Bolton, 2003: 167.

Neoponera Emery, 1901a: 43. Type-species: Formica villosa, by original designation.

Taxonomic history

Neoponera in Pachycondylinae, Pachycondylini: Ashmead, 1905b: 382.

Neoponera in Ponerinae, Ponerini: Wheeler, W.M. 1910g: 135; Emery, 1911d: 70 [subtribe Pachycondylini]; Forel, 1917: 237; Wheeler, W.M. 1922a: 648; all subsequent authors.

Neoponera as junior synonym of Pachycondyla: Brown, 1973b: 183 [provisional]; Snelling, R.R. 1981: 389; Hölldobler & Wilson, 1990: 11; Brown, in Bolton, 1994: 164; Bolton, 2003: 168.

Trachymesopus Emery, 1911d: 84 [as subgenus of Euponera]. Type-species: Formica stigma, by original designation.

Taxonomic history

Trachymesopus in Ponerinae, Ponerini: Emery, 1911d: 84; all subsequent authors.

Trachymesopus as subgenus of Euponera: Emery, 1911d: 84; Arnold, 1915: 74; Forel, 1917: 237; Wheeler, W.M. 1922a: 649; Borgmeier, 1923: 72; Donisthorpe, 1943g: 733; Creighton, 1950a: 46; Chapman & Capco, 1951: 65; Smith, M.R. 1951a: 786; Bernard, 1953b: 195.

Trachymesopus as genus: Wilson, 1958d: 352; Kempf, 1960f: 423; Brown, 1963: 6; Kempf, 1972a: 251; Taylor & Brown, D.R. 1985: 52; Ogata, 1987: 114; Dlussky & Fedoseeva, 1988: 78.

Trachymesopus as junior synonym of Pachycondyla: Brown, 1973b: 185 [provisional]; Snelling, R.R. 1981: 389; Hölldobler & Wilson, 1990: 11; Brown, in Bolton, 1994: 164.

Trachymesopus as junior synonym of Pseudoponera: Bolton, 1995b: 48 (because of synonymous type-species); Bolton, 2003: 168.

[Trachyponera Santschi, 1928a: 43, incorrect subsequent spelling.]

Xiphopelta Forel, 1913a: 108 [as subgenus of Ponera]. Type-species: Ponera (Xiphopelta) arnoldi, by monotypy.

Taxonomic history

Xiphopelta as subgenus of Ponera: Forel, 1913a: 108; Donisthorpe, 1943g: 737.

Xiphopelta as subgenus of Euponera: Forel, 1917: 237; Emery, 1919c: 106; Bernard, 1953b: 191.

Xiphopelta as junior synonym of Mesoponera: Wheeler, W.M. 1922a: 775; all subsequent authors to the following.

Xiphopelta as junior synonym of Pachycondyla: Brown, 1973b: 185 [provisional]; Snelling, R.R. 1981: 389; Hölldobler & Wilson, 1990: 11; Brown, in Bolton, 1994: 164; Bolton, 2003: 168.

[Hiphopelta Forel, 1913a: 108, incorrect subsequent spelling.]

Termitopone Wheeler, W.M. 1936d: 159. Type-species: Ponera laevigata, by original designation.

Taxonomic history

Termitopone in Ponerinae, Ponerini: Donisthorpe, 1943g: 731.

Termitopone as junior synonym of Pachycondyla: Brown, 1973b: 185 [provisional]; Snelling, R.R. 1981: 389; Hölldobler & Wilson, 1990: 11; Brown, in Bolton, 1994: 164; Bolton, 2003: 168.

Syntermitopone Wheeler, W.M. 1936d: 169 [as subgenus of Termitopone.] Type-species: Ponera commutata, by original designation.

Taxonomic history

Syntermitopone as subgenus of Termitopone: Wheeler, W.M. 1936d: 169.

Syntermitopone as genus: Kusnezov, 1956: 15.

Syntermitopone as junior synonym of Termitopone: Borgmeier, 1959a: 312.

Syntermitopone as junior synonym of Pachycondyla: Brown, 1973b: 185 [provisional]; Snelling, R.R. 1981: 389; Hölldobler & Wilson, 1990: 11; Brown, in Bolton, 1994: 164; Bolton, 2003: 168.

Wadeura Weber, 1939a: 102. Type-species: Wadeura guianensis, by original designation.

Taxonomic history

Wadeura in Ponerinae, Ponerini: Donisthorpe, 1943g: 735.

Wadeura as junior synonym of Pachycondyla: Brown, 1973b: 185 [provisional]; Snelling, R.R. 1981: 389; Hölldobler & Wilson, 1990: 11; Brown, in Bolton, 1994: 164; Bolton, 2003: 168.

Pseudoneoponera Donisthorpe, 1943d: 439. Type-species: Pseudoneoponera verecundae, by original designation.

Taxonomic history

Pseudoneoponera in Ponerinae, Pseudoneoponerini: Chapman & Capco, 1951: 77.

Pseudoneoponera as junior synonym of Bothroponera: Wilson, 1958d: 361.

Pseudoneoponera as junior synonym of Pachycondyla: Brown, 1973b: 184 [provisional]; Snelling, R.R. 1981: 389; Hölldobler & Wilson, 1990: 11; Brown, in Bolton, 1994: 164; Bolton, 2003: 168.

Genus Pachycondyla references

Mayr, 1862: 719 (diagnosis); Roger, 1863b: 16, 17, 18 (catalogue); Mayr, 1863: 397, 428, 439, 440 (catalogue); Mayr, 1865: 13, 15 (Bothroponera, Pachycondyla, Megaponera, Paltothyreus diagnoses); Mayr, 1867a: 82 (Bothroponera diagnosis); Mayr, 1870a: 396 (Colombia + Panama (= New Grenada) species key); Emery, 1887b: 18 (footnote) (Asia & Australia Bothroponera species key); Emery, 1890a: 71 (Neotropical species key); Dalla Torre, 1893: 30, 31, 32, 35 (Ectomomyrmex, Megaponera, Ophthalmopone, Paltothyreus, Pachycondyla, Bothroponera catalogues); Forel, 1900d: 321 (India & Sri Lanka Ectomomyrmex species key); Bingham, 1903: 86, 91, 96, 101 (India, Sri Lanka & Burma Ectomomyrmex, Pseudoponera, Bothroponera, Brachyponera species keys); Emery, 1911d: 62 (Paltothyreus diagnosis, catalogue); Emery, 1911d: 67 (Megaponera diagnosis, catalogue); Emery, 1911d: 69 (Megaponera (Hagensia) and Ophthalmopone diagnoses, catalogues); Emery, 1911d: 70 (Neoponera diagnosis, catalogue); Emery, 1911d: 73 (diagnosis, subgenera key, catalogue); Emery, 1911d: 75 (P. (Bothroponera) diagnosis, catalogue); Emery, 1911d: 78 (P. (Ectomomyrmex) diagnosis, catalogue); Emery, 1911d: 80 (Euponera diagnosis, subgenera key, catalogue); Emery, 1911d: 80 (Euponera (Mesoponera) diagnosis, catalogue); Emery, 1911d: 83 (Euponera (Brachyponera) diagnosis, catalogue); Emery, 1911d: 84 (Euponera (Trachymesopus) diagnosis, catalogue); Emery, 1911d: 86 (Pseudoponera diagnosis, catalogue); Emery, 1911d: 88 (Cryptopone diagnosis, catalogue); Arnold, 1915: 43, 46, 49, 53, 63 (Paltothyreus, Megaponera, Ophthalmopone, Pachycondyla, Euponera diagnoses); Arnold, 1915: 50, 55, 64 (South Africa Ophthalmopone, Bothroponera, Euponera (Mesoponera) species keys); Gallardo, 1918b: 56, 66 (Argentina Neoponera, Mesoponera, keys); Wheeler, W.M. 1922a: 72 (Afrotropical Bothroponera species key); Wheeler, W.M. 1922a: 60, 63, 69, 81 (Paltothyreus, Megaponera, Bothroponera, Euponera diagnoses); Wheeler, W.M. 1922a: 763, 766, 768, 769, 774, 780 (Afrotropical Paltothyreus, Megaponera, Ophthalmopone, Bothroponera, Ectomomyrmex, Euponera, Pseudoponera, Cryptopone catalogues); Wheeler, W.M. 1922a: 1007, 1008 (Malagasy Bothroponera, Euponera catalogues); Arnold, 1926: 203 (South Africa Hagensia species key); Wheeler, W.M. 1933g: 5 (Cryptopone all species key); Santschi, 1935b: 259 (Africa Mesoponera, key); Creighton, 1950a: 45 (North America Euponera species key); Arnold, 1951: 53 (Hagensia species key); Chapman & Capco, 1951: 49, 52, 60, 63, 74 (Asia Bothroponera, Cryptopone, Ectmomomyrmex, Euponera, Pseudoponera checklists); Arnold, 1952a: 461 (South Africa Bothroponera, key); Bernard, 1953b: 191 (Guinea Euponera (Xiphopelta) species key); Brown, 1958h: 20 (New Zealand species); Wilson, 1958d: 346, 357, 362 (Melanesia Brachyponera, Cryptopone, Ectomomyrmex species keys); Kempf, 1961b: 192 (Brazil species key); Brown, 1963: 6, 9 (reviews of Cryptopone, Trachymesopus & Ectomomyrmex); Kempf, 1964e: 51 (Neotropical Pachycondyla species key); Kusnezov, 1969: 36 (Argentina Neoponera species key); Kempf, 1972a: 90, 141, 160, 174, 248, 251, 257 (Neotropical Cryptopone, Mesoponera, Neoponera, Pachycondyla, Termitopone, Trachymesopus, Wadeura catalogues); Alayo, 1974: 7 (Cuba species key); Smith, D.R. 1979: 1339, 1341 (North America, Pachycondyla, Brachyponera, Cryptopone catalogues); Taylor & Brown, D.R. 1985: 21, 23, 28, 29, 34, 34, 52 (Australia, catalogue); Taylor, 1987a: 9, 23, 26, 39, 80 (Australia, New Caledonia & New Zealand checklists); Morisita, Kubota, et al. 1989: 18 (Japan Ectomomyrmex, Cryptopone, Brachyponera, Trachymesopus species key); Brandão, 1991: 354, 360, 365, 381, 382 (Neotropical catalogue); Xu, 1994b: 182 (China Brachyponera species key); Bolton, 1994: 164 (synoptic classification); Bolton, 1995a: 1051 (census); Bolton, 1995b: 166, 302 (Cryptopone, Pachycondyla catalogues); Wu, J. & Wang, 1995: 45 (China species key); Xu, 1995b: 104 (China species key); Collingwood & Agosti, 1996: 310 (Cryptopone, Pachycondyla Saudi Arabia species key); Kim, Kim & Kim, 1998: 147 (Korea species key); Shattuck, 1999: 182, 196 (Australia Cryptopone, Pachycondyla fauna synopsis); Zhou, 2001b: 35, 49 (China, Guangxi Cryptopone, Pachycondyla species keys); Dlussky & Rasnitsyn, 2002: 424 (wingless female fossils, diagnosis); Wild, 2002: 4 (Paraguay species key); Bolton, 2003: 161, 165 (Cryptopone, Pachycondyla synopses); Wild, 2005: 5 (P. apicalis complex key); Yamane, 2007: 652 (P. nigrita species group key); Mackay, Mackay, Fernández & Arias-Penna, in Fernández, 2008: 172 (Neotropical species key); Terayama, 2009: 105, 108 (Taiwan Pachycondyla, Cryptopone species keys); Heterick, 2009: 134 (south-western Australia species key); Mackay & Mackay, 2010: 26, 107 (New World Pachycondyla species revision, species complexes key and species key).

Ponerinae Ponerini Plectroctena TRUE TRUE FALSE

16 species

Plectroctena Smith, F. 1858b: 101. Type-species: Plectroctena mandibularis, by monotypy.

Taxonomic history

Plectroctena in Poneridae: Smith, F. 1858b: 101.

Plectroctena in Ponerinae: Mayr, 1862: 713 [Poneridae]; Mayr, 1865: 12 [Poneridae]; Dalla Torre, 1893: 31.

Plectroctena in Pachycondylinae, Pachycondylini: Ashmead, 1905b: 382.

Plectroctena in Ponerinae, Ponerini: Emery, 1895j: 767; Wheeler, W.M. 1910g: 135; Emery, 1911d: 94 [subtribe Plectroctenini]; Arnold, 1915: 84; Forel, 1917: 238; Wheeler, W.M. 1922a: 650; all subsequent authors.

Junior synonym of PLECTROCTENA

Cacopone Santschi, 1914d: 325. Type-species: Cacopone hastifer, by monotypy.

Taxonomic history

Cacopone in Ponerinae, Ponerini: Forel, 1917: 238 [subtribe Plectroctenini]; Wheeler, W.M. 1922a: 650; Santschi, 1924a: 159; Donisthorpe, 1943f: 629.

Cacopone as junior synonym of Plectroctena: Bolton, 1974c: 313.

Genus Plectroctena references

Roger, 1863b: 19 (catalogue); Mayr, 1863: 442 (catalogue); Mayr, 1865: 12 (diagnosis); Dalla Torre, 1893: 31 (catalogue); Emery, 1911d: 94 (diagnosis, catalogue); Arnold, 1915: 84 (diagnosis, South Africa species); Wheeler, W.M. 1922a: 85, 783, 785, 786 (diagnosis, Plectroctena, Myopias, Cacopone catalogues); Santschi, 1924a: 172 (all species revision, key); Bolton, 1974c: 309 (diagnosis, all species revision, key); Bolton, 1994: 164 (synoptic classification); Bolton, 1995a: 1052 (census); Bolton, 1995b: 337 (catalogue); Bolton & Brown, 2002: 11 (all species key); Bolton, 2003: 169 (synopsis).

Ponerinae Ponerini Ponera TRUE TRUE FALSE

Extant: 52 species

Fossil: 5 species

Ponera Latreille, 1804: 179. Type-species: Formica coarctata, by subsequent designation of Westwood, 1840a: 83 (see discussion in Taylor, 1967a: 6).

Taxonomic history

Ponera in Ponérites: Lepeletier de Saint-Fargeau, 1835: 188.

Ponera in Poneridae: Smith, F. 1858b: 83; Smith, F. 1871a: 320.

Ponera in Ponerinae: Mayr, 1855: 386 [Poneridae]; Smith, F. 1857a: 65 [Poneridae]; Mayr, 1861: 54 [Poneridae]; Mayr, 1862: 713 [Poneridae]; Mayr, 1865: 13 [Poneridae]; Emery & Forel, 1879a: 455 [Poneridae]; Dalla Torre, 1893: 37.

Ponera in Ponerinae, Ponerini: Emery, 1895j: 767; Forel, 1899c: 15; Ashmead, 1905b: 382; Wheeler, W.M. 1910g: 135; Emery, 1911d: 88 [subtribe Ponerini]; Wheeler, W.M. 1915h: 38; Arnold, 1915: 77; Forel, 1917: 238; Wheeler, W.M. 1922a: 650; all subsequent authors.

Junior synonyms of PONERA

Pseudocryptopone Wheeler, W.M. 1933g: 12. Type-species: Cryptopone tenuis, by original designation.

Taxonomic history

Pseudocryptopone in Ponerinae, Ponerini: Donisthorpe, 1943g: 722.

Pseudocryptopone as junior synonym of Ponera: Wilson, 1957b: 356.

Selenopone Wheeler, W.M. 1933g: 19. Type-species: Ponera selenophora, by original designation.

Taxonomic history

Selenopone in Ponerinae, Ponerini: Donisthorpe, 1943g: 725.

Selenopone as junior synonym of Ponera: Wilson, 1957b: 356.

Pteroponera Bernard, 1950a: 3. Type-species: Pteroponera sysphinctoides, by monotypy.

Taxonomic history

Pteroponera as junior synonym of Ponera: Brown, 1973b: 184 [provisional]; Brown, in Bolton, 1994: 164.

Genus Ponera references

[References before Taylor, 1967a refer to both Ponera and Hypoponera.]

Mayr, 1855: 386 (diagnosis); Smith, F. 1858b: 83 (diagnosis); Mayr, 1862: 721 (diagnosis); Roger, 1863b: 16 (catalogue); Mayr, 1863: 447 (catalogue); Mayr, 1865: 13 (diagnosis); Mayr, 1867a: 85 (diagnosis); Forel, 1874: 64 (Switzerland species key); Mayr, 1879: 661 (all species key); André, 1882c: 239 (Europe & Algeria species key); Cresson, 1887: 258 (U.S.A. catalogue); Dalla Torre, 1893: 37 (catalogue); Emery, 1895b: 60 (Mediterranean species key); Emery, 1896g: 53 (New World species key); Emery, 1900c: 316 (Australia, Papuasia & Oceania species key); Forel, 1900d: 322 (India & Sri Lanka species key); Bingham, 1903: 89 (India, Sri Lanka & Burma species key); Emery, 1909c: 367 (Palaearctic species key); Emery, 1911d: 88 (diagnosis, catalogue); Arnold, 1915: 76 (diagnosis, South Africa species key); Emery, 1916b: 107 (Italy species key); Bondroit, 1918: 81 (France & Belgium species key); Gallardo, 1918b: 73 (Argentina species key); Mann, 1921: 419 (Fiji species key); Wheeler, W.M. 1922a: 780, 1009 (Afrotropical, Malagasy catalogues); Menozzi, 1931b: 266 (Costa Rica species key); Wheeler, W.M. 1933g: 13 (Pseudocryptopone species key); Smith, M.R. 1936: 420 (North America species key); Stitz, 1939: 58 (Germany species key); Creighton, 1950a: 47 (North America species key); Chapman & Capco, 1951: 68, 73, 74 (Asia Ponera, Pseudocryptopone, Selenopone checklists); Bernard, 1953b: 197 (West Africa species key); Wilson, 1957b: 359, 376 (P. tenuis, P. selenophora groups revisions, keys); Wilson, 1958d: 323, 343 (Melanesia & Moluccas species, Fiji species revisions, keys); Brown, 1958h: 22 (New Zealand species); Taylor, 1960: 180 (P. leae complex, key); Taylor, 1964: 139 (†fossil species, review); Taylor, 1967a: 18 (diagnosis, all species revision, key); Bernard, 1967: 83 (diagnosis, Western Europe species key); Smith, D.R. 1979: 1341 (North America catalogue); Taylor & Brown, D.R. 1985: 38 (Australia catalogue); Taylor, 1987a: 63 (Australia, New Caledonia & New Zealand, checklist); Morisita, Kubota, et al. 1989: 22 (Japan species key); Perrault, 1993: 337 (addition to Taylor, 1967a key); Bolton, 1994: 164 (synoptic classification); Bolton, 1995a: 1052 (census); Bolton, 1995b: 360 (catalogue); Terayama, 1996: 14 (Japan species key); Kim, Kim & Kim, 1998: 149 (Korea species key); Shattuck, 1999: 199 (Australia synopsis); Xu, 2001a: 52 and Xu, 2001c: 218 (China species keys); Zhou, 2001b: 37 (China, Guangxi species key); Bolton, 2003: 170 (synopsis); Yoshimura, Hosoishi, et al. 2009: 200 (Japan species key); Terayama, 2009: 109 (Taiwan species key).

Formicinae Lasiini Acropyga arnoldi TRUE TRUE FALSE

arnoldi. Acropyga (Malacomyrma) arnoldi Santschi, 1926b: 245 (w.q.) SOUTH AFRICA. Prins, 1982: 245 (m.). Senior synonym of rhodesiana: Prins, 1982: 238. See also: LaPolla, 2004a: 35.

Aenictinae Aenictini Aenictus decolor TRUE TRUE FALSE

decolor. Typhlatta decolor Mayr, 1879: 668 (diagnosis in key) (w.) EAST AFRICA. Gotwald & Leroux, 1980: 602 (q.). Combination in Aenictus: Dalla Torre, 1893: 7. Senior synonym of batesi, bidentatus: Gotwald & Leroux, 1980: 600.

Aenictinae Aenictini Aenictus eugenii TRUE TRUE FALSE

2 subspecies

eugenii. Aenictus eugenii Emery, 1895h: 17, pl. 2, figs. 1-4 (w.) SOUTH AFRICA. Gotwald & Cunningham-Van Someren, 1976: 183 (q.). Senior synonym of kenyensis: Gotwald & Cunningham-Van Someren, 1976: 183. Current subspecies: nominal plus caroli, henrii.

Aenictinae Aenictini Aenictus inconspicuus TRUE TRUE FALSE

inconspicuus. Aenictus inconspicuus Westwood, 1845: 85 (m.) SOUTH AFRICA. [Also described as new by Westwood, 1847a: 238.]

Aenictinae Aenictini Aenictus mariae TRUE TRUE FALSE

1 subspecies

mariae. Aenictus mariae Emery, 1895h: 18, pl. 2, figs. 5-7 (w.) SOUTH AFRICA. Current subspecies: nominal plus natalensis.

Aenictinae Aenictini Aenictus rixator TRUE TRUE FALSE

rixator. Aenictus rixator Forel, in Emery, 1901d: 48 (w.) SOUTH AFRICA.

Aenictinae Aenictini Aenictus rotundatus TRUE TRUE FALSE

2 subspecies

rotundatus. Aenictus rotundatus Mayr, 1901b: 1 (w.) SOUTH AFRICA. Current subspecies: nominal plus guineensis, merwei.

Aenictinae Aenictini Aenictus steindachneri TRUE TRUE FALSE

steindachneri. Aenictus steindachneri Mayr, 1901b: 2 (w.) SOUTH AFRICA.

Aenictinae Aenictini Aenictus mariae natalensis TRUE TRUE FALSE

natalensis. Aenictus mariae var. natalensis Forel, in Emery, 1901d: 49 (w.) SOUTH AFRICA.

Aenictinae Aenictini Aenictus rotundatus merwei TRUE TRUE FALSE

merwei. Aenictus rotundatus var. merwei Santschi, 1932a: 382 (w.) SOUTH AFRICA.

Formicinae Plagiolepidini Agraulomyrmex meridionalis TRUE TRUE FALSE

meridionalis. Agraulomyrmex meridionalis Prins, 1983: 2, fig. 1 (w.) SOUTH AFRICA.

Ponerinae Ponerini Anochetus africanus TRUE TRUE FALSE

africanus. Stenomyrmex africanus Mayr, 1865: 11 (footnote) (w.) GHANA. André, 1892a: 47 (q.). Combination in Anochetus: Forel, 1887: 382. Senior synonym of camerunensis: Mayr, 1904b: 2; of pasteuri: Brown, 1978c: 556 (see also p. 602).

Ponerinae Ponerini Anochetus faurei TRUE TRUE FALSE

faurei. Anochetus faurei Arnold, 1948: 215, fig. 4 (w.) SOUTH AFRICA. See also: Brown, 1978c: 609.

Ponerinae Ponerini Anochetus fuliginosus TRUE TRUE FALSE

fuliginosus. Anochetus fuliginosus Arnold, 1948: 214, fig. 3 (w.) SOUTH AFRICA. See also: Brown, 1978c: 605.

Ponerinae Ponerini Anochetus jonesi TRUE TRUE FALSE

jonesi. Anochetus jonesi Arnold, 1926: 216, fig. 61 (w.q.m.) SOUTH AFRICA.

Ponerinae Ponerini Anochetus levaillanti TRUE TRUE FALSE

levaillanti. Anochetus levaillanti Emery, 1895h: 21 (w.) SOUTH AFRICA. Arnold, 1915: 108 (q.). See also: Brown, 1978c: 594.

Ponerinae Ponerini Anochetus madagascarensis TRUE TRUE FALSE

madagascarensis. Anochetus africanus var. madagascarensis Forel, 1887: 382 (w.) MADAGASCAR. Fisher & Smith, 2008: 12 (q.m.). Raised to species and senior synonym of friederichsi: Brown, 1978c: 557.

Ponerinae Ponerini Anochetus natalensis TRUE TRUE FALSE

natalensis. Anochetus natalensis Arnold, 1926: 215 (w.) SOUTH AFRICA. See also: Brown, 1978c: 603.

Ponerinae Ponerini Anochetus obscuratus TRUE TRUE FALSE

obscuratus. Anochetus madagascarensis var. obscurata Santschi, 1911c: 351 (w.q.) TANZANIA. Subspecies of africanus: Emery, 1911d: 108; Santschi, 1914b: 56. Raised to species: Santschi, 1923e: 265. Senior synonym of schoutedeni, ustus: Brown, 1978c: 558 (see also p. 602).

Ponerinae Ponerini Anochetus punctaticeps TRUE TRUE FALSE

punctaticeps. Anochetus punctaticeps Mayr, 1901b: 4 (w.) SOUTH AFRICA. See also: Brown, 1978c: 606.

Ponerinae Ponerini Anochetus talpa TRUE TRUE FALSE

talpa. Anochetus talpa Forel, 1901f: 351 (w.) SOUTH AFRICA.

Formicinae Lasiini Anoplolepis custodiens TRUE TRUE FALSE

3 subspecies

custodiens. Formica custodiens Smith, F. 1858b: 33 (w.) SOUTH AFRICA. Mayr, 1895: 148 (m.); Emery, 1895h: 44 (q.); Wheeler, G.C. & Wheeler, J. 1953c: 135 (l.). Combination in Plagiolepis: Emery, 1892a: 117; in Plagiolepis (Anoplolepis): Santschi, 1914b: 123; in Plagiolepis (Zealleyella): Arnold, 1922: 586; in Anoplolepis: Emery, 1925b: 17. Senior synonym of hendecarthrus: Mayr, 1865: 54; of berthoudi: Forel, 1879a: 91. See also: Prins, 1982: 218. Current subspecies: nominal plus detrita, hirsuta, pilipes.

Formicinae Lasiini Anoplolepis fallax TRUE TRUE FALSE

fallax. Plagiolepis fallax Mayr, 1865: 54 (w.) SOUTH AFRICA. Forel, 1910f: 24 (q.). Combination in Plagiolepis (Zealleyella): Arnold, 1922: 589; in Anoplolepis: Emery, 1925b: 17. Subspecies of custodiens: Mayr, 1893: 5; Arnold, 1922: 589. Revived status as species: Emery, 1925b: 17.

Formicinae Lasiini Anoplolepis nuptialis TRUE TRUE FALSE

nuptialis. Plagiolepis (Anoplolepis) nuptialis Santschi, 1917b: 289 (m.) SOUTH AFRICA. Prins, 1982: 222 (q.). Combination in Anoplolepis: Emery, 1925b: 18; in Anoplolepis (Zealleyella): Santschi, 1926a: 14.

Formicinae Lasiini Anoplolepis rufescens TRUE TRUE FALSE

rufescens. Plagiolepis (Anoplolepis) rufescens Santschi, 1917b: 289 (s.w.) SOUTH AFRICA. Combination in Anoplolepis: Emery, 1925b: 18; in Anoplolepis (Zealleyella): Santschi, 1926a: 14.

Formicinae Lasiini Anoplolepis steingroeveri parsonsi TRUE TRUE FALSE

parsonsi. Anoplolepis (Zealleyella) braunsi st. parsonsi Santschi, 1937g: 83 (w.) SOUTH AFRICA. Currently subspecies of steingroeveri: Bolton, 1995b: 67.

Ponerinae Ponerini Asphinctopone differens TRUE TRUE FALSE

differens. Asphinctopone differens Bolton & Fisher, 2008a: 59, figs. 1c,d; 2b,d (w.) CENTRAL AFRICAN REPUBLIC.

Dolichoderinae Tapinomini Axinidris lignicola TRUE TRUE FALSE

lignicola. Axinidris lignicola Snelling, R.R. 2007: 564, figs. 5, 15, 25 (w.) SOUTH AFRICA.

Dolichoderinae Tapinomini Axinidris mlalu TRUE TRUE FALSE

mlalu. Axinidris mlalu Snelling, R.R. 2007: 566 (w.) CENTRAL AFRICAN REPUBLIC.

Ponerinae Ponerini Boloponera vicans TRUE TRUE FALSE

vicans. Boloponera vicans Fisher, 2006: 113, figs. 1-4, 24 (w.) CENTRAL AFRICAN REPUBLIC.

Myrmicinae Stenammini Calyptomyrmex brunneus TRUE TRUE FALSE

brunneus. Calyptomyrmex brunneus Arnold, 1948: 221 (w.) SOUTH AFRICA. See also: Bolton, 1981a: 70.

Myrmicinae Stenammini Calyptomyrmex claviseta TRUE TRUE FALSE

claviseta. Dicroaspis claviseta Santschi, 1914e: 27 (w.) SOUTH AFRICA. Combination in Calyptomyrmex (Dicroaspis): Emery, 1915g: 15; in Calyptomyrmex: Bolton, 1981a: 71.

Formicinae Camponotini Camponotus angusticeps TRUE TRUE FALSE

angusticeps. Camponotus angusticeps Emery, 1886: 355, pl. 17, fig. 1 (w.) SOUTH AFRICA. Arnold, 1924: 688 (s.). Combination in C. (Myrmosericus): Forel, 1914a: 268.

Formicinae Camponotini Camponotus arnoldinus TRUE TRUE FALSE

arnoldinus. Camponotus (Myrmoturba) arnoldinus Forel, 1914d: 251 (s.w.) SOUTH AFRICA. Combination in C. (Tanaemyrmex): Emery, 1925b: 91. Subspecies of maculatus: Santschi, 1923e: 290. Revived status as species: Emery, 1925b: 91. See also: Arnold, 1922: 639.

Formicinae Camponotini Camponotus baynei TRUE TRUE FALSE

baynei. Camponotus maculatus r. baynei Arnold, 1922: 632 (s.w.) SOUTH AFRICA. Combination in C. (Tanaemyrmex): Emery, 1925b: 92. Raised to species: Emery, 1925b: 92.

Formicinae Camponotini Camponotus berthoudi TRUE TRUE FALSE

berthoudi. Camponotus berthoudi Forel, 1879a: 108, pl. 1, fig. 6 (w.) SOUTH AFRICA. Arnold, 1924: 700. Combination in C. (Myrmamblys): Forel, 1914a: 271; in C. (Myrmopiromis): Emery, 1925b: 128.

Formicinae Camponotini Camponotus bertolonii TRUE TRUE FALSE

bertolonii. Camponotus bertolonii Emery, 1895h: 51 (s.w.q.) SOUTH AFRICA. Mayr, 1901b: 28 (m.). Combination in C. (Myrmoturba): Forel, 1914a: 267; Arnold, 1922: 651; in C. (Myrmotemnus): Emery, 1920b: 258; in C. (Myrmamblys): Santschi, 1914b: 134; Wheeler, W.M. 1922a: 989; Emery, 1925b: 141; in C. (Paramyrmamblys): Santschi, 1926c: 604.

Formicinae Camponotini Camponotus bianconii TRUE TRUE FALSE

bianconii. Camponotus bianconii Emery, 1895h: 52 (s.) SOUTH AFRICA. Combination in C. (Tanaemyrmex): Emery, 1925b: 92.

Formicinae Camponotini Camponotus bifossus TRUE TRUE FALSE

bifossus. Camponotus (Colobopsis) bifossus Santschi, 1917b: 293 (s.) SOUTH AFRICA.

Formicinae Camponotini Camponotus brevisetosus TRUE TRUE FALSE

brevisetosus. Camponotus fulvopilosus subsp. brevisetosus Forel, 1910f: 25 (w.q.) SOUTH AFRICA. Combination in C. (Myrmopiromis): Wheeler, W.M. 1922a: 986. Raised to species: Robertson, 1990: 327. See also: Robertson & Zachariades, 1997: 3.

Formicinae Camponotini Camponotus brookei TRUE TRUE FALSE

brookei. Camponotus (Myrmamblys) brookei Forel, 1914d: 259 (s.w.) SOUTH AFRICA. Arnold, 1924: 706 (s.q.). Combination in C. (Paramyrmamblys): Santschi, 1926c: 604.

Formicinae Camponotini Camponotus buttikeri TRUE TRUE FALSE

buttikeri. Camponotus (Tanaemyrmex) buttikeri Arnold, 1958: 126, fig. 7 (s.w.) SOUTH AFRICA.

Formicinae Camponotini Camponotus caffer TRUE TRUE FALSE

caffer. Camponotus caffer Emery, 1895h: 47, pl. 2, figs. 25, 26 (s.w.) SOUTH AFRICA. Combination in C. (Myrmoturba): Forel, 1914a: 267; in C. (Dinomyrmex): Emery, 1920b: 256; in C. (Tanaemyrmex): Emery, 1925b: 84. Subspecies of maculatus: Forel, 1901h: 69. Revived status as species: Forel, 1914a: 267; Emery, 1920b: 256; Arnold, 1922: 650; Wheeler, W.M. 1922a: 963.

Formicinae Camponotini Camponotus cleobulus TRUE TRUE FALSE

cleobulus. Camponotus (Myrmoturba) cleobulus Santschi, 1919b: 238 (s.w.) SOUTH AFRICA. Combination in C. (Tanaemyrmex): Emery, 1925b: 92.

Formicinae Camponotini Camponotus cognatocompressus TRUE TRUE FALSE

cognatocompressus. Camponotus maculatus r. cognatocompressus Forel, 1904b: 379 (w.q.m.) "AFRICA" (no locality). [First available use of Camponotus rubripes r. cognatus var. cognatocompressus Forel, 1886f: 151; unavailable name.] Junior synonym of xerxes: Karavaiev, 1910b: 9. Revived from synonymy and raised to species: Radchenko, 1997d: 810.

Formicinae Camponotini Camponotus cosmicus TRUE TRUE FALSE

cosmicus. Formica cosmica Smith, F. 1858b: 34 (s.w.) SOUTH AFRICA. Combination in Camponotus: Roger, 1863b: 2; in C. (Myrmosericus): Emery, 1925b: 106. Subspecies of rufoglaucus: Emery, 1895h: 49 (in key); Stitz, 1916: 401; Arnold, 1924: 693. Revived status as species: Santschi, 1935a: 280.

Formicinae Camponotini Camponotus cuneiscapus TRUE TRUE FALSE

cuneiscapus. Camponotus cuneiscapus Forel, 1910f: 29, pl. 1, fig. 2 (s.w.) SOUTH AFRICA. Arnold, 1924: 679 (q.). Combination in C. (Myrmopsamma): Forel, 1914a: 270; Wheeler, W.M. 1922a: 988; Arnold, 1924: 679; in C. (Camponotus): Santschi, 1923e: 292; in C. (Myrmamblys): Emery, 1925b: 142; in C. (Myrmespera): Santschi, 1926b: 248.

Formicinae Camponotini Camponotus detritus TRUE TRUE FALSE

detritus. Camponotus detritus Emery, 1886: 357 (w.) SOUTH AFRICA. Forel, 1910f: 26 (s.). Combination in C. (Myrmepomis): Forel, 1914a: 273; in C. (Myrmopiromis): Wheeler, W.M. 1922a: 986. Subspecies of fulvopilosus: Forel, 1910f: 26; Wheeler, W.M. 1922a: 986; Stitz, 1923: 166; Arnold, 1924: 739. Revived status as species: Forel, 1914a: 273; Emery, 1920b: 258; Emery, 1925b: 128; Robertson, 1990: 327. See also: Robertson & Zachariades, 1997: 5.

Formicinae Camponotini Camponotus dicksoni TRUE TRUE FALSE

dicksoni. Camponotus (Tanaemyrmex) dicksoni Arnold, 1948: 227, fig. 13 (s.w.) SOUTH AFRICA.

Formicinae Camponotini Camponotus emarginatus TRUE TRUE FALSE

emarginatus. Camponotus emarginatus Emery, 1886: 358, pl. 17, fig. 2 (s.w.q.m.) SOUTH AFRICA. Combination in C. (Myrmophyma): Forel, 1914a: 269; Arnold, 1922: 663; in C. (Myrmotemnus): Emery, 1920b: 258; in C. (Myrmamblys): Wheeler, W.M. 1922a: 989; in C. (Myrmespera): Santschi, 1926c: 604.

Formicinae Camponotini Camponotus eugeniae TRUE TRUE FALSE

1 subspecies

eugeniae. Camponotus micans r. eugeniae Forel, 1879a: 78 (s.w.) SOUTH AFRICA. Forel, 1886f: 174 (q.); Arnold, 1924: 687 (m.). Combination in C. (Myrmosericus): Forel, 1912i: 91. Subspecies of rufoglaucus: Mayr, 1893: 195. Raised to species: Forel, 1886f: 174; Emery, 1895h: 50. Current subspecies: nominal plus amplior.

Formicinae Camponotini Camponotus fulvopilosus TRUE TRUE FALSE

fulvopilosus. Formica fulvopilosa De Geer, 1778: 613, pl. 45, figs. 13, 14 (w.) SOUTH AFRICA. Forel, 1879a: 108 (s.); Forel, 1910f: 25 (m.); Arnold, 1924: 737 (q.). Combination in Camponotus: Mayr, 1862: 668; in C. (Myrmepomis): Forel, 1912i: 92; in C. (Myrmopiromis): Wheeler, W.M. 1922a: 985. Senior synonym of pilosa, rufiventris: Lepeletier, 1835: 213; Smith, F. 1858b: 32; Wheeler, W.M. 1922a: 985; of detritoides, flavopilosus: Robertson, 1990: 327. See also: Robertson & Zachariades, 1997: 8.

Formicinae Camponotini Camponotus guanchus TRUE TRUE FALSE

guanchus. Camponotus sicheli var. guancha Santschi, 1908: 534 (w.q.) "North Africa". Cagniant & Espadaler, 1993: 427 (m.). Combination in C. (Myrmentoma): Emery, 1925b: 121. Subspecies of lateralis: Emery, 1925a: 64; Emery, 1925b: 121. Raised to species: Cagniant & Espadaler, 1993: 426.

Formicinae Camponotini Camponotus havilandi TRUE TRUE FALSE

havilandi. Camponotus (Myrmophyma) havilandi Arnold, 1922: 666 (s.w.) SOUTH AFRICA. Combination in C. (Myrmamblys): Emery, 1925b: 140; in C. (Myrmespera): Santschi, 1926c: 604.

Formicinae Camponotini Camponotus importunus TRUE TRUE FALSE

importunus. Camponotus maculatus subsp. importunus Forel, 1911e: 288 (s.w.q.m.) SOUTH AFRICA. [First available use of Camponotus maculatus subsp. sexpunctatus var. importuna Forel, 1910f: 27; unavailable name.] Combination in C. (Myrmoturba): Emery, 1920c: 5; in C. (Tanaemyrmex): Emery, 1925b: 87. Subspecies of maculatus: Emery, 1920c: 5; Arnold, 1922: 635. Raised to species: Baroni Urbani, 1972: 131.

Formicinae Camponotini Camponotus klugii TRUE TRUE FALSE

klugii. Camponotus bedoti var. klugii Emery, 1895h: 51 (w.) SOUTH AFRICA. Combination in C. (Myrmamblys): Wheeler, W.M. 1922a: 989. Raised to species: Emery, 1925b: 141.

Formicinae Camponotini Camponotus knysnae TRUE TRUE FALSE

knysnae. Camponotus (Myrmoturba) maculatus r. knysnae Arnold, 1922: 642 (s.w.q.) SOUTH AFRICA. Combination in C. (Tanaemyrmex): Emery, 1925b: 100. Raised to species: Emery, 1925b: 100 (misspelled as kaysnae).

Formicinae Camponotini Camponotus liogaster TRUE TRUE FALSE

liogaster. Camponotus (Orthonotomyrmex) liogaster Santschi, 1932a: 390 (w.) SOUTH AFRICA.

Formicinae Camponotini Camponotus maculatus TRUE TRUE FALSE

5 subspecies

maculatus. Formica maculata Fabricius, 1782: 491 (w.) WEST AFRICA. Mayr, 1862: 654 (q.m.); Forel, 1891b: 29 (s.w.q.m.); Donisthorpe, 1915a: 221 (redescription of holotype). Combination in Camponotus: Mayr, 1862: 654; in C. (Myrmoturba): Forel, 1912i: 91; in C. (Tanaemyrmex): Emery, 1925b: 86. Subspecies of sylvaticus: Forel, 1879a: 64; of rubripes: Forel, 1886f: 143. Revived status as species: Emery, 1891b: 17. Senior synonym of atramentarius, ballioni, boera, cavallus, cluisoides, cognata, conakryensis, erythraea, flavifemur, flavominor, hannae, hieroglyphicus, intonsus, lacteipennis, liengmei, liocnemis, lividior, lohieri, manzer, mathildae, melanocnemis (and its junior synonym schultzei), nubis, proletaria, radamoides, sarmentus, schereri, semispicatus, sudanicus, thomensis, tuckeri, zumpti and material of the unavailable names calceatus, cataractae, citinus, cluis, contaminatus, diffusus, hansingi, importunoides, madecassa, pessimus referred here: Baroni Urbani, 1972: 125-127. Current subspecies: nominal plus foveolatus (unresolved junior homonym), miserabilis (unresolved junior homonym), obfuscatus, subnudus, sylvaticomaculatus, ugandensis.

Formicinae Camponotini Camponotus mayri TRUE TRUE FALSE

1 subspecies

mayri. Camponotus mayri Forel, 1879a: 92, pl. 1, fig. 1 (w.) SOUTH AFRICA. Combination in C. (Myrmentoma): Forel, 1913a: 146; in C. (Orthonotomyrmex): Forel, 1913b: 347; Emery, 1925b: 125. Current subspecies: nominal plus chimporensis.

Formicinae Camponotini Camponotus mystaceus TRUE TRUE FALSE

2 subspecies

mystaceus. Camponotus mystaceus Emery, 1886: 356, pl. 17, fig. 3 (s.w.m.) SOUTH AFRICA. Arnold, 1924: 675 (q.). Combination in C. (Myrmopsamma): Forel, 1914a: 261. Current subspecies: nominal plus exsanguis, kamae.

Formicinae Camponotini Camponotus nasutus TRUE TRUE FALSE

5 subspecies

nasutus. Camponotus nasutus Emery, 1895h: 53, pl. 2, fig. 27 (w.) SOUTH AFRICA. Combination in C. (Myrmophyma): Forel, 1914a: 269; in C. (Myrmotemnus): Emery, 1920b: 258; in C. (Myrmamblys): Santschi, 1921f: 311; in C. (Myrmespera): Santschi, 1926c: 604. Current subspecies: nominal plus fenestralis, pretiosus, quadridentatus, quinquedentatus, subnasutus.

Formicinae Camponotini Camponotus natalensis TRUE TRUE FALSE

4 subspecies

natalensis. Formica natalensis Smith, F. 1858b: 33 (s.w.q.m.) SOUTH AFRICA. Combination in Camponotus: Roger, 1863b: 2; in C. (Myrmoturba): Forel, 1914a: 267; in C. (Tanaemyrmex): Emery, 1925b: 103. See also: Arnold, 1922: 652. Current subspecies: nominal plus corvus, diabolus, fulvipes, politiceps.

Formicinae Camponotini Camponotus niveosetosus TRUE TRUE FALSE

2 subspecies

niveosetosus. Camponotus niveosetosus Mayr, 1862: 665 (w.) SOUTH AFRICA. Mayr, 1866b: 885 (q.); Forel, 1891b: 41 (s.); Arnold, 1922: 659 (m.). Combination in C. (Myrmoturba): Forel, 1914a: 267; in C. (Myrmepomis): Emery, 1920b: 259; in C. (Myrmopiromis): Wheeler, W.M. 1922a: 986. Current subspecies: nominal plus irredux, madagascarensis.

Formicinae Camponotini Camponotus orites TRUE TRUE FALSE

orites. Camponotus (Myrmamblys) orites Santschi, 1919a: 346, fig. 4 (w.) SOUTH AFRICA. Arnold, 1926: 290 (s.). Combination in C. (Paramyrmamblys): Santschi, 1926c: 604. Senior synonym of orinodromus: Arnold, 1926: 291.

Formicinae Camponotini Camponotus ostiarius TRUE TRUE FALSE

ostiarius. Camponotus (Myrmamblys) ostiarius Forel, 1914d: 258 (s.w.) SOUTH AFRICA. Arnold, 1924: 702 (q.m.). Combination in C. (Paramyrmamblys): Santschi, 1926c: 604.

Formicinae Camponotini Camponotus petersii TRUE TRUE FALSE

1 subspecies

petersii. Camponotus petersii Emery, 1895h: 50, pl. 2, fig. 28 (w.) SOUTH AFRICA. Forel, 1914d: 254 (s.); Arnold, 1924: 684 (q.m.). Combination in C. (Myrmosericus): Forel, 1912i: 91. Current subspecies: nominal plus janus.

Formicinae Camponotini Camponotus postoculatus TRUE TRUE FALSE

postoculatus. Camponotus (Myrmotrema) postoculatus Forel, 1914d: 254 (s.w.) SOUTH AFRICA.

Formicinae Camponotini Camponotus reevei TRUE TRUE FALSE

reevei. Camponotus (Myrmophyma) reevei Arnold, 1922: 671, fig. 58 (s.w.) SOUTH AFRICA. Combination in C. (Myrmamblys): Emery, 1925b: 141; in C. (Myrmespera): Santschi, 1926c: 604.

Formicinae Camponotini Camponotus scalaris TRUE TRUE FALSE

scalaris. Camponotus scalaris Forel, 1901d: 308 (s.w.q.) SOUTH AFRICA. Combination in C. (Myrmosphincta): Forel, 1912i: 92; in C. (Orthonotomyrmex): Emery, 1920b: 258; in C. (Myrmisolepis): Santschi, 1921f: 310; in C. (Myrmopiromis): Emery, 1925b: 129.

Formicinae Camponotini Camponotus sellidorsatus TRUE TRUE FALSE

sellidorsatus. Camponotus (Mayria) sellidorsatus Prins, 1973: 21, figs. 25, 26, 34 (w.) SOUTH AFRICA.

Formicinae Camponotini Camponotus storeatus TRUE TRUE FALSE

storeatus. Camponotus fulvopilosus subsp. storeatus Forel, 1910f: 26 (w.) SOUTH AFRICA. Arnold, 1924: 739 (s.q.). Combination in C. (Myrmopiromis): Wheeler, W.M. 1922a: 986. Raised to species: Robertson, 1990: 327. See also: Robertson & Zachariades, 1997: 15.

Formicinae Camponotini Camponotus traegaordhi TRUE TRUE FALSE

1 subspecies

traegaordhi. Camponotus (Myrmoturba) maculatus st. traegaordhi Santschi, 1914e: 38 (s.w.q.m.) SOUTH AFRICA. Combination in C. (Tanaemyrmex): Emery, 1925b: 85. Subspecies of guttatus: Emery, 1920c: 6; of maculatus: Arnold, 1922: 627. Raised to species: Santschi, 1926b: 252. Current subspecies: nominal plus fumeus.

Formicinae Camponotini Camponotus transvaalensis TRUE TRUE FALSE

2 subspecies

transvaalensis. Camponotus (Myrmespera) transvaalensis Arnold, 1948: 228, fig. 14 (s.w.) SOUTH AFRICA. Current subspecies: nominal plus prinsi, griqua.

Formicinae Camponotini Camponotus trifasciatus TRUE TRUE FALSE

trifasciatus. Camponotus (Myrmespera) debellator var. trifasciatus Santschi, 1926b: 248 (q.) SOUTH AFRICA. Raised to species: Santschi, 1926c: 604.

Formicinae Camponotini Camponotus valdeziae TRUE TRUE FALSE

valdeziae. Camponotus valdeziae Forel, 1879a: 90 (w.q.) SOUTH AFRICA. Arnold, 1924: 696 (s.). Combination in C. (Myrmosericus): Forel, 1914a: 268.

Formicinae Camponotini Camponotus vestitus TRUE TRUE FALSE

8 subspecies

vestitus. Formica vestita Smith, F. 1858b: 32 (q.) SOUTH AFRICA. Arnold, 1924: 694 (s.w.). Combination in Camponotus: Roger, 1863b: 2; in C. (Myrmosericus): Emery, 1925b: 106. Subspecies of rufoglaucus: Emery, 1894i: 47; Emery, 1895h: 48; Forel, 1907e: 16; Santschi, 1914d: 381; Arnold, 1924: 694. Revived status as species: Santschi, 1926b: 258; Santschi, 1930a: 74; Bernard, 1953b: 262. Current subspecies: nominal plus anthracinus, bombycinus, comptus, intuens, lujai, pectitus, perpectitus, strophiatus.

Formicinae Camponotini Camponotus victoriae TRUE TRUE FALSE

victoriae. Camponotus (Myrmopiromis) victoriae Arnold, 1959: 338, fig. 24 (s.w.) SOUTH AFRICA.

Formicinae Camponotini Camponotus rufoglaucus zulu TRUE TRUE FALSE

zulu. Camponotus rufoglaucus subsp. zulu Emery, 1895h: 50 (s.w.) SOUTH AFRICA. Combination in C. (Myrmosericus): Emery, 1925b: 107.

Formicinae Camponotini Camponotus olivieri tenuipilis TRUE TRUE FALSE

tenuipilis. Camponotus (Myrmotrema) olivieri var. tenuipilis Santschi, 1915c: 270 (in key) (w.) SOUTH AFRICA. [First available use of Camponotus foraminosus subsp. olivieri var. tenuipilis Emery, 1895h: 54; unavailable name.]

Formicinae Camponotini Camponotus nasutus subnasutus TRUE TRUE FALSE

subnasutus. Camponotus (Myrmophyma) nasutus var. subnasutus Arnold, 1922: 670 (s.w.) SOUTH AFRICA. Combination in C. (Myrmamblys): Emery, 1925b: 141.

Formicinae Camponotini Camponotus vestitus strophiatus TRUE TRUE FALSE

strophiatus. Camponotus (Myrmosericus) vestitus var. strophiatus Santschi, 1926b: 259 (w.) SOUTH AFRICA.

Formicinae Camponotini Camponotus werthi skaifei TRUE TRUE FALSE

skaifei. Camponotus (Tanaemyrmex) werthi var. skaifei Arnold, 1959: 336, fig. 22 (s.w.) SOUTH AFRICA.

Formicinae Camponotini Camponotus transvaalensis griqua TRUE TRUE FALSE

griqua. Camponotus (Myrmespera) transvaalensis subsp. griqua Arnold, 1952a: 465 (s.w.q.m.) SOUTH AFRICA.

Formicinae Camponotini Camponotus natalensis fulvipes TRUE TRUE FALSE

fulvipes. Camponotus (Myrmoturba) fulvipes Forel, 1914a: 267 (w.) SOUTH AFRICA. [First available use of Camponotus natalensis subsp. corvus var. fulvipes Emery, 1895h: 51; unavailable name.] Santschi, 1914e: 38 (q.m.). Currently subspecies of natalensis: Santschi, 1914e: 38; Arnold, 1922: 654; Emery, 1925b: 103.

Formicinae Camponotini Camponotus cubangensis dofleini TRUE TRUE FALSE

dofleini. Camponotus dofleini Forel, 1911e: 291 (w.) SOUTH AFRICA. Combination in C. (Myrmosphincta): Forel, 1914a: 273; in C. (Orthonotomyrmex): Emery, 1920b: 257. Arnold, 1924: 714 (s.). Senior synonym of ledieui: Emery, 1920c: 29. Currently subspecies of cubangensis: Santschi, 1930b: 81 (in text).

Formicinae Camponotini Camponotus natalensis diabolus TRUE TRUE FALSE

diabolus. Camponotus natalensis r. diabolus Forel, 1879a: 81 (s.w.) SOUTH AFRICA. Combination in C. (Myrmoturba): Forel, 1914a: 267; in C. (Tanaemyrmex): Emery, 1925b: 103. Raised to species: Dalla Torre, 1893: 228; Forel, 1914a: 267. Currently subspecies of natalensis: Emery, 1896d: 373; Wheeler, W.M. 1922a: 961; Emery, 1925b: 103. See also: Arnold, 1922: 646.

Formicinae Camponotini Camponotus arminius bicontractus TRUE TRUE FALSE

bicontractus. Camponotus (Orthonotomyrmex) arminius var. bicontractus Wheeler, W.M. 1922a: 971 (w.) SOUTH AFRICA. [First available use of Camponotus (Orthonotomyrmex) barbarossa r. arminius var. bicontracta Forel, 1914d: 256; unavailable name.] Arnold, 1924: 721 (s.). Combination in C. (Myrmamblys): Emery, 1925b: 142; in C. (Myrmopelta): Santschi, 1926c: 603 (misspelled as biconstrictus).

Formicinae Camponotini Camponotus thraso agricola TRUE TRUE FALSE

agricola. Camponotus maculatus subsp. agricola Forel, 1910e: 454 (s.w.) SOUTH AFRICA. Combination in C. (Tanaemyrmex): Emery, 1925b: 92. Subspecies of maculatus: Arnold, 1922: 647. Currently subspecies of thraso: Emery, 1920c: 6; Emery, 1925b: 92.

Formicinae Camponotini Camponotus auropubens absalon TRUE TRUE FALSE

absalon. Camponotus (Myrmotrema) auropubens var. absalon Santschi, 1915c: 274 (s.w.q.) SOUTH AFRICA.

Formicinae Camponotini Camponotus natalensis corvus TRUE TRUE FALSE

corvus. Camponotus natalensis r. corvus Forel, 1879a: 79 (s.w.q.m.) SOUTH AFRICA. Combination in C. (Tanaemyrmex): Emery, 1925b: 103. Raised to species: Dalla Torre, 1893: 227. Currently subspecies of natalensis: Emery, 1895h: 51; Arnold, 1922: 654; Wheeler, W.M. 1922a: 960.

Formicinae Camponotini Camponotus olivieri infelix TRUE TRUE FALSE

infelix. Camponotus (Myrmotrema) olivieri st. infelix Santschi, 1915c: 270 (s.w.) SOUTH AFRICA. [First available use of Camponotus (Myrmotrema) foraminosus r. lemma var. infelix Forel, 1914d: 254; unavailable name.]

Formicinae Camponotini Camponotus niveosetosus irredux TRUE TRUE FALSE

irredux. Camponotus irredux Forel, 1910f: 30 (s.w.) SOUTH AFRICA. Combination in C. (Myrmopiromis): Wheeler, W.M. 1922a: 987. Currently subspecies of niveosetosus: Forel, 1910e: 451; Arnold, 1922: 659.

Formicinae Camponotini Camponotus petersii janus TRUE TRUE FALSE

janus. Camponotus petersi var. janus Forel, 1911e: 228 (w.) SOUTH AFRICA. Combination in C. (Myrmosericus): Emery, 1925b: 105.

Formicinae Camponotini Camponotus hova pictiventris TRUE TRUE FALSE

pictiventris. Camponotus maculatus subsp. pictiventris Mayr, 1901b: 28 (s.w.) SOUTH AFRICA. Combination in C. (Tanaemyrmex): Emery, 1925b: 86. Subspecies of maculatus: Arnold, 1922: 629; Wheeler, W.M. 1922a: 958. Currently subspecies of hova: Emery, 1920c: 6; Emery, 1925b: 86.

Formicinae Camponotini Camponotus natalensis politiceps TRUE TRUE FALSE

politiceps. Camponotus (Myrmoturba) natalensis var. politiceps Santschi, 1914e: 38 (w.m.) SOUTH AFRICA. Combination in C. (Tanaemyrmex): Emery, 1925b: 103.

Formicinae Camponotini Camponotus nasutus quinquedentatus TRUE TRUE FALSE

quinquedentatus. Camponotus nasutus var. quinquedentata Forel, 1910e: 451 (w.) SOUTH AFRICA. Combination in C. (Myrmamblys): Wheeler, W.M. 1922a: 990.

Myrmicinae Solenopsidini Carebara africana TRUE TRUE FALSE

africana. Aeromyrma africana Forel, 1910f: 15 (s.) BOTSWANA. Combination in Oligomyrmex (Aeromyrma): Emery, 1924d: 216; in Carebara: Fernández, 2004a: 235.

Myrmicinae Solenopsidini Carebara arnoldiella TRUE TRUE FALSE

arnoldiella. Oligomyrmex (Aeromyrma) arnoldiella Santschi, 1919a: 340, fig. 3 (s.w.) SOUTH AFRICA. Combination in Aeromyrma: Wheeler, W.M. 1922a: 882; in Oligomyrmex: Ettershank, 1966: 123; in Carebara: Fernández, 2004a: 235.

Myrmicinae Solenopsidini Carebara donisthorpei TRUE TRUE FALSE

donisthorpei. Oligomyrmex donisthorpei Weber, 1950a: 7, fig. (s.w.) CENTRAL AFRICAN REPUBLIC. Combination in Carebara: Fernández, 2004a: 235.

Myrmicinae Solenopsidini Carebara junodi TRUE TRUE FALSE

junodi. Carebara junodi Forel, 1904a: 154 (q.) SOUTH AFRICA. Forel, 1909b: 53 (m.); Forel, 1913b: 336 (w.). Subspecies of vidua: Forel, 1909b: 53; Forel, 1913b: 336; Arnold, 1916: 252. Revived status as species: Santschi, 1914d: 364 (in key); Wheeler, W.M. 1922a: 883; Emery, 1924d: 220; Santschi, 1935a: 263.

Myrmicinae Solenopsidini Carebara perpusilla TRUE TRUE FALSE

3 subspecies

perpusilla. Pheidologeton perpusillum Emery, 1895h: 26, pl. 2, figs. 8-11 (s.w.) SOUTH AFRICA. Combination in P. (Aneleus): Emery, 1900c: 327; in Aneleus: Santschi, 1914b: 77; in Oligomyrmex: Ettershank, 1966: 124; in Carebara: Fernández, 2004a: 235. Current subspecies: nominal plus arnoldiana, concedens, spinosa. See also: Arnold, 1916: 254.

Myrmicinae Solenopsidini Carebara robertsoni TRUE TRUE FALSE

robertsoni. Paedalgus robertsoni Bolton & Belshaw, 1993: 188, fig. 1 (w.q.) SOUTH AFRICA. Combination in Carebara: Fernández, 2004a: 235.

Myrmicinae Solenopsidini Carebara semilaevis TRUE TRUE FALSE

semilaevis. Solenopsis semilaevis Mayr, 1901b: 15 (w.) SOUTH AFRICA. Combination in Oligomyrmex (Aeromyrma): Emery, 1915c: 59 (in text); in Aeromyrma: Wheeler, W.M. 1922a: 882; in Oligomyrmex: Ettershank, 1966: 124; in Carebara: Fernández, 2004a: 235. Senior synonym of hewitti: Arnold, 1926: 236.

Myrmicinae Solenopsidini Carebara traegaordhi TRUE TRUE FALSE

traegaordhi. Aeromyrma traegaordhi Santschi, 1914e: 20, fig. 2 (w.) SOUTH AFRICA. Combination in Oligomyrmex (Aeromyrma): Santschi, 1919a: 338; in Carebara: Fernández, 2004a: 235. Subspecies of semilaevis: Santschi, 1917b: 285. Revived status as species: Santschi, 1919a: 338; Emery, 1924d: 216; Arnold, 1926: 233.

Myrmicinae Solenopsidini Carebara vidua TRUE TRUE FALSE

1 subspecies

vidua. Carebara vidua Smith, F. 1858b: 179 (m.) SOUTH AFRICA. Mayr, 1866b: 904 (q.); Forel, 1901g: 392 (w.). Senior synonym of colossus, dux: Mayr, 1866b: 904; Santschi, 1913c: 307; of abdominalis: Forel, 1913b: 337; Arnold, 1916: 252; Wheeler, W.M. 1922a: 884. See also: Arnold, 1916: 249; Robertson & Villet, 1989: 121.  Current subspecies: nominal plus fur.

Formicinae Formicini Cataglyphis albicans TRUE TRUE FALSE

6 subspecies

albicans. Formica albicans Roger, 1859: 235 (w.) NORTH AFRICA. André, 1882b: 174 (m.); Forel, 1894d: 11 (q.); Santschi, 1921d: 76 (gynandromorph). Combination in Cataglyphis: Mayr, 1862: 701; Santschi, 1912c: 149; Menozzi, 1922b: 332; Santschi, 1929b: 37; in Myrmecocystus: Emery, 1891b: 17; in Myrmecocystus (Cataglyphis): Karavaiev, 1924: 305. Subspecies of viatica: Emery & Forel, 1879: 449. Revived status as species: André, 1882b: 168; Emery, 1891b: 17; Emery, 1906d: 51. Senior synonym of armena: Radchenko, 1997c: 425. Current subspecies: nominal plus cubica, fezzanensis, franchettii, kairuana, opaca, targuia.

Formicinae Formicini Cataglyphis bicolor TRUE TRUE FALSE

7 subspecies

bicolor. Formica bicolor Fabricius, 1793: 351 (m.) NORTH AFRICA. Wheeler, G.C. & Wheeler, J. 1980: 538 (l.). Combination in Cataglyphis: Roger, 1863b: 12; Mayr, 1863: 402; Emery, 1912f: 98; Emery, 1915g: 22; in Myrmecocystus: Emery & Forel, 1879: 449. Junior synonym of viatica: Smith, F. 1861a: 32; Nasonov, 1889: 14. Revived from synonymy: Emery & Forel, 1879: 449; Emery, 1906d: 58; Forel, 1907e: 15. Subspecies of viatica: Forel, 1907e: 15. Revived status as species: Karavaiev, 1912b: 590; Emery, 1925b: 264; Santschi, 1929b: 41. Senior synonym of fairmairei: Santschi, 1929b: 41; of megalocola: Mayr, 1855: 382; Santschi, 1929b: 41; of rotundinodis: Santschi, 1929b: 41. Biogeography of bicolor group: Wehner, R., Wehner, S. & Agosti, 1994: 163. Current subspecies: nominal plus adusta, basalis, congolensis, pubens, rufiventris, seticornis, sudanica.

Myrmicinae Cataulacini Cataulacus fricatidorsus TRUE TRUE FALSE

fricatidorsus. Cataulacus otii st. fricatidorsus Santschi, 1914e: 26 (w.) SOUTH AFRICA. Raised to species: Bolton, 1974a: 42.

Myrmicinae Cataulacini Cataulacus guineensis TRUE TRUE FALSE

guineensis. Cataulacus guineensis Smith, F. 1853: 225, pl. 20, fig. 5 (w.) "TROPICAL WEST AFRICA". Santschi, 1910c: 390 (q.); Wheeler, W.M. 1922a: 199 (m.). Senior synonym of sulcatus: Forel, 1910e: 421; of alenensis, fernandensis, parallelus, sulcatus, sulcinodis: Bolton, 1974a: 55.

Myrmicinae Cataulacini Cataulacus intrudens TRUE TRUE FALSE

intrudens. Meranoplus intrudens Smith, F. 1876d: 609, pl. 11, fig. 7 (w.q.m.) SOUTH AFRICA. Combination in Cataulacus: Mayr, 1886c: 364. Senior synonym of batonga, baumi, densipunctatus, foveosquamosus, gazanus, hararicus, intermedius, johannae, pseudotrema, rugosus, subrugosus, umbilicatus and material of the unavailable names krugeri, tangana referred here: Bolton, 1974a: 43; of bulawayensis: Bolton, 1995b: 138.

Myrmicinae Cataulacini Cataulacus micans TRUE TRUE FALSE

micans. Cataulacus rugosus subsp. micans Mayr, 1901b: 27 (w.q.m.) SOUTH AFRICA. Raised to species: Forel, 1914d: 219. Senior synonym of tristiculus: Bolton, 1974a: 46.

Myrmicinae Cataulacini Cataulacus traegaordhi TRUE TRUE FALSE

traegaordhi. Cataulacus traegaordhi Santschi, 1914e: 24, fig. 3 (w.q.m.) SOUTH AFRICA. Junior synonym of pygmaeus: Bolton, 1974a: 48. Revived from synonymy and senior synonym of marleyi, suddensis, ugandensis: Bolton, 1982: 359.

Ponerinae Ponerini Centromyrmex ereptor TRUE TRUE FALSE

ereptor. Centromyrmex ereptor Bolton & Fisher, 2008c: 18, figs. 13, 14 (w.) CENTRAL AFRICAN REPUBLIC.

Cerapachyinae Cerapachyini Cerapachys arnoldi TRUE TRUE FALSE

arnoldi. Cerapachys arnoldi Forel, 1914d: 211 (w.) SOUTH AFRICA. Senior synonym of hewitti: Brown, 1975: 22. See also: Arnold, 1926: 191.

Cerapachyinae Cerapachyini Cerapachys braunsi TRUE TRUE FALSE

braunsi. Phyracaces braunsi Emery, 1902c: 27 (w.) SOUTH AFRICA. Combination in Cerapachys: Brown, 1975: 22.

Cerapachyinae Cerapachyini Cerapachys faurei TRUE TRUE FALSE

faurei. Cerapachys faurei Arnold, 1949: 262, fig. 2 (w.) SOUTH AFRICA.

Cerapachyinae Cerapachyini Cerapachys natalensis TRUE TRUE FALSE

natalensis. Cerapachys cribrinodis var. natalensis Forel, 1901f: 335 (w.) SOUTH AFRICA. [Also described as new by Forel, 1910e: 423.] Raised to species: Brown, 1975: 23.

Cerapachyinae Cerapachyini Cerapachys peringueyi TRUE TRUE FALSE

peringueyi. Parasyscia peringueyi Emery, 1886: 360 (w.) SOUTH AFRICA. Emery, 1895j: 757 (q.). Combination in Cerapachys: Emery, 1902c: 24. Senior synonym of latiuscula: Brown, 1975: 23. See also: Arnold, 1915: 13.

Cerapachyinae Cerapachyini Cerapachys validus TRUE TRUE FALSE

validus. Cerapachys validus Arnold, 1960a: 79, fig. 1 (w.) SOUTH AFRICA.

Cerapachyinae Cerapachyini Cerapachys wroughtoni TRUE TRUE FALSE

wroughtoni. Cerapachys wroughtoni Forel, 1910e: 422 (w.) SOUTH AFRICA. Senior synonym of rhodesiana, roberti: Brown, 1975: 24.

Myrmicinae Crematogastrini Crematogaster africana TRUE TRUE FALSE

10 subspecies

africana. Crematogaster africana Mayr, 1895: 142 (w.) CAMEROUN. Santschi, 1937g: 78 (q.). Combination in C. (Atopogyne): Santschi, 1916b: 501. Subspecies of buchneri: Forel, 1913a: 125. Revived status as species: Santschi, 1918d: 183. Current subspecies: nominal plus alligatrix, biemarginata, camena, polymorphica, schumanni, stanleyi, stolonis, thoracica, tibialis, variegata.

Myrmicinae Crematogastrini Crematogaster arnoldi TRUE TRUE FALSE

1 subspecies

arnoldi. Crematogaster arnoldi Forel, 1914d: 238 (w.) SOUTH AFRICA. Combination in C. (Acrocoelia): Emery, 1922e: 145; in C. (Crematogaster): Bolton, 1995b: 166. Subspecies of godfreyi: Arnold, 1920a: 504. Revived status as species: Santschi, 1926b: 214. Current subspecies: nominal plus loveridgei.

Myrmicinae Crematogastrini Crematogaster capensis TRUE TRUE FALSE

2 subspecies

capensis. Crematogaster capensis Mayr, 1862: 764 (w.q.) SOUTH AFRICA. Combination in C. (Crematogaster): Wheeler, W.M. 1922a: 830; in C. (Acrocoelia): Emery, 1922e: 145; in C. (Crematogaster): Bolton, 1995b: 166. Current subspecies: nominal plus calens, tropicorum.

Myrmicinae Crematogastrini Crematogaster castanea TRUE TRUE FALSE

19 subspecies

castanea. Crematogaster castaneus Smith, F. 1858b: 136, pl. 9, fig. 2 (w.q.) SOUTH AFRICA (not Albania). Combination in C. (Crematogaster): Wheeler, W.M. 1922a: 830; in C. (Acrocoelia): Emery, 1922e: 145; in C. (Crematogaster): Bolton, 1995b: 166. Subspecies of tricolor: Forel, 1911f: 279; Santschi, 1913a: 412 (in key). Revived status as species: Emery, 1915g: 12; Arnold, 1920a: 488; Santschi, 1920d: 378. Senior synonym of decolor: Forel, 1907g: 80. Current subspecies: nominal plus adusta, analis, aquila, arborea, bruta, busschodtsi, durbanensis, ferruginea, hararica, insidiosa, inversa, mediorufa, museisapientiae, rufimembrum, rufonigra, simia, tricolor, ulugurensis, yambatensis.

Myrmicinae Crematogastrini Crematogaster cicatriculosa TRUE TRUE FALSE

cicatriculosa. Crematogaster cicatriculosa Roger, 1863a: 207 (w.) SOUTH AFRICA. Mayr, 1866b: 901 (q.). Combination in C. (Crematogaster): Wheeler, W.M. 1922a: 836; in C. (Acrocoelia): Emery, 1922e: 146; in C. (Crematogaster): Bolton, 1995b: 166.

Myrmicinae Crematogastrini Crematogaster constructor TRUE TRUE FALSE

1 subspecies

constructor. Crematogaster constructor Emery, 1895h: 29, pl. 2, figs. 12, 13 (w.) SOUTH AFRICA. [Also described as new by Emery, in Mayr, 1895: 135 (footnote).] Combination in C. (Crematogaster): Wheeler, W.M. 1922a: 836; in C. (Acrocoelia): Emery, 1922e: 146; in C. (Crematogaster): Bolton, 1995b: 166. Current subspecies: nominal plus kirbyella.

Myrmicinae Crematogastrini Crematogaster desperans TRUE TRUE FALSE

desperans. Crematogaster gabonensis var. desperans Forel, 1914d: 233 (w.) SOUTH AFRICA. Combination in C. (Atopogyne): Wheeler, W.M. 1922a: 854; in C. (Acrocoelia): Emery, 1922e: 146; in C. (Crematogaster): Bolton, 1995b: 166. Subspecies of bulawayensis: Arnold, 1920a: 523; Wheeler, W.M. 1922a: 854. Raised to species: Santschi, 1918d: 185; Emery, 1922e: 146.

Myrmicinae Crematogastrini Crematogaster hottentota TRUE TRUE FALSE

1 subspecies

hottentota. Crematogaster kneri subsp. hottentota Emery, 1899e: 482 (w.) SOUTH AFRICA. Combination in C. (Sphaerocrema): Wheeler, W.M. 1922a: 847. Raised to species: Santschi, 1925h: 161 (footnote); Santschi, 1926b: 224. Current subspecies: nominal plus bassuto.

Myrmicinae Crematogastrini Crematogaster laurenti TRUE TRUE FALSE

1 subspecies

laurenti. Crematogaster africana subsp. laurenti Forel, 1909b: 69 (w.) DEMOCRATIC REPUBLIC OF CONGO. Combination in C. (Atopogyne): Wheeler, W.M. 1922a: 852. Subspecies of buchneri: Forel, 1913a: 125; Forel, 1913c: 352; Forel, 1916: 410; of africana: Wheeler, W.M. 1922a: 852. Raised to species: Emery, 1922e: 155; Santschi, 1935a: 262. Current subspecies: nominal plus zeta.

Myrmicinae Crematogastrini Crematogaster magitae TRUE TRUE FALSE

magitae. Crematogaster (Oxygyne) magitae Forel, 1910f: 9 (q.) WEST AFRICA. Combination in C. (Nematocrema): Emery, 1922e: 157.

Myrmicinae Crematogastrini Crematogaster melanogaster TRUE TRUE FALSE

1 subspecies

melanogaster. Crematogaster arborea subsp. melanogaster Emery, 1895h: 29 (w.q.) SOUTH AFRICA. Combination in C. (Crematogaster): Wheeler, W.M. 1922a: 840; in C. (Acrocoelia): Emery, 1922e: 147; in C. (Crematogaster): Bolton, 1995b: 166. Raised to species: Wheeler, W.M. 1922a: 840. Current subspecies: nominal plus homonyma.

Myrmicinae Crematogastrini Crematogaster monticola TRUE TRUE FALSE

monticola. Crematogaster monticola Arnold, 1920a: 517, fig. 40 (w.) SOUTH AFRICA. Combination in C. (Crematogaster): Wheeler, W.M. 1922a: 841.

Myrmicinae Crematogastrini Crematogaster natalensis TRUE TRUE FALSE

2 subspecies

natalensis. Crematogaster sordidula subsp. natalensis Forel, 1910e: 431 (w.q.m.) SOUTH AFRICA. Combination in C. (Orthocrema): Wheeler, W.M. 1922a: 850. Raised to species: Emery, 1912e: 668. Subspecies of sordidula: Arnold, 1920a: 539; Wheeler, W.M. 1922a: 850. Revived status as species: Emery, 1922e: 131; Santschi, 1926b: 212; Menozzi, 1930b: 85. Current subspecies: nominal plus braunsi, dulcis.

Myrmicinae Crematogastrini Crematogaster opaciceps TRUE TRUE FALSE

3 subspecies

opaciceps. Crematogaster opaciceps Mayr, 1901b: 12 (w.) SOUTH AFRICA. Combination in C. (Crematogaster): Wheeler, W.M. 1922a: 842; in C. (Acrocoelia): Emery, 1922e: 148; in C. (Crematogaster): Bolton, 1995b: 166. Current subspecies: nominal plus cacodaemon, clepens, defleta.

Myrmicinae Crematogastrini Crematogaster orobia TRUE TRUE FALSE

orobia. Crematogaster orobia Santschi, 1919a: 343 (w.) SOUTH AFRICA. Combination in C. (Crematogaster): Wheeler, W.M. 1922a: 842.

Myrmicinae Crematogastrini Crematogaster oscaris TRUE TRUE FALSE

oscaris. Crematogaster (Oxygyne) oscaris Forel, 1910f: 7, pl. 1, figs. 5, 6 (w.q.m.) SOUTH AFRICA.

Myrmicinae Crematogastrini Crematogaster peringueyi TRUE TRUE FALSE

4 subspecies

peringueyi. Crematogaster peringueyi Emery, 1895h: 27, figs. 16, 17 (w.q.m.) SOUTH AFRICA. Combination in C. (Crematogaster): Wheeler, W.M. 1922a: 842; in C. (Acrocoelia): Emery, 1922e: 148; in C. (Crematogaster): Bolton, 1995b: 166. Current subspecies: nominal plus angustior, cacochyma, dentulata, gedeon.

Myrmicinae Crematogastrini Crematogaster schultzei TRUE TRUE FALSE

schultzei. Crematogaster schultzei Forel, 1910f: 6 (w.q.) SOUTH AFRICA. Combination in C. (Crematogaster): Wheeler, W.M. 1922a: 843; in C. (Acrocoelia): Emery, 1922e: 148; in C. (Crematogaster): Bolton, 1995b: 166.

Myrmicinae Crematogastrini Crematogaster theta TRUE TRUE FALSE

theta. Crematogaster africana var. theta Forel, 1911f: 278 (w.) DEMOCRATIC REPUBLIC OF CONGO. Forel, 1916: 410 (q.). Combination in C. (Atopogyne): Wheeler, W.M. 1922a: 857. Raised to species: Wheeler, W.M. 1922a: 857; Emery, 1922e: 155; Santschi, 1935a: 262.

Myrmicinae Crematogastrini Crematogaster transvaalensis TRUE TRUE FALSE

1 subspecies

transvaalensis. Crematogaster transvaalensis Forel, 1894b: 96 (w.) SOUTH AFRICA. Combination in C. (Orthocrema): Wheeler, W.M. 1922a: 851. Current subspecies: nominal plus hammi.

Myrmicinae Crematogastrini Crematogaster gerstaeckeri zulu TRUE TRUE FALSE

zulu. Crematogaster (Atopogyne) bulawayensis var. zulu Wheeler, W.M. 1922a: 855 (w.q.) SOUTH AFRICA. [First available use of Crematogaster sjostedti st. bulawayensis var. zulu Santschi, 1920b: 15; unavailable name.] Subspecies of bulawayensis: Arnold, 1926: 289. Currently subspecies of gerstaeckeri: Bolton, 1995b: 165.

Myrmicinae Crematogastrini Crematogaster laurenti zeta TRUE TRUE FALSE

zeta. Crematogaster (Atopogyne) laurenti var. zeta Emery, 1922e: 155 (w.) DEMOCRATIC REPUBLIC OF CONGO. [First available use of Crematogaster africana subsp. laurenti var. zeta Forel, 1909b: 70; unavailable name.] Currently subspecies of laurenti: Santschi, 1937g: 79.

Myrmicinae Crematogastrini Crematogaster kohli winkleri TRUE TRUE FALSE

winkleri. Crematogaster africana var. winkleri Forel, 1909b: 69 (w.) CAMEROUN. Forel, 1913a: 124 (q.). Combination in C. (Atopogyne): Wheeler, W.M. 1922a: 158; Santschi, 1937g: 80. Subspecies of buchneri: Forel, 1913a: 124; Forel, 1913b: 324; Forel, 1913e: 670; of africana: Wheeler, W.M. 1922a: 158. Currently subspecies of kohli: Santschi, 1937g: 80.

Myrmicinae Crematogastrini Crematogaster liengmei weitzeckeri TRUE TRUE FALSE

weitzeckeri. Crematogaster weitzeckeri Emery, 1895h: 28, pl. 2, figs. 14, 15 (w.) SOUTH AFRICA. Combination in C. (Decacrema): Forel, 1910a: 18; Forel, 1910f: 9. Senior synonym of kloofensis: Arnold, 1920a: 549. Currently subspecies of liengmei: Forel, 1910a: 18; Santschi, 1914e: 21; Arnold, 1920a: 548.

Myrmicinae Crematogastrini Crematogaster africana variegata TRUE TRUE FALSE

variegata. Crematogaster africana var. variegata Mayr, 1902: 294 (w.) CAMEROUN. Combination in C. (Atopogyne): Wheeler, W.M. 1922a: 852.

Myrmicinae Crematogastrini Crematogaster africana tibialis TRUE TRUE FALSE

tibialis. Crematogaster (Atopogyne) africana subsp. tibialis Santschi, in Wheeler, W.M. 1922a: 157 (w.) DEMOCRATIC REPUBLIC OF CONGO.

Myrmicinae Crematogastrini Crematogaster africana thoracica TRUE TRUE FALSE

thoracica. Crematogaster (Atopogyne) africana st. thoracica Santschi, 1921c: 118 (w.) DEMOCRATIC REPUBLIC OF CONGO.

Myrmicinae Crematogastrini Crematogaster africana stolonis TRUE TRUE FALSE

stolonis. Crematogaster (Atopogyne) africana st. stolonis Santschi, 1937g: 78, fig. 4 (w.) DEMOCRATIC REPUBLIC OF CONGO.

Myrmicinae Crematogastrini Crematogaster africana stanleyi TRUE TRUE FALSE

stanleyi. Crematogaster (Atopogyne) africana var. stanleyi Wheeler, W.M. 1922a: 852 (w.) CONGO. [First available use of Crematogaster buchneri st. africana var. stanleyi Santschi, 1915c: 253; unavailable name.] [Crematogaster africana var. stanleyi Forel, 1911f: 278. Nomen nudum.] Santschi, 1937g: 79 (q.).

Myrmicinae Crematogastrini Crematogaster africana schumanni TRUE TRUE FALSE

schumanni. Crematogaster africana subsp. schumanni Mayr, 1895: 144 (w.) CAMEROUN. Combination in C. (Atopogyne): Wheeler, W.M. 1922a: 852.

Myrmicinae Crematogastrini Crematogaster castanea rufonigra TRUE TRUE FALSE

rufonigra. Crematogaster tricolor subsp. rufonigra Emery, 1895h: 27 (w.) SOUTH AFRICA. Arnold, 1920a: 492 (q.m.). Combination in C. (Acrocoelia): Emery, 1922e: 146. Currently subspecies of castanea: Santschi, 1916b: 502.

Myrmicinae Crematogastrini Crematogaster gerstaeckeri rufescens TRUE TRUE FALSE

rufescens. Crematogaster (Crematogaster) sjostedti var. rufescens Wheeler, W.M. 1922a: 844 (w.) SOUTH AFRICA. [First available use of Crematogaster sjostedti st. bulawayensis var. rufescens Santschi, 1919b: 236; unavailable name.] Subspecies of bulawayensis: Arnold, 1926: 289. Currently subspecies of gerstaeckeri: Bolton, 1995b: 161.

Myrmicinae Crematogastrini Crematogaster acaciae generosa TRUE TRUE FALSE

generosa. Crematogaster acaciae var. generosa Santschi, 1919a: 343 (w.q.) SOUTH AFRICA.

Myrmicinae Crematogastrini Crematogaster peringueyi gedeon TRUE TRUE FALSE

gedeon. Crematogaster peringueyi var. gedeon Arnold, 1920a: 512 (w.) SOUTH AFRICA. [First available use of Crematogaster peringueyi r. cacodaemon var. gedeon Forel, 1915c: 341; unavailable name.] Combination in C. (Acrocoelia): Emery, 1922e: 148.

Myrmicinae Crematogastrini Crematogaster castanea durbanensis TRUE TRUE FALSE

durbanensis. Crematogaster ferruginea var. durbanensis Forel, 1914d: 234 (w.) SOUTH AFRICA. Arnold, 1920a: 491 (q.). Combination in C. (Acrocoelia): Emery, 1922e: 145. Currently subspecies of castanea: Bolton, 1995b: 152.

Myrmicinae Crematogastrini Crematogaster natalensis dulcis TRUE TRUE FALSE

dulcis. Crematogaster (Orthocrema) natalensis var. dulcis Santschi, 1926b: 212 (w.) SOUTH AFRICA.

Myrmicinae Crematogastrini Crematogaster peringueyi dentulata TRUE TRUE FALSE

dentulata. Crematogaster peringueyi r. dentulata Stitz, 1923: 156 (w.) SOUTH AFRICA.

Myrmicinae Crematogastrini Crematogaster opaciceps defleta TRUE TRUE FALSE

defleta. Crematogaster opaciceps subsp. defletus Forel, 1910e: 434 (w.) SOUTH AFRICA. Combination in C. (Acrocoelia): Emery, 1922e: 148.

Myrmicinae Crematogastrini Crematogaster neuvillei cooperi TRUE TRUE FALSE

cooperi. Crematogaster neuvillei r. cooperi Forel, 1914d: 233 (w.) SOUTH AFRICA. Arnold, 1920a: 498 (q.). Combination in C. (Acrocoelia): Emery, 1922e: 148.

Myrmicinae Crematogastrini Crematogaster santschii clymene TRUE TRUE FALSE

clymene. Crematogaster santschii var. clymene Forel, 1915c: 337 (w.) SOUTH AFRICA. Combination in C. (Oxygyne): Wheeler, W.M. 1922a: 858; in C. (Acrocoelia): Emery, 1922e: 148.

Myrmicinae Crematogastrini Crematogaster delagoensis acutidens TRUE TRUE FALSE

acutidens. Crematogaster delagoense var. acutidens Arnold, 1944: 7, fig. 7 (w.) SOUTH AFRICA.

Myrmicinae Crematogastrini Crematogaster africana alligatrix TRUE TRUE FALSE

alligatrix. Crematogaster buchneri subsp. alligatrix Forel, 1911e: 271 (w.) NIGERIA. Combination in C. (Atopogyne): Wheeler, W.M. 1922a: 852. Currently subspecies of africana: Wheeler, W.M. 1922a: 852.

Myrmicinae Crematogastrini Crematogaster peringueyi angustior TRUE TRUE FALSE

angustior. Crematogaster peringueyi var. angustior Arnold, 1920a: 510, fig. 40 (w.q.m.) SOUTH AFRICA.

Myrmicinae Crematogastrini Crematogaster castanea arborea TRUE TRUE FALSE

arborea. Crematogaster arboreus Smith, F. 1858b: 138, pl. 14 (nest) (w.) SOUTH AFRICA. Combination in C. (Acrocoelia): Emery, 1922e: 145. Junior synonym of tricolor: Mayr, 1895: 136 (footnote); Arnold, 1920a: 506 (note that arborea has priority); of castanea: Wheeler, W.M. 1922a: 830. Revived from synonymy as subspecies of castanea: Emery, 1922e: 145. Senior synonym of decolorata: Santschi, 1930b: 69. Currently subspecies of castanea: Santschi, 1930b: 69.

Myrmicinae Crematogastrini Crematogaster africana biemarginata TRUE TRUE FALSE

biemarginata. Crematogaster africana var. biemarginata Forel, 1910e: 433 (w.) CAMEROUN. Combination in C. (Atopogyne): Wheeler, 1922a: 852.

Myrmicinae Crematogastrini Crematogaster natalensis braunsi TRUE TRUE FALSE

braunsi. Crematogaster (Orthocrema) natalensis var. braunsi Emery, 1922e: 131 (w.) SOUTH AFRICA. [First available use of Crematogaster sordidula subsp. natalensis var. braunsi Forel, 1911f: 277; unavailable name.]

Myrmicinae Crematogastrini Crematogaster castanea bruta TRUE TRUE FALSE

bruta. Crematogaster castanea st. bruta Santschi, 1926b: 213 (w.) SOUTH AFRICA. [First available use of Crematogaster tricolor st. ferruginea var. bruta Santschi, 1913a: 413; unavailable name.]

Myrmicinae Crematogastrini Crematogaster amita bushimana TRUE TRUE FALSE

bushimana. Crematogaster (Sphaerocrema) amita var. bushimana Santschi, 1926b: 226 (w.q.) SOUTH AFRICA.

Myrmicinae Crematogastrini Crematogaster peringueyi cacochyma TRUE TRUE FALSE

cacochyma. Crematogaster peringueyi r. cacochyma Forel, 1914d: 235 (w.q.) SOUTH AFRICA. Combination in C. (Acrocoelia): Emery, 1922e: 512.

Myrmicinae Crematogastrini Crematogaster opaciceps cacodaemon TRUE TRUE FALSE

cacodaemon. Crematogaster peringueyi r. cacodaemon Forel, 1914d: 234 (w.q.m.) SOUTH AFRICA. Combination in C. (Acrocoelia): Emery, 1922e: 148. Currently subspecies of opaciceps: Santschi, 1926b: 216.

Myrmicinae Crematogastrini Crematogaster liengmei caculata TRUE TRUE FALSE

caculata. Crematogaster peringueyi r. caculata Forel, 1915c: 341 (w.) SOUTH AFRICA. Combination in C. (Decacrema): Arnold, 1920a: 550; Wheeler, W.M. 1922a: 861; in C. (Acrocoelia): Emery, 1922e: 148. Currently subspecies of liengmei: Arnold, 1920a: 550; Wheeler, W.M. 1922a: 861.

Myrmicinae Crematogastrini Crematogaster amita caffra TRUE TRUE FALSE

caffra. Crematogaster (Sphaerocrema) amita var. caffra Santschi, 1926b: 226 (w.q.m.) SOUTH AFRICA. [First available use of Crematogaster (Sphaerocrema) kneri st. amita var. caffra Santschi, in Forel, 1922: 94 (footnote); unavailable name.]

Myrmicinae Crematogastrini Crematogaster africana camena TRUE TRUE FALSE

camena. Crematogaster (Atopogyne) africana var. camena Wheeler, W.M. 1922a: 852 (w.) DEMOCRATIC REPUBLIC OF CONGO. [First available use of Crematogaster buchneri r. africana var. camena Forel, 1916: 410; unavailable name.]

Myrmicinae Crematogastrini Crematogaster gerstaeckeri godefreyi TRUE TRUE FALSE

godefreyi. Crematogaster foraminiceps r. godefreyi Forel, 1914d: 237 (w.) SOUTH AFRICA. Combination in C. (Crematogaster): Wheeler, W.M. 1922a: 839; in C. (Acrocoelia): Emery, 1922e: 148. Subspecies of vulcania: Forel, 1916: 406; of sjostedti: Santschi, 1918d: 184; Emery, 1922e: 148. Raised to species: Arnold, 1920a: 503; Wheeler, W.M. 1922a: 839. Currently subspecies of gerstaeckeri: Santschi, 1930b: 69. Senior synonym of carininotum: Santschi, 1918d: 184.

Myrmicinae Crematogastrini Crematogaster melanogaster homonyma TRUE TRUE FALSE

homonyma. Crematogaster (Acrocoelia) melanogaster var. homonyma Emery, 1922e: 147 (w.q.) SOUTH AFRICA.

Myrmicinae Crematogastrini Crematogaster censor junodi TRUE TRUE FALSE

junodi. Crematogaster censor r. junodi Forel, 1916: 406 (w.) SOUTH AFRICA. Combination in C. (Acrocoelia): Emery, 1922e: 146.

Myrmicinae Crematogastrini Crematogaster gambiensis krantziana TRUE TRUE FALSE

krantziana. Crematogaster gambiensis var. krantziana Forel, 1914d: 234 (w.m.) SOUTH AFRICA. Arnold, 1920a: 530 (q.). Combination in C. (Sphaerocrema): Wheeler, W.M. 1922a: 846.

Myrmicinae Crematogastrini Crematogaster castanea mediorufa TRUE TRUE FALSE

mediorufa. Crematogaster tricolor var. mediorufa Forel, 1907g: 81 (footnote) (w.m.) SOUTH AFRICA. Forel, 1913a: 124 (q.); Arnold, 1920a: 493 (w.q.m.). Currently subspecies of castanea: Bolton, 1995b: 157.

Myrmicinae Crematogastrini Crematogaster delagoensis merwei TRUE TRUE FALSE

merwei. Crematogaster (Acrocoelia) delagoensis st. merwei Santschi, 1932a: 385 (w.) SOUTH AFRICA.

Myrmicinae Crematogastrini Crematogaster castanea museisapientiae TRUE TRUE FALSE

museisapientiae. Crematogaster ferruginea subsp. museisapientiae Forel, 1911e: 272 (w.q.m.) "Tropical East Africa". Combination in C. (Acrocoelia): Emery, 1922e: 146.  Subspecies of tricolor: Santschi, 1913a: 413 (in key). Currently subspecies of castanea: Wheeler, W.M. 1922a: 833.

Myrmicinae Crematogastrini Crematogaster brunneipennis omniparens TRUE TRUE FALSE

omniparens. Crematogaster brunneipennis r. omniparens Forel, 1914d: 237 (w.) SOUTH AFRICA. Combination in C. (Acrocoelia): Emery, 1922e: 145.

Myrmicinae Crematogastrini Crematogaster africana polymorphica TRUE TRUE FALSE

polymorphica. Crematogaster (Atopogyne) africana subsp. polymorphica Weber, 1943c: 357 (w.) SUDAN.

Myrmicinae Solenopsidini Diplomorium longipenne TRUE TRUE FALSE

longipenne. Diplomorium longipenne Mayr, 1901b: 18 (w.q.) SOUTH AFRICA. See also: Arnold, 1916: 240; Bolton, 1987: 278.

Proceratiinae Proceratiini Discothyrea hewitti TRUE TRUE FALSE

hewitti. Discothyrea hewitti Arnold, 1916: 160 (q.) SOUTH AFRICA.

Proceratiinae Proceratiini Discothyrea traegaordhi TRUE TRUE FALSE

traegaordhi. Discothyrea traegaordhi Santschi, 1914e: 3 (w.) SOUTH AFRICA. See also: Brown, 1958g: 340.

Dorylinae Dorylini Dorylus erraticus TRUE TRUE FALSE

erraticus. Anomma erratica Smith, F. 1865: 71 (w.) AFRICA (not New Guinea as stated; see: Wilson, 1964a: 443). Combination in Dorylus: Emery, 1910b: 15; in D. (Anomma): Donisthorpe, 1932c: 473.

Dorylinae Dorylini Dorylus faurei TRUE TRUE FALSE

faurei. Dorylus faurei Arnold, 1946: 60, fig. 12 (q.) SOUTH AFRICA.

Dorylinae Dorylini Dorylus fulvus TRUE TRUE FALSE

11 subspecies

fulvus. Typhlopone fulva Westwood, 1839: 219, fig. 86 (w.) NORTH AFRICA. Emery, 1895j: 723 (m.). Combination in Dorylus: Dalla Torre, 1893: 10; in D. (Typhlopone): Emery, 1895j: 723. Senior synonym of kirbii: Smith, F. 1858b: 110; Mayr, 1863: 457; of clausii, dahlbomii, shuckardi, spinolae, thwaitsii: Emery, 1895j: 723. Current subspecies: nominal plus badius, crosi, dentifrons, eurous, glabratus, juvenculus, mordax, obscurior, punicus, ruficeps, saharensis.

Dorylinae Dorylini Dorylus furcatus TRUE TRUE FALSE

furcatus. Dichthadia furcata Gerstäcker, 1872: 267 (q.) SOUTH AFRICA. Combination in Dorylus: Emery, 1888b: 345.

Dorylinae Dorylini Dorylus helvolus TRUE TRUE FALSE

1 subspecies

helvolus. Vespa helvola Linnaeus, 1764: 412 (m.) SOUTH AFRICA. Linnaeus, 1767: 968 (w.); Emery, 1888b: 344 (q.). Combination in Dorylus: Fabricius, 1793: 365. Senior synonym of dorylus, europaea, planifrons, punctata: Emery, 1888b: 350; Emery, 1895j: 714. Current subspecies: nominal plus pretoriae. See also: Arnold, 1915: 116.

Dorylinae Dorylini Dorylus attenuatus acuminatus TRUE TRUE FALSE

acuminatus. Dorylus (Alaopone) attenuatus var. acuminata Emery, 1899e: 462 (m.) SOUTH AFRICA.

Dorylinae Dorylini Dorylus attenuatus australis TRUE TRUE FALSE

australis. Dorylus (Alaopone) montanum st. australis Santschi, 1919b: 231 (w.) SOUTH AFRICA. [Also described as new, but as species, by Santschi, 1920d: 362; based on same specimens.] Subspecies of montanus: Wheeler, W.M. 1922a: 750. Currently subspecies of attenuatus: Santschi, 1939a: 147.

Dorylinae Dorylini Dorylus attenuatus bondroiti TRUE TRUE FALSE

bondroiti. Dorylus (Alaopone) montanus var. bondroiti Santschi, 1912b: 162 (m.) SOUTH AFRICA. Subspecies of montanus: Santschi, 1915c: 248. Currently subspecies of attenuatus: Santschi, 1939a: 148.

Dorylinae Dorylini Dorylus fimbriatus poweri TRUE TRUE FALSE

poweri. Dorylus (Rhogmus) fimbriatus var. poweri Forel, 1914d: 217 (w.) SOUTH AFRICA.

Dorylinae Dorylini Dorylus helvolus pretoriae TRUE TRUE FALSE

pretoriae. Dorylus helvolus subsp. pretoriae Arnold, 1946: 59, fig. 11 (w.q.m.) SOUTH AFRICA.

Dorylinae Dorylini Dorylus emeryi pulsi TRUE TRUE FALSE

pulsi. Anomma emeryi var. pulsi Forel, 1904d: 170 (w.) "West Africa". Santschi, 1923e: 273 (s.). Combination in Dorylus (Anomma): Emery, 1910b: 11.

Dorylinae Dorylini Dorylus nigricans rubellus TRUE TRUE FALSE

rubellus. Anomma rubella Savage, 1849: 196 (w.) "West Africa". Combination in Dorylus (Anomma): Emery, 1895j: 710. Subspecies of burmeisteri: Emery, 1892d: 554. Currently subspecies of nigricans: Emery, 1895j: 701; Emery, 1910b: 12; Forel, 1913h: 348.

Ponerinae Ponerini Hypoponera austra TRUE TRUE FALSE

austra. Hypoponera austra Bolton & Fisher, 2011: 27, figs. 7-9 (w.q.,ergatoid q.) SOUTH AFRICA.

Ponerinae Ponerini Hypoponera blanda TRUE TRUE FALSE

blanda. Hypoponera blanda Bolton & Fisher, 2011: 29, figs. 10-12 (w.q.) CENTRAL AFRICAN REPUBLIC.

Ponerinae Ponerini Hypoponera boerorum TRUE TRUE FALSE

boerorum. Ponera coarctata r. boerorum Forel, 1901f: 339 (w.) SOUTH AFRICA. Combination in Hypoponera: Taylor, 1967a: 12. Raised to species: Santschi, 1938b: 78. See also: Bolton & Fisher, 2011: 30.

Ponerinae Ponerini Hypoponera meridia TRUE TRUE FALSE

meridia. Hypoponera meridia Bolton & Fisher, 2011: 70, figs. 73-75 (w.q.) SOUTH AFRICA.

Ponerinae Ponerini Hypoponera natalensis TRUE TRUE FALSE

natalensis. Ponera coarctata st. natalensis Santschi, 1914e: 7 (w.) SOUTH AFRICA. Combination in P. (Hypoponera): Santschi, 1938b: 79; in Hypoponera: Taylor, 1967a: 12. Raised to species: Santschi, 1938b: 79; Taylor, 1967a: 12. See also: Bolton & Fisher, 2011: 74.

Ponerinae Ponerini Hypoponera perparva TRUE TRUE FALSE

perparva. Hypoponera perparva Bolton & Fisher, 2011: 82, figs. 91-93 (w.) SOUTH AFRICA.

Ponerinae Ponerini Hypoponera spei TRUE TRUE FALSE

spei. Ponera spei Forel, 1910e: 423 (w.) SOUTH AFRICA. Combination in Hypoponera: Bolton, 1995b: 216. Senior synonym of fidelis and material of the unavailable name sancta referred here: Bolton & Fisher, 2011: 101. See also: Arnold, 1915: 77.

Ponerinae Ponerini Hypoponera sulcatinasis TRUE TRUE FALSE

sulcatinasis. Ponera sulcatinasis Santschi, 1914e: 4 (w.) SOUTH AFRICA. Combination in Hypoponera: Bolton, 1995b: 216. Senior synonym of devota: Bolton & Fisher, 2011: 103.

Ponerinae Ponerini Hypoponera traegaordhi TRUE TRUE FALSE

traegaordhi. Ponera traegaordhi Santschi, 1914e: 6 (w.) SOUTH AFRICA. Combination in P. (Hypoponera): Santschi, 1938b: 79; in Hypoponera: Bolton, 1995b: 216. See also: Bolton & Fisher, 2011: 108.

Ponerinae Ponerini Hypoponera transvaalensis TRUE TRUE FALSE

transvaalensis. Ponera transvaalensis Arnold, 1947: 132, fig. 2 (w.) SOUTH AFRICA. Combination in Hypoponera: Bolton, 1995b: 216. See also: Bolton & Fisher, 2011: 108.

Formicinae Plagiolepidini Lepisiota capensis TRUE TRUE FALSE

13 subspecies

capensis. Acantholepis capensis Mayr, 1862: 699 (w.) SOUTH AFRICA. Mayr, 1865: 57 (m.); Emery, 1877b: 366 (q.); Wheeler, G.C. & Wheeler, J. 1980: 542 (l.). Combination in Lepisiota: Xu, 1994c: 233. Current subspecies: nominal plus acholli, anceps, guineensis, issore, junodi, laevis, lunaris, minuta, simplex, simplicoides, specularis, subopaciceps, thoth. See also: Bingham, 1903: 316; Arnold, 1920a: 568.

Formicinae Plagiolepidini Lepisiota crinita TRUE TRUE FALSE

crinita. Acantholepis crinita Mayr, 1895: 149 (w.) SOUTH AFRICA. Combination in Lepisiota: Bolton, 1995b: 227.

Formicinae Plagiolepidini Lepisiota capensis simplicoides TRUE TRUE FALSE

simplicoides. Acantholepis capensis var. simplicoides Forel, 1907g: 86 (footnote) (w.) SOUTH AFRICA. Combination in Lepisiota: Bolton, 1995b: 228.

Formicinae Plagiolepidini Lepisiota spinosior natalensis TRUE TRUE FALSE

natalensis. Acantholepis spinosior var. natalensis Arnold, 1920a: 559 (w.) SOUTH AFRICA. Combination in Lepisiota: Bolton, 1995b: 228.

Formicinae Plagiolepidini Lepisiota capensis minuta TRUE TRUE FALSE

minuta. Acantholepis simplex var. minuta Forel, 1916: 438 (w.) SOUTH AFRICA. Combination in Lepisiota: Bolton, 1995b: 228. Currently subspecies of capensis: Arnold, 1920a: 572.

Formicinae Plagiolepidini Lepisiota capensis junodi TRUE TRUE FALSE

junodi. Acantholepis capensis r. junodi Forel, 1916: 438 (w.) SOUTH AFRICA. Combination in Lepisiota: Bolton, 1995b: 227.

Leptanillinae Leptanillini Leptanilla africana TRUE TRUE FALSE

africana. Leptanilla africana Baroni Urbani, 1977c: 468, figs. 34-37, 43 (m.) NIGERIA.

Leptanillinae Leptanillini Leptanilla australis TRUE TRUE FALSE

australis. Leptanilla australis Baroni Urbani, 1977c: 471, figs. 38, 40-42, 44 (m.) SOUTH AFRICA.

Ponerinae Ponerini Leptogenys attenuata TRUE TRUE FALSE

attenuata. Ponera attenuata Smith, F. 1858b: 91 (w.) SOUTH AFRICA. Arnold, 1915: 98 (m.). Combination in Lobopelta: Roger, 1863b: 19; in Leptogenys: Mayr, 1895: 124. Senior synonym of jaegerskjoeldi: Arnold, 1926: 210. See also: Bolton, 1975a: 274.

Ponerinae Ponerini Leptogenys castanea TRUE TRUE FALSE

castanea. Lobopelta castanea Mayr, 1862: 734 (w.) SOUTH AFRICA. Wheeler, G.C. & Wheeler, J. 1986c: 89 (l.). Combination in Leptogenys: Emery, 1895h: 20. Senior synonym of bellua, dispar, hewitti, parva: Bolton, 1975a: 285.

Ponerinae Ponerini Leptogenys crassinoda TRUE TRUE FALSE

crassinoda. Leptogenys attenuata var. crassinoda Arnold, 1926: 210 (w.) SOUTH AFRICA. Raised to species: Bolton, 1975a: 276.

Ponerinae Ponerini Leptogenys crustosa TRUE TRUE FALSE

crustosa. Leptogenys crustosa Santschi, 1914d: 326, fig. 12 (w.) GUINEA. Senior synonym of africanus, rufipes: Bolton, 1975a: 251.

Ponerinae Ponerini Leptogenys havilandi TRUE TRUE FALSE

havilandi. Leptogenys (Lobopelta) havilandi Forel, 1901f: 332 (w.) SOUTH AFRICA. See also: Bolton, 1975a: 282.

Ponerinae Ponerini Leptogenys intermedia TRUE TRUE FALSE

intermedia. Leptogenys intermedia Emery, 1902c: 32 (w.) SOUTH AFRICA. See also: Forel, 1914d: 214. Junior synonym of nitida Smith: Bolton, 1975a: 289. First available replacement name for nitida Smith, F. 1858b: 92 [Junior primary homonym of †nitida Heer, 1849: 149.]; designated by Bolton, 1995b: 231. Hence senior synonym of tenuis: Forel, 1912n: 52; Bolton, 1975a: 289; of adpressa, aena, capensis, deflocata, gracilis, grandior, insinuata, intermedia, speculans: Bolton, 1975a: 289.

Ponerinae Ponerini Leptogenys mactans TRUE TRUE FALSE

mactans. Leptogenys mactans Bolton, 1975a: 288, fig. 17 (w.) SOUTH AFRICA.

Ponerinae Ponerini Leptogenys peringueyi TRUE TRUE FALSE

peringueyi. Leptogenys (Lobopelta) havilandi st. peringueyi Forel, 1913j: 210 (w.) SOUTH AFRICA. Raised to species: Arnold, 1915: 96. See also: Bolton, 1975a: 282.

Myrmicinae Meranoplini Meranoplus inermis TRUE TRUE FALSE

inermis. Meranoplus inermis Emery, 1895h: 41, pl. 2, fig. 24 (w.) SOUTH AFRICA. Subspecies of nanus: Wheeler, W.M. 1922a: 888. Revived status as species and senior synonym of affinis, kiboshana, nanior, soriculus: Bolton, 1981a: 52.

Myrmicinae Meranoplini Meranoplus peringueyi TRUE TRUE FALSE

peringueyi. Meranoplus peringueyi Emery, 1886: 365, pl. 17, fig. 12 (w.) SOUTH AFRICA. Senior synonym of excisus: Bolton, 1981a: 56.

Myrmicinae Pheidolini Messor arenarius TRUE TRUE FALSE

4 subspecies

arenarius. Formica arenaria Fabricius, 1787: 310 (w.) NORTH AFRICA. André, 1883b: 366 (q.); Emery, 1891b: 12 (q.m.); Forel, 1902a: 148 (m.). Combination in Aphaenogaster: Roger, 1863b: 29; in A. (Messor): Forel, 1890a: lxx; in Messor: Forel, 1894d: 45; in Stenamma (Messor): Emery, 1898c: 125; in Messor: Emery, 1908e: 439. Senior synonym of amaurocyclia, scalpturata: Roger, 1862c: 292; of bugnioni: Forel, 1904f: 424. Current subspecies: nominal plus alfierii, diabolus, ratus, thebaicus. See also: Emery, 1908e: 439; Santschi, 1938a: 32.

Myrmicinae Pheidolini Messor capensis TRUE TRUE FALSE

capensis. Atta capensis Mayr, 1862: 743 (w.) SOUTH AFRICA. Mayr, 1866b: 896 (q.m.). Combination in Aphaenogaster: Roger, 1863b: 30; in Stenamma (Messor): Emery, 1895h: 35; in Messor: Forel, 1910e: 444. Subspecies of barbarus: Emery, 1891b: 12; Forel, 1910e: 444; Arnold, 1920a: 405; Emery, 1922c: 98; Stitz, 1923: 150; Prins, 1963: 100. Status as species: Santschi, 1917e: 94; Wheeler, W.M. 1922a: 804; Bolton, 1982: 345. Senior synonym of braunsi, donisthorpei, pseudoaegyptiaca, schencki: Bolton, 1982: 345.

Myrmicinae Pheidolini Messor decipiens TRUE TRUE FALSE

decipiens. Messor capensis st. decipiens Santschi, 1917e: 94 (w.q.) SOUTH AFRICA. [First available use of Stenamma (Messor) barbarum r. capense var. decipiens Forel, 1905b: 177; unavailable name.] Raised to species and senior synonym of arcistriatus, probus: Bolton, 1982: 348.

Myrmicinae Pheidolini Messor piceus TRUE TRUE FALSE

piceus. Messor piceus Stitz, 1923: 150 (w.q.) SOUTH AFRICA. See also: Bolton, 1982: 352.

Myrmicinae Dacetini Microdaceton exornatum TRUE TRUE FALSE

exornatum. Microdaceton exornatum Santschi, 1913g: 478 (w.) SOUTH AFRICA. [Also described as new by Santschi, 1914e: 33.] Arnold, 1948: 225 (q.). Senior synonym of laevior, leakeyi: Bolton, 1983: 402. See also: Bolton, 2000: 28.

Myrmicinae Solenopsidini Monomorium albopilosum TRUE TRUE FALSE

albopilosum. Monomorium albopilosum Emery, 1895h: 24 (w.) SOUTH AFRICA. Combination in M. (Xeromyrmex): Santschi, 1919b: 235. Senior synonym of clarithorax, fingo, paucipilosa, thales: Bolton, 1987: 335.

Myrmicinae Solenopsidini Monomorium anceps TRUE TRUE FALSE

anceps. Monomorium subopacum var. anceps Emery, 1895h: 24 (w.) SOUTH AFRICA. Combination in M. (Xeromyrmex): Wheeler, W.M. 1922a: 872. Raised to species: Bolton, 1987: 336.

Myrmicinae Solenopsidini Monomorium australe TRUE TRUE FALSE

australe. Monomorium subopacum r. australe Emery, 1886: 363 (w.q.) SOUTH AFRICA. Forel, 1913j: 216 (q.). Combination in M. (Parholcomyrmex): Santschi, 1917b: 282; Wheeler, W.M. 1922a: 873. Subspecies of salomonis: Forel, 1913j: 216; Arnold, 1916: 225. Raised to species: Dalla Torre, 1893: 65; Santschi, 1917b: 282; Bolton, 1987: 337. Material of the unavailable name laeviceps referred here by Bolton, 1987: 337.

Myrmicinae Solenopsidini Monomorium boerorum TRUE TRUE FALSE

boerorum. Monomorium minutum var. boerorum Forel, 1910e: 442 (w.q.) SOUTH AFRICA. Raised to species: Bolton, 1987: 381.

Myrmicinae Solenopsidini Monomorium braunsi TRUE TRUE FALSE

braunsi. Monomorium braunsi Mayr, 1901b: 7 (w.) SOUTH AFRICA. See also: Bolton, 1987: 382.

Myrmicinae Solenopsidini Monomorium excelsior TRUE TRUE FALSE

excelsior. Monomorium tchelichofi var. excelsior Arnold, 1926: 227 (w.m.) SOUTH AFRICA. Raised to species and senior synonym of speculiceps: Bolton, 1987: 344.

Myrmicinae Solenopsidini Monomorium excensurae TRUE TRUE FALSE

excensurae. Monomorium oscaris var. excensurae Forel, 1915c: 342 (w.) SOUTH AFRICA. Raised to species: Bolton, 1987: 386.

Myrmicinae Solenopsidini Monomorium fastidium TRUE TRUE FALSE

fastidium. Monomorium fastidium Bolton, 1987: 389 (w.q.) SOUTH AFRICA.

Myrmicinae Solenopsidini Monomorium fridae TRUE TRUE FALSE

fridae. Monomorium medinae r. fridae Forel, 1905b: 183 (w.) SOUTH AFRICA. Combination in M. (Xeromyrmex): Wheeler, W.M. 1922a: 870. Raised to species: Bolton, 1987: 344.

Myrmicinae Solenopsidini Monomorium havilandi TRUE TRUE FALSE

havilandi. Monomorium havilandi Forel, 1910e: 443 (w.q.) SOUTH AFRICA. Combination in M. (Parholcomyrmex): Santschi, 1917b: 282. Subspecies of australe: Santschi, 1917b: 282. Revived status as species and senior synonym of distinctum, leviceps: Bolton, 1987: 368.

Myrmicinae Solenopsidini Monomorium junodi TRUE TRUE FALSE

junodi. Monomorium salomonis subsp. junodi Forel, 1910e: 441 (w.) SOUTH AFRICA. Combination in M. (Xeromyrmex): Santschi, 1919b: 235. Subspecies of delagoense: Santschi, 1928f: 192; Arnold, 1944: 14. Raised to species and senior synonym of pretoriensis: Bolton, 1987: 346.

Myrmicinae Solenopsidini Monomorium kelapre TRUE TRUE FALSE

kelapre. Monomorium kelapre Bolton, 1987: 395 (w.) SOUTH AFRICA.

Myrmicinae Solenopsidini Monomorium lubricum TRUE TRUE FALSE

lubricum. Monomorium lubricum Arnold, 1948: 218, fig. 7 (w.) SOUTH AFRICA. See also: Bolton, 1987: 398.

Myrmicinae Solenopsidini Monomorium macrops TRUE TRUE FALSE

macrops. Monomorium mediocre subsp. macrops Arnold, 1944: 11, fig. 17 (w.) SOUTH AFRICA. Raised to species: Bolton, 1987: 369.

Myrmicinae Solenopsidini Monomorium mavide TRUE TRUE FALSE

mavide. Monomorium mavide Bolton, 1987: 400 (w.) SOUTH AFRICA.

Myrmicinae Solenopsidini Monomorium mediocre TRUE TRUE FALSE

mediocre. Monomorium mediocre Santschi, 1920d: 376, fig. 13 (w.) SOUTH AFRICA. See also: Bolton, 1987: 349.

Myrmicinae Solenopsidini Monomorium micropacum TRUE TRUE FALSE

micropacum. Monomorium micropacum Bolton, 1987: 350 (w.) SOUTH AFRICA.

Myrmicinae Solenopsidini Monomorium modestum TRUE TRUE FALSE

modestum. Monomorium modestum Santschi, 1914e: 17 (w.) SOUTH AFRICA. Combination in Syllophopsis: Santschi, 1915c: 259; in Monomorium: Bolton, 1987: 423. Senior synonym of smutsi, transwaalensis: Bolton, 1987: 423. See also: Heterick, 2006: 177.

Myrmicinae Solenopsidini Monomorium musicum TRUE TRUE FALSE

musicum. Monomorium oscaris subsp. musicum Forel, 1910e: 442 (w.q.) SOUTH AFRICA. Raised to species: Emery, 1922e: 173; Bolton, 1987: 402.

Myrmicinae Solenopsidini Monomorium notulum TRUE TRUE FALSE

notulum. Monomorium setuliferum var. notula Forel, 1910e: 441 (w.m.) SOUTH AFRICA. Raised to species and senior synonym of dolichops, latior: Bolton, 1987: 370.

Myrmicinae Solenopsidini Monomorium ocellatum TRUE TRUE FALSE

ocellatum. Monomorium (Xeromyrmex) salomonis st. ocelletum Arnold, in Santschi, 1920d: 377 (w.) SOUTH AFRICA. Arnold, 1926: 229 (q.). Raised to species: Santschi, 1936a: 39; Arnold, 1944: 14; Bolton, 1987: 352.

Myrmicinae Solenopsidini Monomorium orangiae TRUE TRUE FALSE

orangiae. Monomorium (Xeromyrmex) orangiae Arnold, 1956: 67, fig. 16 (w.) SOUTH AFRICA. See also: Bolton, 1987: 354.

Myrmicinae Solenopsidini Monomorium pacis TRUE TRUE FALSE

pacis. Monomorium pacis Forel, 1915c: 343 (w.) SOUTH AFRICA. See also: Bolton, 1987: 404.

Myrmicinae Solenopsidini Monomorium paternum TRUE TRUE FALSE

paternum. Monomorium paternum Bolton, 1987: 405, fig. 89 (w.) SOUTH AFRICA.

Myrmicinae Solenopsidini Monomorium rhopalocerum TRUE TRUE FALSE

rhopalocerum. Monomorium rhopalocerum Emery, 1895h: 25, pl. 2, fig 29 (w.) SOUTH AFRICA. Senior synonym of hottentota, leimbachi: Bolton, 1987: 407.

Myrmicinae Solenopsidini Monomorium rotundatum TRUE TRUE FALSE

rotundatum. Monomorium rotundatum Santschi, 1920b: 14, fig. 2 (w.) SOUTH AFRICA. See also: Bolton, 1987: 409.

Myrmicinae Solenopsidini Monomorium schultzei TRUE TRUE FALSE

schultzei. Monomorium schultzei Forel, 1910f: 18 (w.q.) SOUTH AFRICA. See also: Bolton, 1987: 410.

Myrmicinae Solenopsidini Monomorium tablense TRUE TRUE FALSE

tablense. Monomorium altinode st. tablensis Santschi, 1932a: 384, figs. 6, 7 (w.q.) SOUTH AFRICA. Raised to species: Bolton, 1987: 414.

Myrmicinae Solenopsidini Monomorium taedium TRUE TRUE FALSE

taedium. Monomorium taedium Bolton, 1987: 415 (w.) SOUTH AFRICA.

Myrmicinae Solenopsidini Monomorium tchelichofi TRUE TRUE FALSE

tchelichofi. Monomorium tchelichofi Forel, 1914d: 244 (w.) SOUTH AFRICA. See also: Bolton, 1987: 362.

Myrmicinae Solenopsidini Monomorium torvicte TRUE TRUE FALSE

torvicte. Monomorium torvicte Bolton, 1987: 416 (w.) SOUTH AFRICA.

Myrmicinae Solenopsidini Monomorium willowmorense TRUE TRUE FALSE

willowmorense. Monomorium willowmorense Bolton, 1987: 364, fig. 54 (w.) SOUTH AFRICA. Material of the unavailable names willowmorensis, belli referred here by Bolton, 1987: 364. See also: Heterick, 2006: 107.

Myrmicinae Solenopsidini Monomorium xanthognathum TRUE TRUE FALSE

xanthognathum. Monomorium xanthognathum Arnold, 1944: 9, fig. 15 (w.) SOUTH AFRICA. See also: Bolton, 1987: 371.

Myrmicinae Solenopsidini Monomorium zulu TRUE TRUE FALSE

zulu. Monomorium zulu Santschi, 1914e: 18 (w.) SOUTH AFRICA. See also: Bolton, 1987: 365.

Myrmicinae Myrmicariini Myrmicaria anomala TRUE TRUE FALSE

anomala. Myrmicaria anomala Arnold, 1960a: 81, fig. 2 (w.) SOUTH AFRICA.

Myrmicinae Myrmicariini Myrmicaria faurei TRUE TRUE FALSE

faurei. Myrmicaria faurei Arnold, 1947: 134, fig. 3 (w.) SOUTH AFRICA.

Myrmicinae Myrmicariini Myrmicaria foreli TRUE TRUE FALSE

1 subspecies

foreli. Myrmicaria foreli Santschi, 1925c: 163 (w.) SOUTH AFRICA. Current subspecies: nominal plus pallida.

Myrmicinae Myrmicariini Myrmicaria laevior TRUE TRUE FALSE

laevior. Myrmicaria laevior Forel, 1910e: 445 (w.) SOUTH AFRICA.

Myrmicinae Myrmicariini Myrmicaria natalensis TRUE TRUE FALSE

6 subspecies

natalensis. Physatta natalensis Smith, F. 1858b: 172 (q.) SOUTH AFRICA. Santschi, 1925c: 143 (q.m.). Combination in Myrmicaria: Mayr, 1866b: 905. Junior synonym of eumenoides: Mayr, 1866b: 905 (note that natalensis has priority). Revived from synonymy and senior synonym of sulcatus: Emery, 1922e: 123. Current subspecies: nominal plus eumenoides, navicula, nigriventris, obscuriceps, taeniata, verticalis. [M. natalensis referred by error to Myrmica by Santschi, 1925c: 143.]

Myrmicinae Myrmicariini Myrmicaria nigra TRUE TRUE FALSE

nigra. Heptacondylus niger Mayr, 1862: 755 (w.) SOUTH AFRICA. Arnold, 1916: 270 (m.). Combination in Myrmicaria: Dalla Torre, 1893: 155. See also: Mayr, 1865: 110.

Myrmicinae Myrmicariini Myrmicaria foreli pallida TRUE TRUE FALSE

pallida. Myrmicaria foreli var. pallida Arnold, in Santschi, 1925c: 165 (w.m.) SOUTH AFRICA.

Myrmicinae Myrmicariini Myrmicaria natalensis obscuriceps TRUE TRUE FALSE

obscuriceps. Myrmicaria natalensis var. obscuriceps Santschi, 1937d: 228 (w.) SOUTH AFRICA. Currently subspecies of natalensis: Santschi, 1939b: 238.

Myrmicinae Formicoxenini Myrmoxenus africana TRUE TRUE FALSE

africana. Epimyrma (Gonepimyrma) africana Bernard, 1948: 148, fig. 7 (w.) ALGERIA. Combination in Myrmoxenus (Gonepimyrma): new combination (unpublished).

Myrmicinae Formicoxenini Nesomyrmex antoinetteae TRUE TRUE FALSE

antoinetteae. Nesomyrmex antoinetteae Mbanyana & Robertson, 2008: 39, figs. 2g-i (w.) SOUTH AFRICA.

Myrmicinae Formicoxenini Nesomyrmex braunsi TRUE TRUE FALSE

braunsi. Dilobocondyla (Tetramyrma) braunsi Forel, 1912m: 767 (w.) SOUTH AFRICA. Forel, 1913a: 122 (pseudogyne); Arnold, 1917: 359 (q.). Combination in Tetramyrma: Forel, 1913a: 122; in Leptothorax: Bolton, 1982: 325; in Nesomyrmex: Bolton, 2003: 272. See also: Mbanyana & Robertson, 2008: 42.

Myrmicinae Formicoxenini Nesomyrmex cederbergensis TRUE TRUE FALSE

cederbergensis. Nesomyrmex cederbergensis Mbanyana & Robertson, 2008: 42, figs. 4g-i (w.) SOUTH AFRICA.

Myrmicinae Formicoxenini Nesomyrmex denticulatus TRUE TRUE FALSE

denticulatus. Leptothorax denticulatus Mayr, 1901b: 5 (w.q.) SOUTH AFRICA. Combination in L. (Goniothorax): Wheeler, W.M. 1922a: 891; in Nesomyrmex: Bolton, 2003: 272. See also: Bolton, 1982: 328; Mbanyana & Robertson, 2008: 38.

Myrmicinae Formicoxenini Nesomyrmex entabeni TRUE TRUE FALSE

entabeni. Nesomyrmex entabeni Mbanyana & Robertson, 2008: 43, figs. 3d-i (w.) SOUTH AFRICA.

Myrmicinae Formicoxenini Nesomyrmex ezantsi TRUE TRUE FALSE

ezantsi. Nesomyrmex ezantsi Mbanyana & Robertson, 2008: 43, figs. 4m-o (w.) SOUTH AFRICA.

Myrmicinae Formicoxenini Nesomyrmex humerosus TRUE TRUE FALSE

humerosus. Leptothorax (Goniothorax) humerosus Emery, 1896g: 62 (w.) EAST AFRICA (no locality given, probably KENYA). Combination in Nesomyrmex: Bolton, 2003: 272. See also: Bolton, 1982: 329.

Myrmicinae Formicoxenini Nesomyrmex inye TRUE TRUE FALSE

inye. Nesomyrmex inye Mbanyana & Robertson, 2008: 46, figs. 4j-l (w.) SOUTH AFRICA.

Myrmicinae Formicoxenini Nesomyrmex karooensis TRUE TRUE FALSE

karooensis. Nesomyrmex karooensis Mbanyana & Robertson, 2008: 47, figs. 4d-f (w.) SOUTH AFRICA.

Myrmicinae Formicoxenini Nesomyrmex koebergensis TRUE TRUE FALSE

koebergensis. Nesomyrmex koebergensis Mbanyana & Robertson, 2008: 48, figs. 2d-f (w.) SOUTH AFRICA.

Myrmicinae Formicoxenini Nesomyrmex larsenae TRUE TRUE FALSE

larsenae. Nesomyrmex larsenae Mbanyana & Robertson, 2008: 48, figs. 2m-o (w.) SOUTH AFRICA.

Myrmicinae Formicoxenini Nesomyrmex mcgregori TRUE TRUE FALSE

mcgregori. Nesomyrmex mcgregori Mbanyana & Robertson, 2008: 49, figs. 2j-l (w.) SOUTH AFRICA.

Myrmicinae Formicoxenini Nesomyrmex nanniae TRUE TRUE FALSE

nanniae. Nesomyrmex nanniae Mbanyana & Robertson, 2008: 50, figs. 3g-i (w.) SOUTH AFRICA.

Myrmicinae Formicoxenini Nesomyrmex njengelanga TRUE TRUE FALSE

njengelanga. Nesomyrmex njengelanga Mbanyana & Robertson, 2008: 51, figs. 3m-o (w.) SOUTH AFRICA.

Myrmicinae Formicoxenini Nesomyrmex ruani TRUE TRUE FALSE

ruani. Nesomyrmex ruani Mbanyana & Robertson, 2008: 51, figs. 3a-c (w.) SOUTH AFRICA.

Myrmicinae Formicoxenini Nesomyrmex saasveldensis TRUE TRUE FALSE

saasveldensis. Nesomyrmex saasveldensis Mbanyana & Robertson, 2008: 52, figs. 2a-c (w.) SOUTH AFRICA.

Myrmicinae Formicoxenini Nesomyrmex simoni TRUE TRUE FALSE

simoni. Tetramorium simoni Emery, 1895h: 35, pl. 2, fig. 22 (w.) SOUTH AFRICA. Combination in Tetramyrma: Emery, 1924d: 291; in Leptothorax: Bolton, 1982: 331; in Nesomyrmex: Bolton, 2003: 272. See also: Mbanyana & Robertson, 2008: 53.

Myrmicinae Formicoxenini Nesomyrmex stramineus TRUE TRUE FALSE

stramineus. Limnomyrmex stramineus Arnold, 1948: 223, fig. 10 (w.) SOUTH AFRICA. Combination in Leptothorax: Brown, 1971a: 4; in Nesomyrmex: Bolton, 2003: 272. See also: Bolton, 1982: 332; Mbanyana & Robertson, 2008: 38.

Myrmicinae Formicoxenini Nesomyrmex tshiguvhoae TRUE TRUE FALSE

tshiguvhoae. Nesomyrmex tshiguvhoae Mbanyana & Robertson, 2008: 53, figs. 4a-c (w.) SOUTH AFRICA.

Myrmicinae Formicoxenini Nesomyrmex vannoorti TRUE TRUE FALSE

vannoorti. Nesomyrmex vannoorti Mbanyana & Robertson, 2008: 54, figs. 3j-l (w.) SOUTH AFRICA.

Formicinae Plagiolepidini Nylanderia traegaordhi natalensis TRUE TRUE FALSE

natalensis. Prenolepis (Nylanderia) traegaordhi var. natalensis Forel, 1915c: 348 (w.) SOUTH AFRICA. Combination in Paratrechina (Nylanderia): Emery, 1925b: 218; in Nylanderia: LaPolla, Brady & Shattuck, 2010a: 127.

Myrmicinae Pheidolini Ocymyrmex alacer TRUE TRUE FALSE

alacer. Ocymyrmex alacer Bolton & Marsh, 1989: 1290, fig. 9 (w.q.) SOUTH AFRICA.

Myrmicinae Pheidolini Ocymyrmex barbiger TRUE TRUE FALSE

barbiger. Ocymyrmex barbiger Emery, 1886: 364, pl. 17, figs. 9-11 (w.m.) SOUTH AFRICA. [Misspelled as barbatus by Emery, 1892a: 114.] Bolton, 1981b: 261 (q.). See also: Bolton & Marsh, 1989: 1291.

Myrmicinae Pheidolini Ocymyrmex cavatodorsatus TRUE TRUE FALSE

cavatodorsatus. Ocymyrmex cavatodorsatus Prins, 1965c: 1021, figs. 1, 2 (w.) SOUTH AFRICA. Bolton & Marsh, 1989: 1278 (q.). See also: Bolton, 1981b: 266.

Myrmicinae Pheidolini Ocymyrmex cilliei TRUE TRUE FALSE

cilliei. Ocymyrmex cilliei Prins & Roux, in Bolton & Marsh, 1989: 1292 (m.) SOUTH AFRICA.

Myrmicinae Pheidolini Ocymyrmex gariepensis TRUE TRUE FALSE

gariepensis. Ocymyrmex gariepensis Prins & Roux, in Bolton & Marsh, 1989: 1282 (diagnosis in key) (w.) SOUTH AFRICA.

Myrmicinae Pheidolini Ocymyrmex gordoni TRUE TRUE FALSE

gordoni. Ocymyrmex gordoni Prins & Roux, in Bolton & Marsh, 1989: 1282 (diagnosis in key) (w.) SOUTH AFRICA.

Myrmicinae Pheidolini Ocymyrmex ignotus TRUE TRUE FALSE

ignotus. Ocymyrmex ignotus Bolton & Marsh, 1989: 1297 (w.q.) SOUTH AFRICA.

Myrmicinae Pheidolini Ocymyrmex resekhes TRUE TRUE FALSE

resekhes. Ocymyrmex resekhes Bolton & Marsh, 1989: 1301, fig. 17 (w.q.) SOUTH AFRICA.

Ponerinae Ponerini Odontoponera denticulata TRUE TRUE FALSE

denticulata. Ponera denticulata Smith, F. 1858b: 90, pl. 6, figs. 13, 14 (q.) SINGAPORE (not South Africa as originally stated; see Donisthorpe, 1943f: 677). Roger, 1861a: 12 (w.). Combination in Odontoponera: Mayr, 1862: 717. Junior synonym of transversa: Dalla Torre, 1893: 30; Forel, 1900d: 314. Revived from synonymy: Yamane, 2009: 5.

Ponerinae Ponerini Pachycondyla aenigmatica TRUE TRUE FALSE

aenigmatica. Ponera aenigmatica Arnold, 1949: 265, fig. 5 (w.) SOUTH AFRICA. Combination in Pachycondyla: Brown, in Bolton, 1995b: 302.

Ponerinae Ponerini Pachycondyla berthoudi TRUE TRUE FALSE

1 subspecies

berthoudi. Ophthalmopone berthoudi Forel, 1890b: cxiii (w.) SOUTH AFRICA. Forel, 1894b: 76 (m.); Wheeler, G.C. & Wheeler, J. 1971b: 1203 (l.). Combination in Pachycondyla: Brown, in Bolton, 1995b: 303. Current subspecies: nominal plus pubescens. See also: Arnold, 1915: 50; Peeters & Crewe, 1985: 29; Peeters & Crewe, 1987: 201.

Ponerinae Ponerini Pachycondyla caffraria TRUE TRUE FALSE

2 subspecies

caffraria. Ponera caffraria Smith, F. 1858b: 91 (q.) SOUTH AFRICA. Forel, 1913a: 108 (w.); Arnold, 1915: 65 (w.m.); Emery, 1899c: 5 (l.); Wheeler, G.C. & Wheeler, J. 1971b: 1206 (l.). Combination in Euponera (Mesoponera): Emery, 1901a: 46; in Mesoponera: Wheeler, G.C. & Wheeler, J. 1971b: 1206; in Pachycondyla: Brown, in Bolton, 1995b: 303. Senior synonym of guineensis: Mayr, 1895: 125; Emery, 1911d: 81; Wheeler, W.M. 1922a: 775. Current subspecies: nominal plus affinis, caffra.

Ponerinae Ponerini Pachycondyla cavernosa TRUE TRUE FALSE

1 subspecies

cavernosa. Ponera cavernosa Roger, 1860: 288 (w.) SOUTH AFRICA. Combination in Bothroponera: Mayr, 1862: 717; Wheeler, W.M. 1922a: 72; in Pachycondyla (Bothroponera): Emery, 1901a: 45; in Pachycondyla: Brown, in Bolton, 1995b: 304. Current subspecies: nominal plus montivaga.

Ponerinae Ponerini Pachycondyla darwinii TRUE TRUE FALSE

3 subspecies

darwinii. Belonopelta darwinii Forel, 1893d: 460 (q.) AUSTRALIA. Terayama, 1990: 897 (w.). Combination in Pachycondyla (Pseudoponera): Emery, 1900d: 668; Emery, 1901a: 46; in Pseudoponera: Bingham, 1903: 93; in Euponera (Trachymesopus): Emery, 1911d: 86; in Trachymesopus: Wilson, 1958d: 353; in Pachycondyla: Brown, in Bolton, 1995b: 304. Senior synonym of lamarki, rufotestaceus: Brown, 1963: 6. Current subspecies: nominal plus africana, indica, madecassa.

Ponerinae Ponerini Pachycondyla fossigera TRUE TRUE FALSE

fossigera. Euponera fossigera Mayr, 1901b: 3 (w.) SOUTH AFRICA. Combination in E. (Mesoponera): Arnold, 1915: 71; in Pachycondyla: Brown, in Bolton, 1995b: 305.

Ponerinae Ponerini Pachycondyla granosa TRUE TRUE FALSE

granosa. Ponera granosa Roger, 1860: 290 (w.) SOUTH AFRICA. Arnold, 1926: 201 (m.). Combination in Bothroponera: Mayr, 1862: 717; in Pachycondyla (Bothroponera): Emery, 1901a: 45; in Bothroponera: Wheeler, W.M. 1922a: 72; in Pachycondyla: Brown, in Bolton, 1995b: 305.

Ponerinae Ponerini Pachycondyla hartwigi TRUE TRUE FALSE

hartwigi. Cryptopone hartwigi Arnold, 1948: 213, fig. 2 (w.) SOUTH AFRICA. Combination in Hypoponera: Taylor, 1967a: 12; in Cryptopone: Bolton, 1995b: 166; in Pachycondyla: Mackay & Mackay, 2010: 3 (by implication as Cryptopone synonymised with Pachycondyla).

Ponerinae Ponerini Pachycondyla havilandi TRUE TRUE FALSE

3 subspecies

havilandi. Megaloponera (Hagensia) havilandi Forel, 1901f: 333 (w.) SOUTH AFRICA. Arnold, 1915: 68 (m.). Combination in Euponera (Mesoponera): Arnold, 1915: 67; in Hagensia: Arnold, 1926: 203; in Pachycondyla: Brown, in Bolton, 1995b: 305. Senior synonym of sulcigera: Arnold, 1915: 67. Current subspecies: nominal plus fochi, godfreyi, marleyi.

Ponerinae Ponerini Pachycondyla hottentota TRUE TRUE FALSE

hottentota. Pachycondyla hottentota Emery, 1886: 360 (w.) SOUTH AFRICA. Arnold, 1915: 52 (m.). Combination in Ophthalmopone: Emery, 1902c: 30; in Pachycondyla: Brown, in Bolton, 1995b: 306. Senior synonym of lanceolata: Emery, 1902c: 30.

Ponerinae Ponerini Pachycondyla kruegeri TRUE TRUE FALSE

2 subspecies

kruegeri. Pachycondyla (Bothroponera) kruegeri Forel, 1910a: 28 (w.) SOUTH AFRICA. Combination in Bothroponera: Wheeler, W.M. 1922a: 22; in Pachycondyla: Brown, in Bolton, 1995b: 306. Current subspecies: nominal plus asina, rhodesiana. See also: Arnold, 1915: 56.

Ponerinae Ponerini Pachycondyla laevissima TRUE TRUE FALSE

1 subspecies

laevissima. Pachycondyla (Bothroponera) laevissima Arnold, 1915: 58, pl. 2, fig. 15 (w.) SOUTH AFRICA. Combination in Bothroponera: Wheeler, W.M. 1922a: 73; in Pachycondyla: Brown, in Bolton, 1995b: 306. Current subspecies: nominal plus aspera.

Ponerinae Ponerini Pachycondyla peringueyi TRUE TRUE FALSE

1 subspecies

peringueyi. Ponera peringueyi Emery, 1899e: 474 (w.) SOUTH AFRICA. Arnold, 1951: 53 (m.); Wheeler, G.C. & Wheeler, J. 1971b: 1203 (l.). Combination in Euponera (Brachyponera): Emery, 1901a: 47; in E. (Mesoponera): Emery, 1911d: 81; in Hagensia: Arnold, 1926: 202; in Pachycondyla: Brown, in Bolton, 1995b: 308. Current subspecies: nominal plus saldanhae. See also: Arnold, 1915: 66.

Ponerinae Ponerini Pachycondyla pumicosa TRUE TRUE FALSE

1 subspecies

pumicosa. Ponera pumicosa Roger, 1860: 290 (w.) SOUTH AFRICA. Forel, 1901f: 344 (m.); Wheeler, G.C. & Wheeler, J. 1971c: 390 (l.). Combination in Bothroponera: Mayr, 1862: 717; in Pachycondyla (Bothroponera): Emery, 1901a: 45; in Bothroponera: Wheeler, W.M. 1922a: 72; in Pachycondyla: Brown, in Bolton, 1995b: 308. [Misspelled as pumicata by Raffray, 1887: 21.] Current subspecies: nominal plus sculpturata. See also: Arnold, 1915: 62.

Ponerinae Ponerini Pachycondyla strigulosa TRUE TRUE FALSE

strigulosa. Bothroponera strigulosa Emery, 1895h: 19 (w.) SOUTH AFRICA. Combination in Pachycondyla (Bothroponera): Emery, 1901a: 45; in Bothroponera: Wheeler, W.M. 1922a: 72; in Pachycondyla: Brown, in Bolton, 1995b: 310. Senior synonym of berthoudi: Arnold, 1952a: 460.

Ponerinae Ponerini Pachycondyla variolosa TRUE TRUE FALSE

variolosa. Bothroponera variolosa Arnold, 1947: 131, fig. 1 (w.) SOUTH AFRICA. Combination in Pachycondyla: Brown, in Bolton, 1995b: 311.

Ponerinae Ponerini Pachycondyla wroughtonii TRUE TRUE FALSE

1 subspecies

wroughtonii. Euponera (Pseudoponera) wroughtonii Forel, 1901f: 341 (w.m.) SOUTH AFRICA. Arnold, 1926: 204 (q.); Wheeler, G.C. & Wheeler, J. 1971b: 1206 (l.). Combination in E. (Trachymesopus): Emery, 1911d: 85; in Mesoponera: Wheeler, G.C. & Wheeler, J. 1971b: 1206; in Pachycondyla: Brown, in Bolton, 1995b: 311. Current subspecies: nominal plus crudelis.

Ponerinae Ponerini Pachycondyla peringueyi saldanhae TRUE TRUE FALSE

saldanhae. Hagensia peringueyi r. saldanhae Arnold, 1951: 53, fig. 4 (w.) SOUTH AFRICA. Combination in Pachycondyla: Brown, in Bolton, 1995b: 309.

Ponerinae Ponerini Pachycondyla elisae rotundata TRUE TRUE FALSE

rotundata. Ponera elisae var. rotundata Emery, 1895h: 20 (w.) SOUTH AFRICA. Arnold, 1915: 71 (q.). Combination in Euponera (Mesoponera): Emery, 1901a: 46; in E. (Xiphopelta): Forel, 1913j: 206; in Pachycondyla: Brown, in Bolton, 1995b: 309. Senior synonym of arnoldi: Wheeler, W.M. 1922a: 776. Currently subspecies of elisae: Santschi, 1937d: 217.

Ponerinae Ponerini Pachycondyla cavernosa montivaga TRUE TRUE FALSE

montivaga. Bothroponera cavernosa var. montivaga Arnold, 1947: 132 (w.) SOUTH AFRICA. Combination in Pachycondyla: Brown, in Bolton, 1995b: 307.

Ponerinae Ponerini Pachycondyla havilandi marleyi TRUE TRUE FALSE

marleyi. Hagensia havilandi r. marleyi Arnold, 1926: 203 (w.) SOUTH AFRICA. Combination in Pachycondyla: Brown, in Bolton, 1995b: 307.

Ponerinae Ponerini Pachycondyla havilandi godfreyi TRUE TRUE FALSE

godfreyi. Hagensia havilandi r. godfreyi Arnold, 1926: 203 (w.m.) SOUTH AFRICA. Combination in Pachycondyla: Brown, in Bolton, 1995b: 305.

Ponerinae Ponerini Pachycondyla havilandi fochi TRUE TRUE FALSE

fochi. Euponera (Hagensia) havilandi var. fochi Forel, 1918b: 155 (w.) SOUTH AFRICA. Combination in Pachycondyla: Brown, in Bolton, 1995b: 305.

Ponerinae Ponerini Pachycondyla wroughtonii crudelis TRUE TRUE FALSE

crudelis. Euponera (Pseudoponera) wroughtonii var. crudelis Forel, 1901f: 343 (w.m.) SOUTH AFRICA. Combination in E. (Trachymesopus): Emery, 1911d: 85; in Pachycondyla: Brown, in Bolton, 1995b: 304.

Ponerinae Ponerini Pachycondyla laevissima aspera TRUE TRUE FALSE

aspera. Bothroponera laevissima var. aspera Arnold, 1962: 844 (w.) SOUTH AFRICA. Combination in Pachycondyla: Brown, in Bolton, 1995b: 303.

Ponerinae Ponerini Pachycondyla darwinii africana TRUE TRUE FALSE

africana. Euponera (Pseudoponera) darwini var. africana Forel, 1909b: 51 (q.) DEMOCRATIC REPUBLIC OF CONGO. Santschi, 1914b: 51 (m.). Combination in Euponera (Trachymesopus): Emery, 1911d: 86; in Pachycondyla: Brown, in Bolton, 1995b: 302. Currently subspecies of darwinii: Santschi, 1937b: 95.

Myrmicinae Pheidolini Pheidole akermani TRUE TRUE FALSE

akermani. Pheidole akermani Arnold, 1920a: 471 (s.w.) SOUTH AFRICA.

Myrmicinae Pheidolini Pheidole aspera TRUE TRUE FALSE

aspera. Pheidole aspera Mayr, 1862: 746 (s.w.) SOUTH AFRICA. See also: Mayr, 1865: 97.

Myrmicinae Pheidolini Pheidole caffra TRUE TRUE FALSE

5 subspecies

caffra. Pheidole caffra Emery, 1895h: 33 (s.) SOUTH AFRICA. Santschi, 1939b: 239 (w.). Current subspecies: nominal plus abyssinica, amoena, bayeri, montivaga, senilifrons.

Myrmicinae Pheidolini Pheidole capensis TRUE TRUE FALSE

2 subspecies

capensis. Pheidole capensis Mayr, 1862: 746 (s.w.) SOUTH AFRICA. Forel, 1913a: 130 (q.). Subspecies of megacephala: Emery, 1895h: 34. Revived status as species: Forel, 1913a: 130; Arnold, 1920a: 441. Current subspecies: nominal plus dregei, modestior, reddenburgensis.

Myrmicinae Pheidolini Pheidole crassinoda TRUE TRUE FALSE

3 subspecies

crassinoda. Pheidole crassinoda Emery, 1895h: 32 (s.w.) SOUTH AFRICA. Mayr, 1907b: 12 (q.m.). [P. crassinoda Forel, 1894b: 94. Nomen nudum, attributed to Emery.] Current subspecies: nominal plus pluto, ruspolii, sordidula.

Myrmicinae Pheidolini Pheidole foreli TRUE TRUE FALSE

2 subspecies

foreli. Pheidole foreli Mayr, 1901b: 8 (s.w.) SOUTH AFRICA. Forel, 1910f: 12 (q.). Current subspecies: nominal plus pubens. See also: Arnold, 1920a: 433.

Myrmicinae Pheidolini Pheidole hewitti TRUE TRUE FALSE

hewitti. Pheidole hewitti Santschi, 1932a: 382, figs. 1-4 (s.w.) SOUTH AFRICA. Senior synonym of hewitti Arnold: Arnold, 1952a: 462.

Myrmicinae Pheidolini Pheidole irritans TRUE TRUE FALSE

irritans. Myrmica irritans Smith, F. 1858b: 127 (w.) SOUTH AFRICA. Combination in Pheidole: Dixey & Longstaff, 1907: 331.

Myrmicinae Pheidolini Pheidole kitschneri TRUE TRUE FALSE

kitschneri. Pheidole kitschneri Forel, 1910e: 436 (s.w.) SOUTH AFRICA. Santschi, 1914b: 76 (q.).

Myrmicinae Pheidolini Pheidole punctulata TRUE TRUE FALSE

3 subspecies

punctulata. Pheidole punctulata Mayr, 1866b: 899 (s.) SOUTH AFRICA. Arnold, 1920a: 421 (w.q.m.). Subspecies of megacephala: Forel, 1894b: 94; Stitz, 1911b: 385; Emery, 1915j: 235; Emery, 1921f: 85; Wheeler, W.M. 1922a: 132; Santschi, 1930b: 67. Status as species: Forel, 1910e: 441; Santschi, 1914b: 76; Forel, 1914d: 240; Arnold, 1920a: 420; Santschi, 1935a: 256. Current subspecies: nominal plus atrox, spinosa, subatrox.

Myrmicinae Pheidolini Pheidole sculpturata TRUE TRUE FALSE

7 subspecies

sculpturata. Pheidole sculpturata Mayr, 1866b: 897 (s.) SOUTH AFRICA. Emery, 1895h: 31 (w.). Current subspecies: nominal plus areolata, berthoudi, dignata, particeps, rhodesiana, welgelegenensis, zambesiana. See also: Arnold, 1920a: 462.

Myrmicinae Pheidolini Pheidole tenuinodis TRUE TRUE FALSE

2 subspecies

tenuinodis. Pheidole tenuinodis Mayr, 1901b: 10 (s.w.) SOUTH AFRICA. Arnold, 1920a: 430 (q.). Current subspecies: nominal plus bothae, robusta, sipapomae.

Myrmicinae Pheidolini Pheidole capensis dregei TRUE TRUE FALSE

dregei. Pheidole megacephala subsp. dregei Emery, 1895h: 34 (s.) SOUTH AFRICA. Currently subspecies of capensis: Emery, 1915j: 246.

Myrmicinae Pheidolini Pheidole spinulosa conigera TRUE TRUE FALSE

conigera. Pheidole spinulosa subsp. conigera Forel, 1910e: 440 (s.) SOUTH AFRICA.

Myrmicinae Pheidolini Pheidole foreli bothae TRUE TRUE FALSE

bothae. Pheidole foreli r. bothae Forel, 1901h: 62 (s.w.) SOUTH AFRICA. Santschi, 1914e: 22 (q.). Subspecies of tenuinodis: Santschi, 1914e: 22. Junior synonym of tenuinodis: Emery, 1921f: 87. Revived from synonymy as subspecies of tenuinodis: Wheeler, W.M. 1922a: 822.

Myrmicinae Pheidolini Pheidole sculpturata berthoudi TRUE TRUE FALSE

berthoudi. Pheidole sculpturata r. berthoudi Forel, 1894b: 89 (s.w.) SOUTH AFRICA. Raised to species: Santschi, 1921c: 116. Currently subspecies of sculpturata: Emery, 1921f: 89; Wheeler, W.M. 1922a: 819.

Myrmicinae Pheidolini Pheidole sculpturata areolata TRUE TRUE FALSE

areolata. Pheidole sculpturata var. areolata Forel, 1911e: 269 (s.w.) SOUTH AFRICA.

Myrmicinae Pheidolini Pheidole capensis modestior TRUE TRUE FALSE

modestior. Pheidole capensis var. modestior Santschi, 1919a: 337 (s.w.) SOUTH AFRICA.

Myrmicinae Pheidolini Pheidole foreli pubens TRUE TRUE FALSE

pubens. Pheidole foreli var. pubens Forel, 1910f: 12 (s.w.) SOUTH AFRICA. Forel, 1910e: 436 (q.m.).

Myrmicinae Pheidolini Pheidole cuitensis reddenburgensis TRUE TRUE FALSE

reddenburgensis. Pheidole (Allopheidole) cuitensis r. reddenburgensis Forel, 1913a: 135 (s.w.q.m.) SOUTH AFRICA. Subspecies of capensis: Emery, 1915j: 246; Wheeler, W.M. 1922a: 809. [Spelling emended to reddersbergensis by Arnold, 1920a: 450; emendation not accepted by Wheeler, W.M. 1922a: 809.]

Myrmicinae Solenopsidini Pheidologeton hostilis TRUE TRUE FALSE

hostilis. Atta hostilis Smith, F. 1858b: 165 (w.q.) SOUTH AFRICA. Combination in Pheidologeton: Mayr, 1866b: 899.

Formicinae Plagiolepidini Plagiolepis capensis TRUE TRUE FALSE

capensis. Plagiolepis capensis Mayr, 1865: 55 (q.) SOUTH AFRICA.

Formicinae Plagiolepidini Plagiolepis decora TRUE TRUE FALSE

decora. Plagiolepis (Anacantholepis) decora Santschi, 1914e: 36, fig. 9 (w.) SOUTH AFRICA. Arnold, 1922: 582 (q.).

Formicinae Plagiolepidini Plagiolepis deweti TRUE TRUE FALSE

deweti. Plagiolepis deweti Forel, 1904f: 423 (w.q.) SOUTH AFRICA. Combination in P. (Anacantholepis): Forel, 1914d: 249.

Formicinae Plagiolepidini Plagiolepis funicularis TRUE TRUE FALSE

funicularis. Plagiolepis funicularis Santschi, 1919a: 344 (w.) SOUTH AFRICA.

Formicinae Plagiolepidini Plagiolepis fuscula TRUE TRUE FALSE

fuscula. Plagiolepis fuscula Emery, 1895h: 45 (w.) SOUTH AFRICA. Combination in P. (Anacantholepis): Emery, 1925b: 22.

Formicinae Plagiolepidini Plagiolepis intermedia TRUE TRUE FALSE

1 subspecies

intermedia. Plagiolepis pygmaea var. intermedia Emery, 1895h: 45 (w.q.) SOUTH AFRICA. Raised to species: Emery, 1921d: 316. Current subspecies: nominal plus minutula.

Formicinae Plagiolepidini Plagiolepis jouberti TRUE TRUE FALSE

jouberti. Plagiolepis jouberti Forel, 1910f: 23 (w.) SOUTH AFRICA. Combination in P. (Anacantholepis): Santschi, 1914b: 122.

Formicinae Plagiolepidini Plagiolepis puncta TRUE TRUE FALSE

puncta. Plagiolepis pygmaea var. punctum Forel, 1910e: 448 (w.m.) SOUTH AFRICA. Arnold, 1922: 602 (q.). Subspecies of pygmaea: Arnold, 1922: 602; Wheeler, W.M. 1922a: 930. Raised to species: Emery, 1921d: 317; Emery, 1925b: 21.

Formicinae Plagiolepidini Plagiolepis vanderkelleni polita TRUE TRUE FALSE

polita. Plagiolepis (Anacantholepis) vanderkeleni st. polita Santschi, 1914e: 37, fig. 10 (w.) SOUTH AFRICA.

Formicinae Plagiolepidini Plagiolepis intermedia minutula TRUE TRUE FALSE

minutula. Plagiolepis intermedia var. minutula Emery, 1921d: 316, fig. 2 (w.q.) SOUTH AFRICA.

Ponerinae Platythyreini Platythyrea cooperi TRUE TRUE FALSE

cooperi. Platythyrea cooperi Arnold, 1915: 29 (w.m.) SOUTH AFRICA.

Ponerinae Platythyreini Platythyrea lamellosa TRUE TRUE FALSE

lamellosa. Ponera lamellosa Roger, 1860: 295 (w.) SOUTH AFRICA. Wheeler, G.C. & Wheeler, J. 1989b: 51 (l.). Combination in Platythyrea: Roger, 1863a: 173. Senior synonym of grisea: Forel, 1913b: 307; of longinoda, rhodesiana, suturalis: Brown, 1975: 8.

Ponerinae Ponerini Plectroctena mandibularis TRUE TRUE FALSE

mandibularis. Plectroctena mandibularis Smith, F. 1858b: 101, pl. 7, figs. 1-5 (m., ergatoid q., not w.) SOUTH AFRICA. Forel, 1913a: 108 (w.); Arnold, 1915: 86 (w.). Material of the nomen nudum caffra referred here: Roger, 1861a: 41. Senior synonym of major: Emery, 1899e: 469; Bolton, 1974c: 330; of integra and material of the unavailable name strialiventris referred here: Bolton, 1974c: 330; of conjugata: Villet, McKitterick & Robertson, 1999: 282. .

Ponerinae Ponerini Plectroctena strigosa TRUE TRUE FALSE

strigosa. Plectroctena mandibularis var. strigosa Emery, 1899e: 469 (w.) SOUTH AFRICA. Subspecies of cristata: Santschi, 1924a: 161. Raised to species: Bolton, 1974c: 332.

Formicinae Camponotini Polyrhachis arnoldi TRUE TRUE FALSE

arnoldi. Polyrhachis (Myrma) arnoldi Forel, 1914d: 263 (w.) SOUTH AFRICA. See also: Arnold, 1924: 750; Bolton, 1973b: 324.

Formicinae Camponotini Polyrhachis cubaensis TRUE TRUE FALSE

cubaensis. Polyrhachis cubaensis Mayr, 1862: 687 (w.) SOUTH AFRICA (not Cuba; see Mayr, 1893: 195). Combination in P. (Myrma): Wheeler, W.M. 1922a: 994. Senior synonym of gerstaeckeri, striolatorugosa, wilmsi: Bolton, 1973b: 325.

Formicinae Camponotini Polyrhachis durbanensis TRUE TRUE FALSE

durbanensis. Polyrhachis (Myrma) cubaensis r. durbanensis Forel 1914d: 262 (w.q.) SOUTH AFRICA. Raised to species: Bolton, 1973b: 327.

Formicinae Camponotini Polyrhachis gagates TRUE TRUE FALSE

gagates. Polyrhachis gagates Smith, F. 1858b: 71, pl. 4, fig. 14 (w.) SOUTH AFRICA. Forel, 1886f: 194 (q.); Wheeler, G.C. & Wheeler, J. 1953e: 207 (l.). Combination in P. (Myrma): Santschi, 1914b: 139. Subspecies of schistacea: Arnold, 1924: 745. Revived status as species: Emery, 1925b: 199; Bolton, 1973b: 305. Senior synonym of clariseta, congolensis (and its junior synonym indefinita), nigriseta, obsidiana and material of the unavailable name acheron referred here: Bolton, 1973b: 305.

Formicinae Camponotini Polyrhachis gamaii TRUE TRUE FALSE

gamaii. Polyrhachis (Myrma) gamaii Santschi, 1917b: 295 (q.) SOUTH AFRICA. Arnold, 1947: 136 (w.). See also: Bolton, 1973b: 351.

Formicinae Camponotini Polyrhachis militaris TRUE TRUE FALSE

militaris. Formica militaris Fabricius, 1782: 493 (q.) "Tropical Africa". Smith, F. 1858b: 72 (m.); Mayr, 1866b: 886 (w.). Combination in Myrma: Billberg, 1820: 104; in Polyrhachis: Smith, F. 1858b: 72; in P. (Myrma): Santschi, 1914b: 140. Senior synonym of bruta, calabarica, cupreopubescens, ssibangensis, striativentris: Bolton, 1973b: 313; of epinotalis, nkomoensis: Dorow, 1995: 36, 37 respectively. Material of the unavailable names argentatus, dido, pleurata, sankisiana, transversaria referred here by Bolton, 1973b: 313.

Formicinae Camponotini Polyrhachis phidias TRUE TRUE FALSE

phidias. Polyrhachis phidias Forel, 1910e: 450 (w.) "Equatorial Africa". Combination in P. (Myrma): Wheeler, W.M. 1922a: 999. See also: Bolton, 1973b: 316.

Formicinae Camponotini Polyrhachis schlueteri TRUE TRUE FALSE

schlueteri. Polyrhachis militaris r. schlueteri Forel, 1886f: 195 (w.) EAST AFRICA. Arnold, 1924: 748 (q.). Combination in P. (Myrma): Santschi, 1914b: 143. Raised to species: Dalla Torre, 1893: 268. Subspecies of schistacea: Santschi, 1914e: 42; Wheeler, W.M. 1922a: 1003; Arnold, 1924: 747. Status as species: Forel, 1907g: 92; Forel, 1914d: 261; Santschi, 1914b: 143; Emery, 1925b: 200; Bolton, 1973b: 321. Senior synonym of indigens: Arnold, 1924: 747; of plebeia: Bolton, 1973b: 321.

Formicinae Camponotini Polyrhachis viscosa TRUE TRUE FALSE

viscosa. Polyrhachis viscosus Smith, F. 1858b: 71, pl. 4, fig. 41 (w.) SOUTH AFRICA. Combination in P. (Myrma): Santschi, 1914b: 143. Senior synonym of antinorii: Dalla Torre, 1893: 271; of imatongica, spretula: Bolton, 1973b: 330. See also: Arnold, 1924: 749.

Amblyoponinae Amblyoponini Prionopelta aethiopica TRUE TRUE FALSE

aethiopica. Prionopelta descarpentriesi subsp. aethiopica Arnold, 1949: 263, fig. 4 (w.) SOUTH AFRICA. Raised to species: Brown, 1960a: 177.

Myrmicinae Myrmecinini Pristomyrmex africanus TRUE TRUE FALSE

africanus. Pristomyrmex africanus Karavaiev, 1931d: 47, fig. 5 (w.) KENYA. Wang, M. 2003: 424 (q. and ergatoid q.). Combination in Hylidris: Weber, 1952: 17; in Pristomyrmex: Bolton, 1981b: 284. Senior synonym of beni, mbomu, myersi, primus: Bolton, 1981b: 284.

Myrmicinae Myrmecinini Pristomyrmex fossulatus TRUE TRUE FALSE

fossulatus. Tetramorium (Xiphomyrmex) fossulatum Forel, 1910e: 428 (w.) SOUTH AFRICA. Menozzi, 1942: 172 (q.). Combination in Pristomyrmex: Santschi, 1916a: 51; in Hylidris: Weber, 1952: 17; in Pristomyrmex: Bolton, 1981b: 285. See also: Wang, M. 2003: 406.

Proceratiinae Probolomyrmecini Probolomyrmex filiformis TRUE TRUE FALSE

filiformis. Probolomyrmex filiformis Mayr, 1901b: 3 (w.) SOUTH AFRICA. See also: Taylor, 1965d: 353.

Myrmicinae Dacetini Pyramica africana TRUE TRUE FALSE

africana. Glamyromyrmex africanus Bolton, 1983: 322, fig. 27 (w.) GABON. Combination in Pyramica: Bolton, 1999: 1672. See also: Bolton, 2000: 329.

Myrmicinae Dacetini Pyramica agnosta TRUE TRUE FALSE

agnosta. Pyramica agnosta Bolton, 2000: 293, figs. 199, 227 (w.) SOUTH AFRICA.

Myrmicinae Dacetini Pyramica anarta TRUE TRUE FALSE

anarta. Smithistruma anarta Bolton, 1983: 314, fig. 18 (w.) SOUTH AFRICA. Combination in Pyramica: Bolton, 1999: 1673. See also: Bolton, 2000: 323.

Myrmicinae Dacetini Pyramica dictynna TRUE TRUE FALSE

dictynna. Pyramica dictynna Bolton, 2000: 294 (w.) SOUTH AFRICA.

Myrmicinae Dacetini Pyramica emarginata TRUE TRUE FALSE

emarginata. Strumigenys emarginata Mayr, 1901b: 26 (w.) SOUTH AFRICA. Combination in S. (Trichoscapa): Santschi, 1913b: 257; in S. (Cephaloxys): Wheeler, W.M. 1922a: 919; Emery, 1924d: 324; in Smithistruma: Brown, 1948e: 105; in Pyramica: Bolton, 1999: 1673. See also: Brown, 1953g: 126; Bolton, 1983: 291; Bolton, 2000: 301.

Myrmicinae Dacetini Pyramica mira TRUE TRUE FALSE

mira. Pyramica mira Bolton, 2000: 324 (w.) SOUTH AFRICA.

Myrmicinae Dacetini Pyramica noara TRUE TRUE FALSE

noara. Pyramica noara Bolton, 2000: 304 (w.) SOUTH AFRICA.

Myrmicinae Dacetini Pyramica ogyga TRUE TRUE FALSE

ogyga. Pyramica ogyga Bolton, 2000: 324, fig. 213 (w.m.) SOUTH AFRICA.

Myrmicinae Dacetini Pyramica oxysma TRUE TRUE FALSE

oxysma. Smithistruma oxysma Bolton, 1983: 315, fig. 19 (w.q.) SOUTH AFRICA. Combination in Pyramica: Bolton, 1999: 1673. See also: Bolton, 2000: 325.

Myrmicinae Dacetini Pyramica sardonica TRUE TRUE FALSE

sardonica. Pyramica sardonica Bolton, 2000: 326 (w.) SOUTH AFRICA.

Myrmicinae Dacetini Pyramica simoni TRUE TRUE FALSE

simoni. Strumigenys simoni Emery, 1895h: 42, pl. 2, fig. 21 (w.) SOUTH AFRICA. Combination in S. (Trichoscapa): Santschi, 1913b: 258 (in key); in S. (Cephaloxys): Wheeler, W.M. 1922a: 920; Emery, 1924d: 324; in Smithistruma (Serrastruma): Brown, 1948e: 107; in Serrastruma: Brown, 1952e: 82; in Pyramica: Bolton, 1999: 1673. Senior synonym of biconvexa, boerorum, cliens, cognata, escherichi, fusciventris, limbata and material of the unavailable name obscuriventris referred here: Brown, 1952e: 82; of raymondi: Bolton, 1983: 350. See also: Bolton, 2000: 318.

Myrmicinae Dacetini Pyramica transversa TRUE TRUE FALSE

transversa. Strumigenys (Trichoscapa) transversa Santschi, 1913b: 258 (diagnosis in key) (w.) SOUTH AFRICA. Santschi, 1914e: 31 (w.). Combination in S. (Cephaloxys): Wheeler, W.M. 1922a: 920; Emery, 1924d: 324; in Smithistruma: Brown, 1948e: 105; in Pyramica: Bolton, 1999: 1673. See also: Brown, 1953g: 127; Bolton, 1983: 297; Bolton, 2000: 334.

Myrmicinae Tetramoriini Rhoptromyrmex globulinodis TRUE TRUE FALSE

globulinodis. Rhoptromyrmex globulinodis Mayr, 1901b: 20 (w.q.m.) SOUTH AFRICA. Senior synonym of alberti, obscurus: Brown, 1964c: 17. See also: Bolton, 1986a: 13.

Myrmicinae Tetramoriini Rhoptromyrmex transversinodis TRUE TRUE FALSE

transversinodis. Rhoptromyrmex transversinodis Mayr, 1901b: 22 (w.) SOUTH AFRICA. Arnold, 1917: 356 (q.); Bolton, 1986a: 16 (m.). Senior synonym of pretoriae, steini: Brown, 1964c: 16.

Cerapachyinae Cerapachyini Simopone marleyi TRUE TRUE FALSE

marleyi. Simopone marleyi Arnold, 1915: 20 (w.) SOUTH AFRICA.

Myrmicinae Solenopsidini Solenopsis africana TRUE TRUE FALSE

africana. Solenopsis fugax st. africana Santschi, 1914b: 81 (w.m.) KENYA. Raised to species: Emery, 1915c: 59 (in text); Emery, 1922e: 201.

Myrmicinae Solenopsidini Solenopsis capensis TRUE TRUE FALSE

capensis. Solenopsis capensis Mayr, 1866b: 905 (q.) SOUTH AFRICA.

Myrmicinae Solenopsidini Solenopsis punctaticeps TRUE TRUE FALSE

7 subspecies

punctaticeps. Solenopsis punctaticeps Mayr, 1865: 109 (w.) SOUTH AFRICA. Forel, 1910e: 431 (q.m.). Current subspecies: nominal plus caffra, cleptomana, erythraea, fur, indocilis, juba, kibaliensis. See also: Arnold, 1916: 243.

Myrmicinae Solenopsidini Solenopsis punctaticeps caffra TRUE TRUE FALSE

caffra. Solenopsis punctaticeps var. caffra Forel, 1894b: 99 (w.q.) SOUTH AFRICA. Senior synonym of cyclops, diversipilosa: Emery, 1915c: 65.

Cerapachyinae Cerapachyini Sphinctomyrmex chariensis TRUE TRUE FALSE

chariensis. Sphinctomyrmex rufiventris st. chariensis Santschi, 1915c: 245 (m.) CENTRAL AFRICAN REPUBLIC. Combination in Aethiopopone: Santschi, 1930a: 50; in Sphinctomyrmex: Brown, 1975: 33. Raised to species: Brown, 1975: 33.

Myrmicinae Stenammini Stenamma msilanum TRUE TRUE FALSE

msilanum. Stenamma (Stenamma) westwoodi var. msilanum Forel, 1901e: 347 (q.) ALGERIA. Raised to species and senior synonym of africanum, submuticum: DuBois, 1998b: 254.

Ponerinae Ponerini Streblognathus aethiopicus TRUE TRUE FALSE

aethiopicus. Ponera aethiopica Smith, F. 1858b: 91, pl. 6, figs. 19, 20 (w.) SOUTH AFRICA. Emery, 1899e: 468 (m.); Wheeler, G.C. & Wheeler, J. 1989b: 53 (l.). Combination in Streblognathus: Mayr, 1862: 716. See also: Arnold, 1915: 41; Robertson, 2002: 10.

Ponerinae Ponerini Streblognathus peetersi TRUE TRUE FALSE

peetersi. Streblognathus peetersi Robertson, 2002: 11, figs. 1, 2, 3b, 3d, 4 (w.m.) SOUTH AFRICA.

Myrmicinae Dacetini Strumigenys faurei TRUE TRUE FALSE

faurei. Strumigenys faurei Arnold, 1948: 226, fig. 12 (w.) SOUTH AFRICA. Junior synonym of rufobrunea: Brown, 1954k: 17. Revived from synonymy: Bolton, 1983: 371. See also: Bolton, 2000: 602.

Myrmicinae Dacetini Strumigenys havilandi TRUE TRUE FALSE

havilandi. Strumigenys havilandi Forel, 1905c: 13 (footnote) (w.q.) SOUTH AFRICA. See also: Brown, 1954k: 25; Bolton, 1983: 373; Bolton, 2000: 592.

Myrmicinae Dacetini Strumigenys irrorata TRUE TRUE FALSE

irrorata. Strumigenys (Strumigenys) irrorata Santschi, 1913b: 257 (diagnosis in key) (w.) SOUTH AFRICA. Santschi, 1914e: 29 (w.). See also: Brown, 1954k: 33; Bolton, 1983: 375; Bolton, 2000: 593.

Myrmicinae Dacetini Strumigenys marleyi TRUE TRUE FALSE

marleyi. Strumigenys havilandi r. marleyi Arnold, 1914: 31, fig. 10 (w.) SOUTH AFRICA. Raised to species: Arnold, 1926: 286. See also: Brown, 1954k: 25; Bolton, 1983: 378; Bolton, 2000: 599.

Myrmicinae Dacetini Strumigenys pretoriae TRUE TRUE FALSE

pretoriae. Strumigenys pretoriae Arnold, 1949: 267, fig. 8 (w.) SOUTH AFRICA. See also: Brown, 1954k: 15; Bolton, 1983: 385; Bolton, 2000: 603.

Myrmicinae Dacetini Strumigenys rakkota TRUE TRUE FALSE

rakkota. Strumigenys rakkota Bolton, 2000: 595 (w.) SOUTH AFRICA.

Myrmicinae Dacetini Strumigenys syntacta TRUE TRUE FALSE

syntacta. Strumigenys syntacta Bolton, 2000: 597, fig. 364 (w.) SOUTH AFRICA.

Myrmicinae Dacetini Strumigenys traegaordhi TRUE TRUE FALSE

traegaordhi. Strumigenys (Strumigenys) traegaordhi Santschi, 1913b: 257 (diagnosis in key) (w.) SOUTH AFRICA. Santschi, 1914e: 28 (w.); Emery, 1915g: 19 (q.). See also: Brown, 1954k: 27; Bolton, 1983: 396; Bolton, 2000: 598.

Formicinae Plagiolepidini Tapinolepis candida TRUE TRUE FALSE

candida. Anoplolepis (Tapinolepis) candida Santschi, 1928f: 210 (w.) SOUTH AFRICA. Combination in Tapinolepis: Bolton, 2003: 268.

Formicinae Plagiolepidini Tapinolepis decolor TRUE TRUE FALSE

decolor. Plagiolepis decolor Emery, 1895h: 44 (w.) SOUTH AFRICA. Forel, 1910f: 23 (q.m.). Combination in Plagiolepis (Anoplolepis): Arnold, 1922: 599; in Anoplolepis: Emery, 1925b: 17; in Anoplolepis (Mesanoplolepis): Santschi, 1926a: 14; in Tapinolepis: Bolton, 2003: 268.

Formicinae Plagiolepidini Tapinolepis trimenii TRUE TRUE FALSE

2 subspecies

trimenii. Plagiolepis trimenii Forel, 1895g: 430 (w.) SOUTH AFRICA. Arnold, 1922: 596 (q.). Combination in Plagiolepis (Anoplolepis): Arnold, 1922: 595; in Anoplolepis: Emery, 1925b: 18; in Anoplolepis (Mesanoplolepis): Santschi, 1930b: 72; in Tapinolepis: Bolton, 2003: 268. Current subspecies: nominal plus angolensis, karrooensis.

Formicinae Plagiolepidini Tapinolepis trimenii karrooensis TRUE TRUE FALSE

karrooensis. Plagiolepis (Anoplolepis) trimeni var. karrooensis Arnold, 1922: 597 (w.q.) SOUTH AFRICA. Combination in Anoplolepis: Emery, 1925b: 18.

Dolichoderinae Tapinomini Tapinoma albinase TRUE TRUE FALSE

albinase. Technomyrmex nigriventris subsp. albinasis Forel, 1910f: 22 (w.) SOUTH AFRICA. Combination in Tapinoma: Bolton, 2007a: 121. Raised to species: Bolton, 2007a: 121.

Dolichoderinae Tapinomini Tapinoma danitschi TRUE TRUE FALSE

1 subspecies

danitschi. Tapinoma danitschi Forel, 1915c: 345 (w.) SOUTH AFRICA. Current subspecies: nominal plus bevisi.

Dolichoderinae Tapinomini Tapinoma luteum TRUE TRUE FALSE

1 subspecies

luteum. Technomyrmex luteus Emery, 1895h: 43 (w.) SOUTH AFRICA. Mayr, 1907b: 18 (q.); Wheeler, G.C. & Wheeler, J. 1966: 730 (l.). Combination in Tapinoma: Mayr, 1907b: 18. Current subspecies: nominal plus natalicum.

Dolichoderinae Tapinomini Tapinoma arnoldi tectum TRUE TRUE FALSE

tectum. Tapinoma tenue var. tectum Santschi, 1917b: 288 (w.) SOUTH AFRICA. Currently subspecies of arnoldi: Wheeler, W.M. 1922a: 923.

Dolichoderinae Tapinomini Tapinoma danitschi bevisi TRUE TRUE FALSE

bevisi. Tapinoma danitschi var. bevisi Forel, 1915c: 346 (w.q.m.) SOUTH AFRICA.

Dolichoderinae Tapinomini Technomyrmex metandrei TRUE TRUE FALSE

metandrei. Technomyrmex metandrei Bolton, 2007a: 28 (w.) CENTRAL AFRICAN REPUBLIC.

Myrmicinae Formicoxenini Terataner transvaalensis TRUE TRUE FALSE

transvaalensis. Terataner (Tranetera) transvaalensis Arnold, 1952b: 130, fig. (w.) SOUTH AFRICA. See also: Bolton, 1981b: 294.

Myrmicinae Tetramoriini Tetramorium africanum TRUE TRUE FALSE

africanum. Macromischa africana Mayr, 1866a: 507 (w.) GHANA. Mayr, 1895: 131 (q.). Combination in Tetramorium: Emery, 1896a: 103; in Macromischoides: Wheeler, W.M. 1922a: 188; in Tetramorium: Bolton, 1976: 363 (in text); Bolton, 1980: 355. Senior synonym of tessmanni: Brown, 1964c: 12; of lamottei: Bolton, 1980: 355.

Myrmicinae Tetramoriini Tetramorium akermani TRUE TRUE FALSE

akermani. Tetramorium akermani Arnold, 1926: 265, fig. 73 (w.) SOUTH AFRICA. Senior synonym of myersi: Bolton, 1980: 253.

Myrmicinae Tetramoriini Tetramorium avium TRUE TRUE FALSE

avium. Tetramorium avium Bolton, 1980: 276, figs. 58, 63, 64 (w.q.) SOUTH AFRICA.

Myrmicinae Tetramoriini Tetramorium bothae TRUE TRUE FALSE

bothae. Tetramorium simillimum subsp. bothae Forel, 1910e: 425 (w.q.m.) SOUTH AFRICA. Raised to species and senior synonym of guillarmodi: Bolton, 1980: 309.

Myrmicinae Tetramoriini Tetramorium buthrum TRUE TRUE FALSE

buthrum. Tetramorium buthrum Bolton, 1980: 309 (w.) CENTRAL AFRICAN REPUBLIC.

Myrmicinae Tetramoriini Tetramorium capense TRUE TRUE FALSE

capense. Tetramorium capense Mayr, 1865: 89 (w.) SOUTH AFRICA. Emery, 1895h: 36 (q.m.). Senior synonym of braunsi: Santschi, 1914a: 435; Bolton, 1980: 359; of popovici: Bolton, 1980: 359. See also: Prins, 1973: 8.

Myrmicinae Tetramoriini Tetramorium clunum TRUE TRUE FALSE

clunum. Tetramorium setuliferum st. cluna Forel, 1913j: 218 (w.) SOUTH AFRICA. Raised to species: Bolton, 1980: 244.

Myrmicinae Tetramoriini Tetramorium dichroum TRUE TRUE FALSE

dichroum. Tetramorium solidum st. dichroum Santschi, 1932a: 388 (w.) SOUTH AFRICA. Subspecies of peringueyi: Prins, 1973: 11. Raised to species: Bolton, 1980: 245.

Myrmicinae Tetramoriini Tetramorium dominum TRUE TRUE FALSE

dominum. Tetramorium dominum Bolton, 1980: 360, fig. 136, 139 (w.) SOUTH AFRICA.

Myrmicinae Tetramoriini Tetramorium emeryi TRUE TRUE FALSE

emeryi. Tetramorium emeryi Mayr, 1901b: 23 (w.q.m.) SOUTH AFRICA. Senior synonym of cristulatum: Bolton, 1980: 269.

Myrmicinae Tetramoriini Tetramorium erectum TRUE TRUE FALSE

erectum. Tetramorium guineense var. erectum Emery, 1895h: 37 (w.) SOUTH AFRICA. Raised to species and senior synonym of bacchus: Bolton, 1980: 269.

Myrmicinae Tetramoriini Tetramorium frenchi TRUE TRUE FALSE

frenchi. Tetramorium frenchi Forel, 1914d: 229 (w.) SOUTH AFRICA. See also: Bolton, 1980: 278.

Myrmicinae Tetramoriini Tetramorium frigidum TRUE TRUE FALSE

frigidum. Tetramorium akermani var. frigidum Arnold, 1926: 266 (w.) SOUTH AFRICA. Raised to species and senior synonym of drakensbergensis: Bolton, 1980: 256.

Myrmicinae Tetramoriini Tetramorium grandinode TRUE TRUE FALSE

grandinode. Tetramorium grandinode Santschi, 1913c: 308 (w.) SOUTH AFRICA. Senior synonym of hopensis: Bolton, 1980: 247.

Myrmicinae Tetramoriini Tetramorium grassii TRUE TRUE FALSE

grassii. Tetramorium grassii Emery, 1895h: 37 (w.) SOUTH AFRICA. Mayr, 1901b: 25 (q.). Senior synonym of algoa, joffrei, laevigatum, mayri, simulans: Bolton, 1980: 262.

Myrmicinae Tetramoriini Tetramorium inezulae TRUE TRUE FALSE

inezulae. Triglyphothrix inezulae Forel, 1914d: 221 (w.m.) SOUTH AFRICA. Combination in Tetramorium: Arnold, 1917: 343. Senior synonym of hepburni, mashonana: Bolton, 1976: 328. See also: Bolton, 1985: 247.

Myrmicinae Tetramoriini Tetramorium isipingense TRUE TRUE FALSE

isipingense. Tetramorium simillimum r. isipingense Forel, 1914d: 225 (w.) SOUTH AFRICA. Raised to species: Bolton, 1980: 347.

Myrmicinae Tetramoriini Tetramorium jauresi TRUE TRUE FALSE

jauresi. Tetramorium jauresi Forel, 1914d: 226 (w.) SOUTH AFRICA. Senior synonym of latens: Bolton, 1980: 348. See also: Prins, 1973: 17.

Myrmicinae Tetramoriini Tetramorium laevithorax TRUE TRUE FALSE

laevithorax. Tetramorium laevithorax Emery, 1895h: 39 (w.) SOUTH AFRICA. Senior synonym of jeanae: Bolton, 1980: 279.

Myrmicinae Tetramoriini Tetramorium longoi TRUE TRUE FALSE

longoi. Tetramorium longoi Forel, 1915c: 344 (w.) SOUTH AFRICA. See also: Bolton, 1980: 362.

Myrmicinae Tetramoriini Tetramorium microgyna TRUE TRUE FALSE

microgyna. Tetramorium microgyna Santschi, 1918b: 132 (q., not w.) SOUTH AFRICA. See also: Arnold, 1926: 253; Santschi, 1928f: 207; Bolton, 1980: 329.

Myrmicinae Tetramoriini Tetramorium microps TRUE TRUE FALSE

microps. Triglyphothrix microps Mayr, 1901b: 25 (w.q.) SOUTH AFRICA. Combination in Tetramorium: Arnold, 1917: 337; Bolton, 1985: 247. Senior synonym of auropunctatus, bulawayensis, fusciventris, pallens, rhodesiana: Bolton, 1976: 329.

Myrmicinae Tetramoriini Tetramorium parasiticum TRUE TRUE FALSE

parasiticum. Tetramorium parasiticum Bolton, 1980: 281, figs. 65, 66 (q.) SOUTH AFRICA.

Myrmicinae Tetramoriini Tetramorium peringueyi TRUE TRUE FALSE

peringueyi. Tetramorium peringueyi Arnold, 1926: 260 (w.) SOUTH AFRICA. See also: Prins, 1973: 9; Bolton, 1980: 248.

Myrmicinae Tetramoriini Tetramorium phasias TRUE TRUE FALSE

phasias. Tetramorium guineense var. phasias Forel, 1914d: 226 (w.) SOUTH AFRICA. Raised to species and senior synonym of hertigi: Bolton, 1980: 273.

Myrmicinae Tetramoriini Tetramorium platynode TRUE TRUE FALSE

platynode. Tetramorium platynode Bolton, 1980: 258, figs. 35, 38 (w.) SOUTH AFRICA.

Myrmicinae Tetramoriini Tetramorium poweri TRUE TRUE FALSE

poweri. Tetramorium simillimum var. poweri Forel, 1914d: 225 (w.) SOUTH AFRICA. Raised to species: Bolton, 1980: 317.

Myrmicinae Tetramoriini Tetramorium pusillum TRUE TRUE FALSE

pusillum. Tetramorium pusillum Emery, 1895h: 38 (w.q.) SOUTH AFRICA. Senior synonym of ladismithensis, tablensis: Bolton, 1980: 318.

Myrmicinae Tetramoriini Tetramorium quadrispinosum TRUE TRUE FALSE

quadrispinosum. Tetramorium quadrispinosum Emery, 1886: 362, pl. 17, fig. 8 (w.) SOUTH AFRICA. Senior synonym of montanum: Bolton, 1979: 155; of angolense, beirae, calvum, eudoxia (and its junior synonym elegans), otaviensis, repertum and material of the unavailable name benguelense referred here: Bolton, 1980: 330.

Myrmicinae Tetramoriini Tetramorium regulare TRUE TRUE FALSE

regulare. Tetramorium regulare Bolton, 1980: 263, fig. 45 (w.m.) SOUTH AFRICA.

Myrmicinae Tetramoriini Tetramorium rotundatum TRUE TRUE FALSE

rotundatum. Macromischoides africanus var. rotundatus Santschi, 1924b: 209 (q.) DEMOCRATIC REPUBLIC OF CONGO. Combination in Tetramorium: Bolton, 1980: 357. Raised to species: Bolton, 1980: 357.

Myrmicinae Tetramoriini Tetramorium setigerum TRUE TRUE FALSE

setigerum. Tetramorium setigerum Mayr, 1901b: 22 (w.) SOUTH AFRICA. Forel, 1910e: 424 (q.m.). Senior synonym of anteversa, quaerens: Bolton, 1980: 283.

Myrmicinae Tetramoriini Tetramorium setuliferum TRUE TRUE FALSE

setuliferum. Tetramorium setuliferum Emery, 1895h: 36 (w.) SOUTH AFRICA. Forel, 1910f: 19 (q.); Arnold, 1917: 290 (m.). Material of the nomen nudum squamiferum referred here by Wheeler, W.M. 1922a: 903. Senior synonym of cucalense (and its junior syonym triptolemus): Bolton, 1980: 250.

Myrmicinae Tetramoriini Tetramorium signatum TRUE TRUE FALSE

signatum. Tetramorium solidum var. signatum Emery, 1895h: 35 (w.) SOUTH AFRICA. Raised to species and senior synonym of grootensis, lugubre, tuckeri: Bolton, 1980: 251.

Myrmicinae Tetramoriini Tetramorium solidum TRUE TRUE FALSE

solidum. Tetramorium solidum Emery, 1886: 362, pl. 17, fig. 7 (w.q.) SOUTH AFRICA. See also: Prins, 1973: 4; Bolton, 1980: 252.

Myrmicinae Tetramoriini Tetramorium thoth TRUE TRUE FALSE

thoth. Triglyphothrix thoth Bolton, 1976: 337 (w.) SOUTH AFRICA. Combination in Tetramorium: Bolton, 1985: 247.

Myrmicinae Tetramoriini Tetramorium titus TRUE TRUE FALSE

titus. Tetramorium titus Forel, 1910e: 427 (w.) SOUTH AFRICA. See also: Bolton, 1980: 264.

Myrmicinae Tetramoriini Tetramorium traegaordhi TRUE TRUE FALSE

traegaordhi. Tetramorium traegaordhi Santschi, 1914e: 23 (w.q.) SOUTH AFRICA. See also: Bolton, 1980: 292.

Myrmicinae Tetramoriini Tetramorium trimeni TRUE TRUE FALSE

trimeni. Triglyphothrix trimeni Emery, 1895h: 40 (w.q.) SOUTH AFRICA. Combination in Tetramorium: Arnold, 1917: 337. See also: Bolton, 1976: 338; Bolton, 1985: 247.

Myrmicinae Tetramoriini Tetramorium vexator TRUE TRUE FALSE

vexator. Tetramorium vexator Arnold, 1926: 269, fig. 76 (w.) SOUTH AFRICA. See also: Bolton, 1980: 265.

Myrmicinae Tetramoriini Tetramorium warreni TRUE TRUE FALSE

warreni. Tetramorium warreni Arnold, 1926: 268, fig. 75 (w.) SOUTH AFRICA. See also: Bolton, 1980: 294.

Myrmicinae Tetramoriini Tetramorium weitzeckeri TRUE TRUE FALSE

weitzeckeri. Tetramorium (Xiphomyrmex) weitzeckeri Emery, 1895h: 39 (w.) SOUTH AFRICA. Arnold, 1917: 346 (q.). Combination in Xiphomyrmex: Wheeler, W.M. 1922a: 908; in Tetramorium: Bolton, 1980: 233. Senior synonym of ebeninum, edithae, escherichi, nigellus: Bolton, 1980: 233. See also: Hita Garcia, Fischer & Peters, 2010b: 85.

Pseudomyrmecinae Pseudomyrmecini Tetraponera aethiops TRUE TRUE FALSE

aethiops. Tetraponera aethiops Smith, F. 1877b: 71 (w.) SOUTH AFRICA. Santschi, 1911g: 207 (m.); Forel, 1913c: 352 (q.); Wheeler, G.C. & Wheeler, J. 1956: 392 (l.). Combination in Sima: Dalla Torre, 1893: 53; in Sima (Pachysima): Emery, 1912b: 97; in Pachysima: Wheeler, W.M. 1918c: 305; in Tetraponera: Ward, 1990: 487. Senior synonym of spininoda: Emery, 1912b: 97; Wheeler, W.M. 1922a: 801. See also: Arnold, 1916: 175; Wheeler, W.M. 1922a: 114.

Pseudomyrmecinae Pseudomyrmecini Tetraponera ambigua TRUE TRUE FALSE

ambigua. Sima ambigua Emery, 1895h: 23 (w.q.) SOUTH AFRICA. Forel, 1910f: 5 (m.). Combination in S. (Tetraponera): Arnold, 1916: 185; in Tetraponera: Wheeler, W.M. 1922a: 796. [Sima ambigua Forel, 1894b: 89. Nomen nudum, attributed to Emery.] Senior synonym of angolensis, bifoveolata, encephala, erythraea, maculifrons, occidentalis, rhodesiana, syriaca: Ward, 2006: 123.

Pseudomyrmecinae Pseudomyrmecini Tetraponera braunsi TRUE TRUE FALSE

1 subspecies

braunsi. Sima braunsi Forel, 1913a: 112 (w.q.) SOUTH AFRICA. Combination in S. (Tetraponera): Arnold, 1916: 184; in Tetraponera: Wheeler, W.M. 1922a: 797. Subspecies of clypeata: Arnold, 1916: 184; Wheeler, W.M. 1922a: 797. Revived status as species: Ward, 1990: 488. Current subspecies: nominal plus durbanensis.

Pseudomyrmecinae Pseudomyrmecini Tetraponera capensis TRUE TRUE FALSE

capensis. Pseudomyrma capensis Smith, F. 1858b: 160 (w.) SOUTH AFRICA. Combination in Sima: Mayr, 1866b: 906; in Tetraponera: Wheeler, W.M. 1922a: 797.

Pseudomyrmecinae Pseudomyrmecini Tetraponera clypeata TRUE TRUE FALSE

clypeata. Sima clypeata Emery, 1886: 361, pl. 17, figs. 4, 5 (q.) SOUTH AFRICA. Emery, 1895h: 24 (w.); Emery, 1899c: 6 (l.); Wheeler, G.C. & Wheeler, J. 1956: 390 (l.). Combination in S. (Tetraponera): Arnold, 1916: 182; in Tetraponera: Wheeler, W.M. 1922a: 797.

Pseudomyrmecinae Pseudomyrmecini Tetraponera emeryi TRUE TRUE FALSE

emeryi. Sima emeryi Forel, 1911d: 367 (w.) SOUTH AFRICA. Combination in S. (Tetraponera): Arnold, 1916: 187; in Tetraponera: Wheeler, W.M. 1922a: 797.

Pseudomyrmecinae Pseudomyrmecini Tetraponera natalensis TRUE TRUE FALSE

4 subspecies

natalensis. Pseudomyrma natalensis Smith, F. 1858b: 160 (q.) SOUTH AFRICA. Emery, 1895h: 22 (w.); Emery, 1899c: 6 (l.); Wheeler, G.C. & Wheeler, J. 1973d: 208 (l.). Combination in Sima: Emery, 1892e: 237; in Tetraponera: Smith, F. 1877b: 69; Wheeler, W.M. 1922a: 798. Junior synonym of capensis: Gerstäcker, 1871: 355. Revived from synonymy: Smith, F. 1877b: 69. Current subspecies: nominal plus caffra, cuitensis, obscurata, usambarensis. See also: Arnold, 1916: 176.

Pseudomyrmecinae Pseudomyrmecini Tetraponera natalensis obscurata TRUE TRUE FALSE

obscurata. Sima natalensis subsp. obscurata Emery, 1895h: 22 (w.) SOUTH AFRICA. Arnold, 1916: 177 (q.m.). Combination in Tetraponera: Wheeler, W.M. 1922a: 799.

Pseudomyrmecinae Pseudomyrmecini Tetraponera braunsi durbanensis TRUE TRUE FALSE

durbanensis. Sima braunsi var. durbanensis Forel, 1914d: 218 (w.) SOUTH AFRICA. Combination in Tetraponera: Wheeler, W.M. 1922a: 797.

Pseudomyrmecinae Pseudomyrmecini Tetraponera natalensis caffra TRUE TRUE FALSE

caffra. Sima natalensis st. caffra Santschi, 1914e: 15 (w.q.m.) SOUTH AFRICA. Combination in Tetraponera: Wheeler, W.M. 1922a: 799.

Dolichoderinae TRUE TRUE FALSE

Extant: 28 genera, 692 species, 130 subspecies

Fossil: 16 genera, 103 species

Dolichoderidae Forel, 1878: 364. Type-genus: Dolichoderus.

Taxonomic history

Dolichoderinae as family: Emery, 1894g: 378 [Dolichoderidae]; Ashmead, 1905b: 384 [Dolichoderidae]; Novák & Sadil, 1941: 94 [Dolichoderidae]; Bernard, 1951: 1071 [Dolichoderidae]; Bernard, 1953b: 255 [Dolichoderidae].

Dolichoderinae as tribe of Formicidae: André, 1882a: 127 [Dolichoderidae].

Dolichoderinae as subfamily of Formicidae: Forel, 1878: 364 [Dolichoderidae]; Emery & Forel, 1879a: 454 [Dolichoderidae]; Nasonov, 1889: 26 [Dolichoderidae]; Forel, 1892j: 220 [Dolichoderidae]; Forel, 1893a: 165; Dalla Torre, 1893: 156; Forel, 1895b: 107 [Dolichoderidae]; Emery, 1895j: 771 [subfamily spelled Dolichoderini]; Emery, 1896e: 186; Forel, 1899c: 98; Bingham, 1903: 288; Wheeler, W.M. 1910g: 142; Emery, 1913a: 2; Arnold, 1915: 143; Wheeler, W.M. 1915h: 71; Wheeler, 1915g: 811 [Dolichoderides]; Arnold, 1915: 143; Donisthorpe, 1915d: 178; Forel, 1917: 247; Escherich, 1917: 2 [Dolichoderini]; Bondroit, 1918: 86 [Dolichoderitae]; Wheeler, W.M. 1920: 53; Wheeler, W.M. 1922a: 199; Borgmeier, 1923: 80; Karavaiev, 1936: 163; Smith, M.R. 1951a: 833; Clark, 1951: 16; Brown, 1954e: 29; Kempf, 1972a: 266; Wheeler, G.C. & Wheeler, J. 1972a: 41; Brown, 1973b: 169; all subsequent authors.

Dolichoderinae as formicomorph subfamily of Formicidae: Bolton, 2003: 18, 80.

Dolichoderinae as formicoid subfamily of Formicidae: Brady, Schultz, et al. 2006: 18173; Moreau, Bell, et al. 2006: 102.

Dolichoderinae as formicoid dolichoderomorph subfamily of Formicidae: Ward, 2007a: 556.

Tribes (extant) of Dolichoderinae: Bothriomyrmecini, Dolichoderini, Leptomyrmecini, Tapinomini.

Tribes (extinct) incertae sedis in Dolichoderinae: †Miomyrmecini, †Zherichiniini.

Genera (extinct) incertae sedis in Dolichoderinae: †Alloiomma, †Asymphylomyrmex, †Elaeomyrmex, †Elaphrodites, †Eldermyrmex, †Emplastus, †Eotapinoma, †Eurymyrmex, †Kotshkorkia, †Leptomyrmula, †Miomyrmex, †Petraeomyrmex, †Proiridomyrmex, †Protazteca, †Zherichinius.

Subfamily Dolichoderinae and tribes references, world

Forel, 1878: 364, 380 (diagnosis, genera); Dalla Torre, 1893: 156 (catalogue); Emery, 1895j: 771 (synoptic classification); Emery, 1896e: 186 (genera key); Handlirsch, 1907: 869 (†fossil taxa catalogue); Wheeler, W.M. 1910g: 142 (diagnosis); Emery, 1913a: 2, 6 (diagnosis, tribe key, catalogue); Emery, 1913a: 17 (Tapinomini diagnosis, genera key, catalogue); Arnold, 1915: 144, (diagnosis); Gallardo, 1916a: 3 (diagnosis); Forel, 1917: 247 (synoptic classification); Forel, 1921c: 136 (diagnosis); Wheeler, W.M. 1922a: 199, 687, 688 (diagnosis, tribe key, Tapinomini genera key); Brown & Nutting, 1950: 127 (venation, phylogeny); Brown, 1954e: 29 (phylogeny); Pavan, 1955: 135 (gastral organs); Eisner, 1957: 453 (proventriculus morphology); Bernard, 1967: 246 (diagnosis); Gotwald, 1969: 118 (mouthparts morphology); Wheeler, G.C. & Wheeler, J. 1972a: 41 (diagnosis); Brown, 1973b: 169 (genera & distribution); Wheeler, G.C. & Wheeler, J. 1976b: 61 (larvae, review & synthesis); Snelling, R.R. 1981: 401 (synoptic classification); Dazzini Valcurone & Fanfani, 1985: 1 (gastral glands); Wheeler, G.C. & Wheeler, J. 1985: 258 (synoptic classification); Billen, 1986: 173 (Dufour's gland); Billen, 1987: 278 (abdominal glands); Baroni Urbani & Wilson, 1987: 1 (†fossil Leptomyrmecini); Dlussky & Fedoseeva, 1988: 77 (synoptic classification); Hölldobler & Wilson, 1990: 9 (synoptic classification, genera keys); Baroni Urbani, Bolton & Ward, 1992: 316 (phylogeny); Shattuck, 1992b: 199 (higher classification, phylogeny); Shattuck, 1992c: 20 (revision of subfamily, genera key); Shattuck, 1994: 3 (catalogue); Bolton, 1994: 22 (diagnosis, synoptic classification, genera keys); Shattuck, 1995: 217 (phylogeny, genera); Bolton, 1995a: 1038 (census); Bolton, 1995b: 10 (catalogue); Wenseleers, Schoeters, et al., 1998: 121 (cloacal gland); Brandão, Baroni Urbani, et al. 1999: 411 (phylogeny, genera); Chiotis, Jermiin & Crozier, 2000: 108 (phylogeny, genera); Dlussky & Rasnitsyn, 2002: 416 (diagnosis for impression fossils); Bolton, 2003: 18, 80 (diagnosis, synopsis); Dubovikov, 2005a: 91 (tribes of Dolichoderinae); Brady, Schultz, et al. 2006: 18173 (phylogeny); Moreau, Bell, et al. 2006: 102 (phylogeny); Ward, Brady, et al. 2010: 342 (phylogeny, tribe-rank classification).

Regional and national faunas with keys

André, 1882a: 127 (Europe & Algeria); Nasonov, 1889: 50 (Russia); Forel, 1891b: 9 (Madagascar genera); Forel, 1895e: 460 (India & Sri Lanka); Bingham, 1903: 288 (India, Sri Lanka & Burma); Ruzsky, 1905b: 102 (Russian Empire); Bondroit, 1910: 488 (Belgium); Wheeler, W.M. 1910g: 560 (North America genera); Stitz, 1914: 77 (Central Europe); Gallardo, 1915: 34 (Argentina genera); Forel, 1915d: 40 (Switzerland); Arnold, 1915: 145 (South Africa); Donisthorpe, 1915d: 178 (Britain); Gallardo, 1916a: 12 (Argentina); Emery, 1916b: 208 (Italy); Wheeler, W.M. 1916m: 589 (U.S.A., Connecticut); Bondroit, 1918: 86 (France & Belgium); Gallardo, 1919b: 253 (Argentina genera); Soudek, 1922: 58 (Czechoslovakia); Stärcke, 1926: 117 (Netherlands); Karavaiev, 1927c: 271 (Ukraine); Donisthorpe, 1927b: 199 (Britain); Menozzi & Russo, 1930: 172 (Dominican Republic); Arnol'di, 1933b: 600 (Russia); Menozzi, 1933b: 90 (Israel genera); Karavaiev, 1936: 164 (Ukraine); Smith, M.R. 1937: 861 (Puerto Rico); Stitz, 1939: 209 (Germany); Kratochvíl, 1941: 94 (Central Europe); Novák & Sadil, 1941: 94 (Central Europe); Cole, 1942: 370 (U.S.A., Utah); Smith, M.R. 1943f: 309 (U.S.A. males); Buren, 1944a: 290 (U.S.A., Iowa); Smith, M.R. 1947f: 592 (U.S.A. genera); Creighton, 1950a: 330 (North America); Kusnezov, 1956: 27 (Argentina); Brown, 1958h: 29 (New Zealand); Kusnezov, 1959: 40 (Neotropical genera); Gregg, 1963: 341 (U.S.A., Colorado); Wheeler, G.C. & Wheeler, J. 1963: 149 (U.S.A., North Dakota); Bernard, 1967: 246 (Western Europe); Wilson & Taylor, 1967: 17 (Polynesia); Bolton, 1973a: 329 (West Africa genera); Bolton & Collingwood, 1975: 3 (Britain); Boven, 1970b: 24 (Netherlands); Snelling, R.R. & Hunt, 1976: 90 (Chile); Tarbinsky, 1976: 118 (Kyrghyzstan); Boven, 1977: 123 (Belgium); Kutter, 1977c: 170 (Switzerland); Arnol'di & Dlussky, 1978: 546 (former European U.S.S.R.); Collingwood, 1978: 80 (Iberian Peninsula); Collingwood, 1979: 32 (Fennoscandia & Denmark); Greenslade, 1979: 28 (South Australia genera); Schembri & Collingwood, 1981: 420 (Malta); Allred, 1982: 443 (U.S.A., Utah); Baroni Urbani, 1984: 80 (Neotropical genera); Gösswald, 1985: 314 (Germany); Collingwood, 1985: 242 (Saudi Arabia); Wheeler, G.C. & Wheeler, J. 1986g: 55 (U.S.A., Nevada); Agosti & Collingwood, 1987b: 278 (Balkans); Dlussky, Soyunov & Zabelin, 1990: 168 (Turkmenistan); Kupyanskaya, 1990: 152 (Far Eastern Russia); Morisita, Kubota, Onoyama, et al., 1991: 2 (Japan); Atanasov & Dlussky, 1992: 49 (Bulgaria); Lattke, in Jaffe, 1993: 147 (Neotropical genera); Arakelian, 1994: 70 (Armenia); Wu, J. & Wang, 1995: 115 (China genera); Kupyanskaya, 1995: 331 (Far Eastern Russia); Collingwood & Agosti, 1996: 359 (Saudi Arabia); Seifert, 1996b: 161 (Central Europe); Skinner & Allen, 1996: 40 (Britain); Collingwood & Prince, 1998: 20 (Portugal); Kim & Kim, 1999: 18 (Korea); Shattuck, 1999: 26, 64 (Australia genera, synopsis); Andersen, 2000: 59 (northern Australia genera); Zhou, 2001b: 149 (China, Guangxi); Czechowski, Radchenko & Czechowska, 2002: 134 (Poland); Aktaç & Radchenko, 2002: 54 (Turkey genera); Yoshimura & Onoyama, 2002: 425 (Japan genera, males); Mackay & Mackay, 2002: 221 (U.S.A., New Mexico); Palacio & Fernández, in Fernández, 2003d: 239 (Neotropical genera); Radchenko, 2005b: 186 (North Korea); Coovert, 2005: 102 (U.S.A., Ohio); Clouse, 2007b: 190 (Micronesia); Seifert, 2007: 147 (North and Central Europe); Fisher, 2009: 51 (Malagasy genera); Terayama, 2009: 196 (Taiwan); Heterick, 2009: 28 (south-western Australia genera); Boer, 2010: 14 (Benelux); Yoshimura & Fisher, 2011: 7 (Malagasy males key).

Formicinae TRUE TRUE FALSE

Extant: 49 genera, 2,831 species, 839 subspecies

Fossil: 29 genera, 193 species, 1 subspecies

Formicariae Latreille, 1809: 124. Type-genus: Formica.

Taxonomic history

Formicinae as group name: Latreille, 1809: 124 [Formicariae]; Lepeletier de Saint-Fargeau, 1835: 197 [Formicites]; Nylander, 1846a: 877 [Formicae].

Formicinae as family: Smith, F. 1858b: 1 [Formicidae]; Smith, F. 1861b: 36 [Formicidae]; Smith, 1871a: 302 [Formicidae]; André, 1882a: 125 [Formicidae]; Saunders, 1896: 18 [Formicidae]; Novák & Sadil, 1941: 97 [Formicidae]; Bernard, 1951: 1073 [Formicidae]; Bernard, 1953b: 256 [Formicidae].

Formicinae as tribe of Formicidae: André, 1874: 167 [Formicidae].

Formicinae as subfamily of Formicidae: Mayr, 1855: 286, 299 [Formicidae]; Mayr, 1861: 21 [Formicidae]; Mayr, 1862: 651 [Formicidae]; Mayr, 1865: 6 [Formicidae]; Mayr, 1868c: 24 [Formicidae]; Forel, 1870: 307 [Formicidae]; Forel, 1874: 21 [Formicidae]; Emery, 1877a: 70 [Formicidae]; André, 1881b: 54 [Formicidae]; Ashmead, 1905b: 384; Bondroit, 1918: 17 [Formicitae]; Wheeler, W.M. 1920: 53; Wheeler, W.M. 1922a: 210; Emery, 1925b: 2; Karavaiev, 1936: 172; Clark, 1951: 16; Brown, 1954e: 29; Kempf, 1972a: 266; Brown, 1973b: 169; all subsequent authors.

Formicinae as formicomorph subfamily of Formicidae: Bolton, 2003: 20, 93.

Formicinae as formicoid subfamily of Formicidae: Brady, Schultz, et al. 2006: 18173; Moreau, Bell, et al. 2006: 102.

Formicinae as formicoid formicomorph subfamily of Formicidae: Ward, 2007a: 556.

Tribes of Formicinae: Camponotini, Formicini, Gesomyrmecini, Gigantiopini, Lasiini, Melophorini, Myrmecorhynchini, Myrmoteratini, Notostigmatini, Oecophyllini, Plagiolepidini.

Genera (extinct) incertae sedis in Formicinae: †Camponotites, †Drymomyrmex, †Imhoffia, †Kyromyrma, †Leucotaphus, †Protrechina, †Zhangidris.

Hong (2002) genera (extinct) incertae sedis in Formicinae: †Curtipalpulus, †Eoleptocerites, †Eurytarsites, †Fushuniformica, †Huaxiaformica, †Leptogasteritus, †Liaoformica, †Longiformica, †Magnogasterites, †Orbicapitia, †Ovalicapito, †Ovaligastrula, †Sinoformica, †Sinotenuicapito, †Wilsonia (unresolved junior homonym).

Subfamily Formicinae references, world

Mayr, 1862: 651 (genera key); Mayr, 1865: 6 (diagnosis); Handlirsch, 1907: 859 (†fossil taxa catalogue); Dalla Torre, 1893: 171 (catalogue); Emery, 1895j: 772 (synoptic classification); Emery, 1896e: 187 (genera key); Wheeler, W.M. 1910g: 143 (diagnosis); Forel, 1912i: 88 (tribes key); Forel, 1917: 248 (synoptic classification); Arnold, 1920a: 551 (diagnosis); Forel, 1921c: 139 (diagnosis); Wheeler, W.M. 1922a: 210, 691 (diagnosis, tribes key); Emery, 1925b: 2 (diagnosis, tribe key, catalogue); Brown & Nutting, 1950: 127 (venation, phylogeny); Eisner, 1957: 465 (proventriculus morphology); Hung & Brown, 1966: 198 (gastric apex, structure); Bernard, 1967: 267 (diagnosis); Gotwald, 1969: 120 (mouthparts morphology); Wheeler, G.C. & Wheeler, J. 1972a: 41 (diagnosis); Brown, 1973b: 169 (genera, distribution); Wheeler, G.C. & Wheeler, J. 1976b: 62 (larvae, review & synthesis); Snelling, R.R. 1981: 402 (synoptic classification); Wheeler, G.C. & Wheeler, J. 1985: 258 (synoptic classification); Billen, 1986: 173 (Dufour's gland); Dlussky & Fedoseeva, 1988: 77 (synoptic classification); Hölldobler & Wilson, 1990: 9 (synoptic classification, genera keys); Agosti, 1991: 295 (genus group diagnoses); Shattuck, 1992b: 201 (phylogeny); Baroni Urbani, Bolton & Ward, 1992: 317 (phylogeny); Bolton, 1994: 42 (diagnosis, synoptic classification, genera keys); Bolton, 1995a: 1039 (census); Bolton, 1995b: 11 (catalogue); Wenseleers, Schoeters, et al. 1998: 121 (cloacal gland); Dlussky & Rasnitsyn, 2002: 417 (diagnosis for impression fossils); Bolton, 2003: 20, 93 (diagnosis, synopsis); Brady, Schultz, et al. 2006: 18173 (phylogeny); Moreau, Bell, et al. 2006: 102 (phylogeny).

Regional and national faunas with keys

Mayr, 1855: 299 (Austria); Mayr, 1861: 25 (Europe); Mayr, 1868c: 25 (†Baltic Amber); André, 1874: 167 (Europe); Forel, 1874: 22 (Switzerland); Saunders, E. 1880: 203 (Britain); André, 1882a: 126 (Europe & Algeria); Provancher, 1887: 225 (Canada); Cresson, 1887: 94 (U.S.A. genera); Nasonov, 1889: 50 (Russia); Forel, 1891b: 8 (Madagascar genera); Lameere, 1892: 62 (Belgium); Forel, 1892j: 220 (India & Sri Lanka); Bingham, 1903: 308 (India, Sri Lanka & Burma); Ruzsky, 1905b: 100 (Russian Empire); Wasmann, 1906: 7 (Luxemburg); Bondroit, 1910: 481 (Belgium); Wheeler, W.M. 1910g: 560 (North America genera); Stitz, 1914: 80 (Central Europe); Gallardo, 1915: 35 (Argentina genera); Forel, 1915d: 45 (Switzerland); Donisthorpe, 1915d: 184 (Britain); Emery, 1916b: 216 (Italy); Wheeler, W.M. 1916m: 590 (U.S.A., Connecticut); Bondroit, 1918: 17 (France & Belgium); Arnold, 1920a: 552 (South Africa); Kutter, 1920b: 134 (Switzerland); Soudek, 1922: 61 (Czechoslovakia); Lomnicki, 1925a: 160 (Poland); Stärcke, 1926: 118, 146 (Netherlands); Karavaiev, 1927c: 273 (Ukraine); Donisthorpe, 1927b: 205 (Britain); Menozzi & Russo, 1930: 172 (Dominican Republic); Arnol'di, 1933b: 601 (Russia); Menozzi, 1933b: 90 (Israel genera); Karavaiev, 1936: 173 (Ukraine); Smith, M.R. 1937: 865 (Puerto Rico); Stitz, 1939: 230 (Germany); Kratochvíl, 1941: 97 (Central Europe); Novák & Sadil, 1941: 97 (Central Europe); Cole, 1942: 373 (U.S.A., Utah); Smith, M.R. 1943f: 309 (U.S.A., males); Buren, 1944a: 292 (U.S.A., Iowa); Holgersen, 1943b: 173 (Norway); Holgersen, 1944: 199 (Norway); Smith, M.R. 1947f: 599 (U.S.A. genera); Boven, 1947: 181 (Belgium); Creighton, 1950a: 355 (North America); Kusnezov, 1956: 31 (Argentina); Brown, 1958h: 42 (New Zealand); Boven, 1959: 11 (Netherlands); Gregg, 1963: 447 (U.S.A., Colorado); Wheeler, G.C. & Wheeler, J. 1963: 160 (U.S.A., North Dakota); Collingwood, 1964: 104 (Britain); Bernard, 1967: 268 (Western Europe); Wilson & Taylor, 1967: 17 (Polynesia); Boven, 1970b: 26 (Netherlands); Kempf, 1972a: 266 (Neotropical, synoptic classification); Bolton, 1973a: 329 (West Africa genera); Bolton & Collingwood, 1975: 3 (Britain); Snelling & Hunt, 1976: 104 (Chile); Tarbinsky, 1976: 126 (Kyrghyzstan); Boven, 1977: 126 (Belgium); Kutter, 1977c: 183 (Switzerland); Arnol'di & Dlussky, 1978: 548 (former European U.S.S.R.); Collingwood, 1978: 88 (Iberian Peninsula); Collingwood, 1979: 85 (Fennoscandia & Denmark); Greenslade, 1979: 32 (South Australia genera); Schembri & Collingwood, 1981: 436 (Malta); Prins, 1983: 8 (Southern Africa genera); Allred, 1982: 444 (U.S.A., Utah); Verhaeghe, Deligne, et al., 1984: 106 (Belgium genera); Baroni Urbani, 1984: 81 (Neotropical genera); Gösswald, 1985: 263 (Germany); Collingwood, 1985: 273 (Saudi Arabia); Wheeler, G.C. & Wheeler, J. 1986g: 58 (U.S.A., Nevada); Nilsson & Douwes, 1987: 68 (Norway); Agosti & Collingwood, 1987b: 279 (Balkans); Dlussky, Soyunov & Zabelin, 1990: 124 (Turkmenistan); Kupyanskaya, 1990: 162 (Far Eastern Russia); Morisita, Kubota, Onoyama, et al., 1991: 10 (Japan); Atanasov & Dlussky, 1992: 49 (Bulgaria); Shattuck, 1992b: 199 (higher classification, phylogeny); Lattke, in Jaffe, 1993: 150 (Neotropical genera); Arakelian, 1994: 76 (Armenia); Wu, J. & Wang, 1995: 125 (China genera); Kupyanskaya, 1995: 332 (Far Eastern Russia); Collingwood & Agosti, 1996: 361 (Saudi Arabia); Seifert, 1996b: 166 (Central Europe); Skinner & Allen, 1996: 41 (Britain); Collingwood & Prince, 1998: 21 (Portugal); Shattuck, 1999: 25, 86 (Australia genera, synopsis); Andersen, 2000: 68 (northern Australia genera); Zhou, 2001b: 165 (China, Guangxi); Czechowski, Radchenko & Czechowska, 2002: 147 (Poland); Aktaç & Radchenko, 2002: 54 (Turkey genera); Yoshimura & Onoyama, 2002: 425 (Japan genera, males); Mackay & Mackay, 2002: 236 (U.S.A., New Mexico); Palacio & Fernández, in Fernández, 2003d: 242 (Neotropical genera); Radchenko, 2005b: 187 (North Korea); Coovert, 2005: 113 (U.S.A., Ohio); Clouse, 2007b: 190 (Micronesia); Seifert, 2007: 150 (North and Central Europe); Terayama, 2009: 202 (Taiwan); Heterick, 2009: 30 (south-western Australia genera); Boer, 2010: 17 (Benelux).

Myrmicinae TRUE TRUE FALSE

Extant: 141 genera, 6,087 species, 864 subspecies

Fossil: 31 genera, 130 species

Myrmicites Lepeletier de Saint-Fargeau, 1835: 169. Type-genus: Myrmica.

Taxonomic history

Myrmicinae as group name: Lepeletier de Saint-Fargeau, 1835: 169 [Myrmicites]; Nylander, 1846a: 877 [Myrmicae].

Myrmicinae as family: Smith, F. 1851: 4 [Myrmicidae]; Smith, F. 1861b: 45 [Myrmicidae]; Smith, F. 1871a: 324 [Myrmicidae]; André, 1882a: 125 [Myrmicidae]; Cresson, 1887: 93 [Myrmicidae]; Emery, 1894g: 383 [Myrmicidae]; Saunders, 1896: 18 [Myrmicidae]; Ashmead, 1905b: 383 [Myrmicidae]; Novák & Sadil, 1941: 71 [Myrmicidae]; Bernard, 1951: 1058 [Myrmicidae]; Bernard, 1953b: 222 [Myrmicidae].

Myrmicinae as subfamily of Poneridae: Smith, F. 1858b: 114 [Myrmicidae].

Myrmicinae as tribe of Formicidae: André, 1874: 167 [Myrmicidae].

Myrmicinae as subfamily of Myrmicidae: Ashmead, 1905b: 383.

Myrmicinae as subfamily of Formicidae: Mayr, 1855: 290, 299 [Myrmicidae]; Smith, F. 1857a: 70 [Myrmicidae]; Mayr, 1861: 21 [Myrmicidae]; Smith, F. 1862b: 33 [Myrmicidae]; Mayr, 1862: 738 [Myrmicidae]; Mayr, 1865: 17 [Myrmicidae]; Mayr, 1868c: 24 [Myrmicidae]; Forel, 1870: 307 [Myrmicidae]; Forel, 1874: 22 [Myrmicidae]; Emery, 1877a: 70 [Myrmicidae]; Forel, 1878: 367 [Myrmicidae]; Emery & Forel, 1879a: 456 [Myrmicidae]; André, 1881b: 64 [Myrmicidae]; Nasonov, 1889: 28 [Myrmicidae]; Forel, 1891b: 11 [Myrmicidae]; Forel, 1892j: 220 [Myrmicidae]; Forel, 1893a: 163; Dalla Torre, 1893: 53; Emery, 1895j: 768 [subfamily spelled Myrmicini]; Emery, 1896e: 179; Forel, 1899c: 30; Forel, 1902f: 520; Bingham, 1903: 105; Wheeler, W.M. 1910g: 138; Emery, 1914a: 29; Wheeler, 1915g: 806 [Myrmicides]; Wheeler, W.M. 1915h: 40; Donisthorpe, 1915d: 74; Arnold, 1916: 166; Escherich, 1917: 2 [Myrmicini]; Forel, 1917: 240 [subfamily spelled Myrmicini]; Bondroit, 1918: 14 [Myrmicitae]; Wheeler, W.M. 1920: 53; Wheeler, W.M. 1922a: 124; Emery, 1921f: 3; Karavaiev, 1934: 59; Clark, 1951: 16; Brown, 1954e: 28; Wheeler, G.C. & Wheeler, J. 1972a: 40; Brown, 1973b: 166; all subsequent authors.

Myrmicinae as myrmicomorph subfamily of Formicidae: Bolton, 2003: 52, 182.

Myrmicinae as formicoid subfamily of Formicidae: Moreau, Bell, et al. 2006: 102; Brady, Schultz, et al. 2006: 18173.

Myrmicinae as formicoid myrmicomorph subfamily of Formicidae: Ward, 2007a: 556.

Tribes of Myrmicinae: Adelomyrmecini, Ankylomyrmini, Attini, Basicerotini, Blepharidattini, Cataulacini, Cephalotini, Crematogastrini, Dacetini, Formicoxenini, Lenomyrmecini, Liomyrmecini, Melissotarsini, Meranoplini, Metaponini, Myrmecinini, Myrmicariini, Myrmicini, Paratopulini, Phalacromyrmecini, Pheidolini, Solenopsidini, Stegomyrmecini, Stenammini, Tetramoriini.

Genera (extant) incertae sedis in Myrmicinae: Diaphoromyrma, Tyrannomyrmex.

Genera (extinct) incertae sedis in Myrmicinae: †Afromyrma, †Attopsis, †Cephalomyrmex, †Electromyrmex, †Eocenidris, †Eomyrmex, †Fallomyrma, †Miosolenopsis, †Myrmecites, †Solenopsites.

Hong (2002) genera (extinct) incertae sedis in Myrmicinae: †Brachytarsites, †Clavipetiola, †Fushunomyrmex, †Orbigastrula, †Quadrulicapito, †Quineangulicapito, †Sinomyrmex, †Sphaerogasterites, †Wumyrmex.

Collective group name in Myrmicinae: †Myrmicites.

Subfamily Myrmicinae references, world

Mayr, 1865: 17 (diagnosis); Mayr, 1867a: 91 (diagnosis); Forel, 1878: 367 (diagnosis); Handlirsch, 1907: 872 (†fossil taxa catalogue); Dalla Torre, 1893: 53 (catalogue); Emery, 1895j: 768 (diagnosis); Emery, 1896e: 179 (genera key); Wheeler, W.M. 1910g: 138 (diagnosis); Emery, 1912b: 101 (phylogeny); Emery, 1914a: 34 (phylogeny, tribe key); Arnold, 1916: 164 (diagnosis); Forel, 1917: 240 (synoptic classification); Forel, 1921c: 139 (diagnosis); Emery, 1921f: 3 (diagnosis, tribes & genera key, catalogue); Wheeler, W.M. 1922a: 124, 655 (diagnosis, tribes key); Brown & Nutting, 1950: 126 (venation, phylogeny); Brown, 1954e: 28 (phylogeny); Eisner, 1957: 477 (proventriculus morphology); Bernard, 1967: 93 (diagnosis); Gotwald, 1969: 99 (mouthparts morphology); Wheeler, G.C. & Wheeler, J. 1972a: 40 (diagnosis); Brown, 1973b: 166 (genera, distribution); Wheeler, G.C. & Wheeler, J. 1976b: 52 (larvae, review & synthesis); Kugler, C. 1978a: 413 (sting structure); Kugler, C. 1978b: 267 (pygidial glands); Kugler, C. 1979c: 117 (sting, evolution); Snelling, R.R. 1981: 393 (synoptic classification); Caetano, F.H. 1984: 257 (digestive tract, morphology); Wheeler, G.C. & Wheeler, J. 1985: 257 (synoptic classification); Billen, 1986: 167 (Dufour's gland); Dlussky & Fedoseeva, 1988: 79 (synoptic classification); Hölldobler & Wilson, 1990: 9 onward (synoptic classification, genera keys); Baroni Urbani, Bolton & Ward, 1992: 317 (phylogeny); Bolton, 1994: 75 (diagnosis, synoptic classification, genera keys); Bolton, 1995a: 1040 (census); Bolton, 1995b: 13 (catalogue); Hashimoto, 1996: 354 (phylogenetic position); Baroni Urbani, 2000: 480 (phylogeny); Dlussky & Rasnitsyn, 2002: 422 (diagnosis for wingless fossils); Bolton, 2003: 52, 182 (diagnosis, synopsis); Moreau, Bell, et al. 2006: 102 (phylogeny); Brady, Schultz, et al. 2006: 18173 (phylogeny); Ward, 2007a: 556 (classification).

Regional and national faunas with keys

Mayr, 1855: 391 (Austria); Mayr, 1861: 29 (Europe); Mayr, 1868c: 79 (†Baltic Amber); André, 1874: 171 (Europe); Forel, 1874: 29 (Switzerland); Saunders, E. 1880: 213 (Britain); André, 1882c: 256 (Europe & Algeria); Cresson, 1887: 98 (U.S.A. genera); Provancher, 1887: 243 (Canada); Nasonov, 1889: 54 (Russia); Forel, 1891b: 11 (Madagascar genera); Lameere, 1892: 66 (Belgium); Forel, 1902f: 520 (India & Sri Lanka genera); Bingham, 1903: 105 (India, Sri Lanka & Burma); Ruzsky, 1905b: 103 (Russian Empire); Wasmann, 1906: 13 (Luxemburg); Bondroit, 1910: 490 (Belgium); Wheeler, W.M. 1910g: 558 (North America genera); Stitz, 1914: 55 (Central Europe); Gallardo, 1915: 32 (Argentina genera); Forel, 1915d: 8 (Switzerland); Donisthorpe, 1915d: 74 (Britain); Arnold, 1916: 166, 170 (South Africa tribes, genera); Emery, 1916b: 112 (Italy); Wheeler, W.M. 1916m: 581 (U.S.A., Connecticut); Bondroit, 1918: 90 (France & Belgium); Kutter, 1920b: 144 (Switzerland); Soudek, 1922: 20 (Czechoslovakia); Stärcke, 1926: 84 (Netherlands); Karavaiev, 1927c: 256 (Ukraine); Donisthorpe, 1927b: 77 (Britain); Menozzi & Russo, 1930: 170 (Dominican Republic); Gallardo, 1932b: 91 (Argentina, tribes); Arnol'di, 1933b: 596 (Russia); Menozzi, 1933b: 88 (Israel genera); Karavaiev, 1934: 60 (Ukraine); Smith, M.R. 1937: 829 (Puerto Rico); Stitz, 1939: 63 (Germany); Kratochvíl, 1941: 71 (Central Europe); Novák & Sadil, 1941: 71 (Central Europe); Cole, 1942: 360 (U.S.A., Utah); Smith, M.R. 1943f: 291 (U.S.A., males); Holgersen, 1943b: 166 (Norway); Holgersen, 1944: 198 (Norway); Buren, 1944a: 281 (U.S.A., Iowa); Smith, M.R. 1947f: 543 (U.S.A. genera); Boven, 1947: 170 (Belgium); Creighton, 1950a: 83 (North America); Kusnezov, 1956: 15 (Argentina); Brown, 1958h: 25 (New Zealand); Boven, 1959: 7 (Netherlands); Gregg, 1963: 288 (U.S.A., Colorado); Wheeler, G.C. & Wheeler, J. 1963: 92 (U.S.A., North Dakota); Collingwood, 1964: 94 (Britain); Bernard, 1967: 95 (Western Europe); Wilson & Taylor, 1967: 13 (Polynesia); Boven, 1970b: 9 (Netherlands); Kempf, 1972a: 263 (Neotropical, synoptic classification); Bolton, 1973a: 325 (West Africa genera); Bolton & Collingwood, 1975: 3 (Britain); Snelling, R.R. & Hunt, 1976: 70 (Chile); Tarbinsky, 1976: 19 (Kyrghyzstan); Boven, 1977: 69 (Belgium); Kutter, 1977c: 31 (Switzerland); Arnol'di & Dlussky, 1978: 524 (former European U.S.S.R.); Collingwood, 1978: 75 (Iberian Peninsula); Collingwood, 1979: 36 (Fennoscandia & Denmark); Greenslade, 1979: 20 (South Australia genera); Schembri & Collingwood, 1981: 423 (Malta); Allred, 1982: 438 (U.S.A., Utah); Baroni Urbani, 1984: 76 (Neotropical genera); Verhaeghe, Deligne, et al. 1984: 112 (Belgium genera); Gösswald, 1985: 289 (Germany); Collingwood, 1985: 245 (Saudi Arabia); Wheeler, G.C. & Wheeler, J. 1986g: 20 (U.S.A., Nevada); Nilsson & Douwes, 1987: 57 (Norway); Agosti & Collingwood, 1987b: 265 (Balkans); Dlussky, Soyunov & Zabelin, 1990: 181 (Turkmenistan); Kupyanskaya, 1990: 89 (Far Eastern Russia); Ogata, 1991b: 61 (Japan genera); Morisita, Kubota, et al. 1992: 1 (Japan); Atanasov & Dlussky, 1992: 51 (Bulgaria); Lattke, in Jaffe, 1993: 153 (Neotropical genera); Arakelian, 1994: 15 (Armenia); Wu, J. & Wang, 1995: 57 (China genera); Kupyanskaya, 1995: 327 (Far Eastern Russia); Collingwood & Agosti, 1996: 308 (Saudi Arabia); Seifert, 1996b: 108 (Central Europe); Skinner & Allen, 1996: 43 (Britain); Collingwood & Prince, 1998: 10 (Portugal); Shattuck, 1999: 39, 122 (Australia genera, synopsis); Andersen, 2000: 37 (northern Australia genera); Zhou, 2001b: 69 (China, Guangxi); Czechowski, Radchenko & Czechowska, 2002: 135 (Poland); Aktaç & Radchenko, 2002: 55 (Turkey genera); Yoshimura & Onoyama, 2002: 424 (Japan genera, males); Mackay & Mackay, 2002: 58 (U.S.A., New Mexico); Palacio & Fernández, in Fernández, 2003d: 244 (Neotropical genera and synopsis); Coovert, 2005: 32 (U.S.A., Ohio); Radchenko, 2005b: 184 (North Korea); Clouse, 2007b: 190 (Micronesia); Seifert, 2007: 110 (North and Central Europe); Terayama, 2009: 131 (Taiwan); Heterick, 2009: 36 (south-western Australia genera); Boer, 2010: 46 (Benelux); Eguchi, Bui & Yamane, 2011: 8 (Vietnam genera).

Ponerinae TRUE TRUE FALSE

Extant: 27 genera, 1,033 species, 155 subspecies

Fossil: 9 genera, 68 species, 2 subspecies

Ponerites Lepeletier de Saint-Fargeau, 1835: 185. Type-genus: Ponera.

Taxonomic history

Ponerinae as group name: Lepeletier de Saint-Fargeau, 1835: 185 [Ponérites].

Ponerinae as family: Smith, F. 1851: 6 [Poneridae]; Smith, F. 1858b: 76 [Poneridae]; Smith, F. 1861b: 44 [Poneridae]; Smith, F. 1871a: 320 [Poneridae]; André, 1882a: 125 [Poneridae]; Cresson, 1887: 93 [Poneridae]; Emery, 1894g: 379 [Poneridae]; Saunders, 1896: 18 [Poneridae]; Ashmead, 1905b: 382 [Poneridae]; Novák & Sadil, 1941: 70 [Poneridae]; Bernard, 1951: 1042 [Poneridae]; Bernard, 1953b: 185 [Poneridae].

Ponerinae as tribe of Formicidae: André, 1874: 167 [Poneridae].

Ponerinae as subfamily of Poneridae: Ashmead, 1905b: 382.

Ponerinae as subfamily of Formicidae: Mayr, 1855: 289, 299 [Poneridae]; Mayr, 1861: 21 [Poneridae]; Mayr, 1862: 712 [Poneridae]; Smith, F. 1857a: 64 [Poneridae]; Smith, F. 1862b: 31 [Poneridae]; Mayr, 1865: 11 [Poneridae]; Mayr, 1868c: 24 [Poneridae]; Forel, 1870: 307 [Poneridae]; Forel, 1874: 21 [Poneridae]; Emery, 1877a: 70 [Poneridae]; Forel, 1878: 366 [Poneridae]; Emery & Forel, 1879a: 455 [Poneridae]; André, 1881b: 64 [Poneridae]; Forel, 1892j: 220 [Poneridae]; Dalla Torre, 1893: 13; Forel, 1893a: 162; Forel, 1895b: 111 [Poneridae]; Nasonov, 1889: 27 [Poneridae]; Emery, 1895j: 766 [subfamily spelled Ponerini]; Emery, 1896e: 176; Forel, 1899c: 2; Emery, 1901a: 36; Bingham, 1903: 23; Wheeler, W.M. 1910g: 134; Emery, 1911d: 2; Wheeler, 1915g: 805 [Ponerides]; Donisthorpe, 1915d: 65; Wheeler, W.M. 1915h: 25; Arnold, 1915: 9; Escherich, 1917: 2 [Ponerini]; Forel, 1917: 235; Bondroit, 1918: 78 [Poneritae]; Wheeler, W.M. 1920: 53; Wheeler, W.M. 1922a: 56, 632, 640; Borgmeier, 1923: 37; Karavaiev, 1934: 49; Clark, 1951: 15; Brown, 1954e: 24; Wheeler, G.C. & Wheeler, J. 1972a: 39; Brown, 1973b: 165; all subsequent authors.

Ponerinae as poneromorph subfamily of Formicidae: Bolton, 2003: 42, 156.

Ponerinae as poneroid subfamily of Formicidae: Ouellette, Fisher, et al. 2006: 365; Brady, Schultz, et al. 2006: 18173; Moreau, Bell et al. 2006: 102; Ward, 2007a: 555.

Tribes of Ponerinae: Platythyreini, Ponerini, Thaumatomyrmecini.

Genera (extinct) incertae sedis in Ponerinae: †Afropone, †Eogorgites, †Eoponerites, †Furcisutura, †Longicapitia.

Subfamily Ponerinae references, world

Smith, F. 1858b: 76 (diagnosis); Mayr, 1862: 712 (genera key); Mayr, 1865: 11 (Odontomachidae, Poneridae diagnoses), Mayr, 1867a: 79, 81 (Odontomachidae, Poneridae diagnoses); Forel, 1878: 366 (diagnosis); Dalla Torre, 1893: 13 (world catalogue); Emery, 1895j: 766 (diagnosis); Emery, 1896e: 176 (genera key); Handlirsch, 1907: 879 (†fossil taxa catalogue); Wheeler, W.M. 1910g: 134 (diagnosis); Emery, 1911d: 3 (diagnosis, tribes key); Forel, 1917: 235 (synoptic classification); Forel, 1921c: 133 (diagnosis); Wheeler, W.M. 1922a: 56, 636 (diagnosis, tribes key); Brown & Nutting, 1950: 124 (venation, phylogeny); Brown, 1954e: 24 (phylogeny); Eisner, 1957: 475 (proventriculus morphology); Bernard, 1967: 79 (diagnosis); Gotwald, 1969: 25 (mouthparts morphology); Wheeler, G.C. & Wheeler, J. 1972a: 39 (diagnosis); Brown, 1973b: 165 (genera, distribution); Wheeler, G.C. & Wheeler, J. 1976b: 48 (larvae, review & synthesis); Snelling, R.R. 1981: 387 (synoptic classification); Wheeler, G.C. & Wheeler, J. 1985: 256 (synoptic classification); Billen, 1986: 168 (Dufour's gland); Fanfani & Dazzini Valcurone, 1986: 115 (exocrine glands synopsis); Dlussky & Fedoseeva, 1988: 78 (synoptic classification); Hölldobler & Wilson, 1990: 9 onward (synoptic classification, genera keys); Baroni Urbani, Bolton & Ward, 1992: 317 (phylogeny); Bolton, 1994: 153 (diagnosis, synoptic classification, genera keys); Bolton, 1995a: 1042 (census); Bolton, 1995b: 14 (catalogue); Grimaldi, Agosti & Carpenter, 1997: 20 (†Cretaceous genera); Dlussky & Rasnitsyn, 2002: 422 (diagnosis for wingless fossils); Bolton, 2003: 42, 156 (diagnosis, synopsis); Ouellette, Fisher et al. 2006: 359 (phylogeny); Brady, Schultz, et al. 2006: 18173 (phylogeny); Moreau, Bell et al. 2006: 102 (phylogeny); Ward, 2007a: 555 (classification).

.

Regional and national faunas with keys

Mayr, 1861: 28 (Europe); Mayr, 1868c: 69 (†Baltic Amber); André, 1874: 170 (Europe); Forel, 1874: 29 (Switzerland); André, 1882b: 227 (Europe & Algeria); Cresson, 1887: 97 (U.S.A., genera); Provancher, 1887: 238 (Canada); Nasonov, 1889: 54 (Russia); Forel, 1891b: 9 (Madagascar genera); Forel, 1900c: 52 (India & Sri Lanka); Bingham, 1903: 23 (India, Sri Lanka & Burma); Ruzsky, 1905b: 108 (Russian Empire); Wheeler, W.M. 1910g: 557 (North America genera); Bondroit, 1910: 489 (Belgium); Stitz, 1914: 54 (Central Europe); Gallardo, 1915: 31 (Argentina genera); Forel, 1915d: 6 (Switzerland); Arnold, 1915: 10 (South Africa); Donisthorpe, 1915d: 65 (Britain); Emery, 1916b: 98 (Italy); Wheeler, W.M. 1916m: 580 (U.S.A., Connecticut); Bondroit, 1918: 79 (France & Belgium); Gallardo, 1918b: 4 (Argentina); Soudek, 1922: 18 (Czechoslovakia); Stärcke, 1926: 82 (Netherlands); Donisthorpe, 1927b: 68 (Britain); Menozzi & Russo, 1930: 169 (Dominican Republic); Arnol'di, 1933b: 596 (Russia); Karavaiev, 1934: 50 (Ukraine); Smith, M.R. 1937: 823 (Puerto Rico); Stitz, 1939: 58 (Germany); Smith, M.R. 1943f: 278 (U.S.A. males); Buren, 1944a: 279 (U.S.A., Iowa); Smith, M.R. 1947f: 530 (U.S.A. genera); Creighton, 1950a: 31 (North America); Kusnezov, 1956: 11 (Argentina); Brown, 1958h: 11 (New Zealand); Gregg, 1963: 280 (U.S.A., Colorado); Bernard, 1967: 80 (Western Europe); Wilson & Taylor, 1967: 10 (Polynesia); Kempf, 1972a: 262 (Neotropical synoptic classification); Bolton, 1973a: 323 (West Africa genera); Bolton & Collingwood, 1975: 3 (Britain); Boven, 1977: 66 (Belgium); Kutter, 1977c: 21 (Switzerland); Arnol'di & Dlussky, 1978: 522 (former European U.S.S.R.); Collingwood, 1978: 74 (Iberian Peninsula); Collingwood, 1979: 29 (Fennoscandia & Denmark); Greenslade, 1979: 14 (South Australia genera); Francoeur, 1979: 30 (Canada, Québec); Schembri & Collingwood, 1981: 417 (Malta); Allred, 1982: 438 (U.S.A., Utah); Baroni Urbani, 1984: 75 (Neotropical genera); Collingwood, 1985: 236 (Saudi Arabia); Wheeler, G.C. & Wheeler, J. 1986g: 17 (U.S.A., Nevada); Agosti & Collingwood, 1987b: 264 (Balkans); Ogata, 1987: 101 (Japan genera); Morisita, Kubota, et al. 1989: 8 (Japan); Dlussky, Soyunov & Zabelin, 1990: 173 (Turkmenistan); Kupyanskaya, 1990: 85 (Far Eastern Russia); Atanasov & Dlussky, 1992: 51 (Bulgaria); Lattke, in Jaffe, 1993: 166 (Neotropical genera); Arakelian, 1994: 10 (Armenia); Wu, J. & Wang, 1995: 31 (China genera); Collingwood & Agosti, 1996: 308 (Saudi Arabia); Seifert, 1996b: 107 (Central Europe); Collingwood & Prince, 1998: 10 (Portugal); Kim, Kim & Kim, 1998: 145 (Korea); Shattuck, 1999: 52, 179 (Australia genera synopsis); Andersen, 2000: 19 (northern Australia genera); Zhou, 2001b: 22 (China, Guangxi); Czechowski, Radchenko & Czechowska, 2002: 133 (Poland); Aktaç & Radchenko, 2002: 53 (Turkey genera); Yoshimura & Onoyama, 2002: 436 (Japan genera, males); Mackay & Mackay, 2002: 27 (U.S.A., New Mexico); Csösz, 2003: 147 (Carpathian Basin); Palacio & Fernández, in Fernández, 2003d: 256 (Neotropical genera); Radchenko, 2005b: 184 (North Korea); Coovert, 2005: 23 (U.S.A., Ohio); Yoshimura & Fisher, 2007: 27 (Malagasy genera, males); Clouse, 2007b: 190 (Micronesia); Seifert, 2007: 108 (North and Central Europe); Bolton & Fisher, 2008c: 32 (Afrotropical genera); Fernández & Arias-Penna, 2008: 32 (Neotropical genera); Dlussky, 2009: 1071 (Eocene amber); Terayama, 2009: 101 (Taiwan genera); Heterick, 2009: 34 (south-western Australia genera); Boer, 2010: 68 (Benelux).

Dolichoderinae Bothriomyrmecini Bothriomyrmex TRUE TRUE FALSE

34 species, 11 subspecies

Bothriomyrmex Emery, 1869c: 117. Type-species: Bothriomyrmex costae, by monotypy.

Taxonomic history

Bothriomyrmex in Formicinae: André, 1881b: 64 [Formicidae].

Bothriomyrmex in Dolichoderinae: Forel, 1878: 380 [Dolichoderidae]; Emery & Forel, 1879a: 454 [Dolichoderidae]; Dalla Torre, 1893: 170; Emery, 1895j: 771; Forel, 1895e: 469; Wheeler, W.M. 1910g: 142.

Bothriomyrmex in Dolichoderidae: Ashmead, 1905b: 384.

Bothriomyrmex in Dolichoderinae, Tapinomini: Emery, 1913a: 27; Forel, 1917: 248; Bondroit, 1918: 88; Wheeler, W.M. 1922a: 689; all subsequent authors to the following.

Bothriomyrmex in Dolichoderinae, Dolichoderini: Shattuck, 1992c: 54; Bolton, 1994: 26; Bolton, 2003: 84.

Bothriomyrmex in Dolichoderinae, Iridomyrmecini, Bothriomyrmecina: Dubovikov, 2005a: 92.

Bothriomyrmex in Dolichoderinae, Bothriomyrmecini: Ward, Brady, et al. 2010: 361.

Genus Bothriomyrmex references

Forel, 1878: 380 (diagnosis); André, 1882b: 218 (Europe & Algeria species); Dalla Torre, 1893: 170 (catalogue); Forel, 1895e: 469 (India & Sri Lanka species key); Bingham, 1903: 305 (India, Sri Lanka & Burma species key); Emery, 1913a: 27 (diagnosis, catalogue); Wheeler, W.M. 1922a: 690 (subgenera key); Emery, 1925e: 5 (European and Oriental species); Kratochvíl, 1941: 96 (Central Europe species key); Novák & Sadil, 1941: 96 (Central Europe species key); Chapman & Capco, 1951: 187 (Asia checklist); Bernard, 1967: 260 (diagnosis, Western Europe species key); Kutter, 1977c: 175 (Switzerland species key); Arnol'di & Dlussky, 1978: 548 (former European U.S.S.R. species key); Collingwood, 1978: 87 (Iberian Peninsula species key); Gösswald, 1985: 316 (Germany species key); Taylor & Brown, D.R. 1985: 92 (Australia catalogue); Taylor, 1987a: 8 (Australia checklist); Agosti & Collingwood, 1987b: 279 (Balkans species key); Shattuck, 1992c: 54 (diagnosis, review of genus); Shattuck, 1994: 31 (catalogue); Bolton, 1995a: 1048 (census); Bolton, 1995b: 80 (catalogue); Wu, J. & Wang, 1995: 119 (China species key); Seifert, 1996b: 163 (Central Europe species key); Shattuck, 1999: 66 (Australia synopsis); Seifert, 2007: 148 (North and Central Europe species key).

Dolichoderinae Dolichoderini Dolichoderus TRUE TRUE FALSE

Extant: 122 species, 21 subspecies

Fossil: 53 species

Dolichoderus Lund, 1831a: 130. Type-species: Formica attelaboides, by monotypy.

Taxonomic history

Dolichoderus in Formicidae: Smith, F. 1858b: 75.

Dolichoderus in Formicinae: Mayr, 1862: 652 (in key) [Formicidae]; Mayr, 1865: 9 [Formicidae].

Dolichoderus in Dolichoderidae: Ashmead, 1905b: 384.

Dolichoderus in Dolichoderinae: Forel, 1878: 386 [Dolichoderidae]; Emery & Forel, 1879a: 455 [Dolichoderidae]; Dalla Torre, 1893: 156; Forel, 1895b: 107 [Dolichoderidae]; Emery, 1895j: 771; Forel, 1895e: 462; Forel, 1899c: 98; Wheeler, W.M. 1910g: 142.

Dolichoderus in Dolichoderinae, Dolichoderini: Emery, 1913a: 7; Wheeler, W.M. 1915h: 77; Forel, 1917: 247; Wheeler, W.M. 1922a: 688; all subsequent authors.

Junior synonyms of DOLICHODERUS

Hypoclinea Mayr, 1855: 377 (attributed to Foerster). Type-species: Formica quadripunctata, by subsequent designation of Wheeler, W.M. 1911f: 165.

Taxonomic history

Hypoclinea in Formicidae: Smith, F. 1858b: 57.

Hypoclinea in Formicinae: Mayr, 1855: 377 [Formicidae]; Mayr, 1861: 40 [Formicidae]; Mayr, 1862: 652 (in key) [Formicidae]; Mayr, 1865: 9 [Formicidae]; Mayr, 1868c: 53 [Formicidae]; André, 1874: 168 [Formicidae].

Hypoclinea in Dolichoderidae: Ashmead, 1905b: 384.

Hypoclinea in Dolichoderinae, Dolichoderini: Emery, 1913a: 18; Forel, 1917: 247; all subsequent authors.

Hypoclinea as genus: Mayr, 1855: 377; Mayr, 1865: 9; Mayr, 1868c: 53; Mayr, 1870b: 953; Smith, 1871a: 310; Ashmead, 1905b: 384; Kempf, 1972a: 118; Lattke, 1987b: 259; Dlussky & Fedoseeva, 1988: 78; Mackay & Vinson, 1989b: 17; Hölldobler & Wilson, 1990: 17; Wu, J. & Wang, 1995: 120.

Hypoclinea as subgenus of Dolichoderus: Emery, 1894c: 234; Wheeler, W.M. 1910g: 142; Emery, 1913a: 10; Wheeler, W.M. 1915h: 77; Forel, 1917: 247; Borgmeier, 1923: 84; Clark, 1930b: 252; McAreavey, 1949: 17; Creighton, 1950a: 331; Chapman & Capco, 1951: 182; Kusnezov, 1956: 28; Kusnezov, 1959: 50; Smith, D.R. 1979: 1415; Kupyanskaya, 1990: 154.

Hypoclinea as junior synonym of Dolichoderus: Forel, 1878: 386; Emery & Forel, 1879a: 455; Dalla Torre, 1893: 156; Bingham, 1903: 291; Brown, 1973b: 181 [provisional]; Shattuck, 1992c: 66; Mackay, 1993b: 6; Shattuck, 1994: 39; Bolton, 1994: 26; Bolton, 1995b: 31; Bolton, 2003: 85.

Monacis Roger, 1862a: 233. Type-species: Formica bispinosa, by subsequent designation of Wheeler, W.M. 1911f: 167.

Taxonomic history

Monacis in Dolichoderidae: Ashmead, 1905b: 384.

Monacis in Dolichoderinae, Dolichoderini: Emery, 1913a: 9; Forel, 1917: 247; all subsequent authors.

Monacis as genus: Roger, 1862a: 233; Ashmead, 1905b: 384; Brown, 1950e: 249; Kempf, 1959b: 227; Kempf, 1972a: 141; Kempf, 1972c: 253; Dlussky & Fedoseeva, 1988: 78; Mackay & Vinson, 1989b: 20; Hölldobler & Wilson, 1990: 17.

Monacis as subgenus of Dolichoderus: Emery, 1894c: 228; Wheeler, W.M. 1910g: 142; Emery, 1913a: 9; Forel, 1917: 247; Wheeler, W.M. 1922a: 688; Borgmeier, 1923: 81; Kusnezov, 1956: 27; Kusnezov, 1959: 50.

Monacis as junior synonym of Hypoclinea: Mayr, 1862: 704.

Monacis as junior synonym of Dolichoderus: Forel, 1878: 386; Emery & Forel, 1879a: 455; Dalla Torre, 1893: 156; Brown, 1973b: 182 [provisional]; Shattuck, 1992c: 66; Mackay, 1993b: 6; Shattuck, 1994: 39; Bolton, 1994: 26; Bolton, 1995b: 35; Bolton, 2003: 85.

Monoceratoclinea Wheeler, W.M. 1935c: 68 [as subgenus of Dolichoderus]. Type-species: Dolichoderus (Hypoclinea) monoceros, by original designation.

Taxonomic history

Monoceratoclinea as subgenus of Dolichoderus: Wheeler, W.M. 1935c: 68.

Monoceratoclinea as genus: Brown, 1950e: 249; Dlussky & Fedoseeva, 1988: 78; Hölldobler & Wilson, 1990: 17.

Monoceratoclinea as junior synonym of Dolichoderus: Shattuck, 1992c: 66; Shattuck, 1994: 40; Bolton, 1994: 26; Bolton, 1995b: 35; Bolton, 2003: 85.

Diceratoclinea Wheeler, W.M. 1935c: 69 [as subgenus of Dolichoderus]. Type-species: Dolichoderus scabridus, by original designation.

Taxonomic history

Diceratoclinea as subgenus of Dolichoderus: Wheeler, W.M. 1935c: 69; Chapman & Capco, 1951: 181.

Diceratoclinea as genus: Brown, 1950e: 249.

Diceratoclinea as junior synonym of Dolichoderus: Brown, 1973b: 180 [provisional]; Taylor & Brown, D.R. 1985: 93; Shattuck, 1992c: 66; Shattuck, 1994: 40; Bolton, 1994: 26; Bolton, 1995b: 26; Bolton, 2003: 85.

Acanthoclinea Wheeler, W.M. 1935c: 69 [as subgenus of Dolichoderus]. Type-species: Dolichoderus doriae, by original designation.

Taxonomic history

Acanthoclinea in Dolichoderinae, Dolichoderini: Donisthorpe, 1943f: 618.

Acanthoclinea as subgenus of Dolichoderus: Wheeler, W.M. 1935c: 69.

Acanthoclinea as genus: Brown, 1950e: 249.

Acanthoclinea as junior synonym of Dolichoderus: Taylor & Brown, D.R. 1985: 93; Shattuck, 1992c: 66; Shattuck, 1994: 40; Bolton, 1994: 26; Bolton, 1995b: 18; Bolton, 2003: 85.

Karawajewella Donisthorpe, 1944b: 59.

Taxonomic history

[Replacement name for Diabolus Karavaiev, 1926d: 424; junior homonym of Diabolus Gray, J.E. 1841: 400 (Mammalia).]

Karawajewella as junior synonym of Dolichoderus: Brown, 1973b: 181 [provisional]; Shattuck, 1992c: 66; Shattuck, 1994: 39, 40; Bolton, 1994: 26; Bolton, 1995b: 32; Bolton, 2003: 85.

Homonym replaced by Karawajewella

Diabolus Karavaiev, 1926d: 424 [as subgenus of Dolichoderus]. Type-species: Dolichoderus (Diabolus) bifurcatus (junior synonym of Dolichoderus cuspidatus), by original designation.

Taxonomic history

[Diabolus Karavaiev junior homonym of Diabolus Gray, J.E. 1841: 400 (Mammalia).]

Diabolus as subgenus of Dolichoderus: Wheeler, W.M. 1935c: 68.

Genus Dolichoderus references

Roger, 1863b: 10, 14, 15 (Dolichoderus, Hypoclinea, Monacis catalogues); Mayr, 1863: 407, 423 (Dolichoderus, Hypoclinea catalogues); Mayr, 1865: 9 (Dolichoderus, Hypoclinea diagnoses); Mayr, 1867a: 75 (Hypoclinea diagnosis); Mayr, 1868c: 54 (†Baltic Amber species key); Mayr, 1870b: 955 (Hypoclinea species key); Mayr, 1876: 79 (Australia species key); André, 1882b: 225 (Europe & Algeria species); Mayr, 1886d: 435 (U.S.A. species key); Cresson, 1887: 258 (U.S.A. catalogue); Dalla Torre, 1893: 156 (catalogue); Forel, 1895e: 462 (India & Sri Lanka species key); Bingham, 1903: 292 (India, Sri Lanka & Burma species key); Wheeler, W.M. 1905d: 306 (North America species key); Emery, 1913a: 7 (diagnosis, subgenera key, catalogue); Emery, 1913a: 9, 10 (D. (Monacis), D. (Hypoclinea), catalogues); Mann, 1916: 460 (Brazil D. (Dolichoderus) species key); Wheeler, W.M. 1922a: 688 (subgenera key); Clark, 1930b: 253 (Australia D. (Hypoclinea) species key); Creighton, 1950a: 333 (North America D. (Hypoclinea) species key); Chapman & Capco, 1951: 181 (Asia checklist); Kempf, 1959b: 230 (Monacis, all species revision, key); Wheeler, J. 1963: 149 (U.S.A., North Dakota species key); Bernard, 1967: 249 (diagnosis); Kempf, 1969: 292 (Neotropical D. (Dolichoderus) species key); Kempf, 1972a: 98, 118, 141 (Neotropical Dolichoderus, Hypoclinea, Monacis, catalogues); Kempf, 1972c: 253 (Monacis species, additions to 1959a key); Smith, D.R. 1979: 1415 (North America catalogue); Taylor & Brown, D.R. 1985: 93 (Australia catalogue); Taylor, 1987a: 24 (Australia, New Caledonia checklist); Harada, 1987: 602 (Monacis species key); Lattke, 1987b: 263 (Neotropical D. bispinosus group, key); Dlussky & Fedoseeva, 1988: 78 (synoptic classification); Johnson, 1989a: 5 (U.S.A. species key); Hölldobler & Wilson, 1990: 17 (synoptic classification); Brandão, 1991: 340, 347, 356 (Neotropical catalogue); Shattuck, 1992c: 66 (diagnosis, review of genus); Mackay, 1993b: 17 (New World, all species revision, key); Radchenko, 1994b: 112 (South Siberia species key); Shattuck, 1994: 39 (catalogue); Bolton, 1995a: 1049 (census); Bolton, 1995b: 172 (catalogue); Wu, J. & Wang, 1995: 120 (China species key); Xu, 1995a: 34 (China species key); Shattuck, 1999: 68 (Australia synopsis); Xu, 2001b: 356 (China species key); Zhou, 2001b: 153 (China, Guangxi species key); Dill, 2002: 21, 30 (southeast Asia species groups key, D. cuspidatus group revision, key); Dlussky & Rasnitsyn, 2002: 416 (diagnosis for impression fossils); Dlussky, 2002b: 54 (Baltic & Rovno amber species); Dlussky, 2008b: 509 (Europe Eocene amber species key); Heterick, 2009: 45 (south-western Australia species key).

Dolichoderinae Tapinomini Liometopum TRUE TRUE FALSE

Extant: 7 species

Fossil: 9 species

Liometopum Mayr, 1861: 38. Type-species: Formica microcephala, by monotypy.

Taxonomic history

Liometopum in Formicinae: Mayr, 1861: 38 [Formicidae]; Mayr, 1862: 653 (in key) [Formicidae]; Mayr, 1865: 9 [Formicidae]; André, 1874: 168 [Formicidae].

Liometopum in Dolichoderinae: Forel, 1878: 383 [Dolichoderidae]; Emery & Forel, 1879a: 454 [Dolichoderidae]; Dalla Torre, 1893: 163; Emery, 1895j: 771; Forel, 1899c: 104; Wheeler, W.M. 1910g: 142.

Liometopum in Dolichoderidae: Ashmead, 1905b: 384.

Liometopum in Dolichoderinae, Tapinomini: Emery, 1913a: 19; Wheeler, W.M. 1915h: 95; Forel, 1917: 248; Bondroit, 1918: 87; Wheeler, W.M. 1922a: 688; Carpenter, 1930: 46; subsequent authors except those below; then Jaffe, 1993: 9; Ward, Brady, et al. 2010: 361.

Liometopum in Dolichoderinae, Liometopini: Dlussky & Fedoseeva, 1988: 77.

Liometopum in Dolichoderinae, Dolichoderini: Shattuck, 1992c: 121; Bolton, 1994: 26; Bolton, 2003: 89.

Genus Liometopum references

Roger, 1863b: 14 (catalogue); Mayr, 1863: 427 (catalogue); Mayr, 1865: 9 (diagnosis); Forel, 1878: 383 (diagnosis); André, 1882b: 220 (Europe & Algeria species); Cresson, 1887: 258 (U.S.A. catalogue); Dalla Torre, 1893: 162 (catalogue); Bingham, 1903: 289 (diagnosis); Wheeler, W.M. 1905e: 322 (North America species); Emery, 1913a: 19 (diagnosis, catalogue); Creighton, 1950a: 338 (North America species key); Chapman & Capco, 1951: 192 (Asia checklist); Gregg, 1963: 438 (U.S.A., Colorado species key); Bernard, 1967: 253 (diagnosis); Smith, D.R. 1979: 1417 (North America catalogue); Wheeler, G.C. & Wheeler, J. 1986g: 55 (U.S.A., Nevada species key); Shattuck, 1992c: 121 (diagnosis, review of genus); Shattuck, 1994: 128 (catalogue); Bolton, 1995a: 1050 (census); Bolton, 1995b: 247 (catalogue); Mackay & Mackay, 2002: 232 (U.S.A., New Mexico species key); Del Toro, Pacheco & Mackay, 2009: 306 (all species revision, key).

Dolichoderinae Tapinomini Tapinoma TRUE TRUE FALSE

Extant: 64 species, 26 subspecies

Fossil: 4 species

Tapinoma Foerster, 1850a: 43. Type-species: Tapinoma collina (junior synonym of Formica erratica), by monotypy.

Taxonomic history

Tapinoma in Formicidae: Smith, F. 1858b: 55.

Tapinoma in Formicinae: Mayr, 1855: 372 [Formicidae]; Mayr, 1861: 41 [Formicidae]; Mayr, 1862: 653 (in key) [Formicidae]; Mayr, 1865: 10 [Formicidae]; André, 1874: 168 [Formicidae]; André, 1881b: 63 [Formicidae].

Tapinoma in Dolichoderinae: Forel, 1878: 385 [Dolichoderidae]; Emery & Forel, 1879a: 454 [Dolichoderidae]; Forel, 1895b: 109 [Dolichoderidae]; Dalla Torre, 1893: 164; Emery, 1895j: 771; Forel, 1899c: 101; Wheeler, W.M. 1910g: 142.

Tapinoma in Dolichoderidae: Ashmead, 1905b: 384.

Tapinoma in Dolichoderinae, Tapinomini: Emery, 1913a: 38; Arnold, 1915: 152; Forel, 1917: 248; Bondroit, 1918: 89; Wheeler, W.M. 1922a: 690; subsequent authors except those below; then Jaffe, 1993: 9; Dubovikov, 2005a: 91 (implied); Ward, Brady, et al. 2010: 361.

Tapinoma in Dolichoderinae, Dolichoderini: Shattuck, 1992c: 146; Bolton, 1994: 26; Bolton, 2003: 91.

Junior synonyms of TAPINOMA

Micromyrma Dufour, 1857: 60. Type-species: Micromyrma pygmaea, by monotypy.

Taxonomic history

[Type-species not Formica erratica, unjustified subsequent designation by Wheeler, W.M. 1911f: 167.]

Micromyrma as subgenus of Tapinoma: Menozzi, 1925a: 19; Santschi, 1928e: 472; Chapman & Capco, 1951: 192.

Micromyrma as junior synonym of Tapinoma: Mayr, 1863: 455; Forel, 1878: 385; Emery & Forel, 1879a: 454; Dalla Torre, 1893: 164; Emery, 1913a: 38; Wheeler, W.M. 1922a: 923; Smith, M.R. 1951a: 837; Shattuck, 1992c: 146; Shattuck, 1994: 140; Bolton, 1994: 26; Bolton, 2003: 91; Fisher & Bolton, 2007: 65.

Semonius Forel, 1910f: 21. Type-species: Semonius schultzei, by monotypy.

Taxonomic history

Semonius in Dolichoderinae: Wheeler, W.M. 1910g: 142.

Semonius in Dolichoderinae, Tapinomini: Emery, 1913a: 44; Arnold, 1915: 157; Forel, 1917: 247; Wheeler, W.M. 1922a: 688; all subsequent authors.

Semonius as junior synonym of Tapinoma: Shattuck, 1992c: 146; Shattuck, 1994: 140; Bolton, 1994: 26; Bolton, 2003: 91; Fisher & Bolton, 2007: 66.

Pseudaphomomyrmex Wheeler, W.M. 1920: 53. Type-species: Aphomomyrmex emeryi, by original designation.

Taxonomic history

Pseudaphomomyrmex in Formicinae, Dimorphomyrmecini: Donisthorpe, 1943g: 721.

Pseudaphomomyrmex incertae sedis in Formicinae: Dlussky & Fedoseeva, 1988: 77.

Pseudaphomomyrmex in Formicinae, Myrmelachistini: Hölldobler & Wilson, 1990: 18.

Pseudaphomomyrmex in Formicinae, Brachymyrmecini: Bolton, 1994: 50.

Pseudaphomomyrmex in Formicinae, Plagiolepidini: Wheeler, W.M. 1922a: 695; Chapman & Capco, 1951: 214; Wheeler, G.C. & Wheeler, J. 1985: 258; Bolton, 2003: 106.

Pseudaphomomyrmex in Dolichoderinae: LaPolla & Longino, 2006: 305.

Pseudaphomomyrmex as junior synonym of Aphomomyrmex: Emery, 1925b: 44; Donisthorpe, 1943g: 721.

Pseudaphomomyrmex as genus: Wheeler, W.M. 1920: 53; Chapman & Capco, 1951: 214; Bolton, 1994: 50; Bolton, 1995b: 369; Bolton, 2003: 106.

Pseudaphomomyrmex as junior synonym of Tapinoma: Fisher & Bolton, 2007: 66.

Zatapinoma Wheeler, W.M. 1928a: 20. Type-species: Zatapinoma annandalei, by original designation.

Taxonomic history

Zatapinoma in Dolichoderinae, Tapinomini: Donisthorpe, 1943g: 737; all subsequent authors.

Zatapinoma as junior synonym of Tapinoma: Shattuck, 1992c: 146; Shattuck, 1994: 140; Bolton, 1994: 26; Bolton, 2003: 91; Fisher & Bolton, 2007: 66.

Neoclystopsenella Kurian, 1955: 133. Type-species: Neoclystopsenella luffae, by monotypy.

Taxonomic history

Neoclystopsenella in Bethylidae: Kurian, 1955: 133.

Neoclystopsenella in Dolichoderinae: Brown, 1988a: 337.

Neoclystopsenella as junior synonym of Tapinoma: Brown, 1988a: 337; Shattuck, 1992c: 146; Shattuck, 1994: 140; Bolton, 1994: 26; Bolton, 2003: 91; Fisher & Bolton, 2007: 66.

Genus Tapinoma references

Mayr, 1855: 372 (diagnosis); Smith, F. 1858b: 55 (diagnosis); Roger, 1863b: 13, 14 (Tapinoma, Micromyrma catalogues); Mayr, 1863: 455 (catalogue); Mayr, 1865: 10 (diagnosis); Mayr, 1867a: 78 (diagnosis); André, 1882b: 222 (Europe & Algeria species key); Cresson, 1887: 258 (U.S.A. catalogue); Dalla Torre, 1893: 164 (catalogue); Bingham, 1903: 304 (India, Sri Lanka & Burma species key); Emery, 1913a: 38 (diagnosis, catalogue); Emery, 1913a: 44 (Semonius diagnosis, catalogue); Arnold, 1915: 152, 157 (diagnosis, South Africa species key, Semonius diagnosis); Wheeler, W.M. 1922a: 695 (Pseudaphomomyrmex diagnosis, in key). Wheeler, W.M. 1922a: 923, 927 (Afrotropical Tapinoma, Semonius catalogues); Wheeler, W.M. 1922a: 1034 (Malagasy catalogue); Emery, 1925d: 63 (Palaearctic species key); Kuznetsov-Ugamsky, 1927d: 33 (Turkestan species key); Smith, M.R. 1928a: 311 (North America species key); Kratochvíl, Novák & Snoflák, 1944: 79 (Czechoslovakia species key); Creighton, 1950a: 352 (North America species key); Chapman & Capco, 1951: 192, 196 (Asia Semonius, Tapinoma, Zatapinoma checklists); Bernard, 1967: 254 (diagnosis, Western Europe species key); Boven, 1970b: 24 (Netherlands species key); Kempf, 1972a: 246 (Neotropical catalogue); Tarbinsky, 1976: 118 (Kyrghyzstan species key); Kutter, 1977c: 178 (Switzerland species key); Arnol'di & Dlussky, 1978: 547 (former European U.S.S.R. species key); Collingwood, 1978: 87 (Iberian Peninsula species key); Smith, D.R. 1979: 1421 (North America catalogue); Schembri & Collingwood, 1981: 421 (Malta species key); Gösswald, 1985: 317 (Germany species key); Taylor & Brown, D.R. 1985: 106 (Australia catalogue); Agosti & Collingwood, 1987b: 279 (Balkans species key); Taylor, 1987a: 77 (Australia, New Caledonia & New Zealand checklist); Dlussky, Soyunov & Zabelin, 1990: 168 (Turkmenistan species key); Brandão, 1991: 380 (Neotropical catalogue); Shattuck, 1992c: 146 (diagnosis, review of genus); Arakelian, 1994: 72 (Armenia species key); Shattuck, 1994: 140 (catalogue); Bolton, 1995a: 1053 (census); Bolton, 1995b: 399 (catalogue); Wu, J. & Wang, 1995: 116 (China species key); Collingwood & Agosti, 1996: 360 (Saudi Arabia species key); Seifert, 1996b: 165 (Central Europe species key); Skinner & Allen, 1996: 40 (Britain species key); Collingwood & Prince, 1998: 20 (Portugal species key); Shattuck, 1999: 81 (Australia synopsis); Zhou, 2001b: 151 (China, Guangxi species key); Czechowski, Radchenko & Czechowska, 2002: 135 (Poland species key); Seifert, 2007: 149 (North and Central Europe species key); Heterick, 2009: 58 (south-western Australia species key); Yoshimura & Fisher, 2011: 13 (Malagasy males).

Formicinae Lasiini Lasius TRUE TRUE FALSE

Extant: 100 species, 2 subspecies

Fossil: 22 species, 1 subspecies

Lasius Fabricius, 1804: 415. Type-species: Formica nigra, by subsequent designation of Bingham, 1903: 338.

Taxonomic history

[Lasius Jurine, 1801: 164 (Apidae), resurrected by Morice & Durrant, 1915: 421, later suppressed by Act of Commission.]

Lasius in Formicinae: Mayr, 1861: 49 [Formicidae]; Mayr, 1862: 652 (in key) [Formicidae]; Mayr, 1865: 8 [Formicidae]; Mayr, 1868c: 42 [Formicidae].

Lasius in Camponotinae: Forel, 1878: 374 [Camponotidae]; Emery & Forel, 1879a: 452 [Camponotidae]; Forel, 1886f: 206; Dalla Torre, 1893: 181.

Lasius in Camponotinae, Camponotini: Forel, 1886f: 206.

Lasius in Camponotinae, Formicini: Forel, 1893a: 165; Forel, 1895b: 103; Emery, 1895j: 772; Forel, 1899c: 127; Wheeler, W.M. 1910g: 143; Forel, 1912i: 89; Wheeler, W.M. 1915h: 120.

Lasius in Formicinae, Formicini: Wheeler, W.M. 1922a: 698; Carpenter, 1930: 58; Chapman & Capco, 1951: 201; Wheeler, G.C. & Wheeler, J. 1970: 651; Wheeler, G.C. & Wheeler, J. 1976b: 101; Wheeler, G.C. & Wheeler, J. 1985: 258.

Lasius in Formicinae, Acanthomyopsini: Donisthorpe, 1943f: 655.

Lasius in Formicinae, Lasius genus group: Agosti, 1991: 296.

Lasius in Formicinae, Lasiini: Ashmead, 1905b: 384; Emery, 1925b: 226; all subsequent authors except the entries above; Bolton, 1994: 50; Bolton, 2003: 22, 96.

Lasius as junior synonym of Acanthomyops: Forel, 1916: 460; Donisthorpe, 1916c: 276; Morice & Durrant, 1917: 442. Donisthorpe, 1927b: 206; Donisthorpe, 1937d: 132; Donisthorpe, 1943f: 655; Donisthorpe, 1946g: 91.

Lasius as junior synonym of Formicina: Emery, 1916a: 61; Emery, 1916b: 239; Emery, 1917a: 96; Wheeler, W.M. 1916k: 170; Bondroit, 1918: 19.

Lasius as genus: Fabricius, 1804: 415; Wheeler, W.M. 1916k: 172; Emery, 1925b: 226; Wilson, 1955a: 11; all subsequent authors.

Junior synonyms of LASIUS

Donisthorpea Morice & Durrant, 1915: 423. Type-species: Formica nigra, by original designation.

Taxonomic history

[Unnecessary replacement name for Lasius Fabricius.]

Donisthorpea in Formicinae, Acanthomyopsini: Donisthorpe, 1943f: 640; Donisthorpe, 1946g: 91.

Donisthorpea as genus: Morice & Durrant, 1915: 423; Donisthorpe, 1915d: 184.

Donisthorpea as subgenus of Formicina: Emery, 1916a: 11; Emery, 1916b: 240.

Donisthorpea as subgenus of Acanthomyops: Donisthorpe, 1927b: 229; Donisthorpe, 1943f: 640; Donisthorpe, 1946g: 91.

Donisthorpea as junior synonym of Acanthomyops: Forel, 1916: 460; Morice & Durrant, 1917: 442; Donisthorpe, 1927b: 209.

Donisthorpea as junior synonym of Lasius: Wheeler, W.M. 1916k: 172; Emery, 1925b: 226; Wilson, 1955a: 11. [Lasius and Donisthorpea share the same type-species; synonymy is therefore absolute.]

†Tylolasius Zhang, 1989: 295. Type-species: †Tylolasius inflatus, by original designation.

Taxonomic history

†Tylolasius incertae sedis in Formicinae: Bolton, 2003: 28, 131.

†Tylolasius as junior synonym of Lasius: Hong, Wu & Ren, 2001: 348.

Subgenera of LASIUS include the nominal plus the following.

Subgenus LASIUS (ACANTHOMYOPS)

Acanthomyops Mayr, 1862: 652 (diagnosis in key), 699. Type-species: Formica clavigera, by monotypy.

Taxonomic history

Acanthomyops in Formicinae: Mayr, 1862: 652 (in key) [Formicidae]; Mayr, 1865: 8 [Formicidae].

Acanthomyops in Camponotinae, Formicini: Wheeler, W.M. 1910g: 143; Forel, 1917: 249.

Acanthomyops in Formicinae, Formicini: Wheeler, W.M. 1922a: 698; Wheeler, G.C. & Wheeler, J. 1970: 651; Wheeler, G.C. & Wheeler, J. 1976b: 101; Wheeler, G.C. & Wheeler, J. 1985: 258; Jaffe, 1993: 14.

Acanthomyops in Formicinae, Acanthomyopsini: Donisthorpe, 1943f: 618.

Acanthomyops in Formicinae, Lasius genus group: Agosti, 1991: 296.

Acanthomyops in Formicinae, Lasiini: Ashmead, 1905b: 384 (misspelled as Acanthomyrmex); Emery, 1925b: 236; all subsequent authors except for those above.

Acanthomyops as junior synonym of Lasius: Mayr, 1866b: 888; Dalla Torre, 1893: 181.

Acanthomyops as subgenus of Formicina: Emery, 1916a: 11; Forel, 1917: 249.

Acanthomyops as genus: Mayr, 1862: 699; Donisthorpe, 1927b: 209; Donisthorpe, 1943f: 618; Creighton, 1950a: 426; Smith, M.R. 1958c: 150; Wing, 1968: 47; Bolton, 1994: 50; Bolton, 2003: 94.

Acanthomyops as subgenus of Lasius: Emery, 1893i: 642; Wheeler, W.M. 1910g: 143; Wheeler, W.M. 1916k: 172; Wheeler, W.M. 1922a: 698; Emery, 1925b: 236; Buren, 1950: 188; Smith, M.R. 1951a: 852; Gregg, 1963: 453; Ward, 2005: 13.

Subgenus LASIUS (DENDROLASIUS)

Dendrolasius Ruzsky, 1912: 630 [as subgenus of Lasius]. Type-species: Formica fuliginosa, by monotypy.

Taxonomic history

[Dendrolasius also described as new by Ruzsky, 1914a: 59.]

Dendrolasius as subgenus of Formicina: Forel, 1917: 249.

Dendrolasius as subgenus of Acanthomyops: Donisthorpe, 1927b: 211; Donisthorpe, 1943f: 637.

Dendrolasius as subgenus of Lasius: Ruzsky, 1912: 630; Wheeler, W.M. 1922a: 698; Emery, 1925b: 235; Wilson, 1955a: 14; all subsequent authors except the following.

Dendrolasius as junior synonym of Lasius: Brown, 1973b: 180 [provisional].

Subgenus LASIUS (CHTHONOLASIUS)

Chtonolasius Ruzsky, 1912: 630 [as subgenus of Lasius]. Type-species: Formica umbrata, by subsequent designation of Emery, 1925b: 232.

Taxonomic history

[Chtonolasius also described as new by Ruzsky, 1914a: 59. The claim by some authors (e.g. Emery, 1916a: 11; Donisthorpe, 1941f: 37; Donisthorpe, 1943f: 633) that Ruzsky nominated Lasius flavus as type-species is incorrect. Ruzsky (1912: 630) included those species "with the appearance of the yellow Lasius (Las. flavus D.G.) and its closely related species, and races (Las. carniolicus, Las. umbratus, Las. myops, Las. mixtus)". This is not a direct nomination of a type-species, nor is De Geer (= D.G.) the author of flavus: Bolton, 1995b: 24.]

Chthonolasius justified emendation of spelling: Wheeler, W.M. 1916k: 170.

Chthonolasius as junior synonym of Formicina: Wheeler, W.M. 1916k: 172; Emery, 1916a: 11.

Chthonolasius as subgenus of Acanthomyops: Donisthorpe, 1927b: 254.

Chthonolasius as subgenus of Lasius: Ruzsky, 1912: 630; Ruzsky, 1914a: 59; Wheeler, W.M. 1922a: 698; Emery, 1925b: 231; Wilson, 1955a: 13; all subsequent authors except the following.

Chthonolasius as junior synonym of Lasius: Brown, 1973b: 179 [provisional].

Subgenus LASIUS (CAUTOLASIUS)

Cautolasius Wilson, 1955a: 13 [as subgenus of Lasius]. Type-species: Formica flava, by original designation.

Taxonomic history

Cautolasius as junior synonym of Lasius: Brown, 1973b: 179 [provisional].

Cautolasius as subgenus of Lasius: Bolton, 2003: 97.

Subgenus LASIUS (AUSTROLASIUS)

Austrolasius Faber, 1967: 74 [as subgenus of Lasius]. Type-species: Lasius carniolicus, by original designation.

Taxonomic history

Austrolasius as junior synonym of Lasius: Brown, 1973b: 179 [provisional].

Autrolasius as subgenus of Lasius: Bolton, 2003: 97.

Genus Lasius references

Mayr, 1861: 49 (Europe species key); Roger, 1863b: 11 (catalogue); Mayr, 1863: 394, 425 (catalogue); Mayr, 1865: 8 (diagnosis); Mayr, 1867a: 74 (diagnosis); André, 1874: 179 (Europe species key); Forel, 1874: 46 (Switzerland species key); Forel, 1878: 374 (diagnosis); André, 1882b: 191 (Europe & Algeria species key); Cresson, 1887: 257 (U.S.A. catalogue); Provancher, 1887: 236 (Canada species key); Nasonov, 1889: 64 (Russia species key); Dalla Torre, 1893: 181 (catalogue); Emery, 1893i: 637 (North America species key); Bingham, 1903: 339 (India species key); Ruzsky, 1905b: 264 (Russian Empire species key); Wasmann, 1906: 10 (Luxemburg species key); Bondroit, 1910: 484 (Belgium species key); Wheeler, W.M. 1910e: 237 (North America L. umbratus forms, key); Stitz, 1914: 82 (Central Europe species key); Donisthorpe, 1915d: 188 (Britain species key); Emery, 1916b: 243 (Italy species key); Wheeler, W.M. 1916m: 591 (U.S.A., Connecticut species key); Bondroit, 1918: 19 (France & Belgium species key); Wheeler, W.M. 1922a: 698 (subgenera key); Emery, 1925b: 226 (diagnosis, subgenera key, catalogue); Emery, 1925b: 231 (L. (Chthonolasius) diagnosis, catalogue); Emery, 1925b: 235 (L. (Dendrolasius) diagnosis, catalogue); Emery, 1925b: 236 (L. (Acanthomyops) diagnosis, catalogue); Kuznetsov-Ugamsky, 1927e: 186 (Turkestan species key); Karavaiev, 1927c: 277 (Ukraine species key); Donisthorpe, 1927b: 206 (Britain species key); Arnol'di, 1933b: 602 (Russia species key); Karavaiev, 1936: 193 (Ukraine species key); Stärcke, 1937: 57 (Europe L. umbratus group, key); Menozzi, 1939a: 313 (Himalaya & Tibet species key); Stitz, 1939: 267 (Germany species key); Kratochvíl, 1941: 100 (Central Europe species key); Novák & Sadil, 1941: 100 (Central Europe species key); Cole, 1942: 373 (U.S.A., Utah species key); Holgersen, 1943b: 174 (Norway species key); Holgersen, 1944: 199 (Norway species key); Buren, 1944a: 296 (U.S.A., Iowa species key); Stärcke, 1944a: 157 (Europe L. niger group, key); Boven, 1947: 184 (Belgium species key); Creighton, 1950a: 418 (North America Lasius species key); Creighton, 1950a: 428 (North America Acanthomyops species key); Buren, 1950: 188 (Acanthomyops queens, key); Chapman & Capco, 1951: 201 (Asia checklist); Wilson, 1955a: 13 (subgenera review); Wilson, 1955a: 26 (Nearctic Lasius species revision, key); Wilson, 1955a: 28 (Palaearctic species revision, key); Boven, 1959: 11 (Netherlands species key); Wheeler, G.C. & Wheeler, J. 1963: 177 (U.S.A., North Dakota Lasius species key); Wheeler, G.C. & Wheeler, J. 1963: 195 (U.S.A., North Dakota Acanthomyops species key); Gregg, 1963: 453 (U.S.A., Colorado species key); Collingwood, 1963: 155 (Europe L. umbratus group species key); Collingwood, 1964: 107 (Britain species key); Bernard, 1967: 349 (diagnosis, Western Europe species key); Wing, 1968: 53 (Acanthomyops diagnosis, all species revision, key); Bourne, 1973: 19 (Britain species key); Boven, 1970b: 28 (Netherlands species key); Bolton & Collingwood, 1975: 7 (Britain species key); Tarbinsky, 1976: 133 (Kyrghyzstan species key); Boven, 1977: 135 (Belgium species key); Kutter, 1977c: 208 (Switzerland species key); Arnol'di & Dlussky, 1978: 555 (former European U.S.S.R. species key); Collingwood, 1978: 89 (Iberian Peninsula species key); Collingwood, 1979: 92 (Fennoscandia & Denmark species key); Smith, D.R. 1979: 1435 (North America Lasius catalogue); Smith, D.R. 1979: 1440 (North America Acanthomyops catalogue); Yamauchi, 1979: 147 (Japan species keys); Allred, 1982: 444 (U.S.A., Utah Acanthomyops species key); Allred, 1982: 450 (U.S.A., Utah Lasius species key); Collingwood, 1982: 283 (Himalayan species key); Gösswald, 1985: 268 (Germany species key); Wheeler, G.C. & Wheeler, J. 1986g: 64 (U.S.A., Nevada Lasius species key); Wheeler, G.C. & Wheeler, J. 1986g: 68 (U.S.A., Nevada Acanthomyops species key); Nilsson & Douwes, 1987: 70 (Norway species key); Agosti & Collingwood, 1987b: 281 (Balkans species key); Dlussky & Fedoseeva, 1988: 77 (synoptic classification); Seifert, 1988a: 143 (Europe L. (Chthonolasius) species key); Kupyanskaya, 1989: 781 (East Palaearctic L. (Dendrolasius) species key); Dlussky, Soyunov & Zabelin, 1990: 158 (Turkmenistan species key); Seifert, 1990: 1 (supplement to European L. (Chthonolasius)); Agosti & Bolton, 1990b: 149 (characters); Kupyanskaya, 1990: 211 (Far Eastern Russia species key); Morisita, Kubota, Onoyama, et al., 1991: 24 (Japan species key); Agosti, 1991: 295 (Lasius genus group diagnosis); Seifert, 1992b: 48 (Palaearctic L. (Lasius) species key); Atanasov & Dlussky, 1992: 232 (Bulgaria species key); Arakelian, 1994: 111 (Armenia species key); Bolton, 1994: 50 (synoptic classification); Radchenko, 1994b: 114 (South Siberia species key); Bolton, 1995a: 1050 (census); Bolton, 1995b: 53 (Acanthomyops catalogue); Bolton, 1995b: 221 (Lasius catalogue); Douwes, 1995: 92 (Sweden species key); Kupyanskaya, 1995: 361 (Far Eastern Russia species key); Wu, J. & Wang, 1995: 153 (China species key); Seifert, 1996b: 179 (Central Europe species key); Skinner & Allen, 1996: 42 (Britain species key); Collingwood & Prince, 1998: 22 (Portugal species key); Zhou, 2001b: 187 (China, Guangxi species key); Blacker & Collingwood, 2002: 44 (British species key); Czechowski, Radchenko & Czechowska, 2002: 154 (Poland species key); Mackay & Mackay, 2002: 237 (U.S.A., New Mexico Acanthomyops species key); Mackay & Mackay, 2002: 331 (U.S.A., New Mexico Lasius species key); Radchenko, 2005a: 93 (East Palaearctic L. (Dendrolasius) species key); Radchenko, 2005b: 190 (North Korea species key); Seifert, 2007: 159 (North and Central Europe species key); Terayama, 2009: 207 (Taiwan species key).

Formicinae Plagiolepidini Brachymyrmex TRUE TRUE FALSE

41 species, 17 subspecies

Brachymyrmex Mayr, 1868b: 163. Type-species: Brachymyrmex patagonicus, by monotypy.

Taxonomic history

Brachymyrmex in Camponotinae: Forel, 1878: 375 [Camponotidae]; Dalla Torre, 1893: 174.

Brachymyrmex in Camponotinae, Plagiolepidini: Forel, 1893a: 165; Forel, 1895b: 106; Emery, 1895j: 771; Forel, 1899c: 123; Wheeler, W.M. 1910g: 143.

Brachymyrmex in Formicinae, Plagiolepidini: Ashmead, 1905b: 384.

Brachymyrmex in Camponotinae, Myrmelachistini: Forel, 1912i: 89.

Brachymyrmex in Formicinae, Dimorphomyrmecini: Emery, 1925b: 40 [subtribe Brachymyrmecini]; Donisthorpe, 1943f: 628.

Brachymyrmex incertae sedis in Formicinae: Dlussky & Fedoseeva, 1988: 77.

Brachymyrmex in Formicinae, Myrmelachistini: Wheeler, W.M. 1922a: 697; Hölldobler & Wilson, 1990: 18.

Brachymyrmex in Formicinae, Pseudolasius genus group: Agosti, 1991: 296.

Brachymyrmex in Formicinae, Brachymyrmecini: Wheeler, W.M. 1929a: 12; Kempf, 1972a: 37; Wheeler, G.C. & Wheeler, J. 1985: 258; Jaffe, 1993: 13; Bolton, 1994: 50.

Brachymyrmex in Formicinae, Plagiolepidini: Bolton, 2003: 23, 101.

Junior synonym of BRACHYMYRMEX

Bryscha Santschi, 1923b: 652 [as subgenus of Brachymyrmex]. Type-species: Brachymyrmex pilipes, by original designation.

Taxonomic history

Bryscha as junior synonym of Brachymyrmex: Brown, 1973b: 179 [provisional]; Smith, D.R. 1979: 1424; all subsequent authors.

Genus Brachymyrmex references

Forel, 1878: 375 (diagnosis); André, 1882b: 214 (Europe & Algeria species); Cresson, 1887: 255 (U.S.A. catalogue); Dalla Torre, 1893: 174 (catalogue); Wheeler, W.M. 1922a: 1036 (Malagasy catalogue); Santschi, 1923b: 652 (diagnosis, all species key); Emery, 1925b: 40 (diagnosis, catalogue); Emery, 1925b: 43 (B. (Bryscha) diagnosis, catalogue); Creighton, 1950a: 356 (North America species, review); Bernard, 1967: 279 (diagnosis); Kempf, 1972a: 37 (Neotropical catalogue); Alayo, 1974: 26 (Cuba species key); Smith, D.R. 1979: 1424 (North America catalogue); Brandão, 1991: 331 (catalogue); Bolton, 1995a: 1048 (census); Bolton, 1995b: 81 (catalogue); Quirán, et al. 2004: 275 (diagnosis).

Formicinae Plagiolepidini Lepisiota TRUE TRUE FALSE

81 species, 51 subspecies

Lepisiota Santschi, 1926a: 15 [as subgenus of Acantholepis]. Type-species: Plagiolepis rothneyi, by original designation.

Taxonomic history

[Lepisiota first available replacement name for Acantholepis Mayr, 1861 (junior homonym of Acantholepis Kroyer, 1846: 98 (Pisces)), hence valid name of genus: Bolton, 1995b: 33.]

Lepisiota in Form